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Polaszek, Andrew, Lahey, Zachary, Woolley, James B. (2020): Noyesaphytis (Chalcidoidea: Aphelinidae) - an unusual new genus from Madagascar, and a reassessment of Aphelininae classification based on morphology. Journal of Natural History 54 (9): 647-664, DOI: 10.1080/00222933.2020.1773559

Chartocerus hebes Girault, 1929 urn:lsid:zoobank.org:act: B7FBF276-AEC1-4F17-85CD-BA04AED6564E EOL taxon ID: 848102 (Figure 8; DOI:10.5281/zenodo.3893739) Girault (1929), as Matritia hebes (original description). Dahms (1984, p. 682): notes on type material. Type material. LECTOTYPE female [here designated]: QM T4416, slide, with 1 female (‘Hy.235 | Matritia hebes Girault | also wing ♀ cotype | Ent. Div. Dep. Ag. & Stk., Qld. 4416’). PARALECTOTYPES: 5 females. In the original description, Girault mentions ‘three females from spider eggs in a leafnest, Tasmania’ (Girault 1929, p. 311), but based on the labels and the fact the QM specimens are also labelled as ‘cotypes’, Dahms (1984) indicated the type series was actually composed of six specimens, considered syntypes. He reported on material at two institutions: 1 slide at SAM containing 3 females (Label: ‘ TYPE | Matritia hebes Girault | type ♀ S. Aus. Mus. Ent. Div. Dep. Ag. & Stk., Qld.’), plus 1 card and 1 slide labelled as cotypes at QM. The card contains 2 females and a glue spot where a third specimen (likely the slide one) was attached, labelled ‘ Tasmania || From eggs of spider in leaf nest || 4416 || Matritia hebes Girault | Cotype ♀ ’s | T 246546 (left), T 246547 (center)’, and the data from the QM material matches the original description. The slide-mounted female at QM (T 4416) is here designated as lectotype, and the slide has been labelled accordingly. Description. Female. Body dark, legs also dark, but posterior part of fore leg and apical part of metatibia lighter, tarsi yellowish. Wing infuscation differs from other species of Chartocerus because the apical part is darker than the basal part, with darkened areas extending below the stigmal vein. Head round in frontal view with deep punctations along inner margin of eyes, scattered on face and genae. Scape about 3× as long as wide, pedicel about 2× as long as wide, ring segments in ratios of 4:2:2:1, clava about 5× as long as wide. Scutellum about 1/3 of length of scutum. Fore wing with 2 setae on submarginal vein, 1 costal seta, 6 setae on marginal vein. Basitarsus of mid leg about 1/3 of length of mid leg. The propodeum and Mt1 were not cleared well enough in this preparation to be visualised; the same applies to the setation of the mesoscutum and scutellum., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on pages 693-696, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1929. Notes on, and descriptions of, chalcid wasps in the South Australian Museum. Concluding paper. Trans R Soc South Aust. 53: 309 - 346.","Dahms EC. 1984. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: III. Chalcidoidea species F-M with advisory notes. Mem Queensland Mus. 21: 579 - 842."]}

Chartocerus australiensis Ashmead, 1900 urn:lsid:zoobank.org:act: 7C9C7FB7-7B69-4477-9F7B-9AC18FB66A2A EOL taxon ID: 847913 (Figure 1; DOI:10.5281/zenodo.3893692) Ashmead (1900, p. 410), as Signiphora australiensis (original description). Girault (1913a): notes on type material, descriptive notes, host record, distribution record. Girault (1915): notes (republished from 1913a). Type material. LECTOTYPE female [here designated]: USNM Type No. 4771, mounted in balsam. ‘ Signiphora | australiensis Ash. | ♀ Type no. | 4771. USNM | Australia. | Koebele. 12.’ The original description does not provide further information. PARALECTOTYPE females: two specimens on same slide with same label data. Girault (1913a) redescribed the type material, having remounted the specimens on slides. The slide locality label is in Girault’s handwriting. One of the three females is damaged, embedded in resin that leaked outside of the coverslip. The head is detached from one of the other females (but it bears a complete antenna), and it is next to the third female. We here designate the female with detached head as lectotype, and the slide has been labelled accordingly. Description. Females. Length (pronotum to apex of epiproct): 0.5–0.8 mm (n = 3). Body predominantly dark, originally described as black with metallic tones, scutellum ‘with a bluish tinge’ and mesonotum ‘with a bronzy tinge’. It is perceptible in the type slide that the scutellum is slightly darker than most of the mesoscutum, metanotum and medial area of propodeum. Head dark, as in pronotum and most of metasoma, antennal clava and anelli lighter than pedicel and scape. Legs tan, lateroposterior part of protibia and mesotibia lighter, tarsi white. Fore wings infuscated, darker patch extending from base to apex of stigmal vein, lighter infuscation on apex, and two hyaline patches: one at basalposterior area, and one posterior to stigma vein, slightly curved towards base of wing. Head sculpture punctate and longitudinally striate, more conspicuous punctations along outer margin of antennal scrobes and inner margins of eyes, 1.3× as wide as high. Antennal scrobes well defined, extending through 3/4 of height of head. Other sulci absent. Eyes large, glabrous, about 2/3 of head height. Antenna with 4 anelli (1:1:1:1.2), clava 5× as long as wide, scape about ¾ length of clava; pedicel about 1/3 length of clava. Pronotum through metanotum striate or striate to reticulate, fainter in scutellum and metanotum; pronotum barely visible dorsally in the specimen; mesoscutum about 3.5× longer than scutellum, with about 13–15 small setae, most closer to medial-posterior area, scutellum with 7 setae aligned in parallel with posterior margin plus 1 larger seta on top of each axilla; internally marked axillae project anteriorly from scutellum to about ½ of mesoscutum; metanotum about 2/3 length of scutellum; medial triangle of propodeum finely reticulate. Protibia about 2/3 of length of mesotibia. Mesofemur with 3 large apical spines; mesobasitarsus a little more than 1/3 length of mesotibia, mesotibial spur about 2/ 3 of length of basitarsus; metatibia as long as mesotibia. Apex of medial sclerite of propodeum curved, reaching posterior margin of propodeal segment. Propodeal segment reaches but does not overlap the area between the lobes of Mt1. Fore wings about 2.5× as long as wide; longest setae of marginal fringe about ¼ of wing width; discal seta absent, 1 seta in submarginal vein; seta M1 absent; 5 small setae in basal area. Hind wings 4× as long as wide, longest setae of marginal fringe about ½ maximum width of wing; discal seta present; 2 setae on base of marginal vein. Mt1 bilobed, with slightly curved connection between the two lobes. Imbricate sculpture evident on tergites. Plates of ovipositor almost reach base of gaster. Biology. In the original description, Ashmead mentions an unidentified ‘rhynchotan’ (i.e. Hemiptera) as host. Girault (1913a) reported on another slide (USNM: not located) ‘Acanthococcid on Eucalyptus ’ from Koebele material collected in New South Wales., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on pages 683-685, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Ashmead WH. 1900. On the genera of the chalcid-flies belonging to the subfamily Encyrtinae. Proc US Natl Mus. 22: 323 - 412. doi: 10.5479 / si. 00963801.22 - 1202.323.","Girault AA. 1913 a. A systematic monograph of the chalcidoid Hymenoptera of the subfamily Signiphorinae. Proc US Natl Mus. 45: 189 - 233. doi: 10.5479 / si. 00963801.1977.189.","Girault AA. 1915. Australian Hymenoptera Chalcidoidea - VII. The family Encyrtidae with descriptions of new genera and species. Mem Queensland Mus. 4: 1 - 184."]}

Chartocerus ruskini Girault, 1921 urn:lsid:zoobank.org:act: ED82C36C-9356-4A7F-AC82-C5E0103EAD62 EOL taxon ID: 847839 (Figure 10; DOI:10.5281/zenodo.3893747) Girault (1921, p. 188), as Signiphora ruskini (original description). Dahms (1986, p. 491): notes on type material. Type material. Holotype female, QM T4406, on a slide in bad condition, labelled: ‘ Lathromeroidea nigrella ♂,♀, Anaphoidea linnaei 2 ♀’s, Signiphora ruskini Gir. Type. ♀ 4406 | (red label) 4406). There are several coverslip fragments with several other damaged specimens, some of which do not appear to be listed on the label. The slide does not have date information, but in the original description, Girault indicates ‘Nelson, forest, 6 March 1919 ’. The propodeal sclerite was