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Recent studies have documented benefits of small, prescribed fire and wildfire for grassland-dependent wildlife, such as lesser prairie-chickens (

Grazing management recommendations often sacrifice the intrinsic heterogeneity of grasslands by prescribing uniform grazing distributions through smaller pastures, increased stocking densities, and reduced grazing periods. The lack of patch-burn grazing in semi-arid landscapes of the western Great Plains in North America requires alternative grazing management strategies to create and maintain heterogeneity of habitat structure (e.g., animal unit distribution, pasture configuration), but knowledge of their effects on grassland fauna is limited. The lesser prairie-chicken (Tympanuchus pallidicinctus), an imperiled, grassland-obligate, native to the southern Great Plains, is an excellent candidate for investigating effects of heterogeneity-based grazing management strategies because it requires diverse microhabitats among life-history stages in a semi-arid landscape. We evaluated influences of heterogeneity-based grazing management strategies on vegetation structure, habitat selection, and nest and adult survival of lesser prairie-chickens in western Kansas, USA. We captured and monitored 116 female lesser prairie-chickens marked with very high frequency (VHF) or global positioning system (GPS) transmitters and collected landscape-scale vegetation and grazing data during 2013-2015. Vegetation structure heterogeneity increased at stocking densities = 40%. Probability of use was positively affected by increasing pasture area. A quadratic relationship existed between growing season deferment and probability of use. We found that 70% of nests were located in grazing units in which grazing pressure was

Projected twenty first century increases in temperature and precipitation intensity in the U.S. Great Plains may alter playa wetland hydroperiods. Our objective was to identify favorable germination conditions for a common moist-soil grass, Barnyardgrass (Echinochloa crusgalli L.), by evaluating emergence and growth response to various environmental conditions specific to the Northern (Nebraska) and Southern (Texas) range of playas. We used a temperature-controlled growth chamber experiment to evaluate emergence and growth response of Barnyardgrass to three main effects: (i) weekly temperatures representing historical and future conditions under a moderate emissions scenario, (ii) dry, moist, and saturated soil moisture conditions, and (iii) various seed bank densities. In Nebraska samples, projected future temperatures reduced emergence percentage by up to 20%, but increased emergence percentage by up to 15% for Texas samples. For Nebraska samples, plants were 9.6 cm taller under field capacity moisture compared to saturated moisture. Texas plant height was driven by temperature, where historical conditions produced plants that were 13 cm shorter than future warm conditions. These effects may be exacerbated in natural settings over time and when inter-specific competition exists; thus, temperature, soil moisture, and seed bank densities may be important considerations when planning for playa management in future climate conditions.

Temperate‐breeding, or resident, Canada geese were once extirpated in Kansas, USA, but currently provide abundant viewing and hunting opportunities. Kansas Department of Wildlife, Parks, and Tourism (KDWPT) began reintroducing geese in 1980 with a goal of re‐establishing a breeding population. Successful reintroductions led to translocating flocks to regions with no previous records of nesting geese; however, KDWPT continues to translocate individuals from nuisance flocks in urban areas to rural reservoirs to reduce human conflicts with urban geese. Our goal was to determine the effects of such translocations on survival and recovery of adult, sub‐adult, and juvenile temperate‐breeding Canada geese. We used Brownie dead‐recovery models in Program MARK to compare survival and recovery probabilities between translocated and nontranslocated (normal wild) Kansas‐banded Canada geese for 2012–2017. Model‐estimated annual survival differed between status (normal wild S^ = 0.761, 95% CI 0.734–0.785; translocated S^ = 0.598, 95% CI 0.528–0.665). Recovery probability differed between normal and translocated adults (normal wild f^ = 0.074, 95% CI = 0.069–0.078; translocated f^ = 0.138, 95% CI = 0.120–0.158) and juveniles (normal wild f^ = 0.067, 95% CI = 0.059–0.075; translocated f^ = 0.250, 95% CI = 0.199–0.310). Recovery probability did not differ between status in the sub‐adult age class (normal wild f^ = 0.126, 95% CI = 0.115–0.137; translocated f^ = 0.090, 95% CI = 0.055–0.144). Translocation is a viable management option to successfully reduce survival and increase recovery probability of urban nuisance geese in Kansas.

Large-scale declines of grassland ecosystems in the conterminous United States since European settlement have led to substantial loss and fragmentation of lesser prairie-chicken (Tympanuchus pallidicinctus) habitat and decreased their occupied range and population numbers by ∼85%. Breeding season space use is an important component of lesser prairie-chicken conservation, because it could affect both local carrying capacity and population dynamics. Previous estimates of breeding season space use are largely limited to one of the four currently occupied ecoregions, but potential extrinsic drivers of breeding space use, such as landscape fragmentation, vegetation structure and composition, and density of anthropogenic structures, can show large spatial variation. Moreover, habitat needs vary greatly among the lekking/prelaying, nesting, brood-rearing, and postbreeding stages of the breeding season, but space use by female lesser prairie-chickens during these stages remain relatively unclear. We tested whether home range area and daily displacement (the net distance between the first and last location of each day) of female lesser prairie-chickens varied among ecoregions and breeding stages at four study sites in Kansas and Colorado, U.S.A., representing three of the four currently occupied ecoregions. We equipped females with very-high-frequency (VHF) or Global Positioning System (GPS) transmitters, and estimated home range area with kernel density estimators or biased random bridge models, respectively. Across all ecoregions, breeding season home range area averaged 190.4 ha (±19.1 ha se) for birds with VHF and 283.6 ha (±23.1 ha) for birds with GPS transmitters, whereas daily displacement averaged 374.8 m (±14.3 m). Average home range area and daily displacement of bird with GPS transmitters were greater in the Short-Grass Prairie/ Conservation Reserve Program Mosaic and Sand Sagebrush Prairie Ecoregions compared to sites in the Mixed-Grass Prairie Ecoregion. Home range area and daily displacement were greatest during lekking/prelaying and smallest during the brood-rearing stage, when female movements were restricted by mobility of chicks. Ecoregion- and breeding stage-specific estimates of space use by lesser prairie-chickens will help managers determine the spatial configuration of breeding stage-specific habitat on the landscape. Furthermore, ecoregion- and breeding stage-specific estimates are crucial when estimating the amount of breeding habitat needed for lesser prairie-chicken populations to persist.

Spatially distributed populations often rely on large‐scale processes for long‐term population stability. These processes are driven by individuals moving across the landscape through long‐distance dispersal movements. However, as landscapes are continually altered by anthropogenic development, increased fragmentation and avoidance behavior can affect landscape permeability and limit dispersal. Lesser prairie‐chickens (Tympanuchus pallidicinctus) are a species of concern that have lost significant portions (>90%) of their historic distribution in the Southern Great Plains of the United States and are currently being impacted by continued anthropogenic development. Using GPS telemetry locations of 346 lesser prairie‐chickens across their entire geographic distribution, we identified 184 different long‐distance movements that drive population connectivity. We used empirical cumulative distribution functions to create a selection–avoidance–neutral curve and estimated the spatial scale of response to anthropogenic features (i.e., towers and windmills, large transmission and smaller distribution powerlines, oil wells, roads, and fences) during these movements. In addition, we tested for behavioral differences between movement types (e.g., exploratory loops vs. long‐distance movements between home ranges) and for regional differences in response among study areas. We found that during long‐distance movements, lesser prairie‐chickens generally avoided all anthropogenic feature types we tested despite some variation in the reported response distance among study areas. However, they avoided the tallest features (i.e., towers and windmills and transmission powerlines) at much greater distances in comparison with the shorter features in our analysis. Our results show that long‐distance movements are likely affected by responses to functional landscape fragmentation through increased development of anthropogenic features in important connectivity zones. As our estimated response distances during long‐distance movements varied in comparison with previously reported response distances during other behavioral states (e.g., breeding or nesting), using long‐distance or dispersal specific movement data may be more appropriate when asking questions related to connectivity across the landscape.

Loss and degradation of grasslands in the Great Plains region have resulted in major declines in abundance of grassland bird species. To ensure future viability of grassland bird populations, it is crucial to evaluate specific effects of environmental factors among species to determine drivers of population decline and develop effective conservation strategies. We used threshold models to quantify the effects of land cover and weather changes in "lesser prairie‐chicken" and "greater prairie‐chicken" (Tympanuchus pallidicinctus and T. cupido, respectively), northern bobwhites (Colinus virginianus), and ring‐necked pheasants (Phasianus colchicus). We demonstrated a novel approach for estimating landscape conditions needed to optimize abundance across multiple species at a variety of spatial scales. Abundance of all four species was highest following wet summers and dry winters. Prairie chicken and ring‐necked pheasant abundance was highest following cool winters, while northern bobwhite abundance was highest following warm winters. Greater prairie chicken and northern bobwhite abundance was also highest following cooler summers. Optimal abundance of each species occurred in landscapes that represented a grassland and cropland mosaic, though prairie chicken abundance was optimized in landscapes with more grassland and less edge habitat than northern bobwhites and ring‐necked pheasants. Because these effects differed among species, managing for an optimal landscape for multiple species may not be the optimal scenario for any one species., We quantified the effects of land cover and weather changes in Galliformes in Kansas. We found evidence of nonlinear, threshold effects of land cover change in abundance of all four focal species, suggesting that the presence of both grassland and cropland is important in the landscape to simultaneously maintain these populations.

