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QI, SHUO, LYU, ZHI-TONG, WANG, JIAN, MO, YUN-MING, ZENG, ZHAO-CHI, ZENG, YANG-JIN, DAI, KE-YUAN, LI, YUAN-QIU, GRISMER, L. LEE, WANG, YING-YONG (2022): Erratum: SHUO QI, ZHI-TONG LYU, JIAN WANG, YUN-MING MO, ZHAO-CHI ZENG, YANG-JIN ZENG, KE-YUAN DAI, YUAN-QIU LI, L. LEE GRISMER & YING-YONG WANG (2022) Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China. Zootaxa, 5072: 401–438. Zootaxa 5115 (4): 600-600, DOI: 10.11646/zootaxa.5115.4.10, URL: http://dx.doi.org/10.11646/zootaxa.5115.4.10

III. Genus Atympanophrys Tian & Hu, 1983 Suggested common name: Hidden-tympanum horned toads (in English) / ĿHDzfl (in Chinese) Four species: Atympanophrys shapingensis (Liu, 1950) (type species); Atympanophrys gigantica (Liu, Hu & Yang, 1960); Atympanophrys nankiangensis (Liu & Hu, 1966); Atympanophrys wawuensis (Fei, Jiang & Zheng, 2001)., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

2) Boulenophrys minor group Three species: Boulenophrys minor (Stejneger, 1926); Boulenophrys chishuiensis (Xu, Li, Liu, Wei & Wang, 2020) comb. nov.; Boulenophrys jiangi (Liu, Li, Wei, Xu, Cheng, Wang & Wu, 2020) comb. nov.., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 431, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

3) Boulenophrys omeimontis group Fifteen species: Boulenophrys omeimontis (Liu, 1950); Boulenophrys anlongensis (Li, Lu, Liu and Wang, 2020) comb. nov.; Boulenophrys angka (Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy & Che, 2019) comb. nov.; Boulenophrys binchuanensis (Ye & Fei, 1995); Boulenophrys binlingensis (Jiang, Fei & Ye, 2009) comb. nov.; Boulenophrys caobangensis (Nguyen, Pham, Nguyen, Luong & Ziegler, 2020) comb. nov.; Boulenophrys daweimontis (Rao & Yang, 1997) comb. nov.; Boulenophrys jingdongensis (Fei & Ye, 1983) comb. nov.; Boulenophrys lushuiensis (Shi, Li, Zhu, Jiang, Jiang & Wang, 2021) comb. nov.; Boulenophrys palpebralespinosa (Bourret, 1937) comb. nov.; Boulenophrys qianbeinsis (Su, Shi, Wu, Li, Yao, Wang & Li, 2020) comb. nov.; Boulenophrys rubrimera (Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong & Rowley, 2017) comb. nov.; Boulenophrys sangzhiensis (Jiang, Ye & Fei, 2008) comb. nov.; Boulenophrys spinata (Liu & Hu, 1973); Boulenophrys wuliangshanensis (Ye & Fei, 1995)., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 431, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

