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Lutites plicatus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 1713B70A-3C3C-4FCD-9827-BBBA6F989797 Fig. 55; Table 29 Diagnosis Species of Lutites gen. nov. with conch reaching 120 mm dm. Subadult stage with oval, compressed whorl profile (ww/wh=0.75) and rounded venter; about 12 radial plications per volution. Adult stage with subtrapezoidal, compressed whorl profile (ww/wh =0.72), weakly concave flanks, flattened venter and subangular ventrolateral shoulder; numerous weak and densely arranged plications and small ventrolateral nodes. Prongs of external lobe multiply serrated; altogether 20 notches of E, A and L lobes. Etymology From the Latin ‘ plicatus ’, meaning ‘corrugated’, referring to the dense ribbing. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 55A; MB.C.30018. Description Holotype MB.C.30018 is a specimen with 84 mm diameter, of which the last 120 degrees belong to the body chamber (Fig. 55A). Nearly one volution of the phragmocone can be studied; at this stage the whorl profile is oval and compressed with a rounded venter. The ornament consists of nearly 20 faint radial plications, which are coarsest near the umbilicus and in the ventrolateral area, where they form weak nodes. The ornament is weaker on the body chamber, where the plications are more numerous. The whorl profile is subtrapezoidal and compressed (ww/wh=0.72) in the adult stage with a rounded umbilical margin, weakly concave flanks and a flattened venter. The suture line of the holotype has a narrow external lobe with asymmetric, multiply serrated prongs (four and six small notches), an asymmetric and inflated ventrolateral saddle and multiply serrated adventive and lateral lobes (Fig. 55B). Both lobes show notches of various size. Remarks Lutites plicatus gen. et sp. nov. differs in the shape of the flattened, but convex venter from most of the other species of the genus (L. lyriformis gen. et sp. nov., L. profundus gen. et sp. nov., L. alius gen. et sp. nov.), which possess a completely flat or even concave venter. Stratigraphic range Upper part of the Hambast Formation; horizon unknown., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 85-87, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Esfahanites armatus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: DACEC5B2-4BE5-40D0-AA2B-578741DC349C Figs 45–46; Table 22 ? Paratirolites sp. Ghaedi et al., 2009: pl. 2 figs 1, 4. ? Paratirolites waageni – Zakharov & Mussavi Abnavi 2013: text-fig. 12c. Diagnosis Species of Esfahanites gen. nov. with conch reaching 130 mm dm. Subadult stage with strongly trapezoidal, moderately depressed whorl profile (ww/wh =1.50–1.70) and rounded to tectiform venter; about 12 coarse ventrolateral nodes per volution. Adult stage with trapezoidal and moderately depressed whorl profile (ww/wh =1.50–1.70), flattened tectiform venter and angular ventrolateral shoulder; coarse rursiradiate ribs. Prongs of external lobe trifid or multiply serrated; altogether 15–21 notches of E, A and L lobes. Etymology From the Latin ‘ armatus ’, meaning ‘armed’, because of the coarse spiny sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 45; MB.C.29964. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29963 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29965 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29966 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29967 to MB.C.29968. Description Holotype MB.C.29964 is the best-preserved specimen; it has 106 mm conch diameter and allows the study of two whorls (Fig. 45A). The diameter of the phragmocone is 68 mm and the body chamber has a length of about 270 degrees. Throughout the last two preserved volutions, the whorl profile is trapezoidal, but with the flanks less strongly diverging at the end of growth. The venter is broadly rounded in the penultimate whorl and becomes more flattened on the terminal body chamber. The sculpture on the phragmocone has 14 coarse conical ventrolateral nodes on the last volution; these nodes show a rib-like elongation towards the umbilicus. The sculpture on the body chamber shows variably coarse, rounded ribs on the flank; they turn backward near the umbilicus and are strengthened in the ventrolateral area. The suture line of holotype MB.C.29964 is remarkable because of its very small external lobe, which is wider at the base and is constricted in the upper part (Fig. 45B). The prongs are inclined ventrally and possess five little notches each. The ventrolateral saddle is small and inflated. The asymmetric adventive lobe is strongly serrated with little notches being developed on the ventral side up to half of the lobe depth. The lateral lobe is slightly less strongly serrated than the adventive lobe. Paratype MB.C.29966 is an incomplete but otherwise rather well-preserved individual of 101 mm conch diameter; it allows the study of two and a half whorls (Fig. 46A). The body chamber begins at 60 mm conch diameter, meaning that the preserved part of the body chamber has a length of 270 degrees and is nearly complete. Conch shape changes can be observed during the last whorl of the specimen. While the whorl profile is circular with rounded venter at the binning of the last whorl, it changes to tectiform and has, at the end of growth, an almost flat venter, separated from the flat and diverging flanks by an angular ventrolateral shoulder. The sculpture of the phragmocone part of the specimen shows coarse conical nodes in the ventrolateral area, 12 on the last volution and 10 on the penultimate volution. At the beginning of the terminal body chamber, these nodes are transformed into elongate nodes and finally rounded ribs, which turn from the umbilicus in backward direction. Finally, at the end of the body chamber these ribs become weaker and only shallow rounded ribs and weak ventrolateral nodes can be seen. The suture line of paratype MB.C.29966 shows a very small external lobe that has not half the depth of the adventive lobe (Fig. 46B). It is parallel-sided with parallel-sided prongs that possess three small notches. The ventrolateral saddle is inverted U-shaped with a much longer dorsal flank. The adventive lobe has seven small notches and the lateral lobe five, both lobes are asymmetric and separated by a low semi-circular lateral saddle. Remarks Esfahanites armatus gen. et sp. nov. differs in the very small external lobe from all the species of Paratirolites. It can therefore not be confused with any other species. Stratigraphic range Upper part of the Hambast Formation; the only in situ collected specimen comes from 2.00 m below the extinction horizon (Alibashites ferdowsii Zone).

Stoyanowites aspinosus Korn & Ghaderi in Korn et al., 2016 Fig. 62; Table 36 Stoyanowites aspinosus Korn & Ghaderi in Korn et al., 2016: 885, text-fig. 43. Diagnosis Species of Stoyanowites with conch reaching 100 mm dm. Subadult stage with oval, weakly compressed whorl profile (ww/wh =0.70) and rounded venter; 15 narrow and rounded ribs on the flanks. Adult stage with oval and weakly compressed whorl profile (ww/wh = 0.60–0.70) and moderately wide to wide umbilicus (uw/dm =0.40–0.50), rounded venter and rounded ventrolateral shoulder; very weak ribs on the flanks, coarsest in ventrolateral area. Prongs of external lobe bifid; 8–9 notches of E, A and L lobes. Material examined Holotype IRAN • West Azerbaijan, Aras Valley section; Paratirolites Limestone, 0.95 m below the top; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 43a); MB.C.25458. Additional material IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.300978 • 2 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.30098 to MB.C.30099 • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30100 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30101. Description Specimen MB.C.30100 is a fragmentary specimen with 72 mm diameter and shows the wide umbilicus (uw/dm =0.50), the nearly parallel-sided whorl profile and the rounded venter. The ornament of the last half volution consists of numerous (about 25) rather sharp ribles, which extend with a shallow sinus across the flanks and turn forward in the ventrolateral area, where they become coarser and for a hooklike rib (Fig. 62A). The venter is smooth. Remarks Stoyanowites aspinosus differs from S. dieneri in the absence of dorsolateral nodes and the presence of densely spaced curved riblets on the body chamber. Stoyanowites parallelus sp. nov. differs in the flat venter. Stratigraphic range Upper part of the Hambast Formation; 1.80 to 1.60 m below the extinction horizon (Alibashites ferdowsii Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 100-101, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Lutites lyriformis Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 3FC3DB4D-AA14-43A6-B982-0137ED22D6B7 Figs 51–52; Table 26 Diagnosis Species of Lutites gen. nov. with conch reaching 140 mm dm. Subadult stage with parallel-sided, compressed whorl profile (ww/wh = 0.60–0.70) and rounded venter; about 14 ventrolateral nodes per volution. Adult stage with subtrapezoidal, compressed whorl profile (ww/wh =0.50–0.60), concave flanks, concave venter and angular ventrolateral shoulder; numerous weak and densely arranged ribs. Prongs of external lobe bifid to multiply serrated; altogether 13–22 notches of E, A and L lobes. Etymology From the Latin ‘ lyra ’, referring to the shape of the whorl profile. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain A section; Hambast Formation; illustrated in Fig. 51A; MB.C.29995. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29994 • 2 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29996 to MB.C.29997 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29998 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29999 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30000 to MB.C.30001. Description Holotype MB.C.29995 is a rather well-preserved specimen with 96 mm conch diameter (Fig. 51A). It allows the study of three and a half volutions, of which only one third of a volution belongs to the body chamber. Adulthood is indicated by the weak crowding of the last septa at 73 mm diameter. The whorl profile is, at the largest preserved conch diameter, subtrapezoidal with a broadly rounded umbilical margin and concave and converging flanks. The weakly concave venter is separated from the flanks by an angular ventrolateral shoulder. Half a volution earlier, the venter is plane and another half a volution back, the venter is broadly rounded. The sculpture shows significant ontogenetic changes from the subadult stage with 13 coarse and blunt ventrolateral nodes, which originate from weak and rounded radial ribs, per volution. They become weaker and more numerous in the last quarter volution of the phragmocone (about 12 weak ventrolateral nodes within this interval). The body chamber possesses a very weak sculpture consisting only of weak and rounded radial riblets in the inner flank area. The suture line of the holotype possesses a parallel-sided external lobe with prongs being asymmetrically serrated by four and five small notches, respectively. The asymmetric ventrolateral saddle is nearly as wide as high; it is followed by an adventive lobe with oblique base serrated into seven small notches. The weakly asymmetric, inverted U-shaped lateral saddle is nearly as high as the ventrolateral saddle. Finally, the asymmetric lateral lobe has also seven small notches (Fig. 52A). The smaller paratype MB.C.29997 has 74 mm conch diameter, of which the phragmocone has a diameter of 64 mm and one quarter of a whorl belongs to the body chamber (Fig. 51B). The whorl profile is subtrapezoidal with concave, converging flanks and the venter is weakly concave. The sculpture generally resembles the holotype, but the ribs and nodes are weaker. The same is true for the additional paratypes MB.C.29996 (Fig. 51C) and MB.C.29999 (Fig. 51D), which is the conch geometry are nearly identical with the holotype. The suture line of paratype MB.C.29997 shows a parallel-sided external lobe that is deeper than the adventive lobe. The prongs are bifid and parallel-sided. All lobes and thus saddles are parallel-sided and all saddles have the same height. The adventive lobe and the lateral lobe possess six and four small notches (Fig. 52B). Remarks The new species differs in the coarse ventrolateral nodes of the subadult stage from most of the other species of Lutites gen. nov. with compressed whorl profile. Lutites lyriformis gen. et sp. nov. is similar to L. profundus gen. et sp. nov. but differs in the much weaker sculpture of the terminal body chamber, which shows only very weak ribs in L. lyriformis gen. et sp. nov. but ventrolateral nodes in L. profundus gen. et sp. nov. Stratigraphic range Upper part of the Hambast Formation; 1.45 to 1.00 m below the extinction horizon (Lutites profundus Zone to Abichites abichi Zone).

Paratirolites coronatus Korn & Ghaderi in Korn et al., 2016 Fig. 31; Table 14 Paratirolites coronatus Korn & Ghaderi in Korn et al., 2016: 860, text-figs 17–18. Diagnosis Species of Paratirolites with conch reaching 125 mm dm. Subadult stage with trapezoidal, moderately depressed whorl profile (ww/wh =1.60–1.90) and flattened tectiform venter; 15 coarse conical ventrolateral nodes per volution. Adult stage with extremely trapezoidal and moderately depressed whorl profile (ww/wh =1.60–1.90), flattened tectiform venter and subangular ventrolateral shoulder; weak ventrolateral nodes. Prongs of external lobe trifid or multiply serrated; altogether 11–24 notches of E, A and L lobes. Material examined Holotype IRAN • West Azerbaijan, Aras Valley section; Paratirolites Limestone, 2.90 m below the top; illustrated by Korn et al. (2016: text-fig. 17A); MB.C.25095. Additional material IRAN • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29842. Description Specimen MB.C.29842 is a fully chambered specimen with 55 mm conch diameter and allows the study of one and a half whorls (Fig. 31A). It has a strongly trapezoidal, depressed whorl profile (ww/wh =1.75), weakly sinuous flanks, an angular ventrolateral shoulder and a flattened tectiform venter. Half a volution earlier, the venter is weakly flattened but still rounded and another half volution back, the venter is broadly rounded. The sculpture changes during the last volution of the phragmocone; strong conical nodes at the beginning of this volution are transformed into smaller, longitudinally elongated nodes, which have a position at the ventrolateral shoulder. The suture line of specimen MB.C.29842 has a V-shaped external lobe with concave flanks and symmetric, narrowly V-shaped simple prongs. The ventrolateral saddle is asymmetric and dorsally inclined and the asymmetric, nearly parallel-sided adventive lobe has nine little notches. The lateral lobe is nearly flat at its base and subdivided in small equally sized notches (Fig. 31B). Remarks Paratirolites coronatus is the paratirolitid with the widest conch, resulting from the coronate ventrolateral shoulder. Therefore, it cannot be confused with any other species. Paratirolites coronatus differs from P. vediensis and the other species in the more strongly trapezoidal whorl profile of the adult stage. Stratigraphic range Upper part of the Hambast Formation; precise horizon at Baghuk Mountain not known.

