Search

Menoscelis Mulsant, 1850 Menoscelis Mulsant, 1850: 506, 508; Crotch, 1874: 209; Gemminger & Harold, 1876: 3783; Chapin, 1966: 279; El-Ali, 1972: 23; Gordon, 1987: 26; Mill��o et al., 2004: 395; Gordon & Canepari, 2008: 271 ���274. Menoscelis (Thalassa): Chapuis, 1876: 228, 233��� 234. Thalassa (Menoscelis): Korschefsky, 1931: 208; Blackwelder, 1945: 449; Duverger, 1989: 146; F��rsch, 1990 b: 63. Type species. Menoscelis saginata Mulsant, 1850, by subsequent designation of Crotch (1874). Diagnosis. Menoscelis is a Neotropical genus that is structurally similar to Diazonema Weise, 1926, but differs from all other genera of the tribe in the highly modified protibia, angulate near its base (Figs 5 D���E) and the transverse female genital plate with spoon���like setae near the apex. Diazonema presents a combination of short and compressed protibia, and a transverse female genital plate lacking spoon���like setae near the apex. In addition to the generic characters described by Gordon & Canepari (2008), species of this genus have other useful characters that aid in their recognition: body usually large (4.5���7.5 mm) and rounded, moderately to strongly convex. Dorsal surface glabrous or pubescent, maculate or not. Prosternum process about half the coxal diamenter, surface with complete or incomplete carinae joined anteriorly, forming a triangle (Figs 5 C, 6 E). Prosternal carinae often joined, sometimes connected to the anterior margin of the prosternum by a single carina (Figs 8 A���G, 9 A���E). Epipleura large, external margin explanate (Figs 5 H, 6 H). Protibia flattened, medially emarginate or with parallel sides, tibial groove deep, long, not flanged posteriorly, angulate near base (Figs 5 D���E, 6 F���G); meso- and metatibia narrow, medially emarginate or with parallel sides, not flanged, with shallow tibial groove (Figs 5 F, 6 I). Abdominal postcoxal line incomplete, recurved or slightly recurved (Figs 5 G, 6 J). Parameres symmetrical; basal lobe asymmetrical. Sipho slender with well-developed, T-shaped siphonal capsule (Figs 1 E���G, 4 D���F). Coxites transverse with spoon-like setae near apex, styles well developed, with or without apical setae; infundibulum lacking; sperm duct simple, uniform in diameter (Figs 1 H, 2 E, 3 F). Spermatheca with rounded cornu and narrow at base; distal capsule adjacent to cornu or sperm duct (Figs 2 F, 3 G). Remarks. Two species, M. angeloi sp. nov. and M. cordata sp. nov., lack the protibial emargination but are considered members of Menoscelis due to the described characters and male and female genital patterns., Published as part of Santos, Paula B., Churata-Salcedo, Julissa M. & Almeida, L��cia M., 2016, New species and records of Menoscelis Mulsant (Coleoptera: Coccinellidae) from French Guiana, pp. 252-268 in Zootaxa 4078 (1) on page 253, DOI: 10.11646/zootaxa.4078.1.23, http://zenodo.org/record/262121, {"references":["Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles d' Agriculture et d' Industrie, 2, 1 - 1104. [Lyon] http: // dx. doi. org / 10.5962 / bhl. title. 8953","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, 311 pp.","Gemminger, M. & Harold, E. (1876) Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Vol. 12. sumptu E. H. Gummi, Monachii, pp. 3740 - 3818. http: // dx. doi. org / 10.5962 / bhl. title. 9089","Chapin, E. A. (1966) A new species of Myrmecophilous Coccinellidae, with notes on other Hyperaspini (Coleoptera). Psyche, 73, 278 - 283. http: // dx. doi. org / 10.1155 / 1966 / 73641","El-Ali, A. A. (1972) A biosystematic study of Hyperaspini of California with emphasis on the immature stages. Ph. D. dissertation, University of California, Berkeley, 722 pp.","Gordon, R. D. (1987) A Catalog of the Crotch Collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology, 3, 1 - 46.","Milleo, J., Almeida, L. M. & Gordon, R. B. (2004) South American Coccinellidae (Coleoptera). Part X: A systematic revision of Thalassa Mulsant (Hyperaspidinae). Revista Brasileira de Entomologia, 48, 395 - 400. http: // dx. doi. org / 10.1590 / S 0085 - 56262004000300016","Gordon, R. D. & Canepari, C. (2008) South American Coccinellidae (Coleoptera), Part XI: A systematic revision of Hyperaspidini (Hyperaspidinae). Annali del Museo Civico di Storia Naturale \" G. Doria \", 99, 245 - 512.","Chapuis, F. (1876) Famille des phytophages des erotyliens des endomychides et des coccinellides. 12. In: Lacordaire, J. T. & Chapuis, F. (Eds.), Histoire naturelle des insectes. Genera des Coleopteres. Librairie Encyclopedique de Roret, Paris, pp. 1 - 424.","Korschefsky, R. (1931) Coccinellidae. I. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Pars 118. W. Junk, Berlin, pp. 1 - 224.","Blackwelder, R. E. (1945) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185 (3), 1 - 188. http: // dx. doi. org / 10.5479 / si. 03629236.185.3","Duverger, C. (1989) Contribution a l'etude des Hyperaspinae 1 ere note. Bulletin de la Societe Linneene, 17 (3), 143 - 157.","Fursch, H. (1990 b) Corrections and addictions. Coccinella. Newsletter for Systematic research in Coccinellids, 2 (2), 62 - 63. [Universitat Passau]"]}

Zenoria carbo Gonz��lez & Santos, sp. nov. Figs. 32���45 Diagnosis. Z. carbo sp. nov. resembles a group of species with a long subtriangular penis guide, about 3 times longer than wide, and parameres with setae only at the apex. Other species that have these features are Z. circumcincta Gordon, 1971; Z. emarginata Gordon, 1971; Z. linteolata Mulsant, 1850 and Z. purpurea Gordon, 1972. The new species differs from Z. circumcincta in the more elongated phallobase (3 times longer than wide) and the parameres parallel sided with a straight outer margin in ventral view. It is distinguished from the other species by having a penis guide with a sinuous inner margin in lateral view. Apart from this, all other species are predominantly lighter in color (with or without dark spots on the elytral disc), except Z. linteolata and Z. purpurea, which have a dark metallic-green and dark metallic-purple elytra respectively. Description of holotype, male. Length 3.2 mm; width 2.6 mm. Body oval, convex (Figs. 32���33). Head and prothorax pale yellow; antennae and mouthparts brown; pubescence yellowish white (Fig. 34). Pronotum with basal black spot occupying central �� of base and extending to middle; pubescence yellowish white, about half scutellar shield length (Figs. 32���34). Elytra black, shiny with yellowish-white pubescence, except central medial area with inconspicuous dark-brown pubescence (Figs. 32���35). Scutellar shield black (Fig. 32). Epipleuron black, inner carina extending half distance to outer margin. Meso- and metaventrite black. Legs pale yellow. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 36). Posterior margin of ventrite V with smooth emargination. Posterior margin of ventrite VI with wide emargination, accentuated medially, deep (Fig. 37). Phallobase long, �� length of tegminal strut, three times longer than wide (Fig. 37). Penis guide triangular in dorsal view, nearly three times longer than wide, sides almost parallel, slightly narrower in apical fourth; apex rounded (Figs. 39���40); in lateral view triangular, inner margin slightly sinuous (Figs. 42���43). Parameres projecting slightly beyond penis guide (Fig. 39), in ventral view subtriangular with straight lateral margins, directed slightly inward in distal 1/5 (Fig. 41), regularly curved in lateral view and parallel sided. Sparse short setae at end of paramere (Figs. 42���43). Penis normal, wide, no more than 40 times longer than wide, semicircular in basal 3/5, straight in apical 2/5, except apex recurved upward relative to penis capsule, with membranous areas on outer side, fewer on inner side (Figs. 44���45); penis capsule with inner branch slightly longer than outer branch, curved inward, narrow at base and widening to 2/3 of length with membranous areas, outer branch with almost parallel sides, truncated, with outer membranous area (Fig. 44). Female. Unknown. Variation. Length 3.1���3.2 mm; width 2.5���2.6 mm. The pale yellow coloration of the pronotal anterior margins may be reduced to �� or �� of the size. Etymology. From Latin " carbo " meaning coal, referring to the intense black color of the elytra; proposed as a noun in apposition. Type material. Holotype. ��� Per��, Madre de Dios / Tahuamanu, Iberia / 292 mnsm, 10-I-2010 / Leg. J. Mir��, C��tricos ��� ��� Male 1617��� [MEUT]. Paratypes (2): (1) ��� Per��, Madre de Dios / Tahuamanu, Iberia / 292 mnsm, 10-I-2010 / Leg. J. Mir��, C��tricos ��� ��� Male 612��� [MEUT]. (1) ��� Per�� Madre de Dios dept. / CICRA FldStn, trail 6 research / plot, 12.55207��S 70.10962��W / 295m, 11-13.VI.2011, Chaboo / team, canopy Malaise, bottom / sample, PER-11-CMB-002��� ���[bar code] / SEM C0986969 / KUNHM-ENT��� ��� Male 1571��� [UNMSM]. Distribution. Peru, Madre de Dios., Published as part of Santos, Paula B. & Gonz��lez, Guillermo, 2016, New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru, pp. 386-394 in Zootaxa 4205 (4) on pages 390-392, DOI: 10.11646/zootaxa.4205.4.7, http://zenodo.org/record/197192, {"references":["Gordon, R. D. (1971) A revision of the genus Zenoria Mulsant (Coleoptera: Coccinellidae). Smithsonian Contributions to Zoology, 86, 1 - 22.","Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles d' Agriculture et d' Industrie, Lyon, 2, 1 - 1104.","Gordon, R. D. (1972) Additional notes on the taxonomy of the genus Zenoria (Coleoptera: Coccinellidae). Proceedings of the Entomological Society of Washington, 7 (4), 433 - 442."]}