The lesser prairie-chicken Tympanuchus pallidicinctus has experienced significant declines in distribution and abundance since the early 1900s. A severe and prolonged drought from 2009 to 2013 resulted in further declines in population numbers and despite improved environmental and habitat conditions since 2013, populations of lesser prairie chickens have shown little improvement. To investigate whether breeding season survival of lesser prairie-chickens in eastern New Mexico could be driving this response, we developed the following objectives: 1) estimate male and female breeding-season survival; 2) determine whether male and female survival varies temporally among lekking, nesting, and brood-rearing periods; and 3) determine cause-specific mortality during the breeding season. We captured and radiocollared 76 lesser prairie-chickens (50 male, 26 female) during spring of 2014 and 2015 and estimated their survival throughout the breeding season (15 March–31 August). Male survival was nearly double that of females in both years (0.79–0.81 and 0.38–0.45, respectively). Males had similar survival across all periods (lekking, postlekking, late summer: 0.89–0.95). Females had the greatest period-specific survival during lekking and brood rearing (0.87 ± 0.08 and 0.85 ± 0.10, respectively) relative to the nesting period (0.58 ± 0.11). Mammalian predation was the primary cause of mortality in both years. Our results indicate that in New Mexico 1) lesser prairie-chicken breeding season survival was consistent with geographically similar studies, 2) females have lower survival during the nesting period, and 3) female lesser prairie-chicken survival was lower than male survival regardless of time period. Management actions that provide and protect high-quality nesting habitat may help ensure that female survival is maximized during the nesting period.

Conservation of small populations is often based on limited data from spatially and temporally restricted studies, resulting in management actions based on an incomplete assessment of the population drivers. If fluctuations in abundance are related to changes in weather, proper management is especially important, because extreme weather events could disproportionately affect population abundance. Conservation assessments, especially for vulnerable populations, are aided by a knowledge of how extreme events influence population status and trends. Although important for conservation efforts, data may be limited for small or vulnerable populations. Integrated population models maximize information from various sources of data to yield population estimates that fully incorporate uncertainty from multiple data sources while allowing for the explicit incorporation of environmental covariates of interest. Our goal was to assess the relative influence of population drivers for the Lesser Prairie-Chicken ...

Within the U.S. portion of the Central Flyway, the U.S. Fish and Wildlife Service manages waterfowl on numerous individual units (i.e., Refuges) within the National Wildlife Refuge System. Presently, the extent of waterfowl use that Refuges receive and the contribution of Refuges to waterfowl populations (i.e., the proportion of the Central Flyway population registered at each Refuge) remain unassessed. Such an evaluation would help determine to what extent Refuges support waterfowl relative to stated targets, aid in identifying species requiring management attention, inform management targets, and improve fiscal efficiencies. Using historic monitoring data (1954–2008), we performed this assessment for 23 Refuges in Texas, New Mexico, Oklahoma, Kansas, and Nebraska during migration and wintering months (October–March). We examined six dabbling ducks and two diving ducks, plus all dabbling ducks and all diving ducks across two periods (long-term [all data] and short-term [last 10 October–March periods]). Individual Refuge use was represented by the sum of monthly duck count averages for October–March. We used two indices of Refuge contribution: peak contribution and January contribution. Peak contribution was the highest monthly count average for each October–March period divided by the indexed population total for the Central Flyway in the corresponding year; January contribution used the January count average divided by the corresponding population index. Generally, Refuges in Kansas, Nebraska, and New Mexico recorded most use and contribution for mallards Anas platyrhynchos. Refuges along the Texas Gulf Coast recorded most use and contribution for other dabbling ducks, with Laguna Atascosa and Aransas (including Matagorda Island) recording most use for diving ducks. The long-term total January contribution of the assessed Refuges to ducks wintering in the Central Flyway was greatest for green-winged teal Anas crecca with 35%; 12–15% for American wigeon Mareca americana, gadwall Mareca strepera, and northern pintail Anas acuta; and 7–8% for mallard and mottled duck Anas fulvigula. Results indicated that the reliance on the National Wildlife Refuge System decreased for these ducks, with evidence suggesting that, for several species, the assessed Refuges may be operating at carrying capacity. Future analyses could be more detailed and informative were Refuges to implement a single consistent survey methodology that incorporated estimations of detection bias in the survey process, while concomitantly recording habitat metrics on and neighboring each Refuge.

Coastal wetlands along the Gulf of Mexico support a wide diversity of wildlife, are important nurseries for sport and commercial fisheries, provide erosion and flood control, and serve many other ecological functions and services. These marshes have been declining in area and degrading at alarming rates since the 1930s. Effective conservation planning is vital to protect these ecosystems, but decision makers often lack knowledge of expected future conditions to strategically target conservation actions. To address this issue, we focus on a species of conservation concern, the mottled duck (Anas fulvigula), that resides year-round in the coastal marshes of the Gulf of Mexico. We used location data collected from radiomarked hen mottled ducks from 2006 to 2011 to create an ensemble model of habitat selection for 2010. We then projected future habitat states using models of sea-level rise and human development. By combining future predictions with our ensemble model, we predict future habitat for mottled ducks through 2100, in 20-year time steps beginning with 2020. Sea-level rise models predicted reductions in coastal marsh habitats and our ensemble model predict corresponding declines in overall habitat quantity and quality for mottled ducks, with the largest rate of habitat loss predicted within the Chenier Plain of Louisiana, USA at 71%. In some areas, particularly the Texas Mid-Coast, USA, future urbanization and human development is expected to reduce the ability of wetland habitat to migrate inland with rising sea-levels. Our results also highlight areas of coastal marsh particularly vulnerable to sea-level rise; and conversely, identify areas most likely to persist into the future that could be targeted for habitat conservation to help mottled ducks persist on the landscape.

The lesser prairie-chicken (Tympanuchus pallidicinctus), a species of conservation concern with uncertain regulatory status, has experienced population declines over the past century. Most research on lesser prairie-chickens has focused on the breeding season, with little research conducted during the nonbreeding season, a period that exerts a strong influence on demography in other upland game birds. We trapped lesser prairie-chickens on leks and marked them with either global positioning system (GPS) satellite or very high frequency (VHF) transmitters to estimate survival and home-range size during the nonbreeding season. We monitored 119 marked lesser prairie-chickens in 3 study areas in Kansas, USA, from 16 September to 14 March in 2013, 2014, and 2015. We estimated home-range size using Brownian Bridge movement models (GPS transmitters) and fixed kernel density estimators (VHF transmitters), and female survival using Kaplan–Meier known-fate models. Average home-range size did not differ between sexes. Estimated home-range size was 3 times greater for individuals fitted with GPS satellite transmitters (x¯ = 997 ha) than those with VHF transmitters (x¯ = 286 ha), likely a result of the temporal resolution of the different transmitters. Home-range size of GPS-marked birds increased 2.8 times relative to the breeding season and varied by study area and year. Home-range size was smaller in the 2013–2014 nonbreeding season (x¯ = 495 ha) than the following 2 nonbreeding seasons (x¯ = 1,290 ha and x¯ = 1,158 ha), corresponding with drought conditions of 2013, which were alleviated in following years. Female survival (Sˆ) was high relative to breeding season estimates, and did not differ by study area or year (Sˆ = 0.73 ± 0.04 [SE]). Future management could remain focused on the breeding season because nonbreeding survival was 39–44% greater than the previous breeding season; however, considerations of total space needs would benefit lesser prairie-chickens by accounting for the greater spatial requirements during the nonbreeding season. © 2017 The Wildlife Society.

Understanding the relative influence of environmental and intrinsic effects on populations is important for managing and conserving harvested species, especially those species inhabiting changing environments. Additionally, climate change can increase the uncertainty associated with management of species in these changing environments, making understanding factors affecting their populations even more important. Coastal ecosystems are particularly threatened by climate change; the combined effects of increasing severe weather events, sea level rise, and drought will likely have non-linear effects on coastal marsh wildlife species and their associated habitats. A species of conservation concern that persists in these coastal areas is the mottled duck (Anas fulvigula). Mottled ducks in the western Gulf Coast are approximately 50% below target abundance numbers established by the Gulf Coast Joint Venture for Texas and Louisiana, USA. Although evidence for declines in mottled duck abundance is apparent, specific causes of the decrease remain unknown. Our goals were to determine where the largest declines in mottled duck population were occurring along the system of Texas Gulf Coast National Wildlife Refuges and quantify the relative contribution of environmental and intrinsic effects on changes to relative population density. We modeled aerial survey data of mottled duck density along the Texas Gulf Coast from 1986–2015 to quantify effects of extreme weather events on an index to mottled duck density using the United States Climate Extremes Index and Palmer Drought Severity Index. Our results indicate that decreases in abundance are best described by an increase in days with extreme 1-day precipitation from June to November (hurricane season) and an increase in drought severity. Better understanding those portions of the life cycle affected by environmental conditions, and how to manage mottled duck habitat in conjunction with these events will likely be key to persistence of the species under future environmental conditions. © 2017 The Wildlife Society.

