Search

Eumerus colladoi Ricarte & Aguado-Aranda sp. nov. urn:lsid:zoobank.org:act: C63F1D95-7528-4679-9705-FAA919D7CDDD Figs 2–7 Diagnosis Medium sized (6.5–9.5mm) black species (Fig. 2), slightly larger than E. strigatus, which can be distinguished from similar species by the black pilosity on vertical triangle (Fig. 3), the absence of a baso-ventral ridge in the metatibia (Fig. 5B), the shape of sternum IV with a pair of black spinose valves (Fig. 6) and the general morphology of male genitalia (Fig. 7). Etymology This species is named after the Spanish entomologist Juan Gil Collado for his important contribution to the knowledge of the Iberian hoverflies (Gil-Collado 1930). The specific epithet ‘ colladoi ’ refers to his second surname and should be treated as a noun in the genitive case. Material examined Holotype SPAIN • ♂; Alicante, Alcoleja, Sierra de Aitana, Puerto de Tudons, on Thapsia villosa; 18 Jun. 2020; A. Ricarte leg.; CEUA00109561; CEUA-CIBIO. Paratypes PORTUGAL • 1 ♂; Mértola, NE of town; 29S618-416; 22 May 2004; A. v. Eck leg.; coll. A. v. Eck. SPAIN • 1 ♂; Málaga, Yunquera, Parque Natural Sierra de las Nieves, Puerto Saucillo; 36°43′22.9″ N, 04°57′59.8″ W; 1181m a.s.l.; 21 May 2010; M.A. Marcos García leg.; en (‘hovering around’) piornal (‘broom’); CEUA 00107843; CEUA-CIBIO • 1 ♂; Ciudad Real, Parque Nacional de Cabañeros; 2–23 Aug. 2004; A. Ricarte leg.; Malaise trap maA1; CEUA 00109557; CEUA-CIBIO • 1 ♂; Ciudad Real, Parque Nacional de Cabañeros, Finca Garbanzuelo; 27 Jul. 2021; P. Aguado leg.; CEUA 00109777; CEUA-CIBIO • 1 ♂; León, Sena de Luna; 3 Sep. 2010; F. Fresno leg.; N° Cat. 56574; MNCN • 1 ♂; León, Rodiezmo; 27 Aug. 2010; F. Fresno leg.; N° Cat. 57041; MNCN • 1 ♂; Salamanca, Espeja, Campanarios de Azaba; 2 Aug. 2011; García, Ramírez, Moreno leg.; Malaise trap 19; CEUA 00109556; CEUA-CIBIO • 1 ♀; Salamanca. Topas, Castillo del Buen Amor; 11 Jun. 1978; Mª A. Marcos leg.; CEUA 00018013; Mª A. Marcos det. as Eumerus strigatus; CEUA-CIBIO • 1 ♀; Salamanca, Montemayor del Río; 29 Apr. 1980; Mª A. Marcos leg.; CEUA 00018004; Mª A. Marcos det. as Eumerus strigatus; CEUA-CIBIO • 1 ♂; Salamanca, Vallejera de Riofrío, Puerto Vallejera; 1202m a.s.l.; 26 Sep. 1977; S. Fdez. Gayubo leg.; CEUA 00017757; Mª A. Marcos-García det. as Eumerus amoenus; CEUA-CIBIO • 1 ♂; same locality and altitude as for preceding; 24 Aug. 1980; M.A. Marcos leg.; CEUA 00017758; Mª A. Marcos-García det. as Eumerus amoenus; CEUA-CIBIO • 2 ♂♂; Alicante, Agres, Caveta del Voltor; 1200 m a.s.l.; 29 May 2001; Pérez Bañón, Marcos-García & Rojo leg.; CEUA 00109554 to 0010955; CEUA-CIBIO • 5 ♂♂; Alicante, Agres, Foia Ampla; 1060 m.; 29 May 2001; Pérez Bañón, Marcos- García & Rojo leg.; CEUA 00109771 to 00109775; X. Mengual det. as Eumerus amoenus in Mengual (2005); CEUA-CIBIO • 1 ♂; same locality and altitude as for preceding; 17–31 Jul. 2001; Pérez Bañón, Marcos-García and Rojo leg.; Malaise trap; CEUA 00109552; X. Mengual det. as Eumerus amoenus in Mengual (2005); CEUA-CIBIO • 1 ♂; same locality and altitude as for preceding; 23 Apr.–15 May 2002; Pérez Bañón, Marcos-García and Rojo leg.; CEUA 00109553; Malaise trap; X. Mengual det. as Eumerus amoenus in Mengual (2005); CEUA-CIBIO • 1 ♂; Alicante, Agres, Font Retura; 900 m a.s.l.; 30 Aug.- 2 Sep. 2001; Pérez Bañón, Marcos-García and Rojo leg.; Malaise trap; CEUA00109776; X. Mengual det. as Eumerus amoenus in Mengual (2005); CEUA-CIBIO • 1 ♂; same locality as for holotype; 6 Jun. 2020; A. Ricarte leg.; CEUA_S01; CEUA 00107848; CEUA-CIBIO • 1 ♀; same locality as for holotype; 6 Jun. 2020; A. Ricarte; CEUA 00109558; CEUA-CIBIO • 1 ♀; same locality as for holotype; 29 May 2020; A. Ricarte leg.; CEUA 00109559; CEUA-CIBIO • 3 ♀♀; same collection data as for holotype; CEUA 00109560; CEUA 00109562 to 00109563; CEUA-CIBIO • 3 ♀♀; same locality as for holotype; 20 Jun. 2020; A. Ricarte leg.; CEUA_S02, CEUA 00107847; CEUA 00109564 to 00109565; CEUA-CIBIO; Coll. A. v. Eck • 3 ♀♀; same locality as for holotype; 20 Jun. 2020; Z. Nedeljković leg.; CEUA 00109566 to 00109568; CEUA-CIBIO • 1 ♀; same locality as for holotype; 10 Sep. 2020; Z. Nedeljković leg.; CEUA 00109569; CEUA-CIBIO • 2 ♀♀; same locality as for holotype; 16 May 2021; Z. Nedeljković leg.; CEUA_S52, CEUA 00107928; CEUA 00109570; CEUA-CIBIO • 2 ♂♂; same locality as for holotype; 26 May 2021; Z. Nedeljković leg.; CEUA 00108052 to 00108053; CEUA-CIBIO; CSCA • 5 ♀♀; same locality as for holotype; 26 May 2021; Z. Nedeljković leg.; CEUA 00108050 to 00108051; CEUA 00108067 to 00108069; CEUA-CIBIO; CSCA • 1 ♀; same locality as for holotype; 11 Jun. 2021; Z. Nedeljković leg.; CEUA 00108054; CEUA-CIBIO • 1 ♀; Alicante, Alcoleja, Sierra de Aitana, Mas del Piscul; 29 May 2019; M.A. Marcos García leg.; CEUA 00109854; CEUA- CIBIO • 1 ♀; same locality and date as for preceding; A. Ricarte leg.; CEUA 00109855; CEUA-CIBIO • 1 ♀; same locality as for preceding; 20 Jun. 2016; G.J. Souba Dols leg.; INV09526; CEUA 00111027; Ricarte det. as Eumerus sp.; CEUA-CIBIO • 6 ♀♀; same locality as for preceding; 26 May 2021; I. Ballester leg.; CEUA 00109860 to 00109865; CEUA-CIBIO • 1 ♂; Alicante, Benimaurell; 7 May 2007; Marcos-García & Ricarte leg.; CEUA 00029572; CEUA-CIBIO • 1 ♂; Alicante, la Vall d’Ebo; 38°48′16.60″ N, 0°11′59.57″ W; 555 m a.s.l.; 26 May 2021; P. Aguado leg.; CEUA 00108079; CEUA- CIBIO • 1 ♀; same collection data as for preceding; A. Ricarte leg.; CEUA 00108075; CEUA-CIBIO • 1 ♀; same locality as for preceding; 38°48′16.47″ N, 0°11′42.97″ O; 546m a.s.l.; 20 May 2021; A. Ricarte leg.; on Thapsia villosa; CEUA 00109853; CEUA-CIBIO • 3 ♀♀; Alicante, la Vall de Laguar, Venta del Collao; 38°46′10.91″ N, 0°9′4.21″ W; 762 m a.s.l.; 26 May 2021; A. Ricarte leg; CEUA 00108059; CEUA 00108073 to 00108074; CEUA-CIBIO • 1 ♂; Alicante, Cocentaina, Font de la Penya Banyà; 4 Jul. 2021; I. Ballester leg.; en (‘on’) Rubus sp.; CEUA 00108072; CEUA-CIBIO • 1 ♂; Valencia, Bocairent, Mas del Parral; 900 m a.s.l.; 5–19 Jun. 2001; Malaise trap; Pérez Bañón, Marcos-García and Rojo leg.; CEUA 00109851; X. Mengual det. as Eumerus amoenus in Mengual (2005); CEUA-CIBIO • 1 ♂; same locality and altitude as for preceding; 30 Jul.–13 Aug. 2002; Malaise trap; Pérez Bañón, Marcos-García and Rojo leg.; CEUA 00109852; X. Mengual det. as Eumerus amoenus in Mengual (2005); CEUA-CIBIO • 1 ♀; Valencia, Bocairent, Sierra de Mariola, Pla d’Aparici, on Thapsia villosa; 3 Jun. 2020; A. Ricarte leg.; CEUA_S06; CEUA 00107844; CEUA-CIBIO • 1 ♀, Valencia, Sª Mariola, Bocairent, Font del Mas dels Arbres, on Thapsia villosa; 3 Jun. 2020; A. Ricarte leg.; CEUA 00109850; CEUA-CIBIO • 2 ♀♀; Valencia, Alt del Portell, Bocairent, Vall Albaida; 837 m a.s.l.; 12 Jun. 2021; Z. Nedeljković leg.; CEUA 00108056 to 00108057; CEUA-CIBIO • 1 ♂; same collection data as for preceding; CEUA 00108058; CEUA-CIBIO • 1 ♂; same locality, altitude and date as for preceding; A. Ricarte leg.; CEUA 00108055; CEUA-CIBIO • 1 ♂; Valencia, Sª Mariola, Bocairent, Font del Mas dels Arbres, campo de Thapsia villosa; 12 Jun. 2021; Z. Nedeljković leg.; CEUA 00108078; CEUA-CIBIO • 2 ♀♀; same collection data as for preceding; CEUA 00108077 to 00108076; CEUA-CIBIO • 1 ♂; same locality and date as for preceding; A. Ricarte leg.; CEUA 00108071; CEUA-CIBIO • 1 ♂; Sierra Espuña; 37.866° N, 1.447° W; 732 m a.s.l.; 7 Oct. 2020; A. Ricarte, Z. Nedeljković and P. Aguado leg.; CEUA_S03; CEUA 00107846; CEUA-CIBIO • 1 ♀; same collection data as for preceding; CEUA_S04; CEUA 00107842; CEUA-CIBIO • 1 ♂; El Escorial; 14 Jun. 1922; Dusmet leg.; • 1 ♂; El Pardo; Jun. 1908; Arias leg.; MNCN • 1 ♂; Villaviciosa de Odón, Villaviciosa; 14 Apr. 1926; Dusmet leg.; J. Gil Collado det. as Eumerus amoenus in Gil-Collado (1930); MNNC. Additional examined material of other species of Eumerus Eumerus amoenus Loew, 1848 SERBIA • 1 ♂; Dubašnica, Klisura Lazareve Reke; 21 Aug. 1997; Vujić leg.; CEUA 00017760; A. Vujić det.; CEUA-CIBIO • 1 ♀; Malinik, Malinik; 26 Jul. 1997; Dević leg; CEUA 00017763; A. Vujić det.; CEUA-CIBIO. SPAIN • 1 ♂; Ciudad Real, Parque Nacional de Cabañeros; 1–24 Aug. 2004; A. Ricarte leg.; Malaise trap maJ1; CEUA 00084834; A. Ricarte det.; CEUA-CIBIO. TUNISIA • 1 ♂; Djerba, Houmt Souk, 5km S of Midoun; M. Hauser leg.; CEUA 00017759; M. Hauser det.; CEUA-CIBIO. Eumerus chrysopygus Sack, 1941 SOUTH KOREA • 1 ♂; Chungnam Keumsan Nami-myeon Pohyeonsa Pohyeonsa; 36°03.494 N, 127°27.225 E; 1–8 Jun. 2005; P. Tripton leg.; Malaise; CEUA 00103565; M. Hauser det.; CEUA-CIBIO. Eumerus clavatus Becker, 1921 GERMANY • 1 ♂; D-BW Pforzheim Niefern MV91 Lattenold MF; 26 Jun. 1992; Schmid-Egger leg.; D. Doczkal det.; CSCA • 1 ♀; same locality and collector