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The Balkans are considered a major glacial refugium where flora and fauna survived glacial periods and repopulated the rest of Europe during interglacials. While it is also thought to have harboured Pleistocene human populations, evidence linking human activity, paleoenvironmental indicators and a secure temporal placement to glacial periods is scant. Here, we present the first intra-tooth multi-isotope analysis for the European straight-tusked elephant Palaeoloxodon antiquus, on an adult male individual excavated in association with lithic artefacts at the MIS 12 site Marathousa 1 (Megalopolis basin, Greece). The studied find also exhibits anthropogenic modifications, providing direct evidence of hominin presence. We employed strontium, carbon and oxygen isotope analysis on enamel bioapatite to investigate its foraging and mobility behaviour, using a sequential sampling strategy along the tooth growth axis of the third upper molar, to assess ecological changes during the last decade of life. We found a geographically restricted range, in a C3-dominated open woodland environment, and relatively stable conditions over the examined timeframe. Our results show that, despite the severity of the MIS 12 glacial, the Megalopolis basin sustained a mesic habitat, sufficient plant cover and limited seasonal fluctuations in resource availability, pointing to its role as a glacial refugium for both fauna and hominins. [ABSTRACT FROM AUTHOR]

In contrast to a relatively long history of Palaeolithic investigations in western Europe, research on the Palaeolithic period in Greece has lagged behind considerably. This article reviews the last decade of Palaeolithic research in Greece, with the aim of highlighting key aspects of recent developments in the field. Newly discovered Lower Palaeolithic sites, such as Marathousa 1 in Megalopolis, have offered rare, high-resolution windows into hunter-gatherer adaptations during the earliest-known peopling of the Greek peninsula. Palaeolithic sites in insular settings, exemplified by the latest discoveries in Crete and Naxos, have stirred up intriguing discussions about early seafaring but, most importantly, provide support to a revised view of the role of the Aegean in early human dispersals. Zooarchaeological, palaeoenvironmental and dating analyses of Middle and Upper Palaeolithic materials from new and older assemblages have provided valuable insights that help contextualize the information distilled from lithic industries. In sum, recent excavations, surveys and assessments of new and older collections have together contributed to the compilation of an important corpus of novel and significant data. Palaeolithic Greece is no longer a terra incognita, and it carries the potential to become a key player in understanding early human societies in southern Europe. [ABSTRACT FROM AUTHOR]

Abstract We investigated the magnetostratigraphy of the Megalopolis basin in central Peloponnese, Greece, which encompasses a record of Pleistocene lacustrine and lignite-bearing sedimentation, where lithic tools stratigraphically associated with remnants of an almost complete skeleton of Palaeoloxodon antiquus were recently found at the Marathousa 1 site. A magnetic polarity reversal was observed within a ∼10 m-thick lignite seam at the base of the (exposed) stratigraphic sequence, and it was interpreted as a record of the Brunhes/Matuyama boundary (0.78 Ma). Assuming that lignite seams were deposited generally under warm and humid climate conditions, this finding is in agreement with data from the literature indicating that the Brunhes/Matuyama boundary occurs within warm Marine Isotope Stage (MIS) 19. We then attempted to correlate the remainder of the lacustrine and lignite-bearing intervals above the Brunhes/Matuyama boundary to a standard oxygen isotope record of Pleistocene climate variability. Two age models of sedimentation were generated: according to preferred option #1, the artifact-bearing stratigraphic units of the Marathousa 1 site should have an age between ∼0.48 Ma and ∼0.42 Ma, while according to alternative option #2, the archaeological layers would have an age between ∼0.56 Ma and ∼0.54 Ma. Option #1 is at present considered the preferred option as it is in closer agreement with preliminary post-IR IRSL and ESR dates from the Marathousa 1 site. This age model has been exported to other areas of the Megalopolis basin, where additional archaeological and/or palaeontological sites could be present, by means of correlations to lithostratigraphic logs derived from commercial drill cores taken in the 1960s and 1970s for lignite exploitation. [ABSTRACT FROM AUTHOR]

Abstract The PaGE Project survey of the Megalopolis Basin (Arcadia, Greece), conducted in 2012–2013 over a period of two field seasons, was an intensive, target-oriented surface survey of Pleistocene sediments. Implementing a modified version of field methods applied in our survey of cave systems and Pleistocene cave sediments, the main goal of this systematic research focused on the identification of intact stratified remains from either stratigraphic sections or on remnants of exposed ancient surfaces. The project area consisted of the active open-cast lignite mine of Megalopolis, where access to recently exposed sections were abundant, and of the alluvial fans in the surrounding uplands, where hominins exploited the commanding views of the ancient lake. This paper presents the survey results and applied field methods for conducting archaeological research within the remnants of an Early to Middle Pleistocene ancient lake system. The project succeeded in identifying at least five surface and stratified sites, the latter including the Lower Paleolithic remains of Marathousa-1. This site is currently under investigation, but has already yielded the oldest chronometrically dated archaeological remains in Greece. Therefore, PaGE has not only managed to place mainland Greece on the Lower Paleolithic map of Europe, but has also successfully tested a methodological corpus for survey research targeting the earliest phases of the Paleolithic period in a Mediterranean landscape. [ABSTRACT FROM AUTHOR]