1) Boulenophrys boettgeri group Thirty-nine species: Boulenophrys boettgeri (Boulenger, 1899) (type species); Boulenophrys acuta (Wang, Li & Jin, 2014); Boulenophrys baishanzuensis (Wu, Li, Liu, Wang & Wu, 2020) comb. nov.; Boulenophrys baolongensis (Ye, Fei & Xie, 2007); Boulenophrys brachykolos (Inger & Romer, 1961); Boulenophrys caudoprocta (Shen, 1994) comb. nov.; Boulenophrys cheni (Wang & Liu, 2014); Boulenophrys congjiangensis (Luo, Wang, Wang, Lu, Wang, Deng & Zhou, 2021) comb. nov.; Boulenophrys daiyunensis (Lyu, Wang & Wang, 2021) comb. nov.; Boulenophrys daoji (Lyu, Zeng, Wang & Wang, 2021) comb. nov.; Boulenophrys dongguanensis (Wang & Wang, 2019) comb. nov.; Boulenophrys huangshanensis (Fei & Ye, 2005); Boulenophrys insularis (Wang, Liu, Lyu, Zeng & Wang, 2017) comb. nov.; Boulenophrys jinggangensis (Wang, 2012); Boulenophrys jiulianensis (Wang, Zeng, Lyu & Wang, 2019) comb. nov.; Boulenophrys kuatunensis (Pope, 1929); Boulenophrys leishanensis (Li, Xu, Liu, Jiang, Wei & Wang, 2018) comb. nov.; Boulenophrys liboensis (Zhang, Li, Xiao, Li, Pan, Wang, Zhang & Zhou, 2017) comb. nov.; Boulenophrys lini (Wang & Yang, 2014); Boulenophrys lishuiensis (Wang, Liu & Jiang, 2017); Boulenophrys mirabilis (Lyu, Wang & Zhao, 2020) comb. nov.; Boulenophrys mufumontana (Wang, Lyu & Wang, 2019) comb. nov.; Boulenophrys nankunensis (Wang, Zeng & Wang, 2019) comb. nov.; Boulenophrys nanlingensis (Lyu, Wang, Liu & Wang, 2019) comb. nov.; Boulenophrys obesa (Wang, Li & Zhao, 2014); Boulenophrys ombrophila (Messenger & Dahn, 2019) comb. nov.; Boulenophrys sanmingensis (Lyu & Wang, 2021) comb. nov.; Boulenophrys shimentaina (Lyu, Liu, & Wang, 2020) comb. nov.; Boulenophrys shunhuangensis (Wang, Deng, Liu, Wu & Liu, 2019) comb. nov.; Boulenophrys tongboensis (Wang & Lyu, 2021) comb. nov.; Boulenophrys tuberogranulatus (Shen, Mo & Li, 2010); Boulenophrys wugongensis (Wang, Lyu & Wang, 2019) comb. nov.; Boulenophrys wushanensis (Ye & Fei, 1995); Boulenophrys xiangnanensis (Lyu, Zeng & Wang, 2020) comb. nov.; Boulenophrys xianjuensis (Wang, Wu, Peng, Shi, Lu & Wu, 2020) comb. nov.; Boulenophrys yangmingensis (Lyu, Zeng, & Wang, 2020) comb. nov.; Boulenophrys yaoshanensis Qi, Mo, Lyu, Wang & Wang sp. nov.; Boulenophrys yingdeensis Qi, Lyu, Wang & Wang sp. nov.; Boulenophrys yunkaiensis Qi, Wang, Lyu & Wang sp. nov.., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 431, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

1) Brachytarsophrys carinense group Two species: Brachytarsophrys carinensis (Boulenger, 1889) (type species); Brachytarsophrys intermedia (Smith, 1921)., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