Genus Alibashites Korn & Ghaderi in Korn et al., 2016 Type species Xenodiscus (Paratirolites ?) Mojsisovicsi Stoyanow, 1910, by original designation. Included species Alibashites ferdowsii Korn & Ghaderi in Korn et al., 2016; Xenodiscus (Paratirolites?) Mojsisovicsi Stoyanow, 1910; Alibashites uncinatus Korn & Ghaderi in Korn et al., 2016. Diagnosis Representatives of the family Dzhulfitidae with moderately large to large conch; maximum adult diameters 90 mm. Adult stage with subtrapezoidal or quadrate whorl profile. Subadult stage with large conical ventrolateral nodes, adult stage with weak ribs. Suture line with deep external lobe; depths of external lobe and adventive lobe nearly identical. Remarks Alibashites has an intermediate morphological position between Paratirolites and Abichites. It possesses the subadult sculpture with conical ventrolateral nodes of Paratirolites but lacks the trapezoidal whorl profile of that genus. Abichites has a similar adult conch morphology with a rectangular or subtrapezoidal whorl profile but does not possess the strong ventrolateral nodes developed in Alibashites., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 71, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211","Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135."]}

Genus Dzhulfites Shevyrev, 1965 Type species Dzhulfites spinosus Shevyrev, 1965, by original designation. Included species Dzhulfites hebes Korn & Ghaderi in Korn et al. (2016); Dzhulfites nodosus Shevyrev, 1965; Dzhulfites spinosus Shevyrev, 1965; Dzhulfites zalensis Korn & Ghaderi in Korn et al. (2016); Dzhulfites brevisellatus Korn & Hairapetian sp. nov. Diagnosis Representatives of the family Dzhulfitidae with moderately large to large conch; maximum adult diameters are between 80 and 160 mm. Subadult stage with trapezoidal whorl profile, adult stage variable. Subadult stage with small to large conical ventrolateral nodes, adult stage with weakening sculpture. Suture line with external lobe that does not reach the depth of the adventive lobe; prongs of external lobe simple to trifid. Remarks For a discussion of the genus, see Korn & Ghaderi in Korn et al. (2016)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 27, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Paratirolites aduncus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 86D671DC-3110-45AE-A379-2543EA8DEDD0 Figs 35–37; Table 17 Diagnosis Species of Paratirolites with conch reaching 125 mm dm. Subadult stage with circular or weakly depressed whorl profile (ww/wh = 1.00–1.30) and rounded venter; about 10 very coarse conical ventrolateral nodes per volution. Adult stage with weakly trapezoidal and weakly compressed whorl profile (ww/wh =0.75–0.90), flattened tectiform venter and subangular ventrolateral shoulder; about 10 hook-shaped ventrolateral nodes per half volution. Prongs of external lobe usually bifid but rarely trifid or multiply serrated; altogether 11–17 notches of E, A and L lobes. Etymology From the Latin ‘ aduncus ’, meaning ‘incurved’, because of the curved ventrolateral nodes. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain B section; Hambast Formation; illustrated in Fig. 35A; MB.C.29894. Paratypes IRAN • 9 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29880 to MB.C.29888 • 5 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29889 to MB.C.29893 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29895 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29896 to MB.C.29900 • 17 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29901 to MB.C.29917 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29918 • 8 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29919 to MB.C.29926 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29927 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29928 to MB.C.29930. Description Holotype MB.C.29894 (Fig. 35A) is a fairly well-preserved specimen with 80 mm conch diameter. It shows, at a conch diameter of 60 mm, three closely approximated septa, after which the terminal body chamber begins with a sudden weakening of the ventrolateral nodes. While these nodes are coarse and conical on the phragmocone, they almost spontaneously become elongated and hook-shaped on the body chamber. The suture line of holotype MB.C.29894 shows a deep, parallel-sided external lobe with prongs multiply serrated asymmetrically (Fig. 35C). The ventrolateral saddle is nearly symmetric and inverted U-shaped, it is followed by an adventive lobe and a lateral lobe with six little notches each. Both lobes are semicircular at the base and slightly asymmetric. Paratype MB.C.29880 has 83 mm conch diameter; half of the last volution belongs to the body chamber (Fig. 35B). It largely agrees, in conch shape and sculpture, with the holotype. The specimen allows for the study of two whorls, on which transformations in conch shape and sculpture can be observed. The phragmocone has a nearly circular whorl profile (ww/wh about 1.15 at 60 mm conch diameter) with nine coarse conical ventrolateral nodes on its last volution, which sometimes are developed as sharp spines. On the body chamber, the transformation into a weakly trapezoidal whorl profile can be observed; here the slightly concave flanks diverge towards the subangular ventrolateral shoulder that separates the flank from the flattened, weakly tectiform venter. On the body chamber occurs also a change in the sculpture towards more numerous (about 10 per half whorl) short ribs, which are restricted to the outer flank and ventrally end in hook-shaped, sharp nodes. Paratype MB.C.29900 is a specimen of 104 mm conch diameter. It has a fairly well-preserved phragmocone but a weathered body chamber, of which a little more than half a volution is preserved (Fig. 36A). The shape of the body chamber whorl profile had to be reconstructed; it shows a trapezoidal shape with weakly diverging flanks, a rounded ventrolateral shoulder and a flattened tectiform venter. One volution earlier, the whorl profile is strongly trapezoidal with flat, diverging flanks and a flattened, broadly rounded venter. The sculpture is coarse on the phragmocone and weakens in strength significantly on the body chamber. The last volution of the phragmocone has 15 conical, spiny ventrolateral nodes, which are first very coarse and conical but later elongate, weaker and more closely spaced. On the body chamber, weak and rounded ribs with concave course are visible on the outer flank. The suture line of paratype MB.C.29900 shows a parallel-sided external lobe with parallel-sided prongs that are divided by four small notches. The very narrow ventrolateral saddle is weakly inclined towards the umbilicus. Following is a very asymmetric, weakly pouched adventive lobe with eight notches, which are already climbing up the ventral side. The weakly inflated, asymmetric lateral saddle is low and the asymmetric lateral lobe with steep dorsal flank possesses five notches (Fig. 36B). Remarks Paratirolites aduncus sp. nov. differs from the other species of the genus in the presence of the hookshaped ventrolateral nodes of the adult stage. Stratigraphic range Upper part of the Hambast Formation; 2.65 to 1.15 m below the extinction horizon (Paratirolites kittli Zone to Lutites profundus Zone).

Family Dzhulfitidae Shevyrev, 1965 For the composition and diagnosis of the family, see Korn & Ghaderi in Korn et al. (2016)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 27, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Paratirolites trapezoidalis Shevyrev, 1965 Fig. 32; Table 15 Paratirolites trapezoidalis Shevyrev, 1965: 177, pl. 24 fig. 1. Paratirolites trapezoidalis – Shevyrev 1968: 92, pl. 4 fig. 1. — Korn in Ghaderi et al. 2014: text-fig. 7e. — Korn & Ghaderi in Korn et al. 2016: 858, text-fig. 16. Diagnosis Species of Paratirolites with conch reaching 120 mm dm. Subadult stage with trapezoidal, depressed whorl profile (ww/wh= 1.40–1.60) and flattened, slightly tectiform venter; 10–15 coarse ribs, which end in coarse conical ventrolateral nodes, per volution. Adult stage with strongly trapezoidal and weakly depressed whorl profile (ww/wh =1.00–1.40), flattened venter and angular ventrolateral shoulder; weak straight riblets and numerous small ventrolateral nodes. Prongs of external lobe usually unsubdivided, rarely trifid; altogether 12–19 notches of E, A and L lobes. Type material Holotype AZERBAIJAN • Nakhichevan Province, Dorasham 2 section; Paratirolites Limestone; illustrated by Shevyrev (1965: pl. 24 fig. 1); PIN 1252/129. Material examined IRAN • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29843 • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29844 to MB.C.29845 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29846 to MB.C.29848. Description Specimen MB.C.29844 is a moderately well-preserved individual of 71 mm conch diameter and allows the study of one and a half whorls (Fig. 32B). The last segment of 210 degrees belongs to the body chamber, which has a weakly depressed trapezoidal whorl profile (ww/wh =1.05) with weakly concave flanks and a nearly flat venter and an angular ventrolateral shoulder. The sculpture of the subadult stage consists of conical and coarse, spiny ventrolateral nodes, 13 on the last phragmocone volution. The nodes become significantly weaker and more numerous on the body chamber, where about 16 of them are present on half a volution. Specimen MB.C.29844 allows only the study of the suture line on the venter. The external lobe is narrow and parallel-sided with non-serrated, narrow lanceolate prongs. The ventrolateral saddle is very asymmetric and dorsally inclined and the asymmetric adventive lobe has an oblique base that is serrated by six small notches (Fig. 32C). specimen MB.C.29845 (Fig. 32A) is similar in conch shape and sculpture but possesses a suture line with trifid prongs of the external lobe. Remarks Paratirolites vediensis has a similar conch shape, but differs from P. trapezoidalis in the stronger ventrolateral nodes in the adult stage. P. coronatus has an even wider whorl profile (ww/wh = 1.60–1.80 in P. coronatus but only 1.40 in P. trapezoidalis); the species has also less coarse but more numerous ventrolateral nodes in the subadult stage (15 in P. coronatus but only 10 in P. trapezoidalis). Stratigraphic range Upper part of the Hambast Formation; 3.05 to 2.80 m below the extinction horizon (Paratirolites kittli Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 47-49, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Shevyrev A. A. 1968. Triasovye ammonoidei Yuga SSSR. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 119: 1 - 272.","Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. R. 2014. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts, and ammonoids. Fossil Record 15: 41 - 57. https: // doi. org / 10.5194 / fr- 17 - 41 - 2014","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Paratirolites aduncus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 86D671DC-3110-45AE-A379-2543EA8DEDD0 Figs 35–37; Table 17 Diagnosis Species of Paratirolites with conch reaching 125 mm dm. Subadult stage with circular or weakly depressed whorl profile (ww/wh = 1.00–1.30) and rounded venter; about 10 very coarse conical ventrolateral nodes per volution. Adult stage with weakly trapezoidal and weakly compressed whorl profile (ww/wh =0.75–0.90), flattened tectiform venter and subangular ventrolateral shoulder; about 10 hook-shaped ventrolateral nodes per half volution. Prongs of external lobe usually bifid but rarely trifid or multiply serrated; altogether 11–17 notches of E, A and L lobes. Etymology From the Latin ‘ aduncus ’, meaning ‘incurved’, because of the curved ventrolateral nodes. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain B section; Hambast Formation; illustrated in Fig. 35A; MB.C.29894. Paratypes IRAN • 9 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29880 to MB.C.29888 • 5 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29889 to MB.C.29893 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29895 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29896 to MB.C.29900 • 17 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29901 to MB.C.29917 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29918 • 8 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29919 to MB.C.29926 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29927 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29928 to MB.C.29930. Description Holotype MB.C.29894 (Fig. 35A) is a fairly well-preserved specimen with 80 mm conch diameter. It shows, at a conch diameter of 60 mm, three closely approximated septa, after which the terminal body chamber begins with a sudden weakening of the ventrolateral nodes. While these nodes are coarse and conical on the phragmocone, they almost spontaneously become elongated and hook-shaped on the body chamber. The suture line of holotype MB.C.29894 shows a deep, parallel-sided external lobe with prongs multiply serrated asymmetrically (Fig. 35C). The ventrolateral saddle is nearly symmetric and inverted U-shaped, it is followed by an adventive lobe and a lateral lobe with six little notches each. Both lobes are semicircular at the base and slightly asymmetric. Paratype MB.C.29880 has 83 mm conch diameter; half of the last volution belongs to the body chamber (Fig. 35B). It largely agrees, in conch shape and sculpture, with the holotype. The specimen allows for the study of two whorls, on which transformations in conch shape and sculpture can be observed. The phragmocone has a nearly circular whorl profile (ww/wh about 1.15 at 60 mm conch diameter) with nine coarse conical ventrolateral nodes on its last volution, which sometimes are developed as sharp spines. On the body chamber, the transformation into a weakly trapezoidal whorl profile can be observed; here the slightly concave flanks diverge towards the subangular ventrolateral shoulder that separates the flank from the flattened, weakly tectiform venter. On the body chamber occurs also a change in the sculpture towards more numerous (about 10 per half whorl) short ribs, which are restricted to the outer flank and ventrally end in hook-shaped, sharp nodes. Paratype MB.C.29900 is a specimen of 104 mm conch diameter. It has a fairly well-preserved phragmocone but a weathered body chamber, of which a little more than half a volution is preserved (Fig. 36A). The shape of the body chamber whorl profile had to be reconstructed; it shows a trapezoidal shape with weakly diverging flanks, a rounded ventrolateral shoulder and a flattened tectiform venter. One volution earlier, the whorl profile is strongly trapezoidal with flat, diverging flanks and a flattened, broadly rounded venter. The sculpture is coarse on the phragmocone and weakens in strength significantly on the body chamber. The last volution of the phragmocone has 15 conical, spiny ventrolateral nodes, which are first very coarse and conical but later elongate, weaker and more closely spaced. On the body chamber, weak and rounded ribs with concave course are visible on the outer flank. The suture line of paratype MB.C.29900 shows a parallel-sided external lobe with parallel-sided prongs that are divided by four small notches. The very narrow ventrolateral saddle is weakly inclined towards the umbilicus. Following is a very asymmetric, weakly pouched adventive lobe with eight notches, which are already climbing up the ventral side. The weakly inflated, asymmetric lateral saddle is low and the asymmetric lateral lobe with steep dorsal flank possesses five notches (Fig. 36B). Remarks Paratirolites aduncus sp. nov. differs from the other species of the genus in the presence of the hookshaped ventrolateral nodes of the adult stage. Stratigraphic range Upper part of the Hambast Formation; 2.65 to 1.15 m below the extinction horizon (Paratirolites kittli Zone to Lutites profundus Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 52-57, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Abichites ovalis Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 983F405A-033F-49EB-8D75-E5F917F23D86 Fig. 58; Table 32 Diagnosis Species of Abichites with conch reaching 85 mm dm. Subadult stage with subcircular, weakly compressed whorl profile (ww/wh =0.80–1.00) and rounded venter; about 12 shallow and rounded radial ribs. Adult stage with parallel sided or weakly trapezoidal, weakly compressed whorl profile (ww/wh =0.80–1.00), rounded venter and narrowly rounded ventrolateral shoulder; shallow wide ribs on the flank. Prongs of external lobe simple; altogether 7–11 notches of E, A and L lobes. Etymology From the Latin ‘ ovalis ’, meaning ‘oval’, because of the oval whorl profile in the subadult stage. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain B section; Hambast Formation; illustrated in Fig. 58A; MB.C.30059. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.30060 • 3 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30061 to MB.C.30063 • 6 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30064 to MB.C.30069 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30070 to MB.C.30071 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30072. Description Holotype MB.C.30059 is a rather well-preserved specimen with 62 mm conch diameter; the phragmocone diameter is 44 mm and 190 degrees of the body chamber are preserved (Fig. 58A). The adult whorl profile is weakly trapezoidal with gently diverging, slightly concave flanks, a rounded ventrolateral shoulder and a slightly flattened but broadly rounded venter. One volution earlier, the whorl profile is subcircular and weakly compressed (ww/wh = 0.90). The sculpture on the phragmocone consists of 12 shallow and rounded radial ribs per volution. They become weaker on the body chamber but develop to elongate, rather sharp nodes on the outer flank. The suture line of holotype MB.C.30059 shows an external lobe with flanks standing parallel in the lower part and diverging in the upper part. The prongs are very narrow, lanceolate and not secondarily serrated. On the outer flank lies the symmetric adventive lobe with semicircular, strongly serrated base; it is followed by a rather high lateral saddle and a parallel-sided lateral lobe with five small notches (Fig. 58C). Paratype MB.C.30071 has 60 mm conch diameter and is, with its weakly trapezoidal whorl profile and the general conch shape very similar to the previous one but has a more slender conch (ww/wh =0.85). It shows, in the inner whorls, weak ventrolateral nodes connected to the shallow radial ribs (Fig. 58B). Remarks Abichites ovalis sp. nov. closely resembles A. paucinodum, but differs in the whorl profile (diverging flanks in A. ovalis sp. nov. but converging in A. paucinodum), the wider umbilicus in A. ovalis sp. nov. and in the rounded venter in A. ovalis sp. nov. (flat in A. paucinodum). Stratigraphic range Upper part of the Hambast Formation; 0.75 to 0.40 m below the extinction horizon (Abichites stoyanowi Zone to lowest part of the Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 92-93, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Abichites stoyanowi (Kiparisova in Voinova et al., 1947) Fig. 59; Table 33 Kashmirites ? stoyanowi Kiparisova in Voinova et al., 1947: 149, pl. 35 fig. 1. Xenodiscus radians – Stoyanow 1910: 86, pl. 9 fig. 5. Xenodiscus sp. indet. Stoyanow 1910: 87, pl. 9 fig. 6. Abichites stoyanowi – Shevyrev 1965: 179, pl. 24 figs 2–3; 1968: 94, pl. 3 fig. 5, pl. 4 fig. 2. — Korn in Ghaderi et al. 2014: text-fig. 7h. — Korn & Ghaderi in Korn et al. 2016: 881, text-fig. 38. Diagnosis Species of Abichites with conch reaching 70 mm dm. Subadult stage with circular, weakly compressed whorl profile (ww/wh =0.75–1.00) and rounded venter; 16–20 moderately weak ribs per volution. Adult stage with quadrate and weakly compressed whorl profile (ww/wh =0.90–1.00), flat venter and subangular to angular ventrolateral shoulder; altogether 12–14 sharp concavo-convex ribs per half volution on flanks. Prongs of external lobe simple or bifid; 6–11 notches of E, A and L lobes. Material examined IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.30073 • 2 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30074 to MB.C.30075 • 3 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30076 to MB.C.30078 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.30079 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30080 to MB.C.30081 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30082. Description Specimen MB.C.30078 has 62 mm conch diameter and is rather well-preserved with the last half volution belonging to the body chamber (Fig. 59A). This has a subtrapezoidal, weakly compressed whorl profile (ww/wh =0.90) with a broadly rounded umbilical margin, weakly concave converging flanks, an angular ventrolateral shoulder and a weakly concave venter. Two stages of the ornament are visible; the subadult stage has coarse node-like ribs on the flank, which in the adult stage become increasingly weaker and more numerous (15 per half volution at the beginning of the body chamber). The suture line of specimen MB.C.30078 has a large external lobe with weakly diverging flanks and unsubdivided lanceolate prongs. The inverted U-shaped ventrolateral saddle is dorsally inclined and is followed by the pouched adventive lobe that has a very oblique base subdivided into three very shallow blunt notches. In contrast, the three notches of the parallel-sided lateral lobe are pointed denticles (Fig. 59E). The two smaller specimens MB.C.30074 (31 mm dm; Fig. 59D) and MB.C.30081 (34 mm dm; Fig. 59B) have similar conch shapes and proportions; they differ in the somewhat weaker sculpture. Remarks Abichites stoyanowi differs from A. abichi, which is another similar species with moderately coarse sculpture, in the much weaker ribs. Abichites subtrapezoidalis has a similar whorl profile, but only very weak riblets around the umbilicus on the adult body chamber. Stratigraphic range Upper part of the Hambast Formation; 0.80 to 0.05 m below the extinction horizon (Abichites stoyanowi Zone to lowest part of the Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 94, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Voinova E. V., Kiparisova L. D. & Robinson V. H. 1947. Klass Cephalopoda. Golovonogie., Atlas rukovodyashtchikh form iskopaemykh faun SSSR. VIII. Triasovaya sistema: 124 - 176, Moskva-Leningrad","Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135.","Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. R. 2014. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts, and ammonoids. Fossil Record 15: 41 - 57. https: // doi. org / 10.5194 / fr- 17 - 41 - 2014","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Paratirolites robustus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 10C9CACF-9ED4-4E61-8140-FDBB70154F87 Figs 26–27; Table 11 Diagnosis Species of Paratirolites with conch reaching 120 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.30–1.50) and rounded venter; 8–12 coarse, radially elongated midflank nodes per volution. Adult stage with trapezoidal and weakly depressed whorl profile (ww/wh =1.00–1.10), flattened, tectiform venter and angular ventrolateral shoulder; numerous weak ribs on flank. Prongs of external lobe simple to trifid; altogether 12–17 notches of E, A and L lobes. Etymology From the Latin ‘ robustus ’, meaning ‘robust’, because of the coarse ribs. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation, 2.95 m below top; illustrated in Fig. 26A; MB.C.29778. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29776 to MB.C.29777 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29779 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29780 • 4 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29781 to MB.C.29784. Description Holotype MB.C.29778 has 80 mm conch diameter and consists of a rather well-preserved phragmocone and a more or less strongly weathered body chamber (Fig. 26A). Septal crowding indicates that the maximum phragmocone diameter is 48 mm. The conch shape changes during growth; there is a strongly trapezoidal whorl profile of the phragmocone and a weakly trapezoidal profile of the body chamber. The sculpture of the phragmocone shows 12 conical nodes, which are circular and symmetric in section in the penultimate whorl but become asymmetric and radially elongated towards the end of the phragmocone. The suture line of holotype MB.C.29778 has a Y-shaped external lobe, in which the prongs are weakly serrated including two notches. The adventive lobe is asymmetric and ventrally inclined with nearly parallel flanks and asymmetric notching. It is dorsolaterally followed by an asymmetric lateral saddle and a nearly symmetric lateral lobe with seven small notches (Fig. 27A). The slightly smaller paratype MB.C.29782 (73 mm dm) is very similar in conch shape and suture line. The penultimate whorl has eight coarse conical nodes, which transform, at the end of the phragmocone, into weakly curved elongated nodes (Fig. 26B). The suture line has an external lobe with weakly diverging flanks, an asymmetric ventrolateral saddle and a weakly asymmetric, strongly notched adventive lobe with nearly flat base and parallel flanks (Fig. 27B). Remarks Paratirolites robustus sp. nov. differs from the most similar of the Central Iranian species P. lanceolobatus sp. nov. in the stouter whorl profile (ww/wh = 1.20 in P. robustus sp. nov. at 70 mm conch diameter and only 0.85 in P. lanceolobatus). Furthermore, P. lanceolobatus sp. nov. has non-serrated prongs of the parallel-sided external lobe. Paratirolites robustus sp. nov. differs from P. vediensis in the rounded venter and from P. kittli in the radially elongated nodes of the subadult stage and the presence of weak radial ribs of the body chamber. Stratigraphic range Upper part of the Hambast Formation; 3.20 to 2.90 m below the extinction horizon (uppermost part of the Paratirolites lanceolobatus Zone to Paratirolites kittli Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 36-40, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Paratirolites quadratus Korn & Ghaderi in Korn et al., 2016 Figs 38–39; Table 18 Paratirolites quadratus Korn & Ghaderi in Korn et al., 2016: 866, text-fig. 25. Diagnosis Paratirolites with conch reaching 90 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.35–1.60) and broadly rounded venter; about 11–13 ribs ending in spiny ventrolateral nodes. Adult stage with weakly trapezoidal to nearly quadrate and weakly depressed whorl profile (ww/wh =1.00–1.20), weakly concave venter and angular ventrolateral shoulder; coarse and shallow ribs on flanks. Prongs of external lobe multiply serrated; altogether about 14–17 notches of E, A and L lobes. Material examined Holotype IRAN • 1 specimen; East Azerbaijan, Ali Bashi N section; Paratirolites Limestone, 1.70 m below the top; illustrated by Korn et al. (2016: text-fig. 25); MB.C.25296. Additional material IRAN • 3 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29331 to MB.C.29333. Description Specimen MB.C.29932 is a nicely preserved specimen of 72 mm conch diameter (Fig. 38A). It shows weak septal crowding at 48 mm diameter and thus half of the last volution belongs to the body chamber. The whorl profile shape changes from trapezoidal with convex flanks of the subadult stage to parallelsided at the beginning of the body chamber; at the same time the shape of the venter changes from weakly convex to nearly totally flattened. The sculpture of the subadult stage possesses 13 conical ventrolateral nods per volution; they originate from rounded radial ribs. On the body chamber the nodes are weaker and become radially elongated, they are connected with low, rounded dorsolateral ribs. The suture line of specimen MB.C.29932 shows an asymmetric external lobe with weakly concave flanks, of which both prongs are trifid. Also on the venter follows the very narrow, weakly inflated ventrolateral saddle and the asymmetric adventive lobe (Fig. 38B). Specimen MB.C.29933 is a larger specimen with 101 mm conch diameter (Fig. 39A). It shows similar conch and sculpture characteristics but differs from the preceding specimen in the coarser ventrolateral nods and the larger phragmocone (62 mm dm). Its suture line has a parallel-sided external lobe with parallel-sided trifid prongs. The ventrolateral saddle is very narrow and the adventive lobe shows strong serration by small notches, which climb up the ventral flank to more than half of the lobe depth. The lateral saddle is weakly inflated and lower than the ventrolateral saddle and the lateral lobe is nearly parallel-sided and possesses five small notches (Fig. 39B). Remarks Paratirolites quadratus differs from the other species of Paratirolites in the slightly trapezoidal, nearly quadrate whorl profile in the adult conch and the very long, spiny ventrolateral nodes. Stratigraphic range At Baghuk Mountain only known from float from the upper part of the Hambast Formation; Stoyanowites dieneri Zone in the Julfa region.

