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The prevalence of nine indicator lichens found on trees in Northern Europe and Western Russia was used for monitoring air quality. The 4200 mile route of the survey went through eight countries. Surveys were carried out in cities, towns, countryside and forests, and along motorways. The author has conducted tree lichen surveys with pupils from schools and orphanages in Western Russia for many years. As expected, the calculated "lichen score" at each site correlated negatively with the published nitrogen dioxide levels in that area, and nitrogen-sensitive lichens were found to be most prevalent in forests and less-industrialised locations close to the Baltic Sea. (Contains 14 figures and 1 table.)

Copyright of Studia Ecologiae et Bioethicae is the property of Uniwerystet Kardynala Stefana Wyznskiege w Warzawie and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)

In an effort to ameliorate the impacts of climate change, forest managers in Central Europe increasingly turn to conifer species that produce higher yields and are better adapted to projected future climatic conditions. Though small mammals are an important component of the forest ecosystem, the impacts of enriching native broadleaf forests with conifers on small mammal communities are not well understood. We conducted mark–recapture surveys of small mammals to ascertain differences in their community structure among stands of two conifers (native Norway spruce Picea abies and non-native Douglas fir Pseudotsuga menziesii) and the dominant broadleaf in the region, European beech (Fagus sylvatica). After estimating the density of two common species, the yellow-necked mouse Apodemus flavicollis and bank vole Clethrionomys glareolus, we found that the population density of each is positively related to the proportion of beech and negatively to the proportion of conifers in each stand, though these effects of stand composition are smaller than the positive effect of herb cover. Increasing Norway spruce proportion reduced monthly survival of small mammals, while Douglas fir proportion had a positive effect on survival. We conclude that the two conifer species have similar impacts on small mammal density, though overall small mammal survival was significantly lower on plots with Norway spruce. This suggests that increasing the proportion of Douglas fir at the expense of Norway spruce may be possible without significantly changing local patterns of small mammal population density, but further research is necessary to elucidate the exact impacts of these two conifer species on small mammal demography and behavior. [ABSTRACT FROM AUTHOR]

The aim of this study was to understand the needs of European users for successful development and uptake of a digital land-based knowledge platform. The 'discover and define' phases of the design thinking double diamond framework were applied to inform the construction of 16 user-personas and journeys. Qualitative and quantitative data were collected from users across Europe using a multi-step approach and thematically analysed. User-personas and associated user journeys present useful methods for understanding the requirements of key stakeholders in the land-based sectors to aid the development of a digital knowledge reservoir, tailored to their needs. Problem urgency plays a key role in stakeholder touchpoint decisions, with acute problems approached by direct calls or interactions with important advisors. Digital platforms are more likely to be used for chronic problems or general knowledge exchange and upskilling. The touchpoints, challenges and emotions of users are not significantly linked to geographical location but are more likely to be influenced by age, gender and existing digital knowledge. Design thinking can be used to understand the needs of stakeholders in the land-based sectors for the development of tools, products and services as well as understanding specific knowledge requirements. This study provides a theoretical framework for understanding the digital knowledge needs of different land-based users. The development of user-personas and journeys is limited in the land-based sectors, and this research provides a framework for use. [ABSTRACT FROM AUTHOR]

Disturbances catalyse change in forest ecosystems, and a climate-driven increase in disturbance activity could accelerate forest reorganization. Here, we studied post-disturbance forests after the biggest pulse of tree mortality in Central Europe in at least 170 years, caused by drought and bark beetle (Scolytinae) outbreaks in 2018-2020. Our objectives were to characterize the early state of tree regeneration after mortality, quantify patterns of reorganization relative to undisturbed reference conditions and assess how management and patch size affect forest reorganization after disturbance. We surveyed 1244 plots in 120 patches under managed (salvage-logged, often planted) and unmanaged (deadwood remaining on site, no planting) conditions in Germany. We found that regeneration density on disturbed sites was high (median 11 897 stems ha -1 ), resulting from a cohort of advance regeneration. Disturbances were strong drivers of change, with indications for resilience on only 36.3% of patches. Reassembly (i.e. a change in species composition) was the dominant pattern of reorganization (61.5%), and Picea abies forests changed most strongly. Post-disturbance management facilitated forest change, particularly promoting a change in species composition. The strength of reorganization increased with patch size. We conclude that the recent wave of tree mortality will likely accelerate forest change in Central Europe.

