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Deferunda acuminata Chou & Wang, 1985 Figs 27–39 Deferunda acuminata Chou & Wang, 1985 (in Chou et al., 1985): 32. Material examined. 10♁♁, 8♀♀, Diaoluoshan National Natural Reserve, Hainan Province, 16 July 2007, Zheng-Guang Zhang, Qiong-Zhang Song and Ji-Chun Xing; 3♁♁, 6♀♀, DaTian National Natural Reserve, Hainan Province, 10 July 2011, Bing Zhang and Ji-Chun Xing; 4♁♁, 3♀♀, DaTian National Natural Reserve, Hainan Province, 4–6 April 2014, Hai-Yan Sun, Yuan Liu, Zheng-Xiang Zhou and Mei-Na Guo; 3♁♁, 8♀♀, Jianfengling National Natural Reserve, Hainan Province, 12 July 2007, Ji-Chun Xing and Qiong-Zhang Song. Measurements. Length (including forewing): male 3.4–4.1 mm (N = 20), female 3.4–4.5 mm (N = 25). Coloration. General color grayish yellow to brownish (Figs 27, 28). Head (Figs 27, 29) grayish white. Vertex (Figs 27, 29) with 2 long fuscous stripes along midline from apical third to apex, posterior margin with a brown transverse stripe each sides. Frons (Fig. 30) grayish white, with an inverted chevron shaped marking, lateral margins with 2 slender and 1 broad dark brown transverse stripes, middle area white, each side with an oblique dark brown wedge-shaped stripe. Genae (Fig. 28) grayish white, with 3 fuscous transverse stripes before eyes (2 above and 1 below ocelli), 1 short transverse stripe dorsad, area beneath antenna with a yellowish brown marking. Eyes fulvous, ocelli reddish-brown. Antennae yellowish brown to brown. Lateral areas of pronotum (Figs 27, 29) with 1 brown spots on both sides. Mesonotum (Figs 27, 29) with irregular yellowish brown to fuscous markings. Tegula yellowish white to brownish. Forewings (Fig. 31) pale brown, many brown to fuscous spots scattered, veins with many white vesicular markings, callus blackish brown, tinged with light red marking. Hindwings (Fig. 32) brownish, base of veins brown. Abdomen fuscous, lateral margins brown. Head and thorax. Head (Figs 27, 29) rectangular. Vertex medially longer than wide at base (1.47: 1), disk depressed, with middle carina, lateral carinae strongly keeled. Frons (Fig. 30) near rhomboid, longer in midline than wide at widest portion (about 1.32: 1), with a weak median carina, lateral margins strongly foliate. Rostrum with apical as long as subapical segment. Pronotum (Figs 27, 29) short, in midline shorter than vertex (0.55: 1), with 3 longitudinal carinae. Mesonotum (Figs 27, 29) longer in midline than vertex and pronotum combined (1.73: 1), with 3 carinae. Forewing (Fig. 31) slender, longer than maximum width (2.82: 1), CuA 1 obviously convex distad of clavus, not reaching MP. Hindwings (Fig. 32) wide, longer than widest part (1.94: 1). Hind tibiae with a lateral spine. Male genitalia. Pygofer (Figs 36, 37) in lateral view, gradually widened from the back to the abdomen, anterior margin strongly concave in the middle, posterior margin distal half obliquely straight, with an obtuse angular process near the middle, medioventral process with a small obtuse process at the apex of dorsal margin; in ventral view, basal margin of pygofer broadly rounded, medioventral process tapering distally, apical margin angular. Anal tube (Fig. 33) nearly square in dorsal view, slightly wider at base than apex, apical margin with a wide angular depression medially, anal style long, apical margin rounded and convex, obviously extending out apical margin of anal tube. Aedeagus (Figs 38, 39), asymmetrical bilaterally, tubular, membranous, divided into 4 lobes at apex; in ventral view, ventral lobe shorter than dorsal lobe, apical margin angular concave medially, lateral margins serrated, with 7–8 dentate spikes, with a small straight process in the middle toward the base; in dorsal view, right and left dorsal lobes of phallobase separated, apical margins obtuse rounded, each with a row of serrate spines from middle to base, lateral margin of right dorsal lobe with a triangular process medially. Gonostyli (Figs 34, 35) bow-shaped, apex rounded and convex; in ventral view, outer margin of distal half with single, large, hornlike process; in dorsal view, distal half of dorsal margin bearing 1 pointed dentate process oriented inward, middle area with 1 twisted broad enlarged process oriented outward, basal side with an obtuse angular process, outer side with a fingerlike process. Remarks. Chou & Wang (in Chou et al., 1985) described the D. acuminata based on one male specimen from Xinglong, Hainan province, China. In our study we examined topotypic material (20 male specimens) which we identified as D. acuminata based on external morphological characteristics which according to the original description appear completely congruent to those of D. acuminata. Distribution. China (Hainan) (Fig. 40).