Menoscelis cordata sp. nov. (Figures 2, 7, 9, 11) Type Material. ��� Guyane Fran��aise,/ 4 ��02���N 52 �� 41 ���W, R��serve/Naturelle des Nouragues���/Saut Parar��, 0 8. IX. 2009,/vitre, SEAG col.��� ���Female��� [white label]. ��� HOLOTYPE / Menoscelis cordata /Santos, Almeida & Churata-Salcedo, 2015 ��� [red label]. [MNHN]. Description (female). Length 5.01, width 4.58 mm. Body rounded, strongly convex (Figs 2 A���B). Head yellowish with triangular black spot; clypeus with anterior margin black, labrum dark brown. Pronotum metallic greenish black, anterior and lateral margins yellowish (Fig. 2 B). Eyes finely faceted, not emarginate; interfacetal setae absent (Fig. 7 E). Antennal insertion exposed (Fig. 7 E). Antennae 10 -segmented; scape with convex external suface; pedicel distinctly narrower than scape; antepenultimate antennomere about 2.5 times longer than other segments (Fig. 7 E). Clypeus short, transverse, anterior clypeal margin emarginate medially, with small blunt lateral projections (Fig. 2 B). Labrum exposed. Mandibles bifid, with basal tooth. Maxillary palps securiform (Fig. 7 D). Labial palps 3 -segmented (Fig. 7 D). Mentum cordiform (Fig. 7 D). Prosternal process with complete carinae, joined anteriorly forming a triangle, with stem varying in length, reaching anterior margin of prosternum (Figs 7 D, 9 B). Elytra metallic greenish black with large cordiform yellow spot on the disc; epipleura dark brown, explanate and excavated to receive femoral apices (Fig. 7 D). Pro- and mesosternum dark brown; metasternum yellowish. Legs brown with apices of femora yellowish. Protibia slightly angulate near base, sides parallel (Fig. 7 F). Abdomen yellowish, with yellow pubescence (Fig. 2 C���D). Abdominal postcoxal lines recurved and incomplete, next to hind margin of ventrite, oblique line absent (Figs 2 C���D). Genital plate transverse (Fig. 2 E). Spermatheca with rounded cornu and wide apical projection, base slender (Figs 2 E, F). Coxites with spoon���shaped setae; style with indistinct apical setae and tergite with setae curved at apex (Fig. 2 G). Male. Unknown. Etymology. This species is named based on the shape of the elytral spot, cordata L., heart-shaped. Geographical distribution. French Guiana (R��serve Nationalle Naturelle des Nouragues���Saut Parar��) (Fig. 11). Remarks. Menoscelis cordata sp. nov. differs from all other species of the genus in its color pattern and female genitalia. The presence of very short grayish-white semidecumbent pubescence and the shape of the legs of M. cordata sp. nov. resembles only M. angeloi sp. nov., Published as part of Santos, Paula B., Churata-Salcedo, Julissa M. & Almeida, L��cia M., 2016, New species and records of Menoscelis Mulsant (Coleoptera: Coccinellidae) from French Guiana, pp. 252-268 in Zootaxa 4078 (1) on page 255, DOI: 10.11646/zootaxa.4078.1.23, http://zenodo.org/record/262121

Menoscelis flava sp. nov. (Figures 3, 7, 9, 11) Material examined. ��� Guyane Fran��aise,/ 4 �� 44 ��� 56 ���N 2 �� 26 ��� 28 ���W,/Montagne des Chevaux/ 14.I. 2012,/malaise SLAM,/ SEAG col.��� ���Female��� [white label]. ��� HOLOTYPE / Menoscelis flava /Santos, Almeida & Churata-Salcedo, 2015 ��� [red label]. [MNHN]. Description (female). Length 4.5, width 4.17 mm. Body rounded, strongly convex (Figs 3 A���C). Head yellowish with one triangular black spot; clypeus with anterior margin brownish, labrum dark yellowish. Pronotum black with yellowish anterior and lateral margins (Figs 3 A���C). Eyes finely faceted, not emarginate; interfacetal setae absent. Antennal insertion exposed. Antennae 10 -segmented; scape with convex external suface; pedicel distinctly narrower than scape; antepenultimate antennomere about 2.5 times longer than other segments (Fig. 7 G). Clypeus short, transverse, anterior clypeal margin emarginate medially, with small blunt lateral projections. Labrum exposed. Mandibles bifid, with basal tooth. Maxillary palps securiform. Labial palps 3 -segmented. Mentum cordiform. Prosternal process with complete carinae, joined anteriorly forming a triangle, with long stem, reaching anterior margin of prosternum (Fig. 9 A). Each elytron yellowish with narrow black lateral margin; three black spots: first quadrangular, on humeral callus; second on elytral disc, joined at suture; third at apex (Figs 3 A, C). Epipleura yellowish with black lateral margins, explanate and excavated to receive femoral apices. Middle part of pro-, meso- and metasternum black. Legs black, except apices of femora yellowish. Protibia slightly angulate near base and emarginate. Abdomen yellowish with yellow pubescence (Figs 3 D���E). Abdominal postcoxal lines recurved and incomplete, joined from hind margin of ventrite, oblique line absent (Figs 3 D���E). Genital plate transverse (Fig. 3 F). Spermatheca with rounded cornu, and wide, sinuate apical projection, base slender (Figs 3 F���G). Coxites with spoon���shaped setae and style with well-developed apical setae; tergite with long setae curved at apex (Fig. 3 H). Male. Unknown. Etymology. This species is named after the color of its body pubescence, flava L., yellow. Geographical Distribution: French Guiana (Roura, Montagne des Chevaux) (Fig. 11). Remarks. Menoscelis flava sp. nov. differs from all other species of the genus. The elytral color pattern of Menoscelis flava sp. nov. resembles M. stephani and M. angeloi sp. nov., but differs in the size of the discal and lateral spots, and the shape of the spermatheca., Published as part of Santos, Paula B., Churata-Salcedo, Julissa M. & Almeida, L��cia M., 2016, New species and records of Menoscelis Mulsant (Coleoptera: Coccinellidae) from French Guiana, pp. 252-268 in Zootaxa 4078 (1) on pages 255-256, DOI: 10.11646/zootaxa.4078.1.23, http://zenodo.org/record/262121

Santos, Paula B., González, Guillermo (2016): New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru. Zootaxa 4205 (4): 386-394, DOI: http://doi.org/10.11646/zootaxa.4205.4.7

Menoscelis angeloi sp. nov. (Figures 1, 6, 9, 11) Material examined. ��� Guyane Fran��aise, Point/de vue du B��lv��d��re de/Sa��l, 0 9. IX. 2011, vitre,/ SEAG col. ���Male��� [white label]. ��� HOLOTYPE / Menoscelis angeloi /Santos, Churata-Salcedo & Almeida, 2015 ��� [red label]. [MNHN]. ��� Guyane Fran��aise,/ 4 �� 44 ��� 56 ���N 2 �� 26 ��� 28 ���W,/Montagne des Chevaux, / 75 m, 17. III. 2012,/polytrap, SEAG col.��� ��� DZUP / 285311 ��� [white label]. ��� PARATYPE / Menoscelis angeloi /Santos, Almeida & Churata-Salcedo, 2015 ��� [yellow label]. ��� Guyane Fran��aise,/Montagne des Chevaux,/ 08. V. 2011,/vitre,/ SEAG col. ��� ���Female��� ��� DZUP / 289364 ��� [white label]. ��� PARATYPE / Menoscelis angeloi /Santos, Almeida & Churata-Salcedo, 2015 ��� [yellow label]. Description (male). Length 5.01, width 4.42 mm. Body rounded, strongly convex (Fig. 1 B). Head yellowish with triangular black spot; clypeus with anterior margin brownish, labrum dark yellowish. Eyes finely faceted, not emarginate; interfacetal setae absent. Antennal insertion exposed. Antennae 10 -segmented; scape with convex external suface; pedicel distinctly narrower than scape; antepenultimate antennomere about 2.5 times longer than other segments. Clypeus short, transverse, anterior clypeal margin emarginate medially, with small blunt lateral projections. Labrum exposed. Mandibles bifid, with basal tooth. Maxillary palps securiform. Labial palps 3 - segmented. Mentum cordiform. Pronotum black with yellowish anterior and lateral margins; narrow basal pronotal line present; basomedian projection of pronotum truncate. Prosternal process with complete carinae, joined anteriorly forming a triangle, with stem varying in length, reaching anterior margin of prosternum (Fig. 9 D). Each elytron reddish yellow with narrow black lateral margin; three small black spots: first spot smallest, on humeral callus, second on elytral disc, third near lateral margin (Fig. 1 A). Epipleura yellowish, explanate, and excavated to receive femoral apices. Middle part of pro-, meso- and metasternum black. Legs black, except apices of femora yellowish. Protibia slightly angulate near base, with parallel sides. Abdomen yellowish with yellow pubescence. Abdominal postcoxal lines recurved and incomplete, joined from hind margin of ventrite, oblique line absent. Ventrite 5 in male without apparent sexual caracters (Fig. 1 F). Male genitalia with basal lobe asymmetrical, shorter than parameres, apex rounded; parameres symmetrical, straight, apex rounded with short setae (Figs 1 G���H); sipho slender, long, slightly curved; siphonal capsule with outer arm well developed; apex slightly sinuate and wide (Fig. 1 I). Female. Length 5.25, width 4.67 mm. Similar to male (Figs 1 E, 6 A���J, 9 C, E). Each elytron reddish yellow with narrow black lateral margin; four small black spots: first spot smallest, on humeral callus, second on elytral disc, third near lateral margin, and fourth apical, joined at suture (Figs 1 C���D). Genital plate transverse (Fig. 1 J). Spermatheca with rounded cornu and slender base with wide sinuate cornu projection (Fig. 1 J). Coxites with spoon���shaped setae and style with apical setae well developed; tergite with long setae curved at apex (Fig. 1 K). Etymology. This species is named in honor of the dedicated entomologist and specialist on Odonata, Dr. Angelo B. M. Machado. Geographical distribution. French Guiana (Point de vue du B��lv��d��re de Sa��l and Montagne des Chevaux) (Fig. 11). Remarks. Menoscelis angeloi sp. nov. differs from all other species of the genus, except M. cordata sp. nov., based on the indistinct, very short grayish-white semidecumbent pubescence and the tibia slightly angulate near the base, with parallel sides (Figs 6 F���G). This species differs from M. cordata sp. nov. in the color pattern and female genitalia. The elytral color pattern of M. angeloi sp. nov. resembles M. stephani and M. flava sp. nov., but differs from these species in the size of the discal and lateral spots, shape of the spermatheca, and location of the spermathecal gland., Published as part of Santos, Paula B., Churata-Salcedo, Julissa M. & Almeida, L��cia M., 2016, New species and records of Menoscelis Mulsant (Coleoptera: Coccinellidae) from French Guiana, pp. 252-268 in Zootaxa 4078 (1) on page 254, DOI: 10.11646/zootaxa.4078.1.23, http://zenodo.org/record/262121

