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Made available in DSpace on 2022-04-28T19:42:42Z (GMT). No. of bitstreams: 0 Previous issue date: 2021-10-01 Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) Deutsche Forschungsgemeinschaft A new petrified calamitalean is described from the Permian Motuca Formation, Parnaíba Basin, central-north Brazil. In contrast to all other three-dimensionally preserved Brazilian species, it represents a semi-self-supporting woody plant characterized by a distinctive branch system and cellular anatomy. Arthropitys buritiranensis sp. nov. is proposed to accommodate the new material. The branch architecture is composed of at least three types of branches: (i) principal woody side branches, (ii) up to three secondary woody branches per node, scattered along the stem or the principal side branches, and (iii) 8–11 leafy twigs per whorl, at every third, rarely the second node, without secondary growth. The wood is segmented into fascicular wedges and interfascicular rays, clearly visible only within the inner wood of the stem and woody branches. The tracheid walls of the secondary xylem have exclusively scalariform pits. Ray parenchyma cells have pitted horizontal walls. The new calamitalean was part of the riparian vegetation of tree ferns, calamitaleans and gymnosperms recorded in a distal fluvial plain setting. The wood shows irregular growth interruptions and, therefore, points to a seasonally dry climate that shaped even low latitude paleofloras of tropical Pangaea. UNESP - Rio Claro Institute of Geosciences and Exact Sciences, Department of Geology, Caixa Postal 178 Museum für Naturkunde, Moritzstraße 20 TU Bergakademie Freiberg Geological Institute, Bernhard-von Cotta Straße 2 In den Birkengärten 30 UNESP - Rio Claro Institute of Geosciences and Exact Sciences, Department of Geology, Caixa Postal 178 CNPq: 141365/2008-0 Deutsche Forschungsgemeinschaft: RO 1273/3-1 Deutsche Forschungsgemeinschaft: RO 1273/4-1

Made available in DSpace on 2020-12-10T17:34:20Z (GMT). No. of bitstreams: 0 Previous issue date: 2020-01-01 Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES) Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) The genus Leinzia is a typical member of the renowned Artinskian-Wuchiapingian (Permian) endemic bivalve fauna of the Passa Dois Group, Parana Basin, Brazil. The extraordinary shells of Leinzia, characterized by a rostrum extending from the anterior cardinal margin led certain authors to regard them as bivalved arthropods (Spinicaudata). Due to the unusual morphology and typically poor preservation of the available specimens, the taxonomic position of Leinzia still remains obscure. Leinzia has been variously referred either to the Pterioida, the Crassatelloidea, the Sanguinolitidae, or the Megadesmidae, or to the Pholadomyida. Herein, based on a detailed review of the topotype material and description of newly found specimens of Leinzia from the Serrinha Member, Rio do Rasto Formation, southern Brazil, we shed light on the taxonomic position of this genus. The hinge of the right valve with its large, blunt, anteriorly inclined subumbonal tooth and corresponding socket in the left valve coupled with the absence of true lateral teeth indicate close affinities to Pyramus and Cowperesia. Thus, the data here strongly suggest a Pachydomidae (Edmondioidea) rather than a Crassatelloidea affinity for Leinzia. Conversely, Leinzia differs from all other known Pachydomidae due to its anteriorly rostrate and posteriorly elongated shell. Finally, detailed stratigraphic data indicate that the vertical distribution of Leinzia is constrained to the middle part of the Guadalupian Serrinha Member of the Rio do Rasto Formation. Sao Paulo State Univ, Inst Biosci, Botucatu Campus,POB 510, BR-18618689 Botucatu, SP, Brazil Sao Paulo State Univ, Inst Geosci & Exact Sci, Rio Claro Campus, BR-13506900 Rio Claro, Brazil Sao Paulo State Univ, Inst Biosci, Botucatu Campus,POB 510, BR-18618689 Botucatu, SP, Brazil Sao Paulo State Univ, Inst Geosci & Exact Sci, Rio Claro Campus, BR-13506900 Rio Claro, Brazil

This study concerns silicified fertile pecopterid pinnae from the Permian of the Parnaiba Basin, State of Tocantins, northern Brazil, attributed to a new fern taxon of Marattiales, Buritiranopteris costata gen. nov. et sp. nov. This fern bears groups of three or four sporangia radially arranged in closely spaced synangia, as in some species of Scolecopteris and Acitheca, but they are completely enclosed in thick and long down-curved foliar lobes. This morphology may represent a xeromorphic adaptation against long seasonal drought and direct solar irradiation. These pinnae obviously belonged to one of the abundant marattialean tree ferns, most probably Tietea, which were apparently successful as riparian vegetation of ephemeral rivers.