Deferunda dentata sp. nov. Figs 1–13 Measurements. Length (including forewing): male 3.0–4.3 mm (N = 4), female 3.7–4.5 mm (N = 9). Diagnosis. The salient features of the new species include: frons (Fig. 4) centrally with an inverted chevron shaped dark mark; forewing (Fig. 5) with CuA 1 obviously convex distad of clavus, reaching MP; medioventral process of pygofer (Fig. 11) constricted abruptly at end, apical margin rounded and convex in ventral view; aedeagus (Figs 12, 13) asymmetrical bilaterally, ventral lobe cleft medially, each side of midline with two dentate processes at proximal part; gonostyli (Fig. 9) with three processes along the lateral margin. Coloration. General color yellowish white to fuscous (Figs 1–2). Vertex (Figs 1, 3) yellowish brown with 2 long fuscous stripes along midline. Frons (Fig. 4) yellowish white, with an inverted chevron shaped marking, lateral carinae with 3 brownish black transverse bands. Clypeus (Fig. 4) yellowish brown. Genae (Fig. 2) yellowish white, with 3 fuscous transverse stripes before eyes (2 above and 1 below ocelli), 1 short transverse stripe dorsad, area beneath antenna with a fulvous marking. Eyes brown, ocelli reddish-brown. Antennae yellowish brown to brown. Pronotum (Figs 1, 3) yellowish brown, longitudinal carinae yellowish white, lateral areas with 4 fuscous spots, middle area yellowish-white to yellowish-brown. Mesonotum (Figs 1, 3) yellowish brown, with multiple large or small brown to fuscous markings. Tegula yellowish fuscous. Forewings (Fig. 5) pale brown to brown, many white spots scattered, most of them near veins, with narrow brown markings along veins, callus blackish brown, tinged with reddish orange margin. Hindwings (Fig. 6) pale brown, veins brown. Abdomen fuscous, lateral margins brown. Head and thorax. Head (Figs 1, 3) narrow. Vertex medially longer than wide at base (1: 0.71), disk depressed, apex slightly acute, anterior margin slightly truncate, posterior margin sinuate, median carina distinct, lateral carinae strongly ridged. Frons (Fig. 4) near rhomboid, longer in midline than wide at widest portion (about 1.13: 1), median carina with basal half obsolete, lateral margins strongly foliate. Rostrum with apical and subapical segment equally long. Pronotum (Figs 1, 3) short, in midline shorter than vertex (0.61: 1). Mesonotum (Figs 1, 3) longer in midline than vertex and pronotum combined (1.60: 1). Forewings (Fig. 5) slender, longer than maximum width (2.51: 1), CuA 1 obviously convex distad of clavus, reaching MP. Hindwings (Fig. 6) longer than widest part (1.7: 1). Hind tibiae with one lateral spine. Male genitalia. Pygofer (Figs 10, 11) in lateral view, gradually widened from the back to the abdomen, anterior margin strongly concave in the middle, posterior margin with a short angular process near the middle; in ventral view, medioventral process stout, constricted abruptly at end, apical margin rounded and convex. Anal tube (Fig. 7) in dorsal view length nearly equal to width, apical margin emarginate medially, anal style slightly extending out apical margin of anal tube.Aedeagus (Figs 12, 13) longer than phallobase (2.50: 1), asymmetrical bilaterally, tubular, membranous, divided into 4 lobes at apex; in ventral view, lateral margins with two and one angular processes, middle part with a thick spiny process, directed basad, ventral lobe cleft medially, each side of midline with two dentate processes at proximal part; in dorsal view, dorsal lobe rounded and convex at apical part, left side with 5 spines at basal half, right side with 3 spines. Gonostyli (Figs 8, 9) bow-shaped, apex curved inward, apical margin emarginate; in dorsal view, lateral margin with inverted fingerlike, large hornlike and short rounded processes at 4/5, 2/3 and 2/5 of the proximal part, respectively. Type material. Holotype: ♁, CHINA: Laoshan, Qingdao, Shandong Province, 16 August 2011, Jian-Kun Long; Paratypes: 3♁♁, 9♀♀, Laoshan, Qingdao, Shandong Province, 16–17 August 2011, Jian-Kun Long. Etymology. The specific name is derived from the Latin adjective “ dentatus ”, referring to the number of dentate processes on each side of midline of phallobase. Remarks. This species is similar to Deferunda rubrostigmata (Matsumura, 1914), but differs from the latter in: (1) vertex yellowish brown (vertex milk-white in D. rubrostigmata); (2) medioventral process of pygofer constricted abruptly at end, apical margin rounded and convex (medioventral process of pygofer tapering apically, apex angular in D. rubrostigmata); (3) gonostyli with three processes along the lateral margin (gonostyli with single process medially in D. rubrostigmata); (4) ventral lobe of aedeagus each side of midline with two dentate processes at proximal part (ventral lobe of aedeagus each side of midline with one dentate process at proximal part in D. rubrostigmata). Distribution. China (Shandong) (Fig. 40)., Published as part of Lv, Shasha, Chen, Xiangsheng & Long, Jiankun, 2023, Two new species of the planthopper genus Deferunda Distant from China (Hemiptera: Fulgoromorpha: Achilidae), with the first description of the male of Deferunda acuminata Chou & Wang, pp. 483-493 in Zootaxa 5256 (5) on pages 486-488, DOI: 10.11646/zootaxa.5256.5.4, http://zenodo.org/record/7758970, {"references":["Matsumura, S. (1914) Beitrag zur Kenntnis der Fulgoriden Japans. Annales Historico-Naturales Musei Nationalis Hungarici, 12, 261 - 305."]}