Global-scale canopy height mapping is an important tool for ecosystem monitoring and sustainable forest management. Various studies have demonstrated the ability to estimate canopy height from a single spaceborne multispectral image using end-to-end learning techniques. In addition to texture information of a single-shot image, our study exploits multitemporal information of image sequences to improve estimation accuracy. We adopt a convolutional variant of a long short-term memory (LSTM) model for canopy height estimation from multitemporal instances of Sentinel-2 products. Furthermore, we utilize the deep ensembles technique for meaningful uncertainty estimation on the predictions and postprocessing isotonic regression model for calibrating them. Our lightweight model (${\sim }320{\mathrm {k}}$ trainable parameters) achieves the mean absolute error (MAE) of $1.29 ~{\mathrm {m}}$ in a European test area of $79 ~{\mathrm {km}}^{2}$. It outperforms the state-of-the-art methods based on single-shot spaceborne images as well as costly airborne images while providing additional confidence maps that are shown to be well calibrated. Moreover, the trained model is shown to be transferable in a different country of Europe using a fine-tuning area of as low as ${\sim }2 ~{\mathrm {km}}^{2}$ with ${\mathrm {MAE}}=1.94 {\mathrm {m}}$. [ABSTRACT FROM AUTHOR]

Forests provide important ecosystem services (ESs), including climate change mitigation, local climate regulation, habitat for biodiversity, wood and non-wood products, energy, and recreation. Simultaneously, forests are increasingly affected by climate change and need to be adapted to future environmental conditions. Current legislation, including the European Union (EU) Biodiversity Strategy, EU Forest Strategy, and national laws, aims to protect forest landscapes, enhance ESs, adapt forests to climate change, and leverage forest products for climate change mitigation and the bioeconomy. However, reconciling all these competing demands poses a tremendous task for policymakers, forest managers, conservation agencies, and other stakeholders, especially given the uncertainty associated with future climate impacts. Here, we used process-based ecosystem modeling and robust multi-criteria optimization to develop forest management portfolios that provide multiple ESs across a wide range of climate scenarios. We included constraints to strictly protect 10% of Europe's land area and to provide stable harvest levels under every climate scenario. The optimization showed only limited options to improve ES provision within these constraints. Consequently, management portfolios suffered from low diversity, which contradicts the goal of multi-functionality and exposes regions to significant risk due to a lack of risk diversification. Additionally, certain regions, especially those in the north, would need to prioritize timber provision to compensate for reduced harvests elsewhere. This conflicts with EU LULUCF targets for increased forest carbon sinks in all member states and prevents an equal distribution of strictly protected areas, introducing a bias as to which forest ecosystems are more protected than others. Thus, coordinated strategies at the European level are imperative to address these challenges effectively. We suggest that the implementation of the EU Biodiversity Strategy, EU Forest Strategy, and targets for forest carbon sinks require complementary measures to alleviate the conflicting demands on forests., (© 2024 The Author(s). Global Change Biology published by John Wiley & Sons Ltd.)

Canopy openings are increasing in Europe's forests, yet the contributions of anthropogenic and ecological agents of disturbance to this increase remain debated. Here we attribute the root cause of all stand-replacing canopy disturbances identified for Europe in the period 1986-2020 from Landsat data (417,000 km²), distinguishing between planned and unplanned canopy openings (i.e., disturbance by human land use versus by wind, bark beetles, and wildfire). We show that canopy openings by humans dominate the European forest disturbance regime, accounting for 82% of the area disturbed. Both planned and unplanned canopy openings increased in the early 21st century (+24% and +30% relative to the late 20th century). Their changes are linked, with simultaneous increases in planned and unplanned canopy openings on 68% of Europe's forest area. We conclude that an important direction for tackling disturbance change in policy and management is to break the link between planned and unplanned canopy openings in Europe's forests., (© 2024. The Author(s).)