Boulenophrys yunkaiensis sp. nov. Qi, Wang, Lyu & Wang Yunkai Horned Toad / yun kai jiao chan (云开fflDz) Figures 6, 7C Chresonymy. Megophrys sp 32 (SYS a004637–4638, 4694)— Liu et al. 2018 Holotype. SYS a004637 (Figs. 6A, 7C), adult male, collected on 14 April 2016 by Jian Wang, Zhi-Tong Lyu and Ying-Yong Wang from the Yunkaishan Nature Reserve (22.2758°N, 111.1952°E; ca 950 m a.s.l.), Xinyi City, Guangdong Province, China. Paratypes. Eight adult specimens from the same locality as the holotype: males SYS a004636 and SYS a004638 collected on 14 April 2016 by Jian Wang, Zhi-Tong Lyu and Ying-Yong Wang; male SYS a004662/ CIB 116085 and females SYS a004659 (Figs. 6B) and SYS a004660 collected on 15 April 2016 by Jian Wang, Zhi-Tong Lyu and Ying-Yong Wang; female SYS a004691 collected on 16 April 2016 by Jian Wang, Zhi-Tong Lyu and Ying-Yong Wang; males SYS a004986 and SYS a004987 collected on 28 June 2016 by Jian Wang. Etymology: The specific epithet yunkaiensis refers to its type locality, the Yunkaishan Nature Reserve, western Guangdong, China. Diagnosis. (1) Small body size, SVL 35.3–40.0 mm (37.6 ± 1.7, N = 6) in adult males and SVL 45.3–46.1 mm (45.8 ± 0.4, N = 3) in adult females; (2) snout rounded in dorsal view; (3) tympanum boundary clear, ED/TD 1.68–1.91 in males, 1.47–1.80 in females; (4) presence of vomerine ridge and absence of vomerine teeth; (5) margin of tongue rounded, not notched behind; (6) hindlimbs slender, heels overlapping or just meeting and tibio-tarsal articulation reaching forward between tympanum to posterior corner of eye when leg stretched forward; (7) tibia 0.40–0.48 of SVL and foot 0.60–0.68 of SVL in males, while tibia 0.42–0.46 of SVL and foot 0.59–0.66 of SVL in female; (8) a subarticular tubercle present at the base of each fingers; (9) toes without lateral fringes and with only rudimentary webbing; (10) presence of small horn-like tubercle at the edge of upper eyelid; (11) surface around cloaca with large tubercles bearing tiny spines; (12) dorsal skin rough with small granules, a discontinuous “X” or “Y”-shaped ridge with two discontinuous dorsolateral ridges on two side of dorsum; (13) sparse distinct enlarged tubercles on the flanks; (14) single subgular vocal sac in males; (15) presence of villiform black nuptial spines on the dorsal surface of the first and second fingers in adult males; (16) dense tubercles on skin of upper lip, upper eyelid, mandibular articulation, loreal and temporal regions; (17) dense tubercles on skin of ventral surface of thigh, spiny tubercles surrounding the cloaca. Comparisons. Comparative data of Boulenophrys yunkaiensis sp. nov. from B. yaoshanensis sp. nov., B. yingdeensis sp. nov., and the other recognized members of the genus Boulenophrys are listed in Table 3. Having a smaller body size with SVL 35.3–40.0 mm in males, Boulenophrys yunkaiensis sp. nov. is significantly different from congeners whose SVL> 50 mm in males, including B. caudoprocta (81.3 mm in single male), B. jingdongensis (53.0– 56.5 mm in males), B. liboensis (60.5–67.7 mm in males), B. mirabilis (55.8–61.4 mm in males), B. omeimontis (56.0– 59.5 mm in males), B. sangzhiensis (54.7 mm in single male), and B. shuichengensis (102.0– 118.3 mm in males). Having relatively longer shanks with the heels overlapping or meeting when the flexed hindlimbs are held at right angles to the body axis, Boulenophrys yunkaiensis sp. nov. can be easily distinguished from the following nine congeners, B. acuta, B. brachykolos, B. daoji, B. dongguanensis, B. insularis, B. nankunensis, B. obesa, B. ombrophila, and B. wugongensis (vs. all of which have relatively shorter shanks with the heels not meeting). Lacking vomerine teeth, Boulenophrys yunkaiensis sp. nov. differs from B. caudoprocta, B. daiyunensis, B. daweimontis, B. dongguanensis, B. fansipanensis, B. frigida, B. hoanglienensis, B. insularis, B. jingdongensis, B. jinggangensis, B. jiulianensis, B. liboensis, B. nankunensis, B. nanlingensis, B. omeimontis, B. palpebralespinosa, B. qianbeiensis, B. rubrimera, B. sangzhiensis, B. shimentaina, B. tongboensis, and B. yingdeensis sp. nov. (vs. presence of vomerine teeth in these species). Having an unnotched tongue, Boulenophrys yunkaiensis sp. nov. can be distinguished from B. baolongensis, B. binlingensis, B. boettgeri, B. cheni, B. hoanglienensis, B. huangshanensis, B. insularis, B. jingdongensis, B. jiulianensis, B. kuatunensis, B. liboensis, B. lushuiensis, B. minor, B. nanlingensis, B. ombrophila, B. qianbeiensis, B. sangzhiensis, B. sanmingensis, B. shuichengensis B. spinata, and B. tongboensis (vs. tongue notched posteriorly in these species). Lacking lateral fringes on toes, Boulenophrys yunkaiensis sp. nov. differs from B. acuta, B. anlongensis, B. baishanzuensis, B. binchuanensis, B. boettgeri, B. congjiangensis, B. cheni, B. daiyunensis, B. daoji, B. jingdongensis, B. jinggangensis, B. liboensis, B. lini, B. lushuiensis, B. mirabilis, B. mufumontana, B. nanlingensis, B. omeimontis, B. palpebralespinosa, B. qianbeiensis, B. rubrimera, B. sangzhiensis, B. sanmingensis, B. shimentaina, B. shuichengensis, B. spinata, B. xiangnanensis, B. xianjuensis, and B. yangmingensis (vs. presence of lateral fringes on toes in these species); and from B. wushanensis (vs. presence of wide