Abstract The present study describes the sedimentology and formation processes of the archaeological site Marathousa 1, part of a Pleistocene lignite-bearing succession at the Megalopolis basin (Southern Greece). The sedimentary sequence of the site comprises about 4–5 m lacustrine and fluviolacustrine clastic deposits found between Lignite Seam II and III. The lower part of this sequence is characterized by relatively high rate subaqueous sedimentation of bedded sands and silts with local evidence of slumping and liquefaction attributed to a seismic event. This part of the sequence contains fluctuating but generally low organic and carbonate content probably associated with cold and arid conditions of a glacial period. The upper part of the sequence follows a major hiatus attributed to exposure and erosion. A series of erosional bounded depositional units are observed in this sequence suggesting important water-level fluctuations. The subaqueous emplacement of the deposits is attributed to subaerial flood-generated, organic- and carbonate-rich dilute mudflows and hyperconcentrated flows. Organic-rich sedimentation culminates with the formation of the overlying lignite seam. This change is interpreted as a gradual return to warmer and humid conditions of an interglacial period. The fossiliferous and cultural levels at Marathousa 1 are associated with the major erosional and exposed surface found in the upper part of the sequence. This surface was part of an extensive mudflat surrounding the lake shore at that time. The overlying mudflows have locally redistributed and buried the archaeological remains, thus preserving this very important butchering site. [ABSTRACT FROM AUTHOR]

Abstract Marathousa 1 is a Lower Palaeolithic open-air site located in the Megalopolis basin, an area in Southern Greece known for its fossiliferous sediments. Mining activities in the basin uncovered a thick sequence of Middle Pleistocene lacustrine deposits representing the environment of a palaeolake. Marathousa 1 was discovered in 2013 during a targeted palaeoanthropological survey and excavated subsequently by an interdisciplinary team from the Ephoreia of Palaeoanthropology–Speleology of Greece and the University of Tübingen, Germany. This article presents results from the ongoing investigation and reviews the state of knowledge about the site. Systematic excavations during five field seasons have exposed a total of 72 m2 and revealed a partial skeleton of the elephant Palaeoloxodon antiquus and remains of other large mammals in spatial and stratigraphic association with a "small tool" lithic assemblage. Faunal and taphonomic studies indicate the presence of cut-marks and percussion damage on elephant and other large mammal bones. The study of site formation processes, together with taphonomic and geostatistic spatial analyses confirm the association of fossil and hominin activity remains and the stratigraphic integrity of the site. Radiometric dating, geological and biostratigraphical evidence suggest that hominin activity at the site occurred between 0.5 and 0.4 Ma. Marathousa 1 is the oldest currently known archaeological site in Greece and the only Lower Palaeolithic butchering site in the Southern Balkans. It is also a key site for documenting high resolution palaeoclimatic, palaeoenvironmental and cultural records of a geographical area that potentially acted as a refugium during the successive waves of hominin colonization of Europe. [ABSTRACT FROM AUTHOR]

Abstract The technological systems and subsistence strategies of Middle Pleistocene hominins in South-East Europe are insufficiently understood due to the scarcity of well-preserved, excavated assemblages. In this paper, we present first results from the study of the lithic and bone artifacts unearthed at the Lower Palaeolithic site Marathousa 1 (MAR-1), Megalopolis, Greece. The context of the site represents a depositional environment close to a lakeshore, where rapid burial in a very fine-grained matrix ensured extraordinary conditions for preservation. Lithic artifacts occur in spatial and stratigraphic association with remains of the elephant Palaeoloxodon antiquus as well as other mammals. Bones, including those of elephants, show clear anthropogenic flaking scars, cut-marks and fracture patterns indicating deliberate breakage and modification by early humans. The MAR-1 lithic assemblage is composed of small-sized debitage, retouched tools, a few small and exhausted cores, as well as a large number of debris and retouch products, such as chips and resharpening flakes. Currently, there are no indications of Acheulean bifacial debitage, large cutting tools are missing, and a key aspect of the material refers to its 'microlithic' character. The scarcity of cores and primary flakes indicates a fragmented reduction sequence and complex discard patterns that require further investigation. On the basis of the on-going analysis of lithic material from three field seasons, we discuss aspects of assemblage composition and the role of raw material types, the main technological and typological traits of the industry, as well as the potential contribution of the MAR-1 assemblage in broader discussions about Middle Pleistocene lithic techno-complexes and subsistence strategies in Eurasia. Finally, we briefly present a small sample of bone artifacts, which suggest that hominin exploitation of the animal carcasses was not restricted to marrow extraction and bone processing for nutritional needs, but included also the knapping of bones, potentially with the aim of using the knapped products as tools. The MAR-1 archaeological record compares well with other important Eurasian sites yielding 'small tool assemblages', such as Ficoncella, La Polledrara, Bilzingsleben, Schöningen and Vértesszőlős, some of which, like MAR-1, have provided evidence of elephant or other mega-fauna exploitation. [ABSTRACT FROM AUTHOR]

