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Maladera galdaththana Ranasinghe, Eberle, Benjamin & Ahrens, 2020 Material examined SRI LANKA • 1 ♂; “ X-SR2048, Sri Lanka, Kandy District, Gannoruwa FR; 7.28342ºN, 80.59840 ºE; 592, 5 m; 14-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2245, Sri Lanka, Matale District, Riverston, Thelgamu oya bangalow; 7.53635ºN, 80.77234ºE; 509 m; 01-XII- 2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 93, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Ranasinghe S., Eberle J., Benjamin S. P. & Ahrens D. 2020. New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). European Journal of Taxonomy 621: 1 - 20. https: // doi. org / 10.5852 / ejt. 2020.621"]}

Maladera kandyensis Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “X-SR2163, Sri Lanka, Kandy District, Udawattakele FR; 7.29590ºN, 80.64224 ºE; 26-XI-2020; Ranasinghe & Athukorala leg.; Black light”; ZFMK • 1 ♂; “X-SR2045, Sri Lanka, Kandy District, Gannoruwa FR; 7.28368ºN, 80.59874 ºE; 578m; 15-VII-2020; Ranasinghe & Athukorala leg.; Black light”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 93, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Maladera dambullana sp. nov. urn:lsid:zoobank.org:act: 0605777D-D598-4210-BC74-4534D46DC2A2 Figs 6A–D, 7I, 8H Diagnosis The new species is very similar to M. pubescens (Arrow, 1916). Maladera dambullana sp. nov. differs from M. pubescens by the shape of the aedeagus: the ventral distal lobe (i.e., the fused parameres) is not extended mesally but constant in width over its entire length. Etymology The name of the new species is derived from its type locality ‘Dambulla’ (adjective in nominative case singular). Type material Holotype SRI LANKA • ♂; “ X-SR0269, Sri Lanka, Matale District, Dambulla, NIFS Arboretum, 7.85766ºN, 80.67474ºE, 174m, 13-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK. Paratypes SRI LANKA • 1 ♂; “ X-SR0254, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0257, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0258, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0259, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0262, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0264, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0265, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0266, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0267, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0271, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0272, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0279, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0281, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0282, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0283, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0284, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0285, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0286, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0292, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0297, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0300, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0486, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0489, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0490, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0491, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0501, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0507, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0510, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1049, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0341, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0363, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0414, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85796ºN, 80.67554ºE; 181m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ SR0301, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85824ºN, 80.67506ºE; 182m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0985, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0209, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85907ºN, 80.67587ºE; 160m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0210, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85907ºN, 80.67587ºE; 160m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR0261, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X- SR0263, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK. Description MEASUREMENTS. Length: 5.6 mm, length of elytra: 4.1 mm, width: 3.1 mm. HABITUS (Fig. 6D) AND COLOURATION. Body short oval, yellowish brown, antenna yellow, dorsal surface shiny, densely finely setose. HEAD. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anteriorly shallowly sinuate medially; surface slightly convex, finely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture almost invisible and strongly angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and narrow, minutely and superficially punctate, with a single short terminal seta. Frons with fine, dense punctures, with long erect setae in the punctures. Eyes large, ratio of diameter/interocular width: 0.85. Antenna yellow, with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and anteriorly flattened. PRONOTUM. Moderately wide, widest at base, lateral margins weakly convex and evenly narrowed to the anterior third, anteriorly stronger convex, anterior angles strongly produced and sharp, anterior marginal line fine but complete medially, anterior margin weakly produced medially; surface finely and densely punctate, with dense moderately long setae being bent posteriorly on entire disc and with a few sparse longer setae being directed anteriorly; anterior and lateral borders setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fine and dense punctures, with fine and dense adjacent setae. ELYTRA. Short oval, widest shortly behind middle, striae indistinctly impressed, finely and densely punctate, intervals flat, with fine, very dense punctures, with numerous fine setae similar to those of the pronotum and a few sparser ones being longer and erect or directed anteriorly, in particular on lateral intervals; epipleural edge fine, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely setose, metacoxa glabrous except for numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fine and dense punctures bearing each a fine seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/ metacoxa: 1/1.93. Pygidium moderately convex, finely and very densely punctate, without smooth midline, punctures with short and moderately dense, adjacent setae, along the apical margin with a few long erect setae. LEGS. Short and wide; femur with two longitudinal rows of setae, finely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half and very indistinctly serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.18, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at three quarters of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines and beside them a longitudinal serrated line; lateral face almost flat, with dense, fine punctures and with minute setae in the punctures; ventral margin with five strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso- and metatarsomeres finely and densely punctate and setose dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; first metatarsomere as long as following two tarsomeres combined and a little longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. AEDEAGUS. Fig. 6A–C. Variation Length: 5.6–6.7 mm, length of elytra: 4.1–4.6 mm, width: 3.1–3.5 mm. Female Length: 6.0– 6.5 mm, length of elytra: 4.7–4.9 mm, width: 3.7–3.8 mm. Eyes slightly smaller than in male; antennal club little shorter than remaining antennomeres combined. Distribution See Fig. 7I., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on pages 81-85, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Serica lurida Brenske, 1898 Material examined SRI LANKA • 1 ♂; “ X-SR0591, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0615, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0616, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0617, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0618, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0897, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0898, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0899, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0900, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0901, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0902, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0903, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0904, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0905, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0906, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0907, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0908, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0909, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0910, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0911, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0912, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0913, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0914, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 99, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Neoserica pophami sp. nov. urn:lsid:zoobank.org:act: 86E89148-395A-4509-A223-69CBAC480FBE Figs 3I–L, 7C, 8C Diagnosis Neoserica pophami sp. nov. differs from the very similar N. kalaarensis Fabrizi & Ahrens, 2014 in shape of parameres: the right paramere is slightly longer, medially distinctly bent (lateral view) and much wider (dorsal view). Etymology The new species is named after Mr Sam Popham, founder of the NIFS Arboretum (noun in the genitive case). Type material Holotype SRI LANKA • ♂; “ X-SR0346, Sri Lanka, Matale District, Dambulla, NIFS Arboretum, 7.85783ºN, 80.67391ºE, 167m, 13-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK. Paratypes SRI LANKA • 1 ♂; “ X-SR0332, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0333, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0348, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0353, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0357, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0360, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0365, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0369, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0370, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0380, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0383, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0384, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0388, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0393, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0394, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0401, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1081, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0471, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0473, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0474, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0475, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0488, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13- X _2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0417, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85796ºN, 80.67554ºE; 181m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0423, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85796ºN, 80.67554ºE; 181m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1014, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85824ºN, 182m; 80.67506ºE; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X- SR1042, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 174m; 12- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR1056, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 174m; 12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR1067, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 174m; 12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR1068, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 174m; 12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK. Description MEASUREMENTS. Length: 6.1 mm, length of elytra: 4.4 mm, width: 3.5 mm. HABITUS (Fig. 3L) AND COLOURATION. Body oval, light reddish brown, antenna yellow, dorsal completely shiny, except a few setae on head almost glabrous. HEAD. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and convergent anteriorly, anterior angles moderately rounded, anterior margin slightly sinuated medially, all margins moderately reflexed, lateral margins produce an indistinct angle with the ocular canthus; surface weakly convex medially, densely punctate, coarse and fine punctures mixed, with numerous erect setae; frontoclypeal suture indistinctly incised, not elevated and weakly angled medially; smooth area anterior to eye flat, three times as wide as long; ocular canthus moderately long and wide (one third of ocular diameter), finely and densely punctate, with a single terminal seta. Frons with fine, sparse punctures, with a few erect setae beside eyes. Eyes moderately large, ratio diameter/interocular width: 0.75. Antenna with ten antennomeres; club with five antennomeres and straight, as long as the remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. PRONOTUM. Moderately transverse, widest at base, lateral margins in basal half almost straight and moderately convergent anteriorly, in anterior half weakly convex and evenly convergent anteriorly, anterior angles distinctly produced and moderately acute, posterior angles blunt; anterior margin almost not produced medially, with a robust and complete marginal line, base of pronotum without marginal line; surface finely and densely punctate, punctures glabrous; lateral and lateral anterior margin sparsely setose; hypomeron carinate, not ventrally produced. Scutellum wide, triangular, at apex moderately pointed, with fine, moderately dense punctures, glabrous. ELYTRA. Widest shortly behind the middle, striae weakly impressed, finely and moderately densely punctate, intervals flat, with fine and evenly moderately dense punctures, except a few short setae on lateral intervals glabrous; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra with a fine rim of microtrichomes (100 ×). VENTRAL SURFACE. In major part dull, some parts shiny, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae, metacoxa glabrous, with a few long setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta, the row of punctures on the first visible basal sternite fused to a robust transverse carina bearing the setae, before the carina the fine punctation extremely dense but glabrous. Mesosternum between mesocoxae 1.5 times as wide as the width of mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.65. Pygidium strongly convex and shiny, finely and moderately densely punctate, without smooth midline, glabrous except some longer setae along the apical margin. LEGS. Wide and moderately long; femur with two longitudinal row of setae, finely and sparsely punctate; metafemur shiny, behind the posterior longitudinal row of setae punctures finer and slightly denser, anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally, strongly widened, posterior margin finely serrated over its entire length dorsally, with just a few short setae basally. Metatibia wide and flattened, short, widest at apical second third of metatibial length, ratio of width/length: 1/2.8, sharply carinate dorsally, with two groups of spines, basal group at first third, apical group at two thirds of metatibial length, basally with a few short robust single spines, on basal quarter beside dorsal margin with a short serrated line; lateral face weakly longitudinally convex, finely, superficially and sparsely punctate, glabrous, widely smooth along the middle; ventral margin finely serrated, with four robust equidistant setae; medial face smooth and glabrous; apex finely serrated, interiorly near tarsal articulation weakly concavely sinuate. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, ventrally robustly densely setose; metatarsomeres with a strongly serrated ridge and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. AEDEAGUS. Fig. 3I–K. Variation Length: 6.0– 6.8 mm, length of elytra: 4.3–4.7 mm, width: 3.2–3.6 mm. Female Length: 6.8–7.2 mm, length of elytra: 4.6–5.0 mm, width: 3.7–4.0 mm. Eyes as large as in male; antennal club shorter little than remaining antennomeres combined, the basal joint of club equals only ⅓ of the length of the club; pygidium weakly convex. Distribution See Fig. 7C., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on pages 66-69, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Selaserica fabriziae sp. nov. urn:lsid:zoobank.org:act: 12205E52-C260-4128-9A23-4A7F6D9D2067 Figs 3A–D, 7A, 8A Diagnosis Selaserica fabriziae sp. nov. is in shape of aedeagus rather similar to Sel. sericea (Arrow, 1916), it differs by the shiny dorsal body surface and the shape of the parameres and phallobase: the median dorsoapical sinuation of the phallobase is much narrower (than in Sel. sericea), the parameres are bent dorsally at the apex (not bent in Sel. sericea). Etymology The new species is dedicated to Silvia Fabrizi, who passed away too early, in memory of her and for all her efforts for Sericini taxonomy (noun in genitive singular case). We will not forget you, Silvia! Type material Holotype SRI LANKA • ♂; “ X-SR0859, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 26-X-2019; Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK. Paratypes SRI LANKA • 1 ♂; “ X-SR1948, Sri Lanka, Galle District, Kottawa FR, 6.09712ºN, 80.31666ºE; 46m, 29-30-VI-2020, Benjamin & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR1956, Sri Lanka, Galle District, Kottawa FR, 6.09712ºN, 80.31666ºE; 46m, 29-30-VI-2020, Benjamin & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR1957, Sri Lanka, Galle District, Kottawa FR, 6.09712ºN, 80.31666ºE; 46m, 29-30-VI-2020, Benjamin & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR1958, Sri Lanka, Galle District, Kottawa FR, 6.09712ºN, 80.31666ºE; 46m, 29-30-VI-2020, Benjamin & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR1959, Sri Lanka, Galle District, Kottawa FR, 6.09712ºN, 80.31666ºE; 46m, 29-30-VI-2020, Benjamin & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X- SR1960, Sri Lanka, Galle District, Kottawa FR, 6.09712ºN, 80.31666ºE; 46m, 29-30-VI-2020, Benjamin & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X- SR2118, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 11-XII-2020; Ranasinghe & Athukorala, Black light ”; ZFMK • 1 ♂; “ X- SR2119, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 11-XII-2020; Ranasinghe & Athukorala, Black light ”; ZFMK • 1 ♂; “ X- SR2120, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 11-XII-2020; Ranasinghe & Athukorala, Black light ”; ZFMK • 1 ♂; “ X- SR2121, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 11-XII-2020; Ranasinghe & Athukorala, Black light ”; ZFMK • 1 ♂; “ X- SR2122, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 11-XII-2020; Ranasinghe & Athukorala, Black light ”; ZFMK • 1 ♀; “ X- SR1961, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 30-VI-2020, Benjamin & Ranasinghe, Black light ”; ZFMK • 1 ♀; “ X- SR2123, Sri Lanka, Galle District, Kottawa FR, 6.09812ºN, 80.31610ºE, 44m, 11-XII-2020; Ranasinghe & Athukorala, Black light ”; ZFMK • 1 ♀;“ X- SR2104, Sri Lanka, Galle District, Kottawa FR, 6.09712ºN, 80.31666ºE; 46m, 11-XII-2020; Ranasinghe & Athukorala, Black light ”; ZFMK. Description MEASUREMENTS. Length: 8.9 mm, length of elytra: 6.1 mm, width: 5.2 mm. HABITUS (Fig. 3D) AND COLOURATION. Body oval, dark brown, antenna yellow, dorsal surface shiny and glabrous. HEAD. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins weakly convex and convergent to moderately rounded anterior angles, lateral border and ocular canthus producing a blunt angle, margins weakly reflexed, anterior margin shallowly sinuate medially; surface strongly convex medially, weakly shiny, anterior half nearly impunctate, behind finely and densely punctate, distance between punctures subequal to their diameter, with a few fine setae behind anterior margin; frontoclypeal suture very feebly impressed and weakly angled medially; smooth area in front of eye approximately three times as wide as long; ocular canthus moderately long and narrow, impunctate but surface slightly concave, with a single short terminal seta. Frons with fine, moderately dense punctures, posterior half impunctate, surface glabrous except for a few setae anteriorly beside eyes. Eyes moderately large, ratio of diameter/interocular width: 0.7. Antenna yellowish brown, with ten antennomeres; club with four antennomeres, 1.3 times as long as remaining antennomeres combined. Mentum elevated and anteriorly flattened. PRONOTUM. Moderately wide, widest at base, lateral margins evenly convex and narrowed to anterior angles, anterior angles strongly produced and sharp, anterior marginal line widely incomplete medially, anterior margin moderately produced medially; surface densely and coarsely punctate, glabrous; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and but not produced ventrally. Scutellum wide, triangular, with fine and dense punctures, each bearing a single very minute seta. ELYTRA. Oblong, widest in posterior third, striae indistinctly impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures concentrated along striae, punctures partly with minute setae, without longer erect setae; epipleural edge fine, ending at the convex external apical angle of elytra, epipleura densely setose, apical border membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Dull, thorax and metacoxa with large and dense punctures, sparsely finely setose, metacoxa glabrous except for numerous short setae laterally; each abdominal sternite with generally distributed fine and dense punctures, without a distinct transverse row of coarse punctures each bearing a short seta, punctures with very short or minute setae, penultimate sternite apically with a shiny smooth but very short chitinous border. Mesosternum between mesocoxae little wider than mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.15. Pygidium lost in holotype. LEGS. Moderately wide;femur with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior edge acute, with adjacent continuously serrated line, anterior longitudinal row of setae complete; posterior ventral margin almost straight, weakly widened in apical half, nor ventrally nor dorsally serrated, glabrous. Metatibia moderately wide and long, widest behind middle, dorsal and ventral margins in posterior two thirds subparallel, ratio width/length: 1/3.2, dorsally weakly edged, with two groups of spines, basal one at first quarter, apical one at three quarters of metatibial length, basally beside dorsal margin with a very short serrated line; lateral face longitudinally convex, with very sparse and fine punctures, punctures glabrous; ventral margin with three strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation distinctly concave. Tarsomeres impunctate dorsally, ventrally with dense, fine setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is no strong longitudinal carina, first metatarsomere one quarter of its length longer than dorsal tibial spur, subsequent tarsomeres lacking in holotype. Protibia moderately long, tridentate. Protarsomeres lacking in holotype. AEDEAGUS. Fig. 3A–C. Variation Length: 8.2–8.8 mm, length of elytra: 6.1–6.25 mm, width: 5.2–5.3 mm. Pygidium moderately convex, shiny, finely densely punctate, without smooth midline, glabrous except some longer setae along the apical margin. First metatarsomere one quarter of its length longer than dorsal tibial spur, slightly shorter than following two tarsomeres combined. Protibia moderately long, tridentate. Protarsomeres ventrally with some yellow setae, protarsal claws symmetrical, feebly curved and short, with normally developed basal tooth. Female Length: 8.8–9.2 mm, length of elytra: 6.2–7.0 mm, width: 5.5–5.7 mm. Eyes as large as in male; antennal club as long as remaining antennomeres combined. Distribution See Fig. 7A., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on pages 60-63, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Maladera calcarata (Brenske, 1898) Material examined SRI LANKA • 1 ♂; “ X-SR0309, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85824ºN, 80.67506ºE; 182m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 92, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Maladera bandarwelana Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR0666, Sri Lanka, Matale District, Riverston, below the Greenview lodge; 7.53830ºN, 80.7511ºE; 765m; 14-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0881, Sri Lanka, Matale District, Riverston, below the Greenview lodge; 7.54976ºN, 80.75212ºE; 902m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • ZFMK • 1 ♀; “ X-SR0882, Sri Lanka, Matale District, Riverston, below the Greenview lodge; 7.54976ºN, 80.75212ºE; 902m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 92, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Selaserica nuwarana Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR2058, Sri Lanka, Nuwara Eliya District, Hakgala SNR, Seetha Eliya; 6.9299ºN, 80.81359ºE; 1789m; 21-XII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2099, Sri Lanka, Nuwara Eliya District, Hakgala SNR, Seetha Eliya; 6.93074ºN, 80.81341ºE; 1773m; 21-XII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 97, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Maladera hiyarensis sp. nov. urn:lsid:zoobank.org:act: EA7F3C4B-534E-40E5-ADC5-A5FE770CECEE Figs 5E–H, 7H, 8G Diagnosis Maladera hiyarensis sp. nov. is in external appearance similar to M. anderssoni Fabrizi & Ahrens, 2014 and M. romanoi Fabrizi & Ahrens, 2014 being, however, stouter in shape and larger, the aedeagus has the distal part longer and narrower. The aedeagus is also rather similar to that of M. badullana Fabrizi & Ahrens, 2014, but the ventral convexity is in this new species less pronounced and the distal portion is longer and reflexed (lateral view). Etymology The new species is named after its type locality ‘Hiyare’ (adjective in the nominative singular). Type material Holotype SRI LANKA • ♂; “ X-SR1946, Sri Lanka, Galle District, Hiyare FR; 6.05959ºN, 80.31503ºE; 116m; 01-VII-2020; Benjamin & Ranasinghe leg.; Black light ”; ZFMK. Description MEASUREMENTS. Length: 6.8 mm, length of elytra: 4.8 mm, width: 4.1 mm. HABITUS (Fig. 5H) AND COLOURATION. Body oval, reddish brown, frons, disc of pronotum and a few spots on elytra dark brown, antenna and legs yellow, dorsal surface with iridescent shine, densely and shortly setose, elytra with numerous single erect setae. HEAD. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins moderately reflexed; anterior margin very weakly emarginate medially; surface shiny, weakly elevated medially, finely and densely punctate, distance between punctures equal their diameter, with a few erect setae in larger punctures; frontoclypeal suture fine and weakly angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus moderately long and narrow, impunctate, with a single short terminal seta. Frons with toment and iridescent shine, with fine, dense punctures and minute setae in punctures, with a few long, erect setae on disc and beside eyes. Eyes large, ratio of diameter/interocular width: 0.79. Antenna yellow, with ten antennomeres; club with three antennomeres, straight, 1.1 times as long as remaining antennomeres combined. Mentum elevated and anteriorly flattened. PRONOTUM. Moderately wide, widest at base, lateral margins in basal half straight and convergent, in anterior half weakly convex and convergent anteriorly; anterior angles moderately produced and sharp; anterior marginal line fine and complete, anterior margin weakly produced medially; surface finely and densely punctate, with short setae in punctures, otherwise glabrous; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fine and dense punctures, along midline narrowly impunctate. ELYTRA. Short oval, widest at posterior third, striae indistinctly impressed, finely and densely punctate, intervals flat, with fine, dense punctures, with short setae in punctures; odd intervals with a few impunctate dots which are darker and each bear at centre an erect seta; epipleural edge fine, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border membranous, apex covered with a rim of short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely shortly setose; metacoxa with numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fine and dense punctures bearing each a fine seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a narrow, shiny smooth chitinous border. Mesosternum between mesocoxae as wide as mesofemur, with a semicircular ridge bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/2.24. Pygidium weakly convex, finely densely punctate, with short dense setae, and numerous long setae along apical margin. LEGS. Short and wide; femur with two longitudinal rows of setae, finely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose; posterior ventral margin straight, strongly widened in apical half and very indistinctly serrate apically; posterior dorsal margin not serrated, densely setose. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.4, dorsally sharply carinate, with two groups of spines, basal one shortly behind middle, apical one at four fifths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines and beside them a longitudinal serrated line; lateral face longitudinal convex, with dense, fine punctures and with short setae in punctures; ventral margin finely serrate, with five strong spines equidistant from each other; medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso- and metatarsomeres dorsally sparsely and very finely punctate and with minute setae in punctures, ventrally with robust, dense, short setae; metatarsomeres with a strongly serrated ventral ridge, beside it with a strong longitudinal carina; first metatarsomere as long as two following tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth, distal protarsomeres lacking in holotype. AEDEAGUS. Fig. 5E–G. Female Unknown. Distribution See Fig. 7H., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on pages 79-81, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Selaserica sororinitida sp. nov. urn:lsid:zoobank.org:act: 2B2B4EB0-D50B-409B-8F87-4577B9BC235B Figs 3E–H, 7B, 8B Diagnosis Selaserica sororinitida sp. nov. is in shape of aedeagus and external morphology similar to Sel. nitida (Candèze, 1861) and Sel. knucklensis Fabrizi & Ahrens, 2014. Selaserica sororinitida differs by the antennal club, which is longer compared to that of Sel. nitida but shorter in comparison with that of Sel. knucklensis. From both species Sel. sororinitida can also be distinguished by the shape of the aedeagus: the right paramere is in the new species shorter and less narrowed before the apex (lateral view); the phallobase is not widened distally as in Sel. knucklensis. Etymology The name is derived from the Latin noun ‘ soror ’ (sister) with ‘ nitida ’ from the species Sel. nitida, with reference to the strong morphological similarity with Sel. nitida (noun in nominative case). Type material Holotype SRI LANKA • ♂; “ X-SR0670, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33159ºN, 80.86110ºE; 1139m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK. Paratypes SRI LANKA • 1 ♂; “ X-SR0224, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0227, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0228, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0229, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0230, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0231, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0232, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0233, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0234, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0235, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0236, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0237, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0240, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0511, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0512, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0513, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0514, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0515, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0516, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0517, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0518, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0519, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0520, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0521, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0941, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0942, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-18- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0563, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33616ºN, 80.85907ºE; 1197m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0664, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33501ºN, 80. 85966ºE; 1171m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0669, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33159ºN, 80. 86110ºE; 1139m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0915, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33159ºN, 80. 86110ºE; 1139m; 17- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1135, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33501ºN, 80. 85966ºE; 1171m; 21-II-2019; Eberle & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR0186, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 20-II-2019; Eberle & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X- SR0239, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 18-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR0943, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR2194, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 05-XII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♀; “ X-SR2184, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33159ºN, 80. 86110ºE; 1139m; 05-XII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♀; “ X-SR0793, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33501ºN, 80. 85966ºE; 1171m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♀; “ X-SR0795, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33501ºN, 80. 85966ºE; 1171m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK. Description MEASUREMENTS. Length: 10.2 mm, length of elytra: 7.1 mm, width: 5.6 mm. HABITUS (Fig. 3H) AND COLOURATION. Body oblong, reddish brown, antenna yellow, dorsal surface shiny and glabrous, ventral surface dull. HEAD. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to strongly rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed; anterior margin shallowly sinuate medially; surface almost flat, finely and densely punctate, distance between punctures subequal to their diameter, with a few robust setae anteriorly; frontoclypeal suture very feebly impressed and curved medially; smooth area in front of eye approximately three times as wide as long; ocular canthus moderately long and narrow, impunctate, with a single short terminal seta. Frons with fine, sparse punctures, posterior third impunctate, surface glabrous except for a few long setae beside eyes. Eyes large, ratio of diameter/ interocular width: 0.9. Antenna yellowish brown, with nine antennomeres; club with four antennomeres, 1.2 times as long as remaining antennomeres combined. Mentum elevated and anteriorly flattened. PRONOTUM. Moderately wide, widest at base, lateral margins in basal half almost straight and weakly convergent, in anterior half weakly convex and distinctly narrowed towards anterior angles, anterior angles moderately produced and moderately sharp; anterior margin moderately produced medially, its marginal line complete; surface moderately densely and finely punctate, glabrous; anterior and lateral borders sparsely setose; basal margin without marginal line. Hypomeron ventrobasally not carinate. Scutellum wide, triangular, with fine and dense punctures, glabrous, apical portion completely impunctate. ELYTRA. Oblong, widest in posterior third, striae distinctly impressed, finely and densely punctate, intervals flat, with fine, sparse punctures, glabrous, only on penultimate lateral interval a few single long, erect setae; epipleural edge fine, ending well anterior to convex external apical angle of elytra; epipleura densely setose; apical border membranous, apex covered with a narrow rim of short microtrichomes. VENTRAL SURFACE. Dull, thorax and metacoxa with large and dense punctures, with long and dense setae; metacoxa glabrous except for a few long setae laterally; each abdominal sternite with evenly distributed fine and dense punctures, and with a transverse row of coarse punctures each bearing a robust seta, other punctures with a short or minute setae, penultimate sternite apically with a shiny smooth but very short chitinous border. Mesosternum between mesocoxae little narrower than maximum width of mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.28. Pygidium weakly convex, shiny, finely densely punctate, with long fine setae along the apical margin, otherwise glabrous. LEGS. Moderately wide and shiny; femur with two longitudinal rows of setae, finely and moderately densely punctate; metafemur with acute anterior edge, without an adjacent serrated line, anterior longitudinal row of setae complete; posterior ventral margin almost weakly convex, weakly widened in apical half, not serrated, glabrous; posterior dorsal margin smooth, densely shortly setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.3; dorsal margin longitudinally convex, with two groups of spines, basal one at first quarter, apical one at three quarters of metatibial length, without a serrated line; lateral face longitudinally convex, with very sparse and fine punctures, glabrous; ventral margin serrated, with five robust equidistant spines; medial face smooth; apex interiorly near tarsal articulation concavely emarginate. Tarsomeres dorsally impunctate and glabrous, ventrally with dense, long and robust setae; metatarsomeres ventrally with a serrated ridge, beside it without additional strong longitudinal carina; first metatarsomere one third of its length longer than dorsal tibial spur, distinctly shorter than following two tarsomeres combined. Protibia moderately long, tridentate, basal tooth smaller than the two distal ones. Protarsomeres ventrally with long and dense yellow setae forming a setose pad, protarsal claws symmetrical, basal tooth of claws sharply pointed. AEDEAGUS. Fig. 3E–G. Variation Length: 9.9–12.0 mm, length of elytra: 7.1–8.1 mm, width: 5.3–5.8 mm. Female Length: 12.0– 12.2 mm, length of elytra: 8.1–8.3 mm, width: 6.2–6.4 mm. Eyes as smaller than in male; antennal club with three antennomeres and longer than remaining antennomeres combined. Distribution See Fig. 7B.