Deferunda interanea sp. nov. Figs 14–26 Measurements. Length (including forewing): male 3.7–4.2 mm (N = 19), female 3.8–4.7 mm (N = 8). Diagnosis. The salient features of the new species include: frons (Fig. 17) without dark mark; forewing (Fig. 18) with CuA 1 obviously convex distad of clavus, not reaching MP; pygofer (Fig. 23) in lateral view with a short angular process near ventral margin, medioventral process of pygofer with an angular short process proximal to apical inner margin; aedeagus (Figs 25, 26) asymmetrical bilaterally, lateral margins of ventral lobe serrated in ventral view, in dorsal view, right dorsal lobe with a row of spines at base, lateral margin of left lobe serrated at the basal third. Coloration. General color yellowish white to fuscous (Figs 14–15). Vertex (Figs 14, 16) yellowish white with 2 long fuscous stripes along midline from apical 2/3 to apex, lateral carinae yellowish-brown. Frons and clypeus (Fig. 17) yellowish white, apex of postclypeus yellowish brown. Genae (Fig. 15) yellowish white, with 3 fuscous transverse stripes before eyes (2 above and 1 below ocelli), 1 short transverse stripe dorsad, area beneath antenna with a fulvous marking. Eyes fulvous, ocelli reddish-brown. Antennae yellowish brown to brown. Pronotum (Figs 14, 16) yellowish white, lateral areas with 4 fulvous spots, dark brown behind near eyes. Mesonotum (Figs 14, 16) yellowish brown, with irregular brown to fuscous markings, base of middle area yellow. Tegula yellowish fuscous. Forewings (Fig. 18) pale brown to brown, many brown to fuscous spots scattered, veins with many white vesicular markings, callus blackish brown, tinged with light red marking. Hindwings (Fig. 19) pale brown, veins brown. Abdomen fuscous, lateral margins brown. Head and thorax. Head (Figs 14, 16) narrow. Vertex medially longer than wide at base (1.12: 1), disk depressed, apex slightly acute, anterior margin rounded and convex, posterior margin sinuate, middle part concave angularly, median carina with basal half distinct, lateral carinae strongly ridged. Frons (Fig. 17) near rhomboid, longer in midline than wide at widest portion (about 1.23: 1), median carina distinct, dorsal margin truncate, lateral margins strongly foliate. Rostrum with apical and subapical segment equally long. Pronotum (Figs 14, 16) short, in midline shorter than vertex (0.60: 1). Mesonotum (Figs 14, 16) longer in midline than vertex and pronotum combined (1.91: 1). Forewings (Fig. 18) slender, longer than maximal width (2.90: 1), CuA 1 obviously convex distad of clavus, not reaching MP. Hindwings (Fig. 19) longer than widest part (1.67: 1). Hind tibiae with a lateral spine. Male genitalia. Pygofer (Figs 23, 24) in lateral view, gradually widened from the back to the abdomen, anterior margin strongly concave medially, posterior margin with a short angular process near the middle, near ventral margin with a short angular process, medioventral process with an angular short process proximal to apical inner margin; in ventral view, basal part of medioventral process wide, constricted slightly in the middle, apical margin rounded and convex. Anal tube (Fig. 20) in dorsal view longer than wide (0.71: 1), apical margin emarginate medially, anal style stout, obviously extending out apical margin of anal tube. Aedeagus (Figs 25, 26) longer than phallobase (2.32: 1), asymmetrical bilaterally, tubular, membranous, divided into 4 lobes at apex; in ventral view, ventral lobe shorter than dorsal lobe, ventral lobe cleft at apex medially, from the base of the median cleft extends a slender long spike, toward the base, apex of left and right ventral lobes with a curved spur near the lateral margin, lateral margins of ventral lobe serrated, left of phallobase with a broad angular pointed process; in dorsal view, right and left dorsal lobes of phallobase with a broad cleft, right dorsal lobe with a row of spines at base, lateral margin with a large angular process medially; lateral margin of left lobe serrated at the basal third. Gonostyli (Figs 21, 22) bow-shaped, apex rounded and convex, apical margin emarginate; in ventral view, outer margin of distal half with 1 large angular process; in dorsal view, distal half of dorsal margin bearing 2 pointed dentate processes oriented inward, middle area with one twisted broad enlarged process, inner and basal margins with an obtuse angular process, respectively, outer margin with a fingerlike process. Type material. Holotype: ♁, CHINA: Jianfengling National Natural Reserve, Hainan Province, 14 January 2011, Jian-Kun Long; Paratypes: 15♁♁, 4♀♀, Jianfengling National Natural Reserve, Hainan Province, 13–16 January 2011, Jian-Kun Long and Wei-Bing Zheng; 3♁♁, 4♀♀, Jianfengling National Natural Reserve, Hainan Province, 8 April 2015, Jian-Kun Long, Ji-Chun Xing and Yu-Bo Zhang. Etymology. The specific name is derived from the Latin adjective “ interaneus ”, referring to the medioventral process of pygofer with an angular short process proximal to the apical inner margin. Remarks. This species is similar to Deferunda lua Long, Yang & Chen, 2013, but differs from the latter in: (1) apex of postclypeus yellowish brown (postclypeus yellowish white in D. lua); (2) medioventral process of pygofer with an angular short process proximal to apical inner margin (medioventral process of pygofer without an angular short process proximal to apical inner margin in D. lua); (3) middle part of medioventral process constricted in ventral view (apex of medioventral process constricted abruptly in ventral view in D. lua); (4) phallobase with 1 angular process on the left side (phallobase without 1 angular process on the left side in D. lua). Distribution. China (Hainan) (Fig. 40).

