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Genus Zodarion Walckenaer, 1826 Comments. According to Bosmans (2009), the males within the germanicum group are characterized by a conspicuous tegular projection which form is often of diagnostic value at species level. The males within the spinibarbe group have an unmodified tegulum and possess a large retinaculum (tegular apophysis attached to tegulum with membranous support sensu Jocqué 1991), often incised prolaterally, and with several large teeth. The shape of retinaculum is of significant diagnostic value., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on page 248, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295.","Jocque, R. (1991) A generic revision of the spider family Zodariidae (Araneae). Bulletin of the American Museum of Natural History, 201, 1 - 160."]}

Zodarion rubidum Simon, 1914 Zodarion rubidum Simon, 1914: 233, 234, f. 467, 479–480, 484 (D ♂ ♀). Zodarion fulvonigrum Vogel, 1968: 99, f. 1–7 (♂; misidentified per Bosmans, 1994 b: 132). Zodarion rubidum: Bosmans 1994: 132, f. 65–67, 121–122 (♂ ♀). Zodarion rubidum: Verhoogt & IJland 2021: 7, f. 1, 5–8 (♂ ♀). For the full list of references, see WSC (2022). Material examined. 1♂, BULGARIA: Sofia-Center, N42.69 ˚, E23.31 ˚, 560 m, 20.05.2017, 3♂, 3♀, 16.05.2020, 1♂, 3♀, 20.05.2017, 1♂, 07.05.2020, 1♂, 1♀, Sofia, the Southern Park, N42.66 ˚, E23.31 ˚, 575 m, 22.05.2017, 1♂, Sofia, Borisova Gradina Park, N42.6827 ˚, E23.3459 ˚, 567 m, 24.05.2020, 1♀, Lozenska Mt., German village, N42.66 ˚, E23.40 ˚, 544–687 m, 19.08.2017, 1♂, 18.05.2021, 1♂, 05.06.2021, S. Indzhov leg. Distribution. Europe, Asia Minor, North America (Danışman et al. 2022; Vogel 1968; WSC 2022). Remarks. The species was described from South France and since a few decades shows strong spreading tendencies within Europe. In the Balkans, Z. rubidum is known only from Slovenia (Kuralt & Kostanjšek 2016). The new data (Fig. 76) outline the southern limit of its distribution in the Balkans at present., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on page 254, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Simon, E. (1914) Les arachnides de France. Synopsis generale et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, pp. 1 - 308.","Vogel, B. R. (1968) A zodariid spider from Pennsylvania (Araneida: Zodariidae). Journal of the New York Entomological Society, 76, 96 - 100.","Bosmans, R. (1994) Revision of the genus Zodarion Walckenaer, 1833 in the Iberian Peninsula and Balearic Islands (Araneae, Zodariidae). Eos, 69, 115 - 142.","Verhoogt, K. & IJland, S. (2021) De sepia mierenjager Zodarion rubidum nieuw voor Nederland (Araneae: Zodariidae) [The anteating spider Zodarion rubidum new to the Netherlands (Araneae: Zodariidae)]. Nederlandse Faunistische Mededelingen, 57, 7 - 12.","Danisman, T., Kunt, K. B. & Ozkutuk, R. S. (2022) The Checklist of the Spiders of Turkey. Version 2022, Available from: http: // www. spidersofturkey. info (accessed 25 June 2022)","Kuralt, Z. & Kostanjsek, R. (2016) A contribution to the Slovenian spider fauna - III. Natura Sloveniae, 18, 69 - 75."]}

