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Sohail, Kamran, Huang, Weijian, Zhang, Yalin (2020): A new species of the genus Kodaianellissus Wang, Bourgoin & Zhang (Hemiptera: Issidae) with morphological and molecular data. Zootaxa 4731 (2): 270-274, DOI: https://doi.org/10.11646/zootaxa.4731.2.6

Hemisphaeriinae Melichar, 1906 Kodaianellini Wang, Zhang & Bourgoin, 2016, Published as part of Sohail, Kamran, Huang, Weijian & Zhang, Yalin, 2020, A new species of the genus Kodaianellissus Wang, Bourgoin & Zhang (Hemiptera: Issidae) with morphological and molecular data, pp. 270-274 in Zootaxa 4731 (2) on page 271, DOI: 10.11646/zootaxa.4731.2.6, http://zenodo.org/record/3637887, {"references":["Melichar, L. (1906) Monographie der Issiden. (Homoptera). Abhandlungen der Zoologisch-botanischen Gesellschaft in Wien, 3, 327.","Wang, M. L., Zhang, Y. L. & Bourgoin, T. (2016) Planthopper family Issidae (Insecta: Hemiptera: Fulgoromorpha): linking molecular phylogeny with classification. Molecular Phylogenetics and Evolution, 105, 224 - 234. https: // doi. org / 10.1016 / j. ympev. 2016.08.012"]}

Kodaianellissus gibbusis Kamran, Huang et Zhang, sp. nov. Description: Length of male (from apex of head to tip of forewing, N=3) 5.5���6.0 mm. Vertex and frons yellowish brown, anterior margin of frons black with small yellow spots, lateral margins yellowish with brown markings. Clypeus yellowish. Eyes greyish with tanned bands. Gena and antennae brown. Pronotum yellowish, mesonotum light brown. Abdomen dark brown dorsally. Tegmina, legs and longitudinal veins brown, cross veins yellowish. Disc of vertex rectangular, 1.9 times wider than long, centrally and laterally carinate, two rounded markings near lateral margins. Posterior margins concave, anterior margins slightly curved in mid line at apex, lateral margins slightly transverse (Fig. 1D). Frons slightly globose, broader behind the eyes (Fig. 1C), median carina obvious, lateral margins slightly straight, concavely truncated behind eyes, obliquely narrow near clypeus (Fig. 1C). Clypeus with distinct brown transverse strips (Fig. 1C). Large compound eyes (Figs. 1C, D). Gena smooth, oblique in lateral view (Fig. 1B). Pronotum longer than vertex centrally, anterior margin convex, strongly carinate; posterior margin truncate, central carina yellowish, two black incisions near median carina on the disk (Fig. 1D). Mesonotum as long as pronotum, two brown spots near lateral margins and near base on each side of the disc, posterior one obviously smaller than apical one. Tegmina, distinctly longer than wide, ScP+R vein forked near base, ScP+RA vein long reaching ⅓ costal margin, RP vein very long, exceeding 1/3 of costa distad at outer margin of tegmina. MP vein bifurcate near basal �� long surpassing, MP 1+2 and MP 3+4 sinuate at apical margin. Cu forked at apical ⅓, Pcu and A 1 combine at basal half of clavus (Fig. 1E). Hind wing fully developed tri-lobed. Lobe Pcu-A 1 distinctly longer than wide; A 2 lobe narrower distad at ⅓ of Pcu-A 1, A 2 lobe simple, longer not forked (Fig. 1F). Hind tibiae with two lateral spines, spinal formula 8-7-2. Male genitalia. Anal segment in dorsal view (Fig. 1H) rectangular, longer than wide, laterally straight, slightly convex at basal half, anal foramen short, located in basal half (Fig. 1H). Anal tube in lateral view long, convex at posterior ⅓, ventral margins slightly prominent (Fig. 1G). Pygofer in lateral view, long and slender with parallel anterior and posterior margins (Fig. 1G). Genital styles rectangular, widest at basal ⅓, caudoventrally rectangular slightly curved near base of capitilum at posterior margin, dorsolaterally provided with hump-like processes near base of capitulum (Fig. 1J). Capitilum very long, tubular, apically sclerotized slightly bent directed anteriorly, posterior margin with lobe-like process (Fig. 1J). Aedeagus in lateral view elevated at both sides, divided into two lobes, dorsolateral lobe slightly membranous in apical portion, centrally convex. Sharp sclerotized spines directed dorsad on each side of ventral apical part (Fig. 1K). In ventral view, periandrium is long U-shaped, somewhat symmetrical, dorsolateral margins twisted in distal half, pair of long hooks arising in distal half directed downwards (Fig. 1I). Type materials: Holotype: ♂ Rawlakot, Azad Jammu & Kashmir, Pakistan, 33��51���32.18���N, 73�� 45���34.93���E, 1628 m, 14-vii-2018, coll. Kamran Sohail. Paratypes 2♂♂, same data as holotype. Female: Unknown. Etymology: Latin word ��� gibbus ��� means hump, referring to hump-like processes near base of capitilum on dorsolateral side. Host plants: Unknown. Diagnosis: Kodaianellissus gibbusis sp. nov. can be differentiated from K. intorqueus by characters mentioned in the key. Remarks: Phylogenetic analyses of 5 species of Hemisphaeriinae, including two out-groups (Gelastyrella litaoensis and Darwallia barbata), based on ML and BI analyses of nucleotide sequence data of two genes, yielded almost same topologies with strong support (Fig. 2). The molecular analyses (Fig. 2) with morphological characters indicates its placement into the Kodaianellissus., Published as part of Sohail, Kamran, Huang, Weijian & Zhang, Yalin, 2020, A new species of the genus Kodaianellissus Wang, Bourgoin & Zhang (Hemiptera: Issidae) with morphological and molecular data, pp. 270-274 in Zootaxa 4731 (2) on pages 271-273, DOI: 10.11646/zootaxa.4731.2.6, http://zenodo.org/record/3637887

Key to species of Kodaianellissus 1. Male anal segment in dorsal view with straight apical and dorsolateral margins; aedeagus with pair of hooks directed downwards in ventral view........................................................ Kodaianellissus gibbusis sp. nov. - Male anal segment in dorsal view slightly convex in apical margin, laterally with two convex angles; aedeagus with pair of hooks directed outwards in ventral view (Wang et al., 2017, figs 4, 8, 11).................... Kodaianellissus intorqueus, Published as part of Sohail, Kamran, Huang, Weijian & Zhang, Yalin, 2020, A new species of the genus Kodaianellissus Wang, Bourgoin & Zhang (Hemiptera: Issidae) with morphological and molecular data, pp. 270-274 in Zootaxa 4731 (2) on page 271, DOI: 10.11646/zootaxa.4731.2.6, http://zenodo.org/record/3637887, {"references":["Wang, M. L., Bourgoin, T. & Zhang, Y. L. (2017) New Oriental genera in the family Issidae (Hemiptera: Fulgoromorpha). Zootaxa, 4312 (2), 355 - 367. https: // doi. org / 10.11646 / zootaxa. 4312.2.10"]}