Maladera cervicornis Ranasinghe, Eberle, Benjamin & Ahrens, 2020 Material examined SRI LANKA • 1 ♂; “ X-SR0621, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53827ºN, 80.75ºE; 782m; 14-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0622, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53827ºN, 80.75ºE; 782m; 14-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0623, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53827ºN, 80.75ºE; 782m; 14-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0624, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53827ºN, 80.75ºE; 782m; 14-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0625, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53827ºN, 80.75ºE; 782m; 14-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0626, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53827ºN, 80.75ºE; 782m; 14-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂, “ X-SR0653, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.5491ºN, 80.75386ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂, “ X-SR0867, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.5491ºN, 80.75386ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂, “ X-SR0868, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.5491ºN, 80.75386ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂, “ X-SR0869, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.5491ºN, 80.75386ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂, “ X-SR2237, Matale District, Riverston, Thelgamu oya bungalow; 7.5363ºN, 80.7723ºE; 509m; 02-XII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 92, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Ranasinghe S., Eberle J., Benjamin S. P. & Ahrens D. 2020. New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). European Journal of Taxonomy 621: 1 - 20. https: // doi. org / 10.5852 / ejt. 2020.621"]}

Maladera heveli Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR0988, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0989, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0990, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1006, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1089, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR1090, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1091, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1092, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85897ºN, 80.67533ºE; 203m; 11-12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0404, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.86011ºN, 80.67441ºE; 187m; 11-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X- SR0406, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.86011ºN, 80.67441ºE; 187m; 11-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR1037, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.86011ºN, 80.67441ºE; 187m; 11-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 93, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Maladera kishi sp. nov. urn:lsid:zoobank.org:act: 35A181EA-5701-48E5-9A1F-75906A829611 Figs 4E–H, 7E, 8D Diagnosis Maladera kishi sp. nov. is in external morphology very similar to M. haniel sp. nov. and differs by the lack of a tubercle on abdominal sternite III as well as by the shape of parameres: the fused parameres (distal aedeagal process) is simply pointed and not bifurcate, while the lateral process on the left side is more robust. Etymology The new species is named for Suresh Benjamin’s daughter, Kishi P.L. Benjamin (noun in apposition). Type material Holotype SRI LANKA • ♂; “ X-SR0724, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK. Paratypes SRI LANKA • 1 ♂; “ X-SR0730, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0731, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0736, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0737, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0738, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0739, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0740, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0741, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0742, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0743, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0749, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0750, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0697, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0701, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0924, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0925, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0926, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0927, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0929, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m, 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0565, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33616ºN, 80.85907ºE; 1197m; 17-18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0566, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33616ºN, 80.85907ºE; 1197m; 17-18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0936, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33616ºN, 80.85907ºE; 1197m; 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0546, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33501ºN, 80.85966ºE; 1171m; 17-18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♂; “ X-SR0548, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33501ºN, 80.85966ºE; 1171m; 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♀; “ X-SR0545, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33501ºN, 80.85966ºE; 1171m; 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♀; “ X-SR0567, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33616ºN, 80.85907ºE; 1197m; 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♀; “ X-SR0700, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m; 17-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♀; “ X-SR0725, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.3389ºN, 80.8510ºE, 1192m; 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♀; “ X-SR0799, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33501ºN, 80.85966ºE; 1171m; 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK • 1 ♀; “ X-SR0800, Sri Lanka, Kandy District, Deenston, Knuckles South, 7.33501ºN, 80.85966ºE; 1171m; 18-X-2019, Eberle, Bohacz & Ranasinghe, Black light ”; ZFMK. Description MEASUREMENTS. Length: 9.2 mm, length of elytra: 6.6 mm, width: 5.2 mm. HABITUS (Fig. 4H) AND COLOURATION. Body short oval, dark brown, antenna yellow, dorsal surface shiny, finely densely setose. HEAD. Labroclypeus short and trapezoidal, wider than long, widest at base, lateral margins strongly convex and convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anterior margin not sinuate medially; surface slightly convex, finely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture indistinctly incised and bluntly bent medially; smooth area in front of eye approximately three times as wide as long; ocular canthus long and narrow, minutely and superficially punctate, without a short terminal seta. Frons with fine, dense punctures, with a few long erect setae in larger punctures. Eyes large, ratio of diameter/interocular width: 0.81. Antenna yellow, with ten antennomeres; club with three antennomeres, little longer than remaining antennomeres combined. Mentum elevated and anteriorly flattened. PRONOTUM. Wide, widest at base, lateral margins in basal half straight and weakly convergent, in anterior half weakly convex and narrowed to anterior angles, anterior angles moderately produced and sharp, anterior marginal line fine and complete, anterior margin convexly produced medially; surface finely densely punctate, with moderately dense, short and fine setae and dense long erect setae being directed anteriorly; anterior and lateral borders sparsely setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fine and dense punctures, with short, dense, fine setae. ELYTRA. Short oval, widest shortly behind middle, striae distinctly impressed, finely and densely punctate, intervals weakly convex, with moderately fine, dense punctures and with dense, fine, short setae as well as with sparse long erect setae; epipleural edge fine, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Shiny, thorax and metacoxa with large and dense punctures, sparsely setose, metacoxa with minute adjacent setae in the punctures except for numerous long setae laterally, apical margin weakly convex, without a wide rim of long white microtrichomes; each abdominal sternite, in addition to evenly distributed fine and dense punctures bearing each a fine seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/ metacoxa: 1/1.8. Pygidium moderately convex, moderately finely and densely punctate, without smooth midline, punctures with short and dense, adjacent setae as well as with dense, long, erect setae. LEGS. Short and wide; femur with two longitudinal rows of setae, finely and densely punctate; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half, finely indistinctly serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly only very slightly narrowed, ratio width/length: 1/2.4, dorsally sharply carinate, with two groups of spines, basal well behind middle, apical one at four fifths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing a single robust spine and beside them a longitudinal serrated line; lateral face almost flat, with dense, large punctures and with fine setae in the punctures; ventral margin with five strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly concave. Meso- and metatarsomeres finely and sparsely punctate but glabrous dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and as long as dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. AEDEAGUS. Fig. 4E–G. Variation Length: 8.9–9.2 mm, length of elytra: 6.6–7.1 mm, width: 5.1–5.6 mm. Female Length: 9.0– 9.3 mm, length of elytra: 7.0– 7.5 mm, width: 5.4–5.7 mm. Eyes as large as in male; antennal club little shorter than the remaining antennomeres combined. Distribution See Fig. 7E.

Maladera setosa (Brenske, 1896) Material examined SRI LANKA • 1 ♂; “ X-SR1024, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR1035, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1047, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0372, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0457, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0460, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0465, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1039, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1052, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0411, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85796ºN, 80.67554ºE; 181m; 12- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0450, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85796ºN, 80.67554ºE; 181m; 12-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1013, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85824ºN, 80.67506ºE; 182m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0405, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.86011766ºN, 80.67441844ºE; 187m; 11-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 96, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Maladera laterita (Moser, 1915) Material examined SRI LANKA • 1 ♂; “ X-SR0866, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911ºN, 80.75386ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 94, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Maladera rufocuprea (Blanchard, 1850) Material examined SRI LANKA • 1 ♂; “ X-SR0844, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0845, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0846, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X- SR0847, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0848, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0849, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23- X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0850, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0851, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0852, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0853, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0854, Sri Lanka, Galle District, Hiyare FR; 6.05871ºN, 80.31538ºE; 117m; 22-23-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0529, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0530, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0531, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0532, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0533, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0569, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0570, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0571, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0752, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0753, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0754, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0755, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0762, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0763, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0773, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0774, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0775, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0779, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0788, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0652, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.5491115ºN, 80.7538616ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0864, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.5491115ºN, 80.7538616ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0865, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.5491115ºN, 80.7538616ºE; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0877, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54976ºN, 80.75212ºE; 902; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0878, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54976ºN, 80.75212ºE; 902; 15-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1082, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85783ºN, 80.67391ºE; 167m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 94, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Maladera tricuspidata Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR0326, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85824ºN, 80.67506ºE; 182m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 96, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Maladera padaviyaensis Ahrens & Fabrizi, 2016 Material examined SRI LANKA • 1 ♂; “ X-SR1046, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR1915, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85796ºN, 80.67554ºE; 181m; 13-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK.

Selaserica pusilla Arrow, 1916 Material examined SRI LANKA • 1 ♂; “ X-SR0654, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53804ºN, 80.75027ºE; 778m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X- SR0655, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53804ºN, 80.75027ºE; 778m; 15- X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0656, Sri Lanka, Matale District, Riverston, Greenview lodge; 7.53804ºN, 80.75027ºE; 778m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0627, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0631, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0634, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m; 15-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0635, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m; 778m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0636, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m; 778m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0637, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0638, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m; 15-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0639, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0640, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7.54911N, 80.75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0641, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0642, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0643, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0644, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0645, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0650, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0651, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0671, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0672, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0673, Sri Lanka, Matale District, Riverston, Pitawala Pathana; 7,54911N, 80,75386E; 880m; 15-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0252, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0253, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0256, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0287, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0295, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0296, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0298, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0299, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0485, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0487, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0504, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0505, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0506, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0507, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0508, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0509, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13- X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0998, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0999, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1021, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR1033, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12- 13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1048, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.85766ºN, 80.67474ºE; 174m; 12-13-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 98, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Selaserica convexiuscula Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR0858, Sri Lanka, Galle District, Kottawa FR; 6.09712ºN, 80.3166ºE; 49m; 26-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2074, Sri Lanka, Galle District, Kottawa FR; 6.09712ºN, 80.3166ºE; 49m; 11-XII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK.

Selaserica praetexta Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR1973, Sri Lanka, Kandy District, Gannoruwa FR; 7.28329ºN, 80.59819 ºE; 528 m; 16-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1974, Sri Lanka, Kandy District, Gannoruwa FR; 7.28329ºN, 80.59819 ºE; 528 m; 16-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1975, Sri Lanka, Kandy District, Gannoruwa FR; 7.28329ºN, 80.59819 ºE; 528 m; 16-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2042, Sri Lanka, Kandy District, Gannoruwa FR; 7.28329ºN, 80.59819 ºE; 528 m; 15-VII- 2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2208, Sri Lanka, Kandy District, Gannoruwa FR; 7.28329ºN, 80.59819 ºE; 528 m; 25-XI-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2209, Sri Lanka, Kandy District, Gannoruwa FR; 7.28329ºN, 80.59819 ºE; 528 m; 25 Nov. 2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1972, Sri Lanka, Kandy District, Gannoruwa FR; 7.28342ºN, 80.59840 ºE; 592.5 m; 16-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2046, Sri Lanka, Kandy District, Gannoruwa FR; 7.28342ºN, 80.59840 ºE; 592.5 m; 14-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR1970, Sri Lanka, Kandy District, Udawattakele FR; 7.29590ºN, 80.64224ºE; 16-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2159, Sri Lanka, Kandy District, Udawattakele FR; 7.29690ºN, 80.6448ºE; 26-XI-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2160, Sri Lanka, Kandy District, Udawattakele FR; 7.29690ºN, 80.6448ºE; 27-XI- 2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2017, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 24-VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2025, Sri Lanka, Matale District, Riverston, Pitawala Pathana, 7.55223ºN, 80.75293ºE, 888m; 23 -VII-2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 97, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Updated and corrected key to species of the Maladera fistulosa group (♂♂) 1. Pronotum glabrous on disc, sometimes a few sparse setae on lateral pronotal disc. Anterior marginal line of pronotum widely incomplete medially.................................................................................. 2 – Pronotum densely setose................................................................................................................. 13 2. Apical margin of metacoxa slightly concave, glabrous. Metatibia basally with a longitudinal serrated line.................................................................................................................................................... 3 – Apical margin of metacoxa straight or convex. Metatibia basally without a longitudinal serrated line.................................................................................................................................................... 5 3. Metatibia more stout (ratio width/length> 1/3.2)............................................................................ 4 – Metatibia more slender (ratio width/length M. hortonensis Fabrizi & Ahrens, 2014 4. Metatibia ratio width/length: 1/2.8. Antennal club 1.2 times as long as remaining antennomeres together. Apex of aedeagus apically evenly narrowed and dorsoventrally compressed...................................................................................................................... M. lindulana Fabrizi & Ahrens, 2014 – Metatibia ratio width/length: 1/3.2. Antennal club 1.5 times as long as remaining antennomeres together. Apex of aedeagus apically widened (lateral view), with a median hook ventrally..................................................................................................................................... M. dubia (Arrow, 1916) 5 Basis of clypeus dull......................................................................................................................... 6 – Basis of clypeus shiny...................................................................................................................... 9 6. Metatibia wide (ratio width/length: 1/2.4–2.6). Metacoxal apophysis with very dense, evenly short, scale-like setae.................................................................................................................................. 7 – Metatibia slender (ratio width/length: 1/3.0). Metacoxal apophysis without scale-like setae.Aedeagus with a single narrowly extended tube............................. M. schintlmeisteri Fabrizi & Ahrens, 2014 7. Body size smaller 10 mm (ca 8.8 mm)......................................... M. woodi Fabrizi & Ahrens, 2014 – Body size larger 10 mm (ca 11.8 mm).............................................................................................. 8 8. Aedeagus apically deeply bifurcate..................................... M. kuruwitana Fabrizi & Ahrens, 2014 – Aedeagus apically not incised.................................................................... M. coxalis (Moser, 1915) 9. Distal operculum of aedeagus enlarged apically............................................................................ 10 – Apex of aedeagus almost evenly narrowed, distal operculum of aedeagus not significantly enlarged apically.............................................................................................................................................11 10. Aedeagus ventrally with a large convex elevation at middle. Distal operculum of aedeagus moderately large, not half as wide as width of aedeagus......................................... M. fistulosa (Brenske, 1898) – Aedeagus ventrally without a large convex elevation at middle. Distal operculum of aedeagus large, as wide as width of aedeagus.................................................. M. poyagana Fabrizi & Ahrens, 2014 11. Aedeagus ventrally with a large convex elevation at middle............................................................................................................................................................. M. badullana Fabrizi & Ahrens, 2014 – Aedeagus ventrally without a large convex elevation at middle.................................................... 12 12. Aedeagus strongly compressed at middle, its distal part strongly narrowed towards apex (lateral view) and distinctly narrowed a quarter before apex (dorsal view)............................................................................................................................................................ M. weligamana (Brenske, 1900) – Aedeagus moderately compressed at middle, its distal part less narrowed towards apex (lateral view) and less narrowed a quarter before apex (dorsal view)................................... M. deenstana sp. nov. 13. Elytra with a longitudinal row of widely separated impunctate spots each bearing at centre a coarse puncture with an erect seta............................................................................................................. 14 – Elytra without widely separated impunctate spots bearing at centre a coarse puncture with an erect seta.................................................................................................................................................. 17 14. Body shape more elongate. Distal part of aedeagus short and wide (lateral view)........................ 15 – Body shape oval. Distal part of aedeagus longer and narrower (lateral view)............................................................................................................................................................. M. hiyarensis sp. nov. 15. Aedeagus with a ventral hook......................................................................................................... 16 – Aedeagus without a ventral hook.............................................................. M. karunaratnae sp. nov. 16. Pronotum with double pilosity composed of short adjacent and long erect setae. Aedeagus, with a sharp ventral hook before the middle................................... M. anderssoni Fabrizi & Ahrens, 2014 – Pronotum with simple pilosity composed of short adjacent setae. Aedeagus, with a sharp ventral hook before the apex................................................................. M. romanoi Fabrizi & Ahrens, 2014 17. Pronotum and elytra with fine and significantly larger punctures.................................................. 18 – Pronotum and elytra only with fine, dense punctures. Elytra unicoloured..................................... 22 18. Aedeagus simple, without lobes or branches. Elytral intervals with dark stripes......................................................................................................................... M. galdaththana Ranasinghe et al., 2020 – Aedeagus with apical or lateral lobes or branches.......................................................................... 19 19. Apical margin of metacoxa without setae....................................................................................... 20 – Apical margin of metacoxa with dense short setae......................................................................... 21 20 Median apophysis of metacoxa with dense, evenly short setae. Ventral metatibial spur slightly longer than the basal metatarsomere...................................................... M. brincki Fabrizi & Ahrens, 2014 – Median apophysis of metacoxa with moderately dense, unevenly long setae. Ventral metatibial spur almost as long as the two basal metatarsomeres combined .......... M. heveli Fabrizi & Ahrens, 2014 21. Ventral metatibial spur almost straight and not elongated. Median apophysis of metacoxa with moderately dense and unevenly short setae ................. M. uggalkaltotensis Fabrizi & Ahrens, 2014 – Ventral metatibial spur strongly curved and almost as long as the three basal metatarsomeres combined. Median apophysis of metacoxa with very dense and evenly short setae........................................................................................................................ M. diyalumana Fabrizi & Ahrens, 2014 22. Apical margin of metacoxa with dense, short setae........................................................................ 23 – Apical margin of metacoxa without setae....................................................................................... 24 23. Colour reddish brown, body smaller than 7.5 mm.............. M. tricuspidata Fabrizi & Ahrens, 2014 – Colour dark reddish brown, body larger than 8.0 mm........ M. nilaveliensis Fabrizi & Ahrens, 2014 24. Body smaller than 7 mm................................................................................................................. 25 – Body larger than 7 mm................................................................................................................... 28 25. Antennal club as long as remaining antennomeres combined........................................................ 26 – Antennal club slightly longer than the remaining antennomeres combined......................................................................................................................................... M. bisornata Fabrizi & Ahrens, 2014 26. Dorsal distal lobe of aedeagus simple............................................................................................ 27 – Dorsal distal lobe of aedeagus with a long lobiform extension at the base ............ M. windy sp. nov. 27. Ventral distal lobe of aedeagus (i.e., including the fused parameres) extremely extended medially exceeding almost the maximum width of phallobase (dorsal view) ..... M. pubescens (Arrow, 1916) – Ventral distal lobe of aedeagus (i.e., including the fused parameres) not extended mesally but constant in width over its entire length and being much narrower than the phallobase (dorsal view)....................................................................................................................... M. dambullana sp. nov. 28. Aedeagus with at least two apical processes.................................................................................. 29 – Aedeagus with a single principal apical process..................... M. yalaensis Fabrizi & Ahrens, 2014 29. Aedeagus with at two apical processes. 3 rd abdominal sternite without median tubercle.............. 31 – Aedeagus with at three apical processes......................................................................................... 30 30. 3 rd abdominal sternite with a median tubercle. Right distal lobe of aedeagus split before apex.... 34 – 3 rd abdominal sternite without a median tubercle. Right distal lobe of aedeagus not split before apex........................................................................................................................... M. kishi sp. nov. 31. Median apophysis of metacoxa with dense, evenly short setae...................................................... 32 – Median apophysis of metacoxa with moderately dense, unevenly long setae............................................................................................................................... M. mavilluensis Fabrizi & Ahrens, 2014 32. Aedeagus in lateral view strongly enlarged ventrally, with a marsupium-like excavation......................................................................................................................... M. flinti Fabrizi & Ahrens, 2014 – Aedeagus in lateral not enlarged ventrally..................................................................................... 33 33. Right lateral process of aedeagus convex at the tip.............. M. kandyensis Fabrizi & Ahrens, 2014 – Right lateral process of aedeagus acute at the tip....................... M. hastata Fabrizi & Ahrens, 2014 34. Setae on disc of pronotum almost missing. Right distal lobe of aedeagus shorter, shorter than the rest of the basal part of the phallobase, strongly narrowed towards apex ................................................................................................................................................ M. cervicornis Ranasinghe et al., 2020 – Setae on disc of pronotum dense. Right distal lobe of aedeagus extremely long, as long as the rest of the basal part of the phallobase, equal in width over its entire length................... M. haniel sp. nov., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on pages 88-91, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124.","Ranasinghe S., Eberle J., Benjamin S. P. & Ahrens D. 2020. New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). European Journal of Taxonomy 621: 1 - 20. https: // doi. org / 10.5852 / ejt. 2020.621"]}