Lutites profundus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: BB88C301-E69F-4E4B-8B25-F086D3BC6EEC Fig. 53; Table 27 Diagnosis Species of Lutites gen. nov. with conch reaching 125 mm dm. Subadult stage with subcircular, compressed whorl profile (ww/wh =0.70–0.80) and rounded venter; about 13 coarse ventrolateral nodes per volution. Adult stage with subtrapezoidal, compressed whorl profile (ww/wh =0.70–0.75), flat or concave venter and angular ventrolateral shoulder; weak dorsolateral ribs and elongated ventrolateral nodes. Prongs of external lobe trifid; altogether about 19 notches of E,A and L lobes. Etymology From the Latin ‘ profundus ’, meaning ‘deep’, because of the deep external lobe. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 53; MB.C.30003. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30002 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30004 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30005 • 3 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30006 to MB.C.30008. Description Holotype MB.C.30003 is a rather well-preserved specimen with 86 mm conch diameter, in which two whorls can be studied (Fig. 53A). One third of the last volution belongs to the body chamber and the last four septa of the phragmocone are crowded. In the adult stage, the compressed whorl profile (ww/ wh= 0.75) shows a broadly rounded umbilical margin with an oblique umbilical wall. The whorl profile is widest on the inner flank, from where the concave flanks converge towards the angular ventrolateral shoulder. The venter is concave in this growth stage. Half a volution back the venter is flat and another half whorl back, a rounded venter can be seen belonging to a subcircular, compressed whorl section (ww/wh =0.80). The sculpture experiences a change in strength during the last volution. The penultimate volution has ten coarse ventrolateral nodes, which are connected with weak and rounded radial ribs. These nodes become weaker and more numerous at the end of the phragmocone, but they are still present on the body chamber (12 on a distance of 120 degrees). Here, they are elongate and alternate, in their positions, with low umbilical ribs. The suture line of the holotype has a narrow and deep external lobe (deeper than the adventive lobe) with parallel flanks in the lower part. Its prongs are parallel-sided with asymmetric base and possesses three small notches. The ventrolateral saddle is narrow and asymmetric; it borders the adventive lobe that is widest near its base. It shows two groups of five very narrow notches. The nearly symmetric and inverted U-shaped lateral saddle is higher than the ventrolateral saddle. The asymmetric lateral lobe with less steep dorsal flank has also two groups of narrow notches, a ventral group with two and a dorsal group with three notches (Fig. 53B). Remarks Lutites profundus gen. et sp. nov. is similar to L. lyriformis gen. et sp. nov. but differs in the much coarser sculpture of the terminal body chamber, which shows only very weak ribs in the latter species but ventrolateral nodes in L. profundus gen. et sp. nov. All the other species of Lutites gen. nov. with a slender conch also show a much weaker sculpture. Stratigraphic range Upper part of the Hambast Formation; 1.60 to 1.20 m below the extinction horizon (Lutites profundus Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 81-83, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Paratirolites rubens Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: C8D262EC-926D-4CAB-AE08-BB8C158CE276 Fig. 24; Table 9 Diagnosis Species of Paratirolites with conch reaching 110 mm dm. Subadult stage with broadly oval, moderately depressed whorl profile (ww/wh =1.60) and broadly rounded venter; about 8 coarse ventrolateral nodes per volution. Adult stage with broadly oval, weakly depressed whorl profile (ww/wh = 1.30), broadly rounded venter and rounded ventrolateral shoulder; coarse ventrolateral nodes. Prongs of external lobe unserrated; altogether 16 notches of E, A and L lobes. Etymology From the Latin ‘ rubens ’, meaning ‘red’, because of the red limestone horizon where the material comes from. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 24; MB.C.29749. Paratype IRAN • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29750. Description Holotype MB.C.29749 is a rather well-preserved individual of 50 mm conch diameter (Fig. 24A). It is fully chambered with septal crowing at the end; therefore the adult diameter including body chamber has been about 90 mm. The whorl profile is, at 50 mm conch diameter, broadly oval (ww/wh =1.80) with broadly convex flanks and venter. Half a volution earlier, the ww/wh ratio is even higher (1.90). The sculpture consists of eight coarse conical nodes with a position on the midflank of the last volution; the penultimate volution bears only six nodes. The suture line of the holotype has a narrow external lobe that is wider near its base and weakly constricted in the upper part. It possesses very narrow, non-serrated prongs. A narrow and slightly inflated ventrolateral saddle and an asymmetric adventive lobe with ten little notches follow on the sides of the venter, while the continuously rounded lateral saddle and the nearly symmetric lateral lobe (with five small notches) have a position on the flank (Fig. 24B). Remarks Paratirolites rubens sp. nov. is the species of the genus with the lowest number of ventrolateral nodes. Furthermore, the broadly rounded venter is a criterion of clear separation from species such as P. vediensis and P. kittli. There is also some similarity to the species Dzhulfites spinosus and D. hebes, but these differ in the very short external lobe. Stratigraphic range None of the specimens was collected in situ, but according to the intensive red colour of the host sediment, a position in the Hambast Formation about five metres below the extinction horizon is most likely (probably Shevyrevites shevyrevi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 33-34, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Lutites paucis Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 9C85D567-DA54-463F-8CE3-BFDBABD83928 Fig. 50; Table 25 Diagnosis Species of Lutites gen. nov. with conch reaching 70 mm dm. Subadult stage with circular, slightly compressed whorl profile (ww/wh =0.90–1.00) and broadly rounded venter; very coarse conical ventrolateral nodes. Adult stage with subtrapezoidal, slightly compressed whorl profile (ww/wh =0.80), flattened venter and subangular ventrolateral shoulder; without ribs or nodes. Prongs of external lobe simple; altogether 11 notches of E, A and L lobes. Etymology From the Latin ‘ paucis ’, meaning ‘fewer’, because of the low number of lobes and nodes. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 2.45 m below the top; illustrated in Fig. 50A; MB.C.29991. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29992 to MB.C.29993. Description Holotype MB.C.29991 is a partly corroded specimen with 64 mm conch diameter with two thirds of the last volution belonging to the body chamber (Fig. 50A). At the end of the body chamber, the whorl profile is subtrapezoidal with a broadly rounded umbilical margin, converging flanks, a narrowly rounded ventrolateral shoulder and a flattened venter. By contrast, the whorl profile is trapezoidal at the beginning of the body chamber and subcircular in the last portion of the phragmocone. The sculpture of the phragmocone consists of very coarse conical ventrolateral nodes (eight per volution). The body chamber appears to be smooth. The suture line of the holotype has a large external lobe with weakly diverging flanks and simple, narrow and lanceolate prongs. The ventrolateral saddle is narrowly rounded and the lateral saddle is weakly inflated. Both the adventive lobe and the lateral lobe are parallel-sided and both characterised by five notches that are very dissimilar in size (Fig. 50B). Remarks Lutites paucis gen. et sp. nov. differs from the other species of the genus in the stouter conch, the only weakly flattened venter and the lower number of conical nodes. The new species may represent an intermediate form between Alibashites and Lutites gen. nov. Stratigraphic range Upper part of the Hambast Formation; 2.45 m to 1.20 m below the extinction horizon (uppermost part of the Paratirolites kittli Zone to the Lutites profundus Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 77-78, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Genus Clivotirolites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 2F50EBFF-9384-40A5-84D9-573E68455DD2 Type species Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Representatives of the family Dzhulfitidae with large conch; maximum adult diameters are about 120 mm. Adult stage with trapezoidal whorl profile. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs. Suture line protracted on the flanks, with deep external lobe; depths of external lobe and adventive lobe nearly identical. Etymology From the Latin ‘ clivus ’, meaning ‘oblique’, because of the suture line and the similarity with Tirolites and Paratirolites. Included species Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov.; Clivotirolites petilus Korn & Hairapetian gen. et sp. nov. Remarks Clivotirolites gen. nov. differs from Paratirolites and related genera in the oblique protracting outline of the outer suture line., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 61-62, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Abichites alibashiensis Korn & Ghaderi in Korn et al., 2016 Fig. 57; Table 31 Abichites alibashiensis Korn & Ghaderi in Korn et al., 2016: 879, text-fig. 36. Diagnosis Species of Abichites with conch reaching 85 mm dm. Subadult stage with broadly oval, weakly depressed whorl profile (ww/wh =1.10–1.30) and rounded venter; 12–15 ventrolateral nodes per volution. Adult stage with quadrate of weakly subtrapezoidal, weakly compressed whorl profile (ww/wh =0.80–1.25), flattened venter and subangular to angular ventrolateral shoulder; sharp ribs. Prongs of external lobe simple or bifid; altogether 7–11 notches of E, A and L lobes. Material examined Holotype IRAN • East Azerbaijan, Ali Bashi N section; Paratirolites Limestone, 1.40 m below the top; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 36a); MB.C.25399. Additional material IRAN • 2 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.30041 to MB.C.30042 • 3 specimens; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.30043 to MB.C.30045 • 3 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30046 to MB.C.30048 • 5 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30049 to MB.C.30053 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.30054 • 4 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30055 to MB.C.30058. Description Specimen MB.C.30051 has 62 mm conch diameter and is rather well preserved with about 190 degrees of the body chamber visible (Fig. 57A). The whorl profile of the terminal body chamber has a broadly rounded umbilical margin, weakly concave flanks that are gently converging towards the flat venter, which is deliminated by an angular ventrolateral shoulder. In the phragmocone, the whorl profile is oval with a broadly rounded venter. The change of the sculpture from the subadult stage with coarse ventrolateral nodes (12 per volution) towards the adult stage with only weak ribs on the flank happens already before the beginning of the terminal body chamber. Specimen MB.C.30046 is smaller (48 mm dm) and shows the transformation from the subadult stage with rounded ventrolateral margin to the adult stage with quadrate whorl profile, in which the venter is flat and separated from the flanks by an angular margin (Fig. 57B). The sculpture of the subadult stage possesses prominent ribs (about 13 per volution), which weaken out on the terminal body chamber. The suture line of specimen MB.C.30046 shows a simply subdivided external lobe with narrow, V-shaped prongs, a broadly rounded parabolic ventrolateral saddle and asymmetric, weakly serrated adventive and lateral lobes (Fig. 57D). Remarks Abichites alibashiensis differs from A. abichi, A. stoyanowi and A. subtrapezoidalis in the sharp ribs, which tend to end in ventrolateral nodes, in the subadult stage. Species of Alibashites possess subadult ribs but also ventrolateral nodes; these are A. mojsisovicsi (with very weak adult ribs) and A. ferdowsii (with a much more depressed whorl profile). Stratigraphic range Upper part of the Hambast Formation; 1.15 to 0.40 m below the extinction horizon (Abichites abichi Zone to lowest part of the Arasella minuta Zone).