Genus Destinoides Cai & He Destinoides Cai & He, 2000: 57 Type species: Destinoides fasciata Cai & He, by original designation (= D. latifrons (Walker)). Diagnosis. Destinoides is similar to Petalocephala and Destinia but can be distinguished by: crown shorter, about half as long as wide, longer than pronotum in the middle (crown of Destinia shorter than pronotum); body more robust; pronotum without lateral extensions (Tituria, Thlasia, Dusuna and Neotituria with lateral extensions); pronotum not declivous or humped; lateral margins of crown immediately in front of eyes usually parallel to convergent; scutellum longer than pronotum (similar to Destinia, scutellum of Petalocephala usually shorter than pronotum); pygofer without process on upper hind angle (Destinia with a horizontal process); style slender and flat (style of Destinia thick), curved ventrally at apex, with a small subapical tooth laterally (style of Petalocephala without any tooth). Description. Body medium to large (9–15mm) and robust. Lateral margins of head and pronotum dark. Dorsum densely punctate, acinose (Figs. 1–6). Head about as wide as pronotum, flat; longer than anterior margin of scutellum. Crown smooth, slightly convex, produced forward, anterior margin angulate, setae absent. Coronal suture complete and distinct (Figs. 1, 4). Ocelli welldeveloped, near posterior margin of crown, equidistant from eye and each other (Figs. 1, 4). Head approximately as long as half width between eyes, concave in lateral view (Figs. 1, 2, 4, 5). Clypeal suture distinct. Frontoclypeus narrow and convex, broadened medially; frontoclypeus apical extension not well-defined. Anteclypeus small, distinctly convex, parallel-sided and extending beyond gena. Lora and genae flat (Figs. 3, 6). Antennae slightly longer than eye, extended nearly to margin of face; antennal ledges not distinct (Figs. 3, 6). Proepisternum large and quadrate (Figs. 3, 6). Pronotum broad and flat, anterior margin slightly convex, posterior margin slightly concave (Figs. 1–6). Scutellum triangular, shorter than head. Scutellar suture distinct, reaching lateral margin (Figs. 1, 4). Forewing punctate, venation regular in basal area, reticulate apically; appendix reduced (Figs. 2, 5). Hind wings hyaline, fully-developed. Pro- and mesothoracic leg chaetotaxy reduced. Hind femur macrosetal formula 2 + 1; hind tibia with 5 to 8 stout and enlarged setae perpendicular to axis of leg anterodorsally (AD); posterodorsal (PD) setae short and fine; anteroventral (AV) and posteroventral (PV) setae shorter and finer. Male 8th sternite about as long as 7th sternite, posterior margin strongly concave medially (Figs. 13, 20). Male pygofer almost triangular, without macrosetae; ventroposterior margin with dorsally directed process (Figs. 10, 17). Subgenital plates ligulate and depressed, fused at base, tapered to rounded apex, with numerous short setae on ventral surface (Figs. 11, 18). Aedeagal shaft tubular or slightly lamellate, curved dorsally, generally with single and paired processes arising terminally (Figs. 7, 8, 14, 15). Style slender and flat, curved ventrally at apex, with a small subapical tooth laterally (Figs. 7, 8, 14, 15). Connective short, T-shaped (Figs. 7–9, 14–16). Distribution. China, India., Published as part of Huang, Weijian & Zhang, Yalin, 2019, Two new species in the genus Destinoides (Hemiptera: Cicadellidae, Ledrinae) from China, with DNA barcoding data, pp. 571-578 in Zootaxa 4711 (3) on pages 572-573, DOI: 10.11646/zootaxa.4711.3.8, http://zenodo.org/record/3586697, {"references":["Cai, P. & He, J. H. (2000) A new record and a new allied genera of Ledrinae (Homoptera: Cicadellidae) from China. In: Zhang, Y. L. (Ed.), Systematic and Faunistic Research on Chinese Insects. Proceedings of the 5 th National Congress of Insect Taxonomy. China Agriculture Press, Beijing, pp. 56 - 60."]}

Destinoides latifrons (Walker 1851) Ledra latifrons Walker, 1851: 824 Petalocephala latifrons, Stål, 1862: 494 Destinoides latifrons, Sun, Webb & Zhang, 2014: 80 Destinoides fasciata Cai & He, 2000: 57. synonymized by Sun, Webb & Zhang, 2014: 80 Distribution. China (Xizang), Bengal, India (Assam), Sikkim., Published as part of Huang, Weijian & Zhang, Yalin, 2019, Two new species in the genus Destinoides (Hemiptera: Cicadellidae, Ledrinae) from China, with DNA barcoding data, pp. 571-578 in Zootaxa 4711 (3) on page 572, DOI: 10.11646/zootaxa.4711.3.8, http://zenodo.org/record/3586697, {"references":["Walker, F. (1851) List of the Specimens of Homopterous Insects in the Collection of the British Museum. Part 3. Printed by Order of the Trustees, London, pp. 637 - 907.","Stal, C. (1862). Synonymiska och systematiska anteckningar ofver Hemiptera. Ofversigt af Kongliga Vetenskaps-Akadmiens forhandlingar, 19, 479 - 504.","Sun, J., Webb, M. D. & Zhang, Y. L. (2014) Revision of the Oriental leafhopper genus Destinoides Cai & He (Hemiptera: Cicadellidae: Ledrinae) with a new synonym and two new combinations. Zootaxa, 3786 (1), 79 - 84. https: // doi. org / 10.11646 / zootaxa. 3786.1.6","Cai, P. & He, J. H. (2000) A new record and a new allied genera of Ledrinae (Homoptera: Cicadellidae) from China. In: Zhang, Y. L. (Ed.), Systematic and Faunistic Research on Chinese Insects. Proceedings of the 5 th National Congress of Insect Taxonomy. China Agriculture Press, Beijing, pp. 56 - 60."]}