The Pennsylvanian-Permian interval of the Paraná Basin is relatively well-known on its palynological content. However, little has been published from the Permian Serra Alta, Teresina and lower Rio do Rasto formations (Passa Dois Group). A total of thirty seven core samples were collected along a vertical range of 600 meters of these units in two boreholes in the Congonhinhas Municipality, northeastern Paraná State, South Brazil, but only the Teresina Formation and the lower part of the Serrinha Member of the Rio do Rasto Formation yielded identifiable palynomorphs. Of the thirty recorded species, ten species are registered for the first time in the basin, three are probably new, and five were previously documented at stratigraphic levels below the units of the present study. Assemblages are assigned to the Lueckisporites virkkiae Zone, regarded as Middle Permian (Guadalupian) in age, as suggested by other fossil and radiometric data. According to the relative abundance of some spore species, it is possible to discriminate two intervals: the first characterized by the predominance of Laevigatosporites vulgaris (from the Teresina Formation to the basal Serrinha Member), and the second dominated by Thymospora (in higher stratigraphic levels). The great abundance of fern spores in these assemblages is amazing because lycophyte stems are the best represented macrophytofossils of the Teresina Formation, similarly as glossopterid leaves and sphenophyte stems in the Rio do Rasto Formation. These aspects are discussed in the scope of lithofaciological and other paleontological data.

O estudo de fósseis e litologias da Formação Rio do Rasto (Grupo Passa Dois) ao longo de 11 estradas do sul ao nordeste do Estado do Paraná e de 1 estrada no extremo-norte de Santa Catarina deu suporte a correlações entre as colunas de diversas áreas, à definição de biozonas, à interpretação dos possíveis paleoambientes e a novas considerações sobre a história evolutiva da Bacia do Paraná durante o final do Permiano. A parte da seqüência estudada abrange as porções anteriormente designadas Membro Serrinha (cerca de 1/4 da coluna, a partir da base) e Membro do Morro Pelado, cujas características litológicas, no entanto, não parecem sustentar adequadamente estas subdivisões. Do sul ao centro-norte do Estado do Paraná, na área aflorante, a espessura da Formação Rio do Rasto atinge 420 m, diminuindo acentuadamente da região de São Jerônimo da Serra para a Santo Antônio da Platina, onde se encontram porções mais marginais da bacia. As análises dos fósseis (formalmente descritos em outros trabalhos especialmente preparados para esta dissertação) buscaram dados paleoecológicos, de correlação interbacinal, bioestratonômicos e bioestratigráficos; também fundamentaram a maior parte das correlações entre os afloramentos e entre as seções colunares compostas, tendo permitido estimar os mergulhos aproximados das camadas (entre 0,5º e 0,9º) e, em alguns casos, os rejeitos de falhas de gravidade (muito freqüentes na região do Arco da Ponta Grossa). A distribuição, a diversidade e a abundância de conchostráceos dão sustentação a zoneamento bioestratigráfico relativamente detalhado, comportando, em ordem ascendente, as Zonas Cyzicus sp., Monoleaia unicostata micropolygonata, Paranaleaia supina-Asmussia sp., Asmussia regularis (que compreende a Subzona Palaeolimnadiopsis subalata) e o \"Intervalo de Conchostráceos Superior\". Os conchostráceos evidenciam principalmente ambientes continentais de águas rasas, relativamente quentes e são típicos das regiões semi-áridas. Estes fósseis não lograram importantes correlações interbacinais. Os bivalves, relativamente comuns na parte inferior da formação, estão distribuídos nas Zonas Leinzia similis (inferior) e ? Palaeomutela platinensis (superior, com a Subzona Nothoterraia acarinata em sua porção inferior). Aquelas formas da segunda zona, incluindo algumas similares a espécies do Permiano Superior de outras regiões gondvânicas, certamente representam ambientes continentais de águas calmas. Já os bivalves da zona inferior parecem ser endêmicos, de águas um pouco mais agitadas, possivelmente com salinidade mais elevada. Os megafósseis vegetais possibilitaram o estabelecimento das Zonas Pecopteris dolianitii (inferior, com 3 subdivisões) e Schizoneura gondwanensis ( superior, com 2 subdivisões). Os elementos são típicos da última fase de desenvolvimento da Flora Glossopteris, permitindo correlação principalmente com o Grupo Ecca Superior, Grupo Beaufort Inferior e a