as for preceding; 10 Sep. 1992; D. Doczkal det.; CSCA. Eumerus nudus Loew, 1848 SPAIN • 1 ♂; Alicante, Tibi, Sª del Maigmó, observatorio; 1176 m a.s.l.; 26 May 2021; A. Ricarte leg.; CEUA_S64; CEUA 00111026; Z. Nedeljković det.; CEUA-CIBIO. Description Male MEASUREMENTS (mm). Holotype (♂): l = 8.02; w = 2.05; h = 1.91; hw = 2.54; wl = 5.82. HEAD. Eye contiguity virtually of the same length (0.31 mm) as the height of the frontal triangle (0.33 mm). Eye pilosity consisting of rather short, scarce and pale pile, specially conspicuous on the ventral part of eye. Face and frontal triangle (including lunules) densely and white pollinose, covered with white pile (Fig. 3A). Vertical triangle black and with long, erect, black pile (Fig. 3C), pollinose at its anterior apex and with two extremely reduced pollinose spots which are located behind posterior ocelli. Ocellar triangle isosceles. Occiput black and purple iridescent (under artificial white lightning), with pile shorter than those on the vertical triangle; occiput white pollinose along the eye margin. Scape and pedicel dark brown; pedicel with black pile which are longer on the ventral side. Basoflagellomere almost square (ratio length:width = 1:1.1), convex dorsally and slightly concave ventrally, dark brown except baso-ventrally, with a lunulate black fossette at the distal margin. Pedicel and basoflagellomere sparsely white pollinose (pollinosity more obvious in dorsal view and under artificial white lighting). Arista black (Fig. 3E). THORAX. Mesonotum and lateral sides of the thorax black. Anterior and posterior parts of the scutum covered with rather short and white pile, in the medial part white pile intermixed with short and semireclined black pile; scutum with two medial white pollinose vittae extending over the anterior two thirds of the scutum and with a small circular grey pollinose spot posterior to transverse suture; lateral margins of the scutum with green metallic reflections. Notopleural sulcus absent. Disc of scutellum with white pile slightly longer than those on the scutum; posterior margin of scutellum with teeth-like protuberances, each bearing a long black pilis apically. Posterior anepisternum, anterior anepimeron and katepisternum on its postero-lateral area densely long white pilose. Katatergum with discrete bunch of yellow setae. Pleuron grey pollinose except the posterior anepisternum postero-laterally, anterior anepimeron centrally, ventral margin of posterior anepimeron and the dorsal margin of katepisternum. Femora black, basis and apices light brown. Basal half of pro- and mesotibia light brown and apical half black. Basal third of metatibia light brown and apical two thirds black. Dorsal surface of all tarsi black, ventrally yellowish brown. Posterior side of pro- and mesofemora white pilose. Metafemur densely white pilose, ventrally with pile of different lengths including some which are nearly twice as long as the reminder. Metafemur with an anterior row of 6 spinae and a posterior row of 14 spinae curved towards the posterior side of the metafemur (Fig. 5A). Metatibia flattened, without a baso-ventral ridge (Fig. 5B). Wing membrane extensively microtrichose; posterior margin of the wing with dense, short and brown pile; margin of ventral calypter with rather long and yellow pile; margin of dorsal calypter with pile shorter than those of ventral calypter; halter yellow. ABDOMEN. Terga I–IV black. Lateral margin of tergum I grey pollinose and white pilose. Terga II–IV densely white pilose on their lateral margins (pile on anterior corners of tergum II rather long); terga II– IV each with a pair of slightly curved, white pollinose maculae, terga covered with rather short, reclined and white pile. Anterior margins of terga III–IV black pilose. Posterior margin of tergum IV light brown with a marginal line of black pile along it. Sternum IV with two quadrangular, straight projections on the posterior margin; central area of the sternum IV black pilose and protruding in a rounded, nose-like expansion; sternum IV with two black spinose valves under the medial area (Fig. 6). MALE GENITALIA. Epandrium with a simple posterior surstylar lobe convex anteriorly and straight posteriorly (Fig. 7B). Cercus with long pile. Anterior surstylar lobe with two large hyaline lobes and a small third accessory lobe in between (Fig. 7B–C). Hypandrium with a prominence at the base which shows a small indentation (Fig. 7A). Female Same as the male, besides the sexual dimorphism, except for the following characters: eye pilosity more abundant; basoflagellomere 1.0-1.2 × wider than long and larger than in male (Fig. 3D, F); black macula at the distal margin of the basoflagellomere more noticeable (Fig. 3F); white pollinose spots behind the posterior ocelli larger than in male; frons white pollinose along eye margins but grey on its central area (Fig. 3B). Variation Based on the examined material, E. colladoi Ricarte & Aguado-Aranda sp. nov. shows intraspecific variation in body length (♂ = 6.43–8.91 mm, n = 21; ♀ = 7.69–9.58mm, n = 15), length of frontal triangle in males (0.28–0.38 mm), length of eye contiguity in males (0.24–0.41mm), the presence or absence of two reduced white pollinose spots behind posterior ocelli (in males), the inclination of the spinae on the ventral side of the metafemur (from almost not inclined at all to an inclination of 90°) and the extent of the coloration on the metatibia (from light brown to black). Distribution The new species has been recorded from different Spanish and Portuguese localities, all in the Mediterranean region of the Iberian Peninsula (Fig. 1). Biology The flight period of E. colladoi Ricarte & Aguado-Aranda sp. nov. runs from late May to October. Adults can be found in a wide altitudinal range (80–1200 m) mainly associated with mountain areas. The predominant vegetation in these areas are woodlands of pine (Pinus L.) or oak (Quercus L.) where some adults have been sampled feeding on inflorescences of Thapsia villosa L. (Apiaceae). Systematic position of E. colladoi Ricarte & Aguado-Aranda sp. nov. based on DNA evidence Both the NJ and ML trees had virtually identical topologies. In the ML-tree (Fig. 8), high node support values (> 70) were obtained for every species group but for the E. pulchellus group (see Introduction). All barcodes of E. colladoi Ricarte & Aguado-Aranda sp. nov. were similar to each other with just 1 base pair difference. Eumerus nudus is similar to E. colladoi sp. nov. in the general shape of the basoflagellomere, the pilosity of the scutum, the coloration of the posterior margin of tergum IV and shape of male genitalia (see Discussion). The morphological similarities between E. nudus and E. colladoi sp. nov. justified the incorporation of the E. nudus sequence in the molecular analyses. A COI- 5’ sequence of the E. nudus was generated as it had no representation in the consulted BOLD and GenBank databases. The barcode sequence of E. nudus grouped, as expected, with that of Eumerus alpinus Rondani, 1857 but they did not form a clade with E. colladoi sp. nov. In addition, the new species did not cluster with sequences of species of the E. clavatus group, which have even more similar male genitalia in comparison with E. colladoi sp. nov. together with a medium-sized body, a dark body coloration and an elongated and parallel-sided abdomen. The proportion of invariant sites for the first and third codon positions were 0.29 and 0.07, respectively. Level of variation is usually higher in the third codon position as it tends to have faster rate of evolution (Felsenstein 1978)., Published as part of Aguado-Aranda, Pablo, Ricarte, Antonio, Nedeljković, Zorica & Marcos-García, M. Ángeles, 2022, An overlooked case for a century: taxonomy and systematics of a new Iberian species of Eumerus Meigen, 1822 (Diptera, Syrphidae), pp. 35-57 in European Journal of Taxonomy 817 on pages 41-50, DOI: 10.5852/ejt.2022.817.1761, http://zenodo.org/record/6518247, {"references":["Gil-Collado J. 1930. Monografia de los sirfidos de Espana. Trabajos del Museo Nacional de Ciencias Naturales (Serie Zoologica) 54: 1 - 376.","Mengual X. 2005. Diversidad biologica de los sirfidos (Diptera: Syrphidae) asociada a la actividad humana en areas protegidas de la provincia de Alicante. Master Thesis, University of Alicante, Spain.","Becker T. 1921. Neue Dipteren meiner Sammlung. Mitteilungen aus dem Zoologischen Museum in Berlin 10 (1): 1 - 93.","Felsenstein J. 1978. Cases in which parsimony or compatibility methods will be positively misleading. Systematic Biology 27 (4): 401 - 410. https: // doi. org / 10.1093 / sysbio / 27.4.401"]}