The mottled duck (Anas fulgivula) is a non-migratory duck dependent on coastal habitats to meet all of its life cycle requirements in the Western Gulf Coast (WGC) of Texas and Louisiana, USA. This population of mottled ducks has experienced a moderate decline during the past 2 decades. Adult survival has been identified as an important factor influencing population demography. Previous work based on band-recovery data has provided only annual estimates of survival. We assessed seasonal patterns of female mottled duck survival from 2009 to 2012 using individuals marked with satellite platform transmitter terminals (PTTs). We used temperature and movement sensors within each PTT to indicate potential mortality events. We estimated cumulative weekly survival and ranked factors influential in patterns of mortality using known-fate modeling in Program MARK. Models included 4 predictors: week; hunting and non-hunting periods; biological periods defined as breeding, brooding, molt, and pairing; and mass at time of capture. Models containing hunt periods, during and outside the mottled duck season, comprised essentially 100% of model weights where both legal and illegal harvest had a negative influence on mottled duck survival. Survival rates were low during 2009–2011 (12–38% annual rate of survival), when compared with the long-term banding average of 53% annual survival. During 2011, survival of female mottled ducks was the lowest annual rate (12%) ever documented and coincided with extreme drought. Management actions maximizing the availability of wetlands and associated upland habitats during hunting seasons and drought conditions may increase adult female mottled duck survival. © 2017 The Authors. Journal of Wildlife Management Published by Wiley Periodicals, Inc. on behalf of The Wildlife Society.

Since the beginning of the 20th century, the overall occupied range of the lesser prairie-chicken (Tympanuchus pallidicinctus) has declined by 84% commensurate with population trends. Much of this decline has been attributed to the loss and fragmentation of native grasslands throughout the lesser prairie-chicken range. However, quantification of changes in land cover in the distribution of the lesser prairie-chicken is lacking. Our objectives were to (1) document changes in the areal extent and connectivity of grasslands in the identified lesser prairie-chicken range in Kansas, USA, (>60% of extant lesser prairie-chicken population) from the 1950s to 2013 using remotely sensed data and (2) assess the potential of the Conservation Reserve Program (U.S. Department of Agriculture Program converting cropland to permanent cover; CRP) to mitigate grassland loss. Digital land cover maps were generated on a decadal time step through spectral classification of LANDSAT images and visual analysis of aerial photographs (1950s and 1960s). Landscape composition and configuration were assessed using FRAGSTATS to compute a variety of landscape metrics measuring changes in the amount of grassland present as well as changes in the size and configuration of grassland patches. With the exception of a single regional portion of the range, nearly all of the grassland converted to cropland in the lesser prairie-chicken range of Kansas occurred prior to the 1950s. Prior to the implementation of CRP, the amount of grassland decreased 3.6% between the 1950s and 1985 from 18,455 km2 to 17,788 km2. Since 1985, the overall amount of grassland in the lesser prairie-chicken range has increased 11.9% to 19,898 km2 due to implementation of CRP, although the area of grassland decreased between 1994 and 2013 as CRP contracts were not renewed by landowners. Since 1986 grassland in Kansas became more connected and less fragmented in response to the CRP. While the CRP has been successful in increasing grassland quantity and connectivity throughout the lesser prairie-chicken range in Kansas, offsetting loss of grassland since the 1950s, abundance and occupied range of lesser prairie-chickens has declined since the 1980s, suggesting that habitat quality is the principal factor influencing population demography of the species. Although the CRP is contributing to conservation actions for lesser prairie-chickens, efforts to improve habitat quality throughout the range of the lesser prairie-chicken are likely necessary to meet management goals. Continuation of the CRP faces an uncertain future in the face of rising commodity prices, energy development, and reduction in program scope, leaving open the possibility that these areas that have created habitat for lesser prairie-chickens could be lost.

The >2,570,000‐ha Flint Hills ecoregion of Kansas, USA, harbors the largest remaining contiguous tract of tallgrass prairie in North America, a unique system, as the remainder of North America's tallgrass prairie has succumbed to development and conversion. Consequently, the loss and degradation of tallgrass prairie has reduced populations of many North American prairie‐obligate species including the regal fritillary (Speyeria idalia) butterfly. Population abundance and occupied range of regal fritillary have declined >99%, restricting many populations to isolated, remnant patches of tallgrass prairie. Such extensive decline has resulted in consideration of the regal fritillary for protection under the Endangered Species Act. Although it is widely accepted that management practices such as fire, grazing, and haying are necessary to maintain prairie ecosystems, reported responses by regal fritillary to these management regimes have been ambiguous. We tested effects of prescribed fire across short, moderate, and long fire‐return intervals as well as grazing and haying management treatments on regal fritillary density. We also tested the relative influence of habitat characteristics created by these management regimes by measuring density of an obligate host plant (Viola spp.) and canopy cover of woody vegetation, grasses, forbs/ferns, bare ground, and litter. We found density was at least 1.6 times greater in sites burned with a moderate fire‐return interval vs. sites burned with short and long fire‐return intervals. Overall management regardless of fire‐return interval did not have an effect on density. Percent cover of grass had the strongest positive association, while percent cover of woody vegetation had the greatest negative effect on density. Our results indicate that patch‐burning is a viable and perhaps even ideal management strategy for regal fritillary in tallgrass prairie landscapes. Additionally, these results elucidate the importance of fire, particularly when applied at moderate‐return intervals to regal fritillary, and corroborate a growing suite of studies that suggest fire is perhaps not as detrimental to populations of regal fritillary as previously believed.

Lesser prairie-chicken (Tympanuchus pallidicinctus) populations have declined since the 1980s. Understanding factors influencing nest-site selection and nest survival are important for conservation and management of lesser prairie-chicken populations. However, >75% of the extant population is in the northern extent of the range where data on breeding season ecology are lacking. We tested factors influencing fine-scale and regional nest-site selection and nest survival across the northern portion of the lesser prairie-chicken range. We trapped and affixed satellite global positioning system and very high frequency transmitters to female lesser prairie-chickens (n = 307) in south-central and western Kansas and eastern Colorado, USA. We located and monitored 257 lesser prairie-chicken nests from 2013 to 2016. We evaluated nest-site selection and nest survival in comparison to vegetation composition and structure. Overall, nest-site selection in relation to vegetation characteristics was similar across our study area. Lesser prairie-chickens selected nest microsites with 75% visual obstruction 2.0-3.5 dm tall and 95.7% of all nests were in habitat with >= 1 dm and

Sand shinnery oak (Quercus havardii) prairies are increasingly disappearing and increasingly degraded in the Southern High Plains of Texas and New Mexico. Restoring and managing sand shinnery oak prairie can support biodiversity, specific species of conservation concern, and livestock production. We measured vegetation response to four treatment combinations of herbicide (tebuthiuron applied at 0.60 kg/ha) and moderate-intensity grazing (50% removal of annual herbaceous production) over a 10-year period in a sand shinnery oak prairie of eastern New Mexico. We compared the annual vegetation response to the historical climax plant community (HCPC) as outlined by the U.S. Department of Agriculture Ecological Site Description. From 2 to 10 years postapplication, tebuthiuron-treated plots had reduced shrub cover with twice as much forb and grass cover as untreated plots. Tebuthiuron-treated plots, regardless of the presence of grazing, most frequently met HCPC. Tebuthiuron and moderate-intensity grazi...

Long-term population and range declines from habitat loss and fragmentation caused the lesser prairie-chicken (Tympanuchus pallidicinctus) to be a species of concern throughout its range. Current lesser prairie-chicken range in New Mexico and Texas is partially restricted to sand shinnery oak (Quercus havardii; hereafter shinnery oak) prairies, on which cattle grazing is the main socioeconomic driver for private landowners. Cattle producers within shinnery oak prairies often focus land management on shrub eradication using the herbicide tebuthiuron to promote grass production for forage; however, herbicide application alone, and in combination with grazing, may affect nest site selection and nest survival of lesser prairie-chickens through the reduction of shinnery oak and native grasses. We used a controlled, paired, completely randomized design study to assess the influence of grazing and tebuthiuron application and their combined use on nest site selection and nest survival from 2001 to 2010 in Roosevelt County, New Mexico, USA at 2 spatial scales (i.e., treatment and microhabitat) in 4 treatments: tebuthiuron with grazing, tebuthiuron without grazing, no tebuthiuron with grazing, and a control of no tebuthiuron and no grazing. Grazing treatment was a short-duration system in which plots were grazed once during the dormant season and once during the growing season. Stocking rate was calculated each season based on measured forage production and applied to remove ≤25% of available herbaceous material per season. At the treatment scale, we compared nest site selection among treatments using 1-way χ2 tests and nest survival among treatments using a priori candidate nest survival models in Program MARK. At the microhabitat scale, we identified important habitat predictors of nest site selection and nest survival using logistic regression and a priori candidate nest survival models in Program MARK, respectively. Females typically used treatments as expected and we did not detect trends in selection. Nest survival did not differ among treatments. At the microhabitat scale, nest sites had less bare ground (P = 0.001) and greater angles of obstruction (P ≤ 0.001) compared to random sites. There was a high degree of model selection uncertainty among our candidate models at the microhabitat scale and survival estimates were similar among habitat covariates. Results suggest a tebuthiuron application rate of 0.60 kg/ha, short-duration grazing, and a combination of these management techniques were not detrimental to lesser prairie-chicken nest site selection or nest survival. However, intensified management that increases bare ground or reduces overhead cover may negatively affect lesser prairie-chicken nesting habitat and nest survival. © 2016 The Wildlife Society.