Formação Madumabisa do sul da África e a Formação Raniganj da Índia. Estes fósseis ocorrem em relativa abundância, mas são pouco representativos da vegetação global e apresentam diversidade não muito elevada. Há diversos elementos com possíveis adaptações xeromórficas. A análise palinológica de uma amostra da localidade-tipo do Membro Serrinha, praticamente da base da Formação Rio do Rasto, sugere correlação principalmente com os palinomorfos dos depósitos da Zona Tapinocephalus da parte inferior do Grupo Beaufort Inferior. Endossando ainda as correlações anteriores indicadas pelo estudo de tetrápodes da Serra do Cadeado, a deposição da unidade na área estudada pode ter ocorrido no intervalo equivalente ao Kazaniano Superior-Tatariano Inferior, tendo possivelmente cessado um pouco mais cedo na região de Santo Antônio da Platina. Além dos fósseis mencionados, ainda foram constatados estromatólitos nodulares (particularmente na base da formação), pequenos gastrópodes, ostracodes, escamas de peixes (em maior abundância no nordeste do estado), e ainda icnofósseis (principalmente na porção inferior da formação). Estes fósseis apoiam as interpretações ambientais já mencionadas ou, pelo menos, não parecem indicar ambientes estritamente marinhos. Os depósitos sedimentares, quase todos avermelhados, exceto na porção basal da seqüencia, foram classificados em 17 tipos. O conjunto das evidências litológicas, bioestratonômicas e paleoecológicas sugere que a Formação Rio do Rasto representa deposição em grandes lagos rasos, com oscilação freqüente do nível de água, da área de recobrimento e possivelmente também da salinidade, sob condições climáticas variavelmente secas; supõe-se que os lagos eram limitados por terras preponderantemente baixas, com dunas eólicas em extensas áreas, vegetação melhor desenvolvida próximo às margens, e drenagem provavelmente efêmera, apenas efetiva nos eventos de chuvas torrenciais. Os lobos de suspensão e as sigmóides, acumulados nos lagos próximo às desembocaduras dos cursos de água, assim como os turbiditos das regiões de costa afora, devem atestar as maiores descargas durante as cheias. Em certas fases, principalmente durante a deposição da metade inferior da formação, a dinâmica dos lagos parece ter sido influenciada por intensas tempestades; oscilações mais fracas, como as provocadas pelo vento, deixaram marcas em diversos depósitos da base ao topo da seqüencia. Os dados apontam para a incidência de uma fase inicial de condições climáticas bastante secas, provavelmente sucedida por dois importantes intervalos de melhora climática; o primeiro intervalo um pouco mais úmido é caracterizado pelo desenvolvimento expressivo de glossopterídeas, pecopterídeas, bivalves e conchostráceos (M. unicostata micropolygonata); o segundo é representado, principalmente, pela ingressão de Schizoneura gondwanensis e pela máxima diversidade em conchostráceos; durante este intervalo pode ter havido ligeiro levantamento/basculamento tectônico da área de Reserva-Cândido de Abreu, sugerido, em parte, pela ocorrência de grandes intraclastos em dois diamictitos. O terço superior da formação evidencia novo acentuado recrudescimento climático, aparentemente com poucas fases de maior umidade, tendo provocado o desaparecimento de diversas espécies da fauna e da flora. Há registro de invasões cada vez mais freqüentes dos lagos por dunas eólicas, até a sua colmatação definitiva, a qual representa o início da deposição da Formação Pirambóia. Study of the Rio do Rasto Formation (Passa Dois Group) along 11 roads in southern to northeastern Paraná State and 1 road in northern Santa Catarina State, Brazil, has provided the basis for stratigraphical correlation of vertical profiles, establishment of biozones, interpretation of possible paleoenvironments and new inferences on the history of the Paraná Basin at the end of the Permian. The studied sequence comprises the Serrinha Member (the lowest 1/4 of the formation) and the Morro Pelado Member. However, the lithologies observed do not provide reasonable support for this subdivision. The thickness of the formation does not exceed 420 m in the studied area and decreases markedly from São Jerônimo da Serra towards Santo Antônio da Platina near the margin of the basin. Fossils were analyzed for interbasinal correlations and for paleoecological, biostratinomic and biostratigraphic interpretations. The fossils found the most important correlations among outcrops and vertical profiles. They also helped to estimate regional dips of the