dissected; head, legs, wings and antennae were also separated. The head is broken and partially under a corner of the coverslip that is cracked. Description. Female. Length ~ 0.6 mm. Body mostly brown; scutellum, mesoscutum and pronotum slightly lighter than metasoma (the propodeum was dissected, so it appears lighter); the legs were removed, but were described as orange. Fore wing infuscation darker from base to under seta M4; remainder of wing lightly infuscated except for a hyaline basal area posteriorly and the very apex. Face striate, frontovertex with punctations more evident. Scutellum a little less than ½ length of mesoscutum. The fore wing setation seems to be similar to other Chartocerus but is slightly obscured. Fore wing 2.7× as long as wide, marginal fringe as long as slightly less than ½ maximum wing width. Mt1 bilobed., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on pages 697-698, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1921. Miscellaneous species of Chalcid-flies from Australia (Hymenoptera, Chalcididae). Insecutor Inscitiae Menstruus. 9: 186 - 191.","Dahms EC. 1986. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: IV. Chalcidoidea species N-Z and Genera with advisory notes plus addenda and corrigenda. Mem Queensland Mus. 22: 319 - 739."]}

Chartocerus australicus Girault, 1913 urn:lsid:zoobank.org:act: BE035C57-4B1D-4D49-97EF-C4154DFCB0B0 EOL taxon ID: 847848 (Figure 2; DOI:10.5281/zenodo.3894025) Girault (1913a, p. 226), as Signiphora australica (original description). Dahms (1983, p. 91): notes on type material. Type material. Holotype male, QM Hy 773 (T773; T4412), under cracked resin and coverslip, along with a specimen of Aphelinoidea howardi Gir. Transcription of labels as in Dahms (1983): ‘From window of a workmen’s quarters. Sugar farm, Nelson [= Gordonvale], N.Q. XII.21.1911 ’, ‘ Signiphora australica Girault, ♂ Type, Hy/773 4412, 4412 ’, and on the reverse of the slide, ‘Queensland Museum. TYPE, Hy/773 S’. Dahms (1983) lists two other slides at the QM, one topotypical, with the date of 4 December 1911 (paratype, male) and the other without locality information, on the same slide as the type of C. australiensis orbiculatus. The original description refers only to the two males from Nelson (‘in xylol-balsam, December 4 and 21, 1911 ’) and indicates Hy 773 as the type. Dahms (1983) also states that T4412 was a duplicate QM accession number for this specimen and had been cancelled. Description. Male. Length (pronotum to apex of epiproct): 0.6 mm. Body brown; metasoma and propodeum darker than head and mesosoma; antennae, genae and clypeus slightly lighter than surrounding areas. Head approximately round in frontal view, antennal scrobes evident, clava 7× as long as wide, 3 anelli (2:2:1), pedicel 2.5× as long as wide, scape 4× as long as wide, clava 2.5× length of scape. Many structures cannot be observed. Pronotum about 8× as wide as long. The other sclerites were crushed; margins are not discernible. The wings are not visible. Median propodeal sclerite seems to be coarsely reticulated like the remainder of the metasoma. Mt1 visibly bilobed., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on pages 685-686, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1913 a. A systematic monograph of the chalcidoid Hymenoptera of the subfamily Signiphorinae. Proc US Natl Mus. 45: 189 - 233. doi: 10.5479 / si. 00963801.1977.189.","Dahms EC. 1983. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: II. Preamble and Chalcidoidea species A-E with advisory notes. Mem Queensland Mus. 21: 1 - 255."]}

Chartocerus beethoveni Girault, 1915 urn:lsid:zoobank.org:act: B94D903A-6262-4516-A95B-FDB8C9D0C032 EOL taxon ID: 847846 (Figure 4; DOI:10.5281/zenodo.3893696) Girault (1915, p. 71), as Signiphora beethoveni (original description). Dahms (1983, p. 119): notes on type material. Type material. Holotype female, QM Hy 2969 (T2969, T4418), QMB, Chindera, New South Wales, 9 May 1914, A.P. Dodd, ‘sweeping forest on sand-ridges near coast’. Partially crushed; medially broken. On the same slide there are parts of Tetracnemoidea secunda (Girault, 1915) (as Arhopoideus secundus Gir.). A separate label indicates ‘ Signiphora beethoveni Gir. | ♀ type | 4418 ’. Described from Chindera, New South