Zodarion evvoia Bosmans, 2009 Zodarion evvoia: Bosmans et al. 2019: 15. Erroneously reported from Bulgaria (probably lapsus calami) in Bosmans et al. (2019) and then in the WSC ( 2022) and Nentwig et al. (2022). Not documented record from Bulgaria., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on page 253, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295.","Bosmans, R., Pantini, P., Loverre, P. & Addante, R. (2019) New species and new records of ant - eating spiders from Mediterranean Europe (Araneae: Zodariidae). Arachnologische Mitteilungen, 57, 8 - 20. https: // doi. org / 10.30963 / aramit 5703","Nentwig W., Blick, T., Bosmans, R., Gloor, D., Hanggi, A. & Kropf, C. (2022) Spiders of Europe. Version 1.2022. Available from: https: // www. araneae. nmbe. ch (accessed 25 June 2022) https: // doi. org / 10.24436 / 1"]}

Zodarion emilijae Deltshev & Naumova sp. n. Figs 1, 5, 9, 13, 17–19, 29, 33, 37, 41, 45, 48, 51, 54. Z. epirense: Matevski et al. 2020: 12 (misidentified) Z. prope epirense: Matevski et al. 2020: 12 (misidentified). Types. Holotype: ♂ (NMNHS-Zod-0003.1), NORTH MACEDONIA: Kozhuf Mts, N41.22158 ˚, E22.27413 ˚, 1673 m, 05.2016, D. Matevski leg. Paratypes: 2♂, 2♀ (NMNHS-Zod-0003.2) same data as the holotype; 7♀ (NMNHS-Zod-0003.3), Mariovo region, Nidzhe Mts, Kaimakchalan peak, N40.9489 ˚, E21.8000 ˚, 2300 m, 23.06.2009, C. Deltshev, M. Komnenov & E. Stojkoska leg. Comparative material examined. Zodarion blagoevi Bosmans, 2009 (2♂, paratypes (MHNG), BULGARIA: General Todorov village, 400 m, under stones, 04.03.1996, G. Blagoev leg.; 1♀ paratype (MHNG), BULGARIA: Belasitsa Mts, Kongur Peak, 1200 m, under stones, 14.06.1988, C. Deltshev leg.; 1♂, (Lazarov 2005: 150, sub Z. epirense, misidentified) (NMNHS), BULGARIA: Slavyanka Mts, Kalimantsi village, 07.2002); Zodarion epirense Brignoli, 1984 (♂, holotype (MHNG), GREECE: Ipeiros, Ioanina, Metsovo pass, 1620 m, 28.04.1973, V. Mahnert leg.; ♀, paratype (MHNG), Ioanina, Eleousa, 28.07.1969, P. Brignoli leg). Diagnosis: Zodarion emilijae sp. n. belongs to the spinibarbe group (sensu Bosmans 2009) and is similar to Z. blagoevi, Z. epirense and Z. zorba. Males are distinguishable by the following traits: the proximal prolateral tooth of the retinaculum (PPR) 2 times longer than wide in Z. emilijae sp. n. (Fig. 18) vs. 1.2–1.5 times longer than wide in Z. blagoevi (Fig. 21) and Z. epirense (Fig. 24) and about 1.5 times wider than long in Z. zorba (Fig. 27); incision of retinaculum U-shaped in Z. emilijae sp. n. vs. roughly V-shaped in Z. blagoevi and Z. zorba and almost round in Z. epirense, with opening two times longer than PPR width in Z. emilijae sp. n. vs. almost equal in Z. blagoevi and Z. epirense and two times shorter than PPR width in Z. zorba. Females are distinguishable as follows: posterior margin of epigynal plate trapezoid and deeply incised in Z. emilijae sp. n. vs. triangular with a rounded tip and less incised in Z. blagoevi, Z. epirense and Z. zorba; spermathecae separated by less than 2 diameters in Z. emilijae sp. n. (Figs 22, 25, 27) vs. more than 2 diameters in Z. blagoevi and Z. zorba (Figs 24, 26, 28) and 4 diameters in Z. epirense (Fig. 23); spermathecae large and almost spherical in Z. emilijae sp. n. vs. small and two-chambered in Z. epirense. Snow-white coxae and abdominal spots present in both sexes in Z. emilijae sp. n. (Figs 5, 33) vs. whitish or pale beige in Z. blagoevi (Figs 6, 34), Z. epirense (Figs 7, 35) and Z. zorba (Figs 8, 36). Etymology. The specific name is dedicated to Emilija Stojkoska (Skopje, North Macedonia), a malacologist who has a great contribution to the faunistic studies in North Macedonia. Description. Male (holotype). Habitus as in Figs 1, 5. Total length 3.72. Carapace 1.62 long, 1.08 wide; sternum 0.9 long, 0.72 wide; chelicerae 0.42 long, 0.20 wide; clypeus 0.16 high. