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† Pherodactylus rectanguli (Xu, Fang & Wang, 2020) n. comb. Figs. 1–6 = † Chunxiania fania Xu, Wang & Fang, 2022 n. syn. = † Tresdigitus gracilis Jiang, Xu, Jarzembowski & Xiao, 2022 n. syn. General description. Male (based on specimens ECNU-AM-0019 and ECUN-AM-0010, Figs. 1–3). Smallsized (8.07 mm.). Predominantly yellowish coloration with grayish and black spots. Cephalic capsule with dorsal surface (from vertex to fastigium) and upper area of frons with dark brown to black stripes; area surrounding ocelli, mandibles, lower margin of genae, and frons ocher; ocelli yellow; eyes greyish. Frons with a yellow inverted “Y”, * for the data may be doubtful due to different measurement method. starting from vertex, dividing at fastigium, and branching to the inner margin of each lateral ocellus. Pronotal disc greyish brown, with black corner rims and delimiting pronotal disc from lateral lobes; pronotal disk with two distinct large dark “eyespots”. Lateral lobes of pronotum with upper half grayish brown and lower half yellowish. All legs are yellowish with black or grayish spots, irregularly scattered, most noticeable on femora. Tegminae translucent, with veins outlined in light yellow. Abdomen yellowish covered by various black or grayish spots. Body covered with short or mid-sized fine hairs. Head as wide as pronotum, rounded and smooth; almost as wide as high in frontal view, with epistomal suture little concave. Vertex rounded. Eyes ovoid, not protruding. Lateral ocelli circular, central ocellus absent. Eyes and antennal pits located very low on face, close to epistomal suture, almost at the same level. Fastigium wide, almost four times as wide as scape. Maxillary palpi with the 4 th and 5 th longer, apical segment of labial palpi flattened. Thorax. Pronotum longer than wide, covered by several and mid-sized hairs; anterior margin concave and broader than posterior margin, with long bristles; lateral lobules rectangular, wider than high and with a rounded lateral edge, ventral margin almost straight. Wings. Tegmina short and ovoid, reaching to the second abdominal tergite. Mirror in oval shape, longer than wide, with a curved dividing vein; harp with three dividing veins; apical field reduced; lateral field two longitudinal veins from base to apex, and ten transversal veins born from the second longitudinal vein to costal margin. Hind wings no visible. Legs. Coxa and trochanter not elongated. Foreleg robust with long setae, large and oval tympana in both sides, on middle of fore tibiae; three movable dactyls on apical part of fore tibiae, first tarsomere almost as long as fore tibiae, other two tarsomere short. Mid leg slender, coxa robust, with a process partly covering trochanter; femur slightly slenderer than fore femur; tibia slightly inflated, shorter than femur, armed at apex with three inner spurs and two outer spurs. Hind leg elongated, femur strong, tibia with several small spines on dorsal side at apex. Three internal apical spurs long, and two external apical spurs short. The first and third tarsomere long and the second one short. Abdomen cylindrical; tergites with few short hairs; epiproctus subtriangular, apex rounded; subgenital plate short, wider than long, apex rounded; cerci almost as long as the two-thirds of hind tibia, covered by abundant medium-sized hairs. Female (emended description; based on specimens ECNU-AM-0002 and ECUN-AM-0024, Figs. 4, 5). Similar to the males in shape and size, differing by the following characters: the color tones are darker than the males, but keep the same pattern. Tegmina absent. Abdomen with hairs and several long bristles on dorsal side. Epiproct triangular, with rounded posterior margin. Ovipositor almost straight, apex of valves sharp, pointed, and lanceolate in dorsal view. Subgenital plate trapezoidal, apex moderately protruding. Male nymph (based on specimen ECNU-AM-0045, Fig. 6). Similar to the adult males, but no developed wings, with a more fragile and slender appearance. It differs from the adults, because the tympana are smaller in diameter than the other adult specimens examined. The small tympana in immature specimens can happen. This structure increases its size in adults. Measurements (mm). Adult male ECNU-AM-0019 (Fig. 1, 2): BL: 8.07; HL: 1.07; PL: 1.57; PW: 1.43; FWL: 2.18; FFL: 1.61; FTL: 1.04; MFL: 1.41; MTL: 1.09; HFL: 3.15; HTL: 1.82; AL: 5.14; CL: 2.26. Adult male ECNU-AM-0010 (Fig. 3): PL: 1.47; PW: 1.33; FWL: 2.05; FFL: 0.98; FTL: 1.06; Body broken, uncertain for else. Adult female ECNU-AM-0002 (Fig. 4): BL: 8.20; HL: 1.21; PL: 1.69 PW: 1.71 FFL: 1.56; FTL: 0.97; MFL: 1.91; MTL: not visible; HFL: 3.63; HTL: 2.54; AL: 4.63, OL: 1.82. Adult female ECNU-AM-0024 (Fig. 5): BL: 5.24; HL: 1.06; PL: 1.60; PW: uncertain; FFL: 1.45; FTL: 0.95; MTL: 1.42, femur and tarsus incomplete; HFL: 3.34, HTL: 2.56; AL: 3.82, CL: 2.39, OL: 1.72. Nymph ECNU-AM-0045 (Fig. 6): BL: 5.22; HL: 0.87; PL: 1.33, PW: uncertain; FFL: 1.40; FTL: 0.83; MFL: 0.97, MTL: 0.82; HFL: 2.49; HTL: 1.71; AL: 3.25; CL: 1.67. Remarks. Synonymy and new combination at the species level are justified as follows: 1) † P. micromorphus Poinar et al. 2020 is a subadult female specimen, which looks different in the shape of the pronotum in contrast to the holotype of † P. rectanguli n. comb. (a subadult male). However, when the photos of specimens are taken from different angles, an optical effect will be caused, which confuses us to consider them as different shapes. This can be happened in the additional specimens of † P. rectanguli n. comb. studied here. The fore tibia does not vary in both specimens. Only in the holotype of P. rectangulari n. comb., a bending of the ventral edge is observed, but this can happen due to dehydration during fossilization. However, we prefer to keep this species as valid, since the subadult female holotype specimen of † P. micromorphus had a wing pad, which is different from the adult females of † P. rectangulari n. comb. 2) † C. fania n. syn. is an adult male specimen. In the images of the original description, its diagnostic characters coincide with † P. rectanguli n. comb. in terms of the shape of fore tibia, apical spurs organization and shape, and location of the spurs of hind tibia. The hind tibiae are thin and moderately elongated, and the shape of pronotum and its spots coincide in these specimens. Unfortunately, the venation of † C. fania n. syn. is not observed due to preservation.Although the subgenital plate of † C. fania n. syn. dehydrated in fossilization process and coiled dorsally, it still could be observed in a triangular shape, which is similar to those of field crickets in a semicircle shape. 3) † Tresdigitus gracilis n. syn. is a nymph of indefinite sex (possibly female in penultimate or ultimate instar), not as developed as the two “species” previously synonymized. In the key presented in the original description by Jiang et al., 2022, the arrangement of head and the shape of pronotum are some of the characters proposed by the authors to distinguish the species from † P. rectanguli n. comb. The disposition of the head can vary in the way that insects die, and thus remains when fossilized. In contrast to † P. rectanguli n. comb., its head is in a normal position, locating mouthparts down (hypognathous/ hypognathous mandible). Other characters that stand out in † T. gracilis n. syn. is the moderately expanded fore tibia, and the small tympanum. However, it is common for crickets that they are different in these characters in their early stages. Through molting and growth, the tympanum becomes larger, and the tibiae attains the shape final in the last stages of insect development (Table 2; Ball and Cowan, 1978). In this study, we provide well-preserved males, females, and nymphal specimens, in which most of the morphological characters can be studied. It is applicable to verify the similarities and differences with the other recently described fossil taxa of “Mole Crickets” with those specimens. In terms of coloration, it was feasible to show that the coloration of † P. micromorphus is similar to those of the two males studied here. † P. rectanguli n. comb. has the same color pattern, but its image was darker in the original description (possibly due to a lack of brightness or light from the photographic record). For † C. fania n. syn., the same issue happened as mentioned above. All the specimens in the different stages of development have similar coloration, with two distinct large dark “eyespots” of similar shape; lower half of the anterior lobe of the yellow pronotum; shape and location of apical spurs of fore tibia; hind tibia serrulate with small spines; apex with conspicuous slender spines. The number of dorsal spines on legs is not reliable classification characters due to its intraspecific variation, which may be different from one leg to the other leg on one specimen (Cadena-Castañeda et al. 2022b). We consider † Pherodactylus micromorphus and † P. rectanguli n. comb. as valid genus and species. Both genera were described in 2020, and the original description of † P. micromorphus is available on March of 2020, while † Tresdigitus rectanguli is published on August of 2020. It is worth noting that Poinar et al. (2020) proposed a more reasonable taxonomic placement of its genus and species than the other authors, consisting with our results discussed and analyzed here. † Burmagryllotalpa longa is proposed to move under Gryllidae: Gryllinae: Sclerogryllini: Pherodactylina n. subtr. Although this species shares some morphologies with † Pherodactylus species, † B. longa completely lacks tympanum, and its pronotum is dorsally curved without spots on pronotal disc that characterize † Pherodactylus species. Thus, we consider † B. longa as valid.

† Pherodactylus rectanguli (Xu, Fang & Wang, 2020) n. comb. Figs. 1–6 = † Chunxiania fania Xu, Wang & Fang, 2022 n. syn. = † Tresdigitus gracilis Jiang, Xu, Jarzembowski & Xiao, 2022 n. syn. General description. Male (based on specimens ECNU-AM-0019 and ECUN-AM-0010, Figs. 1–3). Smallsized (8.07 mm.). Predominantly yellowish coloration with grayish and black spots. Cephalic capsule with dorsal surface (from vertex to fastigium) and upper area of frons with dark brown to black stripes; area surrounding ocelli, mandibles, lower margin of genae, and frons ocher; ocelli yellow; eyes greyish. Frons with a yellow inverted “Y”, * for the data may be doubtful due to different measurement method. starting from vertex, dividing at fastigium, and branching to the inner margin of each lateral ocellus. Pronotal disc greyish brown, with black corner rims and delimiting pronotal disc from lateral lobes; pronotal disk with two distinct large dark “eyespots”. Lateral lobes of pronotum with upper half grayish brown and lower half yellowish. All legs are yellowish with black or grayish spots, irregularly scattered, most noticeable on femora. Tegminae translucent, with veins outlined in light yellow. Abdomen yellowish covered by various black or grayish spots. Body covered with short or mid-sized fine hairs. Head as wide as pronotum, rounded and smooth; almost as wide as high in frontal view, with epistomal suture little concave. Vertex rounded. Eyes ovoid, not protruding. Lateral ocelli circular, central ocellus absent. Eyes and antennal pits located very low on face, close to epistomal suture, almost at the same level. Fastigium wide, almost four times as wide as scape. Maxillary palpi with the 4 th and 5 th longer, apical segment of labial palpi flattened. Thorax. Pronotum longer than wide, covered by several and mid-sized hairs; anterior margin concave and broader than posterior margin, with long bristles; lateral lobules rectangular, wider than high and with a rounded lateral edge, ventral margin almost straight. Wings. Tegmina short and ovoid, reaching to the second abdominal tergite. Mirror in oval shape, longer than wide, with a curved dividing vein; harp with three dividing veins; apical field reduced; lateral field two longitudinal veins from base to apex, and ten transversal veins born from the second longitudinal vein to costal margin. Hind wings no visible. Legs. Coxa and trochanter not elongated. Foreleg robust with long setae, large and oval tympana in both sides, on middle of fore tibiae; three movable dactyls on apical part of fore tibiae, first tarsomere almost as long as fore tibiae, other two tarsomere short. Mid leg slender, coxa robust, with a process partly covering trochanter; femur slightly slenderer than fore femur; tibia slightly inflated, shorter than femur, armed at apex with three inner spurs and two outer spurs. Hind leg elongated, femur strong, tibia with several small spines on dorsal side at apex. Three internal apical spurs long, and two external apical spurs short. The first and third tarsomere long and the second one short. Abdomen cylindrical; tergites with few short hairs; epiproctus subtriangular, apex rounded; subgenital plate short, wider than long, apex rounded; cerci almost as long as the two-thirds of hind tibia, covered by abundant medium-sized hairs. Female (emended description; based on specimens ECNU-AM-0002 and ECUN-AM-0024, Figs. 4, 5). Similar to the males in shape and size, differing by the following characters: the color tones are darker than the males, but keep the same pattern. Tegmina absent. Abdomen with hairs and several long bristles on dorsal side. Epiproct triangular, with rounded posterior margin. Ovipositor almost straight, apex of valves sharp, pointed, and lanceolate in dorsal view. Subgenital plate trapezoidal, apex moderately protruding. Male nymph (based on specimen ECNU-AM-0045, Fig. 6). Similar to the adult males, but no developed wings, with a more fragile and slender appearance. It differs from the adults, because the tympana are smaller in diameter than the other adult specimens examined. The small tympana in immature specimens can happen. This structure increases its size in adults. Measurements (mm). Adult male ECNU-AM-0019 (Fig. 1, 2): BL: 8.07; HL: 1.07; PL: 1.57; PW: 1.43; FWL: 2.18; FFL: 1.61; FTL: 1.04; MFL: 1.41; MTL: 1.09; HFL: 3.15; HTL: 1.82; AL: 5.14; CL: 2.26. Adult male ECNU-AM-0010 (Fig. 3): PL: 1.47; PW: 1.33; FWL: 2.05; FFL: 0.98; FTL: 1.06; Body broken, uncertain for else. Adult female ECNU-AM-0002 (Fig. 4): BL: 8.20; HL: 1.21; PL: 1.69 PW: 1.71 FFL: 1.56; FTL: 0.97; MFL: 1.91; MTL: not visible; HFL: 3.63; HTL: 2.54; AL: 4.63, OL: 1.82. Adult female ECNU-AM-0024 (Fig. 5): BL: 5.24; HL: 1.06; PL: 1.60; PW: uncertain; FFL: 1.45; FTL: 0.95; MTL: 1.42, femur and tarsus incomplete; HFL: 3.34, HTL: 2.56; AL: 3.82, CL: 2.39, OL: 1.72. Nymph ECNU-AM-0045 (Fig. 6): BL: 5.22; HL: 0.87; PL: 1.33, PW: uncertain; FFL: 1.40; FTL: 0.83; MFL: 0.97, MTL: 0.82; HFL: 2.49; HTL: 1.71; AL: 3.25; CL: 1.67. Remarks. Synonymy and new combination at the species level are justified as follows: 1) † P. micromorphus Poinar et al. 2020 is a subadult female specimen, which looks different in the shape of the pronotum in contrast to the holotype of † P. rectanguli n. comb. (a subadult male). However, when the photos of specimens are taken from different angles, an optical effect will be caused, which confuses us to consider them as different shapes. This can be happened in the additional specimens of † P. rectanguli n. comb. studied here. The fore tibia does not vary in both specimens. Only in the holotype of P. rectangulari n. comb., a bending of the ventral edge is observed, but this can happen due to dehydration during fossilization. However, we prefer to keep this species as valid, since the subadult female holotype specimen of † P. micromorphus had a wing pad, which is different from the adult females of † P. rectangulari n. comb. 2) † C. fania n. syn. is an adult male specimen. In the images of the original description, its diagnostic characters coincide with † P. rectanguli n. comb. in terms of the shape of fore tibia, apical spurs organization and shape, and location of the spurs of hind tibia. The hind tibiae are thin and moderately elongated, and the shape of pronotum and its spots coincide in these specimens. Unfortunately, the venation of † C. fania n. syn. is not observed due to preservation.Although the subgenital plate of † C. fania n. syn. dehydrated in fossilization process and coiled dorsally, it still could be observed in a triangular shape, which is similar to those of field crickets in a semicircle shape. 3) † Tresdigitus gracilis n. syn. is a nymph of indefinite sex (possibly female in penultimate or ultimate instar), not as developed as the two “species” previously synonymized. In the key presented in the original description by Jiang et al., 2022, the arrangement of head and the shape of pronotum are some of the characters proposed by the authors to distinguish the species from † P. rectanguli n. comb. The disposition of the head can vary in the way that insects die, and thus remains when fossilized. In contrast to † P. rectanguli n. comb., its head is in a normal position, locating mouthparts down (hypognathous/ hypognathous mandible). Other characters that stand out in † T. gracilis n. syn. is the moderately expanded fore tibia, and the small tympanum. However, it is common for crickets that they are different in these characters in their early stages. Through molting and growth, the tympanum becomes larger, and the tibiae attains the shape final in the last stages of insect development (Table 2; Ball and Cowan, 1978). In this study, we provide well-preserved males, females, and nymphal specimens, in which most of the morphological characters can be studied. It is applicable to verify the similarities and differences with the other recently described fossil taxa of “Mole Crickets” with those specimens. In terms of coloration, it was feasible to show that the coloration of † P. micromorphus is similar to those of the two males studied here. † P. rectanguli n. comb. has the same color pattern, but its image was darker in the original description (possibly due to a lack of brightness or light from the photographic record). For † C. fania n. syn., the same issue happened as mentioned above. All the specimens in the different stages of development have similar coloration, with two distinct large dark “eyespots” of similar shape; lower half of the anterior lobe of the yellow pronotum; shape and location of apical spurs of fore tibia; hind tibia serrulate with small spines; apex with conspicuous slender spines. The number of dorsal spines on legs is not reliable classification characters due to its intraspecific variation, which may be different from one leg to the other leg on one specimen (Cadena-Castañeda et al. 2022b). We consider † Pherodactylus micromorphus and † P. rectanguli n. comb. as valid genus and species. Both genera were described in 2020, and the original description of † P. micromorphus is available on March of 2020, while † Tresdigitus rectanguli is published on August of 2020. It is worth noting that Poinar et al. (2020) proposed a more reasonable taxonomic placement of its genus and species than the other authors, consisting with our results discussed and analyzed here. † Burmagryllotalpa longa is proposed to move under Gryllidae: Gryllinae: Sclerogryllini: Pherodactylina n. subtr. Although this species shares some morphologies with † Pherodactylus species, † B. longa completely lacks tympanum, and its pronotum is dorsally curved without spots on pronotal disc that characterize † Pherodactylus species. Thus, we consider † B. longa as valid., Published as part of Cadena-Castañeda, Oscar J., Liu, Yi-Jiao, Yu, Zhe-Yuan, Hu, Tian-Hao, Wu, Shi-Yang & He, Zhu-Qing, 2023, Are the recently described fossil Mole Crickets of Myanmar amber real gryllotalpids? (Orthoptera: Gryllotalpidae & Gryllidae), pp. 48-64 in Zootaxa 5311 (1) on pages 52-59, DOI: 10.11646/zootaxa.5311.1.2, http://zenodo.org/record/8090321, {"references":["Wang, H., Lei, X., Zhang, G., Xu, C., Fang, Y. & Zhang, H. (2020) The earliest Gryllotalpinae (Insecta, Orthoptera, Gryllotalpidae) from mid-Cretaceous Burmese amber. Cretaceous Research, 107, 104292. https: // doi. org / 10.1016 / j. cretres. 2019.104292","Xu, C., Wang, H., Fang, Y., Jarzembowski, E. A. & Zhuo, D. (2022) Chunxiania fania: a new genus and species of mole cricket (Orthoptera: Ensifera: Gryllotalpidae) from mid-Cretaceous Kachin amber. Cretaceous Research, 134, 105 - 159. https: // doi. org / 10.1016 / j. cretres. 2022.105159","Jiang, X., Xu, C., Jarzembowski, E. A. & Xiao, C. (2022) A peculiar species of mole cricket (Orthoptera: Gryllotalpidae) from mid-Cretaceous Kachin amber. Cretaceous Research, 139, 1 - 6. https: // doi. org / 10.1016 / j. cretres. 2022.105273","Xu, C., Fang, Y. & Wang, H. (2020) A new mole cricket (Orthoptera: Gryllotalpidae) from mid-Cretaceous Burmese amber. Cretaceous Research, 112, 104428. https: // doi. org / 10.1016 / j. cretres. 2020.104428","Poinar Jr., G., Su, Y. N. & Brown, A. E. (2020) A new genus of crickets (Orthoptera: Gryllidae) in Mid-Cretaceous Myanmar Amber. Biosis: Biological Systems, 1 (1), 33 - 38. https: // doi. org / 10.37819 / biosis. 001.01.0049","Ball, E. E. & Cowan, N. (1978) Ultrastructural study of the development of the auditory tympana in the cricket Teleogryllus commodus (Walker). Journal of Embryology and Experimental Morphology, 46, 75 - 87. https: // doi. org / 10.1242 / dev. 46.1.75","Cadena-Castaneda, O. J., Quintana-Arias, R. F., Trujillo, D. M., Prias, J. P. & Castellanos-Morales, C. A. (2022 b) Studies on Neotropical crickets: Aclodes paz n. sp. a new phalangopsid cricket (Orthoptera: Phalangopsidae) from the Santander caves, Colombia. Zootaxa, 5141 (6), 568 - 580. https: // doi. org / 10.11646 / zootaxa. 5141.6.3"]}

