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The first known Interatheriinae (Interatheriidae, Notoungulata) were described by Florentino Ameghino based on fossils collected by his brother Carlos in diverse Argentinean early Miocene outcrops in the Santa Cruz Province, which are housed at different institutions. In order to perform a systematic study of the subfamily, it is essential to revise as much specimens as possible, but first of all the type material of the species under study. Due to the fact that Florentino Ameghino never referred to any type specimens with a collection number in any of his publications, their identification becomes a complex task. In consequence, when studying the species erected by Ameghino within the genera Interatherium and Icochilus, we identified a lack of correspondence between some materials catalogued as type specimens and the original descriptions. In this contribution, we present the type specimens of 18 out of the 21 Santacrucian species of Interatherium (4) and Icochilus (14) within the paleontological collection of the Museo de La Plata and Ameghino National Collection at the Museo Argentino de Ciencias Naturales "Bernardino Rivadavia". Four case studies are presented: a) specimens correctly identified; b) materials erroneously catalogued as type specimens; c) specimens catalogued as type, whose status cannot be evaluated because they are lost; and d) specimens not catalogued as types, but herein recognised as such. As the selection of the type species of Icochilus was pending, Ic. extensus is herein designated as such. The lectotypes of Ic. extensus and Ic. multidentatus are selected among their identified syntypes.

New rhino remains recovered from Cova del Rinoceront (Castelldefels, Barcelona) confirm the presence of Stephanorhinus hundsheimensis (Toula, 1902) at the site and the taxon's persistence until the late Middle–early Upper Pleistocene in Europe, that is, its latest documented occurrence. The three individuals recovered from the site are compared with specimens of other Pleistocene species, including those of S. hemitoechus, S. kirchbergensis and Coelodonta antiquitatis, but their anatomical characteristics (a long skull, moderate occipital elevation, partial nasal septum, and slender zygomatic arch) do not coincide with the latter's documented features. Certain similarities are found with the most frequently occurring rhinocerotid at that time in the Iberian Peninsula, S. hemitoechus, but the cranial features of the latter differ. The anatomical characteristics of the Cova del Rinoceront individuals coincide most closely with those of S. hundsheimensis (i.e., a high occipital face, with rounded proximolateral angles and oblique lateral borders, as well as the frontoparietal angle, and facial development). Despite the marked overlaps in the general measurements of S. hundsheimensis and S. hemitoechus, many (cranial and postcranial) dimensions of the Cova del Rinoceront individuals coincide more closely with those of the former, although some bone proportions are more similar to those of the latter specimens. Therefore, S. kirchbergensis and C. antiquitatis can be discarded as they tend to be larger, more robust species. [ABSTRACT FROM AUTHOR]

This is an updated contribution to the Quaternary geology and paleontology from central-western Argentina, focused on San Luis Province. It is mostly based on unpublished data; only some fossils had previously been briefly mentioned in broader faunal contexts. The fossil-bearing sediments correspond to eolian and alluvial environments of moderate energy, dominated by sands and sandy silts. They overlie high-energy fluvial cycles and underlie edaphic horizons. They have a wide distribution, and several radiocarbon dates allow their regional correlation. Stratigraphic sequences with the precise origin of fossils allow for the improvement of lithostratigraphic and faunal correlations with the Pampean Region (central and east Argentina; La Pampa and Buenos Aires provinces), where Pleistocene assemblages are better known, but also with central-western (Cuyo Region), northwestern, and northeastern Argentina. Faunal remains correspond to large mammals, represented by xenarthrans (Cingulata and Tardigrada), macraucheniids (Litopterna), gomphotheres (Proboscidea), and equids (Perissodactyla), a typical Pleistocene mixture of native (xenarthrans and litopterns) and immigrant mammals. [ABSTRACT FROM AUTHOR]