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.05, PLE 0.05; AME–AME 0.07, ALE–AME 0.03, PLE–AME 0.03, PME–PME 0.22. Abdomen 1.62 long. Legs: I—(1.15, other segments missing); II—(1.08, other segments missing); III—4.28 (1.08, 0.54, 0.9, 1.08, 0.68); IV—6.04 (1.44, 0.60, 1.47, 1.54, 0.9). Color (Figs 1, 5): carapace and sternum dark-brown. Chelicerae brown. Abdomen dark sepia-brown with white spot, extending a half of the venter. Spinnerets whitish, uniform in color. Legs: coxae snow white, other segments brown to brown-yellow, without annulations. Palp as in Figs 9, 13, 17, 19. Femur 0.54 long, 0.18 wide, dark brown; patella 0.22 long, 0.18 wide, beige; tibia 0.11 long, 0,18 wide, brown; RTA 0.18 long, triangular, slightly curved ventrally, dark brown. Cymbium 0.54 long, 0.40 wide, dark brown with pale tip; with slightly concave bulge retro-basally; bulb 0.43 long, 0.36 wide. Tegulum oval, with distinct prolateral basal bulge; retinaculum 0.47 long, 0.16 wide, with U-shaped incision and 2 unequal teeth, proximal, longer than distal, claw-shaped. Embolus narrow, curved and pointed, originating at a 9 o’clock position and terminating at a 1 o’clock position. Female (paratype). Habitus as in Figs 29, 33. Total length 3.78. Carapace 1.62 long, 1.08 wide; sternum 0.9 long, 0.72 wide; chelicerae 0.54 long, 0.34 wide; clypeus 0.28 high. Eye sizes and interdistances: AME 0.05, ALE 0.07, PME 0.05, PLE 0.05, AME–AME 0.07. ALE–AME 0.03, PLE–AME 0.03, PME–PME 0.22. Abdomen 2.16 long. Legs: I—(1.15, 0.54, 1.44, 1.69, tarsus missing); II—(1.15, other segments missing); III—4.57 (1.04, 0.54, 0.97, 1.18, 0.72); IV—6.26 (1.62, 0.61, 1.44, 1.62, 0.97). Coloration as in male. Epigyne (Figs 22, 25, 27): epigynal plate 2 times wider than long, with trapezoid sclerotized sutures. Posterior margin trapezoid, about 2 times wider than long, not reaching half the length of the spermathecae and with large median incision; spermathecae oval, slightly wider than long (0.11 long, 0.14 wide), separated by less than 2 diameters. Ecology. The scarce data suggest that Z. emilijae sp. n. is a species with spring activity, preferring elevations above 1670 m in the coniferous and subalpine zone of the mountains. Distribution. Known only from the type localities in Kozhuf Mts and Nidzhe Mts, North Macedonia (Fig. 76)., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on pages 248-250, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Matevski, D., Cvetkovska - Gjorgjevska, A., Prellik, D., Hristovski, S., Naumova, M. & Deltshev, C. (2020) Distribution and community structure of araneocoenoses (Araneae) along an altitudinal gradient on Kozuf Mountain (North Macedonia). Springer, Biologia, 1 - 14. https: // doi. org / 10.2478 / s 11756 - 020 - 00474 - 2","Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295.","Lazarov, S. (2005) Spiders (Araneae) from Maquises in South - West Bulgaria) - Part I. Acta Zoologica Bulgarica, 57 (2), 145 - 152.","Brignoli, P. M. (1984) Ragni di Grecia XII. Nuovi dati su varie famiglie (Araneae). Revue Suisse de Zoologie, 91 (2), 281 - 321. https: // doi. org / 10.5962 / bhl. part. 81881"]}