† Pherodactylus Poinar, Su & Brown, 2020 = † Tresdigitus Xu, Fang & Wang, 2020 n. syn. = † Chunxiania Xu, Wang & Fang, 2022 n. syn. Diagnosis. Head round, pronotum elongate, longer than the wide, middle of pronotal disk with two distinct large dark “eyespots”, foreleg robust, with three apical spurs arranged on the inner side of fore tibia. Included species. † Pherodactylus micromorphus Poinar, Su & Brown, 2020 (type species by monotypy and original designation) and † Pherodactylus rectanguli (Xu, Fang & Wang, 2020) n. comb. Remarks. We treated genera † Tresdigitus n. syn. and Chunxiania n. syn. as synonyms of genus Pherodactylus Poinar, Su & Brown, 2020. We propose the genera and their species as synonyms or relocated: genera † Tresdigitus n. syn. and Chunxiania n. syn. as synonyms of genus Pherodactylus Poinar, Su & Brown, 2020, and † Tresdigitus gracilis Jiang, Xu, Jarzembowski & Xiao, 2022 n. syn. and † Chunxiania fania n. syn. as synonyms of † Pherodactylus rectanguli n. comb. The type specimen of each species represents different stages of the same species (Table 2), and they are all from the same locality. Besides, their morphological characters coincide with the diagnostic characters of † Pherodactylus. For additional taxonomic justifications, see † P. rectanguli n. comb. remarks., Published as part of Cadena-Castañeda, Oscar J., Liu, Yi-Jiao, Yu, Zhe-Yuan, Hu, Tian-Hao, Wu, Shi-Yang & He, Zhu-Qing, 2023, Are the recently described fossil Mole Crickets of Myanmar amber real gryllotalpids? (Orthoptera: Gryllotalpidae & Gryllidae), pp. 48-64 in Zootaxa 5311 (1) on page 52, DOI: 10.11646/zootaxa.5311.1.2, http://zenodo.org/record/8090321, {"references":["Poinar Jr., G., Su, Y. N. & Brown, A. E. (2020) A new genus of crickets (Orthoptera: Gryllidae) in Mid-Cretaceous Myanmar Amber. Biosis: Biological Systems, 1 (1), 33 - 38. https: // doi. org / 10.37819 / biosis. 001.01.0049","Wang, H., Lei, X., Zhang, G., Xu, C., Fang, Y. & Zhang, H. (2020) The earliest Gryllotalpinae (Insecta, Orthoptera, Gryllotalpidae) from mid-Cretaceous Burmese amber. Cretaceous Research, 107, 104292. https: // doi. org / 10.1016 / j. cretres. 2019.104292","Xu, C., Wang, H., Fang, Y., Jarzembowski, E. A. & Zhuo, D. (2022) Chunxiania fania: a new genus and species of mole cricket (Orthoptera: Ensifera: Gryllotalpidae) from mid-Cretaceous Kachin amber. Cretaceous Research, 134, 105 - 159. https: // doi. org / 10.1016 / j. cretres. 2022.105159","Jiang, X., Xu, C., Jarzembowski, E. A. & Xiao, C. (2022) A peculiar species of mole cricket (Orthoptera: Gryllotalpidae) from mid-Cretaceous Kachin amber. Cretaceous Research, 139, 1 - 6. https: // doi. org / 10.1016 / j. cretres. 2022.105273"]}

Cadena-Castañeda, Oscar J., Liu, Yi-Jiao, Yu, Zhe-Yuan, Hu, Tian-Hao, Wu, Shi-Yang, He, Zhu-Qing (2023): Are the recently described fossil Mole Crickets of Myanmar amber real gryllotalpids? (Orthoptera: Gryllotalpidae & Gryllidae). Zootaxa 5311 (1): 48-64, DOI: 10.11646/zootaxa.5311.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5311.1.2

Tribe Sclerogryllini Gorochov, 1985 Remarks. This tribe was described by Saussure (1877) as “Légion des Scléropterites”, including Acanthoplistus Saussure, 1877, and Scleropterus Haan, 1844. Subsequently, this taxon was treated as a subfamily by Shiraki (1930), and this status was followed by Chopard (1934, 1936) and Desutter (1987). Gorochov (1985) proposed a new name Sclerogryllus for Scleropterus Haan, 1844, which has been used for Scleropterus Schoenherr, 1825 (Coleoptera). At the same time, Gorochov also established the tribal status as Sclerogryllini, with Sclerogryllus as the type genus under Gryllinae. Otte (1994) argued that according to the nomenclatural change proposed by Gorochov, the subfamily name should be changed to Sclerogryllinae. Otte used the name Sclerogryllinae for the first time, and this was followed by Storozhenko et al. (2015). Recently, Yu et al. (2022) published a complete mitochondrial genome of cricket Sclerogryllus punctatus. By phylogenetic analysis, they verified that Sclerogryllus was grouped within the Gryllinae clade. Therefore, Sclerogryllinae should be treated as a synonym of Gryllinae, and Gorochov’s tribe Sclerogryllini under Gryllinae is recommended rather than a different subfamily from field crickets. Sclerogryllini is a monogeneric tribe, with five species distributed in Asia. Rhabdotogryllus caraboides Chopard, 1954 from Guinea (Africa), is only known from its holotype female, and no additional records have been reported since its description. R. caraboides is superficially similar to Sclerogryllus females, but possessing more conspicuous dorsal spurs on the hind tibia. Thus, the homology of Rhabdotogryllus and Sclerogryllus is still a question, and Sclerogryllini probably only occurs in Asia. Here, we propose a new fossil subtribe that more closely resembles extant Sclerogryllini species in morphology than any other tribal taxon of field crickets. Additionally, a key to separate the taxa of the tribe is provided., Published as part of Cadena-Castañeda, Oscar J., Liu, Yi-Jiao, Yu, Zhe-Yuan, Hu, Tian-Hao, Wu, Shi-Yang & He, Zhu-Qing, 2023, Are the recently described fossil Mole Crickets of Myanmar amber real gryllotalpids? (Orthoptera: Gryllotalpidae & Gryllidae), pp. 48-64 in Zootaxa 5311 (1) on page 51, DOI: 10.11646/zootaxa.5311.1.2, http://zenodo.org/record/8090321, {"references":["Gorochov, A. V. (1985) On the Orthoptera subfamily of Gryllinae (Orthoptera, Gryllidae) from eastern Indochina. In: Medvedev, L. N. (Ed.), Fauna iekologiya nasekomykh Vetnama [The fauna and ecology of insects of Vietnam], 1985, pp. 9 - 17.","Saussure, H. (1877) Melanges orthopterologiques V. fascicule Gryllides. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 25 (1), 169 - 504.","Shiraki, T. (1930) Orthoptera of the Japanese Empire. Part I. (Gryllotalpidae and Gryllidae). Insecta Matsumurana, 4, 181 - 252.","Chopard, L. (1934) Catalogues raisonnes de la faune entomologique du Congo belge. Orthopteres - Gryllides. Annales du Musee du Congo belge Tervueren, Belgique, Zoologie, 3 (2), 4, 1 - 88.","Chopard, L. (1936) The Tridactylidae and Gryllidae of Ceylon. Ceylon Journal of Science (Biological Science), 20, 9 - 87.","Desutter, L. (1987) Structure et evolution du complexe phallique de Gryllidea (Orthopteres) et classification des genres neotropicauxde Grylloidea. Premiere partie. Annales de la Societe Entomologique de France, New Series, 23 (3), 213 - 239.","Otte, D. (1994) Crickets (Grylloidea). Orthoptera Species File. Vol. 1. Orthopterist' Society, Philadelphia, Pennsylvania, iv + 120 pp.","Storozhenko, S. Y., Kim, T. W. & Jeon, M. J. (2015) Monograph of Korean Orthoptera. National Institute of Biological Resources, Incheon, 377 pp.","Yu, Z., Xie, H., Liu, Y., Li, K. & He, Z. (2022) The complete mitochondrial genome of cricket Sclerogryllus punctatus (Orthoptera: Gryllidae) and phylogenetic analysis. Journal of Asia-Pacific Entomology, 25 (3), 1 - 9. https: // doi. org / 10.1016 / j. aspen. 2022.101933"]}

† Pherodactylus Poinar, Su & Brown, 2020 = † Tresdigitus Xu, Fang & Wang, 2020 n. syn. = † Chunxiania Xu, Wang & Fang, 2022 n. syn. Diagnosis. Head round, pronotum elongate, longer than the wide, middle of pronotal disk with two distinct large dark “eyespots”, foreleg robust, with three apical spurs arranged on the inner side of fore tibia. Included species. † Pherodactylus micromorphus Poinar, Su & Brown, 2020 (type species by monotypy and original designation) and † Pherodactylus rectanguli (Xu, Fang & Wang, 2020) n. comb. Remarks. We treated genera † Tresdigitus n. syn. and Chunxiania n. syn. as synonyms of genus Pherodactylus Poinar, Su & Brown, 2020. We propose the genera and their species as synonyms or relocated: genera † Tresdigitus n. syn. and Chunxiania n. syn. as synonyms of genus Pherodactylus Poinar, Su & Brown, 2020, and † Tresdigitus gracilis Jiang, Xu, Jarzembowski & Xiao, 2022 n. syn. and † Chunxiania fania n. syn. as synonyms of † Pherodactylus rectanguli n. comb. The type specimen of each species represents different stages of the same species (Table 2), and they are all from the same locality. Besides, their morphological characters coincide with the diagnostic characters of † Pherodactylus. For additional taxonomic justifications, see † P. rectanguli n. comb. remarks.