Large pulses of tree mortality have ushered in a major reorganization of Europe's forest ecosystems. To initiate a robust next generation of trees, the species that are planted today need to be climatically suitable throughout the entire twenty-first century. Here we developed species distribution models for 69 European tree species based on occurrence data from 238,080 plot locations to investigate the option space for current forest management in Europe. We show that the average pool of tree species continuously suitable throughout the century is smaller than that under current and end-of-century climate conditions, creating a tree species bottleneck for current management. If the need for continuous climate suitability throughout the lifespan of a tree planted today is considered, climate change shrinks the tree species pool available to management by between 33% and 49% of its current values (40% and 54% of potential end-of-century values), under moderate (Representative Concentration Pathway 2.6) and severe (Representative Concentration Pathway 8.5) climate change, respectively. This bottleneck could have strong negative impacts on timber production, carbon storage and biodiversity conservation, as only 3.18, 3.53 and 2.56 species of high potential for providing these functions remain suitable throughout the century on average per square kilometre in Europe. Our results indicate that the option space for silviculture is narrowing substantially because of climate change and that an important adaptation strategy in forestry-creating mixed forests-might be curtailed by widespread losses of climatically suitable tree species., (© 2024. The Author(s), under exclusive licence to Springer Nature Limited.)

Throughout history, humans have relied on wood for constructions, tool production or as an energy source. How and to what extent these human activities have impacted plant abundance and composition over a long-term perspective is, however, not well known. To address this knowledge gap, we combined 44,239 precisely dated tree-ring samples from economically and ecologically important tree species (spruce, fir, pine, oak) from historical buildings, and pollen-based plant cover estimates using the REVEALS model from 169 records for a total of 34 1° × 1° grid cells for Central Europe. Building activity and REVEALS estimates were compared for the entire study region (4-15°E, 46-51°N), and for low (<500 m asl) and mid/high elevations (≥500 m asl) in 100-year time windows over the period 1150-1850. Spruce and oak were more widely used in wooden constructions, amounting to 35 % and 32 %, respectively, compared to pine and fir. Besides wood properties and species abundance, tree diameters of harvested individuals, being similar for all four species, were found to be the most crucial criterion for timber selection throughout the last millennium. Regarding land use changes, from the 1150-1250's onwards, forest cover generally decreased due to deforestation until 1850, especially at lower elevations, resulting in a more heterogeneous landscape. The period 1650-1750 marks a distinct change in the environmental history of Central Europe; increasing agriculture and intense forest management practices were introduced to meet the high demands of an increasing population and intensifying industrialization, causing a decrease in palynological diversity, especially at low elevations. Likely the characteristic vegetation structure and composition of contemporary landscapes originated from that period. We further show that land use has impacted vegetation composition and diversity at an increasing speed leading to a general homogenization of landscapes through time, highlighting the limited environmental benefits of even-aged plantation forestry., Competing Interests: Declaration of competing interest The authors declare that they have no known competing financial interests or personal relationships that could have appeared to influence the work reported in this paper., (Copyright © 2024 The Authors. Published by Elsevier B.V. All rights reserved.)

Ash dieback, acute oak decline (AOD), and Xylella Fastidiosa are emerging infectious diseases (EIDs) that have spread rapidly in European forests during the last decade. Quarantine measurements have mostly failed to repress the outbreaks and millions of trees have already been infected. Identifying infected trees in a nondestructive manner is of high importance for monitoring, managing, and preventing EIDs. The aim of this article is to examine the capabilities of ground penetrating radar (GPR) on evaluating the internal structure of tree trunks and detecting tree decay associated with EIDs. Traditionally used processing schemes tuned for GPR line acquisitions are modified accordingly to be compatible with the new measurement configurations. In particular, a detection framework is presented based on a modified Kirchhoff and a reverse-time migration. Both of the aforementioned methodologies are compatible with measurements taken along closed irregular curves assuming a homogeneous permittivity distribution. To that extent, prior to migration, a novel focal criterion is used that estimates the bulk permittivity of the host medium from the measured B-scans. The suggested detection scheme is successfully tested on both numerical and laboratory measurements, indicating that GPR has the potential to become a coherent and practical tool for detecting tree decay associated with EIDs. [ABSTRACT FROM AUTHOR]