Maladera anderssoni Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR0949, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X- SR0950, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33082ºN, 80.86203ºE; 1108m; 17-X- 2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0704, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33097ºN, 80.85934ºE; 1190m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0705, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33097ºN, 80.85934ºE; 1190m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂;“ X-SR0706, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33097ºN, 80.85934ºE; 1190m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0707, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33097ºN, 80.85934ºE; 1190m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0709, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33097ºN, 80.85934ºE; 1190m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0710, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33097ºN, 80.85934ºE; 1190m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0711, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33097ºN, 80.85934ºE; 1190m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0922, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33159ºN, 80.86110ºE; 1139m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0923, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33159ºN, 80.86110ºE; 1139m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0559, Sri Lanka, Kandy District, Deanston, Knuckles South; 7.33501ºN, 80.85966ºE; 1171m; 18-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 91, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Selaserica impexa Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR0855, Sri Lanka, Galle District, Hiyare FR; 6.0587ºN, 80.3153ºE; 117m; 22- X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X-SR0856, Sri Lanka, Galle District, Kanneliya FR; 6.2509ºN, 80.340130ºE; 56m; 26-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0857, Sri Lanka, Galle District, Kanneliya FR; 6.25013ºN, 80.338ºE; 42m; 26-X-2019; Eberle, Bohacz & Ranasinghe leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 97, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Here, we present the results of our field survey in Sri Lanka and describe ten new species of Sericini: Selaserica fabriziae sp. nov., Sel. sororinitida sp. nov., Neoserica pophami sp. nov., Maladera haniel sp. nov., M. kishi sp. nov., M. windy sp. nov., M. karunaratnae sp. nov., M. hiyarensis sp. nov., M. dambullana sp. nov., and M. deenstana sp. nov. All seven of the newly described species of Maladera belong to the M. fistulosa species group, which is an endemic radiation on the island that is characterized by entirely reduced or fused parameres. An updated key to the Maladera fistulosa group is provided. Further, new locality records for 23 already known species are given. The genitalia and habitus of all new species are illustrated, the distribution of the new species is shown with maps.

Maladera iuga Fabrizi & Ahrens, 2014 Material examined SRI LANKA • 1 ♂; “ X-SR0543, Sri Lanka, Kandy District, Deenston, Knuckles South; 7.35771ºN, 80.85006ºE; 980m; 17-X-2019; Eberle, Bohacz & Ranasinghe leg.; Light sheet ”; ZFMK • 1 ♂; “ X- SR2000, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.58507ºN, 80.74714ºE; 399m; 24-VI- 2020; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR2022, Sri Lanka, Matale District, Dambulla, NIFS Arboretum; 7.5845ºN, 80.7472ºE; 396m; 25-VI-2020; Ranasinghe, Athukorala & Jayatissa leg.; Black light ”; ZFMK.

Maladera windy sp. nov. urn:lsid:zoobank.org:act: 0FAD16C9-9434-4A0F-9D11-42A464F3F2BB Figs 4I–L, 7F, 8E Diagnosis The new species is very similar to M. pubescens (Arrow, 1916) and M. dambullana sp. nov. as well as M. bisornata Fabrizi & Ahrens, 2014. Maladera windy sp. nov. differs from the first two by the distal aedeagal lobe (i.e., the fused parameres) having a basal lateral lobe with grater-like surface, the ventral process of phallobase is shorter, less pointed and not mesally extended; from M. bisornata, M. windy sp. nov. differs by the longer and slightly reflexed distal aedeagal lobe (i.e., the fused parameres). Etymology The name of the new species is derived from ‘Windy Holiday Bungalow’, where the research group stayed during the second expedition in Knuckles region (noun in apposition). Type material Holotype SRI LANKA • ♂; “ X-SR0769, Sri Lanka, Deenston, Knuckles South, 7.35771ºN, 80.85006ºE, 980m, 17-X-2019, Eberle, Bohacz & Ranasinghe, Light sheet ”; ZFMK. Paratypes SRI LANKA • 1 ♂; “ X-SR0757, Sri Lanka, Deenston, Knuckles South, 7.35771ºN, 80.85006ºE, 980m, 17-X-2019, Eberle, Bohacz & Ranasinghe, Light sheet ”; ZFMK • 1 ♂; “ X-SR0790, Sri Lanka, Deenston, Knuckles South, 7.35771ºN, 80.85006ºE, 980m, 17-X-2019, Eberle, Bohacz & Ranasinghe, Light sheet ”; ZFMK • 1 ♂; “ X-SR0580, Sri Lanka, Deenston, Knuckles South, 7.35771ºN, 80.85006ºE, 980m, 16-X-2019, Eberle, Bohacz & Ranasinghe, Light sheet ”; ZFMK • 1 ♀; “ X-SR0572, Sri Lanka, Deenston, Knuckles South, 7.35771ºN, 80.85006ºE, 980m, 16-X-2019, Eberle, Bohacz & Ranasinghe, Light sheet ”; ZFMK • 1 ♀; “ X-SR0574, Sri Lanka, Deenston, Knuckles South, 7.35771ºN, 80.85006ºE, 980m, 16-X-2019, Eberle, Bohacz & Ranasinghe, Light sheet ”; ZFMK • 1 ♀; “ X-SR0575, Sri Lanka, Deenston, Knuckles South, 7.35771ºN, 80.85006ºE, 980m, 16-X-2019, Eberle, Bohacz & Ranasinghe, Light sheet ”; ZFMK. Description MEASUREMENTS. Length: 6.0 mm, length of elytra: 4.1 mm, width: 3.4 mm. HABITUS (Fig. 4L) AND COLOURATION. Body short oval, yellowish brown, antenna yellow, dorsal surface shiny, densely finely setose. HEAD. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to widely rounded anterior angles, lateral border and ocular canthus producing an indistinct blunt angle, margins moderately reflexed, anteriorly distinctly sinuate medially; surface flat, finely and densely punctate, distance between punctures smaller than their diameter, with numerous erect setae in larger punctures; frontoclypeal suture fine and angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and narrow, finely and densely punctate, with a single short terminal seta. Frons with fine, dense punctures, with dense short and sparse long erect setae. Eyes large, ratio of diameter/interocular width: 0.85. Antenna yellow, with ten antennomeres; club with three antennomeres, as long as remaining antennomeres combined. Mentum elevated and anteriorly flattened. PRONOTUM. Moderately wide, widest at base, lateral margins weakly convex and evenly narrowed anteriorly, anterior angles strongly produced and sharp, anterior marginal line fine and complete, anterior margin weakly produced medially; surface finely and densely punctate, with dense moderately long setae being bent posteriorly on entire disc and with a few sparse longer setae being directed anteriorly; anterior and lateral borders setose, basal margin without marginal line; hypomeron ventrobasally carinate and slightly produced ventrally. Scutellum short and wide, triangular, with fine and dense punctures, with fine and dense adjacent setae. ELYTRA. Short oval, widest shortly behind middle, striae indistinctly impressed, finely and densely punctate, intervals flat, with fine, very dense punctures, with numerous fine setae similar to those of the pronotum and a few sparser ones being longer and erect or directed anteriorly, in particular on lateral intervals; epipleural edge fine, ending at the weakly convex external apical angle of elytra, epipleura densely setose; apical border narrowly membranous, apex covered with short microtrichomes. VENTRAL SURFACE. Moderately shiny, thorax and metacoxa with large and dense punctures, densely setose, metacoxa glabrous except for numerous long setae laterally; each abdominal sternite, in addition to evenly distributed fine and dense punctures bearing each a fine seta, with a distinct transverse row of coarse punctures each bearing a long and more robust seta, penultimate sternite apically with a shiny smooth chitinous border, which is a quarter as long as the sternite. Mesosternum between mesocoxae as wide as mesofemur, with a semi-circular ridge bearing robust setae. Ratio of length of metepisternum/ metacoxa: 1/1.9. Pygidium moderately convex, finely and very densely punctate, without smooth midline, punctures with short and moderately dense, adjacent setae, along the apical margin with a few long erect setae. LEGS. Short and wide; femur with two longitudinal rows of setae, finely and densely punctate, densely setose; metafemur shiny, anterior edge acute, lacking an adjacent serrated line, ventral surface densely punctate and setose, posterior ventral margin straight, only little widened in apical half and very indistinctly serrate apically, dorsally not serrated, glabrous. Metatibia short, widest at middle, posteriorly slightly narrowed, ratio width/length: 1/2.6, dorsally sharply carinate, with two groups of spines, basal one shortly behind middle, apical one at four fifths of metatibial length, basally beside dorsal margin with two single punctures with serrated margins, each bearing single spines and beside them a longitudinal serrated line; lateral face almost flat, with dense, fine punctures and with minute setae; ventral margin with five strong spines equidistant from each other, medial face smooth, apex interiorly near tarsal articulation shallowly truncate. Meso- and metatarsomeres finely and densely punctate and setose dorsally, ventrally with sparse, short setae; metatarsomeres ventrally with a strongly serrated ridge, beside which is a strong longitudinal carina; first metatarsomere little shorter than following two tarsomeres combined and a little longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth. AEDEAGUS. Fig. 4I–K. Variation Length: 6.0– 6.4 mm, length of elytra: 4.0– 4.6 mm, width: 3.2–3.4 mm. Female Length: 6.5–6.8 mm, length of elytra: 4.6–5.0 mm, width: 3.6–3.7 mm. Eyes slightly smaller than in male; antennal club little shorter than the remaining antennomeres combined. Distribution See Fig. 7F., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on pages 75-77, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468, {"references":["Fabrizi S. & Ahrens D. 2014. A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, Supplements 61: 1 - 124."]}

Maladera weligamana (Brenske, 1900) Material examined SRI LANKA • 1 ♂; “ X-SR0815, Sri Lanka, Nuwara Eliya District, Hakgala SNR, Seetha Eliya; 6.9307ºN, 80.8134ºE; 1773m; 20-XI-2019; Ranasinghe & Athukorala leg.; Black light ”; ZFMK • 1 ♂; “ X-SR0835, Sri Lanka, Nuwara Eliya District, Hakgala SNR, Seetha Eliya; 6.93045ºN, 80.81356ºE; 1789m; 20-XI-2019; Ranasinghe & Athukorala leg.; Black light ”; ZFMK., Published as part of Ranasinghe, Sasanka, Eberle, Jonas, Athukorala, Namal, Benjamin, Suresh P. & Ahrens, Dirk, 2022, New species of Sericini from Sri Lanka (Coleoptera, Scarabaeidae). Part II, pp. 57-101 in European Journal of Taxonomy 821 (1) on page 96, DOI: 10.5852/ejt.2022.821.1799, http://zenodo.org/record/6595468

Carrhotus albosetosus sp. nov. urn:lsid:zoobank.org:act: 8E87E441-3B3F-43D8-B5B3-9B647FC853B1 Figs 2A–B, 3A–C, 4A–B, 17 Diagnosis Females of C. albosetosus sp. nov. can be separated from other congeners by the white hairs on the legs (Fig. 2A–B). The epigyne of this species is close to that of C. silanthi; however, it differs by the absence of prominent CD and FD as well as in the position of the opening of the accessory glands, which open at the 2’o clock position at the anterior region of the lateral walls of the spermathecae (Fig. 4A–B). Etymology The specific epithet is derived from a combination of Latin words; ‘albi’ meaning ‘white’ and ‘comus’ meaning ‘legs’, loosely referring to the white legs/white leg setae. Material examined Holotype SRI LANKA • ♀; Western Province, Gampaha District, Pillikutuwa FR; 07°03′52.4″ N, 80°03′04″ E; 69 m a.s.l; 28 Sep. 2016; N. Kanesharatnam. leg.; beating; NIFS_SAL_911. Other material SRI LANKA • 1 ♀; Central Province, Matale District, NIFS Arboretum; 07°51′65″ N, 80°40′53″ E; 184 m a.s.l; 11–13 Oct. 2019; S.P. Benjamin et al. leg.; beating; NIFS_SAL_1262. Description Female MEASUREMENTS. TL 4.9, PL 1.85, PW at PLEs 1.68, AL 2.1, AW 1.06. Eye field: diameter of AME 0.48; ALE 0.24; PME 0.22; PLE 0.08; PME–PME 1.4; PLE–PLE 1.44; ALE–PME 0.72; ALE–PLE 0.34. Leg I: Tr 1.1, Fm 0.9, Pt 0.8, Tb 0.5, Mt 0.4; Leg II: Tr 0.9, Fm 0.46, Pt 0.8, Tb 0.6, Mt 0.5; Leg III: Tr 1.5, Fm 0.42, Pt 0.82, Tb 0.34, Mt 0.42; Leg IV: Tr 1.6, Fm 0.5, Pt 1.02, Tb 0.6, Mt 0.34. COLOR AND BODY. Live spider with clypeal region brown; covered with iridescent hairs providing a metallic sheen to green colour; lateral margins of carapace lined by with thin line of white hairs; posterior region black (Fig. 2A–B). ALEs and PLEs surrounded by blackish orbital setae. Chelicerae dark brown with curved outer margins and excavated inner margins: two promarginal and one retromarginal teeth. Sternum oval, brownish; labium and maxillae reddish-brown, with paler outer margins. Abdomen ovoid, densely covered with rusty brown hairs; anterior margin covered with a thin line of white hairs; pattern present on black background with pair of two white spots anteriorly and pairs of transverse stripes and two pairs of white spots following posteriorly. Mid-dorsum covered with scales of metallic sheen; venter yellowish brown, with a thin dark brown scattered pattern on median region. Spinnerets blackish, covered with a patch of white hairs dorsally (Fig. 3A–B). EPIGYNUM. Epigyne with a pair of simple copulatory openings and short copulatory ducts diverge to join semioval-shaped spermathecae. Accessory glands open at 2’o clock position on anterior region of lateral walls of spermathecae (Figs 3C, 4A–B). Distribution and habitat Known only from the two localities mentioned above. The spiders were collected by beating foliage up to a height of 2 m. This species occurs in the lowland secondary rainforest of the dry zone and submontane forest of the central highlands of Sri Lanka (Fig. 17)., Published as part of Satkunanathan, Abira & Benjamin, Suresh P., 2022, Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae), pp. 78-110 in European Journal of Taxonomy 817 (1) on pages 85-86, DOI: 10.5852/ejt.2022.817.1765, http://zenodo.org/record/6518305