Eumerus sulcitibius Rondani, 1868 (New to Madrid) Material examined: 1 female; 1- 6.6.1990. Guild: Phytophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Rondani C. 1868. Diptera Italica non vel minus cognita descripta vel annotata observationibus nonnullis additis. Fasc. III. Atti della Societa italiana di scienze naturali e del Museo civico di storia naturale di Milano 11: 21 - 54."]}

Paragus bicolor (Fabricius, 1794) Material examined: 1 female; 28.7- 4.8.1989. – 1 female; 1- 6.6.1990. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Fabricius J. C. 1794. Entomologia systematica emendata et aucta. C. G. Proft, Hafniae [= Copenhagen], Vol. 4, [6] + 472 + [5] pp."]}

Pelecocera caledonica (Collin, 1940) (New to Madrid) Material examined: 1 female; 21- 28.9.1989. – 1 male; 1- 6.6.1990. Guild: Phytophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Collin J. E. 1940. Notes on Syrphidae (Diptera). IV. The Entomologist's monthly magazine 76: 150 - 158."]}

Episyrphus balteatus (De Geer, 1776) Material examined: 1 male and 1 female; 7- 14.7.1989. – 19 males and 9 females, 14- 21.7.1989. – 1 male and 5 females; 20- 28.7.1989. – 5 females (1 female discarded); 28.7- 4.8.1989. – 2 females; 11- 18.8.1989. – 1 female (discarded); 25- 31.8.1989. – 1 male; 11- 20.4.1990. – 2 males and 2 females; 15- 22.5.1990. – 8 males and 1 female; 23- 31.5.1990. – 1 male; 1- 6.6.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on pages 397-398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Geer C. de 1776. Memoires pour servir a l'histoire des insectes. P. Hesselberg, Stockholm. Vol. 6, viii + 523 pp., 30 pls."]}