Genus Destinoides Cai & He Destinoides Cai & He, 2000: 57 Type species: Destinoides fasciata Cai & He, by original designation (= D. latifrons (Walker)). Diagnosis. Destinoides is similar to Petalocephala and Destinia but can be distinguished by: crown shorter, about half as long as wide, longer than pronotum in the middle (crown of Destinia shorter than pronotum); body more robust; pronotum without lateral extensions (Tituria, Thlasia, Dusuna and Neotituria with lateral extensions); pronotum not declivous or humped; lateral margins of crown immediately in front of eyes usually parallel to convergent; scutellum longer than pronotum (similar to Destinia, scutellum of Petalocephala usually shorter than pronotum); pygofer without process on upper hind angle (Destinia with a horizontal process); style slender and flat (style of Destinia thick), curved ventrally at apex, with a small subapical tooth laterally (style of Petalocephala without any tooth). Description. Body medium to large (9–15mm) and robust. Lateral margins of head and pronotum dark. Dorsum densely punctate, acinose (Figs. 1–6). Head about as wide as pronotum, flat; longer than anterior margin of scutellum. Crown smooth, slightly convex, produced forward, anterior margin angulate, setae absent. Coronal suture complete and distinct (Figs. 1, 4). Ocelli welldeveloped, near posterior margin of crown, equidistant from eye and each other (Figs. 1, 4). Head approximately as long as half width between eyes, concave in lateral view (Figs. 1, 2, 4, 5). Clypeal suture distinct. Frontoclypeus narrow and convex, broadened medially; frontoclypeus apical extension not well-defined. Anteclypeus small, distinctly convex, parallel-sided and extending beyond gena. Lora and genae flat (Figs. 3, 6). Antennae slightly longer than eye, extended nearly to margin of face; antennal ledges not distinct (Figs. 3, 6). Proepisternum large and quadrate (Figs. 3, 6). Pronotum broad and flat, anterior margin slightly convex, posterior margin slightly concave (Figs. 1–6). Scutellum triangular, shorter than head. Scutellar suture distinct, reaching lateral margin (Figs. 1, 4). Forewing punctate, venation regular in basal area, reticulate apically; appendix reduced (Figs. 2, 5). Hind wings hyaline, fully-developed. Pro- and mesothoracic leg chaetotaxy reduced. Hind femur macrosetal formula 2 + 1; hind tibia with 5 to 8 stout and enlarged setae perpendicular to axis of leg anterodorsally (AD); posterodorsal (PD) setae short and fine; anteroventral (AV) and posteroventral (PV) setae shorter and finer. Male 8th sternite about as long as 7th sternite, posterior margin strongly concave medially (Figs. 13, 20). Male pygofer almost triangular, without macrosetae; ventroposterior margin with dorsally directed process (Figs. 10, 17). Subgenital plates ligulate and depressed, fused at base, tapered to rounded apex, with numerous short setae on ventral surface (Figs. 11, 18). Aedeagal shaft tubular or slightly lamellate, curved dorsally, generally with single and paired processes arising terminally (Figs. 7, 8, 14, 15). Style slender and flat, curved ventrally at apex, with a small subapical tooth laterally (Figs. 7, 8, 14, 15). Connective short, T-shaped (Figs. 7–9, 14–16). Distribution. China, India.

Destinoides conspicua (Distant 1907) Petalocephala conspicua Distant, 1907: 189 Destinoides latifrons (Walker), misidentification, Cai & He, 2000: 58 Destinoides conspicua, Sun, Webb & Zhang, 2014: 82 Distribution. China (Hainan; Guangxi; Yunnan), Vietnam, Singapore., Published as part of Huang, Weijian & Zhang, Yalin, 2019, Two new species in the genus Destinoides (Hemiptera: Cicadellidae, Ledrinae) from China, with DNA barcoding data, pp. 571-578 in Zootaxa 4711 (3) on page 572, DOI: 10.11646/zootaxa.4711.3.8, http://zenodo.org/record/3586697, {"references":["Distant, W. L. (1907) Contributions to a knowledge of the Ledrinae. Annales de la Societe Entomologique de Belgique, 51, 185 - 197.","Cai, P. & He, J. H. (2000) A new record and a new allied genera of Ledrinae (Homoptera: Cicadellidae) from China. In: Zhang, Y. L. (Ed.), Systematic and Faunistic Research on Chinese Insects. Proceedings of the 5 th National Congress of Insect Taxonomy. China Agriculture Press, Beijing, pp. 56 - 60.","Sun, J., Webb, M. D. & Zhang, Y. L. (2014) Revision of the Oriental leafhopper genus Destinoides Cai & He (Hemiptera: Cicadellidae: Ledrinae) with a new synonym and two new combinations. Zootaxa, 3786 (1), 79 - 84. https: // doi. org / 10.11646 / zootaxa. 3786.1.6"]}

Genus Eleazara Distant, new record to Malaysia Eleazara Distant, 1908: 182; Cai & He, 1997: 12 Type species: Eleazara aedificatura Distant, 1908 Diagnosis. The genus Eleazara is similar to Ledra Fabricius, 1803. It can be distinguished from the latter by: 1) pronotum longer than crown, with the lateral dilated areas laminately subangular (Figs. 1, 4); 2) scutellum with a sagittal elevated crest-like process (Figs. 1, 2); 3) the long metathoracic tarsomere I (Fig. 3); and 4) dorsum strongly granulose (Figs. 1���6). Description. Body medium to rather large (10.9���17.3mm). Dorsum densely punctate. Head broad, shorter and narrower than pronotum, longer than the width of the anterior margin of the scutellum. Crown foliaceous, punctate, nodulate, not rugose, slightly convex, produced forward, anterior margin angulate, straight in front of eyes and then oblique to apex, median length much greater than width between eyes (Figs. 1, 2, 5). Coronal suture complete and distinct (Figs. 1, 6). Ocelli well-developed, near posterior margin of crown, approximately as close to each other as to eyes (Figs. 1, 6). Face elongate, broadest between eyes. Proepisternum large and collar-like (Fig. 3). Pronotum broad, with paired sublateral longitudinal rows of tubercles and paired submedial carinae variably developed, strongly granulose on posterior area, anterior margin straight or slightly convex, posterior margin slightly or moderately concave (Figs. 1���4). Scutellum and exposed part of mesonotum triangular, shorter than pronotum, with sagittal elevated crest. Scutellar suture distinct, reaching lateral margin (Figs. 1, 2). Forewing granulose, venation reticulate, especially apically, veins elevated and prominent; appendix poorly developed (Figs. 1, 2). Hind wings hyaline, fully-developed. Pro- and mesothoracic leg chaetotaxy reduced, with setae short and fine. Legs of moderate length. Hind femur macrosetal formula 2 + 0; hind tibiae dilated, with 5 to 6 stout and enlarged setae rows perpendicular to axis of leg anterodorsally (AD); posterodorsal (PD) setae short and fine; anteroventral (AV) and posteroventral (PV) setae shorter and finer. Male 8th sternite about as long as or longer than 7th sternite, posterior margin moderately concave medially, with or without numerous short setae apically on ventral surface (Fig. 15). Male pygofer almost triangular, without terminal process; ventroposterior margin with terminally directed process (Figs. 7, 8, 12, 14). Subgenital plate ligulate and depressed, fused at base, tapered to rounded apex, with numerous short setae on ventral surface (Figs. 7, 8, 12, 13). Aedeagal shaft tubular, curved dorsad, with (Eleazara yangi Cai & He) or without (Eleazara viraktamathi Huang & Zhang sp. nov.) paired processes arising subapically (Figs. 7 ���10, 12). Style slender or broad, curved ventrally at apex (Figs. 9, 10). Connective short, T-shaped or trapeziform (Figs. 9���11). Distribution. China, India, Malaysia., Published as part of Huang, Weijian & Zhang, Yalin, 2018, First record of the genus Eleazara Distant (Hemiptera: Cicadellidae, Ledrinae) in Malaysia, with description of a new species, pp. 434-440 in Zootaxa 4532 (3) on page 435, DOI: 10.11646/zootaxa.4532.3.7, http://zenodo.org/record/2615515, {"references":["Distant, W. L. (1908) Rhynchota, Homoptera and Appendix (Pt.). In: Bingham, C. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Vol. IV. Taylor & Francis, London, pp. i-xv + 1 - 501.","Cai, P. & He, J. (1997) A new and a newly recorded genera of Ledridae from China, with descriptions of four new species. Wuyi Science Journal, 13, 8 - 14."]}