Genus Esfahanites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 61EDFE2C-1650-4303-BDC4-3E9F0434CDB8 Type species Esfahanites armatus Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Representative of the family Dzhulfitidae with large conch; maximum adult diameters about 120 mm. Adult stage with trapezoidal whorl profile. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs. Suture line with very small external lobe reaching only half the depth of adventive lobe. Etymology Named after the city of Esfahan. Included species Esfahanites armatus Korn & Hairapetian gen. et sp. nov. Remarks Esfahanites gen. nov. differs from Paratirolites and related genera in the very small external lobe.

Genus Clivotirolites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 2F50EBFF-9384-40A5-84D9-573E68455DD2 Type species Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Representatives of the family Dzhulfitidae with large conch; maximum adult diameters are about 120 mm. Adult stage with trapezoidal whorl profile. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs. Suture line protracted on the flanks, with deep external lobe; depths of external lobe and adventive lobe nearly identical. Etymology From the Latin ‘ clivus ’, meaning ‘oblique’, because of the suture line and the similarity with Tirolites and Paratirolites. Included species Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov.; Clivotirolites petilus Korn & Hairapetian gen. et sp. nov. Remarks Clivotirolites gen. nov. differs from Paratirolites and related genera in the oblique protracting outline of the outer suture line.

Genus Esfahanites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 61EDFE2C-1650-4303-BDC4-3E9F0434CDB8 Type species Esfahanites armatus Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Representative of the family Dzhulfitidae with large conch; maximum adult diameters about 120 mm. Adult stage with trapezoidal whorl profile. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs. Suture line with very small external lobe reaching only half the depth of adventive lobe. Etymology Named after the city of Esfahan. Included species Esfahanites armatus Korn & Hairapetian gen. et sp. nov. Remarks Esfahanites gen. nov. differs from Paratirolites and related genera in the very small external lobe., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 67, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Paratirolites robustus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 10C9CACF-9ED4-4E61-8140-FDBB70154F87 Figs 26–27; Table 11 Diagnosis Species of Paratirolites with conch reaching 120 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.30–1.50) and rounded venter; 8–12 coarse, radially elongated midflank nodes per volution. Adult stage with trapezoidal and weakly depressed whorl profile (ww/wh =1.00–1.10), flattened, tectiform venter and angular ventrolateral shoulder; numerous weak ribs on flank. Prongs of external lobe simple to trifid; altogether 12–17 notches of E, A and L lobes. Etymology From the Latin ‘ robustus ’, meaning ‘robust’, because of the coarse ribs. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation, 2.95 m below top; illustrated in Fig. 26A; MB.C.29778. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29776 to MB.C.29777 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29779 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29780 • 4 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29781 to MB.C.29784. Description Holotype MB.C.29778 has 80 mm conch diameter and consists of a rather well-preserved phragmocone and a more or less strongly weathered body chamber (Fig. 26A). Septal crowding indicates that the maximum phragmocone diameter is 48 mm. The conch shape changes during growth; there is a strongly trapezoidal whorl profile of the phragmocone and a weakly trapezoidal profile of the body chamber. The sculpture of the phragmocone shows 12 conical nodes, which are circular and symmetric in section in the penultimate whorl but become asymmetric and radially elongated towards the end of the phragmocone. The suture line of holotype MB.C.29778 has a Y-shaped external lobe, in which the prongs are weakly serrated including two notches. The adventive lobe is asymmetric and ventrally inclined with nearly parallel flanks and asymmetric notching. It is dorsolaterally followed by an asymmetric lateral saddle and a nearly symmetric lateral lobe with seven small notches (Fig. 27A). The slightly smaller paratype MB.C.29782 (73 mm dm) is very similar in conch shape and suture line. The penultimate whorl has eight coarse conical nodes, which transform, at the end of the phragmocone, into weakly curved elongated nodes (Fig. 26B). The suture line has an external lobe with weakly diverging flanks, an asymmetric ventrolateral saddle and a weakly asymmetric, strongly notched adventive lobe with nearly flat base and parallel flanks (Fig. 27B). Remarks Paratirolites robustus sp. nov. differs from the most similar of the Central Iranian species P. lanceolobatus sp. nov. in the stouter whorl profile (ww/wh = 1.20 in P. robustus sp. nov. at 70 mm conch diameter and only 0.85 in P. lanceolobatus). Furthermore, P. lanceolobatus sp. nov. has non-serrated prongs of the parallel-sided external lobe. Paratirolites robustus sp. nov. differs from P. vediensis in the rounded venter and from P. kittli in the radially elongated nodes of the subadult stage and the presence of weak radial ribs of the body chamber. Stratigraphic range Upper part of the Hambast Formation; 3.20 to 2.90 m below the extinction horizon (uppermost part of the Paratirolites lanceolobatus Zone to Paratirolites kittli Zone).

Paratirolites baghukensis Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 27027DEB-9F15-4F61-8EA3-3F15E997A1D9 Figs 33–34; Table 16 Diagnosis Species of Paratirolites with conch reaching 130 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.00–1.20) and rounded venter; about 12 moderately coarse ribs ending in conical ventrolateral nodes per volution. Adult stage with trapezoidal and weakly compressed whorl profile (ww/wh =0.70–0.90), flattened tectiform venter and subangular ventrolateral shoulder; numerous sharp ribs ending in sharp radially elongated ventrolateral nodes. Prongs of external lobe usually trifid; altogether 13–15 notches of E, A and L lobes. Etymology Named after the occurrence at Baghuk Mountain. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain G section; Hambast Formation; illustrated in Fig. 33A; MB.C.29868. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29849 to MB.C.29850 • 4 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29851 to MB.C.29854 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29855 to MB.C.29859 • 8 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29860 to MB.C.29867 • 7 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29869 to MB.C.29875 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29876 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29877 to MB.C.29879. Description Holotype MB.C.29868 has 70 mm conch diameter and is fairly well-preserved but slightly corroded. The last third of the terminal volution belongs to the body chamber (which is thus only incompletely preserved); adulthood of the specimen is obvious because of the crowding of the last septa of the phragmocone (Fig. 33A). The penultimate of the preserved whorls shows a trapezoidal profile with a rounded venter. The body chamber finally shows a trapezoidal whorl profile with weakly diverging flanks, an angular ventrolateral shoulder and a nearly flat, tectiform venter. The sculpture changes with the beginning of the terminal body chamber, but this change is continuous without sudden breaks. One subadult volution has ten isolated spiny ventrolateral nodes.Already towards the end of the phragmocone, these nodes become radially elongate and form coarse ribs across the flank. At the beginning of the terminal body chamber, the ventrolateral nodes weaken and finally, only sharp ribs, about 15 per quarter of a volution, are present. The suture line of holotype MB.C.29868 shows a slightly pouched external lobe with trifid, asymmetric prongs. The ventrolateral saddle is weakly inflated, the adventive and lateral lobes are flattened at the base and subdivided in five and four notches, respectively (Fig. 33C). Paratype MB.C.29860 (Fig. 33B) is smaller with 61 mm conch diameter but confirms the description given for the holotype. It shows four whorls preserved and thus allows for the study of the sculpture of the inner volutions. The two larger paratypes MB.C.29874 and MB.C.29850 (Fig. 34) show then the transformation into the weakly ornamented adult stage, in which a tendency towards a parallel-sided whorl profile is visible. Remarks Paratirolites baghukensis sp. nov. belongs to the slender species of the genus. It differs from the other species of the genus in the sharp ribs on the adult body chamber. Paratirolites multiconus also has a slender conch, but differs from P. baghukensis sp. nov. in the much more numerous ventrolateral nodes. Stratigraphic range Upper part of the Hambast Formation; 3.10 to 2.35 m below the extinction horizon (Paratirolites kittli Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 49-52, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Lutites profundus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: BB88C301-E69F-4E4B-8B25-F086D3BC6EEC Fig. 53; Table 27 Diagnosis Species of Lutites gen. nov. with conch reaching 125 mm dm. Subadult stage with subcircular, compressed whorl profile (ww/wh =0.70–0.80) and rounded venter; about 13 coarse ventrolateral nodes per volution. Adult stage with subtrapezoidal, compressed whorl profile (ww/wh =0.70–0.75), flat or concave venter and angular ventrolateral shoulder; weak dorsolateral ribs and elongated ventrolateral nodes. Prongs of external lobe trifid; altogether about 19 notches of E,A and L lobes. Etymology From the Latin ‘ profundus ’, meaning ‘deep’, because of the deep external lobe. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 53; MB.C.30003. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30002 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30004 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30005 • 3 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30006 to MB.C.30008. Description Holotype MB.C.30003 is a rather well-preserved specimen with 86 mm conch diameter, in which two whorls can be studied (Fig. 53A). One third of the last volution belongs to the body chamber and the last four septa of the phragmocone are crowded. In the adult stage, the compressed whorl profile (ww/ wh= 0.75) shows a broadly rounded umbilical margin with an oblique umbilical wall. The whorl profile is widest on the inner flank, from where the concave flanks converge towards the angular ventrolateral shoulder. The venter is concave in this growth stage. Half a volution back the venter is flat and another half whorl back, a rounded venter can be seen belonging to a subcircular, compressed whorl section (ww/wh =0.80). The sculpture experiences a change in strength during the last volution. The penultimate volution has ten coarse ventrolateral nodes, which are connected with weak and rounded radial ribs. These nodes become weaker and more numerous at the end of the phragmocone, but they are still present on the body chamber (12 on a distance of 120 degrees). Here, they are elongate and alternate, in their positions, with low umbilical ribs. The suture line of the holotype has a narrow and deep external lobe (deeper than the adventive lobe) with parallel flanks in the lower part. Its prongs are parallel-sided with asymmetric base and possesses three small notches. The ventrolateral saddle is narrow and asymmetric; it borders the adventive lobe that is widest near its base. It shows two groups of five very narrow notches. The nearly symmetric and inverted U-shaped lateral saddle is higher than the ventrolateral saddle. The asymmetric lateral lobe with less steep dorsal flank has also two groups of narrow notches, a ventral group with two and a dorsal group with three notches (Fig. 53B). Remarks Lutites profundus gen. et sp. nov. is similar to L. lyriformis gen. et sp. nov. but differs in the much coarser sculpture of the terminal body chamber, which shows only very weak ribs in the latter species but ventrolateral nodes in L. profundus gen. et sp. nov. All the other species of Lutites gen. nov. with a slender conch also show a much weaker sculpture. Stratigraphic range Upper part of the Hambast Formation; 1.60 to 1.20 m below the extinction horizon (Lutites profundus Zone).