To identify factors contributing to the long-term decline of American woodcock, a holistic understanding of range-wide population connectivity throughout the annual cycle is needed. We used band recovery data and isotopic composition of primary (P1) and secondary (S13) feathers to estimate population sources and connectivity among natal, early fall, and winter ranges of hunter-harvested juvenile American woodcock. We used P1 feathers from known-origin pre-fledged woodcock (n = 43) to create a hydrogen δ2Hf isoscape by regressing δ2Hf against expected growing-season precipitation (δ2Hp). Modeled δ2Hp values explained 79% of the variance in P1 δ2Hf values, indicating good model fit for estimating woodcock natal origins. However, a poor relationship (r2 = 0.23) between known-origin, S13 δ2Hf values, and expected δ2Hp values precluded assignment of early fall origins. We applied the δ2Hf isoscape to assign natal origins using P1 feathers from 494 hunter-harvested juvenile woodcock in the United States and Canada during 2010–2011 and 2011–2012 hunting seasons. Overall, 64% of all woodcock origins were assigned to the northernmost (>44°N) portion of both the Central and Eastern Management Regions. In the Eastern Region, assignments were more uniformly distributed along the Atlantic coast, whereas in the Central Region, most woodcock were assigned to origins within and north of the Great Lakes region. We compared our origin assignments to spatial coverage of the annual American woodcock Singing Ground Survey (SGS) and evaluated whether the survey effectively encompasses the entire breeding range. When we removed the inadequately surveyed Softwood shield Bird Conservation Region (BCR) from the northern portion of the SGS area, only 48% of juvenile woodcock originated in areas currently surveyed by the SGS. Of the individuals assigned to the northernmost portions of the breeding range, several were harvested in the southern extent of the wintering range. Based upon this latitudinal winter stratification, we examined whether woodcock employed a leapfrog migration strategy. Using δ2Hf values and band-recovery data, we found some support for this migration strategy hypothesis but not as a singular explanation. The large harvest derivation of individuals from the northernmost portions of the breeding range, and the difference in breeding distributions within each Management Region should be considered in future range-wide conservation and harvest management planning for American woodcock. © 2016 The Wildlife Society.

Playa wetlands in the Great Plains, USA support a wide variety of plant species not found elsewhere in this agriculturally-dominated region due to the ephemeral presence of standing water and hydric soils within playas. If longer dry periods occur due to climate change or if changes in surrounding land use alter sediment accumulation rates and water storage capacity in playas, plant communities could experience decreased diversity, with lasting effects on ecosystem services provided by playas in the Great Plains and at a continental-level in North America. We quantified potential changes in playa wetland plant community composition associated with predicted changes in precipitation and land use in the Great Plains through the end of the 21st century. We conducted two six-month greenhouse experiments mimicking field conditions using intact mesocosms collected from playas in Nebraska and Texas. In the precipitation experiment, treatments derived from historical precipitation observations and three future moderate emissions (CMIP5 RCP4.5) downscaled climate projections were applied to mesocosms. For the land use experiment, treatments were simulated by nitrogen (N) applications to soil ranging from 0 to 100 mg-N L-1 with each precipitation event under historical rainfall patterns, representing increasing and decreasing area in agricultural use in playa watersheds. Plant communities tended to shift toward more native species under projected future climate conditions, but as N runoff increased, native species richness decreased. Agricultural land-use surrounding playas may have a greater effect on wetland plant communities than future alterations to hydrology based on climate change in the Great Plains; thus, efforts to reduce nutrient runoff into playas would likely mitigate loss in ecosystem function in the coming decades. National Science FoundationNational Science Foundation (NSF) [IIA-1355406, IIA-1430427]; USDA-NIFAUnited States Department of Agriculture (USDA) [MO-HANR0007]; Missouri Department of Conservation; University of Missouri; U.S. Fish and Wildlife ServiceUS Fish & Wildlife Service; U.S. Geological SurveyUnited States Geological Survey; Wildlife Management Institute This research was part of the Missouri EPSCoR project, funded by the National Science Foundation under Award #IIA-1355406 and #IIA-1430427. Partial financial support was also provided by USDA-NIFA through Hatch funding (MO-HANR0007). Any opinions, findings, and conclusions or recommendations expressed in this manuscript are those of the authors and do not necessarily reflect the views of the National Science Foundation or USDA-NIFA. The Missouri Cooperative Fish and Wildlife Research Unit is jointly sponsored by the Missouri Department of Conservation, the University of Missouri, the U.S. Fish and Wildlife Service, the U.S. Geological Survey and the Wildlife Management Institute. Any use of trade, firm or product names is for descriptive purposes only and does not imply endorsement by the U.S. Government. This research would not have been possible without the assistance of Eric Simpson, Janith Chandrasoma, and Amanda Burnett. Finally, we would like to thank the School of Natural Resources Writing Workshop group for conceptual and technical edits which greatly improved the quality of this manuscript. Public domain – authored by a U.S. government employee

Knowledge of landscape and regional circumstances where conservation programs are successful on working lands inagricultural production are needed. Convertingmarginal croplands to grasslands using conservation programs such as the United States Department of Agriculture Conservation Reserve Program (CRP) should be beneficial for many grassland-obligate wildlife species; however, addition of CRP grasslands may result indifferent population effects based on regional climate, characteristics of the surrounding landscape, or species planted or established. Within landscapes occupied by lesser prairie-chickens (Tympanuchus pallidicinctus), CRP may provide habitat only for specific life stages and habitat selection for CRP may vary between wet and dry years. Among all study sites, we captured and fitted 280 female lesser prairie-chickens with very high frequency (VHF)- and global positioning system (GPS) transmitters during the spring lekking seasons of 2013-2015 to monitor habitat selection for CRP in regions of varying climate. We also estimated vital rates and habitat selection for 148 individuals, using sites in northwest Kansas, USA. The greatest ecological services of CRP became apparent when examining habitat selection and densities. Nest densities were approximately 3 times greater in CRP grasslands than native working grasslands (i.e., grazed), demonstrating a population-level benefit (CRP = 6.0 nests/10 km(2) +/- 1.29 [ SE], native working grassland = 1.7 nests/10 km(2) +/- 0.62). However, CRP supporting high nest density did not provide brood habitat; 85% of females with broods surviving to 7 days moved their young to other cover types. Regression analyses indicated lesser prairie-chickens were approximately 8 times more likely to use CRP when 5,000-ha landscapes were 70% rather than 20% grassland, indicating variation in the level of ecological services provided by CRP was dependent upon composition of the larger landscape. Further, CRP grasslands were 1.7 times more likely to be used by lesser prairie-chickens in regions receiving 40 cm compared to 70 cm of average annual precipitation and during years of greater drought intensity. Demographic and resource selection analyses revealed that establishing CRP grasslands in northwest Kansas can increase the amount nesting habitat in a region where it may have previously been limited, thereby providing refugia to sustain populations through periods of extreme drought. Nest survival, adult survival during breeding, and nonbreeding season survival did not vary between lesser prairie-chickens that used and did not use CRP grasslands. The finite rate of population growth was also similar for birds using CRP and using only native working grasslands, suggesting that CRP provides habitat similar to that of native working grassland in this region. Overall, lesser prairie-chickens may thrive in landscapes that are a mosaic of native working grassland, CRP grassland, with a minimal amount of cropland, particularly when nesting and brood habitat are in close proximity. (C) 2018 The Wildlife Society Kansas Wildlife, Parks, and Tourism (Federal Assistance Grant) [KS W-73-R-3]; United States Department of Agriculture (USDA) Farm Services CRP Monitoring, Assessment, and Evaluation [7, KSCFWRU RWO 62]; USDA Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative Any use of trade, firm, or product names is for descriptive purposes only and does not imply endorsement by the United States Government. We thank R. D. Rodgers for providing comments and edits on a previous draft of the manuscript. B. Anderson, S. Baker, S. Bard, G. Brinkman, K. Broadfoot, R. Cooper, J. Danner, J. Decker, E. D. Entsminger, R. M. Galvin, N. Gilbert, A. Godar, G. Gould, B. Hardy, S.P. Hoffman, D. Holt, B. M. Irle, T. Karish, A. Klais, H. Kruckman, K. Kuechle, S. J. Lane, E. A. Leipold, J. Letlebo, E. Mangelinckx, L. McCall, A. Nichter, K. Phillips, J. K. Proescholdt, J. Rabon, T. Reed, A. Rhodes, B. E. Ross, D. Spencer, A. M. Steed, A. E. Swicegood, P. Waldron, B. A. Walter, I. Waters, W. J. White, E. Wiens, J. B. Yantachka, and A. Zarazua, provided much needed assistance with data collection. We greatly appreciate the logistic and technical support provided by J. C. Pitman, J. Kramer, M. Mitchener, D. K. Dahlgren, J. A. Prendergast, C. Berens, G. Kramos, A. A. Flanders, and S. Hyberg. Funding for the project was provided by Kansas Wildlife, Parks, and Tourism (Federal Assistance Grant KS W-73-R-3); United States Department of Agriculture (USDA) Farm Services CRP Monitoring, Assessment, and Evaluation (12-IA-MRE CRP TA#7, KSCFWRU RWO 62); and USDA Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative. Public domain – authored by a U.S. government employee