Carrhotus silanthi Caleb, 2020 Figs 10A–F, 11A–H, 12A–D, 17 Carrhotus silanthi Caleb in Caleb et al., 2020: 58, Figs 33–56. Diagnosis Males of C. silanthi can be separated from other congeners by having the RTA markedly hook-shaped, slanted ventrally, and the distally pointed acuminate embolus (Fig. 12A–B). Females can be readily separated from those of C. albosetosus sp. nov. by the prominent ML and the position of the opening of the accessory glands on the internal spermathecal walls, close to the base of the copulatory ducts (Figs 11G, 12C–D). Type material Carrhotus silanthi Caleb in Caleb et al., 2020: 58, Figs 33–56 (male holotype and female paratype from Madras Christian College, Tambaram, Chennai, India, deposited in NZC-ZSI, Kolkata (NZC-ZSI 6939/18, NZC-ZSI 6940/18)). Not examined. Material examined SRI LANKA – Uva Province • 1 ♂, 1 ♀; Monaragala District, Nilgala FR; 07°11′08″ N, 81°24′24″ E; 122 m a.s.l; 12 Jul. 2017; N. Athukorala and I.S. IIeperuma Arachchi leg.; beating; NIFS_SAL_1042, SAL_1043. – North Central Province • 1 ♂, 1 ♀; Anuradhapura District, Mihintale Sanctuary; 08°20′59″ N, 80°30′20″ E; 119 m a.s.l; 14 Jun. 2016; N. Athukorala et al. leg.; beating; NIFS_SAL_824, SAL_825 • 1 ♂; Allepothana, Kok-ebe FR; 08°26′58″ N, 80°46′39′ E; 88 m a.s.l; 24 Apr. 2017; N. Kanesharatnam leg.; beating; NIFS_SAL_1032. – Northern Province • 2 ♂♂, 1 ♀; Mannar District, Giant’s Tank Sanctuary (Site 1); 27 m a.s.l; 4 Apr. 2018; S.P. Benjamin et al. leg.; beating; NIFS_ SAL_1137 to SAL_1139 • 1 ♀; same collection data as for preceding; NIFS_SAL_1162. – Southern Province • 1 ♂; Hambantota District, Ussangoda NP; 06°05′43″ N, 80°59′15″ E; 22 Nov. 2017; S.P. Benjamin et al. leg.; beating; NIFS_SAL_1116. Description (based on material from Sri Lanka) Male MEASUREMENTS. TL 4.8, PL 2.16, PW at PLEs 1.786,AL 2.19, AW 1.44. Eye field: diameter of AME 0.44; ALE 0.26; PME 0.12; PLE 0.24; PME–PME 1.68; PLE–PLE 1.44; ALE–PME 0.36; ALE–PLE 0.6. Leg I: Tr 1.2, Fm 0.82, Pt 0.78, Tb 1.22, Mt 0.62; Leg II: Tr 1.48, Fm 0.42, Pt 0.62, Tb 1.02, Mt 0.56; Leg III: Tr 1, Fm 0.7, Pt 0.72, Tb 1.06, Mt 0.5; Leg IV: Tr 0.84, Fm 0.6, Pt 1, Tb 1.02, Mt 0.62. COLOR AND BODY. In live spiders, both sexes look similar in general body colour pattern and are clothed with iridescent hairs. Colour varies from metallic sheen to bronze-green as shown in Fig. 10A–F. Carapace: prosoma reddish-brown covered with iridescent hairs providing a metallic sheen; lateral margins of carapace lined by broad patches of white hairs; posterior region black. Hairs sparse. Sternum oval, brownish; labium and maxillae dark brown, with paler outer margins. Clypeal region brownish; eyes surrounded by yellowish orbital setae. Chelicerae dark brown with curved outer margins and excavated inner margins: two promarginal and one retromarginal teeth. Abdomen ovoid, densely covered with rusty brown hairs; anterior margin covered by white scales; pattern present on black background with pair of white spots anteriorly and pairs of transverse stripes following posteriorly (Fig. 11A–B). Middorsum covered with scales of metallic sheen; venter yellowish, with a broad dark brown median region. Spinnerets brownish, covered with a patch of white hairs dorsally. Leg I robust; femora I-IV dark brown dorsally; patellae and tibiae dark brown; tarsi and metatarsi of all legs dark brown. PALP. Dark brown; embolus short and thick with a blunt tip and embolus tip directed in clockwise path, its base separated from tegulum; bulbus with PLP; RTA markedly hook-shaped bent ventrally and angle between RTA and tibia about 45º (Figs 11E–F, 12A–B). Female MEASUREMENTS. TL 5.5, PL 2.04, PW at PLEs 1.95, AL 2.64, AW 2.16. Eye field: diameter of AME 0.48; ALE 0.26; PME 0.12; PLE 0.24; PME–PME 1.5; PLE–PLE 1.46; ALE–PME 0.44; ALE–PLE 0.78. Leg I: Tr 1, Fm 0.92, Pt 0.94, Tb 0.56, Mt 0.38; Leg II: Tr 1.2, Fm 0.78, Pt 1.1, Tb 0.7, Mt 0.46; Leg III: Tr 1.3, Fm 0.8, Pt 1.02, Tb 0.6, Mt 0.42; Leg IV: Tr 1, Fm 0.74, Pt 1.1, Tb 0.4, Mt 0.32. COLOR AND BODY. Colouration pattern as in male, but differs as follows: AMEs surrounded by yellow orbital setae; clypeus covered with white hairs; carapace with white hairs making an arc behind PLEs. Abdomen ovoid, densely covered with rusty brown hairs; anterior margin covered by white scales; pattern present on black background with pair of white spots anteriorly and pairs of transverse stripes following posteriorly (Fig. 11C–D). Mid-dorsum covered with scales of metallic sheen; venter yellowish, with a broad dark brown median region. Spinnerets brownish, covered with a patch of white hairs dorsally. Legs with white border on distal margin of femur I–IV; other segments covered with sparse white hairs and black annulations alternatively. EPIGYNUM. Epigyne with a pair of simple copulatory openings placed in small oval, yellowish depressions; copulatory ducts short, diverge to join stomach-shaped spermathecae.Accessory glands open on internal spermathecal walls close to base of copulatory ducts (Fig. 12C–D). Intraspecific variation The palpal conformation of NIFS _SAL_1116 closely resembles that of C. silanthi. However, it differs by having the PLP shorter and less rounded (Fig. 11F) vs oval-shaped PLP in C. silanthi (Fig. 11E), and a short, cone-shaped embolus (Fig. 11F) in NIFS _SAL_1116 vs a thinner embolus in C. silanthi (Fig. 11E). Distribution and habitat This species occurs in the lowland secondary rainforests of the dry and wet zones of Sri Lanka including the Ussangoda dry zone National Park in Hambantota District in the Southern Province (Fig. 17). Specimens were collected by beating vegetation up to a height of 1– 2 m., Published as part of Satkunanathan, Abira & Benjamin, Suresh P., 2022, Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae), pp. 78-110 in European Journal of Taxonomy 817 (1) on pages 95-98, DOI: 10.5852/ejt.2022.817.1765, http://zenodo.org/record/6518305, {"references":["Caleb J. T. D., Bera C. & Acharya S. 2020. New species and synonymies in the genus Carrhotus Thorell, 1891 from India (Aranei: Salticidae: Salticini). Arthropoda Selecta 29 (1): 51 - 66. https: // doi. org / 10.15298 / arthsel. 29.1.04"]}

Carrhotus taprobanicus Simon, 1902 Figs 2E–F, 13A–G, 14A–D, 17 Carrhotus taprobanicus Simon, 1902: 394. Diagnosis Males of C. taprobanicus can be separated from those of C. viduus by their body colour pattern with no longitudinal stripes on the carapace and abdomen (present in C. viduus) (Fig. 15A, C). Further, the cheliceral front part is more wrinkled in C. viduus (Fig. 13B). The palpal conformation of C. taprobanicus closely resembles that of C. viduus; however, it can be distinguished by the shorter embolus and its rounded base (Fig. 13E). Further, the RTA is more slanted ventrally in C. taprobanicus (Fig. 14A–B). Females can be separated from those of C. viduus by the narrower ML and stouter CD and FD (Fig. 14C–D). Type material Carrhotus taprobanicus Simon, 1902; ♂ ♀ syntypes from Colombo and Kandy, Sri Lanka; should be in MNHN, not examined. Material examined SRI LANKA – Central Province • 1 ♂; Kandy District, Knuckles Range, along Dothalugala Nature Trail; 07°20′19″ N, 80°51′3″ E; 1202 m a.s.l; 3 Mar. 2018; S.P. Benjamin et al. leg.; beating; NIFS_ SAL_1143 • 1 ♂; same collection data as for preceding; 3 May 2018; NIFS_SAL_1155 • 1 ♀; same collection data as for preceding; SAL_1156 • 6 ♀♀; same collection data as for preceding; 17–18 Oct. 2019; NIFS_SAL_1241 to SAL_1246 • 1 ♂; same collection data as for preceding; NIFS_SAL_1248 • 1 ♀; same collection data as for preceding; NIFS_SAL_1275 • 1 ♂; same collection data as for preceding but 17–18 Nov. 2019; NIFS_SAL_1252 • 2 ♀♀; same collection data as for preceding but 18 Jun. 2020; NIFS_SAL_1308, SAL_1309 • 1 ♀; Gomaraya; 07°23′10″ N, 80°44′30″ E; 600 m a.s.l; 2 Feb. 2010; S. Batuwita and P.M.H. Sandamali et al. leg.; hand collection; NIFS_SAL_660 • 1 ♀; Nuwara Eliya District, Victoria, Randenigala, Rantembe Sanctuary; 07°08′42″ N, 80°51′28″ E; 436 m a.s.l; 22 Sep. 2018; N.P. Athukorala et al. leg.; beating; NIFS_SAL_1273. – Uva Province • 1 ♂; Badulla District, Ohiya; 06°50′32″ N, 80°53′05″ E; 26 May 2012; S.P. Benjamin et al. leg.; beating; NIFS_SAL_279. – Western Province • 1 ♂; Panadura, Mahabellana, along Bolgoda South Lake; 06°42′48″ N, 79°54′09″ E; Jul. 2008; S.P. Benjamin et al. leg.; NIFS_SAL_694. Description Male MEASUREMENTS. TL 4.7, PL 2.1, PW at PLEs 1.8, AL 1.95, AW 1.32. Eye field: diameter of AME 0.4; ALE 0.24; PME 0.2; PLE 0.08; PME–PME 1.06; PLE–PLE 1.1; ALE–PME 0.5; ALE–PLE 0.3. Leg I: Tr 1.3, Fm 0.9, Pt 1.4, Tb 0.7, Mt 0.44; Leg II: Tr 1, Fm 0.8, Pt 0.84, Tb 0.4, Mt 0.36; Leg III: Tr 1.04, Fm 0.86, Pt 0.8, Tb 0.7, Mt 0.6; Leg IV: Tr 0.7, Fm 0.8, Pt 0.7, Tb 0.72, Mt 0.6. COLOR AND BODY. Live spider with carapace blackish-brown, covered with iridescent hairs providing a metallic sheen; lateral margins of carapace lined by broad patches of white hairs; posterior region black (Fig. 2E–F). Clypeal region blackish; eyes surrounded by yellowish orbital setae. Chelicerae brown with curved outer margins and excavated inner margins: two promarginal and one retromarginal teeth. Sternum pentagonal, blackish brown; labium and maxillae yellowish brown, with paler outer margins. Abdomen ovoid, densely covered with black hairs; lateral sides of anterior margin with two white longitudinal lines; pattern present on black background with three pairs of white transverse stripes lateral margins of mid-dorsum. Mid-dorsum covered with scales of metallic sheen; venter yellowish, with a broad black median region covered with two broad whitish longitudinal bands. Spinnerets brownish, covered with a patch of white hairs dorsally (Fig. 13A–B). Leg I robust; femora I–IV dark brown dorsally; other segments covered with sparse black hairs and black annulations. PALP. Dark brown; embolus thinner and cone-shaped, its base separated from tegulum. PLP shorter and similar to that of C. viduus, angle between RTA and tibia about 45º vs about early 20º in C. viduus (Figs 13E–F, 14A–B). Female MEASUREMENTS. TL 4.9, PL 2.07, PW at PLEs 1.68, AL 2.1, AW 1.06. Eye field: diameter of AME 0.48; ALE 0.24; PME 0.22; PLE 0.08; PME–PME 1.4; PLE–PLE 1.44; ALE–PME 0.72; ALE–PLE 0.34. Leg I: Tr 1.1, Fm 0.9, Pt 0.8, Tb 0.5, Mt 0.4; Leg II: Tr 0.9, Fm 0.46, Pt 0.8, Tb 0.6, Mt 0.5; Leg III: Tr 1.5, Fm 0.42, Pt 0.82, Tb 0.34, Mt 0.42; Leg IV: Tr 1.6, Fm 0.5, Pt 1.02, Tb 0.6, Mt 0.34. COLOR AND BODY. Abdomen ovoid, sparsely covered with black hairs; lateral sides of anterior margin with four pairs of white herringbone-shaped pattern present on brown background with two pairs of white spots in mid-dorsum. ALEs surrounded by blackish orbital setae; clypeus covered with white hairs (Fig. 13C–D). EPIGYNUM. Epigyne with a pair of copulatory openings, long copulatory ducts move anteriorly and almost sub-parallel to elongated, pear-shaped spermathecae. Accessory glands open at 2 o’clock position of mid-region of lateral walls of spermathecae (Figs 13G, 14C–D). Resembles the epigynes of C. viduus but differs in relative proportions of spermathecae, rather kidney-shaped in C. taprobanicus (Fig. 14C–D), PEB less curved inwards and margins of CO less sclerotized than in C. viduus (Fig. 13G). Remarks Our identification is based on Simon’s (1902) description of the male and female syntypes. Though we were unable to examine the types of C. taprobanicus, the description by Simon (1902) was clear enough for an unambiguous identification of the species, which we translate here to English: “ ♂. Length 6.5 mm. Cephalothorax darkened upper part, bronze-colour-haired, white hair belts at the thoracic margins, short clypeus and yellowish-red hairs around the eyes. Chelicerae long, planar and hairless with bronze and leathery transverse folds; curved outer margins and excavated inner margins: promarginal and one retromarginal teeth. Sternum; black, white-haired. Abdomen shorter and ovalshaped. Venter greyish and covered on both sides with white margins. Legs I, II darkened, tarsi lightened, metatarsi, lower tibia very slightly black-haired, patella inwardly white-haired. Legs: coxa, trochanter, metatarsi darker, metatarsi dark annulated. ♀. Length 7.8 mm. Cephalothorax black, bronze-colour-haired, reddish-white hair belts at the thoracic margins and around the eyes. The males can be distinguished from those of C. viduus (C.L. Koch, 1846) by their body colour pattern with no longitudinal stripes on the carapace and abdomen and the cheliceral front part is more wrinkled in C. viduus ”. Distribution and habitat This species occurs in the montane and submontane forests of the central highlands of Sri Lanka (Fig. 17). Specimens were collected by beating vegetation up to a height of 1– 2 m., Published as part of Satkunanathan, Abira & Benjamin, Suresh P., 2022, Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae), pp. 78-110 in European Journal of Taxonomy 817 (1) on pages 99-102, DOI: 10.5852/ejt.2022.817.1765, http://zenodo.org/record/6518305, {"references":["Simon E. 1902. Etudes arachnologiques. 32 e Memoire. LI. Descriptions d'especes nouvelles de la famille des Salticidae (suite). Annales de la Societe entomologique de France 71: 389 - 421. Available from https: // www. biodiversitylibrary. org / page / 8251793 [accessed 25 Mar. 2022].","Koch C. L. 1846. Die Arachniden: Getreu nach der Natur abgebildet und beschrieben. Vol. 13: 1 - 234, 4334 - 68 (f. 1078 - 1271); Vol. 14: 1 - 88, 4674 - 80, (f. 1272 - 1342). J. L. Lotzbeck, Nurnberg. https: // doi. org / 10.5962 / bhl. title. 43744 [For the correct year of publication see Brignoli 1985]"]}