Eleazara yangi Cai & He Eleazara yangi Cai & He, 1997:12 Description. “Size medium. Body length 10.9 mm. The terminus of abdomen extending beyond forewing. Head length 1.4mm, approximately as long as half width between eyes. Coronal suture complete and distinct. Ocelli placed close to eye as long as half width between each other. Pronotum length 1.9mm, as long as half width. Head concave in front of eyes and then oblique to apex. Scutellum length slightly longer than head, with a central elevated anteriorly directed crest-like process. Anterior margin and posterior margin of forewing parallel. Hind tibiae long and slender, with 6 stout and enlarged setae in rows perpendicular to axis of leg anterodorsally (AD). Male 8th sternite longer than 7th sternite, with posterior margin concave. Subgenital plates salix leaf-like, rather long and narrow, with numerous short setae and numerous apically rusty brown spots. Style rather broad, curved ventrally at apex. Aedeagus long and slender, shaft with a pair of lamellate processes apically. Coloration isabelline; eyes and ocelli grayish brown; posterior area of pronotum gray; basal forewings fuscous; abdomen yellowish ventrally, rusty brown dorsally; legs yellowish-brown, terminal femur and basal tibiae fuscous.” (Cai & He, 1997) Distribution. China (Yunnan). Female. Unknown. Diagnosis. This species resembles E. viraktamathi Huang & Zhang sp. nov. but differs in having an isabelline colored body, a tubular aedeagal shaft, a robust style and pygofer side in lateral aspect tapering posteriorly without posterior process.

Family Cicadellidae Subfamily Ledrinae Tribe Ledrini

Eleazara viraktamathi Huang & Zhang sp. nov. (Figs. 1���4, 9���15) Description. Size rather large. Body length (incl. forewing) 17.3 mm, head width (incl. eyes) 4.1mm, pronotum width 5.2mm, scutellum width 3.3mm. Coloration rusty. Dorsum strongly granulose (Figs. 1���6). Crown not rugose, transverse camber curved in anterior view (Figs. 5, 6). Median longitudinal carina present. Eyes slate-colored, kidney-shaped. Ocelli dusky; protuberance laterad of ocelli present (Fig. 6). Facial setae numerous throughout. Dark facial pigmentation absent (Figs. 3, 5). Clypeal suture distinct. Frontoclypeus long, narrow and convex, broadened medially, without median longitudinal carina and groove; frontoclypeus base with lateral perpendicular ridges; frontoclypeus apical extension well defined. Anteclypeus small, parallel-sided and extended beyond gena. Lora and genae flattish (Fig. 3). Antennae longer than length of eye, apex extending beyond margin of face; antennal ledges distinct (Fig. 3). Proepisternum large and collar-like (Fig. 3). Pronotum flat anteriorly and abruptly declivous posteriorly, with setae ventrally (Fig. 2���5); pronotal tubercules present (Fig. 4). Scutellum with central elevated crest-like process and pair of lateral crestlike processes (Figs. 1, 2). Forewing punctate, granulose, venation coriaceous in basal two-thirds, highly reticulate and hyaline in apically; appendix vanishing (Figs. 1, 2). Pro- and mesothoracic leg chaetotaxy reduced, with fine setae. Hind tibiae extremely dilated, with 5 stout and enlarged setae anterodorsally (AD). Metathoracic tarsomere I columnar, more robust compared with other tarsomeres, in ventral view longer in length with remaining tarsomeres combined (Fig. 3). Male pygofer narrow, without terminal process, with medial short and fine setae (Figs. 8, 12, 14). Male 8th sternite nearly longer than 7th sternite, with posterior margin concave (Fig. 15). Pygofer side in lateral aspect tapering posteriorly with posterior long hook-like process, beyond pygofer (Fig. 14). Style slim, widest medially, acuminate apically, curved ventrally at apex, with 4 setae on the tortuous area (Figs. 9, 10). Aedeagus curved dorsally, shaft slightly lamellate with a pair of lamellate processes medially (Figs. 7, 8). Connective flat, short and wide, trapeziform (Figs. 9���11). Material examined. Holotype: male, Malaysia, Sabah, 20 August 2016, Yisheng Zhao (NWAFU). Female. Unknown. Etymology. The new species is named in honor of Chandra A Viraktamath (University of Agricultural Sciences, Bangalore, Bengalur) and his great contributions to leafhopper taxonomy. Diagnosis. This species resembles E. yangi Cai & He but differs in having a rusty body, a slightly lamellate aedeagal shaft, a slender style and pygofer side in lateral aspect tapering posteriorly with posterior long hook-like process., Published as part of Huang, Weijian & Zhang, Yalin, 2018, First record of the genus Eleazara Distant (Hemiptera: Cicadellidae, Ledrinae) in Malaysia, with description of a new species, pp. 434-440 in Zootaxa 4532 (3) on pages 436-438, DOI: 10.11646/zootaxa.4532.3.7, http://zenodo.org/record/2615515

Family Cicadellidae Subfamily Ledrinae Tribe Ledrini, Published as part of Huang, Weijian & Zhang, Yalin, 2018, First record of the genus Eleazara Distant (Hemiptera: Cicadellidae, Ledrinae) in Malaysia, with description of a new species, pp. 434-440 in Zootaxa 4532 (3) on page 435, DOI: 10.11646/zootaxa.4532.3.7, http://zenodo.org/record/2615515

Eleazara yangi Cai & He Eleazara yangi Cai & He, 1997:12 Description. ���Size medium. Body length 10.9 mm. The terminus of abdomen extending beyond forewing. Head length 1.4mm, approximately as long as half width between eyes. Coronal suture complete and distinct. Ocelli placed close to eye as long as half width between each other. Pronotum length 1.9mm, as long as half width. Head concave in front of eyes and then oblique to apex. Scutellum length slightly longer than head, with a central elevated anteriorly directed crest-like process. Anterior margin and posterior margin of forewing parallel. Hind tibiae long and slender, with 6 stout and enlarged setae in rows perpendicular to axis of leg anterodorsally (AD). Male 8th sternite longer than 7th sternite, with posterior margin concave. Subgenital plates salix leaf-like, rather long and narrow, with numerous short setae and numerous apically rusty brown spots. Style rather broad, curved ventrally at apex. Aedeagus long and slender, shaft with a pair of lamellate processes apically. Coloration isabelline; eyes and ocelli grayish brown; posterior area of pronotum gray; basal forewings fuscous; abdomen yellowish ventrally, rusty brown dorsally; legs yellowish-brown, terminal femur and basal tibiae fuscous.��� (Cai & He, 1997) Distribution. China (Yunnan). Female. Unknown. Diagnosis. This species resembles E. viraktamathi Huang & Zhang sp. nov. but differs in having an isabelline colored body, a tubular aedeagal shaft, a robust style and pygofer side in lateral aspect tapering posteriorly without posterior process., Published as part of Huang, Weijian & Zhang, Yalin, 2018, First record of the genus Eleazara Distant (Hemiptera: Cicadellidae, Ledrinae) in Malaysia, with description of a new species, pp. 434-440 in Zootaxa 4532 (3) on page 436, DOI: 10.11646/zootaxa.4532.3.7, http://zenodo.org/record/2615515, {"references":["Cai, P. & He, J. (1997) A new and a newly recorded genera of Ledridae from China, with descriptions of four new species. Wuyi Science Journal, 13, 8 - 14."]}