lateral fringes on toes in males while lacking in females). Having rudimentary webbing on toes, Boulenophrys yunkaiensis sp. nov. differs from B. baishanzuensis, B. baolongensis, B. daweimontis, B. fansipanensis, B. frigida, B. huangshanensis, B. kuatunensis, B. lishuiensis, B. ombrophila, B. rubrimera, B. tongboensis, and B. wuliangshanensis (vs. absence of webbing on toes in these species); and from B. jingdongensis, B. palpebralespinosa, B. qianbeiensis, B. shuichengensis, and B. spinata (vs. presence of well-developed webbing on toes in these species). For the remaining eight species, Boulenophrys yunkaiensis sp. nov. can be further distinguished by body size with SVL 45.3–46.1 mm in females (vs. 37.5–39.2 mm in B. angka, 39.5–40.4 mm in B. jiangi, 42.3 mm in B. leishanensis, 37.6 mm in B. shunhuangensis, and 50.5 mm in B. tuberogranulatus), the relative finger lengths II B. angka, IV B. caobangensis), and the presence of tiny spines on skin of upper lip, upper eyelid, loreal and temporal regions excluding the tympanum in adult males (vs. absence of such tiny spines in B. angka, B. chishuiensis, B. jiangi, B. leishanensis, B. shunhuangensis, B. tuberogranulatus, and B. yaoshanensis sp. nov.). Description of holotype. Adult male. small body size, SVL 37.0 mm; head width slightly larger than head length, HWD/HDL 1.15; snout rounded in dorsal view, projecting, sloping backward to mouth in profile, protruding well beyond the margin of lower jaw; top of head flat; eyes moderate in size, ED 0.36 of HDL, pupil vertical, near diamond-shaped; nostril oblique-ovoid; canthus rostralis well developed; loreal region slightly oblique; internasal distance slightly larger than interorbital distance; tympanum boundary clear, TD/ED 1.84; large ovoid choanae at the base of the maxilla; vomerine ridge weak, vomerine teeth absent, maxillary teeth present; margin of tongue rounded, not notched behind; presence of a single subgular vocal sac, a pair of slit-like openings at posterior of jaw. Radio-ulnar length 0.22 of SVL and hand 0.27 of SVL; hand without webbing, fingers without lateral fringes, relative finger length II Skin of doesum rough with sparse granules; sparse large tubercles on the flanks; a small, horn-like prominent tubercle on the edge of upper eyelid; clear supratympanic fold curving posteroventrally from posterior corner of eye to a level above insertion of arm; dense tubercles on skin of upper lip, upper eyelid, mandibular articulation, loreal and temporal regions, excluding the tympanum and surface around cloaca, some of them bearing tiny spines; a discontinuous “X” shaped ridge and a few elongated warts arranged in longitudinal rows on two sides at the middorsum; sparse tubercles on the dorsal shank and thigh; ventral surface smooth; small pectoral gland close to axilla; a single femoral gland positioned subequally distant from knees and cloaca on posterior surface of each thigh. Coloration of holotype in life. Dorsal surface of body yellowish brown with an inverted brown triangular marking between eyes; an “X” shaped marking on the mid-dorsum. Forearms and hindlimbs with dark brown transverse bands. Supratympanic fold with a discontinuous white line; a dark vertical band below the eye, from the inferior margin of the eye to the upper lip. Numerous brown patches scattered on lateroventral surface of flanks; groin red-orange. Ventral surface of throat and chest light salmon in color with brown patches, an indistinct longitudinal stripe on throat; ventral surface of body light salmon in color with brown patches and white spots; ventral surface of limbs light salmon in color with dark brown spots and blotches; ventral surfaces of hands and ventral surfaces of feet brown, tips of digits pale brown; metacarpal tubercle and metatarsal tubercle reddish. Pectoral glands and femoral glands white. Iris yellowish brown. Coloration of holotype in preservative. Yellowish brown faded to greyish brown dorsally. Triangular marking between eyes, “X”-shaped marking on the mid-dorsum, transverse bands on dorsal forearms and hind limbs became indistinct. Color of ventral surface faded to greyish white, all bands and spots became indistinct. Variation and sexual dimorphism. Measurement data of type series are listed in Table 6. Females (SVL 45.3–46.1 mm) are distinctly larger than males (SVL 35.3 –40.0 mm). In adult males, skin of upper lip, upper eyelid, mandibular articulation, loreal and temporal regions excluding the tympanum and surface around cloaca with dense tubercles, some of them bearing tiny spine. Tubercles present in females but without tiny spines. Presence of villiform black nuptial spines on the dorsal surface of the first and second fingers in adult males. Distribution and ecology. Currently, Boulenophrys yunkaiensis sp. nov. is known only from Yunkaishan Nature Reserve, Guangdong, China.All individuals were found in evergreen secondary forest, near montane streams and in the nearby leaf litter on the forest floor at elevations between 900–1400 m. Males call on the rocks by the flowing streams from April to June, suggesting their breeding season corresponds to this period. Females were found on the forest floor, and one female was observed feeding on an earthworm. Tadpoles could be found all year-round.

