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Löbl, Ivan (2023): Assessing Earth's biotic diversity in natural history museums - gone with the wind? Phegea 51, No. 2: 50-51, DOI: 10.6084/m9.figshare.22722733, URL: http://dx.doi.org/10.6084/m9.figshare.22722733

Members of the scaphidiine tribe Scaphisomatini collected recently in Tianmushan Biosphere Reserve (Zhejiang) and Wuyishan National Nature Reserve (part of Wuyishan Biosphere Reserve, Fujian), China, are studied. Two species, Scaphisoma krali Löbl, sp. nov. and Scaphisoma sekerkai Löbl, sp. nov., are described from Wuyishan. Baeocera franzi (Löbl, 1973) and Scaphisoma mutator Löbl, 2000 are recorded for the first time from Zhejiang province, and Scaphisoma binhanum (Pic, 1922) and Scaphoxium intermedium Löbl, 1984 are recorded for the first time from Fujian province.

Sapitia versicolor (Pic, 1920) Material examined. 474, Sabah, Batu Punggul Resort env., 24.vi.̅1.vii.1996, 11c, FIT (MHNG); 3, Sabah, Danum Valley, 14̅ 16.ii.1968, G. de Rougemont (MHNG); 1, Sabah, Poring Hot Springs, 500 m, 11.v.1987, Burckhard & Löbl (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05°02.682′ E 117°45.553′, 250 m, iii̅ iv.2005, FIT, primary dipterocarp forest, E. Slade & J. Villanuava lgt. OUMNH 2055 ­062 (OUMNH). Remarks. Thie species appears to be widely distributed and has been reported from Indonesia: Sumatra; East Malaysia: Sabah and Thailand (see LÖBL 2018). It is very similar to S. sumatrana Löbl, 1978 while it differs notably from S. lombokina by the nearly fused mesepimera and the stoud legs., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 16, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2018: Coleoptera: Staphylinidae: Scaphidiinae. World Catalogue of Insects. Volume XVI: i ‾ xvi + 1 ‾ 418 pp."]}

Scaphicoma rufa (Pic, 1923) Material examined. 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 450 m, 22̅ 23.xi.2005, primary dipterocarp forest, west trail area, first inferior FIT, D. Mann, E. Slade & J. Villanuava lgt., OUMNH 2006 ­051 (OUMNH); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, E. Slade & J. Villanuava lgt., selectively logged dipterocarp forest, Yayasan Sabah Logging Concesion, OUMNH 2005 ­052 (MHNG). Remarks. These specimens exhibit characters congruent with the redescription given by LÖBL & LESCHEN (2010). Notable are the widely interrupted basal striae of the elytra, the mesoventrite with distinct mesal ridge, the lateral parts of the metaventrite lacking coarser punctures, and the lateral parts of ventrite I, each bearing a subbasal patch of distinct punctures, in combination with the uniformly rufous, about 1.9 mm long body. The species is very similar to S. arcuatum Champion, 1927 and S. pulex Heller, 1917., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 17, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & LESCHEN R. A. B. 2010: Notes on the Toxidium group (Coleoptera: Staphylinidae: Scaphidiinae). Folia Heyrovskyana Series A 18 (1 - 3): 71 ‾ 93."]}

Pseudobironium rapax sp. nov. (Figs 8 ̅10) Type material. Holotype male, Malaysia, Sabah, Crocker Range, Mawar Waterfall env. 17.VI.1996, 9c, vegetation debris and forest floor litter accumulated along large fallen trees near river (MHNG). Description. Length 1.95 mm, width 1.30 mm. Head and most of body dark reddishbrown, apex of abdomen lighter, yellowish; femora and tibiae roughly as body, tarsi and antennae light brown. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows: III 15: IV 22: V 26: VI 21: VII 26: VIII 26: IX 27: X: 26: XI 35; antennomere XI about 3.5 times as long as wide. Pronotal punctation very fine, similar to that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 4 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation similar to pronotal. Hypomera impunctate, impressed posteriad. Mesoventrite lacking mesal ridge and lacking distinct setiferous punctures. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite lacking microsculpture; median area nearly flat, lacking impressions and tubercle, with two irregular rows of distinct admesal punctures. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, horizontal. Submesocoxal area short, about as fourth shortest interval to metacoxa. Submesocoxal line finely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Ventrite Ilacking microsculpture, with very fine and scattered punctures; following ventrites and exposed tergites with punctulate microsculpture. Male. Aedeagus (Figs 8 ̅10) 0.74 mm long. Median lobe with basal bulb shorter than apical process, the latter slightly inflexed, with sinuate ventral margin (lateral view), and blunt tip; apical part of median lobe lacking distinct valves. Parameres nearly evenly wide, bent in dorsal view, straight in lateral view. Internal sac with pair of strongly sclerotized denticles and mesal sclerite, lacking flagellum and lacking apical spinose structures. Etymology. The species epithet is a Latin noun, referring to the canine­like denticles of the internal sac. Differential diagnosis. The species is as P. irregulare less than 2 mm long and thus conspicuous by its small body­size. It may be easily distinguished from P. irregulare by the evenly very fine elytral punctation, the narrow submesocoxal areas and ventrite I lacking microsculpture. The aedeagal characters suggest relationship with members of the P. subovatum group (see LÖBL & TANG 2013), though the shape of the admesal sclerites of the internal sac is unique. The species falls in the key species to couplet 27, to P. schuhi Löbl & Tang, 2013 from which it differs notably by the parameres evenly narrow, except at their basis. Remarks. The single available specimen has its forelegs broken, probably lost when collected., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 14-15, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Mordelloscaphium Pic, 1915 The genus is monospecific. It was established to accommodate a species possessing characters typical for the termitophilous members of the Baeoceridium group (see LESCHEN & LÖBL 2005) and thus is assumed to live on fungus gardens of termites., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LESCHEN R. A. B. & LOBL I. 2005: Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1 ‾ 63."]}

Bironium borneense Löbl, 1987 Material examined. 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 27̅ 28.x.2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, West trail 10 FIT, OUMNH 2006­051 (OUMNH); 1, with the same data but 21.X.2005, West trail area (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, primary forest, Slade & J. Villanuava lgt. lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005­062 (OUMNH). Remarks. The species is known only from East Malaysia (LÖBL et al. 2020). It was based on specimens found in the Gunung Mulu National Park in Sarawak and subsequently recorted from the Mount Kinabalu National Park, the vicinity of Tambunan and the Batu Punggul Resort in Sabah., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I., LESCHEN R. A. B. & KODADA J. 2020: Review of the Asian species and cladistic analysis of Bironium Csiki (Coleoptera: Staphylinidae: Scaphidiinae) with comments on biogeography. Annales Zoologici (Warszawa) 70 (4): 711 ‾ 736."]}

Vituratella Reitter, 1908 The genus encompasses 31 species, all Afrotropical but the Asian V. termitophila (Champion, 1927) and V. kistneri (Löbl, 1979)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 18, DOI: 10.5281/zenodo.7501705

Scaphicoma Motschulsky, 1863 The genus comprises 19 Afrotropical, Oriental, Melanesian and Australian species. Several, described by J. Achard and M. Pick, still lack revisions. Asingle species is present in samples from Sabah., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 17, DOI: 10.5281/zenodo.7501705

Baeocera incisa (Löbl, 1973) Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57′ E 117°48′, 450 m, 1̅ 5.ix.2005, Slade & Villanuava lgt., lowland mixed dipterocarp forest. Nature trail area 1, FIT, OUMNH (OUMNH). Remarks. The species is widely distributed in Sabah and seems to be common in samples of forest floor litter (see LÖBL 2021: 28)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53 (1): 85 ‾ 89."]}

Baeocera doriae (Pic, 1020) Material examined. 1 male, Sabah, Mt. Kinabalu N. P., 1700 m, 16.vii.02, Kurbatov & Zimina (MHNG); 1 male, Sabah, rd Kota Kinabalu ̅ Tambunan, km 52, 1600̅ 1800 m, 2̅ 5.vii.02, Kurbatov & Zimina (MHNG). Remarks. This species is known from Sarawak and Sabah only. It is poorly represented in collections, a single specimen has been reported from each of these states., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 9, DOI: 10.5281/zenodo.7501705

Baeocera hutan sp. nov. (Figs 1 ̅4) Type material. Holotype male, Sabah: Lahad Datu Ulu Segama forest Reserve, Danum Valley For. Centre, N 04°57.9′ E 117°48.1′, 200 m, 21.xi.2005, D. Mann, E. Slade & J. Villanuava lgt., lowland mix. dipterocarp forest. West trail area 1, FIT, OUMNH 2006­051 (OUMNH). Description. Length 1.28 mm, width 0.76 mm. Head and body nearly evenly reddishbrown, elytra hardly darker than pronotum, apical abdominal segments slightly lighter than pronotum, femora and tibiae as elytra, tarsi and antennae lighter. Head with interocular distance nearly as dorsoventral eye diameter. Head, thorax and elytra lacking microsculpture. Length/width ratios of antennomeres as follows: III 20 /5: IV 20 /6: V 30 /7: VI 27 /7: VII 33 /8: VIII 27 /7: IX 36 /11: X: 37/15: XI 48 /17. Pronotal and elytral punctation even, very fine. Lateral contours of pronotum and elytra continuously, very weakly arcuate. Pronotum with anterior margin bead broadly interrupted, lateral margins sinuate in lateral view, lateral margin striae impunctate. Margin of exposed part of scutellum rounded. Elytra with lateral margins nearly parallel, lateral margins in lateral view slightly sinuate, lateral margin carinae visible in dorsal view, lateral margin striae impunctate; sutural striae starting at base, parallel, deeply impressed, curved along pronotal lobe to form basal striae extended to humeral areas, broadly separated from lateral striae; discal punctation sparse, near base with puncture intervals about 8 to 10 larger than puncture diameters. Hind wing fully developed. Hypomera and mesanepisterna appearing impunctate. Mesoventrite with mesal ridge, punctate along lateral margins. Mesepimeron about 4 times as long as large and 5 times as long as interval to mesocoxa. Median part of metaventrite nearly flat, impunctate mesally, with dense, fairly coarse punctures on posterior third and laterally smooth mesal area, bearing conspicuous, long pubescence. Lateral parts of metaventrite with few scattered extremely fine punctures. Submesocoxal line parallel, margined with row of coarse elongate punctures extended posterior the inner part of mesepimeron. Submesocoxal area about 0.02 mm long. Metanepisterna large, about 0.10 mm wide, twice as long as wide, slightly convex, with inner margin broadly rounded anteriorly, nearly straight posteriad; inner suture impunctate. Protibiae straight. Mesotibiae and metatibiae slightly bent. Ventrite I lacking striae, with row of elongate basal punctures not interrupted in middle, extended laterally behind tip of metepimeron; punctation posterior basal row very fine and sparse. Male. Protarsomeres Ito III strongly widened, bearing long tenent setae; protarsomere I wider than apex of protibia. Ventrite VI with apical lobe subtriangular, about 0.04 mm long. Aedeagus (Figs 1 ̅4) 0.60 mm long. Median lobe and parameres strongly sclerotized, symmetrical, dorsal valves excepted. Articular process not prominent, distinct. Apical process of median lobe somewhat shorter than basal bulb, moderately inflexed, with weakly concave ventral margin, gradually narrowed apically, tip acute in lateral view, obtuse in dorsal view. Parameres robust, gradually narrowed apically in dorsal view, widened apical in lateral view, with broadly rounded apex. Internal sac with complex sclerites, lacking spinose and denticulate structures, with apical section of ejaculatory duct wide, weakly membranous, extruded in repos. Etymology. The species epithet is a noun meaning "forest̏ in Malay. Differential diagnosis. This species is unique in having broadly interrupted pronotal bead. Therefore, it is placed in Baeocera tentatively (the mouth­parts and thorax of the unique available specimen were not dissected). In addition, its aedeagal characters are unusual, in particular by the structure of the internal sac and the asymmetrical valves of the median lobe. The internal sac lacking a flagellar guide sclerite and possessing a membranous ejaculatory duct extruded in repos is shared with the Chinese B. xichangana Löbl, 1999. It differs notably from B. xichangana by the not prominent articular process, the two asymmetrical dorsal valves, and the much shorter, robust parameres. Both species are also very distinct in external characters, in particular by their body­shape and size (B. xichangana is 2.0̅ 2.15 mm long). Remarks. As to the Sabah species, only B. barbara Löbl, 1990 and B. doriae (Pic, 1920) share the wide metanepisterna. These two species have asymmetrical aedeagus and may be easily distinguished by their external characters. The antitheses 7 and 8 have been inversed in key to the Sabah Baeocera given by LÖBL & SMETANA (2021): the thesis 6 (Metanepisternum fused,...) should lead to ̤ 8., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 8-9, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & SMETANA A. 2021: On the Baeocera Erichson (Coleoptera: Staphylinidae: Scaphidiinae) of Sabah, Malaysia, and a tale on mystified biodiversity. Journal of Insect Biodiversity 23 (2): 23 ‾ 42."]}