Climate change adaptation and mitigation strategies (CCAMS) are changes to the management of production forests motivated by the need to mitigate climate change, or adapt production forests to climate change risks. Sweden is employing CCAMS with unclear implications for biodiversity and forest ecosystem services (ES). Here, we synthesized evidence from 51 published scientific reviews, to evaluate the potential implications for biodiversity and a range of provisioning, regulating, and cultural ES, from the adoption of CCAMS relative to standard forestry practice. The CCAMS assessed were the adoption of (i) mixed-species stands, (ii) continuous cover forestry, (iii) altered rotation lengths, (iv) conversion to introduced tree species, (v) logging residue extraction, (vi) stand fertilization, and (vii) altered ditching/draining practices. We highlight the complexity of biodiversity and ES outcomes, identify knowledge gaps, and emphasize the importance of evidence-based decision making and landscape-scale planning when navigating choices involving the widespread adoption of CCAMS., (© 2023. The Author(s).)

Natural forest expansion (NFE), that is, the establishment of secondary forest on non-forested land through natural succession, has substantially contributed to the widespread expansion of forests in Europe over the last few decades. So far, EU policies have largely neglected the potential of NFE for meeting policy objectives on restoration. Synthesising recent interdisciplinary research, this paper assesses the challenges and opportunities of NFE in view of contributing to European forest and ecosystem restoration. Specifically, we discuss the potential for supporting climate change mitigation and adaptation, biodiversity conservation, and forestry and economic use, summarize the current knowledge about societal perceptions and the policymaking on NFE, and make policy recommendations to better use the potential of NFE. We conclude that NFE has the potential to contribute to the European restoration policy agenda if local contexts and possible trade-offs are properly considered., (© 2023. The Author(s).)

The growing demand for timber, in particular for renewable energy, increases pressures on global forests and requires a robust monitoring system to ensure sustainability. This article takes a first step toward more systemic monitoring by asking how the global use of forests by EU consumers can be accounted for. Specifically, this article builds on and develops the method of global land use accounting to account for the EU-27's consumption of primary timber between 2002 and 2011 in terms of both volume and forest area. It assesses international trade flows for around 100 commodities and converts them into a volume of primary raw timber based on conversion values. Results reveal that both imports and exports increased over the assessed time period, with primary EU-27 timber estimated to be around 1 m 3 /cap in 2011. Gaps, uncertainty and a lack of harmonization regarding especially trade data and conversion values are key challenges to further improving the robustness of the method and reliability of results. Future research may focus on improving the method to address in particular recycled and recovered flows as well as the question of whether area or volume is the most appropriate metric for further development of a forest footprint indicator. [ABSTRACT FROM AUTHOR]

This article aims at finding statistics about the forestry sector, both in terms of resources and exploitation rights in the European Union, in 2015, at the territory there were approximately 185 million hectares of forests and other woodland, accounting for about 40% of the total area. Currently, the inventory of forest resources as well as quantitative and qualitative characteristics can capitalize by using various sources, from measurements in situ national administrative statistics. Forest management should ensure the organization and management of forests to their most effective multifunctional state, according to multiple environmental and socio-economic objectives of forestry based on the concept of functional zoning and promoting the concept of systemic forest management [ABSTRACT FROM AUTHOR]

The Natura 2000 appropriate assessment for impacting projects or plans under Article 6(3) HD is the central statutory instrument for the protection of Sites of Community Importance (SCI) and the Special Protection Areas (SPA). The decisive factor in whether or not an appropriate assessment is required depends on the question of whether a project or plan is present within the meaning of Article 6(3) of the Habitats Directive 92/43/EEC1 (HD). The Habitats Directive does not define these terms in any more detail, which is why they must be specified more closely through interpretation. This paper will present the definitions given by the European Court of Justice (ECJ)2 case law and national courts like the German Federal Administrative Court (BVerwG)3 and discuss the consequences and practical scope of the terms. The focus of the following investigation will be on the term "project". This is because for "plans", the envisaged projects are essentially also decisive, given that only these can have significant adverse effects on the conservation objectives. There are a variety of questions regarding when a human activity constitutes a project and under which conditions Member States could exempt activities from the requirement for an assessment. This article will start with an outline of the temporal scope of the appropriate assessment and, following this, briefly explore the scope of plans or projects directly connected with or necessary to the management of the site, as they are not the subject of an assessment. [ABSTRACT FROM AUTHOR]