Abstract Recent excavations at the Middle Pleistocene open-air site of Marathousa 1 have unearthed in one of the two investigated areas (Area A) a partial skeleton of a single individual of Palaeoloxodon antiquus and other faunal remains in spatial and stratigraphic association with lithic artefacts. In Area B, a much higher number of lithic artefacts was collected, spatially and stratigraphically associated also with faunal remains. The two areas are stratigraphically correlated, the main fossiliferous layers representing an en mass depositional process in a lake margin context. Evidence of butchering (cut-marks) has been identified on bones of the elephant skeleton, as well on elephant and other mammal bones from Area B. However, due to the secondary deposition of the main find-bearing units, it is of primary importance to evaluate the degree and reliability of the spatial association of the lithic artefacts with the faunal remains. Indeed, spatial association does not necessarily imply causation, since natural syn- and post-depositional processes may equally produce spatial association. Assessing the degree and extent of post-depositional reworking processes is crucial to fully comprehend the archaeological record, and therefore to reliably interpret past human behaviours. The present study uses a comprehensive set of spatial statistics in order to disentangle the depositional processes behind the distribution of the archaeological and palaeontological record at Marathousa 1. Preliminary results of our analyses suggest that a high-energy erosional process, attributed to a hyperconcentrated flow deposited at the margin of a swamp, reworked an autochthonous, exposed or slightly buried, scatter of lithic artefacts and faunal remains. Minor reworking and substantial spatial association of the lithic and faunal assemblages support the current interpretation of Marathousa 1 as a butchering site. [ABSTRACT FROM AUTHOR]

Southern Europe is hypothesized to have acted as a glacial refugium for hominin populations during the Pleistocene. Of particular importance is South-East Europe, which most likely played a dual role, both as refugium and dispersal corridor, especially during the Middle Pleistocene glaciations, when drastic climatic conditions led to major sea level drops in the Aegean. However, little is known about the palaeoenvironmental conditions at the time of hominin presence in this region, making these hypotheses difficult to test. Here we analyze biomarker data and leaf wax stable isotopic compositions of the MIS 12 Lower Palaeolithic site Marathousa 1 (Megalopolis Basin, Greece) to assess the climatic conditions accompanying the time of hominin presence in the area. Our data indicate a major cooling affecting the north Mediterranean/Aegean domain during this time interval, with lowest temperatures recorded between ∼440–432 ka. The glacial peak is associated with changes in vegetation (i.e., from more forested to more open landscape), reduction of humidity and water availability (i.e., moisture depletion, increased evaporation). Hominins are present at the Marathousa 1 location at the end of this interval (434–432 ka), confirming that the Megalopolis Basin served as a refugium for hunter-gatherer groups during periods of harsh climatic conditions. Additionally, the progressive cooling is associated with an important sedimentary hiatus between ∼465–440 ka reflected in all circum-Mediterranean records (both marine and continental), indicating a regional impact of the MIS 12 glaciation over surface processes. • Hominin presence corelates with the end of the glacial maximum. • Highly evaporative intervals are associated with shifts in moisture and vegetation. • Overlap of regional and global signals. • Megalopolis basin was a refugium for biota during MIS 12. [ABSTRACT FROM AUTHOR]

Abstract The mining activities in the Middle Pleistocene lacustrine basin of Megalopolis (Peloponnesus, Greece) have exposed expanded sections of lacustrine sediments. In particular, the northernmost mine of Kyparíssia has yielded numerous vertebrate fossils, recovered during field surveys and small-scale rescue excavations. The stratified specimens indicate the presence of at least two fossiliferous horizons, which are stratigraphically closely situated. The fossils are generally widely dispersed, but they are much more abundant along the western margin of the palaeolake. Four sites showing denser concentration of fossils were located and partially excavated during the 2004–2008 fieldwork. An additional site was identified and excavated in 2012. The fossils were found in organic-rich sediments (mostly lignites), indicating that they were deposited during a warm and humid (i.e., interglacial) time period, in a richly vegetated environment. The recovered fauna is diverse, dominated by hippopotamuses, deer and elephants. The present study presents the chelonian, avian and suiform finds. The following taxa are recognised: Emys orbicularis , Testudo marginata , Sus scrofa , Hippopotamus antiquus , as well as a diverse sample of aquatic birds. This faunal composition indicates a temperate, woodland/forest environment, with continuous presence of a large water body, whilst the presence of the darter (Anhinga sp.) points to relatively warm conditions. Biochronologically it points to an age within the early part of the Middle Pleistocene. [ABSTRACT FROM AUTHOR]