I. Genus Megophrys Kuhl & Van Hasselt, 1822 Suggested common name: Indonesian horned toads (in English) / fflDzfl (in Chinese) Five species: Megophrys montana Kuhl & Van Hasselt, 1822 (type species); Megophrys acehensis Munir, Nishikawa, Hamidy & Smith, 2021; Megophrys lancip Munir, Hamidy, Farajallah & Smith, 2018; Megophrys parallela Inger & Iskandar, 2005; Megophrys selatanensis Munir, Nishikawa, Hamidy & Smith, 2021., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

VI. Genus Ophryophryne Boulenger, 1903 Suggested common name: Narrow-mouth horned toads (in English) / ���fflDzfl (in Chinese) Seven species: Ophryophryne microstoma Boulenger, 1903 (type species); Ophryophryne elfina Poyarkov, Duong, Orlov, Gogoleva, Vassilieva, Nguyen, Nguyen, Nguyen, Che & Mahony, 2017; Ophryophryne gerti Ohler, 2003; Ophryophryne hansi Ohler, 2003; Ophryophryne pachyproctus Kou, 1985; Ophryophryne poilani Bourret, 1937; Ophryophryne synoria Stuart, Sok & Neang, 2006., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on pages 430-431, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

2) Xenophrys aceras group Two species: Xenophrys aceras Boulenger, 1903; Xenophrys longipes (Boulenger, 1886)., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

1) Xenophrys major group Twelve species: Xenophrys monticola G��nther, 1864 (type species); Xenophrys major (Boulenger, 1908); Xenophrys flavipunctata (Mahony, Kamei, Teeling & Biju, 2018); Xenophrys glandulosa (Fei, Ye & Huang, 1990); Xenophrys himalayana (Mahony, Kamei, Teeling & Biju, 2018); Xenophrys maosonensis (Bourret, 1937); Xenophrys mangshanensis (Fei & Ye, 1990); Xenophrys medogensis (Fei, Ye & Huang, 1983); Xenophrys oreocrypta (Mahony, Kamei, Teeling & Biju, 2018); Xenophrys periosa (Mahony, Kamei, Teeling & Biju, 2018); Xenophrys robusta (Boulenger, 1908); Xenophrys zhangi (Ye & Fei, 1992)., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

