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Abstract: The dispersal of hominins may have been favored by the opening of the landscape during the Early–Middle Pleistocene transition (EMP) in Western Europe. The structure of the small-vertebrate assemblages of the archaeo-paleontological karstic site of Gran Dolina in Atapuerca (Burgos, Spain) shows important environmental and climatic changes in the faunal succession, across the Matuyama–Brunhes boundary at 780 ka. These changes are interpreted to indicate impoverishment of the forests, along with an increase in dry meadows, and open lands in general that entailed a tendency towards the loss of diversity in small-vertebrate communities above the EMP. We evaluate variation in diversity of the faunal succession of Gran Dolina using Shannon’s Second Theorem as an index of ecosystem structure. The long cultural-stratigraphic sequence of Gran Dolina during the EMP is somewhat similar in its completeness and continuity to that in the locality of Gesher Benot Ya''aqov in the Upper Jordan Valley. We also evaluate related data including faunal and floral (pollen) succession. Both localities present cold, dry and humid, warm fluctuations at the transition between the Early and the Middle Pleistocene. Comparisons between these sites present opportunities to understand large-scale climatic changes. [Copyright &y& Elsevier]

Abstract: In addition to the Neanderthals, other mammal species vanished locally or totally in southern Europe during relatively a short period, between about 40 and 10kyr. The late Upper Pleistocene was in general a time of extinction and extirpation in southern Europe, ranging from musk-oxen and reindeer to Nordic voles (extirpations) to Pliomys and Neanderthals (extinctions). The southern regions of Europe have been glacial refugia for several mammalian species (including humans), and when climatic conditions ameliorated, some spread back to the North and East European Plains (reindeer, saiga, the marsh voles and narrow-skull voles, among others), whence they originally had come. Others remained at their refugia in southern France and on the Iberian, Italian and Balkan Peninsulas, and, after periods of survival, eventually became extinct (Homo neanderthalensis, Pliomys lenki). The demise of the Neanderthals is, of course, well known, but that of the extinct vole, P. lenki, is not well known at all. Their long survival in southwestern European regions must be related to the existence of suitable climatic, vegetational and geographic conditions. Did these regions become traps when new competitors arrived? [Copyright &y& Elsevier]