Lv, Shasha, Chen, Xiangsheng, Long, Jiankun (2023): Two new species of the planthopper genus Deferunda Distant from China (Hemiptera: Fulgoromorpha: Achilidae), with the first description of the male of Deferunda acuminata Chou & Wang. Zootaxa 5256 (5): 483-493, DOI: 10.11646/zootaxa.5256.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5256.5.4

Key to species of Deferunda Distant, 1912 1. Frons centrally with an inverted chevron shaped dark mark.................................................... 2 - Frons without dark mark............................................................................... 9 2. Hind tibiae without lateral spine..................................................... D. stigmatica Distant, 1912 - Hind tibiae with a lateral spine........................................................................... 3 3. Anterior margin of vertex truncate; apical third of forewing without milky-white spots (Chen et al. 1989: figs 28A, D).......................................................................... D. truncata Chen, Yang & Wilson, 1989 - Anterior margin of vertex acute; apical third of forewing with milky-white spots................................... 4 4. Vertex length subequal to pronotum in mid line..................................... D. philippina (Melichar, 1914) - Vertex distinctly longer than pronotum in mid line........................................................... 5 5. Lateral margins of frons without narrow dark stripes (Chen & He 2010: fig. 2).................................................................................................................... D. diana Chen & He, 2010 - Lateral margins of frons with several narrow dark stripes...................................................... 6 6. Vertex relatively short, as long in mid line as broad at base, median carina bordered with short longitudinal black stripes apically...................................................................... D. lineola (Matsumura, 1914) - Vertex relatively long, longer in median line than broad at base, median carina bordered with longitudinal dark stripes from apex to base......................................................................................... 7 7. Forewing with CuA 1 straight or slightly convex distad of clavus, not reaching MP (Chou & Wang 1985: fig. 8a)................................................................................... D. acuminata Chou & Wang, 1985 - Forewing with CuA 1 obviously convex distad of clavus, reaching MP............................................ 8 8. Medioventral processes of pygofer tapering to end, apical margin angular; phallobase in ventral view each side of midline with one dentate process at proximal part (Chen et al. 1989: figs 27L, H).................. D. rubrostigma (Matsumura, 1914) - Medioventral processes of pygofer constricted abruptly near apex, apical margin rounded and convex; phallobase in ventral view each side of midline with two dentate processes at proximal part (figs 11, 12)................... D. dentata sp. nov. 9. Forewing with irregular yellowish brown transverse stripes or markings (Wang et al. 2008: figs 10, 20–21)............................................................................................ D. striata Wang & Liu, 2008 - Forewing without above markings....................................................................... 10 10. Lateral margins of frons with 5 narrow dark brown stripes; costal margin of forewing red, apical region with 3 red triangular marks (Wang et al. 2008: figs 1, 2, 18)......................................... D. trimaculata Wang & Peng, 2008 - Lateral margins of frons without dark brown stripes; forewing without above marks............................... 11 11. Forewing with apex of clavus sanguineous with a large bluish-black spot, base of first apical cell sanguineous........................................................................................... D. majella (Kirkaldy, 1906) - Forewing not as above................................................................................ 12 12. Anal tube in dorsal view longer than widest part, apical margin rounded (Long et al. 2013: fig. 8)......................................................................................... D. ellipsoidea Long, Yang & Chen, 2013 - Anal tube in dorsal view shorter than widest part, apical margin slightly incised in middle........................... 13 13. Medioventral processes of pygofer narrow in the middle, thence diverging apically (Dlabola 1961: fig. 32)............................................................................................. D. incompta Dlabola, 1961 - Medioventral processes of pygofer not as above............................................................ 14 14. Clypeus and mesonotum dark brown; each medioventral processes of pygofer as right angled triangles; gonostyli with 1 process on outer margin................................................................ D. albomaculata (Muir, 1922) - Clypeus with apical half brown to fuscous or totally yellowish white; mesonotum yellowish brown, with irregular brown to fuscous markings; each medioventral processes of pygofer not as above; gonostyli with 2 processes................... 15 15. Phallobase in ventral view with 2 large forked processes arising from near middle of lateral margin, lateral margins without dentate process (Chen & He 2010: fig. 21)............................................ D. qiana Chen & He, 2010 - Phallobase in ventral view without 2 large forked processes arising from near middle of lateral margin, lateral margins with dentate process...................................................................................... 