Chrysotoxum sp. near vernale Loew, 1841 (New to Madrid) Fig. 6C Material examined: 1 male, 9- 16.6.1989. – 1 female; 22- 30.6.1989. – 1 female; 14- 21.7.1989. – 1 female; 22- 30.3.1990. – 1 female; 11- 20.4.1990. – 1 male and 2 females; 15- 22.5.1990. – 2 females; 23- 31.5.1990. – 1 male and 3 females; 1- 6.6.1990. Guild: Zoophagous (larva undescribed). Remarks: This species is currently being described by Nedeljković et al. (in press)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Loew H. 1841. Ueber die Gattung Chrysotoxum. Stettiner Entomologische Zeitung 2: 136 - 141, 155 - 160."]}

Meliscaeva auricollis (Meigen, 1822) Material examined: 1 male; 22- 30.6.1989. – 12 males; 7- 14.7.1989. – 65 males and 13 females; 14- 21.7.1989. – 1 male; 20- 28.7.1989. – 3 males; 28.7– 4.8.1989. – 1 female; 1- 7.9.1989. – 1 male and 1 female; 21- 28.9.1989. – 1 male; 23- 31.5.1990. – 1 male; 1- 6.6.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Meigen J. W. 1822. Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Dritter Theil. Schulz- Wundermann, Hamm. x + 416 pp., pls 22 - 32."]}

Cheilosia proxima (Zetterstedt, 1843) (New to Madrid) Material examined: 1 female; 1- 6.6.1990. Guild: Phytophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912

Cheilosia gigantea (Zetterstedt, 1838) (New to Madrid) Material examined: 1 female; 1- 7.5.1990. – 1 female; 15- 22.5.1990. – 1 female; 1- 6.6.1990. Guild: Phytophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Zetterstedt J. W. 1838. Dipterologis Scandinaviae. Sect. 3: Diptera, pp. 477 - 868. Insecta Lapponica, Lipsiae [= Leipzig], vi + 1, 140 pp."]}

Dasysyrphus pauxillus (Williston, 1887) (New to Madrid) Material examined: 1 male; 9- 16.6.1989. – 4 males and 1 female; 23- 31.5.1990. – 2 males; 1- 6.6.1990. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Williston S. W. 1887. Synopsis of the NorthAmerican Syrphidae. Bulletin of the American Museum of Natural History (1886) 31, xxx + 335 pp, 12 pls."]}

Callicera rufa Schummel, 1842 Material examined: 1 female; 1- 7.9.1989. Guild: Saproxylic., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Schummel T. E. 1842. Verzeichniss und Beschreibung der bis jetzt in Schlesien gefangenen Zweiflugler der Syrphen Familie. Uebersicht der Arbeiten und Veranderungen der Schlesischen Gesellschaft fiir Vaterlandische Kultur: 15 - 22."]}

Eupeodes corollae (Fabricius, 1794) Material examined: 2 males and 16 females; 9- 16.6.1989. – 7 females; 22- 30.6.1989. – 2 males and 3 females (2 males and 2 females discarded); 31.6- 6.7.1989. – 65 males and 87 females; 14- 21.7.1989. – 5 males and 26 females; 20- 28.7.1989. – 1 male and 54 females; 7- 14.8.1989. – 12 males and 22 females (4 males and 10 females discarded); 28.7- 4.8.1989. – 1 male and 16 females; 11- 18.8.1989. – 3 females (discarded); 25- 31.8.1989. – 1 female; 1- 7.9.1989. – 1 female (discarded); 11- 25.10.1989. – 1 female; 21- 28.9.1989. – 1 female (discarded); 2- 9.11.1989. – 2 females (discarded); 14- 21.11.1989. – 1 male and 1 female; 11- 20.4.1990. – 1 male and 10 females; 15- 22.5.1990. – 1 male and 7 females; 23- 31.5.1990. – 1 male and 1 female; 1- 6.6.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Fabricius J. C. 1794. Entomologia systematica emendata et aucta. C. G. Proft, Hafniae [= Copenhagen], Vol. 4, [6] + 472 + [5] pp."]}

Merodon unicolor Strobl in Czerny & Strobl, 1909 Material examined: 4 males; 23- 31.5.1990. Guild: Phytophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Czerny L., Strobl P. G. 1909. Spanische Dipteren. III. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 59: 121 - 301."]}

Didea intermedia Loew, 1854 (New to Madrid) Material examined: 1 female (discarded); 25- 31.8.1989. – 1 female; 1- 7.9.1989. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Loew H. 1854. Neue Beitrage zur Kenntniss der Dipteren. Programm der Koniglichen Realschule zu Meseritz 1854: 1 - 24."]}

Parasyrphus punctulatus (Verrall, 1873) (New to Madrid) Material examined: 1 male; 7- 15.5.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Verrall G. H. 1873. Additions and correction to the list of British Syrphidae. Entomologist's Monthly Magazine 9: 251 - 256."]}

Chrysotoxum volaticum Séguy, 1961 Material examined: 3 males; 22- 30.6.1989. – 3 males; 31.6– 6.7.1989. – 3 males; 7- 14.7.1989. – 3 males; 14- 21.7.1989. – 1 female; 11- 18.8.1989. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Seguy E. 1961. Dipteres syrphides d'Europe Occidentale. Memoires du Museum national d'Histoire naturelle Serie (A) 23, 248 pp."]}

Volucella elegans Loew, 1862 Material examined: 1 male; 7- 14.7.1989. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Loew H. 1862. Ueber einige bei Varna gefangene Dipteren. Wiener Entomologische Monatsschrift 6: 161 - 175."]}

Chrysotoxum cisalpinum Rondani, 1845 Material examined: 1 male; 7- 14.7.1989. – 2 males; 11- 18.8.1989. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Rondani C. 1845. Species Italicae generis Chrysotoxi insectis Dipteris, observatae et distinctae. Fragmentum decimum ad inserviendum dipterologiae Italicae. Annales de la Societe entomologique de France (2) 3: 193 - 203."]}

Eumerus sabulonum (Fallén, 1817) Material examined: 1 male and 1 female; 9- 16.6.1989. Guild: Phytophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912

Brachypalpus laphriformis (Fallén, 1816) (New to Madrid) Material examined: 1 male; 7- 14.7.1989. Guild: Saproxylic., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Fallen C. F. 1816 - 1817. Syrphici Sveciae. Berlingianis, Lundae [= Lund], 1 - 14 [1816.06.08], 15 - 22 [1816.06.08], 23 - 30 [1817.05.10], 31 - 42 [1817.05.20], 43 - 54 [1817.05.21], 55 - 62 [1817.05.22] pp."]}

Dasysyrphus albostriatus (Fallén, 1817) Fig. 6D Material examined: 5 females; 9- 16.6.1989. – 1 male; 22- 30.6.1989. – 2 males and 1 female; 14- 21.7.1989; 2 males and 2 females (all discarded); 25- 31.8.1989. – 4 males and 2 females; 1- 7.9.1989. – 1 female (discarded); 14- 21.11.1989. – 1 male and 1 female; 21- 28.9.1989. – 1 male; 23- 31.5.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912

Eristalis arbustorum (Linnaeus, 1758) Material examined: 1 male; 31.6- 6.7.1989. – 1 male and 1 female; 7- 14.7.1989. – 1 female; 14- 21.7.1989. – 1 female; 28.7- 4.8.1989. Guild: Saprophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Linnaeus C. 1758. Systema naturae. Ed. 10, Vol. 1. 824 pp. L. Salvii, Holmiae [= Stockholm]."]}

Paragus pecchiolii Rondani, 1857 (New to Madrid) Material examined: 1 male; 14- 21.7.1989. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Rondani C. 1857. Dipterologiae italicae prodromus. Vol: II. Species italicae ordinis dipterorum in genera characteribus definita, ordinatim collectae, methodo analitica distinctae, et novis vel minus cognitis descriptis. Pars prima. Oestridae: Syrpfhidae: Conopidae. A. Stocchi, Parmae [= Parma], 264 pp."]}