strata (around 0.5°- 0.9°) and, in some cases, the approximate offset of the vertical faults, very common in the region of the Ponta Grossa Arch. The geographical distribution, diversity and abundance of the conchostracans supports the establishment of the following biozones (in ascending order): Cyzicus sp. Zone, Monoleaia unicostata micropolygonata Zone, Paranaleaia supine-Asmussia sp. Zone, Asmussia regularis Zone (including the Palaeolimnadiopsis subalata Subzone) and the \"Upper Conchostracans Interval\". These organisms are typical of freshwater ponds and shallow lakes in warm, semi-arid regions. The conchostracans seem not to be very important in interbasinal correlations. Bivalves, most frequent in the lower part of the formation, are distributed in the Leinzia similis Zone (almost entirely at the base of the formation) and the? Palaeomutela platinensis Zone (including the Nothoterraia acarinata Subzone). The bivalves of the second zone, which comprise some forms similar to other Upper Permian Gondwanic species, probably lived in calm fresh waters, whereas the bivalves of the Leinzia similis Zone seem to be endemic, associated with more agitated waters and possibly with highter salinity.Two zones of fossil plants were defined: the Pecopteris dolianitii Zone (in the lower part, with 3 subdivisions) and the Schizoneura gondwanensis Zone (in the upper part, with 2 subdivisions). The elements are typical of the last phase of development of the Glossopteris Flora, as reported for the Upper Ecca Group, the Lower Beaufort Group and the Madumabisa Formation in southern Africa, and the Raniganj Formation in India. The fossil plants are relatively abundant, but not very representative of the regional vegetation. Neither zone shows great diversity. Elements with possible xeromorphic adaptations are common. Palynological analysis of a single sample from the type-locality of the Serrinha Member, almost at the base of the Rio do Rasto Formation, suggests correlation with forms of the Tapinocephalus Zone from the basal part of the Lower Beaufort Group. In accordance with previous observations based on the study of tetrapods from the Serra do Cadeado, the sequence in the studied area may have been deposited in the Late Kazanian to the Lower Tatarian; in northeastern Paraná State (Santo Antônio da Platina), however, deposition may have ended earlier. Other fossils observed in the formation, such as nodular stromatolites (mainly at the basal part of the unit), small gastropods, ostracodes, abundant fish scales (particularly in northeast Paraná) and trace fossils (mainly in the lower part to the formation), are consistent with the above-mentioned paleoenvironmental interpretations and do not include any strictly marine species. The sedimentary deposits were classified in 17 types. Lithologic, biostratinomic and paleoecological evidence suggests that the predominantly red sequence of the Rio do Rasto Formation accumulated in great shallow lakes characterized by water level and possibly salinity oscillations within and variable arid context. The lakes were surrounded by lowlands, which contained marginal vegetation, eolian dunes and probably ephemeral streams. Flood stages would have caused the deposition of suspension lobes and sigmoidal bars near the mouths of tibutaries and turbidites farther offshore in the lakes. Occasionally, especially during the deposition of the lower part of the sequence, the lakes were influenced by heavy storm episodes; weaker wave action, such as that produced by winds, also left marks in many stata, throughout the formation. The sequence apparently began to accumulate under quite arid conditions, followed by more humid conditions, probably in two distinct intervals. The expressive development of glossopterids, pecopterids, bivalves and conchostracans (M. unicostata micropolygonata) would have occurred in the first interval, whereas Schizoneura gondwanensis and several conchostracans appeared in the second interval. In the latter interval may have occurred mild tectonic uplift of the area near Reserva-Cândido de Abreu, as suggested by the presence of two anomalous diamictites with large intraclasts. The upper third of the formation documents the return of rigorous arid conditions, that alternated with only a few, short, less severe intervals. Many floral and faunal species disappeared at this time, and eolian dunes invaded the lakes with greater frequency, finally filling them up and setting the stage for deposition of the Pirambóia Formation.