4. Key to the Baeocera species of the People’s Republic of China 1 Lateral parts of metaventrite impunctate, or very finely and sparsely punctate...................2 - Lateral parts of metaventrite coarsely, usually densely punctate, or at least with several conspicuous, coarse punctures...........................................................................................14 2 Elytra with basal striae interrupted, not joined to lateral striae. Exposed part of scutellum large....................................................................................................................3 - Elytral with basal striae entire, joined to lateral striae. Scutellum concealed or minute, except in B. breveapicalis...................................................................................................6 3 Apical part of median lobe large, not overlapped, about as long as basal bulb. Male metatibiae each with subapical denticle........................................ B. satana NAKANE, 1963 - Apical part of median lobe small, to part or entirely overlapped, much shorter than basal bulb. Male metatibiae lacking denticles.....................................................................4 4 Aedeagus with parameres simple, hook-like and curved at apices....... B. gilloghyi LÖBL, 1973 - Aedeagus with parameres lobed, not hook-like at apices....................................................5 5 Left parameres with small dorsal lobe and ventrally inflexed apex; right parameres lobed ventrally........................................................................... B. yunnanensis LÖBL, 1999 - Left parameres with large dorsal lobe and hardly inflexed apex; right parameres not lobed............................................................................................ B. pubiventris LÖBL, 1990 6 Antennomere VI short and wide, about 2 times as long as wide and as long as half of antennomere VII..................................................................................................................7 - Antennomere VI long and narrow, about 3 to 5 times as longs as wide, slightly shorter than antennomere VII..........................................................................................................8 7 Aedeagus with outer margin of left paramere sinuate and inner lobe subtriangular............................................................................................................... B. breveapicalis (PIC, 1926) - Aedeagus with outer margin of left paramere convexly rounded and inner lobe blunt................................................................................................................ B. takizawai LÖBL, 1984 8 Aedeagus with parameres strongly expanded apico-dorsally to form large, strongly sclerotized lobes................................................................................. B. kubani LÖBL, 1999 - Aedeagus with parameres not lobed, or each with small subapical lobe.............................9 9 Aedeagus with apices of parameres distinctly widened (dorsal view)....................................................................................................................................... B. huanshana LÖBL, 1999 - Aedeagus with apices of parameres not or hardly widened (dorsal view).........................10 10 Aedeagus lacking strongly sclerotized guide-sclerite, parameres slightly shorter than median lobe................................................................................ B. xichanagna LÖBL, 1999 - Aedeagus lacking with strongly sclerotized guide-sclerite, parameres much shorter than median lobe.......................................................................................................................11 11 Aedeagus with guide-sclerite narrow, angulate basally...................... B. callida LÖBL, 1986 - Aedeagus with guide-sclerite robust, not angulate basally................................................12 12 Aedeagus with guide-sclerite tapering apicad........................................ B. freyi LÖBL, 1966 - Aedeagus with guide-sclerite not tapering apicad.............................................................13 13 Aedeagus with guide-sclerite straight, becoming stouter apicad........ B. cooteri LÖBL, 1999 - Aedeagus with guide-sclerite abruptly curved in middle, widened subapically, acute at tip................................................................................................. B. hammondi LÖBL, 1984 14 Elytra with sutural striae shortened, starting well behind level of scutellar tip, far from elytral base........................................................................................................................15 - Elytra with sutural striae starting at elytral base, usually curved along pronotal lobe and forming basal striae...........................................................................................................16 15 Pronotal punctation coarse and dense, centre of metaventrite with mesal row of coarse punctures and narrow impunctate admesal areas................................ B. karenovicsi nov.sp. - Pronotal punctation very fine and sparse, hardly visible, metaventrite lacking mesal row of coarse punctures....................................................................... B. proclinata nov.sp. 16 Elytra with basal striae widely separated from lateral striae.............................................17 - Elytra with basal striae entire, joined to lateral striae........................................................19 17 Elytral punctation coarse anterior mid-length, very fine posterior mid-length or in apical thirds. Aedeagus with notched parameres...............................................................18 - Elytral punctation coarse on entire discal surfaces. Aedeagus with straight, not notched parameres.................................................................................................. B. repleta nov.sp. 18 Internal sac of aedeagus bearing tuft of spines.................................. B. franzi (LÖBL, 1973) Internal sac of aedeagus lacking tuft of spines.......................... B. proseminata LÖBL, 2013 19 Pronotal and elytral punctation similar, very fine..............................................................20 - Pronotal punctation very fine, elytral punctation in part or entirely coarse.......................21 20 Aedeagus with flagellum curved and thickened basally. Lateroposterior part of metaventrite coarsely punctate................................................... B. sordidoides LÖBL, 1992 - Aedeagus with flagellum almost straight in dorsal view, not thicken basally. Lateroposterior part of metaventrite very finely punctate.................. B. coalita LÖBL, 2013 21 Metanepisterna separated from metaventrite by distinct sutures.......................................................................................................................................... B. serendibense (LÖBL, 1971) - Metanepisterna fused with metaventrite............................................................................22 22 Punctation coarse on most of basal half to two thirds of elytral disc, apical third to half of elytral disc very finely punctate, appearing impunctate................................................23 - Elytra entirely or nearly entirely coarsely punctate...........................................................24 23 Parameres narrow, not folded apically............................................... B. erroris LÖBL, 1990 - Parameres wide, folded apically................................................ B. pseudinculta LÖBL, 1990 24 Ventrite 1 very finely punctate posterior basal puncture row, appearing impunctate............................................................................................................. B. longicornis (LÖBL, 1971) - Ventrite 1 coarsely punctate posterior basal puncture row................................................25 25 Apical halves of parameres distinctly bent and with weakly sclerotized inner margins................................................................................................................... B. pigra (LÖBL, 1971) - - Parameres straight, with inner and outer margins nearly evenly sclerotized...................26 26 Parameres almost as long as median lobe. Internal sac without narrow lateral rod............................................................................................................................ B. vidua LÖBL, 1990 - Parameres much shorter than median lobe. Internal sac with narrow lateral rod.............................................................................................................................. B. lasciva LÖBL, 2013, Published as part of Löbl, Ivan, 2018, Supplement to the knowledge of the genera Baeocera ERICHSON, 1845 and Scaphobaeocera CSIKI, 1909 (Coleoptera, Staphylinidae, Scaphidiinae) of the People's Republic of China, pp. 1295-1303 in Linzer biologische Beiträge 50 (2) on pages 1299-1300, DOI: 10.5281/zenodo.5275102, {"references":["LOBL I. (1999): A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of China, I. - Revue suisse zool. 106: 691 - 744.","LOBL I. (1992): The Scaphidiidae (Coleoptera) of the Nepal Himalaya. - Revue suisse zool. 99: 471 - 627.","LOBL I. (1971): Die Scaphidiidae (Coleoptera) Sudindiens. - Revue s uisse zool. 86: 77 - 129."]}

Pseudobironium subovatum Pic, 1920 Material examined. 1 male, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, primary forest, E. Slade & J. Villanuava lgt. Lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005­062 (OUMNH). Remarks. The species seems to be rather widely distributed. It is currently known from Indonesia (Sumatra and Kalimantan), Malaysia (West Malaysia, Sabah and Sarawak), and Thailand, though only a single Sabah record is published from the Gunung Emas in the Crocket Range (LÖBL & TANG 2013). The species may be easily distinguished by the long protibial setae in males., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 11, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Mordelloscaphium testaceimembre Pic, 1915 Material examined. 184, Sabah, Batu Punggul Resort env., 24.vi. ̅ 1.vii.1996, FIT (MHNG); 1, with the same data but in debris accumulates around large trees near river (MHNG); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 22̅ 23.x.2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, west trail area, FIT, OUMNH 2006­051 (OUMNH); 1, with the same data but 3̅13.xi., D.J. Mann & E. Slade lgt,, forest natural trail area OUMNH 2065­051 (OUMNH); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05°02.682′ E 117°45.553′, 250 m, iii̅ iv.2005, E. Slade & J. Villaneuve lgt., FIT, primary dipterocarp forest, OUMNH 2065­062 (OUMNH). Remarks. The currently known specimens of this species are the syntypes from east Kalimantan, Indonesia., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 10-11, DOI: 10.5281/zenodo.7501705

Pseudobironium Pic, 1920 The genus is Asian in distribution, with most members found in the tropics and subtropics, remaining unknown south of the Wallace line. Currently, six species are known to occur in Sabah (LÖBL & TANG 2013): P. convexum Löbl & Tang, 2013, P. horaki Löbl & Tang, 2013, P. pubiventer Löbl & Tang, 2013, P. sparsepunctatum (Pic, 1915), P. subovatum Pic, 1920, and P. vitalisi (Achard, 1920). Anew record is given, and two additional species are described below., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 11, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Scaphoxium sinuatum sp. nov. Figs 11 ̅13 Type material. Holotype male, Malaysia, Sabah, Batu Punggul Resort env., 24.VI. ̅ 1.VII.1996, 11c. vegetation debris and forest floor litter accumulated around large trees near river (MHNG). Description. Length 1.18 mm, width 0.55 mm, dorsoventral diameter 0.62 mm. Thorax, elytra and most of abdomen uniformly reddish­brown, head somewhat lighter, apical abdominal segments light, nearly yellowish, femora and tibiae roughly as thorax, tarsi lighter, antennae light brown. Length/width ratios of antennomeres as follows: III 22 /5: IV 17 /4: V 22 /4: VI 23 /4: VII 28 /7: VIII 20 /7: IX 28/11: X: 28/11: XI 38 /12. Pronotal and elytral punctation similar, very fine, hardly visible at 40 times magnification. Scutellum concealed. Elytra with sutural striae parallel, shortened, starting about 0.15 mm posterior of margin of pronotal lobe, adsutural areas hardly raised. Hypomeron with scattered, very fine punctured and short oblique stria. Mesoventrite with mesal impression shallow, narrowed posteriad, lacking mesal ridge and lacking microsculpture. Margin of mesocoxal process rounded. Metaventrite lacking microsculpture, very finely punctate, convex in middle. Lateral stria oblique, nearly reaching convex, finely punctate submesocoxal line. Submesocoxal area about 0.03 mm, as long as half of the shortest interval to margin of metacoxa. Metanepisternum flat, with suture fairly narrow, punctate, somewhat rounded. Abdomen very finely punctate, median areas of ventrites with punctulate microsculpture. Male. Segments Ito III of protarsi hardly widened. Aedeagus (Figs 11 ̅13) 0.44 mm long. Parameres narrowed posterior of basal section, gradually weakly widened toward narrow apical section, not forming subapical lobe or process; outer margins of parameres concave and inner margins strongly sclerotized posterior of narrowed apical section. Internal sac bearing two pairs of sclerotized pieces, outer larger and more sclerotized than inner pair, expanded apically, inner pair widened proximally, with margins becoming indistinct apically; scale­like, spine­like or filamentous structures absent. Etymology. The species epithet is a Latin adjective meaning "sinuatȅ, referring to the outer margins of the parameres. Differential diagnosis. The body­size of this new species is significantly smaller than that of other Bornean species and the shape of its parameres is diagnostic for it. The new species is similar to S. bilobum Löbl, 2015 from Lombok. It shares with the latter the simple internal sac of the aedeagus lacking spine­like, denticulate or striate membranous structures in combination with the presence of two pairs of admesal sclerites. However, the outer pair of sclerites is much more expanded and the inner sclerites are much shorter and broader in S. sinuatum, which may be distinguished from B. bilobum also by its wide impression of the mesoventrite, finer punctation of the lateral parts of the metaventrite and shorter submesocoxal areas., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 17-18, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2015: On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands. Revue suisse de zoologie 122 (1): 75 ‾ 120."]}

Baeocera sarawakensis Löbl, 1987 Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°58′ E 117°42′, 450 m, viii.1998, D. Bebber lgt., lowland mixed dipterocarp forest, FIT, OUMNH 2003­050 (OUMNH). Remarks. As B. incisa, this species is widely distributed in Sabah and seems to be common in forest floor litter (see LÖBL 2021: 31)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53 (1): 85 ‾ 89."]}