Elongationa gen. nov. Type Species: Elongationa hyalina sp. nov., here designated. Description. Body large, rufous (Figs. 1, 12). Head narrower than pronotum. Eyes small. Crown triangular, spatulate with apex slightly upturned, longer than width between eyes; coronal suture indistinct, with elevated ochraceous ridge along midline. (Figs. 1, 13). Anterolateral portions of face without setae. Ocelli near posterior margin of crown, nearer to each other than to corresponding eye (Figs. 1, 13). Face distinctly longer than wide (Figs.4, 16); clypeal suture distinct; frontoclypeus narrow, convex apically; anteclypeus broadened medially; genae flat; lora broad; lateral margins between lora/ genae and frontoclypeus distinctly depressed (Figs. 4, 16); antennae short; antennal ledges not distinct (Figs. 4, 16). Pronotum quadrangular, anterior margin slightly convex, lateral margins oblique, slightly divergent posteriorly, posterior margin sinuate, humeral lobe extended only slightly laterad of forewing base in dorsal view (Figs. 1, 2, 13, 14). Proepisternum large and quadrate (Figs. 4, 16). Scutellum triangular (Figs. 1, 2, 13, 14). Fore wing extending beyond apex of abdomen, veins reticulate apically, clavus densely punctate (Figs. 2, 14). Hind wings hyaline, with well defined veins(Figs. 3, 15). Legs short. Fore femur with a few setae. Middle tibia with numerous fine setae. Hind femur with three stout macrosetae at apex. Hind tibiae flattened with five or six stout anterodorsal (AD) setae with spine-like bases; posterodorsal (PD) setae fine; anteroventral (AV) and posteroventral (PV) setae shorter and finer. Metathoracic tarsomere I columnar, more robust than other tarsomeres, as long or slightly longer than remaining tarsomeres combined in ventral view. Male 8th sternite slightly longer than 7th sternite, posterior margin convex and slightly concave medially (Fig. 7). Male pygofer long, abruptly narrowing from dorsal margin at midlength; dorsal margin sinuate; ventroposterior margin with dorsally directed process (Fig. 6, 12). Subgenital plates ligulate, fused at base, tapered distally to rounded apex; with numerous short setae on ventral surface and with row of slightly stouter setae near apical margin (Fig. 10). Aedeagal shaft depressed, curved dorsally, with paired processes arising near base (Figs. 8, 9). Style elongate, apex footlike (Figs. 9). Connective T-shaped (Fig. 9, 11). Female with 1st and 2nd valvulae narrow, curved dorsally; 1st strigate with sculpture apically, 2nd with several small dorsal subapical teeth (Figs. 17, 18, 19, 20). Distribution. China (Sichuan Province and Hainan Province). Etymology. The name of this new genus refers to the prolonged head. Remarks. This new genus is close to the genus Ledropsis White but can be distinguished from the latter by: 1) the shape of head triangulate, not infundibulate (Figs. 1, 13); 2) head upturned apically (Figs. 2, 14); 3) pronotum punctate, not tuberculate (Figs. 1, 13)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on pages 361-362, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728

Midoria emmrichi (Zhang & Yang, 2004) Thlasia emmrichi Zhang & Yang, 2004: 85 ���86, figs. 12���19 Midoria emmrichi (Zhang & Yang, 2004) Li & Li, 2010: 64, figs. 1���3 Distribution. China., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 367, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Zhang, Y. L., Yang, L. H., Webb, M. D. & Sun, Q. (2004) Type specimens of Jacobi's Fukien leafhoppers and description of three new species from China (Insecta: Auchenorrhyncha: Cicadellidae: Ledrinae: Penthimiinae). Entomologische Abhandluagen (Dresden), 62, 83 - 92.","Li, Y. J. & Li, Z. Z. (2010) A taxonomic study of the genus of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) with descriptions of two new species from China. Zootaxa, 2451, 63 - 68."]}

Elongationa hyalina sp. nov. (Figs 1–12) Description. Male. Length (incl. forewing) 13 mm; head width (incl. eyes) 3.2 mm; pronotum width 3.2 mm. Body generally rufous, more yellowish ventrally (Fig. 1). Crown and apical half of face, dark brown, lateral margins of crown basally and pronotum edged yellowish white (Fig. 1). Fore wing with basal costal margin edged with dark brown basally; elongate hyaline band medially from base to apex of wing (Fig. 2). Pronotum slightly depressed laterally. Male pygofer with dorsal edge sinuate and long finger-like process apically (Fig. 6). Style apex triangular laterally (Fig. 9). Aedeagus flat, curved dorsally and tapering apically, with pair of long spinelike processes basally, second pair of lamelliform footlike processes medially (Figs. 8, 9); gonopore apical on dorsal surface (Fig. 8, 9). Type material. Holotype: male, CHINA, Hainan Prov., Mt. Jianfengling, 1100m, 20 March 1980, Wang Shiyong (IZCAS). Etymology. The specific epithet refers to the hyaline fore wing band.

Midoria emmrichi (Zhang & Yang, 2004) Thlasia emmrichi Zhang & Yang, 2004: 85 –86, figs. 12–19 Midoria emmrichi (Zhang & Yang, 2004) Li & Li, 2010: 64, figs. 1–3 Distribution. China.

Midoria hastifera Li & Li, 2011 Midoria hastifera Li & Li, 2011: 63, figs. 21–24, 33, 36 Distribution. China (Yunnan).

Midoria hamulata Li & Li, 2011 Midoria hamulata Li & Li, 2011: 60, figs. 9–12, 27, 30 Distribution. China (Yunnan).

Midoria lamellata Li & Li, 2010 Midoria lamellata Li & Li, 2010: 67, figs. 19���24, 26 Distribution. China (Guizhou)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 368, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2010) A taxonomic study of the genus of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) with descriptions of two new species from China. Zootaxa, 2451, 63 - 68."]}

Midoria curvidentata sp. nov. (Figs. 21���30) Description. Male. Length (incl. forewing) 6.9 mm; head width (incl. eyes) 2.2 mm; pronotum width 2.3 mm. Body yellowish brown, apical half of crown dark brown and anterior edge of pronotum black (Fig. 21). Fore wing with some black spots on veins and yellowish brown band (Fig. 21). Male pygofer with long spinelike process ventroapically (Fig. 27). Style parallel-sided through most of length, apex bent mesad and acute (Fig. 29). Aedeagus compressed basally, curved dorsally and expanded gradually apically in lateral view, with pair of short processes dorsomedially and pair of long processes lateroapically (Figs. 26, 29, 30); gonopore apical on dorsal surface (Figs. 26, 29, 30). Type material. Holotype: male, CHINA, Shaanxi Prov., Ningshan County, Huoditang Forest Farm, 5 June 2009, Xie Sha (NWAFU). Paratype: 1 male, CHINA, Shaanxi Prov., Ningshan County, Huoditang Forest Farm, 12 May 2014, Li Qinglong (NWAFU). Etymology. The species epithet refers to the shape of the long processes on the subapical aedeagus., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 368, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728

Midoria zunyiensis Li & Li, 2011 Midoria zunyiensis Li & Li, 2011: 60, figs 9–12 Distribution. China (Guizhou). Material examined. 1 male, CHINA, Guizhou Prov., Dao Zhen, San Qiao Town, 31 May 2004, Yu Yang (NWAFU).