Specimens of Schizoneura gondwanensis are here described from 13 new outcrops of the Rio do Rasto Formation (Passa Dois Group, Parana Basin, Upper Permian) in Parana State and northern Santa Catarina State. This species was previously known in Brazil from only two other locaiities of the same formation. The foliage shoots, preserved as impressions in red and gray mudstones, siltstones and fine sandstones, generally occur as fragments, which may, however, exhibit well-preserved details such as possible structures of the mesophyll and epidermis, only very rarely recognized in other Gondwana specimens of Schizoneura. The fossils also corroborate Feistmantel's observations made in 1880, but overlooked by many later authors, regarding the embracing of the leaf clusters at the nodes, commissural lines, veins, etc. Although the leaf clusters at one outcrop are oblong-oval and not oval-lanceolate as at the other outcrops, ali are tentatively considered as belonging to the same species. Several of the leaf clusters of S. gondwanensis present ruptures along the commissural lines, here interpreted as the result of taphonomic processes or, possibly, environmental factors that affected the living plant. Ontogenetic aspects of Schizoneura and comparisons with Cathaysian representatives of this genus are also discussed. S. gondwanensis thus has a much broader geographical range in Brazil than previously thought and, consequently, can now be considered as one more important element of Rosler's Taphoflora E

This paper describes a new species of Sphenophyllum (S. paranaensis n. sp.) from the Rio do Rasto Formation, Serrinha Facies, cropping out along the Dorizon-Paulo Frontin road (State of Parana, Brazil), 1.5 km south of the main church of Dorizon. The new species is based on isolated leaves, verticils and articulated stems. A few lascinated leaves which occur dispersed at the same outcrop may be interpreted as a polymorphic variation of the new species, but until further data become available they are here called Sphenovhyllum cf. S. thonii, In fact, both forms of this genus from Dorizon are similar to specimens assigned to S. thonii in Africa but smaller than type material of this species

This paper deals with Pteridophylla and Filicophyta from 23 outcrops of the Rio do Rasto Formation in Parana State and 2 outcrops of the Estrada Nova Formation in Sao Paulo State (Passa Dois Group, Parana Basin, Upper Permian). Equivalent fossils have been described previously from only 3 localities of these same stratigraphic units. The fossils occur as impressions of fragmented fronds and pinnae in red, gray and yellowish mudstones, siltstones and fine sandstones. All the better preserve d pecopterids correspond to new species: Pecopteris taguaiensis, P. dolianitii, P. esperancensis, P. bracatingaensis and P. cadeadensis. Other described specimens are considered as Pecopteris sp.1, P. sp.2, P. sp.3, ?Dizeugotheca bortoluzzii Cazzulo-Klepzig and? Dizeugotheca sp

Conchostracans associated with bivalves, gastropods, ostracodes, fish scales and plant remains (mainly glossopterids and sphenophytes) from 67 outcrops of the Rio do Rasto Formation (Passa Dois Group, Parana Basin, Upper Permian) in Parana and north of Santa Catarina States occur as molds of frequently flattened and not dismembered carapaces (or sometimes isolated valves) in ritmic or massive red, gray and yellowish mudstones, siltstones and fine sandstones. Asmussia regularis and Monoleaia unicostata, previously reported from only a few localities of this formation, in addition to one new genus and species - Paranaleaia supina, are the predominant species and thus have a much broader geographical distribution than previously thought. Other identified species