Prolonged drought and susceptibility to biotic stressors induced an extensive calamity in Norway spruce (Picea abies (L.) Karst.) and widespread crown defoliation in European beech (Fagus sylvatica L.) in Central Europe. For future management decisions, it is crucial to link changes in canopy cover to site conditions. However, current knowledge on the role of soil properties for drought-induced forest disturbance is limited due to the scarcity and low spatial resolution of soil information. We present a fine-scale assessment on the role of soil properties for forest disturbance in Norway spruce and European beech derived from optical remote sensing. A forest disturbance modeling framework based on Sentinel-2 time series was applied on 340 km 2 in low mountain ranges of Central Germany. Spatio-temporal information on forest disturbance was calculated at 10 m spatial resolution in the period 2019-2021 and intersected with high-resolution soil information (1:10,000) based on roughly 2850 soil profiles. We found distinct differences in disturbed area, depending on soil type, texture, stoniness, effective rooting depth and available water capacity (AWC). For spruce, we found a polynomial relationship between AWC (R 2  = 0.7) and disturbance, with highest disturbed area (65 %) for AWC between 90 and 160 mm. Interestingly, we found no evidence for generally higher disturbance on shallow soils, although stands on the deepest soils were significantly less affected. Noteworthy, sites affected first did not necessarily exhibit highest proportions of disturbed area post-drought, indicating recovery or adaptation. We conclude that site- and species-specific understanding of drought impacts benefits from a combination of remote sensing and fine-scale soil information. Since our approach revealed which sites were affected first and most, it qualifies for prioritizing in situ monitoring activities to most vulnerable stands in acute drought conditions as well as for developing long-term strategies for reforestation and site-specific risk assessment for precision forestry., Competing Interests: Declaration of competing interest The authors declare that they have no known competing financial interests or personal relationships that could have appeared to influence the work reported in this paper., (Copyright © 2023 Elsevier B.V. All rights reserved.)

Copyright of International Forestry Review is the property of Commonwealth Forestry Association and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)

Abstract: The study determined the performance of small-scale commercial firewood processing operations under the typical work conditions of Southern Europe. In particular, five units were tested, fed with the same 2.1-m long beech (Fagus sylvatica L.) logs. All machines were tested with sorted and unsorted logs. Productivity varied between 1.1 and 2.1th−1, and cost between 20 and 39€t−1. There were significant differences between machines, which may partly be attributed to operator effect. Feeding the machines with sorted logs had a significant effect on the productivity of all machines on test, increasing productivity by 40% and reducing cost by 34%. Fuel use varied between 1.3 and 2.8lt−1. The energy balance was always very favorable. The ration between output and input was never smaller than 59 to 1 and peaked at 130 to 1. In other words, processing required about 1% of the energy contained in the firewood — or 1.7% in the worst case. The productivity figures reported in this experiment were much lower than reported for Northern Europe, which seems to confirm the significant effect of regional work conditions – especially different wood species – on firewood processing performance. [Copyright &y& Elsevier]

Accelerating introductions of forest insects challenge decision-makers who might or might not respond with surveillance programs, quarantines, eradication efforts, or biological control programs. Comparing ecological controls on indigenous vs. introduced populations could inform responses to new introductions. We studied the European woodwasp, Sirex noctilio, which is not a pest in its native forests, is a serious invasive pest in the southern hemisphere, and now has an uncertain future in North America after its introduction there. Indigenous populations of S. noctilio (in Galicia, Spain) resembled those in New York in that S. noctilio were largely restricted to suppressed trees that were also dying for other reasons, and still only some dying trees showed evidence of S. noctilio: 20–40% and 35–51% in Galicia and New York, respectively. In both areas, P. sylvestris (native to Europe) was the species most likely to have attacks in non-suppressed trees. P. resinosa, native to North America, does not appear dangerously susceptible to S. noctilio. P. radiata, which sustains high damage in the southern hemisphere, is apparently not innately susceptible because in Galicia it was less often used by native S. noctilio than either native pine (P. pinaster and P. sylvestris). Silvicultural practices in Galicia that maintain basal area at 25–40 m2/ha limit S. noctilio abundance. More than 25 species of other xylophagous insects feed on pine in Galicia, but co-occurrences with S. noctilio were infrequent, so strong interspecific competition seemed unlikely. Evidently, S. noctilio in northeastern North America will be more similar to indigenous populations in Europe, where it is not a pest, than to introduced populations in the southern hemisphere, where it is. However, S. noctilio populations could behave differently when they reach forests of the southeastern U.S., where tree species, soils, climate, ecology, management, and landscape configurations of pine stands are different. [ABSTRACT FROM AUTHOR]