The Sri Lankan species of the genus Carrhotus Thorell,1891 have remained taxonomically unrevised. The present study reviews all species of the genus in the island. Here we describe and diagnose three new species: Carrhotus albosetosus sp. nov. (♀), C. atratus sp. nov. (♂♀) and C. lobatus sp. nov. (♂). Further, C. silanthi Caleb, 2020 is reported in Sri Lanka for the first time. Carrhotus taprobanicus Simon, 1902 and C. viduus (C.L. Koch, 1846) are redescribed based on material from Sri Lanka. A key to the Carrhotus species in Sri Lanka is given.

Carrhotus lobatus sp. nov. urn:lsid:zoobank.org:act: B1F383E0-259C-4888-8227-FEB0F27014D3 Figs 8A–D, 9A–B, 17 Diagnosis This species is distinguishable by the distinctive double-lobed PLP and short and hook-shaped embolus tip directed in a clockwise path (Figs 8C–D, 9A–B). Etymology The specific epithet is derived from the Latin word ‘ lobatus ’ meaning ‘lobed’, loosely referring to its distinctive proximal lobe of the tegulum. Material examined Holotype SRI LANKA • ♂; Central Province, Matale District, Gammaduwa, Knuckles Range; 07°27′57″ N, 80°46′24″ E; 900 m a.s.l.; 18 Nov. 2009; S.P. Benjamin and S. Batuwita leg.; hand collection; NIFS_ SAL_282. Other material SRI LANKA – Central Province • 1 ♂; Matale District, Knuckles range, Pitawala Pathana; 07°27′57″ N, 80°46′24″ E; 17 Mar. 2015; N.P. Athukorala leg.; hand collection; NIFS_SAL_403 • 1 ♂; Kandy District, Peradeniya; 07°14′58″ N, 80°36′43″ E; 674 m a.s.l; 22 Dec. 2017; N.P. Athukorala et al. leg.; beating; NIFS_SAL_1114. – Southern Province • 1 ♂; Galle District, Hiyare, Kombala-Kottawa FR; 06°03′53″ N, 80°18′56″ E; 110 m a.s.l.; 24–26 May 2016; N.P. Athukorala et al. leg.; beating; NIFS_ SAL_1285. Description Male MEASUREMENTS. TL 5.15, PL 2.13, PW at PLEs 1.71, AL 2.19, AW 1.71. Eye field: diameter of AME 0.46; ALE 0.28; PME 0.08; PLE 0.26; PME–PME 1.46; PLE–PLE 1.34; ALE–PME 0.24; ALE–PLE 0.6. Leg I: Tr 0.82, Fm 1.08, Pt 1.36, Tb 0.86, Mt 0.5; Leg II: Tr 0.7, Fm 0.82, Pt 0.8, Tb 0.6, Mt 0.4; Leg III: Tr 1, Fm 0.9, Pt 0.74, Tb 0.8, Mt 0.44; Leg IV: Tr 0.92, Fm 0.5, Pt 1.08, Tb 0.74, Mt 0.46. COLOR AND BODY. In ethanol, carapace brown, posterior region blackish-brown. Clypeal region brown; ALEs and PLEs surrounded by blackish orbital setae. Lateral margins of carapace slightly truncated and steep. Chelicerae brownish with curved outer margins and excavated inner margins: two promarginal and one retromarginal teeth. Sternum pentagonal, blackish brown; labium and maxillae brown. Leg I robust; femora I–IV dark brown dorsally; patellae and tibiae dark brown; tarsi and metatarsi of all legs dark brown. Abdomen oblong, sparsely covered with black hairs; anterior margin covered with long black scales with sparse white hairs making an arc; no prominent pattern present on posterior region, beige pairs of transverse lines and pair of beige spots covered with white hairs on mid dorsum; venter blackish brown with a blackish brown median region. Spinnerets blackish brown, covered with a patch of white hairs dorsally (Fig. 8A–B). PALP. Dark brown; embolus short and hook-shaped and embolus tip directed in the clockwise path; its base separated from the tegulum. The double-lobed PLP is very distinctive compared to that of the other species considered here. RTA long and angle between the RTA and tibia is about 45º (Fig. 9A–B). Distribution and habitat This species occurs in the montane and submontane forests of the central highlands of Sri Lanka with limited distribution (Fig. 17). Specimens were collected by beating vegetation up to a height of 1– 2 m., Published as part of Satkunanathan, Abira & Benjamin, Suresh P., 2022, Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae), pp. 78-110 in European Journal of Taxonomy 817 (1) on pages 92-94, DOI: 10.5852/ejt.2022.817.1765, http://zenodo.org/record/6518305

Key to the species of Carrhotus in Sri Lanka 1. Males (males of C. albosetosus sp. nov. unknown).......................................................................... 2 – Females (females of C. lobatus sp. nov. unknown).......................................................................... 6 2. PLP with two lobes (Figs 8C, 9A)........................................................... Carrhotus lobatus sp. nov. – PLP with single lobe......................................................................................................................... 3 3. Embolus straight............................................................................................................................... 4 – Embolus curved................................................................................................................................ 5 4. Embolus long, acuminate, RTA as in Figs 11G–H, 12B................ Carrhotus silanthi (Caleb, 2020) – Embolus short, acute, RTA as in Figs 13F, 14B................... Carrhotus taprobanicus (Simon, 1902) 5. Embolus tapered, RTA tapered (Fig. 7D)................................ Carrhotus viduus (C.L. Koch, 1846) – Embolus bow-shaped, RTA stout (Figs 6E, 7D)....................................... Carrhotus atratus sp. nov. 6. CD and FD prominent...................................................................................................................... 7 – CD and FD not prominent................................................................................................................ 9 7. Spermathecae boat-shaped, CD hook-shaped (Fig. 7D)........................... Carrhotus atratus sp. nov. – Spermathecae kidney-shaped, CD otherwise................................................................................... 8 8. ML prominent (Figs 15G, 16C–D).......................................... Carrhotus viduus (C.L. Koch, 1846) – ML less prominent (Figs 13G, 14C–D)............................... Carrhotus taprobanicus (Simon, 1902) 9. ML and CO as in Fig. 4A, D............................................................. Carrhotus albosetosus sp. nov. – ML and CO as in Fig. 12C–D........................................................ Carrhotus silanthi (Caleb, 2020), Published as part of Satkunanathan, Abira & Benjamin, Suresh P., 2022, Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae), pp. 78-110 in European Journal of Taxonomy 817 (1) on page 107, DOI: 10.5852/ejt.2022.817.1765, http://zenodo.org/record/6518305, {"references":["Caleb J. T. D., Bera C. & Acharya S. 2020. New species and synonymies in the genus Carrhotus Thorell, 1891 from India (Aranei: Salticidae: Salticini). Arthropoda Selecta 29 (1): 51 - 66. https: // doi. org / 10.15298 / arthsel. 29.1.04","Simon E. 1902. Etudes arachnologiques. 32 e Memoire. LI. Descriptions d'especes nouvelles de la famille des Salticidae (suite). Annales de la Societe entomologique de France 71: 389 - 421. Available from https: // www. biodiversitylibrary. org / page / 8251793 [accessed 25 Mar. 2022].","Koch C. L. 1846. Die Arachniden: Getreu nach der Natur abgebildet und beschrieben. Vol. 13: 1 - 234, 4334 - 68 (f. 1078 - 1271); Vol. 14: 1 - 88, 4674 - 80, (f. 1272 - 1342). J. L. Lotzbeck, Nurnberg. https: // doi. org / 10.5962 / bhl. title. 43744 [For the correct year of publication see Brignoli 1985]"]}

Carrhotus viduus (C.L. Koch, 1846) Figs 2C–D, 15A–G, 16A–D, 17 Plexippus viduus C.L. Koch, 1846: 104, fig. 1166. Carrhotus viduus – Thorell 1891: 142. — Andreeva et al. 1981: 103, figs 39–42. — Prószyński 1984 a: 16; 1992b: 169, fig. 7. — Caleb et al. 2020: 61, figs 57–73, 76–78, 80–82. Diagnosis Males of this species are distinguishable by the presence of white longitudinal stripes on the carapace and abdomen, as well as the relatively larger retromarginal cheliceral tooth. The palpal conformation of C. viduus closely resembles that of C. taprobanicus; however, it can be distinguished by the longer, ventrally projecting embolus and its oval base (Fig. 16A). Further, the RTA is less slanted ventrally in C. viduus (Fig. 16A–B). Females can be separated from those of C. taprobanicus by the broader ML and longer CD and FD (Fig. 16C–D). Females can be readily distinguished from those of C. silanthi by the kidney-shaped spermathecae (Figs 12C–D, 16C–D). Type material Plexippus viduus C.L. Koch, 1846: 2 ♂♂; depository unknown. Our identification is based solely on the description and illustrations of Prószyński (2009). He mentioned two possible specimens currently deposited in ZMB. However, their geographic origin is ambiguous (see Prószyński 2009). Remarks According to Sudhin et al. (2021), C. tholpettyensis Sudhin, Nafin, Caleb & Sudhikumar, 2021 differs by the absence of a pair of white longitudinal stripes found on the abdomen of male C. viduus. The mentioned minor gentilic differences could easily be attributed to intraspecific variation. Thus, we consider C. tholpettyensis as a possible junior synonymy of C. viduus. Material examined SRI LANKA – Northern Province • 1 ♂; Jaffna District, Mandaitivu; 09°36′26″ N, 79°59′5″ E; 20–22 Sep. 2016; S.P. Benjamin leg.; beating; NIFS_SAL_958 • 1 ♀; same collection data as for preceding but 26 Jan. 2021; NIFS_SAL_1442. – Central Province • 1 ♂; Kandy District, Gannoruwa FR; 07°17′16″ N, 80°35′47″ E; 575 m a.s.l; 30 Jul. 2016; N.P. Athukorala et al. leg.; beating; NIFS_SAL_888 • 1 ♂; Kandy District, NIFS premises; 600 m a.s.l.; 12 Mar. 2017; S.P. Benjamin leg.; hand collection; NIFS_SAL_1025. – North Western Province • 1 ♂; Kurunegala District, Nikaravatiya; 1–3 Nov. 2007; Ziyard Jaleel leg.; hand collection; NIFS_SAL_506. Description Male MEASUREMENTS. TL 6.85, PL 2.58, PW at PLEs 2.31,AL 1.35,AW 0.87. Eye field: diameter of AME 0.28; ALE 0.18; PME 0.1; PLE 0.14; PME–PME 0.82; PLE–PLE 0.84; ALE–PME 0.32; ALE–PLE 0.42. Leg I: Tr 1, Fm 1.3, Pt 1.6, Tb 0.8, Mt 0.6; Leg II: Tr 1.12, Fm 1.1, Pt 1.06, Tb 0.7, Mt 0.42; Leg III: Tr 1.1, Fm 1.12, Pt 1.4, Tb 0.52, Mt 0.3; Leg IV: Tr 0.76, Fm 0.52, Pt 0.82, Tb 0.84, Mt 0.52. COLOR AND BODY. Live spider with carapace robust and relatively broad, blackish-brown covered with long black hairs sparsely and two longitudinal stripes of white hairs (Fig. 2C–D). Sternum oval, brown; labium and maxillae yellowish brown, with paler outer margins. Clypeal region brownish; eyes surrounded by blackish orbital setae. Chelicerae brown with curved outer margins and excavated inner margins: two promarginal and one retromarginal teeth (Fig. 15B). Abdomen blackish brown, ovoid with two longitudinal belts of white hairs and entire surface covered with a few whitish hairs. Middorsum with four median beige spots in different shapes, covered with black scales; venter greyish with a blackish-brown median region covered with two broad whitish longitudinal bands. Spinnerets brown, covered with a patch of black hairs dorsally. Leg I robust; femora I–IV dark brown dorsally; patellae and tibiae dark brown; tarsi and metatarsi of all legs dark brown. PALP. Dark brown; embolus short and thick with blunt tip; bulbus with PLP; RTA only slightly bent ventroapically, curved and claw-like (Figs 15E–F, 16A–B). Female MEASUREMENTS. TL 5.5, PL 2.04, PW at PLEs 1.95, AL 2.64, AW 2.16. Eye field: diameter of AME 0.48; ALE 0.26; PME 0.12; PLE 0.24; PME–PME 1.5; PLE–PLE 1.46; ALE–PME 0.44; ALE–PLE 0.78. Leg I: Tr 1, Fm 0.92, Pt 0.94, Tb 0.56, Mt 0.38; Leg II: Tr 1.2, Fm 0.78, Pt 1.1, Tb 0.7, Mt 0.46; Leg III: Tr 1.3, Fm 0.8, Pt 1.02, Tb 0.6, Mt 0.42; Leg IV: Tr 1, Fm 0.74, Pt 1.1, Tb 0.4, Mt 0.32. COLOR AND BODY. Colouration pattern as in the male, but differs as follows: AMEs surrounded by yellow orbital setae; clypeus covered with white hairs; carapace with white longitudinal stripes (Fig. 15C). EPIGYNUM. Structure of epigyne of C. viduus (Figs 15G, 16C–D) resembles that of C. taprobanicus but differs in relative proportions of spermathecae, rather elongated and globular in C. viduus (Fig. 16C–D), PEB more curved inwards by forming a prominent central curve, presence of a well-sclerotized midline, margins of CO well sclerotized and shape of CO more prominently C shaped than in C. taprobanicus (Fig. 15G). Distribution and habitat This species occurs in the mangrove forests in the arid zone and lowland secondary rainforests of the dry and wet zones of Sri Lanka (Fig. 17). Specimens were collected by beating vegetation up to a height of 1– 2 m. Key to the species of Carrhotus in Sri Lanka 1. Males (males of C. albosetosus sp. nov. unknown).......................................................................... 2 – Females (females of C. lobatus sp. nov. unknown).......................................................................... 6 2. PLP with two lobes (Figs 8C, 9A)........................................................... Carrhotus lobatus sp. nov. – PLP with single lobe......................................................................................................................... 3 3. Embolus straight............................................................................................................................... 4 – Embolus curved................................................................................................................................ 5 4. Embolus long, acuminate, RTA as in Figs 11G–H, 12B................ Carrhotus silanthi (Caleb, 2020) – Embolus short, acute, RTA as in Figs 13F, 14B................... Carrhotus taprobanicus (Simon, 1902) 5. Embolus tapered, RTA tapered (Fig. 7D)................................ Carrhotus viduus (C.L. Koch, 1846) – Embolus bow-shaped, RTA stout (Figs 6E, 7D)....................................... Carrhotus atratus sp. nov. 6. CD and FD prominent...................................................................................................................... 7 – CD and FD not prominent................................................................................................................ 9 7. Spermathecae boat-shaped, CD hook-shaped (Fig. 7D)........................... Carrhotus atratus sp. nov. – Spermathecae kidney-shaped, CD otherwise................................................................................... 8 8. ML prominent (Figs 15G, 16C–D).......................................... Carrhotus viduus (C.L. Koch, 1846) – ML less prominent (Figs 13G, 14C–D)............................... Carrhotus taprobanicus (Simon, 1902) 9. ML and CO as in Fig. 4A, D............................................................. Carrhotus albosetosus sp. nov. – ML and CO as in Fig. 12C–D........................................................ Carrhotus silanthi (Caleb, 2020) Key to the species of Carrhotus in Sri Lanka 1. Males (males of C. albosetosus sp. nov. unknown).......................................................................... 2 – Females (females of C. lobatus sp. nov. unknown).......................................................................... 6 2. PLP with two lobes (Figs 8C, 9A)........................................................... Carrhotus lobatus sp. nov. – PLP with single lobe......................................................................................................................... 3 3. Embolus straight............................................................................................................................... 4 – Embolus curved................................................................................................................................ 5 4. Embolus long, acuminate, RTA as in Figs 11G–H, 12B................ Carrhotus silanthi (Caleb, 2020) – Embolus short, acute, RTA as in Figs 13F, 14B................... Carrhotus taprobanicus (Simon, 1902) 5. Embolus tapered, RTA tapered (Fig. 7D)................................ Carrhotus viduus (C.L. Koch, 1846) – Embolus bow-shaped, RTA stout (Figs 6E, 7D)....................................... Carrhotus atratus sp. nov. 6. CD and FD prominent...................................................................................................................... 7 – CD and FD not prominent................................................................................................................ 9 7. Spermathecae boat-shaped, CD hook-shaped (Fig. 7D)........................... Carrhotus atratus sp. nov. – Spermathecae kidney-shaped, CD otherwise................................................................................... 8 8. ML prominent (Figs 15G, 16C–D).......................................... Carrhotus viduus (C.L. Koch, 1846) – ML less prominent (Figs 13G, 14C–D)............................... Carrhotus taprobanicus (Simon, 1902) 9. ML and CO as in Fig. 4A, D............................................................. Carrhotus albosetosus sp. nov. – ML and CO as in Fig. 12C–D........................................................ Carrhotus silanthi (Caleb, 2020), Published as part of Satkunanathan, Abira & Benjamin, Suresh P., 2022, Phylogenetic placement of Carrhotus Thorell, 1891 with three new species from Sri Lanka (Araneae: Salticidae), pp. 78-110 in European Journal of Taxonomy 817 (1) on pages 102-107, DOI: 10.5852/ejt.2022.817.1765, http://zenodo.org/record/6518305, {"references":["Koch C. L. 1846. Die Arachniden: Getreu nach der Natur abgebildet und beschrieben. Vol. 13: 1 - 234, 4334 - 68 (f. 1078 - 1271); Vol. 14: 1 - 88, 4674 - 80, (f. 1272 - 1342). J. L. Lotzbeck, Nurnberg. https: // doi. org / 10.5962 / bhl. title. 43744 [For the correct year of publication see Brignoli 1985]","Thorell T. 1891. Spindlar fran Nikobarerna och andra delar af sodra Asien. Kongliga Svenska Vetenskaps- Akademiens Handlingar 24 (2): 11 - 49. Available from https: // www. biodiversitylibrary. org / page / 34324437 [accessed 25 Mar. 2022].","Andreeva E. M., Kononenko A. P. & Proszynski J. 1981. Remarks on genus Mogrus Simon, 1882 (Aranei, Salticidae). Annales Zoologici 36: 85 - 104.","Proszynski J. 1984. Remarks on Viciria and Telamonia (Araneae, Salticidae). Annales Zoologici 37 (18): 417 - 436.","Caleb J. T. D., Bera C. & Acharya S. 2020. New species and synonymies in the genus Carrhotus Thorell, 1891 from India (Aranei: Salticidae: Salticini). Arthropoda Selecta 29 (1): 51 - 66. https: // doi. org / 10.15298 / arthsel. 29.1.04","Proszynski J. 2009. Monograph of the Salticidae (Araneae) of the World (Version October 1 st, 2016). Available from http: // www. salticidae. pl / [accessed 25 Mar. 2022].","Sudhin P. P., Nafin K. S., Caleb J. T. D. & Sudhikumar A. V. 2021. A new spider species of the genus Carrhotus Thorell, 1891 (Aranei: Salticidae: Salticini) from Western Ghats of India. Arthropoda Selecta 30 (4): 551 - 556.","Simon E. 1902. Etudes arachnologiques. 32 e Memoire. LI. Descriptions d'especes nouvelles de la famille des Salticidae (suite). Annales de la Societe entomologique de France 71: 389 - 421. Available from https: // www. biodiversitylibrary. org / page / 8251793 [accessed 25 Mar. 2022]."]}