Sapitia Achard, 1920 The genus includes three South-East Asian species, all reported also from Indonesia though only one of them from the Lesser Sundas. Members of Sapitia are presumably associated with termites (Leschen & Löbl, 2005)., Published as part of Löbl, Ivan, 2015, On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands, pp. 75-120 in Revue suisse de Zoologie 122 (1) on page 92, DOI: 10.5281/zenodo.14582, {"references":["Achard J. 1920. Notes sur les Scaphidiidae de la faune indomalaise. Annales de la Societe Entomologique de Belgique 60: 123 - 136.","Leschen R. A. B., Lobl I. 2005. Phylogeny and classification of Scaphisomatini (Staphylinidae: Scaphidiinae) with notes on mycophagy, termitophily and functional morphology. Coleopterists Society Monographs, Patricia Vaurie series no. 3: 63 pp."]}

Pseudobironium irregulare sp. nov. (Figs 5 ̅7) Type material. Holotype male, SABAH: Poring Hot Springs 600 m 9.V.1987 Burckhardt ̅ Löbl (MHNG); Paratypes: 1 female, SABAH: Crocker Ra. 1350 m, km 60 Kota Kinabalu­Tambunan 17.V.1987 Burckhardt ̅ Löbl; female, BORNEO SABAH Tamis, Hwy AI 10 km NW Kinabalu Park Entr. 1100 m 24.V.87 A. Smetana; 1 female, SABAH: E Mt. Kinabalu 1150 m, rte Ranau­Kota Kinabalu 24.V.1987 Burckhardt ̅ Löbl (all in MHNG). Description. Length 1.78̅ 1.87 mm, width 1.20̅ 1.35 mm. Head and most of body dark reddish­brown, metaventrite somewhat darkened, apex of abdomen lighter, yellowish, femora and tibiae roughly as body, tarsi and antennae lighter, yellowish. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows: III 14: IV 25: V 25: VI 22: VII 26: VIII 26: IX 26: X: 26: XI 31; antennomere XI about 4 times as long as wide. Pronotal punctation about as fine as that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 5 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation on prevailing surface similar to pronotal punctation, conspicuous patch of rather coarse punctures present on anterolateral area. Puncture patch separated from lateral and basal margins by rather broad, very finely punctate areas. Lateral striae coarsely punctate. Hypomera impunctate, flattened along basal margin. Mesoventrite lacking mesal ridge, with distinct setiferous punctures at lateral margins of coxal process, setae long. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite with inconspicuous striate to punctulate microsculpture; median area nearly flat, lacking impressions and lacking tubercle, its punctation fine, arranged to form two sparse admesal rows joined posteriorly. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, inflexed. Submesocoxal area slightly shorter than half of shortest interval to metacoxa. Submesocoxal line coarsely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Exposed ventrites and tergites with distinct punctulate microsculpture, setiferous punctures very fine and scattered. Male. Protarsus shorter than protibia, protarsomeres Ito III hardly widened, narrower than apex of tibiae, bearing short setae. Aedeagus (Figs 5 ̅7) 0.68 mm long, weakly sclerotized. Median lobe with basal bulb shorter than apical process, the latter hardly inflexed, with sinuate ventral margin (lateral view) and blunt tip; apical part of median lobe overlapped by two valves. Parameres widened basally, weakly bent. Internal sac lacking sclerotized pieces, without distinct flagellum, bearing apical spinose structures. Etymology. The species epithet is a Latin adjective meaning "irregulaȑ, referring to the elytral punctation. Differential diagnosis. This species may be readily distinguished from its congeners, P. impressipenne Löbl, 1973 and P. subglabrum Löbl, 1990 excepted, by elytra bearing each an anterolateral patch of coarse punctures. In addition, its small body­size and the structure of the internal sac are diagnostic. It may be readily distinguished from P. impressipenne and P. subglabrum by the parameres widened basally. It falls to the couplet 25 in the key to the species in LÖBL & TANG (2013), to P. feai Pic, though the latter species is 2.90̅ 3.20 mm long and has a black body. The aedeagal characters suggest relationships with members of the P. plagiferum group. It may be distinguished from the three species placed in that group (P. javanum Löbl & Tang, 2013, P. plagiferum Löbl, 1980, and P. spinipes Löbl & Tang, 2013) by the shape of the parameres and apex of the median lobe., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 11-14, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Vituratella kistneri (Löbl, 1979) Materiel examined. 50, Sabah, Batu Punggul Resort, 24.vi̅1.vii.1996, FIT (MHNG); 1, Sabah, Mt. Kinabalu Nat. Park, Poring Hot Springs, 495 m, 30.viii.1988, A. Smetana [B161] (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, selectively logged lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession, E. Slade & J. Villanuava lgt., OUMNH­2005­062 (OUMNH); 1, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 27̅ 28.x.2005, primary forest, D.J. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, west trail 10, forest interior FIT, OUMNH­2006­051 (OUMNH). Remarks. The species was based on several specimens found in North Sumatra, in fungus gardens of Odontotermes takensis Ahmad, 1965. It was not reported after its description while V. termitophila Champion, 1927 is widely distributed and has been referred to many times (see OGAWA & MAETO 2015). The colour pattern of the abdomen used as one of the diagnostic characters proves to be unreliable in the examined material., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 18, DOI: 10.5281/zenodo.7501705, {"references":["OGAWA R. & MAETO K. 2015: The termitophilous Scaphidiinae (Coleoptera: Staphylinidae) from Sulawesi, Indonesia. The Coleopterists Bulletin 69 (2): 301 ‾ 304."]}

Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1-2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705, {"references":["LESCHEN R.A.B. & LOBL I. 2005: Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1‾63.","LOBL I. 2015: On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands. Revue suisse de zoologie 122(1): 75‾120.","LOBL I. 2018: Coleoptera: Staphylinidae: Scaphidiinae. World Catalogue of Insects. Volume XVI: i‾xvi + 1‾418 pp.","LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53(1): 85‾89.","LOBL I. & LESCHEN R.A.B. 2010: Notes on the Toxidium group (Coleoptera: Staphylinidae: Scaphidiinae). Folia Heyrovskyana Series A 18(1-3): 71‾93.","LOBL I., LESCHEN R.A.B. & KODADA J. 2020: Review of the Asian species and cladistic analysis of Bironium Csiki (Coleoptera: Staphylinidae: Scaphidiinae) with comments on biogeography. Annales Zoologici (Warszawa) 70(4): 711‾736.","LOBL I. & SMETANA A. 2021: On the Baeocera Erichson (Coleoptera: Staphylinidae: Scaphidiinae) of Sabah, Malaysia, and a tale on mystified biodiversity. Journal of Insect Biodiversity 23(2): 23‾42.","LOBL I. & TANG L. 2013:Areview of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665‾734.","OGAWA R. & LOBL I. 2016: Areview of the genus Xotidium Lobl, 1992 (Coleoptera, Staphylinidae, Scaphidiinae), with description of five new species. Deutsche entomologische Zeitschrift 63: 155‾169.","OGAWA R. & MAETO K. 2015: The termitophilous Scaphidiinae (Coleoptera: Staphylinidae) from Sulawesi, Indonesia. The Coleopterists Bulletin 69(2): 301‾304.","PROKOFIEV A.M. 2013: Contribution to the knowledge of the scaphidiine genus Scaphisoma Leach of the Bu Gia Map National Park, Vietnam. Calodema 245: 1‾6."]}

Löbl, Ivan (2022): On 'Scaphisoma tricolor Heller' from Japan and Errata to the Descriptions of Scaphisoma konvickai Löbl and Toxidium hartmanni Löbl (Coleoptera: Staphylinidae: Scaphidiinae). Japanese Journal of Systematic Entomology 28 (1): 85-88, DOI: http://doi.org/10.5281/zenodo.6787471, {"references":["Heller, K.M., 1917. Scaphidiidae von den Philippinen. Wiener entomologische Zeitung, 36: 41-50.","Hoshina, H., 2013. Scaphidiinae, pp. 124-127. In: Catalogue of Japanese Staphylinidae. Bulletin of the Kyushu University Museum, 11: 69-218.","Lobl, I., 1970. Uber einige Scaphidiidae (Coleoptera) aus der Sammlung des Museum National d'Histoire Naturelle de Paris. Mitteilungen der schweizerischen entomologischen Gesellschaft, 43: 125-132.","Lobl, I., 1971. Scaphidiidae der Noona Dan Expedition nach den Philippinen und Bismark Inseln (Insecta, Coleoptera). Steenstrupia, 1: 247-253.","Lobl, I., 1972. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) von den Philippinen. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 45: 79-109.","Lobl, I., 1981. Insects of Micronesia Coleoptera Scaphidiidae. Insects of Micronesia, 15: 69-80.","Lobl, I., 1982. Uber die Scaphidiidae (Coleoptera) der japanischen Ryukyu-Inseln. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 55: 101-105.","Lobl, I., 2018. Coleoptera: Staphylinidae: Scaphidiinae. World Catalogue of Insects. Volume 16. Brill, Leiden/Boston, xvi + 418 pp.","Lobl, I. & R. Ogawa, 2016. On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia Achard and Scaphisoma Leach. Linzer biologische Beitrage, 48(2): 1339-1492.","Lobl, I., 2022. On new collections of Scaphidiinae (Coleoptera: Staphylinidae) from China, with description of two new species. Zootaxa, 5092 (4): 487-492.","Morimoto, K., 1985. Scaphidiidae, pp. 252-258. pl. 45. In: Ueno, S., Kurosawa, Y. & Sato, M. (eds.): The Coleoptera of Japan in color. Vol. 2. Hoikusha Publ. Co., Osaka, viii + 479 pp., 80 pls [in Japanese].","Ogawa, R., 2015. Phylogeography of the subfamily Scaphidiinae (Coleoptera, Staphylinidae) in Sulawesi, with its systematic revision. Graduate School of Agricultural Science, Kobe University, 180 pp. [PhD thesis, unpublished].","Schulke, M. & A. Smetana, 2015. Staphylinidae, pp. 304-900. In: Lobl, I. & D. Lobl, (eds), Catalogue of Palaearctic Coleoptera. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Volume 1. Brill, Leiden/Boston, xxv + 900 pp. [Received: February 21, 2022; accepted: June 12, 2022]"]}

Löbl, Ivan (2022): Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas. Revue suisse de Zoologie 129 (1): 103-118, DOI: 10.35929/RSZ.0064