Dzhulfites brevisellatus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 1FDCA9D7-546A-48FA-94E1-0053EB7F7C23 Fig. 23; Table 8 Diagnosis Species of Dzhulfites with conch reaching 160 mm dm. Subadult stage with weakly trapezoidal, moderately depressed whorl profile (ww/wh =1.50–1.80) and broadly rounded venter; 10–13 coarse conical nodes on midflank per volution. Adult stage with numerous weak ribs and small ventrolateral nodes. Prongs of external lobe bifid or trifid; altogether 15–20 notches of E, A and L lobes. Etymology From the Latin ‘ brevis ’, meaning ‘low’ and ‘ sella ’, meaning ‘saddle’, after the low amplitude of lobes and saddles in the suture line. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain H section; Hambast Formation, 5.70 m below top; illustrated in Fig. 23B; MB.C.29748. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29746 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29747. Description The large paratype MB.C.29746 has 125 mm conch diameter, of which half of the last whorl belongs to the strongly weathered body chamber (Fig. 23A). The phragmocone is 90 mm in diameter and is partly crushed; it shows that the whorl profile is trapezoidal with a broadly rounded venter. The sculpture changes dramatically throughout ontogeny. At 40 mm dm there are 13 coarse, spiny ventrolateral nodes per volution; these nodes become weaker and more numerous so that the last 120° of the phragmocone have about 20 weak ventrolateral nodes, which are connected with weak and slightly sinuous radial ribs on the flank. The suture line of the specimen suffered from weathering, but, most probably, the prongs of the very short external lobe are bifid. Holotype MB.C.29748 is a fairly well-preserved, fully septate specimen with 51 mm conch diameter (Fig. 23B). It has a weakly trapezoidal whorl profile with broadly rounded flanks and venter. The sculpture shows about ten coarse conical nodes, which are positioned on the midflank, on the last preserved volution. The suture line of the holotype shows an external lobe with subparallel, weakly diverging flanks and weakly asymmetric, trifid prongs. All three saddles of the outer suture line are similar in shape, being inverted U-shaped and broadly rounded. The adventive lobe and the lateral lobe are similar in shape with parallel flanks and many small notches at the base. They differ in their depth; the adventive lobe is much deeper than the external lobe (Fig. 23C). Remarks Among the species of Dzhulfites, only D. nodosus is similar in conch shape and sculpture, but this species shows an applanate venter in the adult stage. Dzhulfites brevisellatus sp. nov. differs from D. spinosus and the other species of the genus in the suture line, which in D. brevisellatus sp. nov. shows much wider elements; particularly the saddles are wider than the lobes. In the shape of the suture line, D. brevisellatus sp. nov. has some resemblance to representatives of the family Xenodiscidae. Stratigraphic range Upper part of the Hambast Formation; 5.70 to 4.70 m below the extinction horizon (Dzhulfites spinosus Zone to Shevyrevites shevyrevi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 30-32, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Alibashites ferdowsii Korn & Ghaderi in Korn et al., 2016 Figs 48–49; Table 24 Alibashites ferdowsii Korn & Ghaderi in Korn et al., 2016: 871, text-fig. 30. Diagnosis Species of Alibashites with conch reaching 100 mm dm. Subadult stage with circular to subtrapezoidal, weakly depressed whorl profile (ww/wh =1.25–1.50) and broadly rounded venter; 10–12 very coarse ventrolateral nodes. Adult stage with subtrapezoidal and weakly depressed whorl profile (ww/wh =1.20– 1.50) with converging flanks, almost flat venter and angular ventrolateral shoulder; sharp ribs on flanks, forming short dorsolateral nodes and stronger ventrolateral nodes. Prongs of external lobe variable in shape, bifid to multiply serrated; altogether 15–18 notches of E, A and L lobes. Material examined Holotype IRAN • West Azerbaijan, Aras Valley section; Paratirolites Limestone; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 30a); MB.C.25327. Additional material IRAN • 6 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29976 to MB.C.29978 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29979 • 7 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29980 to MB.C.29986 • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29987 to MB.C.29988 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29989 to MB.C.29990. Description Specimen MB.C.29974 is the largest available specimen with 59 mm conch diameter (Fig. 48A). Slightly less than half of a volution belongs to the body chamber. The profile of the aperture is characterised by a broadly rounded umbilical margin, weakly concave flanks, an angular ventrolateral shoulder and a broadly rounded but somehow flattened venter. The whorl profile in this stage is weakly depressed with a ww/wh ratio of 1.35. One volution earlier, the venter is more rounded and the flanks are widely curved, producing a depressed oval whorl profile. The sculpture changes from the subadult stage with coarse ventrolateral nodes per volution into the body chamber sculpture with sharp dorsolateral ribs, which are connected by low and rounded folds with weak ventrolateral nodes. The suture line of specimen MB.C.29974 is characterised by a Y-shaped external lobe, of which the prongs are subdivided by four small notches of varying depth. The ventrolateral saddle is strikingly asymmetric and dorsally inclined. It is followed by the asymmetric adventive lobe with seven small notches, the asymmetric parabolic lateral saddle and a lateral lobe with three notches (Fig. 49A). Specimen MB.C.29987 is a specimen with 51 mm conch diameter. Its profile is subtrapezoidal at the aperture; it shows a convex and oblique umbilical wall and is widest in the dorsolateral portion of the flank and weakly concave in the outer flank area (Fig. 48C). The flattened venter is separated from the flank by an angular ventrolateral shoulder. One whorl earlier is the whorl profile trapezoidal with a rounded ventrolateral shoulder and a broadly rounded venter. The sculpture on the phragmocone consists of twelve coarse conical nodes; on the body chamber these nodes become weaker and are connected with elongate rounded dorsolateral ribs by shallow, rounded folds. The suture line shows a slightly pouched external lobe with lanceolate, unsubdivided prongs. The ventrolateral saddle is turned U-shaped with nearly parallel flanks. Both flanks of the adventive lobe stand parallel, but its base is oblique and serrated into six little notches (Fig. 49D). Remarks The specimens from Baghuk Mountain are morphologically very similar to the material from the type area, and the sizes of the specimens are nearly identical. Differences occur in the degree of sutural subdivision; while the prongs of the external lobe in the material from Julfa are secondarily subdivided (two, three or four notches), many of the specimens from Baghuk Mountain show non-subdivided lanceolate prongs. Alibashites ferdowsii shows an intermediate morphological position between the genera Paratirolites and Abichites. Characteristic for Paratirolites is the intermediate growth stage with trapezoidal whorl profile with the prominent conical ventrolateral nodes; similar to Abichites is the flat venter in the adult stage. The intermediate position of A. ferdowsii is also visible in the suture line; on the one side is the more complex external lobe in Paratirolites (with serrated prongs of the external lobe), and on the other side is Abichites (with a trend towards unserrated prongs of the external lobe). Stratigraphic range Upper part of the Hambast Formation; 2.25 to 0.70 m below the extinction horizon (Alibashites ferdowsii Zone to lower part of the Abichites stoyanowi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 73-76, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Paratirolites coronatus Korn & Ghaderi in Korn et al., 2016 Fig. 31; Table 14 Paratirolites coronatus Korn & Ghaderi in Korn et al., 2016: 860, text-figs 17–18. Diagnosis Species of Paratirolites with conch reaching 125 mm dm. Subadult stage with trapezoidal, moderately depressed whorl profile (ww/wh =1.60–1.90) and flattened tectiform venter; 15 coarse conical ventrolateral nodes per volution. Adult stage with extremely trapezoidal and moderately depressed whorl profile (ww/wh =1.60–1.90), flattened tectiform venter and subangular ventrolateral shoulder; weak ventrolateral nodes. Prongs of external lobe trifid or multiply serrated; altogether 11–24 notches of E, A and L lobes. Material examined Holotype IRAN • West Azerbaijan, Aras Valley section; Paratirolites Limestone, 2.90 m below the top; illustrated by Korn et al. (2016: text-fig. 17A); MB.C.25095. Additional material IRAN • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29842. Description Specimen MB.C.29842 is a fully chambered specimen with 55 mm conch diameter and allows the study of one and a half whorls (Fig. 31A). It has a strongly trapezoidal, depressed whorl profile (ww/wh =1.75), weakly sinuous flanks, an angular ventrolateral shoulder and a flattened tectiform venter. Half a volution earlier, the venter is weakly flattened but still rounded and another half volution back, the venter is broadly rounded. The sculpture changes during the last volution of the phragmocone; strong conical nodes at the beginning of this volution are transformed into smaller, longitudinally elongated nodes, which have a position at the ventrolateral shoulder. The suture line of specimen MB.C.29842 has a V-shaped external lobe with concave flanks and symmetric, narrowly V-shaped simple prongs. The ventrolateral saddle is asymmetric and dorsally inclined and the asymmetric, nearly parallel-sided adventive lobe has nine little notches. The lateral lobe is nearly flat at its base and subdivided in small equally sized notches (Fig. 31B). Remarks Paratirolites coronatus is the paratirolitid with the widest conch, resulting from the coronate ventrolateral shoulder. Therefore, it cannot be confused with any other species. Paratirolites coronatus differs from P. vediensis and the other species in the more strongly trapezoidal whorl profile of the adult stage. Stratigraphic range Upper part of the Hambast Formation; precise horizon at Baghuk Mountain not known., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 46-47, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Stoyanowites parallelus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: E6C0C2D8-02EF-4E72-9E1E-78816A50573E Fig. 63; Table 37 Diagnosis Species of Stoyanowites with conch reaching 100 mm dm. Subadult stage with rectangular, compressed whorl profile (ww/wh =0.70), subangular ventrolateral shoulder and flattened venter; 10 rather coarse ventrolateral spines per half volution. Adult stage with parallel-sided, compressed whorl profile (ww/ wh= 0.60), flat venter and angular ventrolateral shoulder; 15 ventrolateral spines per half volution. Prongs of external lobe simple; altogether 13 notches of E, A and L lobes. Etymology Named after the parallel flanks of the whorl profile. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 1.70 m below the top; illustrated in Fig. 63A; MB.C.30102. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30103 to MB.C.30104 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30105. Description Holotype MB.C.30102 is an incomplete specimen of 72 mm diameter; it allows the study of conch shape, ornament and suture line. The conch is extremely discoidal (ww/dm= 0.19) with a wide umbilicus (uw/ dm =0.47) and a compressed whorl profile (ww/wh =0.61). It has nearly parallel, slightly concave flanks, a prominent subangular ventrolateral shoulder and a weakly concave venter at the largest diameter (Fig. 63A). The sculpture is dominated by coarse ventrolateral spines, which become weaker towards the end of growth, but of which are still 15 are present on half a volution of the body chamber. They are not always connected with short ribs on the inner flank. The suture line of holotype MB.C.30102 shows the short external lobe, which has simple, weakly pouched prongs. The ventrolateral saddle is rather narrow; the adventive and lateral lobes possess six notches (Fig. 63B). Remarks Stoyanowites parallelus sp. nov. differs from S. dieneri and S. aspinosus in the flat venter and the numerous sharp ventrolateral nodes in the adult stage. Stratigraphic range Upper part of the Hambast Formation; 1.70 m below the extinction horizon (Alibashites ferdowsii Zone).

Clivotirolites petilus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 6608C3B9-8B98-4E05-91E6-DA463119748F Fig. 44; Table 21 Diagnosis Species of Clivotirolites gen. nov. with conch reaching 120 mm dm. Subadult stage with trapezoidal, weakly compressed whorl profile (ww/wh=0.90) and flattened tectiform; 15 coarse ventrolateral nodes per volution. Adult stage with trapezoidal, weakly compressed whorl profile (ww/wh =0.70), flat venter and angular ventrolateral shoulder; weak ribs and ventrolateral nodes. Prongs of external lobe bifid; altogether 16 notches of E, A and L lobes. Etymology From the Latin ‘ petilus ’, meaning ‘slim’, because of the slender conch. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 44A; MB.C.29962. Description Holotype MB.C.29962 is a comparatively well-preserved individual with 80 mm conch diameter and allows the study of two whorls (Fig. 44A). Nearly three quarters of the last volution belong to the body chamber. The adult whorl profile is compressed (ww/wh ~ 0.80) and weakly trapezoidal with gently diverging flanks, a subangular ventrolateral shoulder and a flattened but still weakly rounded venter. The sculpture of the last phragmocone volution has 15 coarse ventrolateral nodes. With beginning of the terminal body chamber, these nodes become weaker and more numerous; at the end of the body chamber, ten faint nodes occur in a quarter of a volution. The suture line of the holotype has an external lobe with weakly converging flanks. Its prongs are slightly pouched and possess three denticles. The ventrolateral saddle is inflated and followed by an asymmetric, pouched adventive lobe with oblique, weakly rounded base and seven small notches. The lateral lobe is generally V-shaped with weakly diverging flanks; it has six notches (Fig. 44B). Remarks Clivotirolites petilus gen. et sp. nov. differs in the lower number (about 15 per volution) of ventrolateral nodes from the species of Clivotirolites decorates gen. et sp. nov. Stratigraphic range The single specimen was collected from float of the upper part of the Hambast Formation.

Abichites abichi Shevyrev, 1965 Fig. 56; Table 30 Abichites abichi Shevyrev, 1965: 181, pl. 24 fig. 4. Xenodiscus aff. kapila – Stoyanow 1910: 87, pl. 9 fig. 3. Abichites abichi – Shevyrev 1968: 96, pl. 4 fig. 4. — Korn & Ghaderi in Korn et al. 2016: 878, textfig. 35. Diagnosis Species of Abichites with conch reaching 75 mm dm. Subadult stage with subquadrate, weakly depressed to compressed whorl profile (ww/wh =1.20) and slightly flattened venter; with 15 rounded straight ribs on the flanks. Adult stage with quadrate whorl profile (ww/wh =1.00), parallel flanks and flattened venter; with weak rounded ribs on the flanks. Prongs of external lobe usually simple; 7–12 notches of E, A and L lobes. Type material Holotype AZERBAIJAN • Nakhichevan Province, Dorasham 2 section; Paratirolites Limestone; illustrated by Shevyrev (1965: pl. 24 fig. 4); PIN 1252/137. Material examined IRAN • 4 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.30019 to MB.C.30022 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.30023 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30024 to MB.C.30028 • 6 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30029 to MB.C.30034 • 4 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30035 to MB.C.30038 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30039 to MB.C.30040. Description Specimen MB.C.30039 has 44 mm diameter and is a fragment of half a volution (Fig. 56A). However, it allows the study of the conch shape with a nearly quadrate whorl profile. On half of a volution there are twelve sharp radial ribs, which are coarsest in the outer flank. Specimen MB.C.30019 has 40 mm conch diameter and allows the study of only one volution (Fig. 56B). The whorl profile is subtrapezoidal with broadly rounded umbilical margin, weakly converging flanks, a subangular ventrolateral shoulder and a flat venter. The sculpture is weaker than in the first specimen and the ribs are coarsest on the inner flank. The suture line of specimen MB.C.30019 is characterised by a large external lobe, of which the flanks stand parallel in the middle part. Its prongs are narrowly V-shaped and non-serrated. After a ventrolateral saddle that is as wide as the external lobe follow a small, nearly parallel-sided adventive lobe with three notches, a small lateral saddle and a small parallel-sided lateral lobe with three notches (Fig. 56C). Remarks Abichites abichi differs from the otherwise similar species A. subtrapezoidalis in the quadrate adult whorl profile (subtrapezoidal in A. subtrapezoidalis). The body chamber of A. abichi has stronger ribs than A. subtrapezoidalis does. A. alibashiensis has a similar body chamber, but differs from A. abichi in the coarse ventrolateral nodes of the subadult stage. Stratigraphic range Upper part of the Hambast Formation; 1.20 to 0.05 m below the extinction horizon (Abichites abichi Zone to Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 87-89, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135.","Shevyrev A. A. 1968. Triasovye ammonoidei Yuga SSSR. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 119: 1 - 272.","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Lutites paucis Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 9C85D567-DA54-463F-8CE3-BFDBABD83928 Fig. 50; Table 25 Diagnosis Species of Lutites gen. nov. with conch reaching 70 mm dm. Subadult stage with circular, slightly compressed whorl profile (ww/wh =0.90–1.00) and broadly rounded venter; very coarse conical ventrolateral nodes. Adult stage with subtrapezoidal, slightly compressed whorl profile (ww/wh =0.80), flattened venter and subangular ventrolateral shoulder; without ribs or nodes. Prongs of external lobe simple; altogether 11 notches of E, A and L lobes. Etymology From the Latin ‘ paucis ’, meaning ‘fewer’, because of the low number of lobes and nodes. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 2.45 m below the top; illustrated in Fig. 50A; MB.C.29991. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29992 to MB.C.29993. Description Holotype MB.C.29991 is a partly corroded specimen with 64 mm conch diameter with two thirds of the last volution belonging to the body chamber (Fig. 50A). At the end of the body chamber, the whorl profile is subtrapezoidal with a broadly rounded umbilical margin, converging flanks, a narrowly rounded ventrolateral shoulder and a flattened venter. By contrast, the whorl profile is trapezoidal at the beginning of the body chamber and subcircular in the last portion of the phragmocone. The sculpture of the phragmocone consists of very coarse conical ventrolateral nodes (eight per volution). The body chamber appears to be smooth. The suture line of the holotype has a large external lobe with weakly diverging flanks and simple, narrow and lanceolate prongs. The ventrolateral saddle is narrowly rounded and the lateral saddle is weakly inflated. Both the adventive lobe and the lateral lobe are parallel-sided and both characterised by five notches that are very dissimilar in size (Fig. 50B). Remarks Lutites paucis gen. et sp. nov. differs from the other species of the genus in the stouter conch, the only weakly flattened venter and the lower number of conical nodes. The new species may represent an intermediate form between Alibashites and Lutites gen. nov. Stratigraphic range Upper part of the Hambast Formation; 2.45 m to 1.20 m below the extinction horizon (uppermost part of the Paratirolites kittli Zone to the Lutites profundus Zone).