16 16. Medioventral processes of pygofer constricted slightly in the middle; phallobase with 1 angular process on the left side (figs 24, 25).................................................................................. D. interana sp. nov. - Medioventral processes of pygofer constricted abruptly at end; phallobase without angular process on the left side (Long et al. 2013: figs 26, 29)............................................................ D. lua Long, Yang & Chen, 2013, Published as part of Lv, Shasha, Chen, Xiangsheng & Long, Jiankun, 2023, Two new species of the planthopper genus Deferunda Distant from China (Hemiptera: Fulgoromorpha: Achilidae), with the first description of the male of Deferunda acuminata Chou & Wang, pp. 483-493 in Zootaxa 5256 (5) on pages 485-486, DOI: 10.11646/zootaxa.5256.5.4, http://zenodo.org/record/7758970, {"references":["Distant, W. L. (1912) Descriptions of new genera and species of Oriental Homoptera. The Annals and Magazine of Natural History, Series 8, 9 (50), 181 - 194. https: // doi. org / 10.1080 / 00222931208693118","Chen, C. L., Yang, C. T. & Wilson, M. R. (1989) Achilidae of Taiwan (Homoptera: Fulgoroidea). Taiwan Museum Special Publication Series, 8, 57 - 61.","Chen, X. S. & He, T. T. (2010) Two new species of genus Deferunda Distant (Hemiptera: Fulgoromorpha: Achilidae) from southwest China. Zootaxa, 2335 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 2335.1.6","Matsumura, S. (1914) Beitrag zur Kenntnis der Fulgoriden Japans. Annales Historico-Naturales Musei Nationalis Hungarici, 12, 261 - 305.","Chou, I., Lu, J. S., Huang, J. & Wang, S. Z. (1985) Homoptera, Fulgoroidea. Economic Insect Fauna of China. Fasc. 36. Science press, Beijing, 152 pp.","Wang, Y. L., Peng, L. F. & Liu, H. W. (2008) A taxonomic study on the genus Deferunda Distant from China (Hemiptera, Achilidae). Acta Zootaxonomica Sinica, 33 (4), 775 - 779.","Kirkaldy, G. W. (1906) Leafhoppers and their natural enemies (Pt. IX) Leafhoppers (Hemiptera). Bulletin of the Hawaiian Sugar Planters' Association, Division of Entomology, 1 (9), 271 - 479, pls. 21 - 32.","Long, J. K., Yang, L. & Chen, X. S. (2013) Review of Chinese species of Deferunda (Hemiptera: Fulgoromorpha: Achilidae) with descriptions of two new species. Florida Entomologist, 96 (4), 1263 - 1273. https: // doi. org / 10.1653 / 024.096.0404","Dlabola, J. (1961) Die Zikaden von Zentralasien, Dagestam und Trankav Kasien (Homoptera: Auchenorrhyncha). Acta Entomologica Musei Nationalis Pragae, 34, 241 - 358."]}

Wang, Xiaoya, Zhi, Yan, Chen, Xiangsheng (2020): Key to species of the genus Andixius Emeljanov & Hayashi (Hemiptera Fulgoromorpha: Cixiidae) with descriptions of two new species. Zootaxa 4802 (3): 440-448, DOI: https://doi.org/10.11646/zootaxa.4802.3.2

Arcofacies longispinus sp. nov. (Figs 1–12) Measurements. Body length including forewing: male 3.0– 3.1 mm (N = 2), forewing length: male 2.4–2.5 mm (N = 2). Diagnosis. The salient features of the new species include the following: anal segment of male with lateroapical processes symmetrical (Fig. 7); pygofer in posterior view with 4 spinous medioventral processes (Fig. 7); aedeagus with long spinous process at base, apex acute, directed ventrad, middle ventral margin of aedeagus with small process (Figs 9, 10); apex of genital style not forked (Figs 11, 12). Coloration. General color yellowish brown with green. Frons, vertex, pronotum and mesonotum with median carina white bordered with blackish brown (Figs 3–5). Eyes yellowish to blackish brown; ocelli reddish brown. Antennae with apex of scape blackish brown, pedicel yellowish white (Fig. 4). Lateral margins of gena, vertex and pronotum each with oblique white band bordered with brown or dark brown (Figs 3, 4). Clypeus yellowish white (Fig. 4). Forewings with pale brown over basal third, rest area hyaline, along transverse vein and apical veins with brown stripes as figured (Fig. 6), in dark portion veins with white spots. Wings hyaline with brown veins. Legs yellowish brown. Head and thorax. Vertex (Fig. 3) wider at base than long submedially about 0.97: 1. Frons longer in middle line than wide at widest part about 1.25: 1, widest above ocelli. Antennae surpassing frontoclypeal suture, scape long equal to wide, shorter than pedicel about 0.38: 1 (Fig. 4). Pronotum shorter than vertex about 0.50: 1. Mesonotum longer in middle line than vertex and pronotum combined about 1.55: 1 (Fig. 3). Forewing longer in middle line than wide at widest part about 2.37: 1 (Fig. 6). Male genitalia. Anal segment of male short, lateroapical angles produced into stout spinous process respectively (Fig. 7). Pygofer in posterior view ventral margin with four medioventral processes, finger-like (Fig. 7), in lateral view ventral angles distinctly produced (Fig. 8). Aedeagus with long spinous process at base, apex acute, directed ventrad, middle ventral margin of aedeagus with small process (Figs 9, 10). Genital styles long, reaching ventral margin of anal segment, broad at base, narrowing to apex (Figs 11, 12). Type materials. Holotype ♂, China: Yunnan, Yingjiang County (24°´44N, 97°33´E), on bamboo, 18 Aug. 2015, Xiang-Sheng Chen; paratype, 1♂, same data as holotype, Lin Yang. Remarks. This species is similar to A. varius sp. nov. but differs by: (1) anal segment of male with lateroapical processes symmetrical (Fig. 7) (anal segment with lateroapical processes asymmetrical, left one distinctly longer than right one in A. varius sp. nov.); (2) middle ventral margin of aedeagus with small process (Figs 9, 10) (middle ventral margin of aedeagus without process in A. varius sp. nov.); (3) genital styles in posterior view not distinct divergent (Fig. 7) (genital styles in posterior view distinct divergent in A. varius sp. nov.). Etymology. The specific name is a combination of the Latin words “ longus ” and “ spinus ”, referring to the aedeagus basally with a long spinous process. Host plant. Bamboo. Distribution. China (Yunnan).