Zodarion germanicum (C.L. Koch, 1837) The Balkans records of Z. germanicum from Bulgaria: Vratsa (Drensky 1913: 60, sub Zodarium (Enio) g.), Greece: Attika (Roewer, 1928: 119),Alexandroupolis (Drensky 1928: 187, sub Zodarium g.) and Parnis Oros (Hadjissarantos, 1940: 34, sub Zodarium g.) and from North Macedonia: Gevgelija (Stojićević, 1929: 41, sub Zodarium g.), cannot be checked because the materials are probably lost. All other records from Bulgaria by Drensky (1913, 1921, 1932, 1936a, 1940) have been re-examined and found to refer to Z. pirini Drensky, 1921 (Deltshev 1987). According to Bosmans (1997) the confirmed records of Z. germanicum are located much further north—in northeastern Croatia and northern Serbia (Djakovo and Fruska Gora) and all southern records in the Balkans are very doubtful. We completely agree with his assessment, moreover, all available re-examined specimens are misidentified and no proven records in the last over 80 years; therefore, we remove Z. germanicum from the checklists of Bulgaria, Greece and North Macedonia., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on page 253, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Drensky, P. (1913) Contribution a la faune d'araignee de la Bulgarie. Sbornik na Balgarskata Akademiya na Naukite, 2 (1), 1 - 146. [In Bulgarian, French summary]","Roewer, C. F. (1928) Araneae. In: Zoologische Streifzuge in Attika, Morea, und besonders auf der Insel Kreta, II. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen, 27, 92 - 123.","Drensky, P. (1928) Contribution a l'etude des Arachnides de la Thrace du sud et cotes de la Mer Egee. Travaux de la Societe Bulgare des Sciences Naturelles, 13, 183 - 192.","Hadjissarantos, H. (1940) Les Araignees de I'Attique. Athens, pp. 1 - 132.","Stojicevic, D. (1929) Les araignees de Serbie. Araneae Sund. Glasnevin Museum de Historia Natural, Beograd, 19, 1 - 65.","Drensky, P. (1921) Contribution a l'etude des araignees de la Macedoine orientale et de Pirine planina. Spisanie na Balgarskata Akademiya na Naukite, 23, 1 - 50. [In Bulgarian, French summary]","Drensky, P. (1932) Zur Kenntnis der Spinnenfauna Tscham - Kuria im Rilagebirge (Bulgarien). Travaux de la Societe Bulgare des Sciences Naturelles, 15 / 16, 326 - 332. [In Bulgarian] https: // doi. org / 10.1007 / BF 01610253","Drensky, P. (1936 a) Katalog der echten Spinnen (Araneae) der Balkanhalbinsel. Sbornik na Balgarskata Akademiya na Naukite, 32, 1 - 223.","Drensky, P. (1940) Die Spinnenfauna Bulgariens. IV. Unterordnung Arachnomorphae, II Gruppe Trionichia, Familien: Zodariidae, Dictynidae und Amaurobiidae. Mitteilungen aus den Koniglichen Naturwissenschaftlichen Instituten in Sofia, 13, 169 - 194. [In Bulgarian]","Deltshev, C. (1987) A Critical Review of Genus Zodarion Walckenaer (Araneae, Zodariidae) in Bulgaria. Acta Zoologica Bulgarica, 33, 19 - 25.","Bosmans, R. (1997) Revision of the genus Zodarion Walckenaer, 1833, part II. Western and Central Europe, including Italy (Araneae: Zodariidae). Bulletin of the British Arachnological Society, 10, 265 - 294."]}