Diets during critical brooding and winter periods likely influence the growth of Lesser Prairie-Chicken (Tympanuchus pallidicinctus) populations. During the brooding period, rapidly growing Lesser Prairie-Chicken chicks have high calorie demands and are restricted to foods within immediate surroundings. For adults and juveniles during cold winters, meeting thermoregulatory demands with available food items of limited nutrient content may be challenging. Our objective was to determine the primary animal and plant components of Lesser Prairie-Chicken diets among native prairie, cropland, and Conservation Reserve Program (CRP) fields in Kansas and Colorado, USA, during brooding and winter using a DNA metabarcoding approach. Lesser Prairie-Chicken fecal samples (n = 314) were collected during summer 2014 and winter 2014-2015, DNA was extracted, amplified, and sequenced. A region of the cytochrome oxidase I (COI) gene was sequenced to determine the arthropod component of the diet, and a portion of the trnL intron region was used to determine the plant component. Relying on fecal DNA to quantify dietary composition, as opposed to traditional visual identification of gut contents, revealed a greater proportion of soft-bodied arthropods than previously recorded. Among 80 fecal samples for which threshold arthropod DNA reads were obtained, 35% of the sequences were most likely from Lepidoptera, 26% from Orthoptera, 14% from Araneae, 13% from Hemiptera, and 12% from other orders. Plant sequences from 137 fecal samples were composed of species similar to Ambrosia (27%), followed by species similar to Lactuca or Taraxacum (10%), Medicago (6%), and Triticum (5%). Forbs were the predominant (>50% of reads) plant food consumed during both brood rearing and winter. The importance both of native forbs and of a broad array of arthropods that rely on forbs suggests that disturbance regimes that promote forbs may be crucial in providing food for Lesser Prairie-Chickens in the northern portion of their distribution. Kansas Wildlife, Parks, and Tourism [KS W-73-R-3]; USDA Farm Services CRP Monitoring, Assessment, and Evaluation [12-IA-MRE CRP TA#7, KSCFWRU RWO 62]; USDA Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative Funding for the project was provided by Kansas Wildlife, Parks, and Tourism (Federal Assistance Grant KS W-73-R-3); USDA Farm Services CRP Monitoring, Assessment, and Evaluation (12-IA-MRE CRP TA#7, KSCFWRU RWO 62); and USDA Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative. Public domain – authored by a U.S. government employee

Agriculture and development have caused landscape change throughout the southwestern Great Plains in the range of the lesser prairie-chicken (Tympanuchus pallidicinctus). Landscape alteration within the lesser prairie-chicken range may contribute to range contraction and population losses through decreases in survival rates. Our objectives were to determine if: (1) landscape configuration (i.e., the spatial arrangement of habitat) or composition (i.e., the amount of habitat), at the study.site scale, affected annual survival of females, (2) relationships exist between landscape context (i.e., landscape configuration and composition) and weekly survival to assess effects of landscape composition and configuration on lesser prairie-chicken populations, and (3) anthropogenic features influenced daily mortality risk. We captured 170 female lesser prairie-chickens and used very-high-frequency and GPS (Global Positioning System) transmitters to track their movement and survival for 2 y. We used known-fate survival models to test if landscape configuration or composition within three sites in Kansas were related to differences in female survival among sites. In addition we tested for relationships between weekly survival and landscape configuration or composition within home ranges. Finally, we used Andersen-Gill models to test the influence of distance to anthropogenic features on daily mortality risk. Differences in survival were evident between sites with differing landscape compositions as annual survival in Northwestern Kansas (S=0.27) was half that of Clark County, Kansas (S=0.56), which corresponded with 41.9% more grassland on the landscape in Clark County; landscape configuration did not measurably differ among sites. Survival was greater for prairie-chickens with home-ranges that had greater patch richness and in areas with 30% crop and 57% grassland. Female lesser prairie-chickens also experienced greater mortality risk closer to fences at patch edges. Further conversion of grassland landscapes occupied by lesser prairie-chickens should be avoided to reduce habitat loss and fragmentation thresholds that could affect survival. We suggest continued encouragement of Conservation Reserve Program enrollment in western areas of the lesser prairie-chicken range to maintain or increase the amount of grassland to increase annual survival. Federal Aid in Wildlife Restoration Grant [W-73-R]; US Geological SurveyUnited States Geological Survey; US Dept ofAgriculture (USDA), Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative; Kansas Wildlife, Parks, and Tourism [KS W-73-R-3]; USDA Farm Services CRP Monitoring, Assessment, and Evaluation [7, KSCFWRU RWO 62] Wethank J. Pitman, K. Sexton, J. Kramer, M. Mitchener, D. Dahlgren, J. Prendergast, C. Berens, G. Kramos, A. Flanders, and S. Hyberg for their assistance with the project. Research was funded by the Federal Aid in Wildlife Restoration Grant W-73-R; US Geological Survey; US Dept ofAgriculture (USDA), Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative; Kansas Wildlife, Parks, and Tourism (Federal Assistance Grant KS W-73-R-3); USDA Farm Services CRP Monitoring, Assessment, and Evaluation (12-IA-MRE CRP TA#7, KSCFWRU RWO 62). We also thank two anonymous reviewers for reviewing earlier versions of this manuscript. Any use of trade, firm, or product names is for descriptive purposes only and does not imply endorsement by the U.S. Government. Public domain – authored by a U.S. government employee

Ethical treatment of research animals is required under the Animal Welfare Act. This includes trapping methodologies that reduce unnecessary pain and duress. Traps used in research should optimize animal welfare conditions within the context of the proposed research study. Several trapping techniques are used in the study of lesser prairie-chickens, despite lack of knowledge of trap injury caused by the various methods. From 2006 to 2012, we captured 217, 40, and 144 lesser prairie-chickens Tympanuchus pallidicinctus using walk-in funnel traps, rocket nets, and drop nets, respectively, in New Mexico and Texas, to assess the effects of capture technique on injury and survival of the species. We monitored radiotagged, injured lesser prairie-chickens 7–65 d postcapture to assess survival rates of injured individuals. Injuries occurred disproportionately among trap type, injury type, and sex. The predominant injuries were superficial cuts to the extremities of males captured in walk-in funnel traps. However, we observed no mortalities due to trapping, postcapture survival rates of injured birds did not vary across trap types, and the daily survival probability of an injured and uninjured bird was ≥99%. Frequency and intensity of injuries in walk-in funnel traps are due to the passive nature of these traps (researcher cannot select specific individuals for capture) and incidental capture of individuals not needed for research. Comparatively, rocket nets and drop nets allow observers to target birds for capture and require immediate removal of captured individuals from the trap. Based on our results, trap injuries would be reduced if researchers monitor and immediately remove birds from walk-in funnels before they injure themselves; move traps to target specific birds and reduce recaptures; limit the number of consecutive trapping days on a lek; and use proper netting techniques that incorporate quick, efficient, trained handling procedures.