The systematics and taxonomy of the tropical Asian jumping spiders of the tribe Baviini is reviewed, with a molecular phylogenetic study (UCE sequence capture, traditional Sanger sequencing) guiding a reclassification of the group’s genera. The well-studied members of the group are placed into six genera: Bavia Simon, 1877, Indopadilla Caleb & Sankaran, 2019, Padillothorax Simon, 1901, Piranthus Thorell, 1895, Stagetillus Simon, 1885, and one new genus, Maripanthus Maddison, gen. nov. The identity of Padillothorax is clarified, and Bavirecta Kanesharatnam & Benjamin, 2018 synonymized with it. Hyctiota Strand, 1911 is synonymized with Stagetillus . The molecular phylogeny divides the baviines into three clades, the Piranthus clade with a long embolus ( Piranthus , Maripanthus ), the genus Padillothorax with a flat body and short embolus, and the Bavia clade with a higher body and (usually) short embolus (remaining genera). In general, morphological synapomorphies support or extend the molecularly delimited groups. Eighteen new species are described: Bavia nessagyna , Indopadilla bamilin , I. kodagura , I. nesinor , I. redunca , I. redynis , I. sabivia , I. vimedaba , Maripanthus draconis (type species of Maripanthus ), M. jubatus , M. reinholdae , Padillothorax badut , P. mulu , Piranthus api , P. bakau , P. kohi , P. mandai , and Stagetillus irri , all sp. nov., with taxonomic authority W. Maddison. The distinctions between baviines and the astioid Nungia Żabka, 1985 are reviewed, leading to four species being moved into Nungia from Bavia and other genera. Fifteen new combinations are established: Bavia maurerae (Freudenschuss & Seiter, 2016), Indopadilla annamita (Simon, 1903), I. kahariana (Proszynski & Deeleman-Reinhold, 2013), I. sonsorol (Berry, Beatty & Proszynski, 1997), I. suhartoi (Proszynski & Deeleman-Reinhold, 2013), Maripanthus menghaiensis (Cao & Li, 2016), M. smedleyi (Reimoser, 1929), Nungia hatamensis (Thorell, 1881), N. modesta (Keyserling, 1883), N. papakula (Strand, 1911), N. xiaolonghaensis (Cao & Li, 2016), Padillothorax casteti (Simon, 1900), P. exilis (Cao & Li, 2016), P. flavopunctus (Kanesharatnam & Benjamin, 2018), Stagetillus banda (Strand, 1911), all comb. nov. One combination is restored, Bavia capistrata (C. L. Koch, 1846). Five of these new or restored combinations correct previous errors of placing species in genera that have superficially similar palps but extremely different body forms, in fact belonging in distantly related tribes, emphasizing that the general shape of male palps should be used with caution in determining relationships. A little-studied genus, Padillothorus Proszynski, 2018, is tentatively assigned to the Baviini . Ligdus Thorell, 1895 is assigned to the Ballini .

Kanesharatnam, Nilani, Benjamin, Suresh P. (2021): Phylogenetic relationships and systematics of the jumping spider genus Colopsus with the description of eight new species from Sri Lanka (Araneae: Salticidae). Journal of Natural History 54 (43-44): 2763-2814, DOI: 10.1080/00222933.2020.1869335, URL: http://dx.doi.org/10.1080/00222933.2020.1869335

Genus Pancorius Simon, 1902 Type species: Ergane dentichelis Simon, 1899. Diagnosis This genus is distinguishable from closely related genera Colopsus, Evarcha, Hyllus by sandy brown habitus with pale white central and lateral prosomal bands, serrated longitudinal abdominal band (Figures 12 (e–h), Figures 18 (a,b), Figures 20 (a–g), Figures 21 (a–f), Figures 22 (a,b), Figures 23 (a–h), Figures 24 (a,b), Figures 25 (a,b)), simple palp with rounded or oval bulbus, short embolus, single RTA with pointed tip, epigyne with large central pocket, comparably small membranous window and multi-chambered spermathecae. Description Medium-sized spiders (6–10 mm). Sandy brown, thickset and hirsute habitus (Zhang et al. 2003) Prosoma high (Żabka 1990) with pale white central and lateral bands in both sexes (Figures 12 (e–h), Figures 21 (a–f)). Short, longitudinal fovea with pale white diamond mark behind it. Clypeus narrow. Chelicerae with simple retromarginal teeth (Ikeda 2013). Subparallel ocular quadrangle, first eye row is wider than third row in males, but in females eye row I is narrower than third row (Peckham and Peckham 1907; Ikeda 2013). Longitudinal median band of abdomen with serrated margins (Peng et al. 1998), this pattern is designated as ‘herring-bone pattern’ in Jastrzebski (Jastrzębski 2011). Simple male palp with rounded or oval bulbus with or without posterior lobe. Embolus thin and short to medium sized originating from antero-lateral portion of bulbus (Figures 18 (c,e), Figures 19 (a,e), Figures 22 (c,e), Figures 24 (c,e), Figures 26 (a); Prószyński 2017). RTA with a pointed tip without bifurcation. Palpal tibia medium sized. Epigyne with simple sclerotised plate ending in front of epigastric furrow (Prószyński 1992). Posterior epigynal plate often with a single central pocket (Figures 18 (c,e), Figures 19 (c,d,g,h), Figures 20 (c,d,h,i), Figures 25 (c,d)) sometimes two small lateral pockets at the edges of plate. A pair of small membranous window or sclerotised narrow crevices leading to copulatory openings at anterior half of the epigynum. Sclerotised spermathecae with 2–3 chambers. Composition At present, the genus comprises 34 species (World Spider Catalog 2020). However, some of them may be misplaced and need to be transferred to other closely related or new genera. This study adds three more new species to the genus, all endemic to Sri Lanka. Distribution This genus is primarily distributed in the Oriental region, including Borneo, Bhutan, China, India, Indonesia, Japan, Malaysia, Nepal, Philippines, Singapore, Sri Lanka, Taiwan and Vietnam., Published as part of Kanesharatnam, Nilani & Benjamin, Suresh P., 2021, Phylogenetic relationships and systematics of the jumping spider genus Colopsus with the description of eight new species from Sri Lanka (Araneae: Salticidae), pp. 2763-2814 in Journal of Natural History 54 (43 - 44) on pages 2796-2797, DOI: 10.1080/00222933.2020.1869335, http://zenodo.org/record/5405241, {"references":["Simon E. 1902. Etudes arachnologiques. 32 e Memoire. LI. Descriptions d'especes nouvelles de la famille des Salticidae (suite). Ann De La Societe Entomologique De France. 71: 389 - 421.","Zhang JX, Song DX, Li DQ. 2003. Six new and one newly recorded species of Salticidae (Arachnida: Araneae) from Singapore and Malaysia. Raffles Bull Zool. 51: 187 - 195.","Zabka M. 1990. Salticidae from the Nepal and Bhutan Himalayas. Genera Pancorius Simon 1902, Plexippus C. L. Koch 1846, and Pseudamycus Simon 1885 (Arachnida: Araneae). Senckenb Biol. 70: 161 - 178.","Ikeda H. 2013. Three species of the jumping spiders (Araneae: Salticidae: Nungia, Pancorius and Thyene) new to Japan. Acta Arachnol. 62 (2): 81 - 87. doi: 10.2476 / asjaa. 62.81.","Peckham G, Peckham EG. 1907. The Attidae of Borneo. Trans Wisconsin Acad Sci Arts Lett. 15: 603 - 653.","Peng XJ, Yin CM, Yan HM, Kim JP. 1998. Five jumping spiders of the family Salticidae (Arachnida: Araneae) from China. Korean Arachnol. 14: 36 - 43.","Jastrzebski P. 2011. Salticidae from the Himalayas. The genus Pancorius Simon, 1892 (Arachnida: Araneae). Genus. 22: 181 - 190.","Proszynski J. 2017. Pragmatic classification of the world's salticidae (Araneae). Ecol Montenegrina. 12: 1 - 133. doi: 10.37828 / em. 2017.12.1.","Proszynski J. 1992. Salticidae (Araneae) of India in the collection of the Hungarian National Natural History Museum in Budapest. Ann Zool Warszawa. 44: 165 - 277.","World Spider Catalog (2020). World Spider Catalog. Version 21.5. Natural History Museum Bern. online. accessed 2020 Aug 07. http: // wsc. nmbe. ch"]}

Taxonomic notes on the crab spider genus Talaus Simon, 1886 are provided. One new species, T. beccarii sp. nov. is described. Further four species are redescribed: T. nanus Thorell, 1890, T. triangulifer Simon, 1886, T. oblitus O. Pickard-Cambridge, 1899 and T. opportunus (O. Pickard-Cambridge, 1873). Talaus now contains 13 species.