We describe new remains of notoungulates from the Cerro Azul Formation (late Miocene) in La Pampa Province and reassign others previously studied, helping to precise the taxonomic diversity of 25 fossiliferous localities of this formation. Geomorphological context previously described for the area provides data to place some localities in a temporal sequence. The following Notoungulata are recognised: Protypotherium minutum (Interatheriidae), Paedotherium minor, Tremacyllus impressus, Hemihegetotherium achataleptum, H. torresi, and Hemihegetotherium sp. (Hegetotheriidae), Typotheriopsis silveyrai, T. chasicoensis, and Pseudotypotherium subinsigne (Mesotheriidae), and Pisanodon nazari, Hemixotodon chasicoensis, and cf. Stenotephanos plicidens (Toxodontidae). This study allows adding new localities to the biochronological sequence previously proposed based on the evolutionary pattern of the octodontoid rodents. Chasicoan and early Huayquerian (as herein considered) faunal assemblages present higher diversity of notoungulates than those assigned to late Huayquerian. Some localities lack a representation of taxa significant enough to precise their biochronological position within the late Miocene. The difference in the notoungulates representation along the Chasicoan–Huayquerian ages may be due to climatic changes that affected the centre of Argentina from late Miocene to early Pliocene. [ABSTRACT FROM AUTHOR]

We revise the mammal fossils from the Fray Bentos Formation previously assigned to Notoungulata, as well as some new remains recovered from levels cropping out in Corrientes Province (Argentina). In some cases, previous determinations have been completed to the species level, but others have been substantially modified, identifying different families. At present, identified taxa are: "Prohegetotherium" schiaffinoi (Hegetotheriidae), an undetermined interatheriid that is hypothesized as representing the upper teeth of Eopachyrucos ranchoverdensis, and Argyrohyrax proavus (Interatheriidae), cf. Archaeohyrax suniensis (Archaeohyracidae), cf. Mendozahippus fierensis ("Notohippidae"), a Leontiniidae indet., and some undetermined Notoungulata. The presence of Leontiniidae is based on the specimen CTES-PZ 7871, but those previously assigned to this family are herein discarded as such and reinterpreted (notohippid and undetermined taxon). The Mesotheriidae are discarded among the revised materials, but they are present in the Fray Bentos Formation in Corrientes Province with Trachytherus spegazzinianus (= Ameghinotherium curuzucuatiense). The revised sample presents affinities with the fauna from the type locality of the Fray Bentos Formation, in Uruguay, and the Deseadan assemblages from Quebrada Fiera (Mendoza Province, Argentina), Patagonia (Argentina), and Salla (Bolivia). These similarities are important from a paleobiogeographic point of view, especially by the recognition of cf. M. fierensis, a species only known from Mendoza up to now. Only the finding of new material from Fray Bentos Formation in Corrientes Province will allow us to corroborate these tentative determinations and a possible eastward extension of up-to-now exclusive taxa from Quebrada Fiera. [ABSTRACT FROM AUTHOR]

New research in the petroliferous area of Tupungato, Mendoza Province, central-western Argentina, has revealed a much richer fossil assemblage than previously known for this zone. First mammal remains from this area go back to the 1930s, but only three taxa were recognized since then, two notoungulates, the mesotheriid Typotheriopsis silveyrai and the hegetotheriid Hemihegetotherium achathaleptum , and the caviid rodent Cardiatherium paranense. A recent project, focused on geological and paleontological aspects of the Tupungato – Piedras Coloradas Anticlines, has allowed us to improve the geological knowledge of the area and increase its faunal diversity. This is one of the best localities for the study of the uppermost section of the Cacheuta foreland basin, and we present the first detailed section of the Río de los Pozos Formation (including part of previously considered Mogotes Formation). Concerning fossils, xenarthrans (sloths and armadillos), pachyrukhine hegetotheriids, a toxodontid, and different families of rodents (Chinchillidae, Octodontidae, Caviidae Dolichotinae, and Abrocomidae) are recognized for the first time in this locality, together with new remains of T. silveyrai , H. achathaleptum , and C. paranense. The comparison with Late Miocene–Pliocene Argentinean faunas reveals general similarities with most assemblages assigned to the Huayquerian Stage/Age, somewhat closer to those from the Cerro Azul Formation, and there are also shared taxa with Early Pliocene faunas. The biostratigraphic data presented here agree with published ages obtained for the Río de los Pozos Formation in the La Pilona Anticline. • The first detailed section of the Río de los Pozos Formation is presented. • Mammals from Tupungato–Piedras Coloradas Anticlines (Mendoza) reveal a high diversity. • Similarities with Late Miocene–Early Pliocene faunas indicate the Huayquerian Stage/Age. • Biostratigraphical data agree with absolute ages near the Anticlinal Tupungato area. [ABSTRACT FROM AUTHOR]