Zenoria carbo González & Santos, sp. nov. Figs. 32–45 Diagnosis. Z. carbo sp. nov. resembles a group of species with a long subtriangular penis guide, about 3 times longer than wide, and parameres with setae only at the apex. Other species that have these features are Z. circumcincta Gordon, 1971; Z. emarginata Gordon, 1971; Z. linteolata Mulsant, 1850 and Z. purpurea Gordon, 1972. The new species differs from Z. circumcincta in the more elongated phallobase (3 times longer than wide) and the parameres parallel sided with a straight outer margin in ventral view. It is distinguished from the other species by having a penis guide with a sinuous inner margin in lateral view. Apart from this, all other species are predominantly lighter in color (with or without dark spots on the elytral disc), except Z. linteolata and Z. purpurea, which have a dark metallic-green and dark metallic-purple elytra respectively. Description of holotype, male. Length 3.2 mm; width 2.6 mm. Body oval, convex (Figs. 32–33). Head and prothorax pale yellow; antennae and mouthparts brown; pubescence yellowish white (Fig. 34). Pronotum with basal black spot occupying central ¾ of base and extending to middle; pubescence yellowish white, about half scutellar shield length (Figs. 32–34). Elytra black, shiny with yellowish-white pubescence, except central medial area with inconspicuous dark-brown pubescence (Figs. 32–35). Scutellar shield black (Fig. 32). Epipleuron black, inner carina extending half distance to outer margin. Meso- and metaventrite black. Legs pale yellow. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 36). Posterior margin of ventrite V with smooth emargination. Posterior margin of ventrite VI with wide emargination, accentuated medially, deep (Fig. 37). Phallobase long, ½ length of tegminal strut, three times longer than wide (Fig. 37). Penis guide triangular in dorsal view, nearly three times longer than wide, sides almost parallel, slightly narrower in apical fourth; apex rounded (Figs. 39–40); in lateral view triangular, inner margin slightly sinuous (Figs. 42–43). Parameres projecting slightly beyond penis guide (Fig. 39), in ventral view subtriangular with straight lateral margins, directed slightly inward in distal 1/5 (Fig. 41), regularly curved in lateral view and parallel sided. Sparse short setae at end of paramere (Figs. 42–43). Penis normal, wide, no more than 40 times longer than wide, semicircular in basal 3/5, straight in apical 2/5, except apex recurved upward relative to penis capsule, with membranous areas on outer side, fewer on inner side (Figs. 44–45); penis capsule with inner branch slightly longer than outer branch, curved inward, narrow at base and widening to 2/3 of length with membranous areas, outer branch with almost parallel sides, truncated, with outer membranous area (Fig. 44). Female. Unknown. Variation. Length 3.1–3.2 mm; width 2.5–2.6 mm. The pale yellow coloration of the pronotal anterior margins may be reduced to ½ or ¼ of the size. Etymology. From Latin " carbo " meaning coal, referring to the intense black color of the elytra; proposed as a noun in apposition. Type material. Holotype. “ Perú, Madre de Dios / Tahuamanu, Iberia / 292 mnsm, 10-I-2010 / Leg. J. Miró, Cítricos ” “ Male 1617” [MEUT]. Paratypes (2): (1) “ Perú, Madre de Dios / Tahuamanu, Iberia / 292 mnsm, 10-I-2010 / Leg. J. Miró, Cítricos ” “ Male 612” [MEUT]. (1) “ Perú Madre de Dios dept. / CICRA FldStn, trail 6 research / plot, 12.55207°S 70.10962°W / 295m, 11-13.VI.2011, Chaboo / team, canopy Malaise, bottom / sample, PER-11-CMB-002” “[bar code] / SEM C0986969 / KUNHM-ENT” “ Male 1571” [UNMSM]. Distribution. Peru, Madre de Dios.

Santos, Paula B., Churata-Salcedo, Julissa M., Almeida, Lúcia M. (2016): New species and records of Menoscelis Mulsant (Coleoptera: Coccinellidae) from French Guiana. Zootaxa 4078 (1): 252-268, DOI: http://doi.org/10.11646/zootaxa.4078.1.23

Zenoria miroi González & Santos, sp. nov. Figs. 46–59 Diagnosis. Zenoria miroi sp. nov. is easily distinguishable from other species with the same color pattern, Z. crotchi Gordon, 1971; Z. delicatula Weise, 1910; Z. discoidalis (Kirsch, 1876); Z. lativerpa González & Honour, 2012; Z. luciae González & Honour, 2012; and Z. sylvatica González & Honour, 2012 by the subtriangular black basal spot on the pronotum. The male genitalia with short, inflated parameres, curved in lateral view with sensory setae limited to the apex are distinctly different from other described species. Although the male genitalia resemble Z. annularis Gordon, 1971 this species has broad triangular parameres, angulate at the base and straight on the outer margin in ventral view. Description of holotype, male. Length 3.2 mm; width 2.5 mm. Body oval, sligthly convex (Figs. 46–47). Head pale yellow with yellowish-white pubescence, longer and more abundant toward clypeus; antennae and mouthparts brown (Fig. 48). Pronotum dark yellow with subrectangular basal black spot; pubescence yellowish white (Figs. 46–48). Elytra pale yellow with large black common spot on disc, covering part of humeral callus; pubescence yellowish-white, except elytral disc with dark-brown pubescence (Figs. 46–49). Scutellar shield black (Fig. 46). Epipleuron yellow, inner carina extending less than half distance to outer margin. Meso- and metaventrite black. Legs pale yellow. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 50). Posterior margin of ventrite V arcuate. Posterior margin of ventrite VI with broad and deep semicircular emargination (Fig. 51). Phallobase short, less than half length of tegminal strut, two times longer than wide (Fig. 52). Penis guide mamilliform, basal third broad with convex sides, apex rounded (Figs. 53–55); in lateral view broad at base, outer margin straight, inner margin curved and not sinuous, with tubercles (Figs. 56–57). Parameres projecting beyond penis guide in ventral view; outer margin wide and curved, inner margin parallel-sided (Figs. 53–54); curved in lateral view, apex truncate; short sensory setae at apex (Figs. 56–57). Penis normal, wide, no more than 40 times longer than wide, basal third semicircular, middle third curved, straight in apical third except apex curved downward and tapering upward relative to penis capsule in short, very narrow projection (Fig. 58); apical membranous projections on both sides (Fig. 59); penis capsule with inner branch curved inward, ending in enlargement, and triangular outer branch (Fig. 58). Female. Unknown. Variation. Not observed. Etymology. This species is named after the agronomist Jimmy Miro, collector of this species and many other species of Coccinellidae in Tumbes and Madre de Dios departments, Peru. Holotype. “ Perú, Madre de Dios / Tambopata, Las / Piedras, Alegría, 5-VIII / 2012, leg. Jimmy Miró ” “N: 8660120 E: 486620 / Cítricos: Naranjo ” “ Male 1100” [MEUT]. Distribution. Peru, Madre de Dios.

Zenoria westerduijni Gonz��lez & Santos, sp. nov. Figs. 18���30 Diagnosis. Z. westerduijni sp. nov. is distinguishable from all other species of the genus by the almost entirely black body. It also differs from other species by the slightly angulate postcoxal line and the extremely narrow penis, about 50 times longer than wide, typically 35���40 times in other species of the genus. Description of holotype, male. Length 4.0 mm; width 3.4 mm. Body oval, rounded, convex (Figs. 18���19). Head and pronotum black, latter with narrow yellow stripe on anterior border; antennae and mouthparts brown, apex sometimes darker; pubescence grayish white, short, irregularly distributed (Fig. 20). Prothorax black; pubescence light brown, short. Scutellar shield black. Elytra entirely black, shiny; pubescence light brown except for central area, with inconspicuous dark-brown pubescence; this pubescence lacking in some areas, probably due to rubbing (Figs. 18���21). Epipleuron black, inner carina extending more than half distance to outer margin. Meso- and metaventrite brown. Legs brown. Abdominal postcoxal line curved posterolaterally, reaching posterior margin of ventrite for short distance, then redirected anteriorly toward base of ventrite in lateral third (Fig. 22). Posterior margin of ventrite V with smooth notch. Posterior margin of ventrite VI with deep triangular emargination (Fig. 23). Phallobase short, half length of tegminal strut, two times longer than wide (Fig. 24). Penis guide triangular in dorsal view, two times longer than wide, with rounded apex (Figs. 25���27); in lateral view, triangular in basal 3/5, parallel-sided and narrow in distal 2/5 (Figs. 28���29). Penis guide projecting beyond parameres (Fig. 25), parallelsided in ventral view, slightly projected apically (Fig. 27); regularly curved in lateral view and tapering from base to apex, the latter with small truncated widening (Figs. 28���29). Sparse, very short setae at end of paramere (Fig. 29). Penis narrow, more than 50 times longer than wide, semicircular in basal third, gently curved in middle third, and straight in apical third (Fig. 30), ending in pointed apex, membranous areas in distal 1/20 of apex, on lower side (Fig. 31); inner branch of penis capsule slightly curved inward and truncated, outer branch subtriangular, tapered with hook, inconspicuous membranous projection on outer margin (Fig. 30). Female. Unknown. Variation. Not observed. Etymology. This species is named after the biologist and entomologist Rob Westerduijn, who collected this specimen and several others described in this contribution. Type material. Holotype. ��� Per��, Loreto / Picuroyacu, Pongo / 150 m, 18-XI-2008 Leg. / R. Westerduijn, / Understory of altered / mature forest��� ��� Male 1612��� [UNMSM]. Distribution. Peru, Loreto., Published as part of Santos, Paula B. & Gonz��lez, Guillermo, 2016, New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru, pp. 386-394 in Zootaxa 4205 (4) on pages 389-390, DOI: 10.11646/zootaxa.4205.4.7, http://zenodo.org/record/197192