Fodigryllus bisetus He sp. nov. GGGG (Figs. 2–5) Materials. Holotype: male, CHINA, Yunnan, Honghe, Lüchun, Huanglian Mountain National Nature Reserve (N 22.88 E 102.32), 12-vii-2021, coll. He Zhu-Qing. Paratypes: 3 males, same data as for holotype. (depository: East China Normal University, China) Description. Male. Head: round and almost as wide as the pronotum (Fig. 3A); occiput round and convex, frontal rostrum almost twice wider than the antennal scape; eyes large but not convex, three ocelli disposed as triangle, lateral ocelli large and vertically oval, median ocelli smaller (Fig. 3B); maxillary palps with distal three segments elongated and fifth segment widened. Thorax: pronotum pubescent, slightly wider than length; forewing short with 3 oblique veins; mirror rounded and flattened with dividing vein not conspicuous; apical field nearly absent; external tympanum large and ovoid, internal tympanum absent (Fig. 4A, B); two long hairs on each foretibia, one extended outwards and one extended forwards (Fig. 4C). Hind tibia slightly shorter than hind femur, armed with 4 pairs of dorsal spurs; anal plate concave in the base (Fig. 3C); cercus little shorter than hind tibia. Genitalia: Epiphallus possessing a notch in middle, the lateral lobes of epiphallus curved inwards and upwards, and narrower than ectoparamere in lateral view (Fig. 5A–C). Female. Unknown. Coloration. Body uniformly black with the base of hind tibiae reddish brown (Fig. 2). Measurements (in mm). Male: body length (from head to anus) 16.59–16.90, pronotum length 3.22–3.93, forewing length 9.26–9.68, hind femur length 11.57–12.13. Distribution. China (Yunnan). Natural history. This species digs tunnels on slopes beside a road, and males sing near the entrance at night as Tarbinskiellus and Gymnogryllus species. The hole is as broad as the specimen’s size, and two hairs probably touch the wall of its tunnel for detecting vibration. Etymology. The species name bisetus refers to two filiform hairs on each foretibia., Published as part of Liu, Yi-Jiao & He, Zhu-Qing, 2022, First report of a cricket with filiform hairs on foretibia (Orthoptera: Gryllidae Gryllinae), pp. 583-588 in Zootaxa 5125 (5) on pages 585-586, DOI: 10.11646/zootaxa.5125.5.8, http://zenodo.org/record/6457447

genus Fodigryllus He gen. nov. ŔṞM Type species: Fodigryllus bisetus He sp. nov. Include species: type only. Diagnosis. Among the Gryllinae genera, this new genus described here is average sized with a black coloration. It possesses tympana on outside tibia and well developed forewing with mirror. Two filiform hairs on foretibia are its unique feature. General appearance is close to that of Vietacheta, but the ectoparamere of the new genus is little wider than the lateral lobes of epiphallus in lateral view, while the ectoparamere of Vietacheta is covered by the lateral lobes of epiphallus. The genitalia are also similar to that of Phonarellus, but the epiphallus of Phonarellus does not possess notch in middle. Compared with Gymnogryllus, the new genus is smaller in size. Furthermore, the ectoparamere of Gymnogryllus is longer, while that of the new genus is shorter than the lateral lobe of epiphallus. Its genitalia are also similar to those of Apterosvercus, while the latter has no tympana or wings. Molecular results supported the validation of the new genus Fodigryllus He gen. nov. (Fig. 1). Descriptions see species part. Etymology. The genus name is derived from Latin Fodi- (dig) + gryllus, which refers to its burrowing habit., Published as part of Liu, Yi-Jiao & He, Zhu-Qing, 2022, First report of a cricket with filiform hairs on foretibia (Orthoptera: Gryllidae Gryllinae), pp. 583-588 in Zootaxa 5125 (5) on pages 584-585, DOI: 10.11646/zootaxa.5125.5.8, http://zenodo.org/record/6457447

genus Fodigryllus He gen. nov. ŔṞM Type species: Fodigryllus bisetus He sp. nov. Include species: type only. Diagnosis. Among the Gryllinae genera, this new genus described here is average sized with a black coloration. It possesses tympana on outside tibia and well developed forewing with mirror. Two filiform hairs on foretibia are its unique feature. General appearance is close to that of Vietacheta, but the ectoparamere of the new genus is little wider than the lateral lobes of epiphallus in lateral view, while the ectoparamere of Vietacheta is covered by the lateral lobes of epiphallus. The genitalia are also similar to that of Phonarellus, but the epiphallus of Phonarellus does not possess notch in middle. Compared with Gymnogryllus, the new genus is smaller in size. Furthermore, the ectoparamere of Gymnogryllus is longer, while that of the new genus is shorter than the lateral lobe of epiphallus. Its genitalia are also similar to those of Apterosvercus, while the latter has no tympana or wings. Molecular results supported the validation of the new genus Fodigryllus He gen. nov. (Fig. 1). Descriptions see species part. Etymology. The genus name is derived from Latin Fodi- (dig) + gryllus, which refers to its burrowing habit.

Zilingxi He gen. nov. ������aeoi Diagnosis. A black line above clypeus (in species Z. dian sp. nov., this black line extended to frontal rostrum), vertex with irregular black stripes, large tympanum on the inner side of fore tibia, but no tympanum on outside, vein more or less crossed, cercus long, pseudepiphallic paramere of male genitalia possessing 2���3 branches, ovipositor short (length:width=4.6~4.7:1) and yellow in color, while in other genus, ovipositor long (length:width=8.4~9.5:1), and brown in color. Veins in yellow color while cells in black, a black circle on the apical area of hind femur. Etymology. The specific epithet Zilingxi is for the Chinese phonetic alphabet ������ae, which refers to the black line above clypeus. Type species. Zilingxi danmai sp. nov. described here., Published as part of Li, Shi-Yu, Liu, Yi-Jiao & He, Zhu-Qing, 2022, New and little-known mute Trigonidiinae species from China (Orthoptera: Grylloidea: Trigonidiidae), pp. 210-220 in Zootaxa 5115 (2) on page 214, DOI: 10.11646/zootaxa.5115.2.2, http://zenodo.org/record/6352266

Amusurgus (Amusurgus) bispinosus He, Li, Fang & Liu, 2010 Figs. 2C, 3C, 4C, 5C, 6C, 7C Amusurgus (Amusurgus) bispinosus He et al. 2010; He 2018 Material examined. 1 female, CHINA, Yunnan Prov., Puer, 8-viii-2017, coll. He Zhu-Qing; 5 females, CHINA, Yunnan Prov., Menglun, 19-ix-2017, coll. He Zhu-Qing; 1 male, CHINA, Yunnan Prov., Mengla, 14-vii-2021, coll. He Zhu-Qing. Measurements (in mm). Male: SZ 10.02, PR 0.94, HF 5.82; Female: SZ 10.49���11.72, PR 0.93���1.15, HF 5.83���5.86, OV 3.57���3.92. Distribution. China (Yunnan). Remarks. Some individuals have yellow face (Fig. 4 C-2), while others have reddish line above clypeus (Fig. 4 C-1). He et al. (2020) proceeded a phylogenetic study on Chinese Trigonidiinae species. In their study, only A. (U.) genji and A. (A.) fulvus were provided and they have close relationship. In our result, when A. (A.) bispinosus is included, subgenus Usgmona is nest with subgenus Amusurgus. To confirm their relationship, more species are needed for molecular study in the further., Published as part of Li, Shi-Yu, Liu, Yi-Jiao & He, Zhu-Qing, 2022, New and little-known mute Trigonidiinae species from China (Orthoptera: Grylloidea: Trigonidiidae), pp. 210-220 in Zootaxa 5115 (2) on page 214, DOI: 10.11646/zootaxa.5115.2.2, http://zenodo.org/record/6352266, {"references":["He, Z. - Q., Li, K., Fang, Y. & Liu, X. - W. (2010) A taxonomic study of the genus Amusurgus Brunner von Wattenwyl from China (Orthoptera, Gryllidae, Trigonidiinae). Zootaxa, 2423 (1), 55 - 62. https: // doi. org / 10.11646 / zootaxa. 2423.1.4","He, Z. - Q. (2018) A checklist of Chinese crickets (Orthoptera: Gryllidea). Zootaxa, 4369 (4), 515 - 535. https: // doi. org / 10.11646 / zootaxa. 4369.4.4","He, Z. Q., Shen, C. Z. & Wu, X. (2020) The first phylogenetic study of sword-tail crickets from China inferred from COI, 18 S and 28 S genes, with the establishment of two new genera and description of one new species (Orthoptera: Grylloidea: Trigonidiidae). Zootaxa, 4890 (1), 119 - 128. https: // doi. org / 10.11646 / zootaxa. 4890.1.7"]}

Key to Chinese mute Trigonidiinae genera and subgenera 1. False veins between real veins................................................ Trigonidium Rambur, 1838 ḃaeoi - No false vein......................................................................................... 2 2. Tegmen almost glabrous................................................................................ 3 - Tegmen covered with an abundant fine pubescence.......................................................... 4 3 Veins quite straight and parallel........................................... Rhicnogryllus Chopard, 1925 aüḃoi - Veins oblique and more or less diverging............................................ Metioche Stål, 1877 LJ ḃaeoi 4. Hind femur with black and white stripes..................................... Paranaxipha Chopard, 1925 Ēḃaeoi - Hind femur concolor................................................................................... 5 5. Veins straight and parallel, vertex flattened................................... Metiochodes Chopard, 1932 哑ḃaeoi - Veins oblique and more or less diverging, vertex convex...................................................... 6 6. A black line above clypeus, tympana on inner side only, pseudepiphallic paramere of male genitalia possessing 2–3 branches, ovipositor short............................................................... Zilingxi He gen. nov. ḙḃaeoi - Not fit all features described above....................................................................... 7 7. Males (short wing form) without tympanum................................. Usgmona Furukawa, 1970 DZNJ ḃaeoi - Males (short wing form) with tympanum.......................... Amusurgus Brunner von Wattenwyl, 1893 NJ ḃaeoi

Zilingxi He gen. nov. ḙḃaeoi Diagnosis. A black line above clypeus (in species Z. dian sp. nov., this black line extended to frontal rostrum), vertex with irregular black stripes, large tympanum on the inner side of fore tibia, but no tympanum on outside, vein more or less crossed, cercus long, pseudepiphallic paramere of male genitalia possessing 2–3 branches, ovipositor short (length:width=4.6~4.7:1) and yellow in color, while in other genus, ovipositor long (length:width=8.4~9.5:1), and brown in color. Veins in yellow color while cells in black, a black circle on the apical area of hind femur. Etymology. The specific epithet Zilingxi is for the Chinese phonetic alphabet ḙḃae, which refers to the black line above clypeus. Type species. Zilingxi danmai sp. nov. described here.

Zilingxi dian He sp. nov. ÷ḙḃae Figs. 3F, 4F, 5F, 6F, 7F Holotype. male, CHINA, Yunnan Prov., Menglun, E 101.28, N 21.92, 19-ix-2017, coll. He Zhu-Qing. Paratypes. 2 males & 2 females, CHINA, Yunnan Prov., Menglun, 19-ix-2017, coll. He Zhu-Qing; 1 female, CHINA, Guangxi Prov., Chongzuo, Ningming, Paiyangshan, E 107.17, N 22.06, 6-vii-2021, coll. He Zhu-Qing. Description. Male. Very similar to Z. danmai (Figs. 3F, 5F). Genitalia: pseudepiphallic apical lobe (ps. a. l.) blunt, pseudepiphallic paramere (ps. p.) possessing two branches, inner pair short and inwards curved, outer pair outwards curved, and as long as ps. a. l. (Fig. 6F). Female. Similar to male. Ovipositor similar to that of Z. danmai (Fig. 7F). Coloration. Very similar to Z. danmai. A black line above clypeus, and extended to frontal rostrum (Fig. 4F), mid-femora possessing a black spot near apex (Fig. 3F). Measurements (in mm). Male: SZ 8.94–9.33, PR 0.85, HF 4.25–4.38; Female: SZ 8.95–9.54, PR 0.71–0.82, HF 3.71–4.22, OV 2.95–3.08. Etymology. The specific epithet dian is for the Chinese phonetic alphabet ÷, which refers to the spot near the apex of mid-femora. Distribution. China (Guangxi, Yunnan) Remarks. Z. dian and Z. danmai were sympatric in Menglun, Yunnan (He Z.-Q. personal observation), but they have different color patterns in frontal rostrum, mid-femora possessing a black spot near apex, and the shape of male genitalia.

Zilingxi danmai He sp. nov. �����������ae Figs. 2E, 3E, 4E, 5E, 6E, 7E Holotype. male, CHINA, Yunnan Prov., Menglun, E 101.28, N 21.92, 19-ix-2017, coll. He Zhu-Qing. Paratypes. 3 males & 2 females, CHINA, Yunnan Prov., Menglun, 19-ix-2017, coll. He Zhu-Qing; 1 female, CHINA, Hainan Prov., Ledong, Jianfengling, E 108.87, N 18.72, 9-viii-2018, coll. He Zhu-Qing; 1 male, CHINA, Yunnan Prov., Honghe, Hekou, E 103.84, N 22.76, 11- vii-2021, coll. He Zhu-Qing. Description. Male. Body small and slender (Figs. 2E, 3E). Head little wider than anterior margin of pronotum, vertex convex with strong bristles, frontal rostrum almost as wide as 1st antennal joint, 5th joint of maxillary palpi long (Fig. 4E). Pronotum transverse with strong bristles, posterior margin wider than anterior margin. Tympana on inner side of fore tibiae only (Fig. 5E), hind tibia with three pairs of dorsal spurs. Forewing with pubescence and oblique veins on dorsal field, hindwing longer than forewing. Genitalia: pseudepiphallus with lobe on each outside (ps. a. l.), pseudepiphallic paramere (ps. p.) possessing three branches, but short than ps. a. l., outer pair possessing diverging apex, middle pair curved with pointed apex, inner pair straight with pointed apex (Fig. 6E). Female. Similar to male, ovipositor short and in yellow color (Fig. 7E). Coloration. Body almost in brown. A black line above clypeus, but not extended to frontal rostrum (Fig. 4E), vertex with irregular black lines, dorsal field of pronotum black, separated by a white line in middle and possessing irregular white stripes, a black line from lateral lobes of pronotum to lateral field of forewing, fore- and mid-femora possessing distinct spot near apex (some individual not), veins yellow and cells black, hindwing black (exceeded part), ovipositor yellow. Measurements (in mm). Male: SZ 9.87���10.27, PR 0.81, HF 4.97���5.12; Female: SZ 10.34���11.08, PR 1.03��� 1.11, HF 5.92���6.38, OV 2.96���3.09. Etymology. The specific epithet danmai is for the Chinese phonetic alphabet �����, which refers to the veins in yellow color, while cells in black. Distribution. China (Yunnan, Hainan)., Published as part of Li, Shi-Yu, Liu, Yi-Jiao & He, Zhu-Qing, 2022, New and little-known mute Trigonidiinae species from China (Orthoptera: Grylloidea: Trigonidiidae), pp. 210-220 in Zootaxa 5115 (2) on pages 215-216, DOI: 10.11646/zootaxa.5115.2.2, http://zenodo.org/record/6352266

Zilingxi dian He sp. nov. ��������ae Figs. 3F, 4F, 5F, 6F, 7F Holotype. male, CHINA, Yunnan Prov., Menglun, E 101.28, N 21.92, 19-ix-2017, coll. He Zhu-Qing. Paratypes. 2 males & 2 females, CHINA, Yunnan Prov., Menglun, 19-ix-2017, coll. He Zhu-Qing; 1 female, CHINA, Guangxi Prov., Chongzuo, Ningming, Paiyangshan, E 107.17, N 22.06, 6-vii-2021, coll. He Zhu-Qing. Description. Male. Very similar to Z. danmai (Figs. 3F, 5F). Genitalia: pseudepiphallic apical lobe (ps. a. l.) blunt, pseudepiphallic paramere (ps. p.) possessing two branches, inner pair short and inwards curved, outer pair outwards curved, and as long as ps. a. l. (Fig. 6F). Female. Similar to male. Ovipositor similar to that of Z. danmai (Fig. 7F). Coloration. Very similar to Z. danmai. A black line above clypeus, and extended to frontal rostrum (Fig. 4F), mid-femora possessing a black spot near apex (Fig. 3F). Measurements (in mm). Male: SZ 8.94���9.33, PR 0.85, HF 4.25���4.38; Female: SZ 8.95���9.54, PR 0.71���0.82, HF 3.71���4.22, OV 2.95���3.08. Etymology. The specific epithet dian is for the Chinese phonetic alphabet ��, which refers to the spot near the apex of mid-femora. Distribution. China (Guangxi, Yunnan) Remarks. Z. dian and Z. danmai were sympatric in Menglun, Yunnan (He Z.-Q. personal observation), but they have different color patterns in frontal rostrum, mid-femora possessing a black spot near apex, and the shape of male genitalia., Published as part of Li, Shi-Yu, Liu, Yi-Jiao & He, Zhu-Qing, 2022, New and little-known mute Trigonidiinae species from China (Orthoptera: Grylloidea: Trigonidiidae), pp. 210-220 in Zootaxa 5115 (2) on pages 216-217, DOI: 10.11646/zootaxa.5115.2.2, http://zenodo.org/record/6352266