Elongationa sp. (Figs. 13–20) Description. Female. Body length (incl. forewing) 16.3 mm, head width 3.6 mm (incl. eyes), pronotum width 4 mm. Colour as in previous species but brown marking paler (Figs. 13, 14). Female with 1st and 2nd valvulae narrow, curved dorsally; 1st strigate apically and 2nd elevated medially with one large and three or four small dorsal subapical teeth (Figs. 17, 18, 19, 20). Material examined. 1 female, CHINA, Sichuan Prov., Mt. Emei, 800–1100m, 22 April 1957, Huang Keren (IZCAS). Remarks. We are uncertain if the specimen described represents the female of E. hyalina sp. nov. or is a different species. It has a longer head, paler brown marking and a shorter hyaline band on the forewing than in the male holotype of E. hyalina sp. nov.

Midoria capitata Kato, 1931 Midoria caPitata Kato, 1931: 439, fig. 1 Distribution. China (Taiwan)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 366, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Kato, M. (1931) Japanese Ledrinae. Journal of the Zoological Society of JaPan, 43, 431 - 440, figs. 1 - 10."]}

Midoria huapingensis Li & Li, 2010 Midoria huaPingensis Li & Li, 2010: 66 ���67, figs. 14���18, 25 Distribution. China (Guangxi). Material examined. 1 male, CHINA, Guangxi Prov., Jin Xiu, Mt. Tian Tang, 30 May 1982, Yang Jikun (NWAFU)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 367, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2010) A taxonomic study of the genus of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) with descriptions of two new species from China. Zootaxa, 2451, 63 - 68."]}

Midoria hei Cai & Jiang, 2000 Midoria hei Cai & Jiang, 2000: 414 ���415, figs. 1���8 Midoria hei, Li & Li, 2010: 66, figs. 12���13 Distribution. China (Yunnan)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 367, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Cai, P. & Jiang, J. F. (2000) Two new species of the genus Midoria Kato from China (Homoptera: Cicadellidae: Ledrinae). Acta Entomologica Sinica, 43, 413 - 416.","Li, Y. J. & Li, Z. Z. (2010) A taxonomic study of the genus of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) with descriptions of two new species from China. Zootaxa, 2451, 63 - 68."]}

Midoria deplanata Li & Li, 2011 Midoria dePlanata Li & Li, 2011: 58, figs. 5���8, 26, 29 Distribution. China (Guangxi). Material examined. 1 male, CHINA, Guangxi Prov., Jin Xiu, Jinzhong Road, 1100m, 12 May 1999, Xiao Hui (NWAFU)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on pages 366-367, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2011) Descriptions of six new species of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) from China with a key to the species of the genus. Zootaxa, 2897, 57 - 63."]}

Midoria curvidentata sp. nov. (Figs. 21–30) Description. Male. Length (incl. forewing) 6.9 mm; head width (incl. eyes) 2.2 mm; pronotum width 2.3 mm. Body yellowish brown, apical half of crown dark brown and anterior edge of pronotum black (Fig. 21). Fore wing with some black spots on veins and yellowish brown band (Fig. 21). Male pygofer with long spinelike process ventroapically (Fig. 27). Style parallel-sided through most of length, apex bent mesad and acute (Fig. 29). Aedeagus compressed basally, curved dorsally and expanded gradually apically in lateral view, with pair of short processes dorsomedially and pair of long processes lateroapically (Figs. 26, 29, 30); gonopore apical on dorsal surface (Figs. 26, 29, 30). Type material. Holotype: male, CHINA, Shaanxi Prov., Ningshan County, Huoditang Forest Farm, 5 June 2009, Xie Sha (NWAFU). Paratype: 1 male, CHINA, Shaanxi Prov., Ningshan County, Huoditang Forest Farm, 12 May 2014, Li Qinglong (NWAFU). Etymology. The species epithet refers to the shape of the long processes on the subapical aedeagus.

Midoria hastifera Li & Li, 2011 Midoria hastifera Li & Li, 2011: 63, figs. 21���24, 33, 36 Distribution. China (Yunnan)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 367, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2011) Descriptions of six new species of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) from China with a key to the species of the genus. Zootaxa, 2897, 57 - 63."]}

Midoria Kato Midoria Kato, 1931: 439 Type species: Midoria caPitata Kato, 1931 Description. Body small, brown (Fig. 21). Crown spatulate, shorter and slightly narrower than pronotum (Fig. 21). Anterolateral portions of face without setae. Ocelli nearer to each other than to corresponding eye (Fig. 21). Clypeal suture distinct, genae flat, lora broad, antennal ledges not distinct. Lateral margins between lora/genae and frontoclypeus distinctly depressed. Pronotum swelled postmedially, anterior margin slightly convex, lateral margins oblique, posterior margin sinuate (Fig. 21). Fore wing extending beyond apex of abdomen, clavus densely punctate (Fig. 21). Fore femur with few setae. Middle tibia with numerous fine setae. Hind tibiae flattened with three to five stout anterodorsal (AD) setae with spine-like bases; posterodorsal (PD), anteroventral (AV) and posteroventral (PV) setae short and fine. Male pygofer abroad; dorsal margin sinuate; ventroposterior margin with long process (Fig. 27). Subgenital plates legume-like, tapered distally to rounded apex (Fig. 25). Aedeagal shaft compressed, curved dorsally (Figs. 26, 29, 30). Style elongate (Figs. 29). Connective T-shaped (Fig. 30). Distribution. China (Shaanxi, Sichuan, Guangxi, Guizhou, Fujian, Guangdong, Taiwan, Yunnan). Remarks. This genus can be distinguished from the genus Petalocephala St��l by the smaller body and compressed aedeagal shaft., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 364, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Kato, M. (1931) Japanese Ledrinae. Journal of the Zoological Society of JaPan, 43, 431 - 440, figs. 1 - 10."]}

Midoria torsiva Li & Li, 2011 Midoria torsiva Li & Li, 2011: 60 Distribution. China (Yunnan)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 368, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2011) Descriptions of six new species of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) from China with a key to the species of the genus. Zootaxa, 2897, 57 - 63."]}

Midoria denticulata Li & Li, 2011 Midoria denticulata Li & Li, 2011: 58, figs 1���4, 25, 28 Distribution. China (Guizhou)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 366, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2011) Descriptions of six new species of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) from China with a key to the species of the genus. Zootaxa, 2897, 57 - 63."]}

Midoria zunyiensis Li & Li, 2011 Midoria zunyiensis Li & Li, 2011: 60, figs 9���12 Distribution. China (Guizhou). Material examined. 1 male, CHINA, Guizhou Prov., Dao Zhen, San Qiao Town, 31 May 2004, Yu Yang (NWAFU)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 368, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2011) Descriptions of six new species of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) from China with a key to the species of the genus. Zootaxa, 2897, 57 - 63."]}