Gene flow is a key factor in the evolution of species, influencing effective population size, hybridisation and local adaptation. We analysed local gene flow in eight stands of white oak (mostly Quercus petraea and Q. robur, but also Q. pubescens and Q. faginea) distributed across Europe. Adult trees within a given area in each stand were exhaustively sampled (range [239, 754], mean 423), mapped, and acorns were collected ([17,147], 51) from several mother trees ([3], [47], 23). Seedlings ([65,387], 178) were harvested and geo-referenced in six of the eight stands. Genetic information was obtained from screening distinct molecular markers spread across the genome, genotyping each tree, acorn or seedling. All samples were thus genotyped at 5–8 nuclear microsatellite loci. Fathers/parents were assigned to acorns and seedlings using likelihood methods. Mating success of male and female parents, pollen and seed dispersal curves, and also hybridisation rates were estimated in each stand and compared on a continental scale. On average, the percentage of the wind-borne pollen from outside the stand was 60%, with large variation among stands (21–88%). Mean seed immigration into the stand was 40%, a high value for oaks that are generally considered to have limited seed dispersal. However, this estimate varied greatly among stands (20–66%). Gene flow was mostly intraspecific, with large variation, as some trees and stands showed particularly high rates of hybridisation. Our results show that mating success was unevenly distributed among trees. The high levels of gene flow suggest that geographically remote oak stands are unlikely to be genetically isolated, questioning the static definition of gene reserves and seed stands. [ABSTRACT FROM AUTHOR]

Traditional grazing areas in Europe have declined substantially over the last century. Specifically, in northern Fennoscandia, the grazing land is disturbed by cumulative land-use pressures. Here we analysed the configuration of the grazing land for reindeer and sheep in northern Fennoscandia in relation to the concurrent land-use pressures from tourism, road and railway networks, forestry, industrial and wind energy facilities, together with predator presence and climate change. Our results show that 85% of the region is affected by at least one land-use pressure and 60% is affected by multiple land-use pressures, co-occurring with predator presence and rising temperatures. As such, a majority of the grazing land is exposed to cumulative pressures in northern Fennoscandia. We stress that, if the expansion of cumulative pressures leads to grazing abandonment of disturbed areas and grazing intensification in other areas, it could irreversibly change northern vegetation and the Fennoscandian mountain landscape., (© 2022. The Author(s).)

Forest age structure has become younger and the amount of old forest has decreased due to intensive forestry in Northern Europe. Consequently, large forest-dwelling hawks may lack nest sites as they need large trees to support their nests. Construction of artificial nests has been a globally adopted conservation measure to provide nest sites for several raptors, but the impact of this practice has seldom been evaluated. We compared the nesting success of northern goshawk Accipiter gentilis, common buzzard Buteo buteo, and honey buzzard Pernis apivorus in natural and artificial nests based on long-term data in Finland. The probability of successful nesting of goshawk and common buzzard was lower in artificial nests, whereas no difference was found in the nesting success of honey buzzard between the nest types. Brood size did not differ between natural and artificial nests in successful nests of any of the three species. Finally, distances to anthropogenic features had no effect on nesting success. These results indicate that hawks switching from natural to artificial nests are potentially lured to an ecological trap with higher risk of nesting failure. However, breeding at artificial nests, even with lower success than in natural nests, can still add to the population, if otherwise nonbreeding individuals settle to nest. It should be investigated which part of the population breeds in artificial nests and whether the site selection for artificial nests needs revision. Our results emphasize the importance of monitoring and evaluation of well-intentioned conservation efforts. [ABSTRACT FROM AUTHOR]