Menoscelis insignis Mulsant, 1850 (Figures 4, 7, 8, 10) Material examined. ��� Guyane Fran��aise,/ 3 �� 37 ��� 22 ���N 53 �� 12 ��� 57 ���W /B��lv��d��re de Sa��l, 326 /m 30.VI. 2011, vitre 1,/ SEAG col.��� ���Male��� ��� DZUP / 289045 ���[white label]. ��� Guyane Fran��aise,/ 4 ��05���N 52 �� 41 ���W, R��serve/Naturelle des Nouragues���/Camp Inselberg, 411 m,/ 28.VI. 2013, vitre,/ SEAG col.��� ��� DZUP / 189143 ���[white label]. ��� Guyane Fran��aise,/ 4 �� 44 ��� 56 ���N 52 �� 26 ��� 28 ���W,/Montagne des Chevaux, 18. X. 2014, malaise/ SLAM, SEAG col.��� ��� DZUP / 146707 ��� [white label]. Description (male). For more detailed description, see Gordon & Canepari (2008) and Corr��a, Santos & Almeida (2011). Length: 6.58, width: 5.58 mm. Similar to female in all aspects except for elytral spots larger and almost joined (Fig. 4 A, B). Abdomen yellowish with yellow pubescence (Fig. 4 C). Genitalia with basal lobe asymmetrical, half length of parameres, apex slightly obliquely truncated; parameres straight, apex rounded with long setae (Fig. 4 D, E); sipho slender, long, strongly curved; siphonal capsule with inner and outer arm developed; apex sinuate (Fig. 4 F���H). Geographical Distribution. French Guiana (Cayenne, Montagne des Chevaux, B��lv��d��re de Sa��l and R��serve Naturelle des Nouragues New records) and Brazil (AM, Itacoatiara) (Fig. 10). Remarks. Menoscelis insignis differs widely from all other species of the genus in its color pattern and male genitalia. The shape of the mentum and legs, 11 -segmented antennae, and the slightly curved postcoxal line of M. insignis resemble M. saginata (Figs 5 A���B, D���E, G)., Published as part of Santos, Paula B., Churata-Salcedo, Julissa M. & Almeida, L��cia M., 2016, New species and records of Menoscelis Mulsant (Coleoptera: Coccinellidae) from French Guiana, pp. 252-268 in Zootaxa 4078 (1) on page 256, DOI: 10.11646/zootaxa.4078.1.23, http://zenodo.org/record/262121, {"references":["Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles d' Agriculture et d' Industrie, 2, 1 - 1104. [Lyon] http: // dx. doi. org / 10.5962 / bhl. title. 8953","Gordon, R. D. & Canepari, C. (2008) South American Coccinellidae (Coleoptera), Part XI: A systematic revision of Hyperaspidini (Hyperaspidinae). Annali del Museo Civico di Storia Naturale \" G. Doria \", 99, 245 - 512.","Correa, G. H., Santos, P. B. & Almeida, L. M. (2011) Additional species and new record to Neotropical genus Menoscelis Mulsant (Coleoptera: Coccinellidae: Hyperaspini). Supplement au Bulletin Le Coleopterist. Contribution a L'etude des Coleopteres de Guyane, Tome IV, 32 - 37."]}

Zenoria limitrophi Gonz��lez & Santos, sp. nov. Figs. 1���13 Diagnosis. Zenoria limitrophi sp. nov. has distinctive male genitalia. The structure differs from all other species in that the penis guide is as wide as long, while in other species it is much longer than wide, except Z. lativerpa Gonz��lez & Honour (2012), which is as wide as long, but with its maximum width at the apex and not at the base. Description of holotype, male. Length 3.9 mm; width 3.0 mm. Body oval, elongate, maximum width at basal third of elytra (Fig. 1). Head, antennae, mouthparts, prothorax, legs and abdomen pale yellow; median basal mark on pronotum black, anteriorly rounded. Head with sparse yellowish-white pubescence. Scutellar shield black. Elytra dark metallic green with narrow humeral and apical pale-yellow margin, widened at apex; yellowish-white pubescence, sparse, except discal area with sparse brown pubescence (Figs. 1���4). Margin of elytron broadly explanate, inner carina extending about �� distance to outer margin near elytral base. Meso- and metaventrite black. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 5). Posterior margin of ventrite V smoothly arched, convex. Posterior margin of ventrite VI with wide and deep semicircular emargination (Fig. 6). Phallobase more than half length of tegminal strut, nearly two times longer than wide (Fig. 7). Penis guide mamilliform, as wide as long at base, first third almost parallel-sided, sharply narrowed at midlength, subtriangular with rounded apex, with tubercules in apical third (Fig. 8); curved in lateral view, basal half with keel on inner side, measuring 2/5 of its length, narrower (Figs. 11���12). Parameres projecting beyond penis guide subtriangular in ventral view, curved inward and narrower in distal third, with short setae at apex (Fig. 9), curved in lateral view in basal half and nearly straight in distal half; apex pointed (Figs. 10���11). Penis normal wide, no more than 40 times longer than wide, semicircular in basal half; apical half straight (Fig. 13); apex bifurcate with outer side short and blunt, inner side long, sharp and slightly curved downward (Fig. 14); penis capsule with inner branch curved slightly inward and ending in a club, outer branch narrowing only near end, with outer membranous projection (Fig. 13). Female. Length 3.6 mm; width 3.0 mm. Similar to male except pronotum dark with narrow yellowish anterior and lateral margins, elytra lacking yellow humeral spot, apical spot shorter and narrower than in male (Fig. 15). Ventrite VI with shallow emargination on posterior margin. Spermatheca J-shaped, cornu thickened toward distal end, with lamellar projection and barely visible basal ramus; nodulus extending in same direction as cornu, conical; accessory gland present; sperm duct long; infundibulum absent; apex of bursa copulatrix not modified (Fig. 16). Coxites four times longer than wide, triangular; styli well developed (Fig. 17). Variation. Length 3.6���4.1 mm; width 3.0��� 3.2 mm. Etymology. From the Latin ���limitrophus���, indicating their location near on the border between Brazil, Colombia and Peru. Type material. Holotype. ��� Brasil, Amazonas, / Tabatinga, 50 m / 01-IX-2006, leg. R. / Westerduijn, understory / of open altered forest��� ��� Male 964��� ��� Zenoria sp. nov. 2 / Santos P.B. 2013��� [DZUP]. Paratypes (3): (1) ��� Brasil, Amazonas, / Tabatinga, 26-VIII al / 6-IX-2006, Leg. R. / Westerduijn. Understory / mature altered forest��� ��� Female 357��� ��� Zenoria / sp. 8 / Det.: G. Gonz��lez 2011��� ��� Zenoria sp. nov. 2 / Santos P.B. 2013��� [DZUP]. (2) ��� Brasil, Amazonas, / Tabatinga, 50 m / 01-IX-2006, leg. R. / Westerduijn, understory / of open altered forest��� ��� Male ��� ��� Zenoria / sp. 10 / Det.: G. Gonz��lez 2011��� ��� Zenoria sp. nov. 2 / Santos P.B. 2013��� [DZUP]. Distribution. Brazil, Amazonas., Published as part of Santos, Paula B. & Gonz��lez, Guillermo, 2016, New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru, pp. 386-394 in Zootaxa 4205 (4) on pages 387-389, DOI: 10.11646/zootaxa.4205.4.7, http://zenodo.org/record/197192, {"references":["Gonzalez, G. & Honour, R. (2012) Tres nuevas especies del genero Zenoria (Coleoptera: Coccinellidae) condiseno similar a Z. discoidalis (Kirsch). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 50, 175 - 181."]}