Amusurgus (Usgmona) genji (Furukawa, 1970) Figs. 2A, 3A, 4A, 5A, 6A, 7A Usgmona genji Furukawa 1970 Metiochodes genji Ichikawa et al. 2000 Amusurgus (Usgmona) genji He et al. 2010; He 2018 Material examined. 1 male, CHINA, Shanghai, Chenshan, 9-viii-2010, coll. He Zhu-Qing; 1 nymph (female), CHINA, Zhejiang Prov., Ningbo, 12-viii-2010, coll. Guo Jiang-Li; 1 female, CHINA, Shanghai, Chongming, 23- viii-2010, coll. Fang Yan; 1 male, CHINA, Shanghai, Chongming, 23-viii-2010, coll. Guo Jiang-Li; 8 females, CHINA, Shanghai, Chongming, 23-viii-2010, coll. Ye Shui-Song; 3 males, CHINA, Shanghai, Dajinshandao, 16- viii-2016, coll. He Zhu-Qing; 3 males & 2 females, CHINA, Zhejiang Prov., Hangzhou, 8-iv-2017, coll. He Zhu-Qing; 4 females, CHINA, Zhejiang Prov., Tianmushan, 17-viii-2018, coll. He Zhu-Qing. Measurements (in mm). Male: SZ 6.49, PR 0.92, HF 4.53; Female: SZ 6.97���7.58/12.08���12.89, PR 0.92���1.03, HF 5.42���5.93, OV 3.47���4.3. Distribution. China (Zhejiang, Shanghai), Japan. Remarks. Both long and short wing forms exist in this species (Fig. 3A). Their head, thorax and veins are similar (Fig. 4A), but tympana are varied. We found only female individuals in long wing form. They have inner tympana, and outer ones are very small (Fig. 5 A-3). In short wing form, males have none tympanum on both sides (Fig. 5 A-1); some female individuals have small inner tympana (hind wing shed individuals?), and others have none tympanum as males (Fig. 5 A-2). Discussion of phylogenetics for this subgenus and species see remarks of Amusurgus bispinosus. Amusurgus (Amusurgus) fulvus Brunner von Wattenwyl, 1893 Figs. 2B, 3B, 4B, 5B, 6B, 7B Amusurgus fulvus Brunner von Wattenwyl 1893 Amusurgus (Amusurgus) fulvus He et al. 2010; He 2018 Amusurgus lateralis Chopard, 1969; He et al. 2010 synonym, Published as part of Li, Shi-Yu, Liu, Yi-Jiao & He, Zhu-Qing, 2022, New and little-known mute Trigonidiinae species from China (Orthoptera: Grylloidea: Trigonidiidae), pp. 210-220 in Zootaxa 5115 (2) on page 213, DOI: 10.11646/zootaxa.5115.2.2, http://zenodo.org/record/6352266, {"references":["Furukawa, H. (1970) Two new interesting genera and species of crickets of Japan (Orthoptera). Kontyu, 38, 59 - 66.","Ichikawa, A., Murai, T. & Honda, E. (2000) Monograph of Japanese crickets (Orthoptera; Grylloidea). Bulletin of the Hoshizaki Green Found, 4, 257 - 332.","He, Z. - Q., Li, K., Fang, Y. & Liu, X. - W. (2010) A taxonomic study of the genus Amusurgus Brunner von Wattenwyl from China (Orthoptera, Gryllidae, Trigonidiinae). Zootaxa, 2423 (1), 55 - 62. https: // doi. org / 10.11646 / zootaxa. 2423.1.4","He, Z. - Q. (2018) A checklist of Chinese crickets (Orthoptera: Gryllidea). Zootaxa, 4369 (4), 515 - 535. https: // doi. org / 10.11646 / zootaxa. 4369.4.4","Brunner von Wattenwyl, K. (1893) Revision du systeme des orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 13, 5 - 230. https: // doi. org / 10.5962 / bhl. title. 5121","Chopard, L. (1969) Orthoptera Volume 2 Grylloidea. In: Sewell, R. B. S. (Ed.), The Fauna of India and the Adjacent Countries, 1 - 421."]}

Key to Chinese mute Trigonidiinae genera and subgenera 1. False veins between real veins................................................ Trigonidium Rambur, 1838 ���aeoi - No false vein......................................................................................... 2 2. Tegmen almost glabrous................................................................................ 3 - Tegmen covered with an abundant fine pubescence.......................................................... 4 3 Veins quite straight and parallel........................................... Rhicnogryllus Chopard, 1925 a�����oi - Veins oblique and more or less diverging............................................ Metioche St��l, 1877 LJ ���aeoi 4. Hind femur with black and white stripes..................................... Paranaxipha Chopard, 1925 Ē���aeoi - Hind femur concolor................................................................................... 5 5. Veins straight and parallel, vertex flattened................................... Metiochodes Chopard, 1932 ������aeoi - Veins oblique and more or less diverging, vertex convex...................................................... 6 6. A black line above clypeus, tympana on inner side only, pseudepiphallic paramere of male genitalia possessing 2���3 branches, ovipositor short............................................................... Zilingxi He gen. nov. ������aeoi - Not fit all features described above....................................................................... 7 7. Males (short wing form) without tympanum................................. Usgmona Furukawa, 1970 DZNJ ���aeoi - Males (short wing form) with tympanum.......................... Amusurgus Brunner von Wattenwyl, 1893 NJ ���aeoi, Published as part of Li, Shi-Yu, Liu, Yi-Jiao & He, Zhu-Qing, 2022, New and little-known mute Trigonidiinae species from China (Orthoptera: Grylloidea: Trigonidiidae), pp. 210-220 in Zootaxa 5115 (2) on page 213, DOI: 10.11646/zootaxa.5115.2.2, http://zenodo.org/record/6352266, {"references":["Furukawa, H. (1970) Two new interesting genera and species of crickets of Japan (Orthoptera). Kontyu, 38, 59 - 66.","Brunner von Wattenwyl, K. (1893) Revision du systeme des orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 13, 5 - 230. https: // doi. org / 10.5962 / bhl. title. 5121"]}

Metiochodes minor Li, He & Liu, 2010 Figs. 2D, 3D, 4D, 5D, 6D, 7D Metiochodes minor Li et al. 2010; He 2018. Material examined. 1 male, CHINA, Fujiang Prov., Sanming, Mingxi, Longkeng, 8-viii-2008, coll. He Zhu-Qing; 2 females, CHINA, Zhejiang Prov., Tianmushan, 17-viii-2018, coll. He Zhu-Qing; 3 males, CHINA, Zhejiang Prov., Tianmushan, 11-viii-2019, coll. He Zhu-Qing; 1 male, CHINA, Zhejiang Prov., Tianmushan, 6-ix-2019, coll. He Zhu-Qing. Measurements (in mm). Male: SZ 9.75���9.79, PR 0.70���0.78, HF 4.10���4.23; females: SZ 10.19���11.81, PR 0.75���0.85, HF 4.55���4.82, OV 3.27���3.58. Distribution. China (Fujian, Zhejiang)., Published as part of Li, Shi-Yu, Liu, Yi-Jiao & He, Zhu-Qing, 2022, New and little-known mute Trigonidiinae species from China (Orthoptera: Grylloidea: Trigonidiidae), pp. 210-220 in Zootaxa 5115 (2) on page 214, DOI: 10.11646/zootaxa.5115.2.2, http://zenodo.org/record/6352266, {"references":["Li, K., He, Z. - Q. & Liu, X. - W. (2010) A taxonomic study of the genus Metiochodes Chopard from China (Orthoptera, Gryllidae, Trigonidiinae). Zootaxa, 2506, 43 - 50.","He, Z. - Q. (2018) A checklist of Chinese crickets (Orthoptera: Gryllidea). Zootaxa, 4369 (4), 515 - 535. https: // doi. org / 10.11646 / zootaxa. 4369.4.4"]}

Amusurgus (Amusurgus) bispinosus He, Li, Fang & Liu, 2010 Figs. 2C, 3C, 4C, 5C, 6C, 7C Amusurgus (Amusurgus) bispinosus He et al. 2010; He 2018 Material examined. 1 female, CHINA, Yunnan Prov., Puer, 8-viii-2017, coll. He Zhu-Qing; 5 females, CHINA, Yunnan Prov., Menglun, 19-ix-2017, coll. He Zhu-Qing; 1 male, CHINA, Yunnan Prov., Mengla, 14-vii-2021, coll. He Zhu-Qing. Measurements (in mm). Male: SZ 10.02, PR 0.94, HF 5.82; Female: SZ 10.49–11.72, PR 0.93–1.15, HF 5.83–5.86, OV 3.57–3.92. Distribution. China (Yunnan). Remarks. Some individuals have yellow face (Fig. 4 C-2), while others have reddish line above clypeus (Fig. 4 C-1). He et al. (2020) proceeded a phylogenetic study on Chinese Trigonidiinae species. In their study, only A. (U.) genji and A. (A.) fulvus were provided and they have close relationship. In our result, when A. (A.) bispinosus is included, subgenus Usgmona is nest with subgenus Amusurgus. To confirm their relationship, more species are needed for molecular study in the further.

Genus Phonarellus Gorochov, 1983 Type species: Phonarellus (Phonarellus) minor Chopard, 1959 Key to Phonarellus species from East Asia 1. head red or yellow (Figs. 3 AB, 4ABDE)................................................................... 2 - head black (Figs. 2, 3 CD)............................................................................... 3 2. hind femora yellow (Fig. 4 BF)...................................................... P. (P.) flavipes (WAE��fi) - hind femora almost black with apical part red (Fig. 3 AB)..................................... P. (P.) minor (�����fi) 3. hind femora black (Fig. 3 CD)........................................................ P. (P.) ritsemae (k��fi) - hind femora with black and white stripes (Fig. 2)............................. P. (P.) zebripes He sp. nov. (NJDZ��fi), Published as part of Liu, Yi-Jiao & He, Zhu-Qing, 2022, A new species of genus Phonarellus Gorochov, 1983 from Yunnan, China (Orthoptera: Gryllidae: Gryllinae), pp. 94-100 in Zootaxa 5093 (1) on page 96, DOI: 10.11646/zootaxa.5093.1.7, http://zenodo.org/record/5901145, {"references":["Gorochov, A. V. (1983) To the knowledge of the cricket tribe Gryllini (Orthoptera, Gryllidae). Entomologicheskoe Obozrenie, 62 (2), 314 - 330.","Chopard, L. (1959) Gryllides d'Iran: (Ergebnisse der Entomologischen Reisen Willi Richer, Stuttgart, im Iran 1954 und 1956 - Nr. 22). Staatliches Museum fur Naturkunde."]}

Phonarellus (Phonarellus) zebripes He sp. nov. NJDZā��� Figs. 2, 5���6 Holotype: male, CHINA, Yunnan, Nabang (E 97.86 N 24.02), 25-viii-2019, coll. (nymph when collected) Paratypes: 1 nymph (female), same data as holotype. Description. Male. Head: shining, vertex protruding; frontal rostrum flat and as wide as the 1st antennal joint; ocelli small, the middle one flattened and in line with the lateral ones (Fig. 5C); maxillary palpi normal. Thorax: anterior and posterior margin parallel and almost straight; one oval tympanum on outside of foretibia and one small tympanum on inner side (Fig. 5E, F); forewings well developed with one dividing vein, 2.5���3 times as long as pronotum, but not reaching the apical part of abdomen (Fig. 5A); hind femora longer than hind tibia, hind tibia with 6 dorsal spurs on outer side and 5 spurs on inner side (Fig. 5G); epiproct and cercus typical. Genitalia: a pair of epiphallic posterior lobes with pointed apex directed upwards; ectoparameron not reaching the apex of epiphallus lobes, inner sides of ectoparameron curved downwards (Fig. 6). Nymph (Female). Similar to male, but hind tibia with 6 pairs of dorsal spurs (Fig. 5B, D). Coloration. Adults: Body dark black, the antenna and legs with black and white stripes, the spurs on hind tibiae brown, cercus black with base segments white (Fig. 2). Measurements (in mm). Male: body length 14.34, pronotum length 2.27, hind femur length 8.28, forewing length 7.24. Nymph (Female): body length 13.08, pronotum length 2.22, hind femur length 8.26, forewing length 3.72, ovipositor length 2.92. Distribution. China (Yunnan). Etymology. The name zebripes is used to emphasize the color patterns on its hind femora., Published as part of Liu, Yi-Jiao & He, Zhu-Qing, 2022, A new species of genus Phonarellus Gorochov, 1983 from Yunnan, China (Orthoptera: Gryllidae: Gryllinae), pp. 94-100 in Zootaxa 5093 (1) on pages 96-97, DOI: 10.11646/zootaxa.5093.1.7, http://zenodo.org/record/5901145

2.6 Hemigyrus (Tomomima) spinosus (Bey-Bienko, 1955) Figs. 11A���H Tomomima spinosa: Bey-Bienko, 1955, Zoologicheskii Zhurnal, 34: 1258; Jin & Xia, 1994, Journal of Orthoptera Research, 3: 26. Hemigyrus (Tomomima) spinosus: Gorochov & Voltshenkova, 1998, Zoologicheskii Zhurnal, 77 (7): 863; Kim & Pham, 2014, Zootaxa, 3811 (1): 74., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 112, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676, {"references":["Bey-Bienko, G. Y. (1955) Studies on fauna and systematic superfamily Tettigonioidea (Orthoptera) of China. Zoologicheskii Zhurnal, 34, 1250 - 1271.","Jin, X. B. & Xia, K. L. (1994) An Index-Catalogue of Chinese Tettigoniodea (Orthopteroidea: Grylloptera). Journal of Orthoptera Research, 3, 15 - 41. https: // doi. org / 10.2307 / 3503405","Gorochov, A. V. & Voltshenkova, N. A. (1998) Vietnamese representatives of the genus Hemigyrus (Orthoptera, Tettigoniidae). Zoologicheskii Zhurnal, 77 (7), 860 - 864.","Kim, T. & Pham, H. T. (2014) Checklist of Vietnamese Orthoptera (Saltatoria). Zootaxa, 3811, 53 - 82. https: // doi. org / 10.11646 / zootaxa. 3811.1.3"]}

2.1 Hemigyrus (Hemigyrus) tonkinensis Beier, 1954 Figs. 6A���H Hemigyrus tonkinensis: Beier, 1954, Revision der Pseudophyllinen, 145; Jin & Xia, 1994, Journal of Orthoptera Research, 3: 24; Liu & Yin, 2004, Insects from Mt. Shiwandashan Area of Guangxi, 97. Hemigyrus (Hemigyrus) tonkinensis: Gorochov & Voltshenkova, 1998, Zoologicheskii Zhurnal, 77 (7): 861; Kim & Pham, 2014, Zootaxa, 3811 (1): 74., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 103, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676, {"references":["Beier, M. (1954) Revision der Pseudophyllinen. Instituto EspanTol de Entomologiia, Madrid, 479 pp.","Jin, X. B. & Xia, K. L. (1994) An Index-Catalogue of Chinese Tettigoniodea (Orthopteroidea: Grylloptera). Journal of Orthoptera Research, 3, 15 - 41. https: // doi. org / 10.2307 / 3503405","Liu, X. W & Yin, H. S. (2004) Orthoptera: Tettigonioidea and Stenopelmatoidea. In: Yang, X. K. (Ed.), Insects from Mt. Shiwandashan Area of Guangxi. China Forestry Publishing House, Beijing, pp. 90 - 110.","Gorochov, A. V. & Voltshenkova, N. A. (1998) Vietnamese representatives of the genus Hemigyrus (Orthoptera, Tettigoniidae). Zoologicheskii Zhurnal, 77 (7), 860 - 864.","Kim, T. & Pham, H. T. (2014) Checklist of Vietnamese Orthoptera (Saltatoria). Zootaxa, 3811, 53 - 82. https: // doi. org / 10.11646 / zootaxa. 3811.1.3"]}