Paratirolites serus Korn & Ghaderi in Korn et al., 2016 Fig. 40; Table 19 Paratirolites serus Korn & Ghaderi in Korn et al., 2016: 868, text-fig. 27. Diagnosis Paratirolites with conch reaching 80 mm dm. Subadult stage with weakly to strongly trapezoidal, depressed whorl profile (ww/wh =1.40–1.80) and broadly rounded venter; 12–14 coarse conical ventrolateral nodes. Adult stage with circular or slightly trapezoidal, weakly to moderately depressed whorl profile (ww/wh =1.20–1.60), flattened venter and subangular to angular ventrolateral shoulder; numerous ribs on the flank. Prongs of external lobe simple; altogether 6–9 notches of E, A and L lobes. Material examined Holotype IRAN • East Azerbaijan, Ali Bashi N section; Paratirolites Limestone, 0.65 m below the top; illustrated by Korn & Ghaderi in Korn et al. (2016: text-fig. 27a); MB.C.25300. Additional material IRAN • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29934 • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29935 to MB.C.29936 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29937. Description Specimen MB.C.29936 has 62 mm conch diameter and is the largest available specimen (Fig. 40A). It is corroded but still rather well preserved and allows the study of three whorls. A little more than half a volution belongs to the body chamber. The weakly depressed whorl profile (ww/wh = 1.05) of the body chamber is trapezoidal with weakly diverging flanks and the flattened venter is separated from the flanks by a subangular ventrolateral shoulder. The sculpture changes, on the last portion of the phragmocone, from a subadult stage with 14 coarse ventrolateral nodes per volution to the adult stage with numerous weak radial ribs with weak ventrolateral nodes on the flank. Specimen MB.C.29934 is a smaller individual with 48 mm conch diameter and is better preserved than the preceding one with half a volution of the body chamber preserved (Fig. 40B). It shows a weakly trapezoidal, weakly depressed body chamber whorl profile (ww/wh=1.05) with rounded ventrolateral shoulder and a broadly rounded venter, while the phragmocone has a more depressed whorl profile (ww/ wh= 1.40). The sculpture in the subadult stage has twelve coarse ventrolateral nodes, which become weaker and radially elongated on the body chamber, where they finally disappear and instead shallow radial dorsolateral ribs take place. The suture line of specimen MB.C.29934 has an external lobe with nearly parallel flanks and unserrated, narrow lanceolate prongs. The ventrolateral saddle is weakly asymmetric and the adventive lobe is strongly asymmetric with a steep ventral flank, four little notches and an oblique dorsal side. The lateral lobe possesses only three small notches (Fig. 40D). Remarks Paratirolites serus is the stratigraphically youngest species of the genus in the Baghuk Mountain sections. It differs from the other by its small conch size and the simple suture line with unsubdivided prongs of the external lobe. Subadult specimens of P. serus resemble some stratigraphically older species such as P. kittli and particularly P. vediensis. Paratirolites serus differs from these in the smaller conch diameter (40 mm maximum phragmocone diameter in contrast to 70–80 mm in P. kittli and P. vediensis) and a simpler suture line with unsubdivided prongs of the external lobe. Stratigraphic range Upper part of the Hambast Formation; 0.75 to 0.60 m below the extinction horizon (Abichites stoyanowi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 59-61, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Abichites ovalis Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 983F405A-033F-49EB-8D75-E5F917F23D86 Fig. 58; Table 32 Diagnosis Species of Abichites with conch reaching 85 mm dm. Subadult stage with subcircular, weakly compressed whorl profile (ww/wh =0.80–1.00) and rounded venter; about 12 shallow and rounded radial ribs. Adult stage with parallel sided or weakly trapezoidal, weakly compressed whorl profile (ww/wh =0.80–1.00), rounded venter and narrowly rounded ventrolateral shoulder; shallow wide ribs on the flank. Prongs of external lobe simple; altogether 7–11 notches of E, A and L lobes. Etymology From the Latin ‘ ovalis ’, meaning ‘oval’, because of the oval whorl profile in the subadult stage. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain B section; Hambast Formation; illustrated in Fig. 58A; MB.C.30059. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.30060 • 3 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30061 to MB.C.30063 • 6 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30064 to MB.C.30069 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30070 to MB.C.30071 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.30072. Description Holotype MB.C.30059 is a rather well-preserved specimen with 62 mm conch diameter; the phragmocone diameter is 44 mm and 190 degrees of the body chamber are preserved (Fig. 58A). The adult whorl profile is weakly trapezoidal with gently diverging, slightly concave flanks, a rounded ventrolateral shoulder and a slightly flattened but broadly rounded venter. One volution earlier, the whorl profile is subcircular and weakly compressed (ww/wh = 0.90). The sculpture on the phragmocone consists of 12 shallow and rounded radial ribs per volution. They become weaker on the body chamber but develop to elongate, rather sharp nodes on the outer flank. The suture line of holotype MB.C.30059 shows an external lobe with flanks standing parallel in the lower part and diverging in the upper part. The prongs are very narrow, lanceolate and not secondarily serrated. On the outer flank lies the symmetric adventive lobe with semicircular, strongly serrated base; it is followed by a rather high lateral saddle and a parallel-sided lateral lobe with five small notches (Fig. 58C). Paratype MB.C.30071 has 60 mm conch diameter and is, with its weakly trapezoidal whorl profile and the general conch shape very similar to the previous one but has a more slender conch (ww/wh =0.85). It shows, in the inner whorls, weak ventrolateral nodes connected to the shallow radial ribs (Fig. 58B). Remarks Abichites ovalis sp. nov. closely resembles A. paucinodum, but differs in the whorl profile (diverging flanks in A. ovalis sp. nov. but converging in A. paucinodum), the wider umbilicus in A. ovalis sp. nov. and in the rounded venter in A. ovalis sp. nov. (flat in A. paucinodum). Stratigraphic range Upper part of the Hambast Formation; 0.75 to 0.40 m below the extinction horizon (Abichites stoyanowi Zone to lowest part of the Arasella minuta Zone).

Genus Lutites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 5B2AB929-F903-46B0-B108-A1C57458A1DA Type species Lutites lyriformis Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Genus of the family Dzhulfitidae with large conch; maximum adult diameters about 140 mm. Adult stage with compressed, subtrapezoidal whorl profile; flanks concave, venter flat or concave. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs or delicate plications. Suture line with deep external lobe (deeper than adventive lobe), ventral prongs unsubdivided or notched. Etymology Named after the Lut Desert, which extends to the North-East of Baghuk Mountain. Included species Alibashites stepanovi Korn & Ghaderi in Korn et al., 2016; Lutites paucis Korn & Hairapetian gen. et sp. nov.; Lutites alius Korn & Hairapetian gen. et sp. nov.; Lutites profundus Korn & Hairapetian gen. et sp. nov.; Lutites lyriformis Korn & Hairapetian gen. et sp. nov.; Lutites plicatus Korn & Hairapetian gen. et sp. nov. Remarks Lutites gen. nov. differs from Paratirolites and related genera in the compressed subtrapezoidal, pearshaped whorl profile and in the very deep external lobe., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 76, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 0A7C7606-6D92-4630-BAF3-1A8E30B6486D Figs 41–43; Table 20 Diagnosis Species of Clivotirolites gen. nov. with conch reaching 110 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh=1.10–1.20) and slightly flattened venter; about 20–24 weak ventrolateral spines per volution. Adult stage with parallel or weakly diverging flanks and weakly compressed whorl profile (ww/wh =0.70–0.80), flattened, slightly tectiform venter and angular ventrolateral shoulder; numerous rounded ribs on flank ending in sharp ventrolateral nodes. Prongs of external lobe variable, simple to digitate; altogether 10–17 notches of E, A and L lobes. Etymology From the Latin ‘ decoratus ’, meaning ‘decorated’, because of the sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 1.35 m below the top; illustrated in Fig. 41A; MB.C.29941. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29938 • 2 specimens; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29939 to MB.C.29940 • 14 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29942 to MB.C.29955 • 2 specimens; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29956 to MB.C.29957 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29958 to MB.C.29959 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29960 to MB.C.29961. Description Holotype MB.C.29941 is a rather well-preserved specimen of 64 mm conch diameter and shows all typical characters for the species (Fig. 41A). The specimen shows one and a half whorls preserved. It displays septal crowding and the end of the phragmocone appears at 38 mm diameter.About 210 degrees of the last volution belong to the body chamber, which shows a weakly trapezoidal whorl profile with weakly concave, weakly diverging flanks, a subangular ventrolateral shoulder and a flattened tectiform venter. The whorl belonging to the phragmocone has a more pronounced trapezoidal profile with convex, diverging flanks. The sculpture on the phragmocone part consists of rather small, pointed ventrolateral tubercles, 22 per volution. On the body chamber, these tubercles are connected with weak and rounded radial ribs, which are strongest on the inner flank. Paratype MB.C.29946 is a similar specimen with 58 mm conch diameter and the last septum at 33 mm dm (Fig. 41B). It possesses a nearly quadrate whorl cross section of the body chamber but also shows that the profile is trapezoidal in an earlier growth stage. On the last volution of the phragmocone, 20 sharp ventrolateral nodes can be counted. Larger paratypes such as MB.C.29947 (50 mm phragmocone diameter), MB.C.29944 (52 mm; Fig. 42B), MB.C.29938 (52 mm; Fig. 42A) and MB.C.29958 (53 mm; Fig. 43A) are generally similar but show stronger diverging flanks of the body chamber. Suture lines were drawn from four specimens, all of them show principally similar outlines with some common characters (Fig. 43B–E). The external lobe is not as deep as the adventive lobe, it is wider at its base, from where the flanks converge towards the ventrolateral saddle. All four specimens show asymmetric notching of its prongs; usually there are three notches but also more intense serration is possible. The ventrolateral saddle is usually slightly inflated and the adventive lobe is parallel-sided with five or six notches at its base. Remarks Clivotirolites decoratus gen. et sp. nov. differs in the large number (more than 20 per volution) of ventrolateral nodes from the species of Clivotirolites petilus gen. et sp. nov. (about 15 nodes per volution). Stratigraphic range Upper part of the Hambast Formation; 2.05 to 1.10 m below the extinction horizon (Alinashites ferdowsii Zone to Abichites abichi Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 62-64, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Lutites plicatus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 1713B70A-3C3C-4FCD-9827-BBBA6F989797 Fig. 55; Table 29 Diagnosis Species of Lutites gen. nov. with conch reaching 120 mm dm. Subadult stage with oval, compressed whorl profile (ww/wh=0.75) and rounded venter; about 12 radial plications per volution. Adult stage with subtrapezoidal, compressed whorl profile (ww/wh =0.72), weakly concave flanks, flattened venter and subangular ventrolateral shoulder; numerous weak and densely arranged plications and small ventrolateral nodes. Prongs of external lobe multiply serrated; altogether 20 notches of E, A and L lobes. Etymology From the Latin ‘ plicatus ’, meaning ‘corrugated’, referring to the dense ribbing. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 55A; MB.C.30018. Description Holotype MB.C.30018 is a specimen with 84 mm diameter, of which the last 120 degrees belong to the body chamber (Fig. 55A). Nearly one volution of the phragmocone can be studied; at this stage the whorl profile is oval and compressed with a rounded venter. The ornament consists of nearly 20 faint radial plications, which are coarsest near the umbilicus and in the ventrolateral area, where they form weak nodes. The ornament is weaker on the body chamber, where the plications are more numerous. The whorl profile is subtrapezoidal and compressed (ww/wh=0.72) in the adult stage with a rounded umbilical margin, weakly concave flanks and a flattened venter. The suture line of the holotype has a narrow external lobe with asymmetric, multiply serrated prongs (four and six small notches), an asymmetric and inflated ventrolateral saddle and multiply serrated adventive and lateral lobes (Fig. 55B). Both lobes show notches of various size. Remarks Lutites plicatus gen. et sp. nov. differs in the shape of the flattened, but convex venter from most of the other species of the genus (L. lyriformis gen. et sp. nov., L. profundus gen. et sp. nov., L. alius gen. et sp. nov.), which possess a completely flat or even concave venter. Stratigraphic range Upper part of the Hambast Formation; horizon unknown.

Abichites infirmis Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: DF627B6D-3446-4B02-89B2-81ED420C876D Fig. 60; Table 34 Diagnosis Species of Abichites with conch reaching 50 mm dm. Subadult stage with oval, weakly compressed whorl profile (ww/wh =0.60–0.80) and rounded venter; very weak, rounded nodes. Adult stage with weakly subtrapezoidal, weakly compressed whorl profile (ww/wh =0.60–0.80) with weakly converging flat flanks, weakly flattened venter and narrowly subangular ventrolateral shoulder. Sculpture with faint plications. Prongs of external lobe simple; altogether 5–6 notches of E, A and L lobes. Etymology From the Latin ‘ infirmis ’, meaning ‘barely visible’, because of the weak sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain G section; Hambast Formation, 0.40 m below the top; illustrated in Fig. 60A; MB.C.30086. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.30083 • 1 specimen; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.30084 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30085 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30087 to MB.C.30088. Description Holotype MB.C.30086 is a rather well-preserved specimen with 42 mm conch diameter (Fig. 60A). The whorl profile of the body chamber is subtrapezoidal with a narrowly rounded umbilical margin, flat and weakly converging flanks, a subangular ventrolateral shoulder and a flattened but still broadly rounded venter. Its ww/wh ratio is 0.70 and the whorl embraces the preceding one only weakly. The shell appears to be smooth except for very faint plications. The suture line of the holotype has a parallel-sided external lobe that has a width/depth ratio of ~ 0.75. It is deeper than the adventive lobe and possesses unsubdivided lanceolate prongs. The ventrolateral saddle is inverted U-shaped and symmetric and the weakly asymmetric, parallel-sided adventive lobe is subdivided into two secondary notches. The narrow lateral saddle is also inverted U-shaped and followed by a parallel-sided lateral lobe with two notches (Fig. 60B). Remarks Abichites infirmis differs from most of the species of the genus in the lack of coarse ribs or nodes. Similar species are A. paucinodus (which possesses few coarse nodes on the flanks) and A. terminalis (with an oval whorl profile). Stratigraphic range Upper part of the Hambast Formation; 0.40 to 0.05 m below the extinction horizon (Arasella minuta Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 96-97, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Genus Lutites Korn & Hairapetian gen. nov. urn:lsid:zoobank.org:act: 5B2AB929-F903-46B0-B108-A1C57458A1DA Type species Lutites lyriformis Korn & Hairapetian gen. et sp. nov., Baghuk Mountain. Diagnosis Genus of the family Dzhulfitidae with large conch; maximum adult diameters about 140 mm. Adult stage with compressed, subtrapezoidal whorl profile; flanks concave, venter flat or concave. Subadult stage with conical ventrolateral nodes, adult stage with weak ribs or delicate plications. Suture line with deep external lobe (deeper than adventive lobe), ventral prongs unsubdivided or notched. Etymology Named after the Lut Desert, which extends to the North-East of Baghuk Mountain. Included species Alibashites stepanovi Korn & Ghaderi in Korn et al., 2016; Lutites paucis Korn & Hairapetian gen. et sp. nov.; Lutites alius Korn & Hairapetian gen. et sp. nov.; Lutites profundus Korn & Hairapetian gen. et sp. nov.; Lutites lyriformis Korn & Hairapetian gen. et sp. nov.; Lutites plicatus Korn & Hairapetian gen. et sp. nov. Remarks Lutites gen. nov. differs from Paratirolites and related genera in the compressed subtrapezoidal, pearshaped whorl profile and in the very deep external lobe.