Zenoria miroi Gonz��lez & Santos, sp. nov. Figs. 46���59 Diagnosis. Zenoria miroi sp. nov. is easily distinguishable from other species with the same color pattern, Z. crotchi Gordon, 1971; Z. delicatula Weise, 1910; Z. discoidalis (Kirsch, 1876); Z. lativerpa Gonz��lez & Honour, 2012; Z. luciae Gonz��lez & Honour, 2012; and Z. sylvatica Gonz��lez & Honour, 2012 by the subtriangular black basal spot on the pronotum. The male genitalia with short, inflated parameres, curved in lateral view with sensory setae limited to the apex are distinctly different from other described species. Although the male genitalia resemble Z. annularis Gordon, 1971 this species has broad triangular parameres, angulate at the base and straight on the outer margin in ventral view. Description of holotype, male. Length 3.2 mm; width 2.5 mm. Body oval, sligthly convex (Figs. 46���47). Head pale yellow with yellowish-white pubescence, longer and more abundant toward clypeus; antennae and mouthparts brown (Fig. 48). Pronotum dark yellow with subrectangular basal black spot; pubescence yellowish white (Figs. 46���48). Elytra pale yellow with large black common spot on disc, covering part of humeral callus; pubescence yellowish-white, except elytral disc with dark-brown pubescence (Figs. 46���49). Scutellar shield black (Fig. 46). Epipleuron yellow, inner carina extending less than half distance to outer margin. Meso- and metaventrite black. Legs pale yellow. Abdominal postcoxal line curved posterolaterally, closely paralleling posterior margin of ventrite for short distance, then abruptly redirected anteriorly toward base of ventrite in lateral third (Fig. 50). Posterior margin of ventrite V arcuate. Posterior margin of ventrite VI with broad and deep semicircular emargination (Fig. 51). Phallobase short, less than half length of tegminal strut, two times longer than wide (Fig. 52). Penis guide mamilliform, basal third broad with convex sides, apex rounded (Figs. 53���55); in lateral view broad at base, outer margin straight, inner margin curved and not sinuous, with tubercles (Figs. 56���57). Parameres projecting beyond penis guide in ventral view; outer margin wide and curved, inner margin parallel-sided (Figs. 53���54); curved in lateral view, apex truncate; short sensory setae at apex (Figs. 56���57). Penis normal, wide, no more than 40 times longer than wide, basal third semicircular, middle third curved, straight in apical third except apex curved downward and tapering upward relative to penis capsule in short, very narrow projection (Fig. 58); apical membranous projections on both sides (Fig. 59); penis capsule with inner branch curved inward, ending in enlargement, and triangular outer branch (Fig. 58). Female. Unknown. Variation. Not observed. Etymology. This species is named after the agronomist Jimmy Miro, collector of this species and many other species of Coccinellidae in Tumbes and Madre de Dios departments, Peru. Holotype. ��� Per��, Madre de Dios / Tambopata, Las / Piedras, Alegr��a, 5-VIII / 2012, leg. Jimmy Mir�� ��� ���N: 8660120 E: 486620 / C��tricos: Naranjo ��� ��� Male 1100��� [MEUT]. Distribution. Peru, Madre de Dios., Published as part of Santos, Paula B. & Gonz��lez, Guillermo, 2016, New species of Zenoria Mulsant (Coleoptera: Coccinellidae) from Brazil and Peru, pp. 386-394 in Zootaxa 4205 (4) on pages 392-393, DOI: 10.11646/zootaxa.4205.4.7, http://zenodo.org/record/197192, {"references":["Gordon, R. D. (1971) A revision of the genus Zenoria Mulsant (Coleoptera: Coccinellidae). Smithsonian Contributions to Zoology, 86, 1 - 22.","Gonzalez, G. & Honour, R. (2012) Tres nuevas especies del genero Zenoria (Coleoptera: Coccinellidae) condiseno similar a Z. discoidalis (Kirsch). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 50, 175 - 181."]}

Pseudoryssomus rufomarginatus sp. nov. (Fig. 10) Description of holotype (female). Length 3.08, width 2.25 mm. Form elongate oval, widest at middle of elytra, not flattened dorsoventrally (Fig. 10 C). General color yellowish red, middle of thorax and abdomen yellowish red. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a dark brown basal spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 10 D). Elytra yellowish red, with a large dark brown spot subquadragular, covering most of elytra, reaching the humeral callus, with constriction in the apical and basal third, not reaching elytral margin (Fig. 10 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly, with punctures large, often confluent (Fig. 10 B). Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally. First abdominal ventrite with postcoxal line complete, extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 10 E). Genitalia with genital plates triangular, elongate, styli with setae; spermatheca elongate, with apex bulbous (Fig. 10 F, G). Male. Unknown. Etymology. The name of this species refers to the color of the elytra. Discussion. Pseudoryssomus rufomarginatus sp. nov. is somewhat similar with P. triangulus sp. nov., but differs in the shape of the spot and male and female genitalia. Although the pattern color is similar with Gordonoryssomus limeirai sp. nov., it differs by the 10 -segmented antenna, body not flattened dorsoventrally and tibia widened, flattened and angulated on external margin. Type material. ���Tunapuna/ Trinidad, W.I./ Aug. 26 -31, 1969 /H. & A. Howden��� ��� Pseudoryssomus /n. sp./det. R. Gordon 94 ��� ���♀��� [white label]. ��� HOLOTYPE / Pseudoryssomus rufomarginatus / Almeida & Santos, 2014 ��� [red label]. 1 ex. ��� DZUP / 185793 ��� [DZUP]. Geographical distribution. Tunapuna, Trinidad and Tobago (Fig. 16)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

9. O. rogeri Almeida & Santos, sp. nov. Type material: Holotype, female (BMNH). Type locality: ���Vila Cachoeira, Esp��rito Santo, Brazil ���. Geographical distribution: Brazil (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

5. O. guillermo Almeida & Santos, sp. nov. Type material: Holotype, male (MNHN). Type locality: ���Montagne de Chevaux, French Guiana ���. Geographical distribution: French Guiana (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

5. Gordonoryssomus everardoi Almeida & Santos, sp. nov. Type Material: Holotype, male (DZUP). Type locality: “Nova Friburgo, Brazil ”. Geographical distribution: Brazil (Fig. 17).

Pseudoryssomus triangulus sp. nov. (Fig. 11) Description of holotype (female). Length 5.26, width 3.89 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 11 C). General color yellowish red, middle of thorax and abdomen yellowish red. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 11 D). Elytra yellowish red with a triangular black spot, almost extending to apex, not reaching the humeral callus (Fig. 11 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally (Fig. 11 B). First abdominal ventrite with postcoxal line incomplete, extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 11 E). Genitalia with genital plates triangular, elongate, styli with setae; spermatheca elongate, uniform (Fig. 11 F, G). Male. Unknown. Etymology. The name of this species refers to the shape of the spot on the elytra. Discussion. Pseudoryssomus triangulus sp. nov. is somewhat similar to P. crucifer sp. nov., but is clearly distinguished by the shape of the spot and male and female genitalia. Type material. “Cayenne” “Muséum Paris/ 1930 /Coll. Sicard” “♀” [white label]. “ HOLOTYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [red label]. 1 ex. [MNHN]; “ Guyane Française,/Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 11.IX. 2011,/vitre, SEAG col.” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. [MNHN]; “ Guyane Française,/ Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 25.IX. 2011, /vitre, SEAG col.” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. “ DZUP / 289061 ” [DZUP]; “ Guyane Française, /Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 23.X. 2011, /vitre, SEAG col.” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. “ DZUP / 288989 ” [DZUP]; “ Guyane Française, /Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 07.XI.2011, 20 m/vitre, SEAG col.” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. “ DZUP / 289235 ” [DZUP]; “ Guyane Française,/Commune de Roura,/Montagne des Chevaux,/RN 2 PK22, 20 m,/ 02.X. 2011, solaire/automatique, SEAG col.” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. “ DZUP / 289898 ” [DZUP]; “ Guyane Française,/Commune de Roura, /Montagne des Chevaux, /RN 2 PK22, 20 m,/ 15.X. 2011, solaire/automatique, SEAG col.” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. “ DZUP / 289902 ” [DZUP]; “ Guyane Française,/Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 20 m, / 30.X. 2011, vitre, SEAG col.” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. “ DZUP / 289074 ” [DZUP]; “see notes” “ BRITISH GUIANA:/ Essequibo, R./Moraballi Creek./ 10 -X- 1929 /Oxf. Univ. Expedn. /B.M. 1929 - 485 ” “ 427 ” “ Oryssomus /det. R. Gordon 91 ” “♀” [white label]. “ PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ” [yellow label]. 1 ex. [BMNH]. Geographical distribution. Commune de Roura, Montagne des Chevaux, French Guiana; Essequibo, Guyana (Fig. 16).

3. Oryssomus unimaculatus Almeida & Lima, 1995 Oryssomus unimaculatus Almeida & Lima, 1995: 708���709 (original description, morphological details and male genitalia figured). Type material: Holotype, male (DZUP). Type locality: ���Amazonas, Brazil ���. Geographical distribution: Brazil (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024"]}

Key to species of the Neoryssomus (based on Hofmann (1970)) 1. Elytra with greenish metallic shine; elytral punctation dispersed and disordered; antennomere III three times longer than IV; last antennomere as long as the previous two; length 3.5 to 4.75 mm (Chile)..................... N. germainii (Crotch) 1 ���. Elytra entirely black or black with reddish border; prothorax black; 3 rd antennomere two times longer than the 4 th; length 3.3 to 3.65 mm (Chile).................................................................. N. variabilis (Korchefsky), Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Hofmann, W. (1970) Nuevo genero para un Coccinellidae (Col.) de Chile. Agricultura Tecnica, 30 (4), 218 - 219."]}

Gordonoryssomus everardoi sp. nov. (Fig. 12) Description of holotype (male). Length 3.44, width 2.26 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 12 C). General color yellowish red, middle of thorax and abdomen dark brown. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal triangular black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 12 D). Elytra yellowish red with a large black spot, extending from the base almost to the apex, reaching the middle of humeral callus (Fig. 12 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs yellowish brown. Metaventrite finely, sparsely punctured medially and anteriorly, coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally (Fig. 12 B). First abdominal ventrite with postcoxal line extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 12 E). Genitalia simple; basal lobe as longer than the parameres, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere straight in lateral view, nearly parallel-sided (Fig. 12 G, H); sipho flattened and rounded apex (Fig. 12 F). Female. Unknown. Etymology. This species is named in honor of Dr. Everardo José Grossi, a doctor with a passion for beetles. Discussion. The color pattern of this species is very similar with G. deyrollii, but it differs in body length, and also male genitalia, with basal lobe almost the same length as the parameres, and J-shaped sipho. It also differs from all other species of the genus by the color pattern and male genitalia. Type material. “ BRAZIL, R.J., Nova/Friburgo, Sans Souci/ 21 -I- 2007 1050 m /P. Grossi col.” “♂” [white label]. HOLOTYPE / Gordonoryssomus / everardoi /Almeida & Santos, 2014 ” [red label]. 1 ex. “ DZUP / 131938 ” [DZUP]. Geographical distribution. Nova Friburgo, Sans Souci, Rio de Janeiro, Brazil (Fig. 17).