Baeocera sp. B Material examined: MHNG; 1 female; MALUC: Kai Besar Island, G. Dab, leaf litter, 400 m, 5.IX.1981, D. Agosti, F911046. Remarks: This is the second Moluccan species of Baeocera remaining unnamed, in absence of at least one male specimen (see “ Baeocera sp. A ” in Löbl, 2015a). It is similar to the Papuan B. curta Löbl, 2002, differs by the less arcuate suture of the metanepisterna and the ventrite I lacking basal striae. Thus, it may be an undescribed species. Baeocera curta has a complex aedeagus providing reliable specific characters which are quite unlike those of members of the Baeocera lenta group and of B. agostii. The specimen has slightly prominent apical inner apical angles of the elytra, a character noted for the first time in Baeocera. It may be sexual, as it is in some species of Scaphisoma Leach., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 112, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2015 a. Contribution to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas. Stuttgarter Beitrage zur Naturkunde A, Neue Serie 8: 165 - 187.","Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Baeocera punctigera sp. nov. Figs 14-16 Materiel examined: Holotype; male; MHNG; MALUC: Tanimbar Isl., Kai Besar, G. Dab, 400 m, 5.IX.81, D. Agosti F911046. Etymology: The species epithet is a Latin adjective meaning bearing punctures. Diagnosis: The species is defined by the following characters in combination: Body-length about 1.2 mm long, body dark brown; pronotum coarsely and densely punctate, with many punctures larger than puncture intervals; lateral margins of pronotum and elytra separately arcuate; entire elytron coarsely punctate, sutural stria shortened, basal stria absent; mesepimeron with puncture row along anterior margin; hypomeron, mesanepisternum, metaventrite and ventrite I conspicuously punctate; ventrite I without basal striae or wrinkles; aedeagus symmetrical, with apical process narrowed apically, bent near tip (lateral view), parameres straight and evenly narrow in dorsal view, sinuate and widened toward apical third in lateral view, internal sac with complex sclerites as in Fig. 16, lacking membranous scale or denticle-like structures. Description: Length 1.17 mm, width 0.80 mm. Head and body dark brown, femora and tibiae lighter, reddish-brown, tarsi and antennae yellowish. Head with interocular distance slightly larger than dorsoventral eye diameter. Antennomere II slightly longer than antennomere III. Length/width ratios of antennomeres as: III 28/7: IV 27/6: V 32/6: VI 28/6: VII 33/10: VIII 28/8: IX 35/13: X 35/14: XI 42/15. Pronotum and elytra separately arcuate. Pronotum not microsculptured, pronotal punctation conspicuously coarse and dense, distinct at 15 times magnification, consisting of sharply delimited punctures smaller than puncture intervals; lateral carinae of pronotum hardly visible in dorsal view, with punctures very dense, larger than puncture intervals. Point of scutellum exposed. Elytra covering abdomen, weakly narrowed apically, with lateral margins evenly, weakly rounded; adsutural area flat and narrow; sutural stria punctate, shortened, starting about 0.20 mm posterior of tip of scutellum; basal stria absent; lateral stria punctate. Elytral disc not microsculptured; punctation entirely coarse and dense, punctures larger than those on pronotum and nearly as well delimited, puncture intervals mostly larger than puncture diameters. Hind wings fully developed. Hypomeron not microsculptured, conspicuously punctate except on smooth areas along upper margin and near mesocoxa, with some punctures about as large as puncture intervals and as pronotal punctures. Mesoventrite punctate, without median ridge, fused with metaventral process. Mesanepisternum conspicuously punctate, punctures similar to those on hypomeron. Mesepimeron with row of coarse punctures along anterior margin, about three times as long as wide and three times as long as interval to coxa. Metaventrite flattened in middle, all over coarsely and densely punctate, narrow apical band excepted; punctures on lateral part of metaventrite well delimited, mostly larger than puncture intervals. Submesocoxal lines parallel, with marginal punctures smaller than punctures on lateral area of metaventrite and somewhat extended laterally along margin of metaventrite. Submesocoxal areas about 0.02 mm long, as fifth of shortest distance to metacoxae. Metanepisternum flat, fused with metaventrite, its margin indicated by outer puncture row of metaventrite. Tibiae straight. Abdomen not microsculptured. Ventrite I densely and coarsely punctate, punctures smaller than those on lateral area of metaventrite, basal punctures not enlarged and not elongate, basal striae or wrinkles absent. Following ventrites finely punctate. Male characters: Protarsomeres I to III slightly widened. Aedeagus (Figs 14-16) 0.29 mm long, weakly sclerotized, symmetrical.Apical process gradually narrowed apically, with tip bent and blunt in lateral view. Parameres straight and evenly narrow in dorsal view, sinuate and gradually widened toward apical third in lateral view. Internal sac with complex sclerites, as in Fig. 16 with flagellar guidesclerite widened apically, broad, nearly membranous accessory lateral sclerite, lacking membranous scale- or denticle-like structures. Habitat: South slope of G. Dab, secondary forest on limestone, soil and very humid leaf litter. Distribution: Indonesia: Moluccas, Tanimbar Islands. Remarks: This is another species of the B. lenta group, in external characters very similar to B. notata. It may be easily distinguished from the latter by the mesepimeron bearing a row of coarse punctures along its anterior margin and by the aedeagal characters, in particular by the parameres straight and narrow in dorsal view. In addition, the body-size of B. punctigera is smaller, the antennae are shorter, with the antennomere XI less than 3 times as long as wide. While the aedeagus of B. punctigera is similar to that of the New Guinean B. flagrans Löbl, 2002, the species differs strongly from the latter by its external characters., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on pages 108-110, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Baeocera notata sp. nov. Figs 7-10 Materiel examined: Holotype; MHNG; male; MALUC: Aru Isls: Kola Isl.: Watan, 27.8.81, D. Agosti F91936. Paratype; MHNG; 1 female; MALUC: Aru Isl., Wokam, 22.8.81, D. Agosti F911970. Etymology: The species epithet is a Latin adjective referring to the pronotum. Diagnosis: The species is defined by the following characters in combination: Body-length about 1.35- 1.45 mm, body dark brown; pronotum coarsely and densely punctate, with many punctures larger than puncture intervals; lateral margins of pronotum and elytra separately arcuate; elytron entirely coarsely punctate, with sutural stria shortened, lacking basal stria; hypomeron and mesanepisternum distinctly punctate; metaventrite and ventrite I coarsely punctate; ventrite I without basal striae or wrinkles; aedeagus symmetrical, with apical process narrowed apically, bent near tip in lateral view, apical halves of parameres bent and bearing subapical membranous lobe (dorsal view), widened posterior of basal third in lateral view, internal sac with complex sclerites as in Fig. 9, lacking membranous scale or denticle-like structures. Description: Length 1.36-1.43 mm, width 0.90- 1.0 mm. Head and body dark brown, femora and tibiae lighter, reddish-brown, tarsi and antennae yellowish. Head with interocular distance slightly smaller than dorsoventral eye diameter. Antennomere II slightly longer than antennomere III. Length/width ratios of antennomeres as: III 28/7: IV 27/7: V 37/7: VI 35/7: VII 39/12: VIII 38/10: IX 39/12: X 37/14: XI 48/14. Pronotum and elytra separately arcuate. Pronotum not microsculptured, pronotal punctation conspicuously coarse and dense, distinct at 15 times magnification, consisting of sharply delimited punctures to part larger than puncture intervals; lateral carinae of pronotum hardly visible in dorsal view, bearing punctures very dense, larger than puncture intervals. Point of scutellum exposed. Elytra covering tip of abdomen, weakly narrowed apically, with lateral margins evenly, weakly rounded; adsutural area flat and narrow; sutural stria punctate, shortened, starting about 0.20 mm posterior of tip of scutellum; basal stria absent; lateral stria punctate. Elytral disc not microsculptured; punctation all over coarse and dense, punctures larger than those on pronotum but not as well delimited, puncture intervals mostly about as puncture diameters or slightly larger. Hind wings fully developed. Hypomeron not microsculptured, conspicuously punctate, with punctures about as large as pronotal punctures. Mesoventrite punctate, without median ridge, fused with metaventral process. Mesanepisternum conspicuously punctate, punctures similar to those on hypomeron. Mesepimeron impunctate, about four times as long as wide and three times as long as interval to coxa. Metaventrite flattened in middle, all over coarsely and densely punctate, smooth narrow apical band excepted; punctures on lateral part of metaventrite well delimited, to part as large or larger than puncture intervals. Submesocoxal lines slightly convex, with marginal punctures smaller than punctures on lateral area of metaventrite and somewhat extended laterally along margin of metaventrite. Submesocoxal areas about 0.03 mm long, as third of shortest interval to metacoxae. Metanepisternum flat, fused with metaventrite, its margin indicated by outer puncture row of metaventrite. Tibiae straight. Abdomen not microsculptured. Ventrite I densely and coarsely punctate, punctures smaller than those on lateral area of metaventrite, basal punctures not enlarged and not elongate, basal striae or wrinkles absent. Following ventrites finely punctate. Male characters: Protarsomeres I to III slightly widened. Aedeagus (Figs 7-10) 0.37 mm long, weakly sclerotized, symmetrical.Apical process gradually narrowed apically, with tip bent and acute in lateral view. Parameres straight in basal halves, bent in apical halves, each with weakly sclerotized subapical lobe (dorsal view), narrow near bases and nearly evenly broad in apical two thirds, with slightly concave ventral margins (lateral view). Internal sac with complex sclerites, as in Fig. 9, with flagellar guide-sclerite widened apically, narrow accessory lateral sclerite present, lacking membranous scale- or denticlelike structures. Habitat: Secondary/primary forest 5 km E of Mt. Sungai, shallow leaf litter, on limestone; understory with Pandanus. Distribution: Indonesia: Moluccas, Aru Islands. Remarks: The species is another member of the B. lenta group. It shares with the Moluccan B. kaibesara Löbl, 2014 and B. vicina Löbl, 2015, and the Papuan B. praedicta Löbl, 2002, elytra with shortened sutural striae. It may be readily distinguished from these three species by the hypomera and mesanepisterna conspicuously punctate. In addition, the shape of the parameres lobed subapically is diagnostic for this new species. Several species, such as the New Guinean B. fortepunctata Löbl, 2002 and the Philippine B. hypomeralis Löbl, 2012 also possess punctate hypomera but differs by the elytra having entire sutural striae and the impunctate or extremely finely punctate mesanepisterna. The Himalayan B. cribrata Löbl, 1992, shares the pronotum coarsely punctate, distinctly punctate hypomera and mesanepisterna, and elytra with shortened sutural striae. It may be easily distinguished by the punctation on lateral parts of the pronotum much coarser than that on pronotal centre., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on pages 106-107, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2014. On the Scaphidiinae Leach (Coleoptera, Staphylinidae) of the Moluccas. Mitteilungen der Schweizerischen entomologischen Gesellschaft 87: 49 - 60.","Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Scaphisoma sublimbatum Löbl, 1977 Scaphisoma sublimbatum Löbl, 1977: 57. Material examined: MHNG; 2 males; MALUK: Aru Isl., Wokam, 5 km E. Mt. Sungai, 22.VIII.1981, D. Agosti, F91936. Redescription: Length 1.53-1.75 mm, width 1.02- 1.22 mm. Head, thorax and most abdomen dark brown, apex of abdomen yellowish. Elytra on adsutural area, along basal and lateral margins, and on transverse stripe in middle as dark as pronotum, in apical sixth to fifth light brown, each with two large, irregular, transverse yellowish spots. Anterior spots extending to middle third of elytral length, posterior spot starting about at level of apical third of elytral length. Femora light brown, tibiae, tarsi and antennae yellowish. Length/ width ratios of antennomeres as: III 12/10: IV 27/8: V 50/9: VI 50/9: VII 57/16: VIII 38/10: IX 53/14: X 53/15: XI 60/14. Pronotum not microsculptured, with lateral margins strongly, evenly rounded, lateral margin carinae throughout visible in dorsal view, lateral margin carinae impunctate, discal punctation fine, sparse, most puncture intervals about 2 to 4 times as large as puncture diameters, punctures shallow, not well delimited, visible at 30 times magnification. Exposed point of scutellum minute. Elytra not microsculptured, with lateral margins evenly rounded, lateral margin carinae exposed in dorsal view, lateral margin striae impunctate, apical margins truncate, inner apical angles not prominent, situated in level with outer apical angles, marginal crenulation very fine, distinct, sutural striae deep, bent at bases and extending laterally to form basal striae reaching outer half of basal width, widely separated from lateral striae, sutural striae parallel, impunctate, adsutural areas flat, each with single, very fine and sparse puncture row, discal punctures irregular, fairly fine and dense, coarser than pronotal or adsutural punctures, not forming rows, puncture intervals mostly distinctly larger than puncture diameters. Hind wings fully developed. Hypomera not microsculptured, impunctate. Mesepimon about 4 times as long as wide and somewhat shorter than interval to mesocoxa. Metaventrite not microsculptured. Median part of metaventrite convex between mesocoxae, flattened apically, with three fovea-like apical impressions, middle impression larger and situated slightly anterior lateral impressions, often joint to short mesal stria; punctation indistinct, sparse and extremely fine on centre and on lateral areas of metaventrite. Lateral parts of metaventrite lacking antecoxal puncture rows. Submesocoxal area 0.02 mm long, about as sixth of shortest interval to metacoxa. Submesocoxal line parallel, densely and coarsely punctate, punctures extended laterally along anterior margin of metaventrite. Metanepisterna slightly convex, impressed along nearly straight suture, narrowed anteriad. Tibiae straight. Abdomen not microsculptured, with hardly visible punctation (at 100 times magnification). Ventrite I with submetacoxal area extremely narrow, about 0.01- 0.02 mm long, narrowed mesally and as fifteens of interval to apical margin of ventrite; marginal punctures coarse. Male characters: Protarsomeres I to III and mesotarsomeres I and II strongly widened, basal tarsomeres nearly as wide as apices of tibiae. Apical margin of ventrite VI trilobed, mesal lobe triangular, about 0.10 mm long, lateral lobes shorter and rounded. Habitat: Secondary/primary forest on limestone, leaf litter on rather dry soil, understory Pandanus. Distribution: Australia: Queensland; Indonesia: Moluccas, Aru Islands. Remarks: This species is a member of the S. unifasciatum group defined in Löbl, 1977. In addition to S. sublimbatum, the group includes four New Guinean species: S. audax Löbl, 1975, S. quadripunctatum (Pic, 1956), S. unifasciatum Pic, 1956, and S. unimaculatum Löbl, 1975. An additional species is described below. The original description of the external features S. sublimbatum was brief and limited to comparation to the similar S. audax. Therefore, it is supplement., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 112, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 1977. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) Australiens. Revue suisse de Zoologie 84: 3 - 69.","Lobl I. 1975. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) von Neuguinea. Revue suisse de Zoologie 82: 369 - 420."]}