Midoria annulata Cai & Jiang, 2000 Midoria annulata Cai & Jiang, 2000: 415, figs. 1���8 Midoria annulata Li & Li, 2010: 66, figs. 10���11, 28 Distribution. China (Shaanxi, Guizhou). Material examined. 1 male, CHINA, Shaanxi Prov., Shuang Shi Pu, 6 May 1980, Xiang Longcheng, Ma Ning (NWAFU); 3 males, CHINA, Shaanxi Prov., Shi Quan, 25 April 1980, Xiang Longcheng, Ma Ning (NWAFU); 1 male, CHINA, Shandong Prov., Mt. Tai, 30 May 1956, Wang Linyao (IZCAS)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 366, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Cai, P. & Jiang, J. F. (2000) Two new species of the genus Midoria Kato from China (Homoptera: Cicadellidae: Ledrinae). Acta Entomologica Sinica, 43, 413 - 416.","Li, Y. J. & Li, Z. Z. (2010) A taxonomic study of the genus of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) with descriptions of two new species from China. Zootaxa, 2451, 63 - 68."]}

Elongationa hyalina sp. nov. (Figs 1���12) Description. Male. Length (incl. forewing) 13 mm; head width (incl. eyes) 3.2 mm; pronotum width 3.2 mm. Body generally rufous, more yellowish ventrally (Fig. 1). Crown and apical half of face, dark brown, lateral margins of crown basally and pronotum edged yellowish white (Fig. 1). Fore wing with basal costal margin edged with dark brown basally; elongate hyaline band medially from base to apex of wing (Fig. 2). Pronotum slightly depressed laterally. Male pygofer with dorsal edge sinuate and long finger-like process apically (Fig. 6). Style apex triangular laterally (Fig. 9). Aedeagus flat, curved dorsally and tapering apically, with pair of long spinelike processes basally, second pair of lamelliform footlike processes medially (Figs. 8, 9); gonopore apical on dorsal surface (Fig. 8, 9). Type material. Holotype: male, CHINA, Hainan Prov., Mt. Jianfengling, 1100m, 20 March 1980, Wang Shiyong (IZCAS). Etymology. The specific epithet refers to the hyaline fore wing band., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 362, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728

Midoria annulata Cai & Jiang, 2000 Midoria annulata Cai & Jiang, 2000: 415, figs. 1–8 Midoria annulata Li & Li, 2010: 66, figs. 10–11, 28 Distribution. China (Shaanxi, Guizhou). Material examined. 1 male, CHINA, Shaanxi Prov., Shuang Shi Pu, 6 May 1980, Xiang Longcheng, Ma Ning (NWAFU); 3 males, CHINA, Shaanxi Prov., Shi Quan, 25 April 1980, Xiang Longcheng, Ma Ning (NWAFU); 1 male, CHINA, Shandong Prov., Mt. Tai, 30 May 1956, Wang Linyao (IZCAS).

Elongationa gen. nov. Type Species: Elongationa hyalina sp. nov., here designated. Description. Body large, rufous (Figs. 1, 12). Head narrower than pronotum. Eyes small. Crown triangular, spatulate with apex slightly upturned, longer than width between eyes; coronal suture indistinct, with elevated ochraceous ridge along midline. (Figs. 1, 13). Anterolateral portions of face without setae. Ocelli near posterior margin of crown, nearer to each other than to corresponding eye (Figs. 1, 13). Face distinctly longer than wide (Figs.4, 16); clypeal suture distinct; frontoclypeus narrow, convex apically; anteclypeus broadened medially; genae flat; lora broad; lateral margins between lora/ genae and frontoclypeus distinctly depressed (Figs. 4, 16); antennae short; antennal ledges not distinct (Figs. 4, 16). Pronotum quadrangular, anterior margin slightly convex, lateral margins oblique, slightly divergent posteriorly, posterior margin sinuate, humeral lobe extended only slightly laterad of forewing base in dorsal view (Figs. 1, 2, 13, 14). Proepisternum large and quadrate (Figs. 4, 16). Scutellum triangular (Figs. 1, 2, 13, 14). Fore wing extending beyond apex of abdomen, veins reticulate apically, clavus densely punctate (Figs. 2, 14). Hind wings hyaline, with well defined veins(Figs. 3, 15). Legs short. Fore femur with a few setae. Middle tibia with numerous fine setae. Hind femur with three stout macrosetae at apex. Hind tibiae flattened with five or six stout anterodorsal (AD) setae with spine-like bases; posterodorsal (PD) setae fine; anteroventral (AV) and posteroventral (PV) setae shorter and finer. Metathoracic tarsomere I columnar, more robust than other tarsomeres, as long or slightly longer than remaining tarsomeres combined in ventral view. Male 8th sternite slightly longer than 7th sternite, posterior margin convex and slightly concave medially (Fig. 7). Male pygofer long, abruptly narrowing from dorsal margin at midlength; dorsal margin sinuate; ventroposterior margin with dorsally directed process (Fig. 6, 12). Subgenital plates ligulate, fused at base, tapered distally to rounded apex; with numerous short setae on ventral surface and with row of slightly stouter setae near apical margin (Fig. 10). Aedeagal shaft depressed, curved dorsally, with paired processes arising near base (Figs. 8, 9). Style elongate, apex footlike (Figs. 9). Connective T-shaped (Fig. 9, 11). Female with 1st and 2nd valvulae narrow, curved dorsally; 1st strigate with sculpture apically, 2nd with several small dorsal subapical teeth (Figs. 17, 18, 19, 20). Distribution. China (Sichuan Province and Hainan Province). Etymology. The name of this new genus refers to the prolonged head. Remarks. This new genus is close to the genus Ledropsis White but can be distinguished from the latter by: 1) the shape of head triangulate, not infundibulate (Figs. 1, 13); 2) head upturned apically (Figs. 2, 14); 3) pronotum punctate, not tuberculate (Figs. 1, 13).

Midoria denticulata Li & Li, 2011 Midoria denticulata Li & Li, 2011: 58, figs 1–4, 25, 28 Distribution. China (Guizhou).

Midoria hei Cai & Jiang, 2000 Midoria hei Cai & Jiang, 2000: 414 –415, figs. 1–8 Midoria hei, Li & Li, 2010: 66, figs. 12–13 Distribution. China (Yunnan).

Key to species of Midoria (without M. capitata Kato) 1. Aedeagus expanded apically in lateral view (Fig. 22)...................................... M. curvidentata sp. nov. - Aedeagus not expanded apically in lateral view..............................................................2 2. Aedeagus with two pairs of processes...................................................................... 3 - Aedeagus with one pair of processes......................................................................4 3. Aedeagus with one pair of processes apically............................................. M. huapingensis Li & Li - Aedeagus with two pairs of processes apically................................................. M. hei Cai & Jiang 4. Aedeagus shaft with serrated processes dorsally............................................ M. denticulata Li & Li - Aedeagus shaft without serrated processes dorsally...........................................................5 5. Aedeagus shaft and lateral processes forming H-shaped structure in ventral view...................................6 - Aedeagus shaft and lateral processes not forming H-shaped structure in ventral view................................7 6. Style with single distal projection......................................................... M. hastifera Li & Li - Style with two distal projections.......................................................... M. hamulata Li & Li 7. Processes longer than aedeagus shaft in lateral view..........................................................8 - Processes not longer than aedeagus shaft in lateral view......................................................10 8. Processes on aedeagus curved toward each other.......................................... M. annulata Cai & Jiang - Processes on aedeagus not curved toward each other..........................................................9 9. Apex of processes on aedeagus hornlike...................................................... M. torsiva Li & Li - Apex of processes on aedeagus not hornlike......................................... M. emmrichi (Zhang & Yang) 10. Aedeagus with processes not furcate..................................................................... 11 - Aedeagus with processes furcate.........................................................................12 11. Processes on aedeagus separated from shaft........................................ M. longicornis (Zhang & Yang) - Processes on aedeagus not separated from shaft.............................................. M. deplanata Li & Li 12. Processes on aedeagus tapering in lateral view..............................................................13 - Processes on aedeagus not tapering in lateral view........................................... M. zunyiensis Li & Li 13. Apex of processes on aedeagus curved basally............................................... M. lamellata Li & Li - Apex of processes on aedeagus not curved basally............................................. M. funebris (Jacobi)