During the last decade, the Deseadan (late Oligocene) Quebrada Fiera locality, Mendoza Province, Argentina, has provided a large amount of mammal remains. Taxonomic studies have shown the presence of faunal elements common with other Deseadan associations from Patagonia and lower latitudes (Salla, Bolivia), as well as endemic taxa of different groups of mammals (Notohippidae, Leontiniidae, Homalodotheriidae, Hegetotheriidae, and Metatheria) and even a gastropod. In this work, we present a taxonomic and phylogenetic analysis of a set of postcranial fossils of three families of the suborder Toxodontia (Notoungulata). The postcranial elements are assigned to taxa previously recognized at Quebrada Fiera, such as the leontiniid Gualta cuyana and the homalodotheriid Asmodeus petrasnerus, but also to the family Toxodontidae that is represented by Proadinotherium sp. and another larger toxodontid; a few dental remains of Proadinotherium are also included as this is the first time that toxodontids from Quebrada Fiera are described. In the case of A. petrasnerus, an almost complete calcaneum allows us to expand the diagnosis of this taxon. The bones assigned to G. cuyana enlarge the anatomical knowledge of this species. In turn, the presence of the genus Proadinotherium extends its geographic distribution in Argentina and adds to the extra-Patagonian record of P. saltoni from Salla, Bolivia. Phylogenetic results do not differ much from previous analyses using postcranial characters, and the obtained consensus trees show low node supports. The various phylogenetic analyses performed here provide a more robust framework to interpret the relationships of the studied taxa. [ABSTRACT FROM AUTHOR]

Abstract: This paper deals with the detailed taxonomic study of the first mammal remains from the Mariño Formation in the Divisadero Largo area, Mendoza, central west Argentina. This area was well-known by the fossils recovered from the Triassic levels and particularly by the faunal assemblage from the Cenozoic levels corresponding to the Divisadero Largo Formation. Until recently, no mammal remains had been accurately reported for the Mariño Formation, Miocene in age. The specimen consists in an almost complete upper jugal series (teeth found separated from the maxillary bone except the M3) of a small Mesotheriidae (Notoungulata), which was preliminary identified as Mesotheriinae indet. The main characteristic of these teeth is their small size, significatively lesser than that of the Miocene genera Eutypotherium, Typotheriopsis and Pseudotypotherium, being closer to Mycrotypotherium from Bolivia and the Chilean species from the Chucal Formation. The direct comparison with these latter forms has allowed a more detailed study and the recognition of the material here described as cf. Altitypotherium chucalensis, with all cautions inherent to the incompleteness of the material. [Copyright &y& Elsevier]