Scaphoxium sp. Materiel examined: MHNG; 1 female; MOLUK: Kai Besar, Bombay, G. Dab, 3.IX.1981, D. Agosti, F911019. MHNG; 1 female; MOLUK: Kai Besar, G. Tukram, 300-500 m, 7.IX.1981, D. Agosti F911075. Remarks: These two specimens differ notably from S. lobatum by the very finely punctate lateral parts of the metaventrite. Habitat: Secondary forests on limestone; leaf litter and moss on fallen tree. Distribution: Indonesia: Moluccas, Kai Besar Island., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 117, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022

Baeocera incurva sp. nov. Figs 4-6 Materiel examined: Holotype; MHNG; male; MALUK: Kai Besar, Bombay, G. Dab, 3.IX.81, D. Agosti F911019. Paratypes; MHNG; 1 male, 2 ex. with the same data as the holotype. – MHNG; 2 males, 1 female; MALUK: Kai Besar, G. Tukrau nr. Bombay, 300 -400 m, 7.IX.81 D. Agosti, F91075. Etymology: The species epithet is a Latin adjective meaning bent. Diagnosis: The species is defined by the following characters in combination: Body-length about 1.25- 1.35 mm, body blackish with lighter apical abdominal segments; lateral margins of pronotum and elytra nearly continuously arcuate; pronotum finely punctate; elytron entirely coarsely punctate, lacking basal stria, sutural stria shortened, starting posterior of level of scutellum; hypomeron very finely punctate; mesepimeron, mesanepisterum, metaventrite and ventrite I distinctly punctate; ventrite I without basal wrinkles or striae; aedeagus symmetrical, with apical process in lateral view abruptly narrowed near apex, parameres sinuate, nearly evenly broad and bent dorsally in lateral view, internal sac with complex sclerites, as in Fig. 6, lacking membranous scale or denticle-like structures. Description: Length 1.25-1.33 mm, width 0.85- 0.88 mm. Head and most of body blackish, apical abdominal segments, femora and tibiae reddish-brown, tarsi and antennae yellowish. Head with interocular distance slightly larger than dorsoventral eye diameter. Antennomere II slightly longer than antennomere III. Length/width ratios of antennomeres as: III 24/6: IV 22/6: V 30/7: VI 30/7: VII 37/9: VIII 33/8: IX 41/12: X 41/14: XI 46/15. Pronotum and elytra nearly continuously arcuate. Pronotum not microsculptured, pronotal punctation hardly visible at 50 times magnification; lateral carinae of pronotum concealed in dorsal view. Point of scutellum exposed. Elytra nearly entirely covering abdomen. Elytron weakly narrowed apically, with lateral margin rounded; lateral margin carinae concealed in dorsal view, adsutural area flat and narrow; sutural stria punctate, shortened, starting well posterior of level of tip of scutellum; basal stria absent; lateral stria punctate. Elytral disc not microsculptured, with punctation well delimited, much coarser than on pronotum, nearly regular, puncture intervals mostly about as to three times puncture diameters. Hind wings fully developed. Impressed inferior part of hypomeron with few very fine punctures. Mesoventrite punctate, without median ridge, fused with metaventral process. Mesanepisternum pubescent, rather finely punctate. Mesepimeron punctate, about four times as long as wide and about three times as long as interval to coxa. Median part of metaventrite flattened, fairly coarsely, irregularly and densely punctate. Lateral area of metaventrite with punctation similar to that on elytron; punctures well delimited, clearly smaller than puncture intervals, evanescent near metacoxal margin. Submesocoxal lines slightly convex, with marginal punctures smaller than punctures on lateral area of metaventrite and not extended laterally along margin of metaventrite. Submesocoxal areas about 0.02-0.03 mm long, about as fifth of shortest interval to metacoxae. Metanepisternum flat, fused with metaventrite, its margin indicated by outer puncture row of metaventrite. Tibiae straight. Abdomen not microsculptured. Ventrite I about as coarsely and densely punctate as lateral area of metaventrite, basal punctures not enlarged and not elongate, basal wrinkles or striae absent. Following ventrites finely punctate. Male characters: Protarsomeres I to III slightly widened. Aedeagus (Figs 4-6) 0.35-0.36 mm long, weakly sclerotized, symmetrical. Apical process abruptly narrowed near apex (lateral view), gradually narrowed in dorsal view. Parameres conspicuously sinuate, nearly evenly broad and bent dorsally in lateral view, hardly sinuate in dorsal view. Internal sac with complex sclerites as in Fig. 6, with very short flagellar guide-sclerite, lacking membranous scale- or denticle-like structures. Habitat: Secondary forests on limestone, litter from dead log and rotting leaves. Distribution: Indonesia: Moluccas, Kai Besar Island. Remarks: The species is also member of the B. lenta group, as shown by its aedeagal characters. It falls in the key to the Moluccan species (Löbl, 2015a) to B. vicina Löbl, 2015. It may be easily distinguished from the latter as from other congeners by the shape of the parameres and by the internal sac possessing a short flagellum and reduced flagellar-guide sclerite. The New Guinean B. praedicta Löbl, 2002 possesses parameres rather similar to those of B. incurva. It differs notably in external characters, in particular by the not fused metanepisterna and the very finely punctate metaventrite., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on pages 104-106, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2015 a. Contribution to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas. Stuttgarter Beitrage zur Naturkunde A, Neue Serie 8: 165 - 187.","Lobl I. 2002. On Baeocera (Coleoptera: Staphylinidae: Scaphidiinae) of New Guinea. Mitteilungen der Schweizerischen entomologischen Gesellschaft 75: 1 - 20."]}

Scaphobaeocera sp. 2 Material examined: MHNG; 1 female; MALUK: Tanimbar Isl., Yamdena, 22 km N Samlaki via Aram, Desa Lorulun, 11.IX.1981, D. Agosti. Remarks: This specimen would fall in the key to the New Guinean Scaphobaeocera (Löbl, 2017) under the couplet 5, to S. antennalis Löbl, 1975 and S. remota Löbl, 1981, and it shares with S. remota the elytra darkened apically and the antennomeres XI more than twice as long as the antennomeres X. Habitat: Melaleuca open forest on limestone, leaf litter heavily infested by mycorrhiza. Distribution: Indonesia: Moluccas, Tanimbar Island., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 115, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2017. New species and records of Scaphobaeocera Csiki from New Guinea (Coleoptera: Staphylinidae: Scaphidiinae). Mitteilungen der Munchner entomologischen Gesellschaft 107: 33 - 41.","Lobl I. 1975. Beitrag zur Kenntnis der Scaphidiidae (Coleoptera) von Neuguinea. Revue suisse de Zoologie 82: 369 - 420.","Lobl I. 1981. Scaphidiidae. Insects of Micronesia 15 (2): 69 - 80."]}

Scaphisoma donati sp. nov. Figs 20-23 Material examined: Holotype; MHNG; male; MALUC: Tanimbar Isl., Sangliat Krawai E Yamdena, 19.IX.81 D. Agosti F911120. Etymology: The species is named in honour of its collector, Donat Agosti (Bern, Switzerland). Diagnosis: The species is defined by the following characters in combination: Body-length nearly 2 mm, body dark reddish-brown, not microsculptured; elytra with sutural striae parallel, basal striae entire, joined to lateral striae; mesepimeron shorter than interval between its tip and mesocoxa; metaventrite distinctly punctate on transverse area anterior of metacoxal process, lacking fovea-like impressions; submesocoxal and submetacoxal areas very narrow, with parallel lines; aedeagus as Figs 20-23, strongly asymmetrical, parameres lobed, overlapping apex of median lobe, internal sac strongly sclerotized. Description: Length 1.94 mm, width 1.33 mm. Head and most of body nearly evenly dark reddish-brown. Apex of abdomen and appendages lighter. Antennomere II about four times as long as antennomere III. Length/ width ratios of antennomeres as: III 9/10: IV 38/8: V 56/9: VI 52/8: VII 56/13: VIII 44/8: IX 57/15: X 52/14: XI 58/15. Pronotum and elytra not microsculptured. Pronotum with lateral margins rounded, lateral margin carinae not visible in dorsal view, discal punctation dense and very fine, consisting of poorly delimited punctures. Minute point of scutellum exposed. Elytra moderately narrowed apically, lateral margin rounded, lateral margin carina clearly exposed in dorsal view only in basal third, inner apical angle about in level with outer apical angle, sutural margin not raised, adsutural area flat, not narrowed apically, with very fine puncture row; sutural striae curved at base and extended laterally to form basal striae joined to lateral striae, discal punctation irregular, fine, punctures not well delimited, puncture intervals mostly about twice to five times as large as puncture diameters. Hind winds reduced. Exposed tergites very finely punctate, not microsculptured. Hypomeron smooth. Mesepimeron about four times as long as wide, distinctly shorter than interval between its tip and mesocoxa. Metaventrite short, not microsculptured, slightly convex in middle. Mesocoxal process concealed with mesoventrite. Centre of metaventrite impunctate, fairly coarse punctures present on transverse area anterior metacoxal process, fovea-like impressions absent. Lateral parts of metaventrite very finely and sparsely punctate, punctures on lateral areas hardly visible, antecoxal puncture rows absent. Submesocoxal lines parallel, fairly coarsely punctured; submesocoxal area about 0.02 mm long, as tenth of shortest distance to metacoxa. Metanepiternum narrowed anteriad. Ventrite I with punctation very fine, similar to lateral parts of metaventrite; submetaxocal lines parallel, margined with coarse punctures; submetacoxal areas hardly 0.02 mm, about as tenth of shortest distance to apical margin. Male characters: Ventrite VI with mesal process triangular, about 0.15 mm long. Protarsomeres and mesotarsomeres I to III distinctly widened, with tenant setae, protarsomeres I narrower than apices of protibiae. Aedeagus (Figs 20-23) 1.25 mm long, strongly sclerotized and strongly asymmetrical. Apical process of median lobe slightly inflexed, obliquely truncate at apex. Left paramere extended posterior of right paramere and overlapping most of apical process. Internal sac with robust, strongly sclerotized and irregularly narrowed rod. Habitat: Kanary evergreen primary forest on limestone, 50 m, thick leaf litter with heavy mycorrhiza. Distribution: Indonesia: Moluccas, Yamdena Island. Remarks: This is a member of the S. sublimbatum group. It may be easily distinguished from the other five members of the group by the elytra lacking spots, the metaventrite lacking mesal stria and fovea-like impressions, the antennomeres XI about as large as the antennomeres IX, and the basal striae of the elytra entire, joined to the lateral striae, in combination. The aedeagal characters suggest close relationships to S. quadripunctatum (Pic, 1956), the shape of the truncate tip of the apical process of the median lobe is unique in the new species. Its reduced hind wings suggest narrow endemism., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 114, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022

Scaphobaeocera sp. 1 Material examined: MHNG; 1 male; MALUK: Aru Isl., Wokam, 5 km E Mt. Sungai, 22.8.1981, D. Agosti. Remarks: This specimen possesses aedeagal characters similar to those of S. notata Löbl, 2017, from Papua New Guinea, and S. lombokensis Löbl, 2015. It differs from S. notata quite conspicuously by the uniformly light brown elytra, and from S. lombokensis by the parameres nearly evenly broad in lateral view and the ventrally bent tip of the median lobe. More material is needed to assess the variability of these characters. Habitat: Secondary/primary forest on limestone, sea level, rather dry leaf litter; understory Pandanus palms. Distribution: Indonesia: Moluccas, Aru Island., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 115, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2017. New species and records of Scaphobaeocera Csiki from New Guinea (Coleoptera: Staphylinidae: Scaphidiinae). Mitteilungen der Munchner entomologischen Gesellschaft 107: 33 - 41."]}