Gordonoryssomus limeirai sp. nov. (Fig. 13) Description of holotype (male). Length 3.71, width 2.78 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 13 C). General color and middle of thorax and abdomen brownish yellow. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal triangular black spot, translucent at apex; finely, densely punctured; pubescence of semierect yellowish white hairs (Fig. 13 D). Elytra yellowish red with a large black spot, extending from the base almost to the apex, reaching the middle of humeral callus. With a brownish yellow spot in sutural margin along almost 2 / 3 of the suture (Fig. 13 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent (Fig. 13 B). Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally. First abdominal ventrite with postcoxal line almost touching the posterior margin of the first abdominal ventrite; precoxal area coarsely punctured on base of cavity (Fig. 13 E). Genitalia simple; basal lobe equal in length to the paramere, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere straight in lateral view, nearly parallel-sided (Fig. 13 G, H); sipho flattened, sinuous at apex (Fig. 13 F). FIGURE 13. Gordonoryssomus limeirai sp. nov. (A���E) Habitus: (A) dorsal; (B) ventral; (C) lateral; (D) frontal view. (E) Abdomen; (F���H) male genitalia: (F) sipho; (G) Tegmen, ventral; (H) lateral view. (I���J) Female terminalia: (I) spermatheca; (J) coxites. Female. Similar to male in all aspects externally. Genitalia with genital plates triangular, not elongated, styli with setae; spermatheca elongate, not uniform, with narrow apex (Fig. 13 I, J). Etymology. This species is named in honor of the dedicated entomologist, Dr. Francisco Limeira de Oliveira. Discussion. The color pattern is similar to that of G. delicatus, but it differs in length and the size of the sutural spot. Female genitalia with genital plate not elongate, somewhat triangular as in G. deyrollii, G. delicatus and G. mirnae. G. limeirai differs from all other species of the genus by the postcoxal line which almost touches the posterior margin of the first abdominal ventrite and male genitalia with wider basal lobe. Type material. ��� BRAZIL (PI), Caracol/Parque Nacional Serra das/Confus��es, Andorinha, 515 m,/ 09��08��� 27.8 ���S / 43 �� 33 ��� 42.1 ���W / CZMA ��� ���Suspensa dupla (20 metros)/ 20-30.IX. 2013, J.A. Rafael,/F. Limeira de Oliveira &/T.T.A. Silva cols./ CZMA ��� ���♂��� [white label]. ��� HOLOTYPE / Gordonoryssomus / limeirai /Almeida & Santos, 2014 ��� [red label]. 1 ex. [CZMA]; ��� BRAZIL (PI), Caracol/Parque Nacional Serra das/Confus��es, Andorinha, 515 m,/ 09��08��� 27.8 ���S / 43 �� 33 ��� 42.1 ���W / CZMA ��� ���Suspensa dupla (20 metros)/ 20-30.IX. 2013, J.A. Rafael, / F. Limeira de Oliveira & /T.T.A. Silva cols./ CZMA ��� ���♀��� [White label].��� PARATYPE / Gordonoryssomus / limeirai / Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 245996 ��� [DZUP]. Geographical distribution. Brazil (PI) (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Oryssomus rhombus sp. nov. (Fig. 6) Description of holotype (male). Length 5.07, width 3.84 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 6 C). General color brownish yellow. Outline of pronotum and elytron discontinuous. Pronotum brownish yellow, with a transverse basal black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 6 D). Elytra brownish yellow with a rhomboidal black spot on the middle of the dorsum (Fig. 6 A). Elytral punctures coarse, dense, slightly larger than on pronotum and sparse in the discal area. Antenna and mouthparts brownish yellow, hypomera and epipleura yellow, legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent (Fig. 6 B). Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally on first and second ventrite (Fig. 6 E). First abdominal ventrite with postcoxal line 2 / 3 distant from the posterior margin of the ventrite. Genitalia simple; basal lobe shorter than paramere, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere slightly curved in lateral view, nearly parallel-sided (Fig. 6 G, H); sipho slightly sinuous and flattened, rounded apex (Fig. 6 F). Female. Unknown. Etymology. The name of this species refers to the shape of the macula in the middle of the elytra. Discussion. Oryssomus rhombus sp. nov. differs from its congeners by its color, shape of spot and male genitalia. Type material. ��� Guyane Fran��aise,/B��lv��d��re de Sa��l,/ 29.X. 2010, vitre 1,/ SEAG col.��� ���♂��� [white label]. ��� HOLOTYPE / Oryssomus rhombus /Almeida & Santos, 2014 ��� [red label]. 1 ex. [MNHN]. Geographical distribution. B��lv��d��re de Sa��l, French Guiana (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Rhizoryssomus Hofmann, 1972 Rhizoryssomus Hofmann, 1972: 82 (original description). Type species. Rhizoryssomus ruficollis Hofmann, 1972. Diagnosis. Oryssomini with posterior angle of pronotum acute; antenna 11 -segmented; all tibiae slender, unmodified; postcoxal line extending 4 / 5 of the width of the ventrite. For more details see Hofmann (1972)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Hofmann, W. (1972) Die Chilenischen Cranophorini (Col. Coccinellidae). Mitteilunger der Munchner Entomologischen Gesellschaft, 62, 66 - 86."]}

Oryssomus rogeri sp. nov. (Fig. 7) Description of holotype (female). Length 4.92, width 3.61 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 7 C). General color brownish yellow. Outline of pronotum and elytron discontinuous. Pronotum brownish yellow, with a small basal black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 7 D). Elytra brownish yellow with a rhomboidal black spot on the middle of the dorsum (Fig. 7 A). Elytral punctures coarse, dense, slightly larger than on pronotum and sparse in the discal area. Antenna and mouthparts brownish yellow, hypomera and epipleura yellow, legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent (Fig. 7 B). Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally on first and second ventrites (Fig. 7 E). First abdominal ventrite with postcoxal line almost touching the posterior margin of the first abdominal ventrite. Genitalia with genital plates triangular, elongate, styli with setae; spermatheca very elongate, rounded at apex (Fig. 7 F). Male. Unknown. Etymology. This species is named in honor of Dr. Roger G. Booth, a noted specialist in Coccinellidae systematics. Discussion. Oryssomus rogeri sp. nov. resembles O. rhombus sp. nov. in color pattern, but the spots are differently shaped; it also differs in the form of the male and female genitalia. Type material. “ Oryssomus /sp./det. R.G. Booth 1986 ” “♀” [white label]. “ HOLOTYPE / Oryssomus rogeri / Almeida & Santos, 2014 ” [red label]. 1 ex. [BMNH]. Geographical distribution. Espírito Santo, Vila Cachoeira, Brazil (Fig. 15).

4. Gordonoryssomus delicatus Almeida & Lima, 1995 Gordonoryssomus delicatus Almeida & Lima, 1995: 715���717 (original description, morphological details and male genitalia figured). Type material: Holotype, male (DZUP); paratypes (4 total), 1 Rio de Janeiro and 1 Minas Gerais (USNM), 1 (BMNH), 1 Santa Catarina (DZUP). Type locality: ���Santa Catarina, Brazil ���. Geographical distribution: Brazil (RJ, MG, SC, RS) (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024"]}

2. Neoryssomus variabilis (Korschefsky, 1934) Oryssomus variabilis Korschefsky, 1934: 35 ���36 (original description); Blackwelder, 1945: 446 (check list). Neoryssomus variabilis: Hofmann, 1972: 83 ���84 (new combination; description; morphological details and male genitalia figured). Type material: Deutsches Entomologisches Institut, Berlin-Dahlen,���Prov. Concepcion, Chile, August 1903, P. Herbst leg.���. Geographical distribution: Chile: Aconcagua to Concepcion (Hofmann 1972). Remarks. Hofmann (1972), recognized some shared characteristics of these two species, Oryssomus variabilis and Cranoryssus germainii, mainly in the shape of the body very flattened, downward pronotum and posterior angles strongly rounded, placing them in a new genus, Neoryssomus. It differs from Oryssomus by having 11 antennomeres and from Gordonoryssomus by the flattened body., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Korschefsky, R. (1934) Weitere Bemerkungen Uber Crotch'sche Typen und Beschreibung einier neuen Oryssomus - Art aus dem Deutschen Entomologischen Institut. Arbeiten uber morphologische und taxonomische entomologie aus Berlin- Dahlem, 1, 35 - 36.","Blackwelder, R. E. (1945) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185 (3), 1 - 188.","Hofmann, W. (1972) Die Chilenischen Cranophorini (Col. Coccinellidae). Mitteilunger der Munchner Entomologischen Gesellschaft, 62, 66 - 86."]}