Chrysotoxum octomaculatum Curtis, 1837 Material examined: 1 male; 22- 30.6.1989. – 2 males; 31.6– 6.7.1989. – 6 males; 7- 14.7.1989. – 6 males; 14- 21.7.1989. – 2 females; 28.7- 4.8.1989. – 1 male; 15- 22.5.1990. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Curtis J. 1837. British entomology. Vol. 14, pls 650 - 653. Privately published, London."]}

Syritta pipiens (Linnaeus, 1758) Material examined: 1 female; 28.7- 4.8.1989. Guild: Saprophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Linnaeus C. 1758. Systema naturae. Ed. 10, Vol. 1. 824 pp. L. Salvii, Holmiae [= Stockholm]."]}

Sphaerophoria scripta (Linnaeus, 1758) Material examined: 6 males and 9 females; 9- 16.6.1989. – 11 males and 25 females; 22- 30.6.1989. – 21 males and 39 females (discarded); 31.6- 6.7.1989. – 50 males; 7- 14.7.1989. – 156 males and 203 females; 14- 21.7.1989. – 112 males and 165 females; 20- 28.7.1989. – 70 males and 130 females (39 males and 66 females discarded); 28.7– 4.8.1989. – 51 males; 7- 14.8.1989. – 1 male and 3 females; 11- 18.8.1989. – 1 female; 14- 22.3.1990. – 2 females; 7- 15.5.1990. – 3 males and 23 females; 15- 22.5.1990. – 18 males and 35 females; 23- 31.5.1990. – 9 males and 6 females; 1- 6.6.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Linnaeus C. 1758. Systema naturae. Ed. 10, Vol. 1. 824 pp. L. Salvii, Holmiae [= Stockholm]."]}

Syrphus ribesii (Linnaeus, 1758) Material examined: 1 female; 9- 16.6.1989. – 2 females; 22- 30.6.1989. – 1 female; 31.6- 6.7.1989. – 1 male and 1 female; 7- 14.7.1989. – 1 male and 1 female; 14- 21.7.1989. – 1 male; 20- 28.7.1989. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Linnaeus C. 1758. Systema naturae. Ed. 10, Vol. 1. 824 pp. L. Salvii, Holmiae [= Stockholm]."]}

Neoascia podagrica (Fabricius, 1775) Material examined: 2 females; 15- 22.5.1990. Guild: Saprophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Fabricius J. C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Kortii, Flensbvrgi et Lipsiae [= Flensburg & Leipzig]. [32] + 832 pp."]}

Dasysyrphus tricinctus (Fallén, 1817) (New to Madrid) Material examined: 1 male; 1- 6.6.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912

Chrysotoxum intermedium Meigen, 1822 Material examined: 1 male; 9- 16.6.1989. – 2 males; 7- 14.7.1989. – 2 females; 15- 22.5.1990. – 1 male; 1- 6.6.1990. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Meigen J. W. 1822. Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Dritter Theil. Schulz- Wundermann, Hamm. x + 416 pp., pls 22 - 32."]}

Eristalis tenax (Linnaeus, 1758) Material examined: 1 male; 7- 14.7.1989. – 1 male; 14- 21.7.1989. – 1 male (discarded); 14- 21.11.1989. Guild: Saprophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Linnaeus C. 1758. Systema naturae. Ed. 10, Vol. 1. 824 pp. L. Salvii, Holmiae [= Stockholm]."]}

Paragus strigatus Meigen, 1822 Material examined: 1 female; 9- 16.6.1989. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Meigen J. W. 1822. Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Dritter Theil. Schulz- Wundermann, Hamm. x + 416 pp., pls 22 - 32."]}

Chrysotoxum festivum (Linnaeus, 1758) Material examined: 1 male; 7- 14.7.1989. – 1 male; 14- 21.7.1989. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Linnaeus C. 1758. Systema naturae. Ed. 10, Vol. 1. 824 pp. L. Salvii, Holmiae [= Stockholm]."]}

Syrphus torvus Osten Sacken, 1875 Material examined: 2 females; 9- 16.6.1989. – 1 male and 2 females; 14- 21.7.1989. – 1 male; 20- 28.7.1989. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Osten Sacken C. R. 1875. On the North American species of the genus Syrphus (in the narrowest sense). Proceedings of the Boston Society of Natural History (1875 - 1876) 18: 135 - 153."]}

Xanthogramma dives (Rondani, 1857) (New to Madrid) Material examined: 1 female; 11- 18.8.1989. – 1 female; 21- 28.9.1989. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Rondani C. 1857. Dipterologiae italicae prodromus. Vol: II. Species italicae ordinis dipterorum in genera characteribus definita, ordinatim collectae, methodo analitica distinctae, et novis vel minus cognitis descriptis. Pars prima. Oestridae: Syrpfhidae: Conopidae. A. Stocchi, Parmae [= Parma], 264 pp."]}

Merodon ibericus Vujić in Popović et al., 2015 (New to Madrid) Material examined: 1 male and 2 females; 7- 14.7.1989. – 3 males; 14- 21.7.1989. Guild: Phytophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Popovic D., Acanski J., Djan M., Obreht D., Vujic A., Radenkovic S. 2015. Sibling species delimitation and nomenclature of the Merodon avidus complex (Diptera: Syrphidae). European Journal of Entomology 112 (4): 790 - 809."]}

Cheilosia laticornis Rondani, 1857 Material examined: 1 male; 11- 20.4.1990. – 1 female; 1-6.vi.1990. Guild: Phytophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Rondani C. 1857. Dipterologiae italicae prodromus. Vol: II. Species italicae ordinis dipterorum in genera characteribus definita, ordinatim collectae, methodo analitica distinctae, et novis vel minus cognitis descriptis. Pars prima. Oestridae: Syrpfhidae: Conopidae. A. Stocchi, Parmae [= Parma], 264 pp."]}

Xanthandrus comtus (Harris, 1780) Material examined: 1 male; 14- 21.7.1989. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 399, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912

Eupeodes latifasciatus (Macquart, 1829) (New to Madrid) Material examined: 3 females; 9- 16.6.1989. – 2 males and 1 female; 7- 14.7.1989. – 3 females; 21- 28.9.1989. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Macquart P. J. M. 1829. Insectes dipteres du nord de la France. Syrphies. Lille. \" 1827 \" 223 pp., 4 pls."]}

Callicera macquarti Rondani, 1844 (New to Madrid) Material examined: 1 female; 1- 7.5.1990. Guild: Saproxylic (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Rondani C. 1844. Species Italicae Generis Callicerae ex insectis Dipteris, distinctae et descriptae; Fragmentum octavum ad inserviendum Dipterologiae Italicae. Annales de la Societe entomologique de France (2) 2: 61 - 68."]}

Melanostoma scalare (Fabricius, 1794) Material examined: 1 female; 9- 16.6.1989. – 4 females; 22- 30.6.1989. – 5 males and 10 females; 14- 21.7.1989. – 1 male and 4 females; 20- 28.7.1989. – 1 male and 2 females; 28.7- 4.8.1989. – 2 males and 12 females; 7- 14.8.1989. – 1 male and 1 female; 11- 18.8.1989. – 1 female; 1- 7.9.1989. – 1 male and 1 female; 15- 22.5.1990. – 7 males and 10 females; 23- 31.5.1990. – 1 female; 1- 6.6.1990. Guild: Zoophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Fabricius J. C. 1794. Entomologia systematica emendata et aucta. C. G. Proft, Hafniae [= Copenhagen], Vol. 4, [6] + 472 + [5] pp."]}