2.4 Hemigyrus (Hemigyrus) acutifolius Brunner von Wattenwyl, 1895 Figs. 9A���H Hemigyrus acutifolius: Brunner von Wattenwyl, 1895, Verhandlungen der Kaiserlich-K��niglichen Zoologisch-Botanischen Gesellschaft in Wien, 45: 48; Kirby, 1906, A Synonymic Catalogue of Orthoptera (Orthoptera Saltatoria, Locustidae vel Acridiidae), 2: 297; Beier, 1954, Revision der Pseudophyllinen, 146; Liana, 1999, Bulletin of the Museum and Institute of Zoology PAS, 2: 62. Hemigyrus (Hemigyrus) acutifolius: Gorochov & Voltshenkova, 1998, Zoologicheskii Zhurnal, 77 (7): 860., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 107, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676, {"references":["Brunner von Wattenwyl, C. (1895) Monographie der Pseudophylliden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 45, 47 - 48.","Kirby, W. F. (1906) A Synonymic Catalogue of Orthoptera. Vol. 2. Orthoptera Saltatoria. Part I. Achetidae et Phasgonuridae. British Museum, London, 502 pp.","Beier, M. (1954) Revision der Pseudophyllinen. Instituto EspanTol de Entomologiia, Madrid, 479 pp.","Liana, A. (1999) The type material of Pseudophyllidae (Orthoptera) in the Museum and Institute of Zoology PAS, Warsaw. Bulletin of the Museum and Institute of Zoology PAS, 2, 45 - 62.","Gorochov, A. V. & Voltshenkova, N. A. (1998) Vietnamese representatives of the genus Hemigyrus (Orthoptera, Tettigoniidae). Zoologicheskii Zhurnal, 77 (7), 860 - 864."]}

2.7 Hemigyrus (Tomomima) spinosus robustus subsp. nov. Xie, Wang & He Figs. 12A���H Material examined: Holotype ♂, CHINA, Guangxi Prov., Jinxiu (110.10��E, 24.15��N), 19-vii-2015, coll. He Zhu- Qing & Lu Hui. Paratypes: 1♂, CHINA, Guangxi Prov., Jinxiu (110.10��E, 24.15��N), 21-vii-2015, coll. He Zhu-Qing & Lu Hui; 1♀, CHINA, Guangxi Prov., Jinxiu (110.10��E, 24.15��N), 20-vii-2015, coll. He Zhu-Qing & Lu Hui. Description: Male: Size medium. Pronotum with sparse tubercles. Side lobes abdominal edge with tumors (Figs.12B, C). Front corners of mesosternum protruding tooth-shaped (Fig. 12D). Fore femora on the ventral inside edge with 5���9 spines, 3���9 spines on the outside edge. Hind femora with 12 lower outer, 8���13 lower inner and 11 upper median spines. Fore femora and median femora dark ventrally. Tegmina with almost truncated apical part (Figs. 5I, J). Epiproct nearly oval, with intermediate longitudinal ridge (Fig. 12E). Cerci robust, almost conical (Figs. 12E, F). Female: Similar to male in general appearance, but larger than male. Subgenital plate subtriangular, notched in terminal (Fig. 12G). Ovipositor wide relatively, distal half of dorsal margin armed with fine teeth (Fig. 12H). Comparison. H. (T.) spinosus robustus subsp. nov. is very similar to H. (T.) spinosus spinosus (Figs. 11A���H; 12A���H), but differs in the size (length of pronotum, tegmina, and hind femora of male in H. (T.) spinosus robustus subsp. nov. are 7.7���7.9, 35.8���36.2 and 18.9���20.6mm respectively, but these are 7.3���8.3, 30.3���34.2 and 17.6���19.6 mm in H. (T.) spinosus spinosus, respectively). H. (T.) spinosus robustus subsp. nov. have more spines on hind femora dorsally than H. (T.) spinosus spinosus (11 vs. 7���9). Measurements (in mm): Male: SZ 45.0���47.9, PR 7.7���7.9, FW 35.8���36.2, HF 18.9���20.6; Female: SZ 59.2, PR 10.5, FW 47.9, HF 22.3, OV 19.4. Etymology: Subspecies name robustus means it���s stronger than H. (T.) spinosus spinosus. Distribution: China (Guangxi)., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 112, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676

Genus Hemigyrus Brunner von Wattenwyl, 1893 Type species: Hemigyrus (Hemigyrus) amplus Brunner von Wattenwyl, 1893, Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 101, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676, {"references":["Brunner von Wattenwyl, C. (1893) Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Serie 2, 13 (33), 175 - 176. https: // doi. org / 10.5962 / bhl. title. 5121"]}

2.7 Hemigyrus (Tomomima) spinosus robustus subsp. nov. Xie, Wang & He Figs. 12A–H Material examined: Holotype ♂, CHINA, Guangxi Prov., Jinxiu (110.10°E, 24.15°N), 19-vii-2015, coll. He Zhu- Qing & Lu Hui. Paratypes: 1♂, CHINA, Guangxi Prov., Jinxiu (110.10°E, 24.15°N), 21-vii-2015, coll. He Zhu-Qing & Lu Hui; 1♀, CHINA, Guangxi Prov., Jinxiu (110.10°E, 24.15°N), 20-vii-2015, coll. He Zhu-Qing & Lu Hui. Description: Male: Size medium. Pronotum with sparse tubercles. Side lobes abdominal edge with tumors (Figs.12B, C). Front corners of mesosternum protruding tooth-shaped (Fig. 12D). Fore femora on the ventral inside edge with 5–9 spines, 3–9 spines on the outside edge. Hind femora with 12 lower outer, 8–13 lower inner and 11 upper median spines. Fore femora and median femora dark ventrally. Tegmina with almost truncated apical part (Figs. 5I, J). Epiproct nearly oval, with intermediate longitudinal ridge (Fig. 12E). Cerci robust, almost conical (Figs. 12E, F). Female: Similar to male in general appearance, but larger than male. Subgenital plate subtriangular, notched in terminal (Fig. 12G). Ovipositor wide relatively, distal half of dorsal margin armed with fine teeth (Fig. 12H). Comparison. H. (T.) spinosus robustus subsp. nov. is very similar to H. (T.) spinosus spinosus (Figs. 11A–H; 12A–H), but differs in the size (length of pronotum, tegmina, and hind femora of male in H. (T.) spinosus robustus subsp. nov. are 7.7–7.9, 35.8–36.2 and 18.9–20.6mm respectively, but these are 7.3–8.3, 30.3–34.2 and 17.6–19.6 mm in H. (T.) spinosus spinosus, respectively). H. (T.) spinosus robustus subsp. nov. have more spines on hind femora dorsally than H. (T.) spinosus spinosus (11 vs. 7–9). Measurements (in mm): Male: SZ 45.0–47.9, PR 7.7–7.9, FW 35.8–36.2, HF 18.9–20.6; Female: SZ 59.2, PR 10.5, FW 47.9, HF 22.3, OV 19.4. Etymology: Subspecies name robustus means it’s stronger than H. (T.) spinosus spinosus. Distribution: China (Guangxi).

2.3 Hemigyrus (Hemigyrus) minor Gorochov & Kang, 2003 Figs. 8A���H Hemigyrus (Hemigyrus) minor: Gorochov, Kang & Liu, 2003, Oriental Insects, 37: 494., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 107, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676, {"references":["Gorochov, A. V., Kang, L. & Liu, C. X. (2003) Two new species of Hemigyrus Brunner von Wattenwyl (Orthoptera: Tettigoniidae: Pseudophyllinae) from China. Oriental Insects, 37, 491 - 495. https: // doi. org / 10.1080 / 00305316.2003.10417368"]}

2.5 Hemigyrus (Tomomima) major Gorochov & Kang, 2003 Figs 10A–H Hemigyrus (Tomomima) major: Gorochov, Kang & Liu, 2003, Oriental Insects, 37: 492.

2.3 Hemigyrus (Hemigyrus) minor Gorochov & Kang, 2003 Figs. 8A–H Hemigyrus (Hemigyrus) minor: Gorochov, Kang & Liu, 2003, Oriental Insects, 37: 494.

2.5 Hemigyrus (Tomomima) major Gorochov & Kang, 2003 Figs 10A���H Hemigyrus (Tomomima) major: Gorochov, Kang & Liu, 2003, Oriental Insects, 37: 492., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 110, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676, {"references":["Gorochov, A. V., Kang, L. & Liu, C. X. (2003) Two new species of Hemigyrus Brunner von Wattenwyl (Orthoptera: Tettigoniidae: Pseudophyllinae) from China. Oriental Insects, 37, 491 - 495. https: // doi. org / 10.1080 / 00305316.2003.10417368"]}

2.2 Hemigyrus (Hemigyrus) amplus Brunner von Wattenwyl, 1893 Figs.7A���H Hemigyrus amplus: Brunner von Wattenwyl, 1893, Annali del Museo Civico di Storia Naturale di Genova Serie 2, 13 (33): 175; Brunner von Wattenwyl, 1895, Monographie der Pseudophylliden, 45: 47; Kirby, 1906, A Synonymic Catalogue of Orthoptera (Orthoptera Saltatoria, Locustidae vel Acridiidae), 2: 297; Beier, 1954, Revision der Pseudophyllinen, 144; Jin & Xia, 1994, Journal of Orthoptera Research, 3: 24. Hemigyrus (Hemigyrus) amplus: Ingrisch & Shishodia, 1998, Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71: 367; Shishodia, Chandra & Gupta, 2010, Records of the Zoological Survey of India, 314: 315., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Zong, Jing-Song, Li, Kai & He, Zhu-Qing, 2022, Taxonomic revision of the genus Hemigyrus Brunner von Wattenwyl, 1893 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 97-115 in Zootaxa 5092 (1) on page 106, DOI: 10.11646/zootaxa.5092.1.5, http://zenodo.org/record/5869676, {"references":["Brunner von Wattenwyl, C. (1893) Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Serie 2, 13 (33), 175 - 176. https: // doi. org / 10.5962 / bhl. title. 5121","Brunner von Wattenwyl, C. (1895) Monographie der Pseudophylliden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 45, 47 - 48.","Kirby, W. F. (1906) A Synonymic Catalogue of Orthoptera. Vol. 2. Orthoptera Saltatoria. Part I. Achetidae et Phasgonuridae. British Museum, London, 502 pp.","Beier, M. (1954) Revision der Pseudophyllinen. Instituto EspanTol de Entomologiia, Madrid, 479 pp.","Jin, X. B. & Xia, K. L. (1994) An Index-Catalogue of Chinese Tettigoniodea (Orthopteroidea: Grylloptera). Journal of Orthoptera Research, 3, 15 - 41. https: // doi. org / 10.2307 / 3503405","Ingrisch, S. & Shishodia, M. S. (1998) New species and records of Tettigoniidae from India (Ensifera). Mitteilungen der Munchner Entomologischen Gesellschaft, 71, 355 - 371.","Shishodia, M. S., Chandra, K. & Gupta, S. K. (2010) An annotated checklist of Orthoptera (Insecta) from India. Records of the Zoological Survey of India, 314, 1 - 366."]}

Genus Ptosoproctus Shen, Yin & He gen. nov. 折尾ỡffl Type species. Ptosoproctus lanzhouensis Shen, Yin & He, here designated Description. Body small. Fastigium of vertex distinctly wider than scape of antenna. Disc of pronotum with a broadly obtuse concavity in the middle on both sides, lateral carina faintly indicated, lateral lobes longer than high, with weakly humeral sinus. Prosternum unarmed. Brachypterous, tegmina around same length as pronotum in male; distinctly shorter than pronotum and only reaches 1 st abdominal tergite anterior margin, clearly visible and discontiguous dorsally in female. Fore-tibiae dorsally without internal spines; mid-tibiae dorsally with spines on both sides. Male 10 th abdominal tergite fused with epiproct, forming a down-folded membranous projection on tergite posterior margin, projection with a median longitudinal furrow. Cercus short and stout, apex short and obtusely conical; inner tooth near cercus apex, robust, recurved and apically acute. Subgenital plate with short styli. Titllators bear a row of denticles on the apical arms. Female cercus conical and pielous, 10 th abdominal tergite fused or not fused with epiproct, posterior margin with a median notch. 7 th abdominal sternite without hump or tubercule. Subgenital plate posterior margin with a median deep, acutely angled concave. Ovipositor shorter than hind femora, distinctly upcurved. Remarks. This new genus is closely related to Eulithoxenus but differs from the latter by: male 10 th abdominal tergite fused with epiproct, and female 7 th abdominal sternite without hump or tubercle. This new genus is also closely related to Novadrymadusa as indicated by the confluent epiproct, but differs from latter by: prosternum unarmed, male epiproct not strongly extended posteriorly, in the middle without 4 cross-shaped contiguous plates, and female 7 th sternite without an acute, spine-like projection posteriorly. Etymology. The new genus name refers to the Greek prefix “πτώσ-” (ptôs-, falling) and Greek “πρωκτός” (prôktós, anus), representing the down-folded epiproct., Published as part of Shen, Zi-Hao, Yin, Zi-Xu, Lee, Michael, Liu, Yi-Jiao, He, Zhu-Qing, Wang, Zi-Fan & Wang, Tong-Xin, 2021, Ptosoproctus gen. nov., a new genus with two new species of Shield-back Katydid with the first record of genus Eulithoxenus Bey-Bienko, 1951 from China (Orthoptera: Tettigoniidae: Tettigoniinae: Drymadusini), pp. 548-568 in Zootaxa 5067 (4) on pages 550-551, DOI: 10.11646/zootaxa.5067.4.4, http://zenodo.org/record/5683472