Paratirolites baghukensis Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 27027DEB-9F15-4F61-8EA3-3F15E997A1D9 Figs 33–34; Table 16 Diagnosis Species of Paratirolites with conch reaching 130 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh =1.00–1.20) and rounded venter; about 12 moderately coarse ribs ending in conical ventrolateral nodes per volution. Adult stage with trapezoidal and weakly compressed whorl profile (ww/wh =0.70–0.90), flattened tectiform venter and subangular ventrolateral shoulder; numerous sharp ribs ending in sharp radially elongated ventrolateral nodes. Prongs of external lobe usually trifid; altogether 13–15 notches of E, A and L lobes. Etymology Named after the occurrence at Baghuk Mountain. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain G section; Hambast Formation; illustrated in Fig. 33A; MB.C.29868. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29849 to MB.C.29850 • 4 specimens; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29851 to MB.C.29854 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29855 to MB.C.29859 • 8 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29860 to MB.C.29867 • 7 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29869 to MB.C.29875 • 1 specimen; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29876 • 3 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29877 to MB.C.29879. Description Holotype MB.C.29868 has 70 mm conch diameter and is fairly well-preserved but slightly corroded. The last third of the terminal volution belongs to the body chamber (which is thus only incompletely preserved); adulthood of the specimen is obvious because of the crowding of the last septa of the phragmocone (Fig. 33A). The penultimate of the preserved whorls shows a trapezoidal profile with a rounded venter. The body chamber finally shows a trapezoidal whorl profile with weakly diverging flanks, an angular ventrolateral shoulder and a nearly flat, tectiform venter. The sculpture changes with the beginning of the terminal body chamber, but this change is continuous without sudden breaks. One subadult volution has ten isolated spiny ventrolateral nodes.Already towards the end of the phragmocone, these nodes become radially elongate and form coarse ribs across the flank. At the beginning of the terminal body chamber, the ventrolateral nodes weaken and finally, only sharp ribs, about 15 per quarter of a volution, are present. The suture line of holotype MB.C.29868 shows a slightly pouched external lobe with trifid, asymmetric prongs. The ventrolateral saddle is weakly inflated, the adventive and lateral lobes are flattened at the base and subdivided in five and four notches, respectively (Fig. 33C). Paratype MB.C.29860 (Fig. 33B) is smaller with 61 mm conch diameter but confirms the description given for the holotype. It shows four whorls preserved and thus allows for the study of the sculpture of the inner volutions. The two larger paratypes MB.C.29874 and MB.C.29850 (Fig. 34) show then the transformation into the weakly ornamented adult stage, in which a tendency towards a parallel-sided whorl profile is visible. Remarks Paratirolites baghukensis sp. nov. belongs to the slender species of the genus. It differs from the other species of the genus in the sharp ribs on the adult body chamber. Paratirolites multiconus also has a slender conch, but differs from P. baghukensis sp. nov. in the much more numerous ventrolateral nodes. Stratigraphic range Upper part of the Hambast Formation; 3.10 to 2.35 m below the extinction horizon (Paratirolites kittli Zone).

Esfahanites armatus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: DACEC5B2-4BE5-40D0-AA2B-578741DC349C Figs 45–46; Table 22 ? Paratirolites sp. Ghaedi et al., 2009: pl. 2 figs 1, 4. ? Paratirolites waageni – Zakharov & Mussavi Abnavi 2013: text-fig. 12c. Diagnosis Species of Esfahanites gen. nov. with conch reaching 130 mm dm. Subadult stage with strongly trapezoidal, moderately depressed whorl profile (ww/wh =1.50–1.70) and rounded to tectiform venter; about 12 coarse ventrolateral nodes per volution. Adult stage with trapezoidal and moderately depressed whorl profile (ww/wh =1.50–1.70), flattened tectiform venter and angular ventrolateral shoulder; coarse rursiradiate ribs. Prongs of external lobe trifid or multiply serrated; altogether 15–21 notches of E, A and L lobes. Etymology From the Latin ‘ armatus ’, meaning ‘armed’, because of the coarse spiny sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 45; MB.C.29964. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29963 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29965 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29966 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29967 to MB.C.29968. Description Holotype MB.C.29964 is the best-preserved specimen; it has 106 mm conch diameter and allows the study of two whorls (Fig. 45A). The diameter of the phragmocone is 68 mm and the body chamber has a length of about 270 degrees. Throughout the last two preserved volutions, the whorl profile is trapezoidal, but with the flanks less strongly diverging at the end of growth. The venter is broadly rounded in the penultimate whorl and becomes more flattened on the terminal body chamber. The sculpture on the phragmocone has 14 coarse conical ventrolateral nodes on the last volution; these nodes show a rib-like elongation towards the umbilicus. The sculpture on the body chamber shows variably coarse, rounded ribs on the flank; they turn backward near the umbilicus and are strengthened in the ventrolateral area. The suture line of holotype MB.C.29964 is remarkable because of its very small external lobe, which is wider at the base and is constricted in the upper part (Fig. 45B). The prongs are inclined ventrally and possess five little notches each. The ventrolateral saddle is small and inflated. The asymmetric adventive lobe is strongly serrated with little notches being developed on the ventral side up to half of the lobe depth. The lateral lobe is slightly less strongly serrated than the adventive lobe. Paratype MB.C.29966 is an incomplete but otherwise rather well-preserved individual of 101 mm conch diameter; it allows the study of two and a half whorls (Fig. 46A). The body chamber begins at 60 mm conch diameter, meaning that the preserved part of the body chamber has a length of 270 degrees and is nearly complete. Conch shape changes can be observed during the last whorl of the specimen. While the whorl profile is circular with rounded venter at the binning of the last whorl, it changes to tectiform and has, at the end of growth, an almost flat venter, separated from the flat and diverging flanks by an angular ventrolateral shoulder. The sculpture of the phragmocone part of the specimen shows coarse conical nodes in the ventrolateral area, 12 on the last volution and 10 on the penultimate volution. At the beginning of the terminal body chamber, these nodes are transformed into elongate nodes and finally rounded ribs, which turn from the umbilicus in backward direction. Finally, at the end of the body chamber these ribs become weaker and only shallow rounded ribs and weak ventrolateral nodes can be seen. The suture line of paratype MB.C.29966 shows a very small external lobe that has not half the depth of the adventive lobe (Fig. 46B). It is parallel-sided with parallel-sided prongs that possess three small notches. The ventrolateral saddle is inverted U-shaped with a much longer dorsal flank. The adventive lobe has seven small notches and the lateral lobe five, both lobes are asymmetric and separated by a low semi-circular lateral saddle. Remarks Esfahanites armatus gen. et sp. nov. differs in the very small external lobe from all the species of Paratirolites. It can therefore not be confused with any other species. Stratigraphic range Upper part of the Hambast Formation; the only in situ collected specimen comes from 2.00 m below the extinction horizon (Alibashites ferdowsii Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 68-71, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Ghaedi M., Mousavi N. & Yazdi M. 2009. Scrutiny and biozonation of Permian - Triassic boundary in Benarizeh area, north of Abadeh. Second Conference of the Iranian Paleontological Association: 1 - 4."]}

Genus Stoyanowites Korn in Ghaderi et al., 2014 Type species Paratirolites dieneri Stoyanow, 1910, by original designation Included species Stoyanowites aspinosus Korn & Ghaderi in Korn et al., 2016; Paratirolites dieneri Stoyanow, 1910; Stoyanowites parallelus Korn & Hairapetian sp. nov. Diagnosis Genus of the family Dzhulfitidae with moderately large conch; maximum adult diameters are between 80 and 95 mm. Subadult and adult stages with rectangular and compressed whorl profile. Subadult stage with weak to moderately strong lateral ribs; adult stage with weak ornament. Suture line with an external lobe that is much shorter than the adventive lobe. Remarks Stoyanowites is separated from most of the other genera of the family Dzhulfitidae by the short external lobe, which does not reach the depth of the adventive and lateral lobes. Only Dzhulfites is similar in this respect, but this genus is characterised by coarse ventrolateral nodes absent in Stoyanowites., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 97, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Ghaderi A., Leda L., Schobben M., Korn D. & Ashouri A. R. 2014. High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts, and ammonoids. Fossil Record 15: 41 - 57. https: // doi. org / 10.5194 / fr- 17 - 41 - 2014","Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135.","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Clivotirolites decoratus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 0A7C7606-6D92-4630-BAF3-1A8E30B6486D Figs 41–43; Table 20 Diagnosis Species of Clivotirolites gen. nov. with conch reaching 110 mm dm. Subadult stage with trapezoidal, weakly depressed whorl profile (ww/wh=1.10–1.20) and slightly flattened venter; about 20–24 weak ventrolateral spines per volution. Adult stage with parallel or weakly diverging flanks and weakly compressed whorl profile (ww/wh =0.70–0.80), flattened, slightly tectiform venter and angular ventrolateral shoulder; numerous rounded ribs on flank ending in sharp ventrolateral nodes. Prongs of external lobe variable, simple to digitate; altogether 10–17 notches of E, A and L lobes. Etymology From the Latin ‘ decoratus ’, meaning ‘decorated’, because of the sculpture. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 1.35 m below the top; illustrated in Fig. 41A; MB.C.29941. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29938 • 2 specimens; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29939 to MB.C.29940 • 14 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29942 to MB.C.29955 • 2 specimens; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29956 to MB.C.29957 • 2 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29958 to MB.C.29959 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29960 to MB.C.29961. Description Holotype MB.C.29941 is a rather well-preserved specimen of 64 mm conch diameter and shows all typical characters for the species (Fig. 41A). The specimen shows one and a half whorls preserved. It displays septal crowding and the end of the phragmocone appears at 38 mm diameter.About 210 degrees of the last volution belong to the body chamber, which shows a weakly trapezoidal whorl profile with weakly concave, weakly diverging flanks, a subangular ventrolateral shoulder and a flattened tectiform venter. The whorl belonging to the phragmocone has a more pronounced trapezoidal profile with convex, diverging flanks. The sculpture on the phragmocone part consists of rather small, pointed ventrolateral tubercles, 22 per volution. On the body chamber, these tubercles are connected with weak and rounded radial ribs, which are strongest on the inner flank. Paratype MB.C.29946 is a similar specimen with 58 mm conch diameter and the last septum at 33 mm dm (Fig. 41B). It possesses a nearly quadrate whorl cross section of the body chamber but also shows that the profile is trapezoidal in an earlier growth stage. On the last volution of the phragmocone, 20 sharp ventrolateral nodes can be counted. Larger paratypes such as MB.C.29947 (50 mm phragmocone diameter), MB.C.29944 (52 mm; Fig. 42B), MB.C.29938 (52 mm; Fig. 42A) and MB.C.29958 (53 mm; Fig. 43A) are generally similar but show stronger diverging flanks of the body chamber. Suture lines were drawn from four specimens, all of them show principally similar outlines with some common characters (Fig. 43B–E). The external lobe is not as deep as the adventive lobe, it is wider at its base, from where the flanks converge towards the ventrolateral saddle. All four specimens show asymmetric notching of its prongs; usually there are three notches but also more intense serration is possible. The ventrolateral saddle is usually slightly inflated and the adventive lobe is parallel-sided with five or six notches at its base. Remarks Clivotirolites decoratus gen. et sp. nov. differs in the large number (more than 20 per volution) of ventrolateral nodes from the species of Clivotirolites petilus gen. et sp. nov. (about 15 nodes per volution). Stratigraphic range Upper part of the Hambast Formation; 2.05 to 1.10 m below the extinction horizon (Alinashites ferdowsii Zone to Abichites abichi Zone).

Dzhulfites brevisellatus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: 1FDCA9D7-546A-48FA-94E1-0053EB7F7C23 Fig. 23; Table 8 Diagnosis Species of Dzhulfites with conch reaching 160 mm dm. Subadult stage with weakly trapezoidal, moderately depressed whorl profile (ww/wh =1.50–1.80) and broadly rounded venter; 10–13 coarse conical nodes on midflank per volution. Adult stage with numerous weak ribs and small ventrolateral nodes. Prongs of external lobe bifid or trifid; altogether 15–20 notches of E, A and L lobes. Etymology From the Latin ‘ brevis ’, meaning ‘low’ and ‘ sella ’, meaning ‘saddle’, after the low amplitude of lobes and saddles in the suture line. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain H section; Hambast Formation, 5.70 m below top; illustrated in Fig. 23B; MB.C.29748. Paratypes IRAN • 1 specimen; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29746 • 1 specimen; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29747. Description The large paratype MB.C.29746 has 125 mm conch diameter, of which half of the last whorl belongs to the strongly weathered body chamber (Fig. 23A). The phragmocone is 90 mm in diameter and is partly crushed; it shows that the whorl profile is trapezoidal with a broadly rounded venter. The sculpture changes dramatically throughout ontogeny. At 40 mm dm there are 13 coarse, spiny ventrolateral nodes per volution; these nodes become weaker and more numerous so that the last 120° of the phragmocone have about 20 weak ventrolateral nodes, which are connected with weak and slightly sinuous radial ribs on the flank. The suture line of the specimen suffered from weathering, but, most probably, the prongs of the very short external lobe are bifid. Holotype MB.C.29748 is a fairly well-preserved, fully septate specimen with 51 mm conch diameter (Fig. 23B). It has a weakly trapezoidal whorl profile with broadly rounded flanks and venter. The sculpture shows about ten coarse conical nodes, which are positioned on the midflank, on the last preserved volution. The suture line of the holotype shows an external lobe with subparallel, weakly diverging flanks and weakly asymmetric, trifid prongs. All three saddles of the outer suture line are similar in shape, being inverted U-shaped and broadly rounded. The adventive lobe and the lateral lobe are similar in shape with parallel flanks and many small notches at the base. They differ in their depth; the adventive lobe is much deeper than the external lobe (Fig. 23C). Remarks Among the species of Dzhulfites, only D. nodosus is similar in conch shape and sculpture, but this species shows an applanate venter in the adult stage. Dzhulfites brevisellatus sp. nov. differs from D. spinosus and the other species of the genus in the suture line, which in D. brevisellatus sp. nov. shows much wider elements; particularly the saddles are wider than the lobes. In the shape of the suture line, D. brevisellatus sp. nov. has some resemblance to representatives of the family Xenodiscidae. Stratigraphic range Upper part of the Hambast Formation; 5.70 to 4.70 m below the extinction horizon (Dzhulfites spinosus Zone to Shevyrevites shevyrevi Zone).