Scaphoxium lobatum sp. nov. Figs 28, 29 Material examined: Holotype; MHNG; male; MALUK: Aru Isl., Wokam, 5 km E Mt. Sungai, 22.8.1981, D. Agosti F91936. Etymology: The species epithet is a Latin adjective meaning lobed. Diagnosis: Body-length 1.35 mm; elytral punctation well delimited; hypomeron with longitudinal ridge; mesoventrite and metaventrite grooved mesally; lateral parts of the metaventrite bearing coarse punctures; submesocoxal area as long as shortest interval to metacoxa; abdomen with punctulate microsculpture; aedeagus as Figs 28, 29, with parameres broadly lobed at level of tip on median love, slightly narrowed posterior of lobe, internal sac lacking distinct sclerites and without scale-like or spine-like structures. Description: Length 1.35 mm, width 0.67 mm, dorsoventral diameter 0.73 mm. Body uniformly dark brown, appendages lighter. Length/width ratios of antennomeres as: III 15/6: IV 15/6: V 25/6: VI 23/7: VII 38/10: VIII 28/8: IX 35/11: X 35/12: XI 43/13. Pronotal punctation dense and fine, distinct at 50 times magnification. Tip of scutellum exposed. Sutural striae starting posterior of basal fourth of elytra, evanescent in anterior 0.20 mm of elytral length. Elytral punctation and pubescence dense, visible at 25 times magnification, consist of sharply delimited punctures. Hypomeron very finely punctate on somewhat convex upper part, impunctate on flattened lower part; lower and upper parts separated by ridge extended anteriorly to level of procoxal mid-length. Mesoventrite grooved mesally. Mesanepisternum very finely punctate. Metaventrite grooved mesally, in middle very finely punctate. Lateral parts of metaventrite with punctation coarse and dense near submesocoxal lines, with punctures to parts larger than puncture intervals; punctation becoming sparser and fines laterad, areas along metanepisterna impunctate. Submesocoxal lines strongly arcuate, finely punctate. Submesocoxal areas 0.05 mm, about as long as shortest intervals to metacoxae. Metanepisterna not fused, with straight, finely punctate suture nearly touching margin of metepimera. Visible abdominal segments with distinct punctulate microsculpture. Exposed ventrites with very fine punctation, coarse row of basal punctures of ventrite I excepted. Male characters: Protarsomeres I to III hardly widened. Aedeagus (Figs 28, 29) 0.51 mm long. Parameres each with large, rounded lobe at level of tip of median lobe, narrow and weakly bent posterior lobe, internal sac with plate-like structure, lacking spines or scales. Habitat: Secondary/primary forest on limestone, rather dry leaf litter; understory Pandanus palms. Distribution: Indonesia: Moluccas, Aru Islands. Remarks: This species resembles S. bilobum Löbl, 2015 from Lombok which shares the shape of the parameres and the coarse punctation on lateral parts of the metaventrite. These two species differ in elytral punctation which is notably denser and consist of sharply delimited punctures in S. lobatum. The more relevant feature of S. lobatum is the internal sac lacking two pairs of strongly sclerotized pieces. The antennomeres III and IV appear slightly shorter in S. lobatum, a character not to be used as diagnostic until more material available to assess its variability more reliably. The length of the antennomere VI of S. bilobum was given in the original description as equal to that of the antennomere VII by error (Löbl, 2015b: 117), it is equal to that of the antennomere V., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas, pp. 103-118 in Revue suisse de Zoologie 129 (1) on page 117, DOI: 10.35929/RSZ.0064, http://zenodo.org/record/6461022, {"references":["Lobl I. 2015 b. On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Island. Revue suisse de Zoologie 122: 65 - 120."]}

Scaphisoma jacqi Löbl & Ramage, n. sp. http://zoobank.org/F092C8CB-A78E-46E8-9CF2-0FEFE201450E HOLOTYPE: ♂, French Polynesia, Society Islands, Raiatea Island, Faaroa Valley, 16°50’51.2’’S 151°25’47.7’’W, 57 m, Malaise trap, 1-8.III.2021, leg. Frédéric Jacq; MHNG-ENTO 0093802 (MHNG). Description. – Length 1.28 mm, width 0.87 mm. Head and pronotum blackish (fig. 1-2). Elytra blackish along bases, suture, lateral margins and on transverse area posterior of mid-length and anterior apical sixth. Area anterior of blackish transverse band dark reddish brown, apical sixth yellowish. Venter of thorax and ventrite I blackish, following abdominal segments and appendages light, ochraceous or yellowish. Pronotum not microsculptured, with lateral margins evenly arcuate, lateral margins carinae visible in dorsal view, lateral striae finely punctate, discal punctation dense and fine, consisting of well delimited punctures much smaller than puncture intervals. Exposed tip of scutellum minute, triangular. Elytra not microsculptured, moderately narrowed apically, with lateral margins nearly evenly rounded, lateral margins striae impunctate near bases, densely punctate posterior basal third of lateral length; apical margins rounded, inner apical angles situated in level with outer apical angles; sutural margin not raised, sutural striae starting at level of scutellar tip, straight, gradually converging apically; adsutural areas flat, combined 0.16 mm wide at level of scutellar tip, each with three irregular puncture rows in basal third reduced to a single puncture row in apical third. Hind wings fully developed. Hypomeron not microsculptured, smooth, lacking traces of punctation. Mesepimeron nearly four times as long as wide and slightly longer than interval to mesocoxa. Median part of metaventrite slightly convex, lacking impressions. Lateral areas of metaventrite not microsculptured, without antecoxal puncture row, sparsely and very finely punctate. Submesocoxal area about 0.05 mm, as fourth of interval to metacoxa. Submesocoxal line arcuate, fairly coarsely punctate. Metanepisternum flat, narrowed anteriad, with slightly sinuate suture, not microsculptured, with punctation as that on sides of metaventrite. Tibiae straight. Abdomen with transversely striate microsculpture. Ventrite I as finely and sparsely punctate as lateral parts of metaventrite; submetacoxal area about 0.08 mm, as two thirds of interval to apical margin; submetacoxal line convex, fairly coarsely punctate. Male characters. Protarsomeres I to III distinctly widened, narrower than apices of tibiae. Mesotarsomeres not widened.Apical process of ventrite VI triangular, acute, 0.02 mm long.Aedeagus (fig. 3-5) 0.49 mm long. Basal bulb large, about twice as long as apical process, with robust, ventrally prominent articular process. Ventral branch of apical process sinuate, in lateral view, lacking denticles. Parameres sinuate in lateral view, arcuate and narrowed posterior of mid-length in dorsal view. Internal sac with two apical bunches of long, straight denticles, single large, straight mesal denticle and two basal vesicles containing dense, curved denticles appearing in dorsal view as irregularly striate structures. Etymology. – The specific epithet refers to the family name of the collector of the species, Frédéric Jacq, a great naturalist and photographer, and for his huge contribution to the knowledge of the entomofauna of French Polynesia. The species epithet is to be treated as a noun in the genitive case. Habitat. – The Malaise trap (fig. 6-7) was set along an abandoned trail, in a Talipariti tiliaceum (L.) Fryxell secondarized forest, degraded with Syzygium cumini (L.) Skeels, Ardisia elliptica Thunb., and Passiflora laurifolia L. The herbaceous understory was composed of Sphagneticola trilobata (L.) Pruski, Christella sp., Zingiber zerumbet (L.) Roscoe ex Sm., Nephrolepis hirsutula (G.Forst.) C.Presl and Centotheca lappacea (L.) Desv. Comments. – The species is a member of the Scaphisoma haemorrhoidale group, defined by the aedeagi symmetrical with a trifid apical process, narrow, simple parameres usually lacking lobes and a complex internal sac lacking flagellum (LÖBL, 1970). The aedeagal characters suggest relationships with S. perkinsi Scott, 1908, described from Hawaii, redescribed and subsequently recorded from the Bonin (Ogasawara) Islands (LÖBL, 1981). The new species may be readily distinguished from S. perkinsi by the pattern of elytral colour (the latter has elytra uniformly black between bases and the light apical area) and by the internal sac of the aedeagus bearing an elongate mesal denticle absent from S. perkinsi, and lacking an elongate V-shaped sclerite., Published as part of Ramage, Thibault & Löbl, Ivan, 2022, A new species of Scaphisoma Leach from the Society Islands with commentary on Staphylinidae of the French Polynesia (Coleoptera, Staphylinidae, Scaphidiinae), pp. 55-60 in Bulletin de la Société entomologique de France 127 (1) on pages 57-59, DOI: 10.32475/bsef_2224, http://zenodo.org/record/6461191, {"references":["LOBL I., 1970. - Revision der palaarktischen Arten der Gattungen Scaphisoma Leach und Caryoscapha Ganglbauer der Tribus Scaphisomini (Col. Scaphidiidae). Revue suisse de zoologie, 77: 727 - 799. https: // doi. org / 10.5962 / bhl. part. 75925","LOBL I., 1981. - Insects of Micronesia Coleoptera Scaphidiidae. Insects of Micronesia, 15: 69 - 80."]}

Scaphisoma konvickai ��sp.��nov. (Figs 1���3) Type�� material. HolotYPE:�� CHINA: male: China, N. Sichuan, Xiao-Zhaizi Nt. Nature Reserve, 7 km of Qingpianxian, Xiaozhaizi, 32��1���25������N, 103��56���21������E, 27. VI.���1.VII.2017, 1560��� 1700 m, leg. O. Konvička (NMPC). Etymology. Patronymic. The species is named in honour of its collector, the coleopterist Ondřej Konvička (Zl��n, Czech Republic). Description. Length 1.65 mm, width 1.17 mm. Head with blackish frons. Pronotum dark brown on large median area, ochraceous along anterior margin and on lateral areas. Elytra ochraceous, each darkened along base, with dark basal area expanded along suture to form triangular spot reaching sutural mid-length, and with narrow subapical darkened stripe. Hypomera ochraceous. Mesoventrite and metaventrite with anepisterna and epimera dark brown to blackish. Ventrites I to IV dark brown, apical abdominal segments yellowish. Femora and tibiae ochraceous, tarsi and antennae yellowish. Length/width ratios of antennomeres as: III 15/8: IV 25/7: V 30/8: VI 48/9: VII 55/16: VIII 47/11: IX 52/16: X 50/15 (left antennomeres X and XI and right antennomere XI broken off and missing). Pronotum lacking miscrosculpture, with lateral margins nearly evenly rounded, lateral margin carinae concealed in dorsal view, these carinae very finely and densely punctate, discal punctation fine and dense, most puncture intervals usually two to four times as large as puncture diameters, punctures well delimited, clearly visible at 25 times magnification. Exposed point of scutellum minute. Elytra not microsculptured, with lateral margins evenly rounded, carinae of lateral margin exposed in dorsal view, lateral margin striae finely and densely punctate, apical margins rounded, inner apical angles not prominent, about in level with outer apical angles, marginal crenulation absent, sutural margin not raised, sutural striae shallow, parallel, each with single puncture row, bent at bases and extended laterally to form basal striae reaching elytral mid-width; discal punctures fine and dense, coarser than pronotal or adsutural punctures, not forming rows, puncture intervals mostly about as larger to two times as large as puncture diameters. Exposed tergites with punctation hardly visible at 100 times magnification. Hypomera not microsculptured, appearing impunctate. Mesepimon about three times as long as wide and distinctly shorter than interval to mesocoxa. Metaventrite with striate microsculpture on median and apicolateral areas, not microsculptured on most of lateral surface. Median part of metaventrite weakly convex, not impressed apically, lacking separate impressions near metacoxae. Punctation on metaventrite sparse and very fine, consisting of well delimited punctures; antecoxal puncture rows absent. Submesocoxal area 0.05 mm long, about as fourth of shortest interval to metacoxa. Submesocoxal line convex, very finely punctate. Metanepisterna slightly convex, narrowed anteriad, with straight suture. Abdomen with transversely striate microsculpture, except on basolateral areas of ventrite I, and very finely and sparsely punctate. Ventrite I very finely punctate, submetacoxal area 0.10 mm long, nearly as long as interval to apical margin of ventrite. Tibiae straight. Male characters. Protarsomeres I to III slightly widened. Mesotarsomeres not widened. Apex of ventrite VI gradually narrowed, not forming distinctive lobe. Aedeagus as in Figs 1���3, 0.68 mm long, moderately sclerotized. Basal bulb large, convex and lacking articular tubercle at base of parameres. Apical process of median lobe strongly inflexed, with dorsal branch only moderately shorter than ventral branch, ventral branch gradually narrowed and with acute tip. Parameres weakly arcuate, as long as half of median lobe. Internal sac consisting of convoluted, spinose tube. Differential��diagnosis. This new species leads to S. latro L��bl, 2000, couplet 24, in the key to the Chinese Scaphisoma (L��bl 2019a). However, the latter species is unrelated (based on aedeagal characters) and differs notably by its colour pattern and the antennomeres V and VI similar in length. The aedeagal characters of S. konvickai suggest relationships to S. invertum L��bl, 2000 from Guangxi and Yunnan, S. invisum L��bl, 1990 from Thailand, S. discretum L��bl, 1986 from Northeast India, and S. binaluanum Pic, 1947 from the Philippines. The new species is distinguished from all of these by the short apical process of the median lobe, compared to the basal bulb, and the complexly convoluted internal sac, similar to that of the Sulawesian S. bugi L��bl, 1983. It differs in external characters from S. invertum notably by the elytra having basal striae, the mesepimeron shorter than the interval to the mesocoxa, and the strigulate abdominal microsculpture. Scaphisoma invertum possesses elytra darkened along the base, in the middle and near the apices, lacks basal striae, and the exposed abdominal segments have punctulate microsculpture. Scaphisoma invisum differs by its body being only 1.05���1.15 mm long, the antennomere V being longer than the antennomere VI and the submesocoxal area longer than the submetacoxal area. Scaphisoma discretum may be easily distinguished by its antennomeres VI being about as long as the antennomeres III to V, combined with the elytra lacking basal striae. Scaphisoma binaluanum also has elytra without basal striae, and is characterized by the well-defined transverse band posterior to the elytral mid-length, that provides a distinctive colour pattern. Scaphisoma bugi has each elytron with two reddish spots, lacking basal striae and an apicoventrally prominent basal bulb of the aedeagus (L��bl 1983), by which it may be readily distinguished from S. konvickai. FIGURES��1���7. Fig. 1���3. Scaphisoma konvicka sp.��nov., (1) aedeagus in dorsal and lateral views, scale = 0.1 mm; (2) apical part of aedeagus; (3) paramere in ventral view; scale = 0.1 mm for (2, 3). Figs 4���7. Toxidium hartmanni sp.��nov., (4, 5) aedeagus in dorsal and lateral views, scale = 0.2 mm; (6) paramere in ventral view, scale = 0.1 mm; (7) internal sac, scale = 0.1mm., Published as part of L��bl, Ivan, 2022, - On-new-collections-of-Scaphidiinae- (Coleoptera: - Staphylinidae) - from-China, - with-description-of-two-new-species, pp. 487-492 in Zootaxa 5092 (4) on pages 487-489, DOI: 10.11646/zootaxa.5092.4.8, http://zenodo.org/record/5886555, {"references":["Lobl, I. (2019 a) New species and records of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae) from the People's Republic of China. Annales Zoologici, 69, 241 - 292. https: // doi. org / 10.3161 / 00034541 ANZ 2019.69.2.002","Lobl, I. (1983) Sechs neue Scaphidiidae (Coleoptera) von Sulawesi, Indonesien. Mitteilungen des Schweizerischen Entomologischen Gesellschaft, 56, 285 - 293."]}