Ptosoproctus lanzhouensis Shen, Yin & He sp. nov. 兰州折尾ỡ Figs. 1–5, 9, 14–15 Description. General. Body small. Head frons flat, slightly oblique (Fig. 2 AB). Fastigium of vertex distinctly wider than scape of antenna (Fig. 2B). Eyes broadly round and projecting outward. Disc of pronotum both sides with a broadly obtuse concavity on middle part, anterior and posterior margin of pronotum slightly concaved, median carina and posterior transvers sulcus obscure, central transvers sulcus distinctly Chinese characters “八”-shaped, lateral carina faintly indicated; lateral lobes longer than high, with weakly humeral sinus (Fig. 2 BCEF). Prosternum unarmed (Fig. 2D). Mesosternum with a pair of obtuse triangular lobe-shaped processes, wider than long (Fig. 2D). Metasternum with a pair of arched lobe-shaped processes, wider than long (Fig. 2D). Dorsal surface of fore coxae with 1 long spine; length of fore femora nearly equal to pronotum, dorsally unarmed, ventrally with 0–2 internal spinules and externally unarmed, genicular lobes with 1–2 internal spinules and externally unarmed; fore tibiae both sides with a crack-shape tympana, dorsally with 3–4 externally spines and internally unarmed, ventrally with 6 pair of spines, a pair at the apical. Middle femora dorsally unarmed, ventrally with 0–1 internal spinules and externally unarmed, genicular lobes with 1–2 external spinules and 1 internal spinule; middle tibiae dorsally with 4–5 internal spines and 2–4 external spines, ventrally with 6 pair of spines, a pair at the apical. Hind femora dorsally unarmed, ventrally with 0–1 external spinules and 2–3 internal spinules, genicular lobes with 0–1 spinules on both sides; hind tibiae dorsally with 23–34 spines on both sides and 1 pair of apical spurs, ventrally with 8–12 spines on both sides and 2 pair of apical spurs. Male. Tegmina brachypterous, slightly shorter than pronotum, slight beyond or extending to 3 th abdominal tergite posterior margin. Left tegmina ventrally with stridulatory file about 30 teeth (Fig. 3E). Right tegmina ventrally with vestigial stridulatory file about 19 teeth (Fig. 3F). Mirror on right tegmina pentagonal, distinctly longer than wide (Fig. 3 CF). Hind wings rudimentary (Fig. 3D). 10 th abdominal tergite fused with epiproct, posterior margin of 10th abdominal tergite with a broadly triangle notch at the joint, form a strongly down-folded membranous projection on tergite posterior margin, apex broadly rounded, almost sightless from above, projection with a median longitudinal furrow which basal narrow and apical widish. Cercus pileous, short and stout, external margin almost straight; apex short and obtusely conical; inner tooth near cercus apex, robust, recurved, wide at base, gradually narrow towards the apical part, apex acute, raise first and then decurved. Subgenital plate about 1.25 times longer than width, with lateral carinae, middle part of posterior margin with a weak triangular excision, styli short (Fig. 4C). Titillators with broad basal arms, apical inflated connect with short apical arms; apical arms dorsally bearing one rows denticles, with acute apex (Fig. 4E). Female. Tegmina slightly shorter than quarter pronotum, extending to 1 th abdominal tergite anterior margin, clearly visible and discontiguous in dorsal view (Fig. 2 EF). Cercus conical and hirsute (Fig. 4F). 10 th abdominal tergite not fused with epiproct, posterior margin of 10 th abdominal tergite with a shallow broadly round notch on middle part (Fig. 4F). Subgenital plate wider than length, with lateral carinae, middle of posterior margin with a deep angle concave (Fig. 4G). Ovipositor shorter than hind femora, distinctly upcurved (Fig. 4H). Coloration. Body laterally and dorsally generally whitish gray, with dark brown or black marmoration, ventrally yellowish (greenish in life). Lower and hind margin of pronotum lateral lobes whitish gray with dark brown spots. Tegmina orange, costal area whitish gray with dark brown spots, radius area with dark brown spots. Spines of each leg black. Apex of male cercus inner tooth black. Ovipositor apical part black. Measurements(mm). Body (from head to tip of abdomen): ♂ 16.67–19.30, ♀ 17.12; pronotum: ♂ 4.46–4.60, ♀ 5.40; tegmina: ♂ 3.74–4.42, ♀ 1.30; mirror of right tegmina (from fore to hind): ♂ 1.90; fore femora: ♂ 4.46–4.80, ♀ 5.15; middle femora: ♂ 4.90–5.54, ♀ 6.10; hind femora: ♂ 17.42–19.20, ♀ 20.65; fore tibiae: ♂ 5.40–5.90, ♀ 6.62; middle tibiae: ♂ 6.0–6.77, ♀ 7.40; hind tibiae: ♂ 17.42–18.42, ♀ 19.70; male titillators (straight-line distance from base to end): 1.50; ovipositor: 10.90. Material examined. Holotype: ♂, Lanzhou University Yuzhong Campu, Xiaguanying town, Yuzhong country, Lanzhou city, Gansu Province, China (104°8'8.00"E, 35°56'40.10"N, alt. 1800m), 30-vi-2020, leg. Zifan Wang. Paratypes: 2♂, same data as in holotype; 2♂ 1♀, Lanzhou University Yuzhong Campu, Xiaguanying town, Yuzhong country, Lanzhou city, Gansu Province, China (104° 8'50.97"E, 35°57'2.75"N, alt. 1850m), 2-vii-2019, leg. Tongxin Wang. Remarks. See Ptosoproctus baishishanicus sp. nov. Songs. The adults make calling songs at night. The song consists of echemes with syllables. Echeme sequence duration for 23–34 s with 7–9 s interval. In each echeme sequence, there are about 40–50 echemes with rate as 1.76 echeme/s. Each echeme consist 1–2 syllables irregularly, echeme duration for 50–150 ms, echeme interval for 350–780 ms. Each syllable duration for 50–55 ms, syllables interval for 35–45 ms. Etymology. The specific epithet is named after Lanzhou, the type locality. Distribution: China (Gansu). Known only from the type locality., Published as part of Shen, Zi-Hao, Yin, Zi-Xu, Lee, Michael, Liu, Yi-Jiao, He, Zhu-Qing, Wang, Zi-Fan & Wang, Tong-Xin, 2021, Ptosoproctus gen. nov., a new genus with two new species of Shield-back Katydid with the first record of genus Eulithoxenus Bey-Bienko, 1951 from China (Orthoptera: Tettigoniidae: Tettigoniinae: Drymadusini), pp. 548-568 in Zootaxa 5067 (4) on pages 551-555, DOI: 10.11646/zootaxa.5067.4.4, http://zenodo.org/record/5683472

Genus Uvarovina Ramme, 1939 Type species. Bergiola daurica Uvarov,1928, Published as part of Shen, Zi-Hao, Yin, Zi-Xu, Lee, Michael, Liu, Yi-Jiao, He, Zhu-Qing, Wang, Zi-Fan & Wang, Tong-Xin, 2021, Ptosoproctus gen. nov., a new genus with two new species of Shield-back Katydid with the first record of genus Eulithoxenus Bey-Bienko, 1951 from China (Orthoptera: Tettigoniidae: Tettigoniinae: Drymadusini), pp. 548-568 in Zootaxa 5067 (4) on page 565, DOI: 10.11646/zootaxa.5067.4.4, http://zenodo.org/record/5683472, {"references":["Ramme, W. (1939) Beitrage zur Kenntnis der palaearktischen Orthopterenfauna (Tettig. u. Acrid.) III. Mitteilungen aus dem Zoologischen Museum in Berlin, 24, 62 - 94.","Uvarov, B. P. (1928) Orthoptera Palaearctica critica VI. Genus Bergiola Stschelk. (Tettig.). Eos, Revista Espanola de Entomologia, 4 (2), 243 - 251."]}

Genus Ptosoproctus Shen, Yin & He gen. nov. 折尾ỡffl Type species. Ptosoproctus lanzhouensis Shen, Yin & He, here designated Description. Body small. Fastigium of vertex distinctly wider than scape of antenna. Disc of pronotum with a broadly obtuse concavity in the middle on both sides, lateral carina faintly indicated, lateral lobes longer than high, with weakly humeral sinus. Prosternum unarmed. Brachypterous, tegmina around same length as pronotum in male; distinctly shorter than pronotum and only reaches 1 st abdominal tergite anterior margin, clearly visible and discontiguous dorsally in female. Fore-tibiae dorsally without internal spines; mid-tibiae dorsally with spines on both sides. Male 10 th abdominal tergite fused with epiproct, forming a down-folded membranous projection on tergite posterior margin, projection with a median longitudinal furrow. Cercus short and stout, apex short and obtusely conical; inner tooth near cercus apex, robust, recurved and apically acute. Subgenital plate with short styli. Titllators bear a row of denticles on the apical arms. Female cercus conical and pielous, 10 th abdominal tergite fused or not fused with epiproct, posterior margin with a median notch. 7 th abdominal sternite without hump or tubercule. Subgenital plate posterior margin with a median deep, acutely angled concave. Ovipositor shorter than hind femora, distinctly upcurved. Remarks. This new genus is closely related to Eulithoxenus but differs from the latter by: male 10 th abdominal tergite fused with epiproct, and female 7 th abdominal sternite without hump or tubercle. This new genus is also closely related to Novadrymadusa as indicated by the confluent epiproct, but differs from latter by: prosternum unarmed, male epiproct not strongly extended posteriorly, in the middle without 4 cross-shaped contiguous plates, and female 7 th sternite without an acute, spine-like projection posteriorly. Etymology. The new genus name refers to the Greek prefix “πτώσ-” (ptôs-, falling) and Greek “πρωκτός” (prôktós, anus), representing the down-folded epiproct.

Uvarovina chinensis Ramme, 1939 Fig. 16 Supplemental description. Prosternum unarmed (Fig. 16D). Mesosternum with a pair of obtuse triangular lobeshaped processes, slight wider than long (Fig. 16D). Metasternum with a pair of obtuse triangular lobe-shaped processes, noticeable wider than long (Fig. 16D). Titillators with longer apical arms, dorsally bearing one rows denticles; basal arms with a right-angle curve on middle part, apical part inflated in renal shape (Fig. 16H). Material examined. 1♂ 1♀, Jiangshuiquan reservoir park, Jinan city, Shandong province, China (117°4'35"E, 36°37'10"N, alt. 273m), 20-viii-2020, leg. Sizhong Shen., Published as part of Shen, Zi-Hao, Yin, Zi-Xu, Lee, Michael, Liu, Yi-Jiao, He, Zhu-Qing, Wang, Zi-Fan & Wang, Tong-Xin, 2021, Ptosoproctus gen. nov., a new genus with two new species of Shield-back Katydid with the first record of genus Eulithoxenus Bey-Bienko, 1951 from China (Orthoptera: Tettigoniidae: Tettigoniinae: Drymadusini), pp. 548-568 in Zootaxa 5067 (4) on page 565, DOI: 10.11646/zootaxa.5067.4.4, http://zenodo.org/record/5683472, {"references":["Ramme, W. (1939) Beitrage zur Kenntnis der palaearktischen Orthopterenfauna (Tettig. u. Acrid.) III. Mitteilungen aus dem Zoologischen Museum in Berlin, 24, 62 - 94."]}

Eulithoxenus Bey-Bienko, 1951 优岩ỡffl Type species. Bergiola mongolicus Uvarov, 1928, Published as part of Shen, Zi-Hao, Yin, Zi-Xu, Lee, Michael, Liu, Yi-Jiao, He, Zhu-Qing, Wang, Zi-Fan & Wang, Tong-Xin, 2021, Ptosoproctus gen. nov., a new genus with two new species of Shield-back Katydid with the first record of genus Eulithoxenus Bey-Bienko, 1951 from China (Orthoptera: Tettigoniidae: Tettigoniinae: Drymadusini), pp. 548-568 in Zootaxa 5067 (4) on page 559, DOI: 10.11646/zootaxa.5067.4.4, http://zenodo.org/record/5683472, {"references":["Bey-Bienko, G. Y. (1951) Investigations on grasshoppers (Orthoptera, Tettigoniidae) of USSR and neighbouring countries. Trudy Russkago Entomologicheskago Obshchestva, 43, 129 - 170.","Uvarov, B. P. (1928) Orthoptera Palaearctica critica VI. Genus Bergiola Stschelk. (Tettig.). Eos, Revista Espanola de Entomologia, 4 (2), 243 - 251."]}

Ptosoproctus baishishanicus Shen, Yin, Lee & He sp. nov. 白石山折尾ỡ Figs. 1, 6–9, 14–15 Description. General. Body small. Head frons flat. Fastigium of vertex distinctly wider than scape of antenna (Fig. 6 AB). Head dorsally with an obscure and continuous median carina. Eyes broadly round and projecting outward. Disc of pronotum both sides with a broadly obtuse concavity on middle part, anterior margin of pronotum slightly concaved and posterior margin nearly straight, median carina continuous and distinctly indicated, central transvers sulcus distinctly Chinese characters “八”-shaped, lateral carina faintly indicated; lateral lobes longer than high, with weakly humeral sinus (Fig. 6 BC). Prosternum unarmed (Fig. 6D). Mesosternum with a pair of obtuse triangular lobe-shaped processes, wider than long (Fig. 6D). Metasternum with a pair of obtuse triangular lobe-shaped processes, extremely wider than long (Fig. 6D). Dorsal surface of fore coxae with 1 long spine; fore femora 1.15–1.2 times longer than pronotum, dorsally unarmed, ventrally with 0–2 internal spinules and externally unarmed, genicular lobes with 1–2 internal spinules and 1 external spinule; fore tibiae both sides with a crack-shape tympana, ventrally with 6 pair of spines, a pair at the apical. Middle femora dorsally unarmed, ventrally both sides with 0–1 spinules, genicular lobes with 1 internal spinules and 1–3 external spinules; middle tibiae ventrally with 6 pair of spines, a pair at the apical. Hind femora dorsally unarmed, ventrally with 0–4 spinules on both sides, genicular lobes with 0–1 spinules on both sides; hind tibiae dorsally with 18–27 spines on both sides and 1 pair of apical spurs, ventrally with 9–10 spines on both sides and 2 pair of apical spurs. Male. Tegmina brachypterous, slightly longer than pronotum, extending to 2 th abdominal tergite posterior margin. Left tegmina ventrally with stridulatory file about 36 teeth (Fig. 6G). Right tegmina ventrally with vestigial stridulatory file about 28 teeth (Fig. 6H). Mirror on right tegmina slightly longer than wide (Fig. 6 FH). Hind wings rudimentary (Fig. 6I). Fore tibiae dorsally armed 2–3 external spines and internally unarmed. Middle tibiae dorsally with 2 external spines and 1–2 internal spines (one at apical). 10 th abdominal tergite fused with epiproct, posterior margin of 10th abdominal tergite with a broadly triangle notch at the joint, form a down-folded membranous projection on tergite posterior margin, projection with a gourd-shaped median longitudinal furrow, apex broadly rounded. Cercus conical, pileous, short and stout, external margin straight; apex short and obtusely conical; inner tooth near cercus apex, robust, recurved and slightly incurve, apical part suddenly narrowly, apex acutely (Fig. 7 AB). Subgenital plate length as long as wide, with lateral carinae, middle part of posterior margin with a triangular excision, styli slender and short (Fig. 7C). Titillators with short apical arms, dorsally bearing one row of denticles; basal arms broad, with a right-angle curve on middle part (Fig. 7E). Female. Tegmina slightly shorter than half pronotum, extending to 1 th abdominal tergite anterior margin, clearly visible and discontiguous in dorsal view (Fig. 6 CJ). Fore tibiae dorsally armed 3 external spines and internally unarmed. Middle tibiae dorsally with 4 internal spines (one at apical) and 2 external spines. Cercus conical and hirsute (Fig. 7F). 10th abdominal tergite partly fused with epiproct, posterior margin of 10th abdominal tergite with a broadly round notch at the joint (Fig. 7F). Subgenital plate wider than length, middle of posterior margin with a deep angle concave (Fig. 7G). Ovipositor shorter than hind femora, distinctly upcurved (Fig. 7H). Coloration. Body laterally and dorsally generally dark brown, with whitish gray and dark green marmoration, ventrally mint green. Lower and hind margin of pronotum lateral lobes whitish gray with dark brown spots. Tegmina orange. Spines of each leg black. Male cercus inner tooth black. Ovipositor apical part black. Measurements(mm). Body (from head to tip of abdomen): ♂ 17.47, ♀ 20.60; pronotum: ♂ 5.04, ♀ 5.66; tegmina: ♂ 5.17, ♀ 2.73; mirror of right tegmina (from fore to hind): ♂ 2.66; fore femora: ♂ 6.00, ♀ 6.57; middle femora: ♂ 6.56, ♀ 7.45; hind femora: ♂ 17.60, ♀ 20.54; fore tibiae: ♂ 6.90, ♀ 7.66; middle tibiae: ♂ 7.62, ♀ 8.20; hind tibiae: ♂ 17.60, ♀ 20.54; male titillators (straight-line distance from base to end): 2.24; ovipositor: 12.67. Material examined. Holotype: ♂, Inside crevices on rock cliff of Baishi Mountain, Baishishan town, Laiyuan country, Baoding city, Hebei Province, China (114°41’41.56”E, 39°12‘16.94”N, alt. 1600m), 11-viii-2019, leg. Michael Lee. Paratype: 1♀, same data as in holotype. Remarks. The new species differ from Ptosoproctus lanzhouensis sp. nov. by: stridulatory file on male tegmina with less teeth; male 10th abdominal tergite posterior membranous projection strongly down-folded, with a gourdshaped median longitudinal furrow; male cercus inner tooth recurved and slightly incurve, apical part suddenly narrowly; male subgenital plate with a deeper triangular excision; titillators with longer apical arms, basal arms nearly right-angle curve; female ovipositor short and slightly curved, upper edge of apical part without wrinkles. Songs. The adults make calling songs at night. The song consists of echemes with syllables. Each echeme consist three syllables, echeme duration for 110–190 ms, echeme interval for 420–470 ms. Each syllable duration for 10–20 ms. Short syllables interval last for 10–15 ms. Etymology. The specific epithet is named after Baishi Mountain, the type locality. Distribution. China (Hebei). Known only from the type locality. Biology. This species inhabits Mt. Baishi’s steep marble cliff-face at around 1600m altitude. Type materials are collected from crevices on the cliff face, of which their body color well camouflaged against. A final-instar female nymph was also found (Fig. 9F). The collector returned to Mt. Baishi in mid-September 2021 to collect more material but returned in no avail, suggesting adult of the species may be short-lived due to Mt. Baishi’s low temperature., Published as part of Shen, Zi-Hao, Yin, Zi-Xu, Lee, Michael, Liu, Yi-Jiao, He, Zhu-Qing, Wang, Zi-Fan & Wang, Tong-Xin, 2021, Ptosoproctus gen. nov., a new genus with two new species of Shield-back Katydid with the first record of genus Eulithoxenus Bey-Bienko, 1951 from China (Orthoptera: Tettigoniidae: Tettigoniinae: Drymadusini), pp. 548-568 in Zootaxa 5067 (4) on pages 555-559, DOI: 10.11646/zootaxa.5067.4.4, http://zenodo.org/record/5683472