We used the lesser prairie‐chicken (Tympanuchus pallidicinctus), an iconic grouse species that exhibits a boom–bust life history strategy, on the Southern High Plains, USA, as a bioindicator of main and interactive effects of severe drought and grazing. This region experienced the worst drought on record in 2011. We surveyed lesser prairie‐chicken leks (i.e., communal breeding grounds) across 12 years that represented 7 years before the 2011 drought (predrought) and 4 years during and following the 2011 drought (postdrought). Grazing was annually managed with the objective of achieving ≤50% utilization of aboveground vegetation biomass. We used lek (n = 49) count data and covariates of weather and managed grazing to: (a) estimate long‐term lesser prairie‐chicken abundance and compare abundance predrought and postdrought; (b) examine the influence of annual and seasonal drought (modified Palmer drought index), temperature, and precipitation on long‐term lesser prairie‐chicken survival and recruitment; and (c) assess and compare the influence of grazing on lesser prairie‐chicken population predrought and postdrought. Lesser prairie‐chicken abundance was nearly seven times greater predrought than postdrought, and population declines were attributed to decreased survival and recruitment. The number of days with temperature >90th percentile had the greatest effect, particularly on recruitment. The population exhibited a substantial bust during 2011 and 2012 without a boom to recover in four postdrought years. Adaptive grazing positively influenced the population predrought, but had no effects postdrought. Results suggest that the severe drought in 2011 may have been beyond the range of environmental conditions to which lesser prairie‐chickens, and likely other species, have adapted. Land management practices, such as grazing, should remain adaptive to ensure potential negative influences to all species are avoided. Increasing habitat quantity and quality by reducing habitat loss and fragmentation likely will increase resiliency of the ecosystem and individual species.

For millennia grasslands have provided a myriad of ecosystem services and have been coupled with human resource use. The loss of 46% of grasslands worldwide necessitates the need for conservation that is spatially, temporally, and socioeconomically strategic. In the Southern Great Plains of the United States, conversion of native grasslands to cropland, woody encroachment, and establishment of vertical anthropogenic features have made large intact grasslands rare for lesser prairie-chickens (Tympanuchus pallidicinctus). However, it remains unclear how the spatial distribution of grasslands and anthropogenic features constrain populations and influence conservation. We estimated the distribution of lesser prairie-chickens using data from individuals marked with GPS transmitters in Kansas and Colorado, USA, and empirically derived relationships with anthropogenic structure densities and grassland composition. Our model suggested decreased probability of use in 2-km radius (12.6 km(2)) landscapes that had greater than two vertical features, two oil wells, 8 km of county roads, and 0.15 km of major roads or transmission lines. Predicted probability of use was greatest in 5-km radius landscapes that were 77% grassland. Based on our model predictions, similar to 10% of the current expected lesser prairie-chicken distribution was available as habitat. We used our estimated species distribution to provide spatially explicit prescriptions for CRP enrollment and tree removal in locations most likely to benefit lesser prairie-chickens. Spatially incentivized CRP sign up has the potential to provide 4189 km2 of additional habitat and strategic application of tree removal has the potential to restore 1154 km(2). Tree removal and CRP enrollment are conservation tools that can align with landowner goals and are much more likely to be effective on privately owned working lands. Kansas Wildlife, Parks, and Tourism (Federal Assistance Grant) [KS W-73-R-3]; United States Department of Agriculture (USDA) Farm Services CRP Monitoring, Assessment, and Evaluation [12-IA-MRE CRP TA, KSCFWRU RWO 62, 7]; USDA Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative We thank Kent Fricke, three anonymous reviewers, and the associate editor for providing reviews that improved the quality of the manuscript. We thank K. Schultz and A. Chappell for capturing and providing GPS data from lesser prairie-chickens captured on the Cimarron National Grasslands. B. Anderson, S. Baker, S. Bard, G. Brinkman, K. Broadfoot, R. Cooper, J. Danner, J. Decker, E. D. Entsminger, R. M. Galvin, N. Gilbert, A. Godar, G. Gould, B. Hardy, S.P. Hoffman, D. Holt, B. M. Irle, T. Karish, A. Klais, H. Kruckman, K. Kuechle, S. J. Lane, E. A. Leipold, J. Letlebo, E. Mangelinckx, L. McCall, A. Nichter, K. Phillips, J. K. Proescholdt, J. Rabon, T. Reed, A. Rhodes, B. E. Ross, D. Spencer, A. M. Steed, A. E. Swicegood, P. Waldron, B. A. Walter, I. Waters, W. J. White, E. Wiens, J. B. Yantachka, and A. Zarazua, provided much needed assistance with data collection. We greatly appreciate the logistic and technical support provided by J. C. Pitman, J. Kramer, M. Mitchener, D. K. Dahlgren, J. A. Prendergast, C. Berens, G. Kramos, and A. A. Flanders. Funding for the project was provided by Kansas Wildlife, Parks, and Tourism (Federal Assistance Grant KS W-73-R-3); United States Department of Agriculture (USDA) Farm Services CRP Monitoring, Assessment, and Evaluation (12-IA-MRE CRP TA#7, KSCFWRU RWO 62); and USDA Natural Resources Conservation Service, Lesser Prairie-Chicken Initiative. Any use of trade, firm, or product names is for descriptive purposes only and does not imply endorsement by the U.S. Government. Public domain – authored by a U.S. government employee

Sericea lespedeza (Lespedeza cuneata) is a high-tannin, late-season invasive forb species that reduces biodiversity in tallgrass prairie ecosystems. The largest tallgrass prairie remnant exists in the Flint Hills of Kansas and Oklahoma, where the most common grazing management practice involves prescribed fire in early spring followed by intensive stocking with yearling beef cattle from April to July. Sericea has continued to spread under this management regime. From 2013 to 2016, in Kansas Flint Hills tallgrass prairie, we tested the effects of using spring burning with early-season steer grazing, followed by late-season sheep grazing (Steer+Sheep) compared to spring burning followed by steer grazing only (Steer) on sericea vigor, grassland birds, and pollinators. Density and nest success of Grasshopper Sparrows (Ammodramus savannarum) and Eastern Meadowlarks (Sturnella magna) were not negatively affected by Steer+Sheep relative to Steer treatments, whereas there was evidence of a negative effect in these same metrics for Dickcissels (Spiza americana). Abundance of butterflies and their nectar sources were similar between treatments but abundance of grassland specialist butterfly species was low, overall. Comprehensively, Steer+Sheep effectively controls the spread of sericea but may not create habitat for all tallgrass prairie wildlife species.

The Snowy Plover (Charadrius nivosus) is unique in being a determinate layer of an odd modal clutch size and in having a variable mating system in which female brood desertion occurs regularly. These traits make determining Snowy Plover offspring sex ratios important not only for long-term population stability, as the species is of conservation concern, but also for application to sex allocation theory. In this study, we determined Snowy Plover offspring sex ratios, examined differential costs of producing male and female offspring, and evaluated sex ratio variation in relation to maternal condition, habitat condition, and time during the nesting season on saline lakes of the Southern High Plains of Texas. Examination of 245 chicks from 118 clutches during 1999–2000 and 2008–2009 showed that male offspring were more costly to produce than female offspring; however, offspring sex ratio did not differ from parity, but was slightly male-biased in most years. The probability of producing a male offspring was greater both earlier and later in the breeding season than in the middle. As the availability of saline lake surface water and the subsequent availability of food vary unpredictably throughout the breeding season, depending on precipitation events, we suggest that sex ratio adjustment in unpredictable environments may not be straightforward and may follow nonlinear models and/or vary annually The effects such changes in sex ratios may have on population growth and stability remain unknown.

Isolated wetlands are often degraded by agriculture, increasing sediment accretion and altering plant composition. Two common opposing wetland restoration practices are self-design vs. intensive revegetation. Self-design restores hydrogeomorphology without inoculating wetland taxa into restoration sites. Self-design may not meet restoration targets if dispersal-limited plants do not colonize restoration sites. Alternatively, intensive revegetation (hydrogeomorphic restoration combined with revegetation) is costly and time consuming. We investigated plant dispersal-limitation in 309 isolated wetlands among two agricultural landscapes in the U.S. Great Plains (the western High Plains (WHP) and the Rainwater Basin (RWB)) and three land-uses (reference, croplands, and previous croplands) to predict optimal restoration practices. We present analytical tools predicting whether self-design or intensive revegetation will be more successful elsewhere. In the WHP and RWB, perennial wetland cover was 61% and 31% greater in reference than in other land-uses. Distance to the nearest reference wetland explained perennial wetland richness in both regions, and area of reference wetlands within 15 km also was important in the WHP. Annual wetland species were over-represented in previous cropland wetlands and were less influenced by landscape isolation. We analytically identified dispersal-limited and cosmopolitan species in reference wetlands, with distance to reference wetlands and area of surrounding reference wetlands important in determining composition. Further, dispersal-limited plants in reference wetlands had greater cover in clustered than isolated wetlands in previous croplands. Plant community patterns in reference conditions may predict composition in restored wetlands. This aids selection of self-design or revegetation restoration approaches for individual plant species in isolated wetlands.

We quantified changes in long-term Snowy Plover (Charadrius nivosus) survey counts and return rates, estimated current sex and age-specific apparent survival and encounter rates, and calculated recruitment thresholds needed to maintain a stable population in the Southern High Plains of Texas. Mean survey counts of adult Snowy Plovers decreased by 78% at one saline lake (from 80 adults/survey to 18 adults/survey) from 1999–2000 to 2008–2010 but remained consistent at an alternate lake (from 45 adults/survey to 41 adults/survey). Adult and juvenile return rates have similarly declined within this time frame by 25 and 62%, respectively. Long-term declines in return rates may be the result of increased mortality from declining habitat conditions either within or outside the breeding season. Current estimates of adult (65%) and juvenile (12%) apparent survival are lower than most other estimates for Snowy Plovers throughout their range. Current estimates of adult and juvenile apparent survival and ret...