2) Brachytarsophrys orientalis group Five species: Brachytarsophrys orientalis Li, Lyu, Wang & Wang, 2020; Brachytarsophrys chuannanensis Fei, Ye & Huang, 2001; Brachytarsophrys feae (Boulenger, 1887); Brachytarsophrys platyparietus Rao & Yang, 1997; Brachytarsophrys popei Zhao, Yang, Chen, Chen & Wang, 2014., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

Specimens of genus Boulenophrys examined in this study. B. acuta (10): China: Guangdong: Fengkai: Heishiding Nature Reserve (type locality): SYS a000168, 0517, 0521, 3257, 2159, 2266���2269, 2276. B. binlingensis (2): China: Sichuan: Hongya: Mt. Wawu (type locality): SYS a005313���5314. B. boettgeri (16): China: Fujian: Wuyishan: Mt. Wuyi (type locality): SYS a002480, 4149���4151; Jiangxi: Guixi: Yangjifeng Nature Reserve: SYS a000312, 0315, 0328���0330, 0376, 0378; Guangfeng: Tongboshan Nature Reserve: SYS a001671��� 1673, 1683, 1700. B. brachykolos (12): China: Hong Kong (type locality): SYS a001502���1503; Guangdong: Shenzhen: Mt. Yangtai: SYS a002051���56; Dapeng Peninsula: SYS a002406���2408, 2410. B. caudoprocta (3): China: Hunan: Sangzhi: Badagongshan Nature Reserve (type locality): SYS a004281, 4308���4309. B. cheni (19): China: Jiangxi: Jinggangshan: Mt. Jinggang (type locality): SYS a001427���1429, 1871���1873; Hunan: Yanling: Taoyuandong Nature Reserve: SYS a002123���2127, 2140���2145. B. daiyunensis (7): China: Fujian: Quanzhou: Daiyun Village (type locality): SYS a001730���1733, 6002; Jiuxianshan: SYS a006000, 6003. B. daoji (6): China: Zhejiang: Taizhou: Mt Tiantai (type locality): SYS a006209���6214 B. huangshanensis (13): China: Anhui: Huangshan: Mt. Huangshan (type locality): SYS a002702���2707; Jiangxi: Wuyuan: Mt. Dazhang: SYS a001622���1623, 3705���3707; Zhejiang: Lin���an: Mt. Tianmu: SYS a002684���2685. B. jingdongensis (24): China: Yunnan: Jingdong: Mt. Wuliang (type locality): SYS a003909, 3928���3929; Zhenyuan: Mt. Ailao: SYS a001778, 2988, 2989���2991, 2993���2994, 3005���3006, 3903���3904; Guangxi: Tianlin: Mt. Cenwanglao: SYS a005160���5165, 5184, 5968���5970. B. jinggangensis (11): China: Jiangxi: Jinggangshan: Mt. Jinggang (type locality): SYS a001413���1416, 1430, 4028; Hunan: Yanling: Taoyuandong Nature Reserve: SYS a001859���1863. B. kuatunensis (3): China: Fujian: Wuyishan: Guadun: SYS a001579, 1590; Jiangxi: Guixi: Yangjifeng Nature Reserve: SYS a000241. B. lini (27): China: Hunan: Yanling: Taoyuandong Nature Reserve (type locality): SYS a002128; Jiangxi: Jinggangshan: Mt. Jinggang: SYS a001417��� 1424, 2375���2386; Suichuan: Nanfengmian Nature Reserve: SYS a002369���2374. B. minor (5): China: Sichuan: Dujiangyan: Mt. Qingcheng (type locality): SYS a003209���3213. B. mirabilis (4): China: Guangxi: Longsheng: Huaping Nature Reserve (type locality): SYS a002192���2193, 2289, 2917. B. obesa (4): China: Guangdong: Fengkai: Heishiding Nature Reserve (type locality): SYS a002270���2272, 3047. B. omeimontis (11): China: Sichuan: Emeishan: Mt. Emei (type locality): SYS a001798���1801, 1940���1941, 5301; Hongya: Mt. Wawu: SYS a005330���5331; Pingshan: Mt. Laojun: SYS a002740���2741. B. sangzhiensis (6): China: Hunan: Sangzhi: Badagongshan Nature Reserve (type locality): SYS a004306���4307, 4313���4316. B. sanmingensis (7): China: Fujian: Sanming: Mt. Longtou (type locality): SYS a002493���2496, 2498���2500. B. shimentaina (11): China: Guangdong: Yingde: Shimentai Nature Reserve (type locality): SYS a002077, 2081���2085, 4172��� 4173, 4710, 5992���5993. B. spinata (2): China: Guizhou: Leishan: Mt. Leigong (type locality): SYS a002226���2227. B. tongboensis (5): China: Jiangxi: Shangrao: Mt. Tongbo (type locality): SYS a001911, 3225���3228. B. tuberogranulatus (1): China: Hunan: Sangzhi: Badagongshan Nature Reserve (type locality): SYS a004310. B. wushanensis (5): China: Hubei: Shennongjia: Shennongjia Nature Reserve (type locality): SYS a003008���3011, 3013. B. wuliangshanensis (5): China: Yunnan: Jingdong: Mt. Wuliang (type locality): SYS a003924���3925; Zhenyuan: Mt. Ailao: SYS a002983���29. B. xiangnanensis (11): China: Hunan: Shuangpai: Mt. Yangming (type locality): SYS a002874���2875, 2878���2886. B. yangmingensis (7): China: Hunan: Shuangpai: Mt. Yangming (type locality): SYS a002877, 2887���2889, 2891���2892., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on pages 431-432, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