In 1889, Florentino Ameghino erected Patriarchus palmidens, based on a specimen from a western outcrop of the Santa Cruz Formation (early Miocene) and associated it with the 'Notohippidian stage'. Later, Ameghino added seven other species to the genus from eastern localities of the same formation. Since 1900, Patriarchus has been considered to be a junior synonym of Protypotherium. Here, we present a systematic revision and recognize the genus Patriarchus as monospecific, with Pa. palmidens characterized by i1–c mesiodistally expanded, labially convex, and similar in size; i1–c with a short lingual sulcus that does not extend to the base of the teeth and 'V'-shaped in occlusal view; p1 labially and lingually bilobate, with trigonid larger than talonid; and i1–p1 in a continuous series without overlapping. Five of the remaining species are recognized as belonging to Protypotherium. Therefore, Patriarchus is presently monospecific (Pa. palmidens) and restricted to the western region of Santa Cruz Province, particularly to the Río Bote locality. Patriarchus is included for the first time in a cladistic analysis to discover its phylogenetic relationships within the Interatheriidae. It is nested within the most derived Interatheriinae and is recovered as the sister group of the Laventan Miocochilius; this clade is, in turn, the sister group of Cochilius plus Interatherium. These results support the hypothesis that the origin of the Miocene Interatheriines was a single evolutionary event. [ABSTRACT FROM AUTHOR]

The specimens described herein come from the Río Bote locality, Santa Cruz Formation (early Miocene), Santa Cruz Province, Argentina. They include the most complete cranial material of the 'Notohippidae' Notohippus toxodontoides and other remains that increase knowledge of the mandible and dentition. The new remains allow extension of the original description proposed by Ameghino in the 19th century and provide data on ontogenetic variation in the dental traits of this species. Notohippus differs from the Deseadan–Colhuehuapian genus Argyrohippus in the following features: discontinuous dentition; C and p1 absent; shorter P2–M3/p2–m3 series; crown of I1 highly curved; upper molars with less developed parastyle and paracone fold, less undulating ectoloph, deeper distolingual groove of M3 directed more lingually and presence of a very shallow distolabial groove; incisiform lower canine; and lower molars with posterolabial angle of talonid more rounded and narrower trigonid valley. As stated in previous works, our phylogenetic analysis indicates that the 'Notohippidae' do not constitute a monophyletic group, because Argyrohippus, Notohippus, and Toxodontidae are recovered as a clade. Internal relationships within this clade are not resolved in any of the phylogenetic analyses performed here. The presence of N. toxodontoides in the faunal assemblage found at the Río Bote locality adds to its previous records in Santa Cruz (Kar-Aiken and Cerro Centinela localities) Province, Argentina, and Sierra Baguales locality in Chile. [ABSTRACT FROM AUTHOR]

Cerro Patagua is a locality with fossiliferous outcrops of the Cerro Azul Formation in La Pampa Province (central Argentina). First fossil remains were known in 1999 and assigned to the Chasicoan Stage/Age (Late Miocene, Tortonian) after the presence of the Octodontoidea rodent Chasichimys bonaerense. Later references to vertebrate remains were scarce and mostly limited to some mentioned taxa in broader faunal contexts. Now, we present a detailed taxonomic study of the entire mammal assemblage from Cerro Patagua, which evidences that most taxa are typical of the Chasicoan Stage/Age. A peculiar enterolith-like structure, recovered from loess levels, is also described, and several taphonomic aspects observed on fossils are related to their level of provenance. In addition, we offer stratigraphical interpretations of the Cerro Azul Formation and a geomorphological analysis of the Cerro Patagua area, and provide a detrital zircon dating of 12.3 ± 1.8 Ma. This result reflects the time interval 14.1–10.5 Ma, a temporal range that includes the Serravalian Stage (late Middle Miocene) and could indicate that the oldest synorogenic deposits could have begun to accumulate at the end of this stage. Even considering the minimum age of this interval (Tortonian Stage), this dating is the oldest obtained for the Cerro Azul Formation, compared with previous published data from Arroyo Chasicó locality (9.43–9.07 Ma). This assignment could explain that taxa from Cerro Patagua show affinities with Late Miocene mammal assemblages from central Argentina, as well as with older faunas from Patagonia and Bolivia. • The oldest exposed level of Cerro Azul Formation (central Argentina) is described. • SynOrogenic deposits at Cerro Patagua begun to accumulate around 12 Ma. • Taxonomic evaluation indicates a Chasicoan Stage/Age for the Cerro Patagua assemblage. [ABSTRACT FROM AUTHOR]