Arcofacies varius sp. nov. (Figs 13–24) Measurements. Body length including forewing: male 2.9–3.0 mm (N = 2), forewing length: male 2.3–2.4 mm (N = 2). Diagnosis. The salient features of the new species include the following: anal segment of male with lateroapical processes asymmetrical, left one distinctly longer than right one (Fig. 21); pygofer in posterior view with 4 spinous medioventral processes (Fig. 19); aedeagus with long spinous process at base, apex acute, directed ventrad, middle ventral margin of aedeagus without process (Fig. 22); apex of genital style not forked (Figs 23, 24). Coloration. General color yellowish brown with green. Frons, vertex, pronotum and mesonotum with median carina white bordered with blackish brown (Figs 15–17). Eyes dark brown to blackish brown; ocelli reddish brown. Antennae with dorsal and ventral margins and apex of scape, base and near apex of pedicel dark brown to blackish brown (Fig. 16). Lateral margins of frons, gena, vertex and pronotum each with oblique white band bordered with brown or dark brown (Figs 15, 16). Cypeus yellowish white (Fig. 16). Forewings with yellowish brown over basal third, rest area hyaline, along transverse vein and apical veins with brown stripes as figured (Fig. 18), in dark portion veins with white spots. Wings hyaline with brown veins. Legs yellowish brown. Abdomen with dorsum dark brown. Head and thorax. Vertex (Fig. 15) wider at base than long submedially about 1.55: 1. Frons longer in middle line than wide at widest part about 1.26: 1, widest just above ocelli.Antennae surpassing frontoclypeal suture, scape long equal to wide, shorter than pedicel about 0.38: 1 (Fig. 16). Pronotum shorter than vertex about 0.70: 1. Mesonotum longer in middle line than vertex and pronotum combined about 1.84: 1 (Fig. 15). Forewing in middle line longer than wide at widest part about 2.34: 1 (Fig. 18). Male genitalia. Anal segment of male short, lateroapical angles produced into stout spinous process respectively, asymmetric, left one distinctly longer than right one (Fig. 21). Pygofer in posterior view ventral margin with four medioventral processes, finger-like (Fig. 19), in lateral view ventral angles distinctly produced (Fig. 20). Aedeagus (Fig. 22) with long spinous process at base, S-shaped curved, apex acute, directed ventrad. Genital styles divergent, long, reaching ventral margin of anal segment, broad at base, narrowing to apex (Figs 23, 24). Type materials. Holotype ♂, China: Yunnan, Mengla County (21°55´N, 101°15´E), on bamboo, 26 Apr. 2015, Zheng-Xiang Zhou; paratype, 1♂, same data as holotype, Qiang Luo. Remarks. This species is similar to A. strigatipennis Ding, 1990 but differs by: (1) anal segment of male with lateroapical processes asymmetrical, left one distinctly longer than right one (Fig. 21) (anal segment with lateroapical processes symmetrical in A. strigatipennis); (2) pygofer in posterior view with 4 spinous medioventral processes (Fig. 19) (pygofer in posterior view without medioventral process in A. strigatipennis); (3) aedeagus with long spinous process at base, middle ventral margin without process (Fig. 22) (aedeagus broad at base, nearly quadrate, middle of dorsal margin and ventral margin each with process in A. strigatipennis). Etymology. The specific name is derived from the Latin word “ varius ” (meaning different, changeable), referring to the anal segment with two asymmetrical processes. Host plant. Bamboo. Distribution. China (Yunnan).