Paratirolites lanceolobatus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: EA9569C5-F88F-4487-A528-878A60FB4CE1 Fig. 25; Table 10 Diagnosis Species of Paratirolites with conch reaching 90 mm dm. Subadult stage with weakly trapezoidal, weakly depressed whorl profile (ww/wh =1.10–1.30) and broadly rounded venter; about 12 radial ribs ending in coarse conical ventrolateral nodes per volution. Adult stage with rectangular and weakly compressed whorl profile (ww/wh = 0.85–1.00), flattened, weakly tectiform venter and subangular ventrolateral shoulder; numerous rounded ventrolateral nodes. Prongs of external lobe lanceolate and usually unsubdivided; altogether 11–15 notches of E, A and L lobes. Etymology From the Latin’ lancea ’, meaning ‘lance’, because of the lanceolate prongs of the external lobe. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain B section; Hambast Formation; illustrated in Fig. 25A; MB.C.29769. Paratypes IRAN • 18 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29751 to MB.C.29768 • 3 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29770 to MB.C.29772 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29773 • 2 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29774 to MB.C.29775. Description Holotype MB.C.29769 is a comparatively well-preserved individual with 73 mm conch diameter; a little more than half of the last whorl belongs to the partly corroded body chamber. The last two and a half whorls of the phragmocone are better preserved (Fig. 25A). The beginning of the terminal body chamber is at 50 mm diameter; it is well identifiable by the crowding of two septa. This boundary is not identical with spontaneous changes in conch shape and sculpture. At the beginning of the body chamber, the whorl profile is trapezoidal with a flattened, broadly rounded venter; at the end, the whorl profile is slightly trapezoidal with weakly diverging flanks. The rounded, weakly tectiform venter is delimited from the flanks by rounded ventrolateral shoulders. The sculpture shows three ontogenetic stages: (1) The inner whorls (up to 10 mm dm) appear to be smooth without ribs or nodes. (2) The intermediate stage (between 10 and 50 mm dm) shows radial ribs that end in coarse conical ventrolateral nodes (12 per volution in the smaller growth stage and more densely arranged in the larger stage), which weaken at the beginning of the body chamber. (3) The main portion of the body chamber shows smoothening of the sculpture. The suture line of holotype MB.C.29769 possesses a parallel-sided external lobe with very narrow, lanceolate non-serrated prongs. The ventrolateral saddle is rather narrow (much narrower than the external lobe) and weakly inflated; it is followed by a rather broad, drop-shaped adventive lobe that is multiply serrated, a wide and broadly rounded lateral saddle and a V-shaped, weakly serrated lateral lobe (Fig. 25E). The smaller paratype MB.C.29770 with 55 mm conch diameter resembles the holotype in conch proportions and sculpture, but appears to enter the adult growth stage at a smaller conch diameter with a maximum phragmocone diameter of 38 mm (Fig. 25C). Very coarse ventrolateral nodes are present in an interval of about 270 degrees of the phragmocone. The suture line of this specimen also shows nonserrated prongs of the external lobe. Suture lines of the further two paratypes MB.C.29751 and MB.C.29757 (both at about 11 mm whorl height) demonstrate, when compared with the holotype, the variability within the Material examined Common to all three specimens are the simple prongs of the external lobe, which in the paratypes MB.C.29751 and MB.C.29757 is less deep as the adventive lobe. All three specimens show an asymmetric adventive lobe that appears to be dorsally inclined (Fig. 25E–G). Remarks The new species shows some similarities to species of the genus Dzhulfites, particularly in the reduced depth of the external and lateral lobes. It may thus represent a link between the two genera. The low stratigraphic position at the base of the Paratirolites Limestone supports this hypothesis. Paratirolites lanceolobatus sp. nov. differs in the non-serrated prongs of the external lobe from most of the other species of the genus. Similar species are the Transcaucasian P. trapezoidalis and P. birunii, but both possess an adult body chamber with numerous small nodes at the angular ventrolateral shoulder. Paratirolites robustus sp. nov. is the most similar of the Central Iranian species but differs in the wider whorl profile (ww/wh= 1.20 at 70 mm dm) from P. lanceolobatus sp. nov. (ww/wh =0.85). Furthermore, P. robustus sp. nov. has bifid prongs of the external lobe. Stratigraphic range Upper part of the Hambast Formation; 3.85 to 2.80 m below the extinction horizon (Paratirolites lanceolobatus Zone to lower part of the Paratirolites kittli Zone).

Stoyanowites parallelus Korn & Hairapetian sp. nov. urn:lsid:zoobank.org:act: E6C0C2D8-02EF-4E72-9E1E-78816A50573E Fig. 63; Table 37 Diagnosis Species of Stoyanowites with conch reaching 100 mm dm. Subadult stage with rectangular, compressed whorl profile (ww/wh =0.70), subangular ventrolateral shoulder and flattened venter; 10 rather coarse ventrolateral spines per half volution. Adult stage with parallel-sided, compressed whorl profile (ww/ wh= 0.60), flat venter and angular ventrolateral shoulder; 15 ventrolateral spines per half volution. Prongs of external lobe simple; altogether 13 notches of E, A and L lobes. Etymology Named after the parallel flanks of the whorl profile. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain C section; Hambast Formation, 1.70 m below the top; illustrated in Fig. 63A; MB.C.30102. Paratypes IRAN • 2 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.30103 to MB.C.30104 • 1 specimen; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.30105. Description Holotype MB.C.30102 is an incomplete specimen of 72 mm diameter; it allows the study of conch shape, ornament and suture line. The conch is extremely discoidal (ww/dm= 0.19) with a wide umbilicus (uw/ dm =0.47) and a compressed whorl profile (ww/wh =0.61). It has nearly parallel, slightly concave flanks, a prominent subangular ventrolateral shoulder and a weakly concave venter at the largest diameter (Fig. 63A). The sculpture is dominated by coarse ventrolateral spines, which become weaker towards the end of growth, but of which are still 15 are present on half a volution of the body chamber. They are not always connected with short ribs on the inner flank. The suture line of holotype MB.C.30102 shows the short external lobe, which has simple, weakly pouched prongs. The ventrolateral saddle is rather narrow; the adventive and lateral lobes possess six notches (Fig. 63B). Remarks Stoyanowites parallelus sp. nov. differs from S. dieneri and S. aspinosus in the flat venter and the numerous sharp ventrolateral nodes in the adult stage. Stratigraphic range Upper part of the Hambast Formation; 1.70 m below the extinction horizon (Alibashites ferdowsii Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 101-102, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Clivotirolites petilus Korn & Hairapetian gen. et sp. nov. urn:lsid:zoobank.org:act: 6608C3B9-8B98-4E05-91E6-DA463119748F Fig. 44; Table 21 Diagnosis Species of Clivotirolites gen. nov. with conch reaching 120 mm dm. Subadult stage with trapezoidal, weakly compressed whorl profile (ww/wh=0.90) and flattened tectiform; 15 coarse ventrolateral nodes per volution. Adult stage with trapezoidal, weakly compressed whorl profile (ww/wh =0.70), flat venter and angular ventrolateral shoulder; weak ribs and ventrolateral nodes. Prongs of external lobe bifid; altogether 16 notches of E, A and L lobes. Etymology From the Latin ‘ petilus ’, meaning ‘slim’, because of the slender conch. Material examined Holotype IRAN • Esfahan Province, Baghuk Mountain E section; Hambast Formation; illustrated in Fig. 44A; MB.C.29962. Description Holotype MB.C.29962 is a comparatively well-preserved individual with 80 mm conch diameter and allows the study of two whorls (Fig. 44A). Nearly three quarters of the last volution belong to the body chamber. The adult whorl profile is compressed (ww/wh ~ 0.80) and weakly trapezoidal with gently diverging flanks, a subangular ventrolateral shoulder and a flattened but still weakly rounded venter. The sculpture of the last phragmocone volution has 15 coarse ventrolateral nodes. With beginning of the terminal body chamber, these nodes become weaker and more numerous; at the end of the body chamber, ten faint nodes occur in a quarter of a volution. The suture line of the holotype has an external lobe with weakly converging flanks. Its prongs are slightly pouched and possess three denticles. The ventrolateral saddle is inflated and followed by an asymmetric, pouched adventive lobe with oblique, weakly rounded base and seven small notches. The lateral lobe is generally V-shaped with weakly diverging flanks; it has six notches (Fig. 44B). Remarks Clivotirolites petilus gen. et sp. nov. differs in the lower number (about 15 per volution) of ventrolateral nodes from the species of Clivotirolites decorates gen. et sp. nov. Stratigraphic range The single specimen was collected from float of the upper part of the Hambast Formation., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 66-67, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715

Paratirolites kittli Stoyanow, 1910 Fig. 30; Table 13 Paratirolites Kittli Stoyanow, 1910: 82, pl. 9 figs 1–2. Stephanites ? Waageni Stoyanow, 1910: 89, pl. 8 fig. 3. Pseudotirolites azariani Rostovtsev in Kotlyar et al., 1983: 154, pl. 15 fig. 5. ? Stephanites sp. indet. Stoyanow, 1910: 89, pl. 7 fig. 8. Paratirolites kittli – Spath 1934: 366, text-fig. 125a–d. — Voinova et al. 1947: 169, pl. 40 fig. 4, textfig. 67. — Kummel 1957: 179, text-fig. 1a–b. — Shevyrev 1965: 174, pl. 22 fig. 4; 1968: 90, pl. 3 fig. 1. — Korn & Ghaderi in Korn et al. 2016: 862, text-figs 20–22. Paratirolites waageni – Spath 1934: 367. — Shevyrev 1965: 175, pl. 22 figs 5–6; 1968: 90, pl. 2 figs 6–7. Stephanites ? waageni – Voinova et al. 1947: 167, pl. 40 fig. 3. Diagnosis Species of Paratirolites with conch reaching 190 mm dm. Subadult stage with rounded trapezoidal, weakly to moderately depressed whorl profile (ww/wh =1.30–1.60) and rounded venter; 10–12 very coarse ventrolateral nodes per volution. Adult stage with strongly trapezoidal and weakly depressed whorl profile (ww/wh =1.00–1.30), flattened tectiform venter and subangular ventrolateral shoulder; weak ventrolateral nodes. Prongs of external lobe usually multiply serrated; altogether 12–21 notches of E, A and L lobes. Material examined IRAN • 6 specimens; Esfahan Province, Baghuk Mountain 1 section; Hambast Formation; MB.C.29813 to MB.C.29818 • 1 specimen; Esfahan Province, Baghuk Mountain A section; Hambast Formation; MB.C.29819 • 1 specimen; Esfahan Province, Baghuk Mountain B section; Hambast Formation; MB.C.29820 • 5 specimens; Esfahan Province, Baghuk Mountain C section; Hambast Formation; MB.C.29821 to MB.C.29825 • 3 specimens; Esfahan Province, Baghuk Mountain E section; Hambast Formation; MB.C.29826 to MB.C.29828 • 1 specimen; Esfahan Province, Baghuk Mountain F section; Hambast Formation; MB.C.29829 • 6 specimens; Esfahan Province, Baghuk Mountain G section; Hambast Formation; MB.C.29830 to MB.C.29835 • 2 specimens; Esfahan Province, Baghuk Mountain H section; Hambast Formation; MB.C.29836 to MB.C.29837 • 4 specimens; Esfahan Province, Baghuk Mountain; Hambast Formation; MB.C.29838 to MB.C.29841. Description The large specimen MB.C.29833 (107 mm dm; Fig. 30A) shows septal crowding at a phragmocone diameter of 70 mm, which indicates that the maximum conch diameter of this specimen was about 130 mm. The body chamber of the specimen is corroded but shows that the shape of the whorl profile is trapezoidal in the subadult and adult stage. The venter becomes nearly flat with an angular ventrolateral shoulder during formation of the terminal body chamber but is broadly rounded in the phragmocone. This specimen shows a sculpture of the phragmocone that consist of 12 coarse conical nodes on the outer flanks. These nodes become continuously weaker on the terminal body chamber. The suture line of specimen MB.C.29833 has an external lobe with weakly diverging flanks, nearly parallel-sided prongs with three small notches. The ventrolateral saddle is narrow and dorsally inclined; it continues into the strongly asymmetric adventive lobe, which is asymmetrically serrated with numerous small notches climbing high up the ventral flank. By contrast, the lateral lobe is trifid and nearly symmetric (Fig. 30B). Remarks Paratirolites kittli differs from the otherwise similar P. vediensis in the more rounded whorl profile of the subadult stage, which in the latter is trapezoidal with a flattened venter and flattened diverging flanks. Stratigraphic range Upper part of the Hambast Formation; 3.10 to 2.40 m below the extinction horizon (Paratirolites kittli Zone)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on pages 44-46, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Stoyanow A. A. 1910. On the character of the boundary of Palaeozoic and Mesozoic near Djulfa. Zapiski Imperatorskago St. - Peterburgskago Mineralogiceskago Obscestva [= Verhandlungen der Russisch- Kaiserlichen Mineralogischen Gesellschaft zu St. Petersburg], 2 nd Series 47: 61 - 135.","Kotlyar G. V., Zakharov Y. D., Koczyrkevicz B. V., Kropatcheva G. S., Rostovcev L. O., Chedija I. O., Vuks G. P. & Guseva E. A. 1983. Posdnepermskiy etap evolyutsii organicheskogo mirA. Dzhulficheskiy i dorashamskiy yarusy SSSR. NAUKA, Leningrad.","Spath L. F. 1934. Catalogue of the Fossil Cephalopoda in the British Museum (Natural History). Part IV. The Ammonoidea of the Trias. British Museum (Natural History), Department of Geology, London. https: // doi. org / 10.5962 / bhl. title. 112429","Voinova E. V., Kiparisova L. D. & Robinson V. H. 1947. Klass Cephalopoda. Golovonogie., Atlas rukovodyashtchikh form iskopaemykh faun SSSR. VIII. Triasovaya sistema: 124 - 176, Moskva-Leningrad","Kummel B. 1957. Suborder Ceratitina Hyatt, 1884. In: Moore R. C. (ed.) Treatise on Invertebrate Paleontology. Part L, Mollusca, Ammonoidea 4: L 130 - L 185.","Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}

Genus Abichites Shevyrev, 1965 Type species Kashmirites ? stoyanowi Kiparisova in Voinova et al., 1947, by original designation. Included species Abichites abichi Shevyrev, 1965; Abichites alibashiensis Korn & Ghaderi in Korn et al., 2016; Abichites ariaeii Korn & Ghaderi in Korn et al., 2016; Abichites paucinodus Korn & Ghaderi in Korn et al., 2016; Abichites shahriari Korn & Ghaderi in Korn et al., 2016; Kashmirites ? stoyanowi Kiparisova in Voinova et al., 1947; Abichites subtrapezoidalis Korn & Ghaderi in Korn et al., 2016; Abichites terminalis Korn & Ghaderi in Korn et al., 2016; Abichites ovalis Korn & Hairapetian sp. nov.; Abichites infirmus Korn & Hairapetian sp. nov. Diagnosis Genus of the family Dzhulfitidae with small to moderately large conch; maximum adult diameters are between 30 and 90 mm. Adult stage with quadrate, rectangular or oval whorl profile. Subadult stage with weak to moderately strong lateral ribs; adult stage with weak ornament. Suture line with deep external lobe; the depths of external lobe and adventive lobe are nearly identical. Remarks The morphological range of the genus Abichites was discussed by Korn & Ghaderi (in Korn et al. 2016)., Published as part of Korn, Dieter, Hairapetian, Vachik, Ghaderi, Abbas, Leda, Lucyna, Schobben, Martin & Akbari, Amir, 2021, The Changhsingian (Late Permian) ammonoids from Baghuk Mountain (Central Iran), pp. 1-106 in European Journal of Taxonomy 776 on page 87, DOI: 10.5852/ejt.2021.776.1559, http://zenodo.org/record/5604715, {"references":["Shevyrev A. A. 1965. Nadortyad Ammonoidea. In: Ruzhencev V. E. & Sarytcheva T. G. (eds) Rasvitie i smena morskikh organizmov na rubezhe Paleozoya i Mezozoya. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR 108: 166 - 182.","Voinova E. V., Kiparisova L. D. & Robinson V. H. 1947. Klass Cephalopoda. Golovonogie., Atlas rukovodyashtchikh form iskopaemykh faun SSSR. VIII. Triasovaya sistema: 124 - 176, Moskva-Leningrad","Korn D., Ghaderi A., Leda L., Schobben M. & Ashouri A. R. 2016. The ammonoids from the Late Permian Paratirolites Limestone of Julfa (East Azerbaijan, Iran). Journal of Systematic Palaeontology 14: 841 - 890. https: // doi. org / 10.1080 / 14772019.2015.1119211"]}