Wales, 9 May 1914, A.P. Dodd, ‘sweeping forest on sand-ridges near coast’. It is unambiguous that T2969 refers to the S. beethoveni specimen as it is cited in the original description. Dahms (1983) states that T4418 was a duplicate QM accession number for this specimen and had been cancelled. Description. Female. Length (pronotum to apex of epiproct): 1.15 mm. Head and body appear uniformly light brown; head (frontovertex) slightly darker; fore legs and tarsi of all legs lighter brown to almost transparent. The wings are faded, mostly hyaline, but were described as with an infuscated band medially under the marginal vein. Head with frontovertex finely transversely striate, punctations typical of most Chartocerus not visible. Antennal scrobes well delimited. The morphology of the antenna is very distinctive, similar to that described for males of C. kerrichi: clava wide, ellipsoid, 3× longer than maximum width; 4 anelli (4:2:2:1), pedicel 2× as long as wide, scape 4× as long as wide and about ¾ of length of clava. Pronotum, mesoscutum, scutellum and metanotum transversally striate to finely, faintly reticulate; axillae reticulate, medial area of propodeum reticulate at lateral margins to striate medially, with cells 2–3× wider than long. Mesoscutum about 3.5× length of scutellum. Fore wing 3× as long as wide, marginal fringe slightly longer than ½ of maximum width of wing, 2 setae in submarginal vein, M1 lacking, 5 small setae in basal area of wing, 1 wing in costal cell. Mesotibia 2× as long as protibia; metatibia only slightly longer than protibia, mesofemur with 3 spines, mesobasitarsus about ½ length of mid tibia and mesotibial spur about same length of basitarsus, with 5 teeth. Mt1 bilobed., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on pages 687-689, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1915. Australian Hymenoptera Chalcidoidea - VII. The family Encyrtidae with descriptions of new genera and species. Mem Queensland Mus. 4: 1 - 184.","Dahms EC. 1983. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: II. Preamble and Chalcidoidea species A-E with advisory notes. Mem Queensland Mus. 21: 1 - 255."]}

Chartocerus gratius (Girault, 1932), described as Matritia gratia (Girault 1932) was not examined. Dahms (1984) notes that the syntypes are in Perth (DEAP) and that the type series is a single slide containing 10 damaged syntypes of both sexes. We have not been able to examine this material. Girault’s (1932) original description states ‘From hebes. Wing 1 sans cross-band. Middle tibial spur all pale; hind fringe wing 2 ½ widest. Western Australia, on Dactylopius, L.J. Newman’., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on page 700, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1932. New lower Hymenoptera from Australia and India. Brisbane: Private publication; p. 6.","Dahms EC. 1984. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: III. Chalcidoidea species F-M with advisory notes. Mem Queensland Mus. 21: 579 - 842."]}

Chartocerus delicatus Girault, 1933 urn:lsid:zoobank.org:act: C9E2E3E9-6AD8-41E0-8AE2-4A8B7438EF0E EOL taxon ID: 848109 (Figure 6; DOI:10.5281/zenodo.3893702) Girault (1933, p. 2), as Matricia (sic) delicata (original description). Dahms (1983, p. 216): notes on type material. Type material. Holotype female, QM T8736; partially broken, the head and one antenna removed. Dahms (1983) mentions the slide was cracked and mended with paper, and that it contains the female holotype together with several other insects. The label data reads: ‘ Matritia delicata Girault, Type ♀ || On adhesive bracts of Passaflora [sic], Taringa, 26 November 1928 | Ent. Div. Dep. Ag. & Stk., Qld.’ The original description mentions only ‘Taringa, No. 26, 1928’. Dahms (1983) provides some more information from Girault’s unpublished manuscript: ‘One female found upon the adhesive bracts of Passaflora [= Passiflora] foetida in my home garden was the type.’ Description. Female. Length 0.74 mm. Body light brown; legs and tarsi yellowish or light brown, syntergum and ovipositor sheath seem slightly darker than the rest of the body. Wing infuscation seems to cover the entire wing; it suggests a medial hyaline area, but the infuscation only gets lighter close to the end of the stigmal vein, and then also noticeably infuscated at the apex of the wing, with another small hyaline area on the anterior portion. Head roughly rounded in frontal view; punctate sculpturing. Pronotum, mesoscutum, scutellum and metanotum transversely striate; axillae reticulate, medial area of propodeum reticulate. Mesoscutum with about 14 small setae arranged somewhat symmetrically in posterior half of mesoscutum. Scutellum with 9–10 fine setae in posterior margin of scutellum and 1 on top of each axilla. Protibia about 2/3 of length of mesotibia; mesotibia about 2/3 of length of metatibia; mesobasitarsus less than half of length of mesotibia, mesotibial spur with 5 teeth. Fore wing with 2 setae on submarginal vein, lacking M1, as described above for other Australian species of Chartocerus. Hind wing 6.5× as long as wide, marginal fringe a little less than ½ width of wing. Mt1 bilobed, although faintly visible. Remarks. Girault (1933) mentions it differs from C. australicus by shorter marginal fringe in wings, and metafemur infuscated only apically, which might have been an oversight, since several other species of Chartocerus present such a pattern., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on pages 691-693, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1933. Some beauties inhabitant not of the boudoirs of commerce but of natures bosom - new insects. Brisbane: Private publication; p. 2.","Dahms EC. 1983. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: II. Preamble and Chalcidoidea species A-E with advisory notes. Mem Queensland Mus. 21: 1 - 255."]}

Chartocerus thusanoides Girault, 1915 urn:lsid:zoobank.org:act: 7158060D-5DCC-41FB-8D23-1A4A790FF251 EOL taxon ID: 847837 (Figure 11; DOI:10.5281/zenodo.3893753) Girault (1915, p. 71), as Signiphora thusanoides (original description). Dahms (1986, p. 588): notes on type material. Remarks. Holotype female, QM Hy 2968 (T2968), on a slide in bad condition with two coverslip fragments, labelled ‘TYPE || 4415 || (fading) Signiphora thusanoides Gir. ♀ type || Holotype Signiphora thusanoides, Gir. Det. J. B. Woolley “79”’. The specimen was crushed, with the mesosoma broken in half, but otherwise most structures are observable; the head is broken in several pieces. The other coverslip fragment contains another ♀ but there is no indication of identification. Dahms (1986) states that number T4415 was a duplicate QM accession number for this type and had been cancelled. The number QM Hy 2968 is given in the original description. Description.Female. Length about 1.4 mm. Body mostly brown, legs lighter, with apical tarsomeres of metatibia tan; posterior face of mesotibia yellowish white. The head capsule was completely crushed; some punctations are visible on the fragments of the genae. Scape 4× as long as wide, pedicel a little less than 2× as long as wide, 4 anelli (2:3:4:7). Clava about 4× as long as wide. Pronotum about 5× as wide as long, mesoscutum 2.5× longer than scutellum. Fore wing wide, 2.3× as long as wide. Hind wing about 3× as long as wide, marginal fringe about ¼ of width of mesotibia. Mt1 slightly bilobed, the two lobes very close to each other, giving Mt1 the same trapezoid appearance as propodeum. Ovipositor short, the outer plates not reaching ½ length of the metasoma., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on pages 698-700, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1915. Australian Hymenoptera Chalcidoidea - VII. The family Encyrtidae with descriptions of new genera and species. Mem Queensland Mus. 4: 1 - 184.","Dahms EC. 1986. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: IV. Chalcidoidea species N-Z and Genera with advisory notes plus addenda and corrigenda. Mem Queensland Mus. 22: 319 - 739."]}