Key to species of Midoria (without M. capitata Kato) 1. Aedeagus expanded apically in lateral view (Fig. 22)...................................... M. curvidentata sp. nov. - Aedeagus not expanded apically in lateral view..............................................................2 2. Aedeagus with two pairs of processes...................................................................... 3 - Aedeagus with one pair of processes......................................................................4 3. Aedeagus with one pair of processes apically............................................. M. huapingensis Li & Li - Aedeagus with two pairs of processes apically................................................. M. hei Cai & Jiang 4. Aedeagus shaft with serrated processes dorsally............................................ M. denticulata Li & Li - Aedeagus shaft without serrated processes dorsally...........................................................5 5. Aedeagus shaft and lateral processes forming H-shaped structure in ventral view...................................6 - Aedeagus shaft and lateral processes not forming H-shaped structure in ventral view................................7 6. Style with single distal projection......................................................... M. hastifera Li & Li - Style with two distal projections.......................................................... M. hamulata Li & Li 7. Processes longer than aedeagus shaft in lateral view..........................................................8 - Processes not longer than aedeagus shaft in lateral view......................................................10 8. Processes on aedeagus curved toward each other.......................................... M. annulata Cai & Jiang - Processes on aedeagus not curved toward each other..........................................................9 9. Apex of processes on aedeagus hornlike...................................................... M. torsiva Li & Li - Apex of processes on aedeagus not hornlike......................................... M. emmrichi (Zhang & Yang) 10. Aedeagus with processes not furcate..................................................................... 11 - Aedeagus with processes furcate.........................................................................12 11. Processes on aedeagus separated from shaft........................................ M. longicornis (Zhang & Yang) - Processes on aedeagus not separated from shaft.............................................. M. deplanata Li & Li 12. Processes on aedeagus tapering in lateral view..............................................................13 - Processes on aedeagus not tapering in lateral view........................................... M. zunyiensis Li & Li 13. Apex of processes on aedeagus curved basally............................................... M. lamellata Li & Li - Apex of processes on aedeagus not curved basally............................................. M. funebris (Jacobi), Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 366, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728

Midoria lamellata Li & Li, 2010 Midoria lamellata Li & Li, 2010: 67, figs. 19–24, 26 Distribution. China (Guizhou).

Midoria deplanata Li & Li, 2011 Midoria dePlanata Li & Li, 2011: 58, figs. 5–8, 26, 29 Distribution. China (Guangxi). Material examined. 1 male, CHINA, Guangxi Prov., Jin Xiu, Jinzhong Road, 1100m, 12 May 1999, Xiao Hui (NWAFU).

Elongationa sp. (Figs. 13���20) Description. Female. Body length (incl. forewing) 16.3 mm, head width 3.6 mm (incl. eyes), pronotum width 4 mm. Colour as in previous species but brown marking paler (Figs. 13, 14). Female with 1st and 2nd valvulae narrow, curved dorsally; 1st strigate apically and 2nd elevated medially with one large and three or four small dorsal subapical teeth (Figs. 17, 18, 19, 20). Material examined. 1 female, CHINA, Sichuan Prov., Mt. Emei, 800���1100m, 22 April 1957, Huang Keren (IZCAS). Remarks. We are uncertain if the specimen described represents the female of E. hyalina sp. nov. or is a different species. It has a longer head, paler brown marking and a shorter hyaline band on the forewing than in the male holotype of E. hyalina sp. nov., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 364, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728

Midoria funebris (Jacobi, 1944) Tlasia [sic] funebris Jacobi, 1944: 40 Thlasia funebris, Zhang et al, 2004: 86, figs. 20���30 Midoria funebris (Jacobi, 1944) Li & Li, 2010: 65, figs. 7���9, 27 Distribution. China (Guangxi, Guizhou, Fujian). Material examined. 1 male, CHINA, Guangxi Prov., Tianlin, Liangzhuo, 30 May 1982, Yang Jikun (CAU); 1 male, CHINA, Fujian Prov., Mt. Wuyi, Da Zhu Lan, 19 May 2004, Yuan Caixia, Li Jing (HBU)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 367, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Jacobi, A. (1944) Die Zikadenfauna der Provinz Fukien in Sudchina und ihre tiergeographischen Beziehungen. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 5 - 66.","Zhang, Y. L., Yang, L. H., Webb, M. D. & Sun, Q. (2004) Type specimens of Jacobi's Fukien leafhoppers and description of three new species from China (Insecta: Auchenorrhyncha: Cicadellidae: Ledrinae: Penthimiinae). Entomologische Abhandluagen (Dresden), 62, 83 - 92.","Li, Y. J. & Li, Z. Z. (2010) A taxonomic study of the genus of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) with descriptions of two new species from China. Zootaxa, 2451, 63 - 68."]}

Midoria longicornis (Zhang & Yang, 2004) Thlasia longicornis Zhang & Yang, 2004: 87 ���88, figs. 40���47 Midoria longicornis (Zhang & Yang, 2004), Li & Li, 2010: 87 ���88, figs. 40���47 Distribution. China., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 368, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Zhang, Y. L., Yang, L. H., Webb, M. D. & Sun, Q. (2004) Type specimens of Jacobi's Fukien leafhoppers and description of three new species from China (Insecta: Auchenorrhyncha: Cicadellidae: Ledrinae: Penthimiinae). Entomologische Abhandluagen (Dresden), 62, 83 - 92.","Li, Y. J. & Li, Z. Z. (2010) A taxonomic study of the genus of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) with descriptions of two new species from China. Zootaxa, 2451, 63 - 68."]}

Midoria hamulata Li & Li, 2011 Midoria hamulata Li & Li, 2011: 60, figs. 9���12, 27, 30 Distribution. China (Yunnan)., Published as part of Sun, Jing, Huang, Weijian & Zhang, Yalin, 2017, Taxonomic studies of two genera, Elongationa gen. nov. and Midoria Kato (Hemiptera: Cicadellidae: Ledrinae), with two new species from China, pp. 361-370 in Zootaxa 4294 (3) on page 367, DOI: 10.11646/zootaxa.4294.3.6, http://zenodo.org/record/832728, {"references":["Li, Y. J. & Li, Z. Z. (2011) Descriptions of six new species of Midoria Kato (Hemiptera: Cicadellidae: Ledrinae) from China with a key to the species of the genus. Zootaxa, 2897, 57 - 63."]}