Genus Arcofacies Muir, 1915 (Figs 1���24) Type species. Arcofacies fullawayi Muir, 1915 Arcofacies Muir, 1915: 319; Kuoh et al. 1983: 45; Yang & Yang 1986: 34; Ding 1990: 74; Ding et al. 1999: 442; Ding 2006: 115; Chen et al. 2007: 684; Hou & Chen 2010: 52. Diagnosis. General color yellowish green to yellowish brown. Frons, vertex, pronotum and mesonotum with median carina white bordered with dark brown or black. Lateral parts of pronotum each with oblique white band bordered with brown or dark brown. Forewings with light brown in basal third, apical portion hyaline, speckled with dark brown markings, in dark portion veins with white spots. Wings hyaline with brown veins. Head including eyes narrower than pronotum. Vertex trapeziform, with margins more or less well defined, wider at base than long submedially (1.70���1.88: 1), apical margin distinctly emarginate at both sides of median point, lateral carinae concave, submedian carinae transverse. Y-shaped carina without stalk, with very short arms, connecting submedian carinae which forms a small cell, in lateral view vertex and frons at right angle. Frons in middle line longer than wide at widest point (1.75���2.17: 1), widest at level of ocelli or at apex, lateral carinae convex at base, nearly straight below level of ocelli, median carina not well developed throughout, forked at extreme base. Postclypeus slightly wider at base than frons at apex, at right angle to frons, tricarinate. Rostrum almost extending to mesotrochanters. Eyes in dorsal view with lateral margin emarginated medially. Lateral ocelli present. Antennae cylindrical, scape distinctly longer than wide (1.60���2.00: 1), shorter than pedicel (0.52���0.59: 1). Pronotum with lateral carinae extending to hind margin, converging apically, median carina weak. Forewings tectiform at rest. M and Sc 1 of wing with a long common stalk, Cu 2 arising from tip of cross vein or basad. Spinal formula of hind leg 5-6-4. Anal segment of male collar-shaped, lateroapical angles produced into spinous processes or not. Pygofer in posterior view with opening longer than wide (1.29���1.6: 1), lateral margins strongly produced caudad medially or not, with a small medioventral process or not. Aedeagus tubular or flat, with spinous process or not, orifice subapical. Diaphragn armature sclerotized and pigmented, V-shaped. Diaphragm wide, membraneous. Genital styles long, simple, broad at base, narrowing apically, basal angle intumescent, apex twisting outward more or less (Hou & Chen 2010). Remarks. This genus is similar to Tropidocephala St��l and Arcofaciella Fennah but differs from them in the shape of the frons, vertex and forewings and in the relative size of the antennae (Liang & Jiang 2005 (2004)). Distribution. China (Fujian, Taiwan, Chongqing, Hong Kong, Hainan, Guizhou, Yunnan), Ceylon, Indonesia, Japan (Ryukyu Islands), Malaysia (Penang), Philippines (Luzon), Singapore., Published as part of Li, Hongxing, Yang, Lin & Chen, Xiangsheng, 2019, Two new species of the bamboo-feeding planthopper genus Arcofacies Muir (Hemiptera: Fulgoroidea: Delphacidae) from China, pp. 384-390 in Zootaxa 4706 (2) on page 385, DOI: 10.11646/zootaxa.4706.2.11, http://zenodo.org/record/3567318, {"references":["Muir, F. (1915) A contribution towards the taxonomy of the Delphacidae. Canadian Entomologist, 47, 317 - 320. https: // doi. org / 10.4039 / Ent 47317 - 10","Kuoh, C. - L., Ding, J. - H., Tian, L. - X. & Huang, C. - L. (1983) Economic insect fauna of China fasc. 27 Homoptera Delphacidae. Science Press, Beijing, xx + 166 pp.","Yang, J. - T. & Yang, C. - T. (1986) Delphacidae of Taiwan (I) Asiracinae and the tribe Tropidocephalini (Homoptera: Fulgoroidea). Taiwan Museum Special Publication Series, 6, 1 - 79.","Ding, J. - H. (1990) Notes on the genus Arcofacies in China (Homoptera: Delphacidae). Journal of Bamboo Research, 9 (1), 74 - 77.","Ding, J. - H., Huang, B. - K. & Zhou, W. - X. (1999) Delphacidae of Fujian (Homoptera: Fulgoroidea). In: Huang, B. - K. (Ed.), Fauna of Insects in Fujiang Province of China. Vol. 2. Fujian Science and Technology Publishing House, Fuzhou, pp. 432 - 464.","Ding, J. - H. (2006) Fauna Sinica Insecta. Vol. 45. Homoptera Delphacidae. Science Press, Beijing, xx + 775 pp.","Chen, X. - S., Yang, L. & Tsai, H. J. (2007) Review of the Bamboo delphacid genus Arcofacies (Hemiptera: Fulgoroidea: Delphacidae) from China, with description of one new species. Florida Entomologist, 90 (4), 683 - 689. https: // doi. org / 10.1653 / 00154040 (2007) 90 [683: ROTBDG] 2.0. CO; 2","Hou, X. - H. & Chen, X. - S. (2010) Description of one new species of oriental bamboo planthopper genus Arcofacies Muir (Hemiptera: Fulgoroidea: Delphacidae) from Yunnan, China. Acta Zootaxonomica Sinica, 35 (1), 52 - 56.","Liang, A. - P. & Jiang, G. - M. (2005 [2004]) Discovery of the genus Arcofaciella Fennah (Hemiptera: Fulgoroidea: Delphacidae) from the mainland China. Journal of the New York Entomological Society, 112 (4), 221 - 226. https: // doi. org / 10.1664 / 0028 - 7199 (2004) 112 [0221: DOTGAF] 2.0. CO; 2"]}