Zodarion graecum (C. L. Koch, 1843) The reports of this species from Bulgaria: Rila Mts, (Borovets resort) by Drensky (1932: 330), Belasitsa Mts by Drensky (1936a: 31) and Slavyanka Mts by Drensky (1940: 174), as well as from North Macedonia (Udovo, near Gevgelija) by Stojićević (1929: 41) are doubtful, as the certified species records are from Greece and Croatia only (Bosmans 2009). The material cannot be reviewed (probably lost) and as there are no subsequent reports of the species in Bulgaria and North Macedonia, it is hereby removed from their checklists., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on page 254, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Drensky, P. (1932) Zur Kenntnis der Spinnenfauna Tscham - Kuria im Rilagebirge (Bulgarien). Travaux de la Societe Bulgare des Sciences Naturelles, 15 / 16, 326 - 332. [In Bulgarian] https: // doi. org / 10.1007 / BF 01610253","Drensky, P. (1936 a) Katalog der echten Spinnen (Araneae) der Balkanhalbinsel. Sbornik na Balgarskata Akademiya na Naukite, 32, 1 - 223.","Drensky, P. (1940) Die Spinnenfauna Bulgariens. IV. Unterordnung Arachnomorphae, II Gruppe Trionichia, Familien: Zodariidae, Dictynidae und Amaurobiidae. Mitteilungen aus den Koniglichen Naturwissenschaftlichen Instituten in Sofia, 13, 169 - 194. [In Bulgarian]","Stojicevic, D. (1929) Les araignees de Serbie. Araneae Sund. Glasnevin Museum de Historia Natural, Beograd, 19, 1 - 65.","Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295."]}

Zodarion weissi Deltshev & Naumova sp. n. Figs 61–75. Types. Holotype: ♂ (NMNHS-Zod-0004.1), BULGARIA: Sofia Mt., Kremikovtsi ward, N42.7882 ˚, E23.4812 ˚, 645 m, 23.06.2019, M. Naumova leg. Paratypes: 2♂, 2♀ (NMNHS-Zod-0004.2) same data as the holotype. Comparative material examined. Zodarion abantense Wunderlich, 1980 (♂ holotype, 1♀ paratype (SMF 30055, 58853), TURKEY, Bolu village, Abant Mountains, 29.04– 20.05.1976, H. Korge leg.; Z. pirini (1♂, 2♀ (IBER), BULGARIA, Eastern Rhodopes Mts, Blagun village, N41.3698 ˚, E25.7515 ˚, 423 m, 24.05.2018, M. Naumova leg.; Z. turcicum Wunderlich, 1980 (1♂ (IBER), BULGARIA, South Black Sea coast, Primorsko resort, N42.2544 ˚, E27.7487 ˚, 3 m, 27.04– 07.06.2016, T. Teofilova, T. Ljubomirov & N. Karaivanov leg.). Diagnosis. Zodarion weissi sp. n. belongs to the germanicum species group (sensu Bosmans 2009) and resembles Z. aurorae Weiss, 1982. Males differ by the following traits: tegular projection elongated triangular in lateral view in Z. weissi sp. n., vs. oblong with a broad rounded tip in Z. aurorae; basal tegular bulge reaching to the half of the tegular projection in Z. weissi sp. n. (Figs 63–65, 71, 72), vs. reaching more than 2/3 of the tegular projection length in Z. aurorae (see in Weiss 1982: 80, fig. 1) in lateral view. Females differ as follows: posterior margin of the epigynal plate with triangular incision with broad and shallow hood, 2.5–3 times wider than long in Z. weissi sp. n. (Figs 68–70, 73–75), vs. narrow and deep anterior hood, clearly longer than wide in Z. aurorae (see in Weiss 1982: 80, fig. 3). Etymology. The species is dedicated to Ingmar Weiss (Grafenau, Germany) a well-known arachnologist who has great contribution to araneological research in Romania. Description. Male (holotype). Total length 3.3. Carapace 1.54 long, 1.22 wide; sternum 0.9 long, 0.72 wide; chelicerae 0.54 long, 0.32 wide; clypeus 0.28 high. Eye sizes and interdistances: AME 0.11, ALE 0.07, PME 0.07, PLE 0.07, AME–AME 0.07, ALE–PLE 0.03, PME–PME 0.22. Abdomen 1.8 long. Legs: I—5.22 (1.26, 0.54, 1.26, 1.26, 0.9); II—4.86 (1.15, 0.54, 1.26, 1.08, 9.82); III—4.46 (1.26, 0.36, 0.97, 1.15, 0.72); IV—5.97 (1.69, 0.54, 1.22, 1.62, 0.9). Color (Fig. 61, 62): Carapace, sternum and chelicerae brown. Abdomen dark sepia-brown with a thick U-shaped, ventrolateral whitish