Eumerus consimilis Šimić & Vujić, 1996 (New to Madrid) Material examined: 1 male and 1 female; 1- 7.9.1989. Guild: Phytophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Simic S., Vujic A. 1996. A new species of the genus Eumerus Meigen, 1822 (Diptera: Syrphidae. Acta Entomologica serbica 1: 1 - 14."]}

Chrysotoxum latifasciatum Becker, 1921 Material examined: 1 male; 9- 16.6.1989. – 1 male; 31.6- 6.7.1989. – 2 males; 7- 14.7.1989. – 1 male; 14- 21.7.1989. – 1 female; 28.7- 4.8.1989. – 1 female; 1- 6.6.1990. Guild: Zoophagous (larva undescribed)., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 397, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Becker T. 1921. Neue Dipteren meiner Sammlung. Mitteilungen aus dem Zoologischen Museum in Berlin 10: 1 - 93."]}

Eristalis horticola (De Geer, 1776) Material examined: 1 female; 7- 14.7.1989. – 2 males; 14- 21.7.1989. Guild: Saprophagous., Published as part of Lorenzo, Daniel, Ricarte, Antonio, Nedeljković, Zorica, Nieves-Aldrey, José Luis & Marcos-García, Maria Ángeles, 2020, Hoverflies (Diptera: Syrphidae) of El Ventorrillo Biological Station, Madrid province, Spain: a perspective from a late twentieth century inventory, pp. 393-412 in Revue suisse de Zoologie 127 (2) on page 398, DOI: 10.35929/RSZ.0029, http://zenodo.org/record/5743912, {"references":["Geer C. de 1776. Memoires pour servir a l'histoire des insectes. P. Hesselberg, Stockholm. Vol. 6, viii + 523 pp., 30 pls."]}

Remarks on Eumerus lyneborgi sp. nov. and similar species Eumerus lyneborgi sp. nov. is similar to E. vestitus Bezzi, 1912 in body size and constitution, predominantly pollinose frons, with punctured pollinosity (females), swollen metafemur, with two ventral rows of short black spinae, one antero-apically and other postero-apical, lateral margins of terga III and IV pollinose, and tergum IV widely pollinose posteriorly. Bezzi (1912) described E. vestitus based on males and females from ��� Guinea Portoguese��� (nowadays, Guinea-Bissau), supposedly the male and three females the authors of the present paper found in the MCSNG collection. These specimens are all labelled as ���syntypus��� and a female has an additional label of ���Typus���. Bezzi (1912) did not mention a holotype or type specimen for his new species in the description. Thus, according to articles 73.1.1 and 73.1.2 of the International Commission on Zoological Nomenclature (1999), this ���Typus��� is not a valid holotype and therefore all specimens are syntypes. In addition, no subsequent type designation for E. vestitus is known to the authors of the present paper. In the past, curators sometimes labelled arbitrarily as ���Typus��� the best looking specimen within the type series (M.A. Alonso-Zarazaga in lit.), and this is likely to be the case for this ���Typus��� specimen. Thus, lectotype designation is possible for this nominal species in order to stabilise this species concept, especially because it is a mixed type series and the newly described species in this paper is similar to E. vestitus. Thus, we here designate the male specimen as lectotype (Fig. 9). All other specimens (females) become automatically paralectotypes (Figs 10, 11). All specimens of the type series, except for one are recognised to be conspecific. The outlier specimen (female paralectotype) has (1) denser and longer eye pilosity (eye with very short and scattered pile in the other two females) (Fig. 10C, D), (2) slight but obvious pollinosity surrounding posterior ocelli (this same area is shiny or nearly so in the other two females), (3) individual dots of frontal pollinosity very small (larger in the other females) (Fig. 10A, C), (4) basoflagellomere tapering dorsally for the apical two thirds (for the apical half or less, in the other two females) (Fig. 10C, D), (5) metatibia bumped ventrally (less bumped, tending to straight, in the other two females), (6) apex of metatibia without short black spinae (apex of metatibia with two short black spinae in the other two females) (Fig. 11). This outlier female is similar to the female of E. obliquus (Fabricius 1805) (widespread in Africa) and E. figurans Walker, 1859 (not recorded from Africa). However, it differs from that of E. obliquus in the pollinose vertex (broadly shiny in E. obliquus), wide pollinose posterior margin of scutellum (much narrower to almost absent in E. obliquus), narrow diagonal vittae of terga III and IV (wider in E. obliquus), and shiny posterior margin of tergum IV (extensively pollinose in E. obliquus); and differs from the female of E. figurans in the pollinose vertex and occiput (vertex and occiput shiny in E. figurans), the densely and homogeneously pollinose frons (frons with a medial line of sparser pollinosity in E. figurans), and the short spinae of the anteroapical row of metafemur (longer spinae in E. figurans). The outlier specimen did not key out with Lyneborg���s manuscript key to the Afrotropical species of Eumerus, and might represent an undescribed sister species of E. vestitus. However, we decided not to describe it as a separate taxon due to the absence of other specimens, including males with conspecific morphology. Additional examined material of other Eumerus species. Type series of the nominal species, Eumerus vestitus Bezzi, 1912. Lectotype: 1��, GUINEA PORTOGUESE, Rio Cassine, XII.1899 ��� IV.1900. L. Fea (part of the date crossed out as indicated) / SYNTYPUS �� Eumerus vestitus Bezzi, 1912 (on pink label). Paralectotypes: 1♀, GUINEA POR- TOGUESE, Rio Cassine, XII.1899 ��� IV.1900. L. Fea (part of the date crossed out as indicated) / vestitus Bezzi / TYPUS (printed in red) / Eumerus vestitus n. sp. (handwritten on a pink label; ��� n. sp. ��� is an interpretation of the actual label lettering) / SYNTYPUS ♀ Eumerus vestitus Bezzi, 1912 (on pink label) / Museo Civico di Genova; 2♀, GUINEA PORTOGUESE, Rio Cassine, XII.1899 ��� IV.1900. L. Fea (part of the date crossed out as indicated) / SYNTYPUS ♀ Eumerus vestitus Bezzi, 1912 (on pink label) / Museo Civico di Genova [MCSNG]. The male syntype lacks the antennae and the right prolegs, and the head is pasted to thorax in its original position. The female syntype labelled as ���typus��� lacks the left basoflagellomere, while another female lacks the left metatarsus. There were specimens from Egypt, donated by Becker to Bezzi and found by this latter author that they were erroneously identified as E. obliquus, mentioned in the original description, which we could not locate. Additional material of Eumerus vesti-tus: 2��, 1♀, Egypt, Cairo, Gizera, 24.ix.1992, leg. M. Hauser [CSCA]; 1��, Egypt, Luxor, Westbank of Nile river 25.694N 32,628E, 1.iv.2018 leg Schmid-Egger [CSCA]; 1��, Tunisia, Monastir, 15km S Sousse, 28.vi.1994, leg. M. Hauser (first record of E. vestitus from Tunisia) [CSCA]. Eumerus obliquus: AFRICA. 1♀, ���Cap. B. Spei.��� [South Africa, Cape of Good Hope], Coll. H. Loew, obliquus F (hand written); 1��, Africa, Coll. H. Loew [ZMB]; 1♀ [published in Marcos-Garc��a et al. (2013)], ��le de la R��union (France), Les Avirons, 24.vi.2010, Leg.: N. Estela Ribera, Det. as E. obliquus by A. Ricarte & M.A. Marcos-Garc��a in 2010 (CEUA00105083) [CEUA]; 1��, 2♀, Mozambique, Sofala Prov. Gorongosa Park, small lake, 18��56���39��S 34��26���35��E, 300m, ex Malaise, 19���30.iv.2015 leg. M. Hauser & A. Rung [CSCA]; 1��, Zambia Southern Prov., Livingston, 17.842 S 15.857 E, 960m, 1.v.2016, leg M. Hauser & CJ Borkent [CSCA]; 1��, Zambia, Northern Prov. 8.8 km WSW Kakumbi, S Luangwa NP, 22���26.iv. 2016, 525m, 13.115 S 31.726 E, Malaise trap, leg. M. Hauser, CJ Borkent & DM Ndalamei [CSCA]; 1��, Mali 30 km N Bamako, 20.vii.1991, leg. M. Schwarz [CSCA]; 1��, Ghana, Northern Region, Mole National Park, 165m, 09��15���33��N 01��51���43��W, Malaise trap, 28���30.iv.2014 leg. S. Gaimari & M. Hauser [CSCA]; 1��, Tunisia, Monastir, 15km S Sousse, 28.vi. 1994, leg. M. Hauser [CSCA]. AUSTRA-LIA. 1♀ with puparium, Palmwoods, nr Nambour, Qld, C. Hayward, emerged 17.v.1986, ex rotting guava infested with larvae of Dacus tryoni (UQIC Reg #94996) [CSCA]. EUROPE. 1♀, Spain (mainland), Alicante, San Juan, 01.iv.2020, Leg. M.A. Marcos; 1♀ [published in Ricarte et al. (2008)], Spain, Balearic Islands, Mallorca, Ses Salines, P/ 29.x.2005, Leg.: M.A. Marcos-Garc��a (#6844), Det. as E. obliquus by A. Ricarte in 2006 (CEUA00084841). Eumerus obliquus is widespread all over Africa, also found in the Canary and various Mediterranean islands, as well as in mainland Europe: Spain (first records in the present paper), southern France and Italy (Speight 2020). This species is also introduced in Australia and South America (Garcete-Barrett et al. 2020). A female of Eumerus punctifrons Loew, 1857 with the following data: Tunis, 62285 [ZMB]. Photos of the holotype of Eumerus figurans Walker, 1859 at the Natural History Museum, London, available at https://www.nhm.ac.uk/., Published as part of Ricarte, Antonio, Hauser, Martin, Kinnee, Scott & Marcos-Garc��a, ��ngeles, 2020, A new Eumerus hoverfly (Diptera: Syrphidae) from Namibia and South Africa with notes on similar species, pp. 493-508 in Zootaxa 4890 (4) on pages 502-503, DOI: 10.11646/zootaxa.4890.4.3, http://zenodo.org/record/4306501, {"references":["Bezzi, M. (1912) Ditteri raccolti da Leonardo Fea durante il suo viaggio nell' Africa occidentale. Parte 1 ª: Syrphidae. Annali del Museo civico di storia naturale di Genova, 45, 400 - 453.","International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp.","Fabricius, J. C. (1805) Systema antliatorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Reichard, Brunsvigae, 373 pp. https: // doi. org / 10.5962 / bhl. title. 15806","Walker, F. (1859) Catalogue of the dipterous insects collected at Makessar in Celebes, by Mr. A. R. Wallace, with descriptions of new species. Journal of the proceedings of the Linnean Society, 3 - 4, 90 - 172. https: // doi. org / 10.1111 / j. 1096 - 3642.1859. tb 00089. x","Marcos-Garcia, M. A., Ricarte, A. & Estela, N. (2013) An updated hoverfly checklist (Diptera: Syrphidae) of the Mascarene Island of Reunion, France. Journal of the Entomological Research Society, 15, 59 - 68.","Ricarte, A., Marcos-Garcia, M. A. & Rotheray, G. E. (2008) The early stages and life histories of three Eumerus and two Merodon species (Diptera: syrphidae) from the Mediterranean region. Entomologica Fennica, 19, 129 - 141. https: // doi. org / 10.33338 / ef. 84424","Speight, M. C. D. (2020) Species accounts of European Syrphidae, 2020. Syrph the Net, the database of European Syrphidae (Diptera). Syrph the Net publications, Dublin, 104, 1 - 314.","Garcete-Barrett, B. R., Morales, M. N., Hauser, M., Smit, J. T., Gonzalez, L., Ramirez De Lopez, M. B., Arias, O., Adorno, M., Sormanti, G. & Mereles, A. (2020) New geographical records and key to the species of Eumerus Meigen, 1823 (Diptera, Syrphidae) introduced into the Americas and Hawaii. Revista Brazileira de Entomologia, 64 (1), 1 - 6. https: // doi. org / 10.1590 / 1806 - 9665 - rbent- 2019 - 0016","Loew, H. (1857) Nachricht uber syrische Dipteren. Verhandlungen des zoologisch-botanischen Vereins in Wien, 7, 79 ‾ 86."]}