Gordonoryssomus mirnae sp. nov. (Fig. 14) Description of holotype (male). Length 5.64, width 3.82 mm. Form elongate oval, widest at middle of elytra, not flattened dorsoventrally (Fig. 14 C). General color reddish yellow (Fig. 14 A, B). Outline of pronotum and elytron discontinuous. Pronotum and elytra without spots. Pronotum translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 14 D). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs reddish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites finely punctured, punctures separated by about a diameter or more, punctures becoming thicker laterally. First abdominal ventrite with postcoxal line slightly angulate, extending beyond middle of first ventrite (Fig. 14 E). Genitalia simple; basal lobe longer than paramere, lateral margin thickened and abruptly widened in apical half, apex rounded; paramere slightly curved in lateral view, nearly parallel-sided (Fig. 14 G, H); sipho curved and apex acuminate (Fig. 14 F). Female. Similar to male in all aspects externally. Genitalia with genital plates triangular, not elongate, styli with setae; spermatheca elongate, uniform (Fig. 14 I). Etymology. This species is named in honor of a great lepidopterist, Dr. Mirna Martins Casagrande. Discussion. G. m i r n a e differs from the other species of the genus because it is the only species with uniform color pattern reddish yellow. FIGURE 14. Gordonoryssomus mirnae sp. nov. (A���E) Habitus: (A) dorsal; (B) ventral; (C) lateral; (D) frontal view. (E) Abdomen; (F���H) male genitalia: (F) sipho; (G) tegmen, ventral; (H) lateral view. Female terminalia: (I) coxites. Type material. ��� PERU, M. de Dios, Par-/que Manu, Pakitza 340m / 11 �� 55 ¨ 48 ���S 71 �� 15 ¨ 18 ���W/ 27 Sep 1991 / Leg. M. Casagrande��� ���♂��� [white label]. ��� HOLOTYPE / Gordonoryssomus / mirnae /Almeida & Santos, 2014 ��� [red label]. 1 ex. ��� DZUP / 188196 ��� [DZUP]; ���Chuani, Dept./La Paz/ Bol��via /May, 1925,/G. L. Harrington��� ���♀��� [white label]. ��� PARATYPE / Gordonoryssomus / mirnae /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. [USNM]. Geographical distribution. Peru and Bolivia (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

7. P. triangulus Almeida & Santos, sp. nov. Type Material: Holotype, female (MNHN). Type locality: “Cayenne, French Guiana ”. Geographical distribution: French Guiana (Fig. 16).

6. Gordonoryssomus limeirai Almeida & Santos, sp. nov. Type Material: Holotype, male (CZMA); paratype (1), same data of holotype (DZUP). Type locality: “Caracol/ Parque Nacional Serra das/Confusões, Andorinha, Piauí, Brazil ”. Geographical distribution: Brazil (Fig. 17).

Oryssomus paschoali sp. nov. (Fig. 5) Description of holotype (female). Length 3.85, width 2.59 mm. Form elongate oval, widest at middle of elytra, flattened dorsoventrally (Fig. 5 C). General color black, middle of thorax and abdomen dark brown. Outline of pronotum and elytron discontinuous. Antenna, mouthparts, hypomera, epipleura and legs yellowish brown. Clypeus and frons joined at abrupt angle. Pronotum black with yellowish brown triangular spots, causing the black markings to appear w-shaped; finely, densely punctured, punctures uniform in size; pubescence of semidecumbent yellowish hairs (Fig. 5 D). Elytra black with a narrow yellowish brown border on the posterior 1 / 3 of elytra (Fig. 5 A). Elytral punctures coarse, dense, slightly larger than on pronotum and similarly dense, uniform in size. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent (Fig. 5 B); each puncture with long, yellow, decumbent seta. Abdominal ventrites coarsely punctured, punctures separated by about twice a diameter, punctures becoming coarser laterally and on first ventrite. First abdominal ventrite with postcoxal line extending beyond middle of first ventrite (Fig. 5 E). Genitalia with genital plates triangular, elongate, styli with setae; spermatheca elongate, slender at apex (Fig. 5 F). Male. Unknown. Etymology. This species is named in honor of the dedicated entomologist, Paschoal Coelho Grossi. Discussion. Oryssomus paschoali sp. nov. differs from other species in the genus by the elytral and pronotal color. The flattened body shape and the sides almost parallel are as in Neoryssomus germainii and N. variabilis, but these differ in color pattern, number of antennomeres and body size. Type material. ��� Brazil /Paran����� ���Fry Coll./ 1905.100 ��� ��� 40403 ��� ��� Oryssomus sp. /det. R.G. Booth, 1986 ��� ���♀���[white label]. ��� HOLOTYPE / Oryssomus paschoali /Almeida & Santos, 2014 ��� [red label]. 1 ex. [BMNH]; ��� Brasil, Paran��/Pinhais (Expotrade), 20 -II- 2008 /P. Grossi (leg.)��� ���♀��� [white label]. ��� PARATYPE / Oryssomus paschoali /Almeida & Santos, 2014 ��� [yellow label]. 2 ex. " DZUP / 131936 and 144184 ��� [DZUP]. Geographical distribution. Pinhais, Paran��, Brazil (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

1. Rhizoryssomus ruficollis Hofmann, 1972 Rhizoryssomus ruficollis Hofmann, 1972: 84 ���85 (original description; morphological details and male genitalia figured). Type material: Holotype, 1 ex., Hofmann Collection, Chile. Geographical distribution: Chile. Remarks. Hofmann (1972) reports that the type material was from Chile, but without exact locality and is somewhat damaged, lacking the last abdominal ventrites. The shape and color of this species are characteristic, as well as pubescence. He also commented that this genus could be near Coccidulini, especially Dioria, rather than Cranophorini. According to Gonz��lez in his web site Coccinellidae de Chile, the type is lost., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Hofmann, W. (1972) Die Chilenischen Cranophorini (Col. Coccinellidae). Mitteilunger der Munchner Entomologischen Gesellschaft, 62, 66 - 86."]}

Almeida, Lúcia M., Santos, Paula B. (2014): Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima. Zootaxa 3846 (1), DOI: http://dx.doi.org/10.11646/zootaxa.3846.1.2

3. Pseudoryssomus brevipilosus Gordon, 1974 Pseudoryssomus brevipilosus Gordon 1974: 152 ���153 (original description; morphological details and male genitalia figured). Type material: Holotype, female (USNM). Type locality: ���Amazonas, Venezuela.��� Geographical distribution: Venezuela (Fig. 16)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Gordon, R. D. (1974) A review of the Oryssomini, a new tribe of Neotropical Coccinellidae (Coleoptera). The Coleopterists Bulletin, 28 (3), 145 - 154."]}

5. Gordonoryssomus everardoi Almeida & Santos, sp. nov. Type Material: Holotype, male (DZUP). Type locality: ���Nova Friburgo, Brazil ���. Geographical distribution: Brazil (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Pseudoryssomus crucifer sp. nov. (Fig. 9) Description of holotype (female). Length 3.50, width 2.48 mm. Form elongate oval, widest at middle of elytra, not flattened dorsoventrally (Fig. 9 C). General color yellowish red, middle of thorax and abdomen yellowish red. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal dark brown spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 9 D). Elytra yellowish red, with a large and irregular dark brown spot roughly in the shape of a cross, not reaching the humeral callus, with a lateral band in the middle of elytra and a smaller band before the apex (Fig. 9 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs reddish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent (Fig. 9 B). Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally. First abdominal ventrite with postcoxal line complete, extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 9 E). Genitalia with genital plates triangular, elongate, styli with setae; spermatheca elongate, uniform (Fig. 9 F). Male. Unknown. Etymology. The name of this species refers to the cross-shaped spot on the elytra. Discussion. Pseudoryssomus crucifer sp. nov. resembles P. b r ul e i sp. nov. in the color pattern, but differs in the shape of the spot and male and female genitalia. Type material. ��� Guyane Fran��aise,/Commune de Roura,/Montagne des Chevaux,/RN 2 PK22, 20 m,/ 14.VIII. 2011, malaise/ SLAM, SEAG col.��� ���♀��� [white label]. ��� HOLOTYPE / Pseudoryssomus crucifer /Almeida & Santos, 2014 ��� [red label]. 1 ex. [MNHN]. Geographical distribution. Montagne des Chevaux, French Guiana (Fig. 16)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

4. O. calix Almeida & Santos, sp. nov. Type material: Holotype, male (MNHN). Type locality: ���Montagne de Chevaux, French Guiana ���. Geographical distribution: French Guiana (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Key to species of the Neoryssomus (based on Hofmann (1970)) 1. Elytra with greenish metallic shine; elytral punctation dispersed and disordered; antennomere III three times longer than IV; last antennomere as long as the previous two; length 3.5 to 4.75 mm (Chile)..................... N. germainii (Crotch) 1 ’. Elytra entirely black or black with reddish border; prothorax black; 3 rd antennomere two times longer than the 4 th; length 3.3 to 3.65 mm (Chile).................................................................. N. variabilis (Korchefsky)

6. P. rufomarginatus Almeida & Santos, sp. nov. Type Material: Holotype, female (DZUP). Type locality: ���Tunapuna, Trinidad and Tobago ���. Geographical distribution: Trinidad and Tobago (Fig. 16)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

2. Oryssomus lineatus Gordon, 1974 Oryssomus lineatus Gordon, 1974: 148 ���149 (original description, morphological details and male genitalia figured); Almeida & Lima, 1995 (description, morphological details and male genitalia figured). Type material: Holotype, male (USNM); paratypes (19), Trinidad and Tobago (USNM and CNC). Type locality: ���Santa Cruz, Bolivia ���. Geographical distribution: Trinidad and Tobago, Bolivia (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Gordon, R. D. (1974) A review of the Oryssomini, a new tribe of Neotropical Coccinellidae (Coleoptera). The Coleopterists Bulletin, 28 (3), 145 - 154.","Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024"]}

7. Gordonoryssomus mirnae Almeida & Santos, sp. nov. Type Material: Holotype, male (DZUP); paratype (1), La Paz, Bolivia (USNM). Type locality: ���M. de Dios, Parque Manu, Peru ���. Geographical distribution: Peru and Bolivia (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Neoryssomus Hofmann, 1972 Neoryssomus Hofmann, 1972: 82 (original description). Type species. Oryssomus germainii Crotch, 1874. Diagnosis. Oryssomini with flattened body; antenna 11 -segmented; pronotum semicircular, covering the head, with posterior angles rounded; all tibiae slender, unmodified; postcoxal line complete; female genitalia with genital plate elongate; spermatheca simple, curved. For more details see Hofmann (1972)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Hofmann, W. (1972) Die Chilenischen Cranophorini (Col. Coccinellidae). Mitteilunger der Munchner Entomologischen Gesellschaft, 62, 66 - 86.","Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, 311 pp."]}