spot on each side. Spinnerets bright, uniform in color. Coxae whitish, first couple light-brown. Other segments of the legs brown to brown-yellow, without annulations. Palp as in Figs 63–65, 71, 72. Femur 0.54 long, 0.18 wide, brown; patella 0.18 long, 0.18 wide, beige; tibia 0.11 long, 0,18 wide, brown, with elongated triangular retrolateral apophysis 0.18 long. Cymbium 0.65 long, 0.48 wide, brown with lightening tip; bulb 0.36 long, 0.29 wide. Tegulum with large basal bulge extended to the half of the tegular projection in prolateral view; tegular projection 0.18 long, 0.11 wide, elongated triangle in lateral view, perpendicular to the tegular surface, apically thinned; retinaculum 0.11 long, 0.07 wide, rhomboid-shaped, same sized as the bulge in ventral view; embolus thin, apically pointed, originating at a 9 o’clock position and terminating at a 1 o’clock position. Female. Total length 4.78. Carapace 1.9 long, 1.26 wide; sternum 1.08 long, 0.82 wide; chelicerae 0.54 long, 0.32 wide; clypeus 0.36 high. Eye sizes and interdistances:AME 0.09, ALE 0.06, PME 0.05, PLE 0.05, AME–AME 0.02, ALE–PLE 0.02, PME–PME 0.13. Abdomen 2.88 long. Legs: I—5.4 (1.36, 0.61, 1.26, 1.26, 0.9); II—5.07 (1.33, 0.61, 1.08, 1.26, 0.79); III—5.11 (1.33, 0.54, 1.08, 1.44, 0.72); IV—6.78 (1.8, 0.61, 1.62, 1.8, 0.9). Coloration as in male. Epigyne (Figs 68–70, 73–75): epigynal plate 1.5 times wider than long, broadly ovate. Posterior margin with large triangular incision with a base as wide as a distance between spermathecae and a tip reaching their half. Hood wide and shallow. Spermathecae rounded, separated by 2.5 diameters. Ecology. Found only in June, under stones in an open, dry and sunny place with very sparse low vegetation. Distribution. Known only from the type locality in Sofia Mt., Bulgaria (Fig. 76)., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on pages 256-258, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Wunderlich, J. (1980) Zur Gattung Zodarion Walckenaer 1847 mit Neubeschreibungen (Arachnida: Araneae: Zodariidae). Senckenbergiana Biologica, 60, 229 - 240.","Bosmans, R. (2009) Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). Contributions to Natural History, 12, 211 - 295.","Weiss, I. (1982) Konstruktions - und Funktionsanalyse der Kopulationsorgane von Zodarion aurorae n. sp. aus Rumanien (Arachnida, Araneae, Zodariidae). Reichenbachia, 20, 77 - 83."]}

Zodarion italicum (Canestrini, 1868) All reports of Z. italicum from Bulgaria and North Macedonia were by Drensky (1913: 60, 1929: 48, 1935: 100, 1936a: 31 (erroneously after Drensky 1932), 1936b: 94, 1940: 173) and Stojićević (1929: 41) and no specimen has been reported over the next more than 80 years. The concerning material (probably lost) has not been examined by the subsequent authors, who either considered the occurrence of the species in the region as doubtful (Deltshev 1987) or completely omitted it (Bosmans 1997, map 11); therefore, we remove this species from the lists of Bulgaria and North Macedonia., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on page 254, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Drensky, P. (1913) Contribution a la faune d'araignee de la Bulgarie. Sbornik na Balgarskata Akademiya na Naukite, 2 (1), 1 - 146. [In Bulgarian, French summary]","Drensky, P. (1929) Spinnen (Araneae) aus Mittel und Sud - West Mazedonien. Spisanie na Balgarskata