include: Asmussia sp., Cyzicus (Euestheria) sp., C. (Lioestheria) sp., Cyzicus sp., Gabonestheria sp., Palaeolimnadiopsis subalata, cf. Palaeolimnadiopsis sp. and Palaeolimnadia (Palaeolimnadia) sp.. Probable egg capsules were found in association with a carapace of Monoleaia unicostata. The postero-dorsal margin of this species is sometimes preserved as a narrow area with fine longitudinal grooves, possibly related to the articulation of the valves. Two very distinct types of carapace sculpture present in different specimens of this same species may support a separation in two new subspecies: M. unicostata timbcensis and M. unicostata micropolygnata

O enfoque da presente tese consiste no estudo dos fósseis das formações Corumbataí e Estrada Nova, no Estado de São Paulo, como subsídio ao conhecimento paleontológico e bioestragráfico do Grupo Passa Dois. Na análise paleontológica são descritos: espículas de espongiários, bivalves, ostracodes, ictiodontes, escamas e coprólitos, principalmente de paleoniscídeos, estruturas biogênicas de afinidade taxionômica desconhecida, oogônios de carófitas e estromatólitos. Foi também realizada análise palinológica de duas amostras do Membro Serra Alta e uma amostra da Formação Corumbataí. A distribuição dos bivalves possibilitou mudanças nos esquemas bioestratigráficos já estabelecidos para as unidades, com redefinições das biozonas Barbosaia angulata - Anhembia froesi, Pinzonella illusa - Plesiocyprinella carinata e Pinzonella neotropical - Jacquesia brasiliensis. Com o objetivo de correlacionar as condições paleoambientais às associações fossilíferas, foram realizadas análises geo químicas, para determinas as variações de boro, vanádio, gálio, rubídio, estrôncio, níquel e cromo ao longo de perfis e segundo às litologias, além das porcentagens de cálcio e magnésio. Também foram selecionadas cinco amostras do Membro Teresina para determinações da razão 87Sr/86Sr. No Membro Teresina, a associação das fácies carbonáticas, fósseis e dados geoquímicos indica ambiente costeiro, parcialmente isolado, de águas rasas e calmas. Os bioclastos e oóides presentes nas lamas carbonáticas foram transportados para esse corpo de água semi-confinado durante tempestades. This thesis aims at the study of the fossil assemblages from Corumbataí and Estrada Nova formations in the State of São Paulo and is a contribution to the understanding of the paleontology and biostratigraphy of the Passa Dois Group. Sponge spicules, bivalves, ostracodes, ichthyodonts, scales and fecal pellets, mainly of paleoniscids, biogenic structures of unknown taxonomic affinity, charophyte oogonia and stromatolites are described. lt is also presented the palinologic analysis of two samples from Serra AIta Member and one sample from the Corumbataí Formation. The distribution of bivalves allowed the redefinition of fhe Barbosaia angulate - Anhembia froesi, Pinzonella illusa - Plesiocyprinella carinata and Pinzonella neotropica - Jacquesia brasiliensis biozones proposed in the biostratigraphic schemes previously defined. With the objective to correlate the paleoenvironmental conditions with the fossil associations, geochemical analyses were carried out in order to determine boron, vanadium, galium, rubidium, strontium, nickel, lithium and chromium variations along profiles and according to lithologies in addition to the percentages of calcium and magnesium. Five samples were selected for 87Sr/86Sr ratio determinations. ln the Teresina Member, the association of the carbonatic facies, fossils and geochemical data indicate a partial isolated, coastal environment of calm, shallow waters. The bioclasts and ooids present in the carbonatic muds were transported to this semi-confined water body during storms.