6. Gordonoryssomus limeirai Almeida & Santos, sp. nov. Type Material: Holotype, male (CZMA); paratype (1), same data of holotype (DZUP). Type locality: ���Caracol/ Parque Nacional Serra das/Confus��es, Andorinha, Piau��, Brazil ���. Geographical distribution: Brazil (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Pseudoryssomus triangulus sp. nov. (Fig. 11) Description of holotype (female). Length 5.26, width 3.89 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 11 C). General color yellowish red, middle of thorax and abdomen yellowish red. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 11 D). Elytra yellowish red with a triangular black spot, almost extending to apex, not reaching the humeral callus (Fig. 11 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally (Fig. 11 B). First abdominal ventrite with postcoxal line incomplete, extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 11 E). Genitalia with genital plates triangular, elongate, styli with setae; spermatheca elongate, uniform (Fig. 11 F, G). Male. Unknown. Etymology. The name of this species refers to the shape of the spot on the elytra. Discussion. Pseudoryssomus triangulus sp. nov. is somewhat similar to P. crucifer sp. nov., but is clearly distinguished by the shape of the spot and male and female genitalia. Type material. ���Cayenne��� ���Mus��um Paris/ 1930 /Coll. Sicard��� ���♀��� [white label]. ��� HOLOTYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [red label]. 1 ex. [MNHN]; ��� Guyane Fran��aise,/Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 11.IX. 2011,/vitre, SEAG col.��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. [MNHN]; ��� Guyane Fran��aise,/ Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 25.IX. 2011, /vitre, SEAG col.��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 289061 ��� [DZUP]; ��� Guyane Fran��aise, /Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 23.X. 2011, /vitre, SEAG col.��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 288989 ��� [DZUP]; ��� Guyane Fran��aise, /Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 07.XI.2011, 20 m/vitre, SEAG col.��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 289235 ��� [DZUP]; ��� Guyane Fran��aise,/Commune de Roura,/Montagne des Chevaux,/RN 2 PK22, 20 m,/ 02.X. 2011, solaire/automatique, SEAG col.��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 289898 ��� [DZUP]; ��� Guyane Fran��aise,/Commune de Roura, /Montagne des Chevaux, /RN 2 PK22, 20 m,/ 15.X. 2011, solaire/automatique, SEAG col.��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 289902 ��� [DZUP]; ��� Guyane Fran��aise,/Commune de Roura,/Montagne des Chevaux, /RN 2 PK22, 20 m, / 30.X. 2011, vitre, SEAG col.��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 289074 ��� [DZUP]; ���see notes��� ��� BRITISH GUIANA:/ Essequibo, R./Moraballi Creek./ 10 -X- 1929 /Oxf. Univ. Expedn. /B.M. 1929 - 485 ��� ��� 427 ��� ��� Oryssomus /det. R. Gordon 91 ��� ���♀��� [white label]. ��� PARATYPE / Pseudoryssomus / triangulus /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. [BMNH]. Geographical distribution. Commune de Roura, Montagne des Chevaux, French Guiana; Essequibo, Guyana (Fig. 16)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

6. O. paschoali Almeida & Santos, sp. nov. Type material: Holotype, female (BMNH). Type locality: ���Paran��, Brazil ���. Geographical distribution: Brazil (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Key to species of Oryssomus Mulsant 1. Pronotum and elytra yellow with dark brown rounded discal spot Brazil (AM).......... O. unimaculatus Almeida & Lima 1 ���. Dorsal color pattern not as described above................................................................. 2 2. Pronotum with an elongate, median, black spot extending from base nearly to apex; elytron black with large, red spot on apical third (Mexico, Belize, Guatemala, El Salvador, Honduras, Costa Rica, Panama, Trinidad and Tobago, Venezuela, Colombia, French Guiana, Brazil)............................................................. O. subterminatus Mulsant 2 ���. Dorsal color pattern not as described above................................................................. 3 3. Elytra black with humeral area and diagonal stripe red (Bolivia and Trinidad and Tobago)............. O. lineatus Gordon 3 ���. Dorsal color pattern not as described above................................................................ 4 4. Elytra black with a slender yellowish brown line at border on the posterior 1 / 3 of elytra (Brazil)........ O. paschoali sp. nov. 4 ���. Dorsal color pattern not as described above................................................................. 5 5. Elytra reddish brown with chalice-shaped black spot in the middle that does not reach humeral callus (French Guiana)...... ....................................................................................... O. calix sp. nov. 5 ���. With a different color pattern........................................................................... 6 6. Elytra reddish brown with black large spot in base comprising 2 / 3 of total length of the elytra (French Guiana)............................................................................................... O. guillermo sp. nov. 6 ���. Dorsal color pattern not as described above................................................................. 7 7. Postcoxal line extending just beyond middle of first ventrite; elytra brownish yellow with a rhomboidal black spot on the mid- dle of the dorsum (French Guiana)........................................................ O. rhombus sp. nov. 7 ���. Postcoxal line almost touching the posterior margin of the first abdominal ventrite; elytra brownish yellow with black area not rhomboidal......................................................................................... 8 8. Elytra with a triangular black spot almost extending to apex (French Guiana).................... O. guyanensis sp. nov. 8 ���. Elytra with a large black spot arrow shaped (Brazil)............................................ O. rogeri sp. nov., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

7. O. guyanensis Almeida & Santos, sp. nov. Type material: Holotype, female (MNHN). Type locality: ���Montagne de Chevaux, French Guiana ���. Geographical distribution: French Guiana (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

2. Gordonoryssomus nigripilosus Almeida & Lima, 1995 Gordonoryssomus nigripilosus Almeida & Lima, 1995: 713���714 (original description, morphological details and male genitalia figured). Type material: Holotype, male (MNHN). Type locality: ���Esp��rito Santo, Brazil ���. Geographical distribution: Brazil (ES) (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024"]}

Oryssomus calix sp. nov. (Fig. 2) Description of holotype (male). Length 3.82, width 2.81 mm. Form elongate oval, widest at middle of elytra, flattened dorsoventrally (Fig. 2 C). Dorsal and ventral general color reddish brown (Figs. 2 A, B). Outline of pronotum and elytron discontinuous. Antenna, mouthparts, hypomera, epipleura and legs reddish brown. Clypeus and frons joined at abrupt angle. Pronotum translucent; finely, densely punctured, punctures uniform in size, with a slight depression at lateral sides; pubescence of semidecumbent yellowish hairs; with black spot at base (Fig. 2 D). Elytra reddish brown with a black spot chalice-shaped in the middle, not reaching humeral callus (Fig. 2 A). Elytral punctures coarse, dense, slightly larger than on pronotum and similarly dense, uniform in size. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent; each puncture with long, yellow, decumbent seta. Abdominal ventrites coarsely punctured, punctures separated by about twice a diameter, punctures becoming coarser laterally and on last ventrite. First abdominal ventrite with postcoxal line extending beyond middle of ventrite (Fig. 2 E). Genitalia with basal lobe equal in length to paramere, lateral margin slightly thickened, parameres slightly curved in lateral view (Figs. 2 G, H); sipho sinuous, apex nearly membranous, unmodified (Fig. 2 F). Female. Length 4.21, width 3.02 mm. Similar to male in all aspects externally. Genitalia with genital plates elongate, styli with setae; spermatheca elongate, unmodified (Fig. 2 I). Etymology. The name of this species refers to the shape of black spot of the elytra. Discussion. Oryssomus calix sp. nov. is similar to O. guillermo sp. nov. and O. unimaculatus, but is clearly distinguished by the chalice-shaped spot on the elytra, and the genitalia. Type material. ��� Guyane Fran��aise,/Commune de Roura,/Montagne des Chevaux,/RN 2 PK22, 17.III. 2013,/ solaire authomatique,/ SEAG col.��� "♂" [white label]. ��� HOLOTYPE / Oryssomus calix /Almeida & Santos, 2014 ��� [red label]. 1 ex. [MNHN]; ��� Guyane Fran��aise,/Commune de Roura,/Montagne des Chevaux,/RN 2 PK22, 22.VIII. 2010,/vitre, SEAG col.��� "♀" [white label]. ��� PARATYPE / Oryssomus calix /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ZUP/ 145669 ���. Geographical distribution. Montagne des Chevaux, French Guiana (Fig. 15)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Gordonoryssomus everardoi sp. nov. (Fig. 12) Description of holotype (male). Length 3.44, width 2.26 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 12 C). General color yellowish red, middle of thorax and abdomen dark brown. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal triangular black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 12 D). Elytra yellowish red with a large black spot, extending from the base almost to the apex, reaching the middle of humeral callus (Fig. 12 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs yellowish brown. Metaventrite finely, sparsely punctured medially and anteriorly, coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally (Fig. 12 B). First abdominal ventrite with postcoxal line extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 12 E). Genitalia simple; basal lobe as longer than the parameres, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere straight in lateral view, nearly parallel-sided (Fig. 12 G, H); sipho flattened and rounded apex (Fig. 12 F). Female. Unknown. Etymology. This species is named in honor of Dr. Everardo Jos�� Grossi, a doctor with a passion for beetles. Discussion. The color pattern of this species is very similar with G. deyrollii, but it differs in body length, and also male genitalia, with basal lobe almost the same length as the parameres, and J-shaped sipho. It also differs from all other species of the genus by the color pattern and male genitalia. Type material. ��� BRAZIL, R.J., Nova/Friburgo, Sans Souci/ 21 -I- 2007 1050 m /P. Grossi col.��� ���♂��� [white label]. HOLOTYPE / Gordonoryssomus / everardoi /Almeida & Santos, 2014 ��� [red label]. 1 ex. ��� DZUP / 131938 ��� [DZUP]. Geographical distribution. Nova Friburgo, Sans Souci, Rio de Janeiro, Brazil (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860