Chartocerus australiensis orbiculatus Girault, 1915 urn:lsid:zoobank.org:act: E1B6555A-B559-41F5-BACE-26F20E018452 (Figure 3; DOI:10.5281/zenodo.3893660) Girault (1915, p. 68), as Signiphora australiensis orbiculata (original description). Dahms (1983, p. 92, 111): notes on type material. Type material. Holotype male, QM Hy 2965 (T2965). The label, in Girault’s handwriting, indicates: ‘Type | Hy/2965 | A.A. Girault || Signiphora | australica Gir. || (red label) 4409 | Signiphora | australiensis | orbiculata | Gir. ♀ ’. The description indicates the locality as Gordonvale (Cairns), North Queensland, but this locality information is not provided in the slide labels. Dahms (1983, p. 92) describes the slide which contains the holotype as ‘slide #3’ in his discussion of C. australicus: ‘2 coverslip fragments; the one closest to “ TYPE ” label contains ♀ Signiphora australiensis orbiculata Girault; the other fragment contains a ♀ Signiphora australica, head, wings and some legs separated’. The female indication in the label is a lapsus (holotype is a male). Dahms (1983) states that T 4409 is the QM number for the holotype. Description. Male. Length (pronotum to apex of epiproct): 2.2 mm. Body brown; legs slightly lighter, tarsi of mid leg and hind leg white, and apical tarsomere of hind leg tan. Fore wings infuscated, interrupted by a median hyaline band; the anterior part of the infuscation extending from the base to the stigmal vein is slightly darker than the posterior. Head (crushed in the specimen; some structures not clearly visible), scape wide (approximately 3× as long as wide, pedicel 2× as long as wide, anelli not visible, clava fairly short and wide (4× as long as wide). Mesoscutum about 2× as long as scutellum, with 12 setae: 8 setae in posterior margin of scutellum plus 1 on top of each axilla. Mesotibia ½ length of metatibia, mesobasitarsus about ½ length of mesotibia, mesotibial spur slightly shorter than basitarsus, with 5 teeth. Mesofemur with 3 spines. Fore wings 3× as long as wide, marginal setae about 1/3 of maximum wing width, 1 seta in submarginal vein. Hind wing 4× as long as wide, marginal fringe about ½ width of wing, 1 seta in marginal vein. Mt1 bilobed, with a transverse area separating the two lobes. Remarks. This name was made available at the subspecies level. Girault (1915) defined this subspecies based on differences of the wing infuscation. The distal portion of the wing is faintly infuscated as in C. australiensis australiensis., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on page 686, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1915. Australian Hymenoptera Chalcidoidea - VII. The family Encyrtidae with descriptions of new genera and species. Mem Queensland Mus. 4: 1 - 184.","Dahms EC. 1983. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: II. Preamble and Chalcidoidea species A-E with advisory notes. Mem Queensland Mus. 21: 1 - 255."]}

Chartocerus funeralis Girault, 1913 urn:lsid:zoobank.org:act: 56888B8B-F5B3-47FF-8E1B-02BC10624260 EOL taxon ID: 848103 (Figure 7; DOI:10.5281/zenodo.3893731) Girault (1913a, p. 224), as Signiphora funeralis (original description). Dahms (1984, p. 582, 634; 1986, p. 568): notes on type material. Type material. Holotype female, QM Hy 771 (T4410). Dahms (1984) describes one slide with a coverslip fragment with the holotype of S. funeralis, complete, along with other chalcidoids (identified as Anagrus armatus, Abbella subflavella and the holotype of Pseudogramma fasciatipenne Gir.) (label: ‘From windows of empty dwellings, Herberton, N.Q., XII.28.1911, Hy 771 | 3484, 4410, 3484’), and a second slide (not type material) with a fragment containing a male S. funeralis partially dissected (head, antenna and one leg) (label: ‘ Signiphora funeralis Gir. | Indooroopilly | window | VII.5.1933)’ and a separate coverslip with a specimen of Chartocerus hebes (as M. hebes). Dahms (1984) states that T4410 was a duplicate QM accession number for the type specimen and had been cancelled. T3484 refers to the specimen of Abbella subflavella. Description. Female. Length 0.74 mm. The holotype has faded to a yellowish tone, but it was originally described as being dark like the other species of Chartocerus, with metallic tinge on the head and mesoscutum. Fore wing infuscated except for a small basal area, posteriorly. Head with punctations scattered on face and genae. Scape about 4.5× as long as wide, pedicel 2× as long as wide, ring segments in ratios of 6:4:2:1, resembling C. kerrichi, clava 5× as long as wide. Mesoscutum a little more than 3.5× length of scutellum, with scattered small setae; scutellum with 6 faint setae along posterior margin. Fore wing almost 3× as long as wide, marginal fringe roughly ½ width of the wing. Mesobasitarsus less than ½ length of mesotibia, mesotibial spur about 1/3 of the basitarsus. Mt1 bilobed, the two lobes separated smoothly by a concave curve; propodeal sclerite projecting posteriorly gradually., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on page 693, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1913 a. A systematic monograph of the chalcidoid Hymenoptera of the subfamily Signiphorinae. Proc US Natl Mus. 45: 189 - 233. doi: 10.5479 / si. 00963801.1977.189.","Dahms EC. 1984. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: III. Chalcidoidea species F-M with advisory notes. Mem Queensland Mus. 21: 579 - 842."]}

Noyesaphytis lasallei Polaszek & Woolley sp. n. urn:lsid:zoobank.org:act: 46A980C5-5B13-45EB-AC20-698C4C207A6A Description In addition to the genus-level characters above, the following characters are likely to be of species-level significance if additional species of Noyesaphytis are discovered: Length (female holotype): 1.1 mm (to ovipositor tip). Colour: Head brown, pale areas between the toruli and compound eyes, and around the ocelli. Genae darker brown than remainder of head. Mesoscutum largely dark brown with the following pale: a narrow longitudinal streak from central pronotum to base of scutellum; sutures bordering mesoscutal mid and side lobes and axillae, axillulae, hind margin of scutellum, metanotum, central pronotum. Mesosoma setation pale (Figure 1). Metasoma entirely dark brown. All femora and tibiae largely brown; femora pale for their distal 1/3-1/4; tibiae pale proximally and distally for 1/4–1/5 of their length. Tarsomeres pale, except all distal segments and fore and hind basitarsus brown. Fore wing distinctly infuscate below parastigma, along hind margin of marginal vein and below stigmal vein, the latter forming a faint cloud that reaches almost to the hind margin of the wing. Sculpture: Frons with aciculate sculpture below the stemmaticum, transverse sculpture at the level of the lower compound eyes (Figure 5); between the eyes and toruli the sculpture is diagonally elongate-reticulate. Mid lobe of mesoscutum and scutellum with fine aciculate sculpture; side lobes and axillae reticulate; axillulae each with 4–5 longitudinal striae. Propodeum reticulate centrally and at sides. Metasoma with longitudinally reticulate sculpture laterally. Additional characters Submarginal vein with five robust setae directed distally, i.e. towards the wing apex; ‘costal cell’ with a complete line of fine setae medially, and a row of five stronger setae in its distal upper corner. Mid tibial spur 0.7x corresponding basitarsus. Ovipositor projecting beyond metasoma; length 2.4x mid tibia. 2nd valvifers 2.9x 3rd valvulae. Male (Figures 10–13): Length: 1.0 mm Colour: Largely as in female, with the following differences: Pale longitudinal streak absent from mid lobe of mesoscutum; metanotum and propodeum uniformly brown; legs more evenly brown than in female, sharply contrasting pale and brown areas absent. Wing less strongly infuscated. Sculpture: Largely as in female, but dorsal mesosomal sculpture tending more towards reticulate rather than aciculate. Additional characters Antennal formula 1,1,4,1,1. Four anelli present, followed by a single funicle and clava (Figures 10–12). Mid tibial spur equal in length to corresponding basitarsus. Genitalia as in Figure 13, phallobase elongate, with digiti and claspers present. Material examined Holotype female: MADAGASCAR: Tuléar (currently Toliara), Berenty 24º59 ʹ 53”S 46º19 ʹ 53”E v.1983 (JS Noyes) BM 1983–201; NHMUK010370379; DNA1157. Holotype dissected and slide-mounted in Canada balsam. Paratypes 2 males: MADAGASCAR: Tuléar (currently Toliara), Bereboka 19º59 ʹ 53”S 44º35 ʹ 59”E 18–23.v.1983 (JS Noyes) BM 1983–201 (NHMUK); DNA1158-9. Holotype and paratype specimens are deposited in NHM, London. Etymology The genus name Noyesaphytis is derived from the collector’s family name (Noyes) and the generic name Aphytis. The use of various prefixes before - aphytis for genera considered to be related to Aphytis has several precedents; e.g. Punkaphytis Kim and Heraty (2012); Wallaceaphytis Polaszek & Fusu, in Polaszek et al. (2013). The specific epithet lasallei is based on, and in memory of,our friend and colleague,Dr John La Salle.