Scaphisoma serosum ��L��bl,��2000 Scaphisoma serosum L��bl, 2000: 614. Type locality: China: Sichuan, Wolong Nat. Res., 1000 m. Material��examined. 1 ex. N-Sichuan, Xiao-Zhaizi Nat. Nature Reserve, 15 km W of Qingpianxiang, Dayaochang, 31��58���18������N 103��51���16������E, 1���2.VII.2017, 1900���2100 m, leg. O. Konvička (PCOK). Note. This species was based on material coming from Wolong National Nature Reserve, in the Chinese Province of Sichuan. It has not been reported since its description., Published as part of L��bl, Ivan, 2022, - On-new-collections-of-Scaphidiinae- (Coleoptera: - Staphylinidae) - from-China, - with-description-of-two-new-species, pp. 487-492 in Zootaxa 5092 (4) on page 492, DOI: 10.11646/zootaxa.5092.4.8, http://zenodo.org/record/5886555

Episcaphium watanabai ��L��bl,��2002 Episcaphium watanabai L��bl, 2002: 289. Type locality: China: Sichuan, Ya���an, Xiaoxiang Ling, side valley above Nanya Cun, 11 km south Shimian, 1250 m. Material�� examined. 10 exx. North Sichuan, Xiao-Zhaizi Nat. Nature Reserve, 4 km NNE of Qingpianxiang, Zhenghecun 32��3���27������N 103��59���37������E, 23���26. VI.2017, 1350��� 1850 m, leg. O. Konvička (PCOK, MHNG); 1 ex. North Sichuan, Xiao-Zhaizi Nat. Nature Reserve, 7 km W of Qingpianxiang, Xiaozhaizi, 32��1���25������N 103��56���21������E, 27. VI.���1.VII.2017, 1560��� 1700 m, leg. O. Konvička (PCOK). Note. This species was based on specimens from Xiaoxiang Ling in the Province of Sichuan and from Taiwan (L��bl 2002). It has not been reported since its description., Published as part of L��bl, Ivan, 2022, - On-new-collections-of-Scaphidiinae- (Coleoptera: - Staphylinidae) - from-China, - with-description-of-two-new-species, pp. 487-492 in Zootaxa 5092 (4) on page 491, DOI: 10.11646/zootaxa.5092.4.8, http://zenodo.org/record/5886555, {"references":["Lobl, I. (2002) Two new species of Episcaphium Achard (Coleoptera, Staphylinidae, Scaphidiinae). Special Bulletin of the Japanese Society of Coleopterology, 5, 289 - 295."]}

Löbl, Ivan (2022): -On-new-collections-of-Scaphidiinae-(Coleoptera:-Staphylinidae)-from-China,- with-description-of-two-new-species. Zootaxa 5092 (4): 487-492, DOI: https://doi.org/10.11646/zootaxa.5092.4.8

Baeocera franzi (Löbl, 1973) Material examined. 1 ♂: Zhejiang Prov., Lin’an county, West Tianmushan Nat. Res., 30.3255°N, 119.4406°E, 430 m, (WT04) sift #02, large bamboo, bushes above Chanyuan Temple, 24.vi.2017, J. Hájek & J. Růžička leg. (NMPC). Comments. The species was described from Thailand and subsequently recorded from Fujian, Hubei, Jiangsu, Shaanxi, Sichuan and Yunnan provinces of China (L ÖBL 1999, 2003). First record from Zhejiang.

Scaphisoma binhanum (Pic, 1922) Material examined. 6 ♂♂ 9 ♀♀: Fujian Prov., Wuyishan Mts. NNR: Sangang vill., 27°45.0′N, 117°40.7′E, 720 m, river valley, mixed forest and bamboo, 24.v.–3.vi.2018, J. Hájek, D. Král, J. Růžička & L. Sekerka leg. (IZCAS, NMPC); 1 ♀: Fujian Prov., Wuyishan Mts. NNR: Guadun vill., 27°44.0–2′N, 117°38.3–7′E, 1075–1250 m, mixed forest + bamboo, on vegetation, 25. + 29.v.2018, J. Hájek, D. Král, J. Růžička & L. Sekerka leg.; 1 ♀: Fujian Prov., Wuyishan Mts. NNR, 1.3 km E Legaduc vill., Yaoyuanyu Valley, 27°43.693′N, 117°43.056′E, (W18) 640 m, sift #11, detritus along path, mixed broad-leaved forest with Cunninghamia, 28.v.2018, D. Král & J. Růžička leg. (NMPC, MHNG). Comments. The species was based on specimens coming from Vietnam, and subsequently reported from India, Nepal, Thailand, Indonesia, and China (see L ÖBL 2018b). At present, it is known from the Chinese provinces of Guangxi, Hainan and Jiangxi (L ÖBL 2019a). First records from Fujian.

Scaphisoma geminatum Löbl, 1986 Material examined. 28 spec.: Fujian Prov., Wuyishan Mts. NNR, 0.5 km SE Tongmu-Guadun, 27°43.981′N, 117°38.375′E, 1230 m, (W09) sift #05, detritus with mycelia, dense short bamboo forest with inter-mixed high evergreen trees, 25.v.2018, D. Král & J. Růžička leg. (IZCAS, NMPC); 10 spec., Fujian Prov., Wuyishan Mts. NNR, 4.8 km SW Tongmu-Sangang, Xiaofeng Mt., 27°42.707′N, 117°39.128′E, 1150 m, (W14) sift #08, detritus under dense shrubs, mixed broad-leaved forest with bamboo, 27.v.2018, D. Král & J. Růžička leg. (NMPC); 1 ♂, Fujian Prov., Wuyishan Mts. NNR, 1.3 km E Legaduc vill., Yaoyuanyu valley, 27°43.693′N, 117°43.056′E, 640 m, (W18) sift #11, detritus along path, mixed broad-leaved forest with Cunninghamia, 28.v.2019, D. Král & J. Růžička leg. (NMPC). Comments. This species was described from Meghalaya, northern India, and was subsequently reported from Fujian, Guangdong and Jiangxi provinces of China (L ÖBL 2000, 2003).

Scaphisoma sekerkai Löbl, sp. nov. (Figs 2, 5–8) Type locality. China: Fujian Prov.,Wuyishan Mts.NNR,Guadun vill.env., 27°44.6′N, 117°38.1′E, 1335 m. Type material. H OLOTYPE ♂: CHINA: FUJIAN Prov. 29.v.2018 Wuyishan Mts. NNR, Guadun vill. env., mixed forest with flowering Castanopsis, individually from vegetation, dead wood, fungi, 27°44.6′N, 117°38.1′E, 1335 m J.Hájek, D.Král, J.Růžička & L.Sekerka lgt. (IZCAS).P ARATYPES: 1 ♀: with the same data as the holotype (NMPC); 2 ♂♂ 2 ♀♀: CHINA: FUJIAN Prov. 23.v.–3.vi.2018 Wuyishan Mts.NNR:Sangang 27°45.0′N, 117°40.7′E, 720 m; mixed forest + bamboo; on vegetation Hájek, Král, Růžička & Sekerka lgt. (MHNG, NMPC). Description. Length 1.51–1.67 mm, width 0.95–1.05 mm. Head blackish. Pronotum entirely dark brown to blackish, or somewhat lighter to ochraceous along lateral margin. Hypomera ochraceous. Elytra reddish-brown, each slightly to strongly darkened along suture, with dark brown to blackish base, apices, and transverse band situated anterior to elytral midlength, latter reaching to or separated from sutural stria. Mesoventrite and metaventrite with anepisterna and epimera dark brown to blackish, ventrites light brown to yellowish, exposed abdominal tergites and appendages yellowish. Length/width ratios of antennomeres as: III 14/9: IV 28/8: V 48/8: VI 43/8: VII 50/12: VIII 42/10: IX 58/12: X 50/12: XI 60/12. Pronotum not microsculptured, with lateral margins nearly evenly rounded, lateral margin carinae exposed in dorsal view, lateral margin carinae finely and densely punctate, discal punctation fine and dense, most puncture intervals slightly larger to three times larger than puncture diameters, punctures well delimited, clearly visible at 15 times magnification. Exposed point of scutellum minute. Elytra not microsculptured, with lateral margins rounded in basal third, oblique posteriad, apical margins rounded, lateral margin carinae exposed in dorsal view, lateral margin striae finely and densely punctate, inner apical angles not prominent, about on level with outer apical angles, marginal crenulation very fine, sutural striae shallow, not or hardly bent at bases, parallel in basal halves, converging posteriad, adsutural areas flat, each with single puncture row, basal striae absent, discal punctures fairly fine and dense, coarser than pronotal or adsutural punctures, not forming rows, puncture intervals mostly about two to three times as large as puncture diameters. Exposed tergites with few very fine punctures. Hypomera with longitudinally striate microsculpture, appearing impunctate. Mesepimeron about 5 times as long as wide and 2.0 to 2.5 times as long as interval to mesocoxa. Metaventrite with transversely striate microsculpture. Median part of metaventrite weakly convex, slightly impressed apically, lacking separate impressions near metacoxae, with punctation distinct, sharply delimited, sparse on centre, fairly dense on apical area. Lateral parts of metaventrite each with dense antecoxal puncture row, punctation very fine and sparse on remaining surface. Submesocoxal area 0.04 mm long, about fourth of shortest interval to metacoxa; submesocoxal line convex, densely punctate. Metanepisterna slightly convex, impressed along arcuate suture, narrowed anteriad. Abdomen with transversely striate microsculpture.Ventrite I with few distinct basomedian punctures, very finely punctate on prevailing surface, submetacoxal area 0.08 mm long, about half of interval to apical margin of ventrite. Following ventrites very finely and sparsely punctate. Protibiae straight, mesotibiae and metatibiae weakly bent. Male characters. Protarsomeres I distinctly widened, II slightly widened, III hardly widened. Mesotarsomeres I weakly widened, II and III hardly widened.Apex of ventrite VI gradually narrowed to form about 0.05 mm long triangle. Aedeagus (Figs 5–8) 0.67–0.70 mm long. Differential diagnosis. Member of the Scaphisoma rouyeri group, as defined in L ÖBL (1981a) and subsequently extended to include species lacking parameral lobes (e.g., L ÖBL & OG AWA 2016). The species is characterized by the following features, in combination: the elytra are darkened at bases, on the transverse stripe posterior to midlength and at the apices, the elytral punctation is not arranged to form distinct striae, the sutural striae start near pronotal lobe, the submesocoxal areas are about as long as one fourth of the shortest interval between them and the metacoxae, the submetacoxal areas are twice as long as the submesocoxal areas and half as long as the shortest intervals between them and the apical margin of the ventrite I, the metaventrite has dense antecoxal puncture rows, the ventrite I is very finely punctate except for its basomedian area, the apical process of the median lobe is curved and tapering, the parameres are strongly expanded near bases and sinuate in dorsal view and lack mesal lobes, the internal sac is tubular, strigulate in proximal section, and bears rows of marginal denticles. This new species falls in the key to the Chinese Scaphisoma (L ÖBL 2019a) under the couplet 54, to S. delictum Löbl, 1981. It may be readily distinguished from the latter by the elytral colour pattern, the elytra lacking coarse puncture rows, and the parameres lacking mesal lobes. The parameres of this new species are similar to those of S. ruzickai Löbl, 2019. These two species differ notably in their elytral colour pattern and punctation (much finer in S. sekerkai sp. nov.), and the structures of the internal sac (with a median tuft of robust teeth in S. ruzickai, with long marginal rows of short denticles and a striate basal tube in S. sekerkai sp. nov.). Collection circumstances. At the type locality, Scaphisoma sekerkai sp. nov. was collected on the border of tea plantation and mixed forest with flowering Castanopsis sp. (Fig. 13); the precise collection circumstances are not known. Etymology. Patronymic. The species is named in honour of one of its collectors, the well-known leaf beetle expert Lukáš Sekerka (NMPC). Distribution. So far known only from two very close localities in the Wuyishan Mts. NNR, Fujian, China.