Ptosoproctus baishishanicus Shen, Yin, Lee & He sp. nov. 白石山折尾ỡ Figs. 1, 6–9, 14–15 Description. General. Body small. Head frons flat. Fastigium of vertex distinctly wider than scape of antenna (Fig. 6 AB). Head dorsally with an obscure and continuous median carina. Eyes broadly round and projecting outward. Disc of pronotum both sides with a broadly obtuse concavity on middle part, anterior margin of pronotum slightly concaved and posterior margin nearly straight, median carina continuous and distinctly indicated, central transvers sulcus distinctly Chinese characters “八”-shaped, lateral carina faintly indicated; lateral lobes longer than high, with weakly humeral sinus (Fig. 6 BC). Prosternum unarmed (Fig. 6D). Mesosternum with a pair of obtuse triangular lobe-shaped processes, wider than long (Fig. 6D). Metasternum with a pair of obtuse triangular lobe-shaped processes, extremely wider than long (Fig. 6D). Dorsal surface of fore coxae with 1 long spine; fore femora 1.15–1.2 times longer than pronotum, dorsally unarmed, ventrally with 0–2 internal spinules and externally unarmed, genicular lobes with 1–2 internal spinules and 1 external spinule; fore tibiae both sides with a crack-shape tympana, ventrally with 6 pair of spines, a pair at the apical. Middle femora dorsally unarmed, ventrally both sides with 0–1 spinules, genicular lobes with 1 internal spinules and 1–3 external spinules; middle tibiae ventrally with 6 pair of spines, a pair at the apical. Hind femora dorsally unarmed, ventrally with 0–4 spinules on both sides, genicular lobes with 0–1 spinules on both sides; hind tibiae dorsally with 18–27 spines on both sides and 1 pair of apical spurs, ventrally with 9–10 spines on both sides and 2 pair of apical spurs. Male. Tegmina brachypterous, slightly longer than pronotum, extending to 2 th abdominal tergite posterior margin. Left tegmina ventrally with stridulatory file about 36 teeth (Fig. 6G). Right tegmina ventrally with vestigial stridulatory file about 28 teeth (Fig. 6H). Mirror on right tegmina slightly longer than wide (Fig. 6 FH). Hind wings rudimentary (Fig. 6I). Fore tibiae dorsally armed 2–3 external spines and internally unarmed. Middle tibiae dorsally with 2 external spines and 1–2 internal spines (one at apical). 10 th abdominal tergite fused with epiproct, posterior margin of 10th abdominal tergite with a broadly triangle notch at the joint, form a down-folded membranous projection on tergite posterior margin, projection with a gourd-shaped median longitudinal furrow, apex broadly rounded. Cercus conical, pileous, short and stout, external margin straight; apex short and obtusely conical; inner tooth near cercus apex, robust, recurved and slightly incurve, apical part suddenly narrowly, apex acutely (Fig. 7 AB). Subgenital plate length as long as wide, with lateral carinae, middle part of posterior margin with a triangular excision, styli slender and short (Fig. 7C). Titillators with short apical arms, dorsally bearing one row of denticles; basal arms broad, with a right-angle curve on middle part (Fig. 7E). Female. Tegmina slightly shorter than half pronotum, extending to 1 th abdominal tergite anterior margin, clearly visible and discontiguous in dorsal view (Fig. 6 CJ). Fore tibiae dorsally armed 3 external spines and internally unarmed. Middle tibiae dorsally with 4 internal spines (one at apical) and 2 external spines. Cercus conical and hirsute (Fig. 7F). 10th abdominal tergite partly fused with epiproct, posterior margin of 10th abdominal tergite with a broadly round notch at the joint (Fig. 7F). Subgenital plate wider than length, middle of posterior margin with a deep angle concave (Fig. 7G). Ovipositor shorter than hind femora, distinctly upcurved (Fig. 7H). Coloration. Body laterally and dorsally generally dark brown, with whitish gray and dark green marmoration, ventrally mint green. Lower and hind margin of pronotum lateral lobes whitish gray with dark brown spots. Tegmina orange. Spines of each leg black. Male cercus inner tooth black. Ovipositor apical part black. Measurements(mm). Body (from head to tip of abdomen): ♂ 17.47, ♀ 20.60; pronotum: ♂ 5.04, ♀ 5.66; tegmina: ♂ 5.17, ♀ 2.73; mirror of right tegmina (from fore to hind): ♂ 2.66; fore femora: ♂ 6.00, ♀ 6.57; middle femora: ♂ 6.56, ♀ 7.45; hind femora: ♂ 17.60, ♀ 20.54; fore tibiae: ♂ 6.90, ♀ 7.66; middle tibiae: ♂ 7.62, ♀ 8.20; hind tibiae: ♂ 17.60, ♀ 20.54; male titillators (straight-line distance from base to end): 2.24; ovipositor: 12.67. Material examined. Holotype: ♂, Inside crevices on rock cliff of Baishi Mountain, Baishishan town, Laiyuan country, Baoding city, Hebei Province, China (114°41’41.56”E, 39°12‘16.94”N, alt. 1600m), 11-viii-2019, leg. Michael Lee. Paratype: 1♀, same data as in holotype. Remarks. The new species differ from Ptosoproctus lanzhouensis sp. nov. by: stridulatory file on male tegmina with less teeth; male 10th abdominal tergite posterior membranous projection strongly down-folded, with a gourdshaped median longitudinal furrow; male cercus inner tooth recurved and slightly incurve, apical part suddenly narrowly; male subgenital plate with a deeper triangular excision; titillators with longer apical arms, basal arms nearly right-angle curve; female ovipositor short and slightly curved, upper edge of apical part without wrinkles. Songs. The adults make calling songs at night. The song consists of echemes with syllables. Each echeme consist three syllables, echeme duration for 110–190 ms, echeme interval for 420–470 ms. Each syllable duration for 10–20 ms. Short syllables interval last for 10–15 ms. Etymology. The specific epithet is named after Baishi Mountain, the type locality. Distribution. China (Hebei). Known only from the type locality. Biology. This species inhabits Mt. Baishi’s steep marble cliff-face at around 1600m altitude. Type materials are collected from crevices on the cliff face, of which their body color well camouflaged against. A final-instar female nymph was also found (Fig. 9F). The collector returned to Mt. Baishi in mid-September 2021 to collect more material but returned in no avail, suggesting adult of the species may be short-lived due to Mt. Baishi’s low temperature.

Eulithoxenus emeljanovi Mishchenko, 1968, New record for China 埃氏优岩ỡ Figs. 1, 10–15 Eulithoxenus emeljanovi Mishchenko, 1968: 295 Redescription. General. Body small. Head frons flat, slightly oblique (Fig. 10A). Fastigium of vertex distinctly wider than scape of antenna. Eyes broadly round and projecting outward. Disc of pronotum both sides with a broadly obtuse concavity on middle part, anterior and posterior margin of pronotum slightly concaved, median carina and posterior transvers sulcus obscure, central transvers sulcus distinctly Chinese characters “八”-shaped, lateral carina faintly indicated; lateral lobes longer than high, with weakly humeral sinus (Fig. 10 BC). Prosternum unarmed (Fig. 10D). Mesosternum with a pair of obtuse triangular lobe-shaped processes, wider than long (Fig. 10D). Metasternum with a pair of obtuse triangular lobe-shaped processes, wider than long (Fig. 10D). Dorsal surface of fore coxae with 1 long spine; length of fore femora about 0.87–0.97 times shorter than pronotum, dorsally and ventrally unarmed, genicular lobes with 0–2 internal spinules and 0–1 external spinules; fore tibiae both sides with a crackshape tympana, dorsally with 3 externally spines and internally unarmed, ventrally with 6 pair of spines, a pair at the apical. Middle femora dorsally and ventrally unarmed, genicular lobes with 1–2 spinules on both sides; middle tibiae dorsally with 4 internal spines and 1–2 external spines, ventrally with 6 pair of spines, a pair at the apical. Hind femora dorsally and ventrally unarmed, genicular lobes with 0–1 spinules on both sides; hind tibiae dorsally with 24–27 spines on both sides and 1 pair of apical spurs, ventrally with 6–10 spines on both sides and 2 pair of apical spurs. Male. Tegmina brachypterous, about 0.6–0.7 times shorter than pronotum, extending to 2 th abdominal tergite middle part. Left tegmina ventrally with stridulatory file about 35 teeth (Fig. 10J). Right tegmina ventrally with vestigial stridulatory file about 30 teeth (Fig. 10L). Mirror on right tegmina pentagonal, slight longer than wide (Fig. 10 KL). Hind wings rudimentary (Fig. 10F). Middle part of 10 th abdominal tergite with a pair of obtuse lobes on posterior margin, furrow between lobes narrowly. Epiproct hidden under 10 th abdominal tergite lobes. Cercus hirsute, broad and flat, subbasal part with a raised ruffle; apex short and obtuse; inner margin most part with a broad inner tooth; inner tooth lager and wide, strongly recurved; inner margin near straight, outer margin arcuate; apical part suddenly narrow with an acute apex. Subgenital plate length slightly shorter than width, with lateral carinae, middle part of posterior margin with a triangular excision, styli short (Fig. 11C). Titillators broad at basal arms; apical arms outer margin bearing two rows denticles, with obtuse apex (Fig. 11E). Female. Tegmina brachypterous, extending to 1 th abdominal tergite anterior margin, almost invisible in dorsal view (Fig. 10 GH). Cercus conical and pileous. Posterior margin of 10 th abdominal tergite with a median narrow notch (Fig. 11F). 7 th abdominal sternite with a strong round hump. Subgenital plate wider than length, with a median hourglass hump, middle of posterior margin with a deep angle concave (Fig. 11G). Ovipositor shorter than hind femora, distinctly upcurved (Fig. 11H). Coloration. Body laterally and dorsally generally whitish gray and yellowish, with dark brown or black marmoration, ventrally whitish. Eyes white with dark brown spots. Lower and hind margin of pronotum lateral lobes whitish gray with dark brown spots. Tegmina orange with dark brown spots. Spines of fore and middle leg black, hind leg spines black at apex, hind femora middle part with a black spot on both sides. Male cercus inner tooth black. Ovipositor apical part black. Variations. Very few individual bodies laterally and dorsally generally reddish orange, except the black spot on hind femora middle part, without distinctly spots. Measurements(mm). Body (from head to tip of abdomen): ♂ 17.43–18.55, ♀ 20.54; pronotum: ♂ 4.50–4.78, ♀ 5.15; tegmina: ♂ 2.90–3.37, ♀ 0.88; mirror of right tegmina (from fore to hind): ♂ 1.93; fore femora: ♂ 4.00–4.30, ♀ 4.50; middle femora: ♂ 4.76–5.12, ♀ 5.50; hind femora: ♂ 16.74, ♀ 18.28; fore tibiae: ♂ 5.15–5.40, ♀ 6.12; middle tibiae: ♂ 5.60–6.50, ♀ 6.73; hind tibiae: ♂ 15.20–15.76, ♀ 17.11; male titillators (straight-line distance from base to end): 0.88; ovipositor: 12.88. Material examined. 2♂ 1♀, Beiwudang National Park, Dawukou District, Shizuishan city, Ningxia Hui Autonomous Region, China (106°19'52.70"E, 39°3'7.48"N, alt. 1223m), 7-vii-2020, leg. Tong Lou. Remarks. Eulithoxenus emeljanovi differs from Eulithoxenus mongolica (Uvarov, 1928) by: posterior margin of male 10 th abdominal tergite with a pair of obtuse lobes, which have an extremely narrow furrow; male cercus broad and flat, apex short and obtuse, inner margin most part with a broad inner tooth; posterior margin of female 7 th abdominal sternite without notch. Based on the photos of type specimen in OSF and description of E. mongolica (Cigliano et al. 2021; Storozhenko 2004; Uvarov 1928), male 10 th abdominal tergite concave widely and without lobes; male cercus slightly incurved, inner tooth near the apex, short and acute, directed obliquely towards the base; posterior margin of female 7 th abdominal sternite with a notch. We concluded that the two species may not belong to the same genus, and furthermore specimens need to be checked to find out their relationships. Songs. The adults make calling songs at night. The song consists of echemes with syllables. Each echeme consist two syllables, echeme duration for 80–100 ms, echeme interval for 240–300 ms. Each syllable duration for 30–50 ms, syllables interval for 10–20 ms. Distribution. Mongolia; Tuva; China (Ningxia)., Published as part of Shen, Zi-Hao, Yin, Zi-Xu, Lee, Michael, Liu, Yi-Jiao, He, Zhu-Qing, Wang, Zi-Fan & Wang, Tong-Xin, 2021, Ptosoproctus gen. nov., a new genus with two new species of Shield-back Katydid with the first record of genus Eulithoxenus Bey-Bienko, 1951 from China (Orthoptera: Tettigoniidae: Tettigoniinae: Drymadusini), pp. 548-568 in Zootaxa 5067 (4) on pages 559-565, DOI: 10.11646/zootaxa.5067.4.4, http://zenodo.org/record/5683472, {"references":["Uvarov, B. P. (1928) Orthoptera Palaearctica critica VI. Genus Bergiola Stschelk. (Tettig.). Eos, Revista Espanola de Entomologia, 4 (2), 243 - 251.","Storozhenko, S. Y. (2004) Long-horned orthopterans (Orthoptera: Ensifera) of the Asiatic part of Russia. Dalnauka, Vladivostok, 280 pp. [in Russian]"]}

Genus Rhicnogryllus Chopard, 1925 Type species: Rhicnogryllus fascipes Chopard, 1925, Published as part of Liu, Yi-Jiao & He, Zhu-Qing, 2021, Two new species of the genus Rhicnogryllus Chopard, 1925 from Hainan, China (Orthoptera: Trigonidiidae: Trigonidiinae), pp. 594-598 in Zootaxa 5048 (4) on page 595, DOI: 10.11646/zootaxa.5048.4.8, http://zenodo.org/record/5556843, {"references":["Chopard, L. (1925) Descriptions de Gryllides nouveaux (Orthopteres). Annales de la Societe Entomologique de France, 94, 291 - 332."]}

Rhicnogryllus xuandan He sp. nov. ������fldz�� Fig. 1A&B, 2A���F Holotype: male, CHINA, Hainan, Baoting, Xian���anshilin (E 109.43 N 18.60), 22-iii-2019, coll. He Zhu-Qing. Paratypes: 4 males and 7 females, same data as holotype. COI gene sequences: OK047283 & OK047284 from the holotype and one paratype. Description. Male. Head: little wider than pronotum, vertex not flattened, with a few strong long setae, frontal rostrum almost as wide as antennal scape, apical segment of maxillary palpi truncated (Fig. 2C). Pronotum: pubescent, anterior margin almost straight while posterior margin little convex, legs pubescent (Fig. 2A), tympanum absent, hind tibia with three pairs of dorsal spurs and three pairs of apical spurs (outer ones small with the middle one longest, dorsal and middle inner ones long). Tegmen coriaceous, almost extending to apex of abdomen. Genitalia: ectophallic apodeme slender and shaped into acute angle, ramus straight, pseudepiphallus separated into two lateral parts joined by a straight wide but feebly sclerotized bridge, posterior apex of pseudepiphallus forming flattened lophi with serrated inner margins, pseudepiphallic parameres triangular, nearly 2/3 of epiphallus apical lobes but wider and smoother than pseudepiphallus (Fig. 2D, E). Female. Similar to male (Fig. 2B), ovipositor little curved (Fig. 2F), finely denticulate on dorsal side (1/2 of total length) and ventral side (1/5 of total length). Coloration. Overall light yellow and black. Brownish red in head and anterior part of pronotum. Cerci yellow and ovipositor dark brown in ventral side. Measurement (mm). Male: SZ 4.60���5.16, PR 0.96���1.07, HF 4.55���4.61; Female: SZ 4.62���5.07, PR 0.82��� 1.02, HF 4.09���4.31, OV 1.83���1.98. Distribution. China (Hainan). Etymology. The specific epithet xuandan is for the Chinese phonetic alphabet ������, which is for black and red color, respectively., Published as part of Liu, Yi-Jiao & He, Zhu-Qing, 2021, Two new species of the genus Rhicnogryllus Chopard, 1925 from Hainan, China (Orthoptera: Trigonidiidae: Trigonidiinae), pp. 594-598 in Zootaxa 5048 (4) on pages 595-596, DOI: 10.11646/zootaxa.5048.4.8, http://zenodo.org/record/5556843