Late Oligocene caviomorphs represent one of the most important early adaptive radiations of this group of rodents. They show a high taxonomic diversity and a wide geographic distribution, with those found in the upper Oligocene of Salla (Bolivia) and Patagonia (Argentina) being particularly diverse. However, caviomorphs from the Deseadan South American Land Mammal Age in other latitudes are still scarcely known. In this contribution, we describe new caviomorph rodents from the Deseadan levels of Quebrada Fiera locality (Mendoza Province; central-western Argentina) and evaluate their systematic and paleobiogeographic significance. The taxonomic composition of the caviomorphs from Quebrada Fiera reveals a complex biogeographic history, as we recognize one endemic Cavioidea (Asteromys puelche sp. nov.) and two Chinchilloidea: one with Patagonian affinities (Cephalomys sp.) and another with a wide distribution in South America (Incamys bolivianus). Asteromys puelche sp. nov., shows closer phylogenetic affinities with the Bolivian A. bolivianus than with the Patagonian A. punctus. These rodents, in common with the whole mammalian association from Quebrada Fiera, show a mix of endemic taxa and others shared with higher and lower latitudes. [ABSTRACT FROM AUTHOR]

We present two new Paleogene fossiliferous sites that provide the first mammal remains for the Malargüe Group in the Neuquén basin of southeastern Mendoza Province, Argentina. Identified taxa from Agua de Flores-Agua de Isaac: Kibenikhoria sp., Oldfieldthomasiidae indet., ?Henricosborniidae indet., and a large Notoungulata probably related to the Isotemnidae. Taxa from Liu Malal: cf. Kibenikhoria and Notopithecidae gen. et sp. nov. Based on facies composition and geological distribution, we consider that the Cenozoic deposits from these fossiliferous localities correspond to the already defined Puesto Fortunata Formation. This unit is correlated to the Coihueco Formation, the top of the Malargüe Group in the northern Neuquén basin, and its mammal content supports the proposed hiatus between the ~40 Ma and 20 Ma based on chronostratigraphy. The presence of Kibenikhoria in the local faunas of southeastern Mendoza extends the geographic range of this genus so far known in Patagonia to central-west areas of Argentina and supports an early-middle Eocene age for the fossil-bearing levels of the Puesto Fortunata Formation. • First report of Eocene mammal for the Malargüe Group, Neuquén basin, Mendoza. • The fossil bearing levels belong to the Puesto Fortunata Formation. • Fossils are recognized as henricosborniid, notopithecid, oldfieldthomasiid, and isotemnid. • These mammal groups imply a faunal relationship between Mendoza and Patagonia. • Evidence reinforce a early–middle Eocene age for the fossil-bearing levels. [ABSTRACT FROM AUTHOR]

Abstract: This paper deals with the taphonomic analysis of the late Miocene bone assemblage from the Cerro Azul Formation at Telén (La Pampa Province, Argentina). The faunal assemblage was assigned to the Huayquerian mammal age (late Miocene). The fossiliferous section shows a homogeneous lithology, and is interpreted as a loess deposit with two similar and slightly developed palaeosols, classified as calcic vertisols. The studied sample comprises 5598 remains anatomically and taxonomically determined. They were collected from an area of about 48,000m2, appearing randomly distributed through the section and with low density. Most remains are small- to very small-sized, disarticulated, and very fragmented. Different taphonomical histories are inferred for microvertebrates and macromammals. The microvertebrate assemblage is interpreted as the result of predator activities. After a brief period of pre-burial exposure, remains were dispersed from the original depositional area. On the other hand, a natural and gradual death process is envisaged for macromammals, followed by a long period of exposure to weathering and dispersal by physical agents. Remains of both groups, once buried, suffered the diagenetic processes of the host rock. Consequently, the fossil assemblage from Telén would represent a condensed assemblage corresponding to two distinct time spans, i.e., the accumulation of microvertebrates took place in a short time interval whereas that of macromammals occurred over a longer period, coincident with the development of both soils. [Copyright &y& Elsevier]