Key to the African species of the Eumerus obliquus group The Eumerus obliquus group as defined in the discussion includes E. incilis Smit in Smit et al. (2017), E. lyneborgi sp. nov., E. obliquus, E. unicolor Loew, 1858 [= E. wainwrighti (Curran, 1938)] and E. vestitus. 1. Face below antennae polished black, without pollinosity; scutum shiny, without a pattern of pollinose markings or only with a faint vestigial pattern; tarsomeres 4 and 5 of all legs black, contrasting conspicuously with the reddish brown tarsomeres 1���3; male eyes separated by a distance equalling the width of the anterior ocellus.............................. E. unicolor - Face below antennae always pollinose; scutum with a conspicuous pattern of pollinose markings, more reduced but still conspicuous in E. incilis [see figure 39 and 40 in Smit et al. (2017)]; tarsi of all legs either uniform in colour or with a dark gradient towards the apex; male eyes holoptic...................................................................... 2 2. Metatarsomere 2 with a conspicuous apical extension (Fig. 3A, B); male metabasotarsomere unmodified.................................................................................................... E. lyneborgi sp. nov. - Metatarsomere 2 without apical extensions; male metabasotarsomere strongly modified in most species, simple in E. punctifrons............................................................................................... 3 3 Metatibia with 2���3 short black apical spinae (Fig. 11A); posterior half of tergum IV extensively pollinose............... 4 - Metatibia without apical spinae (Fig. 11B); tergum IV shiny between the two diagonal pollinose vittae and the posterior margin of tergum............................................................................................ 5 4 Eye with short sparse pile; metabasotarsomere brown (Fig. 9B) and, in male, with a small basal tooth in the sulcus [see figure 30 in Smit et al. (2017)]......................................................................... E. vestitus - Eye usually bare; metabasotarsomere black and, in male, without teeth in the dorsal sulcus [see figure 27 in Smit et al. (2017)]....................................................................................... E. incilis 5 Vertex with black pile and extensive areas free of pollinosity or sparsely pollinose [see figure 4A in Garcete-Barrett et al. (2020)]; male metabasotarsomere laterally compressed, with a dorsal ridge [see figure 4E in Garcete-Barrett et al. (2020)].............................................................................................. E. obliquus - Vertex without black pile, covered in dense pollinosity except for a narrow area surrounding each ocellus [see figure 57 in Smit et al. (2017)]; male metabasotarsomere without ridge.............................................. E. punctifrons, Published as part of Ricarte, Antonio, Hauser, Martin, Kinnee, Scott & Marcos-Garc��a, ��ngeles, 2020, A new Eumerus hoverfly (Diptera: Syrphidae) from Namibia and South Africa with notes on similar species, pp. 493-508 in Zootaxa 4890 (4) on page 503, DOI: 10.11646/zootaxa.4890.4.3, http://zenodo.org/record/4306501, {"references":["Smit, J. T., van Harten, A. & Ketelaar, R. (2017) Order Diptera, family Syrphidae. The hoverflies of the Arabian Peninsula. In: van Harten, A. (Ed.), Arthropod fauna of the UAE. Vol. 6. Department of the President's Affairs, Abu Dhabi, pp. 572 - 612.","Curran, C. H. (1938) Records and descriptions of African Syrphidae. - I (Diptera). American Museum Novitates, 1009, 1 ‾ 15.","Garcete-Barrett, B. R., Morales, M. N., Hauser, M., Smit, J. T., Gonzalez, L., Ramirez De Lopez, M. B., Arias, O., Adorno, M., Sormanti, G. & Mereles, A. (2020) New geographical records and key to the species of Eumerus Meigen, 1823 (Diptera, Syrphidae) introduced into the Americas and Hawaii. Revista Brazileira de Entomologia, 64 (1), 1 - 6. https: // doi. org / 10.1590 / 1806 - 9665 - rbent- 2019 - 0016"]}