Li, Hongxing, Yang, Lin, Chen, Xiangsheng (2019): Two new species of the bamboo-feeding planthopper genus Arcofacies Muir (Hemiptera: Fulgoroidea: Delphacidae) from China. Zootaxa 4706 (2): 384-390, DOI: https://doi.org/10.11646/zootaxa.4706.2.11

Tetricodes songae Zhang & Chen, sp. nov. (Figs 11–19, 23–27) Description. Body length (from apex of coryphe to tip of forewings): ♁ 6.7 mm ♀ 6.8 mm; forewing length: ♁ 5.5 mm ♀ 5.6 mm. Coloration. Body (Figs 23, 24, 27) brown with greenish spots. Coryphe (Figs 23, 24) light green. Eyes brown. Metope (Fig. 26) dark brown,basal protuberance black. Clypeus and rostrum dark brown with light brown marks. Antenna light green. Pronotum light green, mesonotum pale brown. Forewings (Figs 23, 24) yellowish brown, lateral margins with dark brown marks, veins light green; hindwings dark brown. Legs light brown with dark brown or light green spots or stripes. Abdomen ventrally pale brown with pale greenish areas. Head and thorax. Coryphe (Fig. 11) wider than long (1.9:1). Metope (Fig. 12) longer than wide (1.4:1). Pronotum (Fig. 11) without median keel, each side convex, in midline slightly longer than vertex (1.7:1). Scutellum (Fig. 11) longer than pronotum (1.6:1) in midline. Forewing (Fig. 13) 2.6 times longer than widest part, Sc and R convergent 1/4 near base, each reaching to margin of wing. M bifurcate 1/3 near base. MA bifurcate 2/3 near base. MA1 and MA2 reaching to margin of wing, MP bifurcate near middle, MP1 only reaching to the union of MA1 and MA2. MP2 reaching to margin. Hind wings (Fig. 14) with length of incision about 0.3 times of hindwings. Hind tibiae with 2 teeth near apex. Male genitalia. Anal segment (Fig. 16) in dorsal view triangular, anterior margin weakly convex, apex with two finger-shaped lobes. Anal column near middle. Pygofer (Fig. 15) narrow, hind margin convex in middle. Aedeagus (Fig. 17) laterally each with a finger-like process near base and a oval flake 1/3 near apex. Aedeagus with ventral, lateral and dorsal lobes, apical margin of dorsal lobe straight, apically with a short ventral hook, lateral lobes apically with acute hook directed cephalad. Genital styles (Fig. 18) in lateral view oval, with a hook-shaped process in middle of dorsal margin. Type material. Holotype: ♂, Leigongshan National Natural Reserve (26°28´N, 108°17´E), Leishan, Guizhou Province, China, 1620–2178 m, 1 Aug. 2004, Q. Z. Song; Paratypes: 2♂♂, 1♀, same data as holotype (IEGU); 1♂, Leigongshan National Natural Reserve, Leishan, Guizhou Province, 17 Sep. 2005, Z. G. Zhang (IEGU); 1♀, Pianma (26°01´N, 98°37´E), Lushui, Yunnan Province China, 2600–2900 m, 17 Aug. 2000, X. S. Chen (IEGU). Etymology. The species name is derived from the last name of the first collector of the species, Q. Z. Song. Host plant. Unknown. Distribution. South China (Guizhou, Yunnan). Remarks. This new species is similar to T. polyphemus Fennah, 1956, but differs from the latter in: (i) larger body length including forewing ♂ 6.7 mm, ♀ 6.8 mm (♂ 5.8 mm, in T. polyphemus Fennah) (ii) M bifurcate 1/3 near base, MA bifurcate 2/3 near base, MA1 and MA2 reaching to margin of wing, MP bifurcate near middle, MP1 only reaching to the union of MA1 and MA2, MP2 reaching to margin; (iii) anal segment in dorsal view triangular, anterior margin weakly convex, apex with two finger-shaped lobes (anal segment oval laterally without finger-shaped processes at apex in T. polyphemus Fennah); (iv) aedeagus laterally with an oval flake 1/3 near apex, apical margin of dorsal lobe straight, apically with a short ventral hook, lateral lobes apically with an acute hook directed cephalad (aedeagus laterally with a square flake 1/3 near apex, apical margin of dorsal lobe convex in middle, apically without any ventral hooks, lateral lobes apically truncate but not hooked in the T. polyphemus Fennah)., Published as part of Zhang, Zhengguang & Chen, Xiangsheng, 2009, Review of the oriental Issid genus Tetricodes Fennah (Hemiptera: Fulgoroidea: Issidae) with the description of one new species, pp. 16-22 in Zootaxa 2094 on pages 19-22, {"references":["Fennah, R. G. (1956) Fulgoroidea from southern China. Proceedings of the California Academy of Sciences, 28 (4), 512 - 514."]}