Citation: Earl, J. E., Fuhlendorf, S. D., Haukos, D., Tanner, A. M., Elmore, D., & Carleton, S. A. (2016). Characteristics of lesser prairie-chicken (Tympanuchus pallidicinctus) long-distance movements across their distribution. Ecosphere, 7(8). doi:10.1002/(ISSN)2150-8925

Managing for species using current weather patterns fails to incorporate the uncertainty associated with future climatic conditions; without incorporating potential changes in climate into conservation strategies, management and conservation efforts may fall short or waste valuable resources. Understanding the effects of climate change on species in the Great Plains of North America is especially important, as this region is projected to experience an increased magnitude of climate change. Of particular ecological and conservation interest is the lesser prairie‐chicken (Tympanuchus pallidicinctus), which was listed as “threatened” under the U.S. Endangered Species Act in May 2014. We used Bayesian hierarchical models to quantify the effects of extreme climatic events (extreme values of the Palmer Drought Severity Index [PDSI]) relative to intermediate (changes in El Niño Southern Oscillation) and long‐term climate variability (changes in the Pacific Decadal Oscillation) on trends in lesser prairie‐chicken abundance from 1981 to 2014. Our results indicate that lesser prairie‐chicken abundance on leks responded to environmental conditions of the year previous by positively responding to wet springs (high PDSI) and negatively to years with hot, dry summers (low PDSI), but had little response to variation in the El Niño Southern Oscillation and the Pacific Decadal Oscillation. Additionally, greater variation in abundance on leks was explained by variation in site relative to broad‐scale climatic indices. Consequently, lesser prairie‐chicken abundance on leks in Kansas is more strongly influenced by extreme drought events during summer than other climatic conditions, which may have negative consequences for the population as drought conditions intensify throughout the Great Plains.

Snowy plover (Charadrius nivosus) populations have declined throughout their range, in part because of habitat degradation and poor nest success, making information regarding regionally specific nest site selection and spatial patterns important when considering habitat conservation and management guidelines. We determined nest site selection characteristics (n = 180) and examined spatial patterns (n = 215) of snowy plover nests in saline lakes in the Southern High Plains (SHP) of Texas. At 104 nests, we examined the influence of substrate type on nest temperatures and heat mitigation. Snowy plover nests were more likely to be found near an object, on pebble substrate, and with fewer plants than random sites. High use areas were generally located in areas with pebble substrate and on human-made or natural islands, berms, and peninsulas. Overall, nests placed on pebble substrate had lower temperatures during the day than nests placed on sand substrates. Nest placement on pebble substrate may be valuable to nesting snowy plovers, providing thermal advantages to incubating adults and depressing potentially high nest predation rates. Management guidelines for this region should emphasize the importance of addressing key elements of snowy plover nesting habitat including the presence of pebble substrate and reducing vegetation encroachment. © 2012 The Wildlife Society.

Little is known about mate selection and lek dynamics of Lesser Prairie-Chickens (Tympanuchus pallidicinctus). We collected data on male territory size and location on leks, behavior, and morphological characteristics and assessed the importance of these variables on male Lesser Prairie-Chicken mating success during spring 2008 and 2009 in the Texas Southern High Plains. We used discrete choice models and found that males that were less idle were chosen more often for mating. Our results also suggest that males with smaller territories obtained more copulations. Morphological characteristics were weaker predictors of male mating success. Peak female attendance at leks occurred during the 1-week interval starting 13 April during both years of study. Male prairie-chickens appear to make exploratory movements to, and from, leks early in the lekking season; 13 of 19 males banded early (23 Feb–13 Mar) in the lekking season departed the lek of capture and were not reobserved (11 yearlings, 2 adults). T...

Depressional wetlands are predominant surface hydrological features providing critical societal ecosystem services in the semiarid United States High Plains. Critical wetland properties may be threatened because this 30 million ha short-grass prairie largely was converted from grassland to cropland. Further, the United States Department of Agriculture enrolled marginal cropland into the Conservation Reserve Program (CRP). CRP reduces topsoil erosion by planting permanent cover on croplands. In the High Plains, introduced tall-grasses primarily were planted in CRP, possibly reducing precipitation runoff, an important hydroperiod driver in wetlands. We assessed land-use influence on important wetland processes (wetland area, inundation, and plant composition) in 261 depressional wetlands called playas (87 each in native grassland, CRP, and cropland). Surveys spanned six states within three High Plains sub-regions (southern, central and northern). Playas averaged 8 ha in cropland and 16 ha in other land-uses. Plant composition in grassland playas was predominately native perennials, and upland plant cover equaled wetland plant cover. Cropland playas had fewer species/ha, generally more annuals than perennials and 80% greater exposed ground than other land-uses. CRP playas had 400% greater cover of introduced species (mostly upland perennial tall-grasses), which possibly inhibited catchment runoff, as CRP playas were inundated 56% less often than other land-uses. Therefore, tall grasses should not be planted in short-grass prairie CRP catchments, as they alter inundation frequency and vegetation communities in embedded wetlands. Conservation programs containing provisions to protect playas, including planting common native species and using grass buffers to control erosion into wetlands, should be promoted.

Lesser prairie-chickens (Tympanuchus pallidicinctus) are traditionally monitored by spring road- based lek surveys and counts of males attending leks. Several weaknesses exist with ground-based monitoring methods such as the bias of restricting surveys to roads, unknown probability of lek detection, and man-hours required to survey large tracts of habitat. We evaluated aerial surveys to locate lesser prairie-chicken leks in Texas and New Mexico using a Cessna 172 airplane (C172), R-22 Beta II helicopter (R-22), and R-44 Raven II helicopter (R-44) during spring 2007-2008. We determined lek activity during surveys with remote cameras placed on leks and cross-referenced time on the photo frame to time on our Global Positioning System flight log. From remote cameras we found that 305 leks were available for detection during survey flights. We determined lek detectability was 32.7% (95% CI ¼ 20.3-47.1%) in the C172, 72.3% (64.50- 79.14%) in the R-22, and 89.8% (82.0-95.0%) in the R-44. We created 16 a priori logistic regression models incorporating aircraft platform, distance to lek, survey date, lek size, and lek type to explain lek detection from aerial surveys. Our top ranked model included platform, distance, and lek type (model weight; wi ¼ 0.288). We had four competitive models and model averaged to draw inferences. Model averaging showed that detectability was generally greatest with the R-44, followed by the R-22, and lowest with the C172, with a slight deviation from this ranking at increased distances. Within our transect width, model averaging also suggested that detectability decreased as distance from the transect to the lek increased during helicopter surveys, and detectability increased as distance from the transect to the lek increased during C172 surveys. Furthermore, man-made leks were more likely to be detected than natural leks and large leks were more likely to be detected than medium or small leks. Aerial surveys effectively locate new leks and monitor lek density, and alleviate weaknesses associated with ground-based monitoring. We recommend using the R-44 to conduct lek surveys while flying at an altitude of 15 m at a speed of 60 km/hr on sunny mornings. 2011 The Wildlife Society.

Mottled ducks Anas fulvigula are nonmigratory and primarily depend on habitats associated with coastal marshes of the Gulf Coast for their entire life cycle. Much of remaining available coastal marsh habitats of the Texas Gulf Coast are managed by state and federal agencies. The Texas Gulf Coast breeding population of mottled ducks has recently declined dramatically. To aid in development of management plans for breeding mottled ducks, we estimated the number of potential pair ponds available on the Texas Chenier Plain National Wildlife Refuge Complex relative to marsh type (fresh, intermediate, brackish, and saline), estimated occupancy, compared habitats of used and unused pair ponds, and evaluated pair pond selection relative to marsh type during spring 2004 and 2005. Greater than 18,000 ponds were recorded in >22,400 ha of coastal marsh with >74% of potential pair ponds in the intermediate marsh type. Average pond size of 425 randomly selected ponds across marsh types was 0.09 ha. Pond occupancy was estimated as 1.3% and 2.5% in 2004 and 2005, respectively, in 634 ponds surveyed for presence of mottled duck pairs. Mottled ducks selected for ponds in fresh marsh with short surrounding vegetation associated with recent cattle grazing, but avoided recently (