O centro-leste do Estado de São Paulo expõe boa parte da seção neopaleozóica da Bacia do Paraná, com sedimentos do Grupo Tubarão (Subgrupo Itararé e Formação Tatuí) e o Grupo Passa Dois (formações Irati e Corumbataí). Na região entre Leme e Capivari e na região do Domo de Pitanga, entre Piracicaba e Ipeúna, a transição entre as formações Tatuí e Irati é caracterizada comumente por arenitos relativamente grossos e mal selecionados (arenitos com grânulos, arenitos conglomeráticos e conglomerados), abundantemente fossilíferos em contato abrupto com os folhelhos síltico-argilosos da Formação Irati. O presente trabalho concentrou-se no estudo de paleovertebrados paleozóicos aplicado à interpretação paleontológica, sedimentológica e estratigráfica da transição entre os grupos Tubarão e Passa Dois. O conteúdo fóssil identificado compreende escamas, dentes e partes ósseas de vários tipos de peixes, de água doce e salgada, e de possíveis tetrápodes. Os Chondrichthyes são representados por duas variedades de dentes cladodontes; vários tipos de dentes do Xenacanthiformes ?Pleuracanthus? albuquerquei Silva Santos, 1946; dentes de Orodontiformes; dentes de duas espécies de Petalodontiformes, a primeira, e mais comum, Itapyrodus punctatus Silva Santos, 1990 e a segunda, representada por um único exemplar, uma espécie indeterminada. O material ainda inclui um espinho de nadadeira dorsal de Ctenacanthiformes adulto e uma possível escama placóide. A fauna de Osteichthyes é a mais abundante, sendo composta de dentes e escamas ganóides de Paleonisciformes e escamas cosmóides de Actinistia. São observados grandes dentes labirintodontes (provenientes de tetrápodes primitivos e/ou? peixes Rhipidistia) e partes ósseas atribuídas a peixes e/ou possíveis anfíbios. Os fósseis estão normalmente dispersos e desarticulados e os elementos ósseos fragmentados e desgastados. Mesmo assim, foram encontrados dentes de animais continentais - labirintodontes e Xenacanthiformes, com diferentes tipos de preservação, o que indicaria uma influência fluvial. A mistura de elementos marinhos, como Petalodontes, e continentais sugere um caráter marinho costeiro com forte influência continental para a transição entre as unidades. Os estudos tafonômicos sugerem que a camada de estudo é um depósito residual, resultado direto do retrabalhamento por ondas. Portanto, a transição Tatuí - Irati faz parte da evolução deposicional da Formação Irati. Sugere-se que ela seja reconhecida como uma fácies basal local, de grande importância paleontológica e estratigráfica no centro-leste do Estado de São Paulo. A good part of the Neopaleozoic section of the Paraná basin is well-exposed in central-east Sao Paulo, Brazil, represented by sedimentary rocks of the Tubarão Group (Itararé Subgroup and Tatuí Formation) and the Passa Dois Group (Irati and Corumbataí formations). Between Leme and Capivari and in the area of the Pitanga Dome, between Piracicaba and Ipeúna, the transition between the Tatuí and Irati formations commonly is characterized by abundantly fossiliferous, poorly sorted, relatively coarse sandstones (sandstones with granules, conglomeratic sandstones and conglomerate) in abrupt contact with silty shale of the Irati Formation. The present work concentrated on the study of Palaeozoic palaeovertebrates and their significance for paleontological, sedimentological and stratigraphical interpretations of the transition between the Tubarão and Passa Dois groups. The identified fossil content consists of scales, teeth and bony parts from several types of fish, from fresh and salt water, as well as from possible tetrapods. Chondrichthyes are represented by two varieties of cladodont teeth; several types of teeth of the xenacanth ?Pleuracanthus? albuquerquei Silva Santos, 1946; teeth of Orodontiformes; teeth of two species of Petalodontiformes, the first, and more common, Itapyrodus punctatus Silva Santos, 1990, and the other, represented by a single specimen, an unidentified species. The material also includes a fin spine of an adult Ctenacanthiformes and a possible placoid scale. The fauna of Osteichthyes is more abundant, being composed of teeth and ganoid scales of Paleonisciformes and cosmoid scales of Actinistia. Also observed are large labyrinthodont teeth (from primitive tetrapods and/or rhipidistid fish) and bony parts attributed to fish and/or possible amphibians. The fossils are usually disarticulated and dispersed and the bony elements fragmented and abraded. Even so, teeth of continental animals - labyrinthodonts and Xenacanthiformes, were found with different types of preservation, which is indicative of a fluvial influence. The mixture of marine elements, like petalodonts, and continental elements suggests a nearshore character with a strong continental influence for the transition between the units. Taphonomic studies suggest that the studied layer is a residual (lag) deposit that resulted directly from reworking by waves. Thus, the Tatuí - Irati transition is part of the depositional evolution of the early Irati Formation. It is therefore suggested that the transition be recognized as a local basal facies of this formation of great paleontological and stratigraphical importance in the central-eastern part of the state of Sao Paulo.