Kasibaeocera mussardi (Löbl, 1971) Material examined. 1 spec.: Zhejiang Prov., Lin’an county, West Tianmushan Nat. Res., 30.3255°N, 119.4406°E, 430 m, (WT04) sift #02, large bamboo, bushes above Chanyuan Temple, 24.vi.2017, J. Hájek & J. Růžička leg. (NMPC). Comments. Widely distributed Oriental species described from Sri Lanka and subsequently recorded from India, Nepal, Bhutan, China and Thailand (LÖBL 2018b).

Scaphisoma mutator Löbl, 2000 Material examined. 1 ♀: Zhejiang Prov., Lin’an County, West Tianmushan Nat. Res., 100 m SE below of top of Immortal Peak, 30.3491°N, 119.4244°E, 1470 m, (WT13) sift #07, dwarf forest, under bamboo and shrubs, near water source, 28.vi.2017, J. Hájek & J. Růžička leg. (NMPC); 1 ♀: same data but 350 m NWW of Original Temple of Lion Sect, 30.3426°N, 119.4302°E, 1190 m, (WT14) sift #13, broad-leaved deciduous forest, detritus along rocks and fallen logs near brook, 2.vii.2017 (NMPC); 1 ♂, 1 ♀: same data but 400 m N Tianmu vill., 30.3157°N, 119.4441°E, 340–350 m, (WT02) sift #01, mixed broad-leaved forest with Cunninghamia and Liquidambar near brook, 23.vi.2017 (MHNG, NMPC). Comments. This species was based on specimens found in Sichuan, and was subsequently reported from Guangdong, Guangxi and Shaanxi provinces of China (L ÖBL 2019a). First records from Zhejiang.

Pselaphogenius Reitter, 1910 This genus comprises 76 species (NEWTON 2020), many of them described from the temperate part of the Euro -Asian continent. The group was redefined by NOMURA (1998) and its east Asian members were keyed in NOMURA (2003). Two species have been collected in Nepal, one is described below, the second represented by two females, is probably also new. We decline to describe it in absence of at least one available male.

Nepallaphus dve n. sp. (Figs 28, 29) Holotype ♂: NEPAL (Prov. Bagmati) Pokhare NE Barahbise 2800 m 2.V.81 LÖbl & Smetana (MHNG). Paratypes: 1 ♂, 2 ♀, NEPAL, Bagmati Zone, Yardang Ridge NE Barahbise, 3250 m, 5.IV.1981, I. LÖbl & A. Smetana (MHNG); 1 ♂, Bagmati Zone, Kalingchok NE Barahbise, Amatal Khola, Tingsang La, 3100 m, 27.X..1981, P. Cassagnau (MHNG); 1 ♀, Bagmati Zone, Kalingchok NE Barahbise, Yardang Ridge, 3250 m, 28.X.1981, P. Cassagnou (MHNG). Etymology. The species epithet is the numbertwo in Sanskrit. Description. Length 1.60-1.82 mm. Head about 1.7 times as long as wide. Lower margin of frontoclypeus rounded, with long, prominent setae.Anterior marginof frontal rostrum rounded. Frontal sulcus between antennal tubercles narrow, widened posteriad, with carinate inner margins, extended to level of eye mid-length. Frontal carinae oblique, reticulate. Antennal tubercles prominent. Vertex convex, with mesal impression very shallow, narrowed posteriad, mesal carina near neck absent. Eyes prominent, with diameter about as half of interval to neck, consisting of seven facets. Genae reticulate. Temples weakly rounded, contours in dorsal view nearly oblique. Frontal carinae bearing rowsof short appressed setae. Temples and vertex with short appressed setae. Maxillary palpomere IV long about as head, with peduncle somewhat shorter than thickened apical part. Length/width ratio of antennomeres as: I 22/10: II 12/8: III 9/6: IV 7/6: V 7/6: VI 8/6: VII 8/6: VIII 8/6: IX 10/7: X 12/8: XI 24/13. Gular mound longer than wide, narrowed posteriad. Pronotum similar to that in N. ekam. Elytra nearly 1.4 times as long as pronotum, with lateral margins weakly rounded, at apices about 3.5 times as wide asat bases, not microsculptured, with three basal foveae, median longitudinal carina with margins well delimited, reaching nearly elytral mid-length, bearing double rows of setae, adsutural carina with single row of setae; disc with additional, outer double row of setae, intervals between rows of setae glabrous. Hind wings atrophied. Abdomen with tergite I slightly wider than elytra, broad paratergites, rounded lateral margins. Male characters. Metaventrite as in N. ekam. Aedeagus (Figs 28, 29) 0.35-0.37 mm long, with basal bulb not widened apically, its apicodorsal band narrowly notched in middle, dorsal capsule oval. Measurements of the holotype (in mm). Total length 1.82; length/width of head 0.41/0.24, pronotum 0.33/0.30, elytra 0.45/0.70 (0.20), tergite I 0.51/0.75; length of maxillary palpomere IV 0.40, length of aedeagus 0.37. Diagnosis. Length 1.60-1.82 mm. Anterior margin of frontal rostrum rounded. Frontal sulcus rather wide, distinctly widened posteriad. Maxillary palpomere IV long about as head, with peduncle shorter than thickened apical part. Elytron with discal carina short, extended to mid-length of disc, interval between carinae and laterally discal carina glabrous. Aedeagus with basal bulb not widened apically, oval apicodorsal capsule, and minute notch in middle of apicodorsal band. Distribution. Central Nepal. Comments. See key below to distinguish this species from its congeners.

Nepallaphus ekam n. sp. (Figs 26, 27) Holotype ♂: NEPAL (Prov. Bagmati) near Mere Dara, 3000 m, 7.IV.1981 LÖbl & Smetana (MHNG). Paratypes: 5 ♂, 5 ♀, NEPAL, with the same data as the holotype (MHNG, PCPH); 1 ♂, 1 ♀, Bagmati Zone, Mere Dara, 3200 m, north slope, 8.IV.1981, I. LÖbl & A. Smetana (MHNG); 1 ♂, 7 ♀, Bagmati Zone, below Thare Pati, 3300 m, 9.IV.1981, I. LÖbl & A. Smetana (MHNG, PCVB). Etymology. The species epithet is the number one in Sanskrit. Description. Length 1.70-1.90 mm. Head about 1.6 times as long as wide. Lower margin of frontoclypeus rounded, with long, prominent setae. Anterior margin of frontal rostrum prominent, notched in middle, bidentate. Frontal sulcus narrow, with carinate inner marginsextended to level of eye mid-length. Frontal carinae horizontal, reticulate.Antennal tubercles prominent. Vertex convex, lacking distinct mesal impression, mesal carina near neck absent. Eyes prominent, with diameter as half of interval to neck, consisting of seven to eight facets. Genae reticulate. Temples weakly rounded, in dorsal view with nearly oblique contours. Frontal carinae bearing rows of short appressed setae. Temples and vertex with short appressed setae. Maxillary palpomere IV longer than head, with peduncle about as long as thickened apical part. Length/width ratio of antennomeres as: I 24/10: II 12/9: III 10/6: IV 9/6: V 9/6: VI 9/6: VII 10/6: VIII 10/6: IX 13/8: X 14/9: XI 28/14. Pronotum slightly longer than wide and slightly shorter than head, with punctiform median antebasal fovea, distinct lateral antebasal foveae, punctation sparser than on head. Elytra about 1.4 times as long as pronotum, with lateral margins rounded, at apices well 4 times as wide as at bases, not microsculptured, with three basal foveae, median longitudinal carina with margins well delimited, extending to or nearly to apical third of elytral disc, adsutural and median setose rows simple, disc with outer double row od setae, intervals between rows of setae glabrous. Hind wings atrophied. Abdomen with tergite Islightly wider than elytra, withbroad paratergites and roundedlateral margins. Male characters. Middle of metaventrite raised to form a flat, circular area inflexed apically and completely covered by short, appressed setae. Aedeagus (Figs. 26, 27) 0.49-0.53 mm long, with basal bulb not widened apically, its sclerotized apicodorsal band not notched, dorsal capsule cordiform. Measurements of the holotype (in mm). Total length 1.76; length/width of head 0.39/0.25, pronotum 0.35/0.32, elytra 0.49/0.76 (0.17), tergite I 0.50/0.83; length of maxillary palpomere IV 0.43, length of aedeagus 0.50. Diagnosis. Length 1.70-1.90 mm. Anterior margin of frontal rostrum prominent andnotched in middle. Frontal sulcus narrow, parallel-sided. Maxillary palpomere IV longer than head, with peduncle about as long as thickened apical part. Elytron with discal carina extending to or nearly to apical third of disc. Interval between carinae and laterally discal carina glabrous. Aedeagus with basal bulb not widened apically, its apicodorsal band not notched, dorsal capsule cordiform. Distribution. Central Nepal. Comments. See key below to distinguish this species from its congeners.

Nepallaphus treeni n. sp. (Figs 30, 31) Holotype ♂: Népal 15.VII.1981 Langtang N. Park près Gosaikunda Lake 4400 m Beron [handwritten] (NMNS) Etymology. The species epithet is the number three in Sanskrit. Description. Length 2.10 mm. Headabout 1.6 longer than wide. Lower margin of frontoclypeus rounded, with long, prominent setae. Middle of anterior margin of frontal rostrum shallowly notched, bidentate. Frontal sulcus between antennal tubercles narrow, parallel to level of anterior eye margins, widened between eyes, with carinate inner margins, extended to level of eye mid-length. Frontal carinae oblique, reticulate.Antennal tubercles prominent. Vertex convex, with mesal impression very shallow, narrowed posteriad, mesal carina near neck absent. Eyes prominent, with diameter about as half of interval to neck, consisting of eight facets. Genae reticulate. Temples weakly rounded, in dorsal view with nearly oblique contours. Frontal carinae bearing rows of short appressed setae. Temples and vertex with short appressed setae. Maxillary palpomere IV long as head, with peduncle somewhat shorter than thickened apical part. Length/width ratio of antennomeres as: I 26/11: II 12/9: III 8/6: IV 8/6: V 8/6: VI 8/6: VII 8/6: VIII 8/6: IX 10/7: X 12/9: XI 26/15. Gular mound wider than long, nearly circular. Pronotum similar to that of N. ekam. Elytra about 1.3 times as long as pronotum, with lateral margins oblique in basal halves, weakly rounded posteriad mid-length, at apices nearly 4 times as wide as at bases, not microsculptured, with three basal foveae, inner two foveae in elongate impression, median longitudinalcarina absent, adsutural carina with single row of setae; entire disc setose. Abdomen with tergite I slightly wider than elytra, broad paratergites, rounded lateral margins. Male characters. Metaventrite as in N. ekam. Aedeagus (Figs 30, 31) 0.62 mm long, with basal bulb widened apically, its apicodorsal band deeply notched in middle, dorsal capsule oval. Measurements of the holotype (in mm). Total length 2.10; length/width of head 0.41/0.26, pronotum 0.36/0.31, elytra 0.48/0.75 (0.20), tergite I 0.64/0.84; length of maxillary palpomere IV 0.40, length of aedeagus 0.62. Diagnosis. Length 2.10 mm. Anterior margin of frontal rostrum bidentate. Frontal sulcus rather wide, distinctly widened posteriad. Maxillary palpomere IV long as head, with peduncle shorter than thickened apical part. Elytron without discal carina, pubescence covering entire elytral disc. Basal bulb of aedeagus widened apically, its apicodorsalband deeply notched. Distribution. Central Nepal. Comments. The state of the hind wings was not examined. The narrow elytral bases and the small eyes suggest reduced hind wings. See key to distinguish the species from its congeners.