Akademiya na Naukite, 39, 1 - 76. [In Bulgarian]","Drensky, P. (1935) Uber die von Dr. Stanko Karaman in Jugoslavien und besonders in Mazedonien gesammelten Spinnen (Araneae). Mitteilungen aus den Koniglichen Naturwissenschaftlichen Instituten in Sofia, 8, 97 - 110.","Drensky, P. (1936 a) Katalog der echten Spinnen (Araneae) der Balkanhalbinsel. Sbornik na Balgarskata Akademiya na Naukite, 32, 1 - 223.","Drensky, P. (1932) Zur Kenntnis der Spinnenfauna Tscham - Kuria im Rilagebirge (Bulgarien). Travaux de la Societe Bulgare des Sciences Naturelles, 15 / 16, 326 - 332. [In Bulgarian] https: // doi. org / 10.1007 / BF 01610253","Stojicevic, D. (1929) Les araignees de Serbie. Araneae Sund. Glasnevin Museum de Historia Natural, Beograd, 19, 1 - 65.","Deltshev, C. (1987) A Critical Review of Genus Zodarion Walckenaer (Araneae, Zodariidae) in Bulgaria. Acta Zoologica Bulgarica, 33, 19 - 25.","Bosmans, R. (1997) Revision of the genus Zodarion Walckenaer, 1833, part II. Western and Central Europe, including Italy (Araneae: Zodariidae). Bulletin of the British Arachnological Society, 10, 265 - 294."]}

Zodarion epirense Brignoli, 1984 Recently reported from Bulgaria: Slavyanka Mts (Lazarov 2005: 150), Maleshevska Mt. (Lazarov 2007: 140) and Samokov Plain (Kalushkov et al. 2008: 66). The reviewed materials (see also under Z. emilijae sp. n. and Z. zorba) reveal that these records refer to Z. blagoevi (in Lazarov 2005) and Z. zorba (in Kalushkov et al. 2008). The material from Maleshevska Mt. (Lazarov 2007) cannot be checked (probably lost) and is doubtful. Therefore, we remove Z. epirense from the Bulgarian checklist., Published as part of Deltshev, Christo, Naumova, Maria, Matevski, Dragan & Indzhov, Simeon, 2022, New taxonomic and faunistic data on the genus Zodarion Walckenaer, 1826 (Araneae: Zodariidae) in the Balkans, with the descriptions of two new species, pp. 247-261 in Zootaxa 5174 (3) on page 253, DOI: 10.11646/zootaxa.5174.3.3, http://zenodo.org/record/6986267, {"references":["Brignoli, P. M. (1984) Ragni di Grecia XII. Nuovi dati su varie famiglie (Araneae). Revue Suisse de Zoologie, 91 (2), 281 - 321. https: // doi. org / 10.5962 / bhl. part. 81881","Lazarov, S. (2005) Spiders (Araneae) from Maquises in South - West Bulgaria) - Part I. Acta Zoologica Bulgarica, 57 (2), 145 - 152.","Lazarov, S. (2007) Spiders (Araneae) from the Maleshevska Mountain (SW Bulgaria). Part I. Acta Zoologica Bulgarica, 59 (2), 133 - 144.","Kalushkov, P., Blagoev, G. & Deltshev, C. (2008) Biodiversity of Epigeic Spiders in Genetically Modified (Bt) and Conventional (non - Bt) Potato Fields in Bulgaria. Acta Zoologica Bulgarica, 60 (1), 61 - 69."]}

Zodarion weissi Deltshev & Naumova sp. n. Figs 61–75. Types. Holotype: ♂ (NMNHS-Zod-0004.1), BULGARIA: Sofia Mt., Kremikovtsi ward, N42.7882 ˚, E23.4812 ˚, 645 m, 23.06.2019, M. Naumova leg. Paratypes: 2♂, 2♀ (NMNHS-Zod-0004.2) same data as the holotype. Comparative material examined. Zodarion abantense Wunderlich, 1980 (♂ holotype, 1♀ paratype (SMF 30055, 58853), TURKEY, Bolu village, Abant Mountains, 29.04– 20.05.1976, H. Korge leg.; Z. pirini (1♂, 2♀ (IBER), BULGARIA, Eastern Rhodopes Mts, Blagun village, N41.3698 ˚, E25.7515 ˚, 423 m, 24.05.2018, M. Naumova leg.; Z. turcicum Wunderlich, 1980 (1♂ (IBER), BULGARIA, South Black Sea coast, Primorsko resort, N42.2544 ˚, E27.7487 ˚, 3 m, 27.04– 07.06.2016, T. Teofilova, T. Ljubomirov & N. Karaivanov leg.). Diagnosis. Zodarion weissi sp. n. belongs to the germanicum species group (sensu Bosmans 2009) and resembles Z. aurorae Weiss, 1982. Males differ by the following traits: tegular projection elongated triangular in lateral view in Z. weissi sp. n., vs. oblong with a broad rounded tip in Z. aurorae; basal tegular bulge reaching to the half of the tegular projection in Z. weissi sp. n. (Figs 63–65, 71, 72), vs. reaching more than 2/3 of the tegular projection length in Z. aurorae (see in Weiss 1982: 80, fig. 1) in lateral view. Females differ as follows: posterior margin of the epigynal plate with triangular incision with broad and shallow hood, 2.5–3 times wider than long in Z. weissi sp. n. (Figs 68–70, 73–75), vs. narrow and deep anterior hood, clearly longer than wide in Z. aurorae (see in Weiss 1982: 80, fig. 3). Etymology. The species is dedicated to Ingmar Weiss (Grafenau, Germany) a well-known arachnologist who has great contribution to araneological research in Romania. Description. Male (holotype). Total length 3.3. Carapace 1.54 long, 1.22 wide; sternum 0.9 long, 0.72 wide; chelicerae 0.54 long, 0.32 wide; clypeus 0.28 high. Eye sizes and interdistances: AME 0.11, ALE 0.07, PME 0.07, PLE 0.07, AME–AME 0.07, ALE–PLE 0.03, PME–PME 0.22. Abdomen 1.8 long. Legs: I—5.22 (1.26, 0.54, 1.26, 1.26, 0.9); II—4.86 (1.15, 0.54, 1.26, 1.08, 9.82); III—4.46 (1.26, 0.36, 0.97, 1.15, 0.72); IV—5.97 (1.69, 0.54, 1.22, 1.62, 0.9). Color (Fig. 61, 62): Carapace, sternum and chelicerae brown. Abdomen dark sepia-brown with a thick U-shaped, ventrolateral whitish spot on each side. Spinnerets bright, uniform in color. Coxae whitish, first couple light-brown. Other segments of the legs brown to brown-yellow, without annulations. Palp as in Figs 63–65, 71, 72. Femur 0.54 long, 0.18 wide, brown; patella 0.18 long, 0.18 wide, beige; tibia 0.11 long, 0,18 wide, brown, with elongated triangular retrolateral apophysis 0.18 long. Cymbium 0.65 long, 0.48 wide, brown with lightening tip; bulb 0.36 long, 0.29 wide. Tegulum with large basal bulge extended to the half of the tegular projection in prolateral view; tegular projection 0.18 long, 0.11 wide, elongated triangle in lateral view, perpendicular to the tegular surface, apically thinned; retinaculum 0.11 long, 0.07 wide, rhomboid-shaped, same sized as the bulge in ventral view; embolus thin, apically pointed, originating at a 9 o’clock position and terminating at a 1 o’clock position. Female. Total length 4.78. Carapace 1.9 long, 1.26 wide; sternum 1.08 long, 0.82 wide; chelicerae 0.54 long, 0.32 wide; clypeus 0.36 high. Eye sizes and interdistances:AME 0.09, ALE 0.06, PME 0.05, PLE 0.05, AME–AME 0.02, ALE–PLE 0.02, PME–PME 0.13. Abdomen 2.88 long. Legs: I—5.4 (1.36, 0.61, 1.26, 1.26, 0.9); II—5.07 (1.33, 0.61, 1.08, 1.26, 0.79); III—5.11 (1.33, 0.54, 1.08, 1.44, 0.72); IV—6.78 (1.8, 0.61, 1.62, 1.8, 0.9). Coloration as in male. Epigyne (Figs 68–70, 73–75): epigynal plate 1.5 times wider than long, broadly ovate. Posterior margin with large triangular incision with a base as wide as a distance between spermathecae and a tip reaching their half. Hood wide and shallow. Spermathecae rounded, separated by 2.5 diameters. Ecology. Found only in June, under stones in an open, dry and sunny place with very sparse low vegetation. Distribution. Known only from the type locality in Sofia Mt., Bulgaria (Fig. 76).