Unlike other North American prairie-nesting dabbling ducks, northern pintail (Anas acuta) populations have not increased since the early 1990s and remain well below the long-term average for traditional survey areas. Previously reported estimates of annual survival and recovery rates for pintails did not investigate any spatial or temporal factors to explain annual variation of these rates. We used band- recovery data from 1970 to 2003 to test the influence of temporal periods defined by differing harvest regulations and habitat conditions of breeding grounds with spatially delineated regions on survival and recovery rates of northern pintails in North America. We separated regions based on a multiresponse permutation procedure to identify banding blocks with dissimilar recovery distributions based on a cluster analysis. We categorized time by grouping years into temporal periods based on bag limits, season lengths, or overflight versus nonoverflight years. We used the Brownie approach in Program MARK to evaluate 46 a priori models estimating survival and recovery rates. The best approximating model indicated that survival varied with age, sex, and region with additive time and interactive time-by-age and time-by-region effects. Recovery rate was best represented by a fully interactive term comprised of age, sex, region, and year. There were no statistical differences among average annual survival point estimates between age and sex classes within each region, and our estimates were similar to previous unpublished studies. We found the eastern region had decreased survival and increased recovery rates compared to other regions. Trends in pintail survival suggest that variation in annual survival was not the cause of the initial decrease in the northern pintail population and is unlikely the dominant factor preventing the population from increasing. The influence of other population parameters, such as recruitment rate, should be investigated to further evaluate other causes for the population status of northern pintails. Use of the top-ranked model to estimate annual survival and recovery rates for northern pintails in North America, which indicated that annually varying estimates of survival rates were better supported by the data than grouping years into temporal classes (i.e., based on bag limits, season lengths, and overflight yr) can be used by managers and policy makers when considering annual harvest regulations and effects of conservation efforts. Managers should incorporate these estimates into future demographic studies of pintails as well as consider using the top-ranked model for future analyses of band-recovery data.

Sand shinnery oak (Quercus havardii) communities are a unique component of grassland bird habitat in eastern New Mexico and have been impacted by human activities for decades. These communities are frequently managed with livestock grazing and herbicide application for shrub control, strategies that potentially can be used to restore the historical shrub–grass composition of this plant community. During spring migration and the breeding seasons of 2004 and 2005, we compared density and community structure of grassland bird species among four combinations of tebuthiuron application and grazing treatments that were being evaluated for restoration of shinnery oak communities. We performed biweekly point transects on sixteen 65-ha study plots in these communities. Density of all avian species combined did not differ between grazed and ungrazed plots. Tebuthiuron-treated plots had a 40% higher average density for combined species than untreated plots. There was a 41% higher average density of all species during spring 2005 than 2004, but density was similar during the breeding season of both years. These trends were predominantly influenced by densities of migratory Cassin’s Sparrow (Aimophila cassinii), which were greater in tebuthiuron-treated plots in both years. Densities of resident Meadowlarks (Sturnella spp.) exhibited little response to tebuthiuron or grazing treatments. Avian species richness, evenness, and diversity were only minimally affected by the tebuthiuron and grazing treatments. This study occurred over a period of highly variable precipitation, so future assessments, spanning longer wet–dry cycles and maturing plant communities, may be necessary to completely determine avian response to these restoration efforts.

Sand shinnery oak (Quercus havardii) communities, a major component of grassland habitat of birds in eastern New Mexico, frequently are managed with livestock grazing and herbicide application for control of shrubs. We examined nest density, nest-site selection, and daily rate of survival of nests of grassland birds among four combinations of treatments with tebuthiuron (0.75 kg/ha) and a short-duration, rotational-grazing system being used to restore sand shinnery oak communities in eastern New Mexico. During breeding seasons 2004 and 2005, we searched 4-ha subplots in four tebuthiuron-grazing-combination replicates for nests, measured vertical and overhead cover at each nest site and an associated random point, and estimated daily rate of survival of nests using program MARK. Density of nests was similar among all treatments but greater in 2005 than 2004. Although vertical cover differed among treatments and between years, it did not affect selection or success of nest sites. Overhead cover als...

Disturbance is considered one of the main factors influencing plant species composition and diversity. We conducted a field study to address the plant community response in a coastal marsh to a major disturbance. In 1992, muskrats (Ondatra zibethicus L.) completely removed vegetation within a 450 ha area of intermediate coastal marsh in Texas, USA. We used vegetation data collected prior to the disturbance (1989-1991) as a baseline for comparison to that recorded annually for a decade (1992-2002) following the disturbance. We compared species diversity, richness, relative abundance, evenness, and species similarity between pre and postdisturbance periods to evaluate the temporal response of the disturbed plant community. Plant diversity in the study area returned to predisturbance levels after 10 years. Species diversity in the study area had two peaks following the main disturbance. These peaks are associated with fluctuations of the water levels in the area. Our results suggest that it is possible to control the sedge or grass dominance in a community by subjecting the area to a carefully timed willful disturbance (e.g., grazing or fluctuating water levels) to achieve management goals. However, vegetation composition in the area changed from a grass-dominated (predisturbance) to sedge-dominated (postdisturbance) community. At the conclusion of sampling in 2002, plant species abundance, evenness, dominance, and richness conditions reached levels similar to predisturbance. However, the species composition after a decade postdisturbance differed from that during the predisturbance period. Therefore, even though we are able to predict the return of species diversity, evenness, and richness of a community after a period following disturbance, the actual species composition of an intermediate marsh following recovery is difficult to determine accurately, as it is contingent on several biotic and abiotic conditions that prevail while the system recovers from disturbance.

Overwinter body condition (e.g., fat) provides an index to the health of northern pintail (Anas acuta) populations and may be a factor in the decline of the continental pintail population that has been previously overlooked or understated. We compared body condition between 1984–1985 and 2002–2003, and found that body condition of pintails arriving during early winter in the Playa Lakes Region (PLR) of Texas, USA, has declined by an average of 32%. Body fat levels declined at varying levels with juvenile males showing the largest decline of 41%, followed by adult females with 39%, juvenile females with 30%, and adult males with an overall 18% decline. Declines are likely related to declines in migration and wintering habitat quantity and quality within the PLR and potentially across the Central Flyway. We recommend further acquisition and management of playas to ensure that these valuable habitats remain available to provide critical habitats for migrating and wintering pintails and other waterfo...

In response to continued low population numbers of Northern Pintail (Anas acuta) in North America and to increase knowledge of the geographic variation in pintail survival rates, we estimated 126-day (27 October–2 March) survival for male and female pintails wintering in the Middle Rio Grande Valley, New Mexico, during 2001–2002 (SY1) and 2002–2003 (SY2). Sixty-nine adult male and female pintails were marked with radio-transmitters and tracked throughout the study period. Weekly relocation data in relation to study year, sex, time (week), body condition at capture, and hunting seasons were modeled using the known-fate procedure in Program MARK. Year, sex, time, and body condition covariates did not improve model performance in estimating survival, so we used the most parsimonious model to produce an overall winter survival estimate of 0.597 ± 0.077 (95% C.I. = 0.442–0.735). Weekly survival estimates did not differ between hunting and nonhunting seasons. Male and female point estimates did not dif...

Shorebirds migrating through the Southern Great Plains (SGP), USA, use freshwater playas and saline lakes as stopovers. The importance of playas is well documented, but the role of saline lakes is not clearly understood. During 2002 and 2003, we conducted surveys to determine the extent to which the saline lakes serve as stopovers. Twenty-eight species were recorded, and total seasonal abundance ranged from 6779 to 29,924 birds. Potential shorebird abundance for extant saline lakes was estimated at 37,000-71,000 shorebirds annually. American Avocets (Recurvirostra americana), Western Sandpipers (Calidris mauri), Baird's Sandpipers (C. bairdi), Least Sandpipers (C. minutilla), Snowy Plovers (Charadrius alexandrinus), Killdeer (Charadrius vociferus), and Wilson's Phalaropes (Phalaropus tricolor) were the most abundant species. Community composition of shorebirds differed between saline lakes and regional freshwater playas. Peak spring abundance was generally in April, whereas summer/fall migration was more protracted and shorebird abundance peaked during 6-8 weeks in August and September. Migration chronologies differed among morphologically similar species, and among representative species from different guilds. Such patterns of temporal separation permit partitioning of resources by shorebirds migrating through the SGP. The saline lakes of the SGP should be regarded as stopover sites of regional and international value. To ensure that saline lakes function as stopovers and to help maintain those unique communities that inhabit them, conservation of saline lakes should focus on preserving spring flows and conserving water. SINOPSIS. Composici´

The continental population of northern pintails (Anas acuta; hereafter pintails) has declined since the late 1970s, possibly due to poor breeding propensity, low nest success, and low survival rates in response to habitat loss. Survival estimates are unavailable for several winter and migration areas including the Playa Lakes Region (PLR) of northwestern Texas, USA. We investigated winter survival rates, documented periods of mortality, and identified possible causes of mortality for 159 and 168 radiotagged female pintails during 23 October 2002–18 February 2003 and 10 October 2003–18 February 2004, respectively. We located all radiotagged pintails at least once weekly to verify survival status and used-known fate modeling in program MARK to test the influence of capture period, female age class, body mass, and capture location on survival rates. Cumulative survival for the 119-day period in 2002–2003 was 0.925 (95% CI = 86.0–96.3). During 2003–2004, for a 134-day period, survival estimates decli...