Rhicnogryllus fallax He sp. nov. ffi���fldz�� Fig. 1C&D, 3A���F Holotype: male, CHINA, Hainan, Jianfengling (E 109.27 N 22.86), 9-viii-2018, coll. He Zhu-Qing. Paratypes: 6 males and 5 females, same data as holotype. COI gene sequences: OK047279, OK047280, OK047281 & OK0472 from the holotype and 3 paratypes. Description. Male. This species has many characteristics in common with R. xuandan (Fig. 3A, C). The characteristics below are evidently distinct. Pronotum: pronotum slenderer rather than with more obvious spots derived from shed setae. Genitalia: ectophallic apodeme and ramus are similar to Rhicnogryllus xuandan, joined by an arc-shaped and obviously sclerotized bridge, pseudepiphallus separated into two lateral parts and posterior apex of pseudepiphallus forming feebly sclerotized lophi with dentated inner margins, pseudepiphallic parameres triangular (Fig. 3D, E). Female. Similar to male (Fig. 3B), ovipositor little curved (Fig. 3F), finely denticulate on both sides. Coloration. Overall body colour light yellow appearing glossy red in dorsal view. Basal part of pronotum brownish red, other parts pale yellow with red parallel stripes. Cerci yellow. Ovipositor dark brown ventrally. Measurement (mm). Male: SZ 4.17���4.31, PR 0.82���1.05, HF 3.52���3.77; Female: SZ 4.57���5.49, PR 0.82��� 1.02, HF 3.85���4.58, OV 1.77���1.83. Distribution. China (Hainan). Etymology. The new specific name is derived from Latin fallax, meaning very similar to above species., Published as part of Liu, Yi-Jiao & He, Zhu-Qing, 2021, Two new species of the genus Rhicnogryllus Chopard, 1925 from Hainan, China (Orthoptera: Trigonidiidae: Trigonidiinae), pp. 594-598 in Zootaxa 5048 (4) on pages 596-597, DOI: 10.11646/zootaxa.5048.4.8, http://zenodo.org/record/5556843

Loxoblemmus (Asonicogryllus) bronzus He sp. nov. ffiů濯Dz Figs 1–5 Holotype: male, CHINA, Yunnan, Dehong, Ruili (E 97.86 N 24.02), 26-iv-2018, coll. He Zhu-Qing. Paratypes: 3 females & 2 males, same data as holotype; 1 female, CHINA, Yunnan, Dehong, Longchuan (E 97.79 N 24.18), Alt. 1600 m, 10-vi-2018, coll. Bi Wen-Xuan. Description. Male. Head: shining, vertex little protruding, frontal rostrum flat, 2 times as wide as the 1st antennal joint, median ocellus lower than lateral ocelli, maxillary palpi normal (Fig. 2 CDE); Thorax: pronotum setose, anterior and posterior margin almost straight, without border between disc and lateral lobes, one oval tympanum on outside of foretibia and one small tympanum on inner side (Fig. 2 FG); forewings well developed, 2–2.5 times as long as pronotum (Fig. A). Hind tibia with 5 pairs of dorsal spines, three inner apical spurs and three outer spurs (Fig. 2 H), epiproct normal (Fig. 2 I). Genitalia: posterior epiphallic lobes elongate and apically little curved upwards, mesal posterior epiphallus protruding but little concave in middle, posterior edge of ectoparameron large and swollen; as long as posterior epiphallic lobes (Fig. 3 ABC). Female. Similar to male; forewing 1.5 times the length of pronotum (Fig. 2 B), ovipositor long and straight (Fig. 2 J). Coloration. Hatchlings: head light yellow and antennae white (Fig. 4 A). Nymphs: posterior part of head with a white stripe, antennae white with black apex (Fig. 4 B). Adults: almost black, forewing brown (Fig. 4 CD). Etymology. The name bronzus is used to emphasize the color of its forewings. Measurements (in mm). Male: body length 12.02–13.06, pronotum length 2.59–2.88, hind femur length 6.80–7.40, forewing length 6.01–6.38 Female: body length 11.87–13.32, pronotum length 2.54–2.80, hind femur length 6.74–7.05, forewing length 3.57–3.63, ovipositor length 6.61–7.17. Song. The song rate is about 98 pulse groups/minute. One pulse group consists of 9.029±1.243 pulses. The duration of pulse group is 0.419± 0.068s. The interval of pulse group is 0.205± 0.032s, as in Fig. 5. Distribution. China (Yunnan)., Published as part of Xu, Jing-Yi, Liu, Yi-Jiao, Li, Shi-Yu & He, Zhu-Qing, 2021, A new species of subgenus Asonicogryllus He, 2019 from Yunnan (Orthoptera: Gryllidae: Gryllinae), pp. 238-246 in Zootaxa 5040 (2) on pages 242-245, DOI: 10.11646/zootaxa.5040.2.4, http://zenodo.org/record/5531315

genus Orophyllus Beier, 1954 Type species: Orophyllus montanus Beier, 1954 Generic diagnosis. Medium to large size. Vertex wide and slightly extended forward. Fastigium of vertex tapered, with a feeble medial longitudinal sulcus. Anterior margin of pronotum is slightly arcuate, posterior margin broadly rounded, with two transverse grooves. Posterior transverse groove located behind the middle. The lateral lobes higher than width. Prosternum unarmed, meso-and metasternum strongly transverse, the former with entire, smooth anterior and lateral margin. Sc and R separated from the base of the tegmina, the Rs emerges from the proximal part of the R. The hind wings shorter than tegmina. Each leg armed on the internal ventral margin of the femur. Tympana similar on both sides of fore tibiae. Cerci relatively straight, curved inwards dorsally. Apex of male subgenital plate stalk-shaped, with long styli. Ovipositor saber-shaped, with oblique folds at the distal end of the lateral surface of the dorsal valve. Key to species of Orophyllus 1 Internal surface of hind femur without stripes............................................................... 2 - Lower internal portion of hind femur with a longitudinal patch................................................. 3 2 Distance from head to the apex of tegmen less than 47 mm, the length of ovipositor less than 20mm.......... O. montanus - Distance from head to the end of tegmen more than 52 mm, the length of ovipositor more than 28mm.................................................................................................. O. grandivalvatus sp. nov. 3 Subgenital plate unicolourous....................................................... O. nigrisartorius sp. nov. - Subgenital plate bearing markings........................................................................ 4 4 Subgenital plate with 2 black stripes on both sides of base............................ O. supeciliarilamellatus sp. nov. - Subgenital plate basally with a triangular marking............................................ O. guttatus sp. nov., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Li, Kai & He, Zhu-Qing, 2021, Taxonomy of genus Orophyllus Beier, 1954 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 147-159 in Zootaxa 4990 (1) on page 150, DOI: 10.11646/zootaxa.4990.1.9, http://zenodo.org/record/4983016, {"references":["Beier, M. (1954) Revision der Pseudophyllinen. Instituto EspanTol de Entomologiia, Madrid, 479 pp."]}

Orophyllus nigrisartorius Xie, Wang & He sp. nov. Figs. 3A–J Material examined: Holotype: 1 male, CHINA, Guizhou Prov., Leigongshan (108.18°E, 26.37°N), alt 1500m, 14- viii-2020, coll. Xie Hui-Cong (ECNU). Paratype: 2 females, CHINA, Guizhou Prov., Leigongshan (108.18°E, 26.37°N), alt 1500m, 14-viii-2020, coll. Xie Hui-Cong (ECNU); 4 males, CHINA, Sichuan Prov., Mount Emei (103.40°E, 29.57°N), alt 700m, 2–4-viii-2007, coll. Liu, Huang, Zhou & Bi (SEM); 1 male, CHINA, Sichuan Prov., Qingchengshan (103.55°E, 30.90°N), 7-v- 1985, coll. Liu Xian-Wei (SEM). Description: Male: Medium size, larger than O. montanus. Fastigium of vertex tapered, slightly exceeding the base of antennal socket, with a longitudinal groove posteriorly. Frons relatively flat and nearly square (Fig. 3A). Anterior margin of pronotum slightly convex, posterior margin broad and rounded; surface of disk almost smooth, with 2 transverse grooves, posterior transverse groove positioned behind the middle (Figs. 3B, C). The anterior margin of tegmina arched while posterior margin nearly straight, the length from Rs to R greater than to M of the tegmina, and the length from M to Cu as same as to Rs. The middle portion of Sc close to R, and the tip of tegmina blunt. Hind wings shorter than tegmina and transparent. Fore coxae with a spine. Fore femur longer than pronotum. Both dorsal carinae unarmed. Fore femur ventrally with 4–7 inner spines, mid femur ventrally with 6–9 inner spines, hind femur ventrally with 9–13 inner spines. Hind femur with black stripe on internal surface (Fig. 3F). Prosternum unarmed, meso- and metasternum strongly transverse, the former with entire, non-crenellated anterior margin (Fig. 3D). Epiproct ovoid, length almost equal to width, slightly concave apically. Cerci relatively longer and slender, longer than epiproct, and curved inwards apically. Subgenital plate gradually narrows toward the apex, base with membranous part and gaped in dry specimen, the tip stalk-shaped. Styli very long, wide at the base, narrowing towards the end, and pointed in terminal (Figs. 3G, H). Female: Much larger than the male. Tegmina slightly recurved at apex. Epiproct long tongue-shaped or rectangular, nearly 1.5 times as long as wide. Cerci elongate and conical, curved inwards apically. Ovipositor robust, longer than length of hind femur, with a curved ventral edge, the apical area with 4 fine, sharp angled folds. Dorsal margin of the ovipositor not as straight as in O. montanus, basal half of the dorsal valve straight, the dorsal margin of the apical half narrowred toward the ventral surface; distal half of dorsal margin armed with fine teeth (Fig. 3J). The basal panels of subgenital plate expanded at inner apex to triangular lobules, the apical lobe of the subgenital plate with a narrow but deep notch (Fig. 3I). Measurements (in mm): Male: SZ 44.3–52.28, PR 6.2–7.15, FW 36.5–42.64, HF 16.9–19.95; Female: SZ 69.27–71.94, PR 8.93–9.13, FW 58.87–60.39, HF 24.64–24.65, OV 33.36–34.26. Etymology: Species name nigrisartorius means a black stripe inside the hind femur, its position just like the sartorius muscle to human thigh. Distribution: Guizhou, Sichuan., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Li, Kai & He, Zhu-Qing, 2021, Taxonomy of genus Orophyllus Beier, 1954 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 147-159 in Zootaxa 4990 (1) on pages 152-153, DOI: 10.11646/zootaxa.4990.1.9, http://zenodo.org/record/4983016

Orophyllus supeciliarilamellatus Xie, Wang & He sp. nov. Figs. 4A–I Material examined: Holotype: 1 male, CHINA, Guangxi Prov., Maoershan (110.48°E, 25.88°N), alt 1200m, 11-viii-2020 coll. Xie Hui-Cong (ECNU). Paratype: 2 males, CHINA, Guangxi Prov., Jinxiu (110.25°E, 24.16°N), 19-vii-2015, coll. He Zhu- Qing (ECNU); 2 males and 3 females, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 18-vii-2015, coll. Lin & Zhu (SEM); 4 males, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 20-vii-2015, coll. Lin & Zhu (SEM); 2 males, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 21-vii-2015, coll. Lin & Zhu (SEM); 1 female, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 22-vii-2015, coll. Lin & Zhu (SEM); 2 females, CHINA, Guangxi Prov., Damingshan, alt. 1250m, 19~ 25-vii-2013, coll. Zhu, Liu, Wang & Zhang (SEM); 3 males, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), 6-viii-2020, coll. Xie Hui-Cong (ECNU); 3 males, CHINA, Sichuan Prov., Mengdingshan (103.03°E, 30.06°N), alt 1456m, 31-vii–1-viii-2007, coll. Liu, Huang, Zhou & Bi (SEM); 1 female, CHINA, Guizhou Prov., Fanjingshan, alt. 1000~ 1800m, 6-viii-2014, coll. Sun & Li (SEM). Description. Very similar to preceding species but smaller, tegmina shorter and broader, not recurved as preceding species at apex in female (Fig. 8). Male cerci thicker, and styli only about half length of subgenital plate which bearing eyebrow-shaped marking basally (Fig. 4F, G). The basal panels of female subgenital plate each with a blunt conical tubercle at outer apex (Fig. 4H). Ovipositor shorter (Fig. 4I). Measurements (in mm): Male: SZ 43.10–51.57, PR 6.27–8.20, FW 35.56–41.91, HF 17.09–19.56. Female: SZ 53.92–58.15, PR 6.93–8.03, FW 48.25–49.53, HF 19.54–21.25, OV 23.56–25.52. Etymology: Species name supeciliarilamellatus means 2 stripes on the ventral surface of the subgenital plate. Distribution: Guangxi, Guizhou, Jiangxi, Sichuan.

Orophyllus supeciliarilamellatus Xie, Wang & He sp. nov. Figs. 4A–I Material examined: Holotype: 1 male, CHINA, Guangxi Prov., Maoershan (110.48°E, 25.88°N), alt 1200m, 11-viii-2020 coll. Xie Hui-Cong (ECNU). Paratype: 2 males, CHINA, Guangxi Prov., Jinxiu (110.25°E, 24.16°N), 19-vii-2015, coll. He Zhu- Qing (ECNU); 2 males and 3 females, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 18-vii-2015, coll. Lin & Zhu (SEM); 4 males, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 20-vii-2015, coll. Lin & Zhu (SEM); 2 males, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 21-vii-2015, coll. Lin & Zhu (SEM); 1 female, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), alt 1200m, 22-vii-2015, coll. Lin & Zhu (SEM); 2 females, CHINA, Guangxi Prov., Damingshan, alt. 1250m, 19~ 25-vii-2013, coll. Zhu, Liu, Wang & Zhang (SEM); 3 males, CHINA, Guangxi Prov., Yangmeiao (108.64°E, 25.19°N), 6-viii-2020, coll. Xie Hui-Cong (ECNU); 3 males, CHINA, Sichuan Prov., Mengdingshan (103.03°E, 30.06°N), alt 1456m, 31-vii–1-viii-2007, coll. Liu, Huang, Zhou & Bi (SEM); 1 female, CHINA, Guizhou Prov., Fanjingshan, alt. 1000~ 1800m, 6-viii-2014, coll. Sun & Li (SEM). Description. Very similar to preceding species but smaller, tegmina shorter and broader, not recurved as preceding species at apex in female (Fig. 8). Male cerci thicker, and styli only about half length of subgenital plate which bearing eyebrow-shaped marking basally (Fig. 4F, G). The basal panels of female subgenital plate each with a blunt conical tubercle at outer apex (Fig. 4H). Ovipositor shorter (Fig. 4I). Measurements (in mm): Male: SZ 43.10–51.57, PR 6.27–8.20, FW 35.56–41.91, HF 17.09–19.56. Female: SZ 53.92–58.15, PR 6.93–8.03, FW 48.25–49.53, HF 19.54–21.25, OV 23.56–25.52. Etymology: Species name supeciliarilamellatus means 2 stripes on the ventral surface of the subgenital plate. Distribution: Guangxi, Guizhou, Jiangxi, Sichuan., Published as part of Xie, Hui-Cong, Wang, Han-Qiang, Li, Kai & He, Zhu-Qing, 2021, Taxonomy of genus Orophyllus Beier, 1954 (Orthoptera: Tettigoniidae: Pseudophyllinae), pp. 147-159 in Zootaxa 4990 (1) on page 153, DOI: 10.11646/zootaxa.4990.1.9, http://zenodo.org/record/4983016