Talaus triangulifer Simon, 1886 (Figs 2C, H–J, 4E, 7A–B) Talaus triangulifer Simon, 1886: 172. Type material: Holotype: female, INDONESIA, Sumatra, no more locality data given (MNHN 8289, examined by C. L. Deeleman; see remarks below). Other material examined. INDONESIA: East Kalimantan: 4♂, 1♀, Sepaku (40 km N of Balikpapan), de- graded remains of stand of primary rainforest, general hand collecting, 16 July 1979, P. R. & C. L. Deeleman (RMNH.ARA.17832). West Sumatra: 1♀, Mt. Singalang, Anai, 480–520 m, general foliage, 22 June 1994, S. Djo- josudharmo (RMNH.ARA.17834). MALAYSIA: Sabah: 1♀, Sorsinsim, 6° 06′N 116° 50′E, 5 year old adjacent secondary forest loc 21, canopy fogging Melochia umbellata (Stercul.), 19 February 1997, 500– 700 m, A. Floren (RMNH.ARA.17835); 1♀, 5° 26′N 116° 08′E, Crocker Range, 20 year old isolated secondary forest, CR.II.5, canopy fogging, Melanolepis glandulosa (Euphorbiaceae), 18 February 2001, A. Floren (RMNH.ARA.17833). Diagnosis. This species is very similar to T. xiphosus, which was described by Zhu and Ono (2007). A much larger series of speciemns are required to review the specific differences, if any. Males of this species are also simi- lar to those of T. opportunus, but are distinguished by the oval tegulum, the longer, broad-based E and configuration of the tibial apophyses. Females may be distinguished from congeners by the oval S, relatively long C shaped CD (Fig. 4E). Description. Male (RMNH.ARA.17832): Total length: 8.0; prosoma length: 4.6, width: 3.8. Leg I: femur 3.0, patella 1.0, tibia 2.8, metatarsus 2.4, tarsus 1.8. Prosoma reddish brown, with some long setae, surface with markings. ALE, PLE on tubercles, not connected to each other. ALE>PLE>AME>PME, MOA wider than long, distance between AMEs one diameter of AME. Distance between PME more than 4x AME. Palp: VTA large, apically hookshaped; RTA digitiform and well sclerotised. Bulb oval, embolus broad-based, stout, apically pointed (Figs 2 I–J, 7A–B). Female RMNH.ARA.17832: Total length: 10.2; prosoma length: 4.6, width: 4.0. Leg I: femur 3.4, patella 1.0, tibia 3.0, metatarsus 2.4, tarsus 1.8. In general, similar to male except for slightly lighter colour, dark spots of the opisthosoma (Fig. 2C). Epigynum and vulva as in Figs 2H, 4C. Distribution. Known from Indonesia (East Kalimantan) and Malaysia (Sabah). Remarks. The type specimen was examined and drawn by C. L. Deeleman. These drawings were compared with the examined material. The type locality is Sumatra, probably it is Mt Singalang (C. L. Deeleman, pers. comm.). Material from that locality has been examined for this study. The perceived differences in length and curvature of the CD between T. xiphosus and T. triangulifer mentioned in Zhu & Ono (2007) are minor and thus, these species might turn out to be synonyms. Additionally, T. elegans Thorell, 1890 might also turn out to be a junior synonym of T. triangulifer. The sketches of the female type done by C. L. Deeleman during a visit to MCSN in 1987 indicate slight differences from T. triangulifer. Unfortunately, I have been unable to examine the type as it has been misplaced since (M. Tavano, MCSN, pers. comm.)., Published as part of Benjamin, Suresh P., 2020, Distributional and taxonomic notes on the crab spider genus Talaus Simon, 1886 with description of a new species (Araneae: Thomisidae), pp. 405-416 in Zootaxa 4858 (3) on pages 414-415, DOI: 10.11646/zootaxa.4858.3.6, http://zenodo.org/record/4412520, {"references":["Simon, E. (1886) Especes et genres nouveaux de la famille des Thomisisdae. Actes de la Societe Linneenne de Bordeaux, 40, 167 - 187.","Zhu, M. S. & Ono, H. (2007) New record of the spider genus Talaus from south China, with description of a new species (Araneae: Thomisidae). Acta Arachnologica, 56, 81 - 83. https: // doi. org / 10.2476 / asjaa. 56.81","Thorell, T. (1890) Diagnoses aranearum aliquot novarum in Indo-Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30, 132 - 172."]}

Talaus oblitus O. Pickard-Cambridge, 1899 Figs 2D, 5 A–B Talaus oblitus O. Pickard-Cambridge, 1899: 526, pl. 30, fig. 8. Type material. Holotype: male, SRI LANKA, no more locality data given, leg. G. H. K. Thwaites (OUMNH b 1242 t.8, examined). Diagnosis. Easily distinguishable from congeners by the filiform embolus winding 2/3 around tegulum, the short, hooked VTA (lateral view) and the enlarged, truncated RTA. Other material examined. SRI LANKA: Central province: 1♂, Kandy District, Peradeniya Botanical Gardens, 460 m, 7° 17’ 57”N 80° 38’ 29”E, 29 December 2011, litter, Yuri Marusik (ZFMK, IFS _Tho_129,); 1♂, same locality, Hantana, 07° 14’ 57” N 80° 36’ 50” E, 18 December 2012, beating, N. Athukorala et al. (ZFMK, IFS _Tho_178,). Description. Male (from Peradeniya Botanical Gardens): Total length: 2.5; prosoma length: 1.2, width: 1.1. Leg I: femur 1.2, patella 0.4, tibia 1.1, metatarsus 1.0, tarsus 0.6. Prosoma reddish brown, with some long setae (prominent along the posterior margin), surface without markings. ALE, PLE on tubercles, not connected to each other. ALE>PLE>AME=PME, MOA wider than long, distance between AMEs three diameter of AME. Distance between PME more than 4x AME. Opisthosoma yellow-brown, with uniformly distributed black flakes. Palp as in Figs 5 A–B, filiform embolus, large, truncate RTA. Female: Unknown. Distribution. Sri Lanka., Published as part of Benjamin, Suresh P., 2020, Distributional and taxonomic notes on the crab spider genus Talaus Simon, 1886 with description of a new species (Araneae: Thomisidae), pp. 405-416 in Zootaxa 4858 (3) on page 411, DOI: 10.11646/zootaxa.4858.3.6, http://zenodo.org/record/4412520, {"references":["Pickard-Cambridge, O. (1899) On some new species of exotic Araneidea. Proceedings of the Zoological Society of London, 67, 518 - 532, pls. XXIX-XXX. https: // doi. org / 10.1111 / j. 1469 - 7998.1899. tb 06872. x"]}

Talaus opportunus (O. Pickard-Cambridge, 1873) (Figs 2E, F, 6 A–D) Thomisus opportunus O. Pickard-Cambridge, 1873: 120, plate. 13, fig. 5. Talaus opportunus (O. Pickard-Cambridge). Simon 1895: 997, fig. 1061; Simon, 1906: 288; Murphy and Murphy 2000: 457, fig 27 Type material. Syntypes: 1♂ and 4♀, SRI LANKA, no more locality data given (OUMNH b1239, examined). Diagnosis. This species is similar to T. triangulifer, but is distinguished by the shorter, broad-based E and the VTA and RTA which are equal in length. Females may be distinguished from congeners by the oval S, relatively longer C shaped CD (Figs 6C, D). Description. Male: Prosoma reddish brown, with some long setae, surface with markings. ALE, PLE on tubercles, not connected to each other. ALE>PLE>AME>PME, MOA wider than long, distance between AMEs one diameter of AME. Distance between PME more than 4x AME. Palp: Tibia with VTA large, apically hook-shaped; RTA digitiform and well sclerotised. Bulb kidney-shaped, embolus broad-based, stout, apically pointed (Figs 6 A–B). Female: In general, similar to male except for slightly lighter colour, dark spots of the opisthosoma (Fig 7). Epigynum and vulva as in Figs 6 C–D. Distribution. Sri Lanka. Remarks. According to Pickard-Cambridge (1873) the original series consisted of 2 males and 4 females. However, only one male and 4 females were found in OUMNH. Series label and first description gives the locality as Ceylon (now Sri Lanka). The record of this species from India by Simon (1906) cannot be confirmed. Roewer (1955) and WSC (2020), both list the type locality as India., Published as part of Benjamin, Suresh P., 2020, Distributional and taxonomic notes on the crab spider genus Talaus Simon, 1886 with description of a new species (Araneae: Thomisidae), pp. 405-416 in Zootaxa 4858 (3) on pages 411-412, DOI: 10.11646/zootaxa.4858.3.6, http://zenodo.org/record/4412520, {"references":["Pickard-Cambridge, O. (1873) On some new genera and species of Araneida. Proceedings of the Zoological Society of London, 41, 112 - 129, pls. XII-XIV.","Simon, E. (1895) s. n. Histoire naturelle des araignees. Vol. 1. Roret, Paris, pp. 761 - 1084.","Simon, E. (1906) Arachnides (2 e partie). In: Voyage de M. Maurice Maindron dans l'Inde meridionale. 8 e Memoire. Annales de la Societe Entomologique de France, 75, 279 - 314.","Murphy, F. & Murphy, J. (2000) An introduction to the spiders of South East Asia with notes on all the genera. Malaysian Nature Society, Kuala Lumpur, 625 pp.","Roewer, C. F. (1955) Katalog der Araneae von 1758 bis 1940, bzw. 1954. Vol. 2. Institut Royal des Sciences naturelles de Belgique, Bruxelles, 923 pp.","WSC (2020) World Spider Catalog. Version 19.0. Natural History Museum, Bern. Available from: http: // wsc. nmbe. ch (accessed 29 April 2020)"]}

Talaus beccarii sp. nov. (Figs 1 A–F) Type material. Holotype: male, MALAYSIA: Sabah: North Borneo, Kinabalu National Park, Poring hot springs, 6º2’N 116º50’E, 500-700 m, primary forest, canopy fogging Aglaia sp. (Meliaceae), 27 March 1998, leg. A. Floren (RMNH.ARA.17837). Other material examined. MALAYSIA: Sabah: 1♀, Kinabalu National Park, park headquarters, 6º04’N 116º33’E, 1550 m, primary rainforest, general hand collecting, 29 June 1979, leg. P. R. and C. L. Deeleman (RMNH. ARA.17836). Diagnosis. Easily distinguishable from congeners by the curved embolus (winding 0.25 times around tegulum) and shape of RTA and VTA (short, stout, truncate). Females can be distinguished by the comparatively small epigynal hood, and irregularly shaped CD and S (Figs 1D, E). Etymology. Named for Odoardo Beccari (16 November 1843 – 25 October 1920), an Italian naturalist and collector of many spiders from South East Asia described by Teodor Thorell (3 May 1830 – 22 December 1901). Description. Male: Total length: 8.2; prosoma length: 2.4, width: 2.4. Leg I: femur 3.0, patella 1.0, tibia 2.4, metatarsus 1.8, tarsus 1.4. Prosoma rectangular. Prosoma, chelicera, endites, labium, sternum reddish brown. ALE, PLE on tubercles, not connected to each other. ALE>PLE>AME>PME, MOA wider than long. Palp as in Figs 1 A–B. Female: Total length: 9.2; prosoma length: 4.4, width: 4.0. Leg I: femur 3.0, patella 1.0, tibia 2.2, metatarsus 2.0, tarsus 1.4. In general similar to male except for slightly larger size and lighter colour. Epigynum and vulva as in Figs 1 D–E. Distribution. Known only from Kinabalu National Park (Malaysia, Borneo)., Published as part of Benjamin, Suresh P., 2020, Distributional and taxonomic notes on the crab spider genus Talaus Simon, 1886 with description of a new species (Araneae: Thomisidae), pp. 405-416 in Zootaxa 4858 (3) on page 406, DOI: 10.11646/zootaxa.4858.3.6, http://zenodo.org/record/4412520

Genus: Talaus Simon, 1886 Type species: Talaus triangulifer Simon, 1886. Diagnosis. Small to medium sized spiders, 1.9 to 9.0 mm. Leg formula 2143, Femur 1 with prolateral spines, femora 1 and 3 with dorsal spines. Male palp with RTA and VTA and in some species with DTA. Embolus spiniform, short to long (1x the circumference of tegulum), with broad-base and stout in some species. Epigynum slightly sclerotized, median septum lacking, CD short, tubular to irregular in shape, spermathecae rounded to irregular in shape. For a detailed diagnosis see Zhu & Ono (2007). Species composition. The World Spider Catalog (WSC 2020) lists twelve species: T. dulongjiang Tang, Yin, Ubick & Peng, 2008, T. elegans Thorell, 1890, T. limbatus Simon, 1895, T. nanus Thorell, 1890, T. niger Tang, Yin, Ubick & Peng, 2008, T. oblitus O. Pickard-Cambridge, 1899, T. opportunus (O. Pickard-Cambridge, 1873), T. samchi Ono, 2001, T. semicastaneus Simon, 1909, T. sulcus Tang & Li, 2010, T. triangulifer Simon, 1886 and T. xiphosus Zhu & Ono, 2007. The 13 th species, Talaus beccarii sp. nov. is described in this paper. Distribution. South-East Asia (Bhutan, China, India, Indonesia, Myanmar, Sri Lanka and Vietnam). T. limbatus from South Africa is probably misplaced in this genus., Published as part of Benjamin, Suresh P., 2020, Distributional and taxonomic notes on the crab spider genus Talaus Simon, 1886 with description of a new species (Araneae: Thomisidae), pp. 405-416 in Zootaxa 4858 (3) on page 406, DOI: 10.11646/zootaxa.4858.3.6, http://zenodo.org/record/4412520, {"references":["Simon, E. (1886) Especes et genres nouveaux de la famille des Thomisisdae. Actes de la Societe Linneenne de Bordeaux, 40, 167 - 187.","Zhu, M. S. & Ono, H. (2007) New record of the spider genus Talaus from south China, with description of a new species (Araneae: Thomisidae). Acta Arachnologica, 56, 81 - 83. https: // doi. org / 10.2476 / asjaa. 56.81","WSC (2020) World Spider Catalog. Version 19.0. Natural History Museum, Bern. Available from: http: // wsc. nmbe. ch (accessed 29 April 2020)","Tang, G., Yin, C. M., Ubick, D. & Peng, X. J. (2008) Two new species of the crab spider genus Talaus (Araneae: Thomisidae) from Yunnan Province, China. Zootaxa, 1815 (1), 62 - 68. https: // doi. org / 10.11646 / zootaxa. 1815.1.6","Thorell, T. (1890) Diagnoses aranearum aliquot novarum in Indo-Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30, 132 - 172.","Simon, E. (1895) s. n. Histoire naturelle des araignees. Vol. 1. Roret, Paris, pp. 761 - 1084.","Pickard-Cambridge, O. (1899) On some new species of exotic Araneidea. Proceedings of the Zoological Society of London, 67, 518 - 532, pls. XXIX-XXX. https: // doi. org / 10.1111 / j. 1469 - 7998.1899. tb 06872. x","Pickard-Cambridge, O. (1873) On some new genera and species of Araneida. Proceedings of the Zoological Society of London, 41, 112 - 129, pls. XII-XIV.","Ono, H. (2001) Crab spiders of the family Thomisidae from the Kingdom of Bhutan (Arachnida, Araneae). Entomologica Basiliensis, 23, 203 - 236.","Simon, E. (1909) Etude sur les arachnides du Tonkin (1 re partie). Bulletin Scientifique de la France et de la Belgique, 42, 69 - 147. https: // doi. org / 10.5962 / bhl. part. 24151","Tang, G. & Li, S. Q. (2010) Crab spiders from Xishuangbanna, Yunnan Province, China (Araneae, Thomisidae). Zootaxa, 2703 (1), 1 - 105. https: // doi. org / 10.11646 / zootaxa. 2703.1.1"]}

Kanesharatnam, Nilani, Benjamin, Suresh P. (2020): First record of Synagelides Strand, 1906 (Araneae: Salticidae) from Sri Lanka description of four endemic species from tropical wet forest of the island. Zootaxa 4790 (1): 43-56, DOI: https://doi.org/10.11646/zootaxa.4790.1.2