The ecological impacts of changing forest management practices in Europe are poorly understood despite European forests being highly managed. Furthermore, the effects of potential drivers of forest biodiversity decline are rarely considered in concert, thus limiting effective conservation or sustainable forest management. We present a trait-based framework that we use to assess the detrimental impact of multiple land-use and management changes in forests on bird populations across Europe. Major changes to forest habitats occurring in recent decades, and their impact on resource availability for birds were identified. Risk associated with these changes for 52 species of forest birds, defined as the proportion of each species' key resources detrimentally affected through changes in abundance and/or availability, was quantified and compared to their pan-European population growth rates between 1980 and 2009. Relationships between risk and population growth were found to be significantly negative, indicating that resource loss in European forests is an important driver of decline for both resident and migrant birds. Our results demonstrate that coarse quantification of resource use and ecological change can be valuable in understanding causes of biodiversity decline, and thus in informing conservation strategy and policy. Such an approach has good potential to be extended for predictive use in assessing the impact of possible future changes to forest management and to develop more precise indicators of forest health. [ABSTRACT FROM AUTHOR]

Abstract: Industrial forestry has caused large biodiversity changes in European boreal forests. One recently introduced conservation measure in production forestry is retention of trees at clearcutting to benefit flora and fauna. Aspen Populus tremula is often retained for conservation purposes since it is a key tree species for biodiversity with many associated species, a number of which are red-listed. Still, the importance to biodiversity of aspen trees retained at harvest is largely unknown. In 1994, a transplantation experiment with the old-growth forest indicator lichen Lobaria pulmonaria was set up on 280 aspens at 35 sites in east-central Sweden with a total of 1120 transplants, with the aim to assess the habitat suitability of retained aspens following harvest. After 14years 23% of L. pulmonaria transplants remained, with a significantly higher survival on retained aspens than on aspens in the surrounding forest, especially on the northern side of stems. Transplants were also more vital on northern than on southern sides of stems. There was no difference in survival or vitality of transplants between dispersed aspens and aspens in groups. Results largely agreed with a re-inventory made already after two years but the importance of the north side of retention trees became evident for species survival only after 14years, indicating that to gain deeper insights longer time-spans may be necessary. This study, which is the longest lichen transplantation time-series from a well replicated experiment so far published, shows that retention of trees at harvest may be an efficient conservation action. [Copyright &y& Elsevier]

Developing adaptation measures in forestry is an urgent task because the forests regenerated today will have to cope with climate conditions that may drastically change during the life of the trees in the stand. This paper presents a comprehensive review of potential adaptation options in forestry in Europe based on three pillars: a review of the scientific literature, an analysis of current national response strategies, and an expert assessment based on a database compiled in the COST Action ECHOES (Expected Climate Change and Options for European Silviculture). The adaptation measures include responses to both risks and opportunities created by climate change and address all stages of forestry operations. Measures targeted to reduce vulnerability to climate change may either aim to reduce forest sensitivity to adverse climate change impacts or increase adaptive capacity to cope with the changing environmental conditions. Adaptation measures mitigating drought and fire risk such as selection of more drought resistant species and genotypes are crucial. For adaptation to be successful it is of the utmost importance to disseminate the knowledge of suitable adaptation measures to all decision makers from the practice to the policy level. The analysis of the ECHOES database demonstrates that this challenge is well recognized in many European countries. Uncertainty about the full extent of climate change impacts and the suitability of adaptation measures creates a need for monitoring and further research. A better understanding of how to increase adaptive capacity is also needed, as well as regional vulnerability assessments which are crucial for targeting planned adaptation measures. [ABSTRACT FROM AUTHOR]