VIII. Incertae sedis with Megophryinae Two species: “ Megophrys ” dringi Inger, Stuebing & Tan, 1995; “ Megophrys ” feii Yang, Wang & Wang, 2018.

V. Genus Xenophrys G��nther, 1864 Suggested common name: Strange horned toads (in English) / ���fflDzfl (in Chinese), Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

II. Genus Pelobatrachus Beddard, 1908 Suggested common name: Clay horned toads (in English) / ���fflDzfl (in Chinese) Seven species: Pelobatrachus nasuta (Schlegel, 1858) (type species); Pelobatrachus baluensis (Boulenger, 1899); Pelobatrachus edwardinae (Inger, 1989); Pelobatrachus kalimantanensis (Munir, Hamidy, Matsui, Iskandar, Sidik & Shimada, 2019); Pelobatrachus kobayashii (Malkmus & Matsui, 1997); Pelobatrachus ligayae (Taylor, 1920); Pelobatrachus stejnegeri (Taylor, 1920)., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

IV. Genus Brachytarsophrys Tian & Hu, 1983 Suggested common name: Short-legged toads (in English) / ���DZDzfl (in Chinese), Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

3) Xenophrys lekaguli group Three species: Xenophrys lekaguli (Stuart, Chuaynkern, Chanard & Inger, 2006); Xenophrys parva (Boulenger, 1893); Xenophrys takensis (Mahony, 2011)., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979

5) Xenophrys vegrandis group Three species: Xenophrys vegrandis (Mahony, Teeling & Biju, 2013), Xenophrys yeae (Shi, Zhang, Xie, Jiang, Liu, Ding, Luan & Wang, 2020) comb. nov., Xenophrys zhoui (Shi, Zhang, Xie, Jiang, Liu, Ding, Luan & Wang, 2020) comb. nov.., Published as part of Qi, Shuo, Lyu, Zhi-Tong, Wang, Jian, Mo, Yun-Ming, Zeng, Zhao-Chi, Zeng, Yang-Jin, Dai, Ke-Yuan, Li, Yuan-Qiu, Grismer, L. Lee & Wang, Ying-Yong, 2021, Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China, pp. 401-438 in Zootaxa 5072 (5) on page 430, DOI: 10.11646/zootaxa.5072.5.1, http://zenodo.org/record/5748979