Temporal comparisons of a site's biodiversity depend on the availability of at least two asynchronous data sets on a bioindicator group. The Sierra de Guadarrama, a mountain range in central Spain has a high biodiversity and representative sites such as El Ventorrillo Biological Station (Madrid province) play an important role in research and monitoring of the Sierra biodiversity. Although unevenly and randomly, hoverflies of the Sierra de Guadarrama have been surveyed for decades, but never at El Ventorrillo. This important group of pollinators, aphid predators and bioindicators was sampled with a Malaise trap at El Ventorrillo from June 1989 to June 1990. A total of 51 species were identified (20 spp. of Eristalinae and 31 spp. of Syrphinae), 22 of which were new to the Sierra de Guadarrama and 17 to the Madrid province. The genera Brachypalpus and Didea were also new to the Madrid province. The Sierra de Guadarrama now has 126 species recorded and the Madrid province 150. Within the Sierra de Guadarrama, El Ventorrillo becomes the second locality in number of hoverfly species recorded after El Escorial (64 spp.). In the Malaise trap catch, hoverflies with zoophagous larvae had a higher representation than those of other guilds, both in species richness and abundance. The results provided here for El Ventorrillo represent a baseline for future biodiversity studies at this site with a similar experimental design, for example, to detect changes in hoverfly diversities along time., This study belongs to the ‘Fauna Ibérica’ Project PGC2018-095851-A-C65A (Ministerio de Ciencia, Innovación y Universidades, Spain).

Quichuana nigropilosa Montoya & Ricarte sp. n. Figures 1, 4B, 4 D, 6A���D. Length (n = 1). Body 13.4 mm; wing 11.4 mm. Diagnosis. Large species (13.4 mm) with a broad black abdomen. The species is characterized by three faint, medial, grayish pollinose vittae on the scutum reaching the posterior margin; pleuron black pilose; wing extensively bare of microtrichia on posterior half; capitulum white; coxae black pilose; terga I���V black pilose, with a shiny black pollinose band on posterior margin. Description. FEMALE. Head (Figs 6 A���B). Face black, white pollinose, white pilose; tubercle distinct, with a narrow medial shiny vitta not reaching the antennae and the mouth edge. Gena broad, shiny black. Lunule black; frontal triangle mainly black pilose, except for the golden yellow pile along eye margin. Antennae black, black pilose, basoflagellomere dark brown, round to oval, white pollinose, bf = 1.3; scape black pilose; pedicel golden yellow pilose ventrally, black pilose dorsally; arista orange, bare. Eye yellowish pilose, bare along posterior margin; frontal vertical triangle whitish gray pollinose on anterior corner; ocelli reddish, ocellar triangle black pilose; occiput white pollinose, with a line of long black pile intermixed on dorsal half. Thorax (Fig. 6 C). Black, scutum with a faint gray-brown pollinosity, with black and yellowish pile intermixed, scutum with three faint, medial, grayish pollinose vittae reaching the posterior margin, this area with black pile; postpronotum, notopleuron and postalar callus with golden yellow and black pile intermixed; posterior anepisternum black pilose dorsally, with some white pile intermixed dorsally; pleuron dull grayish pollinose; katepisternum black pilose; katatergum short brownish pilose; scutellum opaque brownish, shiny black on marginal corner, long black pilose; metasternum black pilose; calypter brown, brown pilose marginally; plumula brown; spiracular fringe brownish; halter pedicel white; capitulum white. Wing (Figs 4 C, 6C���D). Microtrichose, except for extensive bare areas on posterior half (cells cua1, cup, dm, bm and anal lobe), slightly dark infuscate on anterior margin (cells c, sc, r1, r4+5, dm and bm); tegula and basicosta with black pile, alula bare medially. Legs (Figs 4 D, 6B). Black, extensively black pilose; coxae and trochanters black, black pilose, pro and mesotrochanter white pilose dorsally; femora black, black pilose, except for the pro and mesofemur golden pilose on anterior edge; metafemur swollen, with long black pile ventrally; tibiae black, black pilose, pro-tibia golden pilose antero-ventrally, meso-tibia white pilose anteroventrally, metatibia extensively with black pile, except for the white pile on anterior and posterior edge; tarsi black, black pilose dorsally, orange pilose ventrally, mesotarsus with rows of ventral black pile. Abdomen (Fig. 6 C). Black, opaque, without markings, extensively covered with black pile; terga I���V with a posterior shiny black margin; tergum I with a moustache arrangement of sparse black pile; tergum II yellow pilose on basal fourth, black pilose posteriorly; terga III���V extensively black pilose; sterna black, sterna I���V with long golden pile, in addition to some laterally black pile intermixed. MALE. Unknown. Material examined. HOLOTYPE: female, deposited at the CEUA, and labelled: COLOMBIA: ��� Antioquia, Betulia, Vereda El Indio, La Mina, Alto de San Jos�� / 6.251447, -76.044872, 3092m, transici��n / 2013. xii.14-18, C. Henao; J. Cogollo. Leg. (CEUA 87092)���. ��� Holotype / Quichuana nigropilosa / Montoya & Ricarte 2017���. [red, handwritten except first line]. Distribution. Eastern slope of Cordillera Occidental, Antioquia, Colombia (3092 m. a. s. l.) (Fig. 1). Etymology. The epithet ���nigropilosa��� (from the Latin ��� nigro ���, black, and the Greek ��� pilos ���, pile) refers to the typical black pile of the body of this singular species. Taxonomic notes. Although Q. nigropilosa sp. n. is described from a single female, we evidenced that the species differs significantly from Q. citara sp. n., Q. melas and Q. atra (see ���Taxonomic notes��� under Q. nigropilosa and Table 1). Another important feature is its restricted distribution, which raises the need for further expeditions to neighboring areas of the type locality, in order to increase the knowledge about species distribution, as well as finding male specimens. Habitat and co-occurrence with other flower fly species. Quichuana nigropilosa sp. n. co-occurs in the paramo de Frontino with the species Argentinomyia browni Fluke, A. luculenta Fluke, A. rex Fluke, Dasysyrphus lotus (Williston), Fazia fascifrons (Macquart) and Ocyptamus eblis (Hull). TABLE]. Morphological đifferences between Quichuana citara sp. n., Q. nigropilosa sp. n., Q. melas Ricarte & Marcos-Garc��a anđ Q. atra Ricarte & Marcos-Garc��a., Published as part of Montoya, Augusto L., Ricarte, Antonio & Wolff, Marta, 2017, Two new species of Quichuana Knab (Diptera: Syrphidae) from the paramo ecosystems in Colombia, pp. 390-402 in Zootaxa 4244 (3) on pages 396-397, DOI: 10.11646/zootaxa.4244.3.7, http://zenodo.org/record/429885

Two new species of Quichuana Knab (Diptera: Syrphidae), Quichuana citara Montoya & Wolff sp. n. and Quichuana nigropilosa Montoya & Ricarte sp. n. are described from highlands of the Colombian Andes. Images of type material, including drawings of male genitalia are provided. An adjustment for the latest identification key for the Quichuana species and distribution maps for those species occurring in Colombia are given.