Chartocerus corvinus Girault, 1913 urn:lsid:zoobank.org:act: F8E2494C-599C-4BC9-BCF1-08255B0A2082 EOL taxon ID: 847843 (Figure 5; DOI:10.5281/zenodo.3893699) Girault (1913a, p. 225), as Signiphora corvina (original description). Dahms (1983, p. 198; 1984, p. 698, 701; 1986, p. 409, 568): notes on type material. Type material. Holotype female, QM Hy 772, same slide as type of Gonatocerus huxleyi Girault and other specimens identified as Signiphora australiensis, Abbella subflava, Ufens piceipes and Aphelinoidea howardii, ‘from a window in a granary and barn on a wheat farm at Roma, Queensland, 6 October 1911 ’. The accession number Hy 772 is not indicated on the slide; number 4407 is indicated on a red label. According to Dahms (1983), this female is in fact the holotype: ‘QM: Slide – 1 cracked, complete coverslip (partly missing over 1 specimen) containing the Holotype = (head separated) of Signiphora corvina and other specimens as per labels “ Gonatocerus huxleyi Girault, ♀ Type 1040. From windows of a granary, Roma, Q., 6 October 1911 AAG 3691, 3691” [...]’. An arrow was added by JBW to indicate the holotype. Dahms (1983) states that number T4407 was a duplicate QM accession number for this type and had been cancelled. T3691 refers to the Gonatocerus specimen. Description. Female. Length about 1.2 mm. Body dark, femora and tibia slightly lighter, protibia especially light distally. Tarsi of all legs lighter than the rest of the legs. The basal infuscation of the fore wing extends in the anterior-basal portion of the wing to the stigmal vein. Head round in frontal view, genae reticulate, punctations lateral to antennal scrobes; interantennal area seems lighter than clypeus and genae. Eyes relatively small, about 1/3 height of head. Antennal scape about 5× as long as wide, pedicel and ring segments cannot be observed, club about 2.6× as long as wide. Mesoscutum transversally striate; scutellum, metanotum and basal part of medial propodeal sclerite striate to reticulate towards margins. Mesoscutum at least 2× length of scutellum. It is not possible to count the setae on the mesosomal terga. Fore wing very broad, at the broadest part about ½ length of wing. The wing vein reaches about half of the distance to the apex. Marginal fringe very short, not more than 1/8 of wing width, 2 setae on submarginal vein, M1 missing (M6 is broken in this type but the socket is visible). Hind wing also wide towards the apex, about 3.5× as long as wide, marginal fringe 1/5 of wing width, with discal seta. Mesobasitarsus less than ½ length of mesotibia; mesotibial spur slightly shorter than basitarsus and with 4 teeth. Mesofemur with 2 visible spines., Published as part of Molin, Ana Dal & Woolley, James B., 2020, Notes on types of Australian Chartocerus Motschulsky (Hymenoptera: Signiphoridae), pp. 681-702 in Journal of Natural History 54 (9) on page 689, DOI: 10.1080/00222933.2020.1785573, http://zenodo.org/record/4290435, {"references":["Girault AA. 1913 a. A systematic monograph of the chalcidoid Hymenoptera of the subfamily Signiphorinae. Proc US Natl Mus. 45: 189 - 233. doi: 10.5479 / si. 00963801.1977.189.","Dahms EC. 1983. A checklist of the types of Australian Hymenoptera described by Alexandre Arsene Girault: II. Preamble and Chalcidoidea species A-E with advisory notes. Mem Queensland Mus. 21: 1 - 255."]}