Abstract: The purpose of this paper is to provide estimated Armington elasticities for selected European countries and for three forest biomass commodities of main interest in many energy models: roundwood, chips & particles and wood residues. The Armington elasticity is based on the assumption that a specific forest biomass commodity is differentiated by its origin. The statistically significant estimated Armington elasticities range from 0.52 for roundwood in Hungary to approximately 4.53 for roundwood in Estonia. On average, the statistically significant Armington elasticity for chips & particles over all countries is 1.7 and for wood residues and roundwood 1.3 and 1.5, respectively. These elasticities can provide benchmark values for simulation models trying to assess trade patterns of forest biomass commodities and energy policy effects for European countries or for the EU as a whole. [ABSTRACT FROM AUTHOR]

Risks can generally be described as the combination of hazard, exposure and vulnerability. Using this framework, we evaluated the historical and future development of risk of fire and wind damage in European forestry at the national level. Fire risk is expected to increase, mainly as a consequence of an increase in fire hazard, defined as the Fire Weather Index in summer. Exposure, defined as forest area, is expected to increase slightly as a consequence of active afforestation and abandonment of marginal agricultural areas. Adaptation options to fire risk should therefore aim to decrease the vulnerability, where a change in tree species from conifers to broadleaves had most effect. Risk for wind damage in forests is expected to increase mainly as a consequence of increase in exposure (total growing stock) and vulnerability (defined by age class and tree species distribution). Projections of future wind climate indicate an increase in hazard (storminess) mainly over Western Europe. Adaptation options should aim to limit the increase in exposure and vulnerability. Only an increase in harvest level can stop the current build-up of growing stock, while at the same time it will lower vulnerability through the reduction of the share of old and vulnerable stands. Changing species from conifers to broadleaves helps to reduce vulnerability as well. Lowering vulnerability by decreasing the rotation length is only effective in combination with a high demand for wood. Due to data limitations, no forecast of future fire area or damaged timber amount by storms was possible. [ABSTRACT FROM AUTHOR]

Overviewing the European carbon (C), greenhouse gas (GHG), and non-GHG fluxes, gross primary productivity (GPP) is about 9.3 Pg yr−1, and fossil fuel imports are 1.6 Pg yr−1. GPP is about 1.25% of solar radiation, containing about 360 × 1018 J energy – five times the energy content of annual fossil fuel use. Net primary production (NPP) is 50%, terrestrial net biome productivity, NBP, 3%, and the net GHG balance, NGB, 0.3% of GPP. Human harvest uses 20% of NPP or 10% of GPP, or alternatively 1‰ of solar radiation after accounting for the inherent cost of agriculture and forestry, for production of pesticides and fertilizer, the return of organic fertilizer, and for the C equivalent cost of GHG emissions. C equivalents are defined on a global warming potential with a 100-year time horizon. The equivalent of about 2.4% of the mineral fertilizer input is emitted as N2O. Agricultural emissions to the atmosphere are about 40% of total methane, 60% of total NO-N, 70% of total N2O-N, and 95% of total NH3-N emissions of Europe. European soils are a net C sink (114 Tg yr−1), but considering the emissions of GHGs, soils are a source of about 26 Tg CO2 C-equivalent yr−1. Forest, grassland and sediment C sinks are offset by GHG emissions from croplands, peatlands and inland waters. Non-GHGs (NH3, NO x) interact significantly with the GHG and the C cycle through ammonium nitrate aerosols and dry deposition. Wet deposition of nitrogen (N) supports about 50% of forest timber growth. Land use change is regionally important. The absolute flux values total about 50 Tg C yr−1. Nevertheless, for the European trace-gas balance, land-use intensity is more important than land-use change. This study shows that emissions of GHGs and non-GHGs significantly distort the C cycle and eliminate apparent C sinks. [ABSTRACT FROM AUTHOR]

Abstract: This study compiles and summarizes the existing knowledge about observed and projected impacts of climate change on forests in Europe. Forests will have to adapt not only to changes in mean climate variables but also to increased variability with greater risk of extreme weather events, such as prolonged drought, storms and floods. Sensitivity, potential impacts, adaptive capacity, and vulnerability to climate change are reviewed for European forests. The most important potential impacts of climate change on forest goods and services are summarized for the Boreal, Temperate Oceanic, Temperate Continental, Mediterranean, and mountainous regions. Especially in northern and western Europe the increasing atmospheric CO2 content and warmer temperatures are expected to result in positive effects on forest growth and wood production, at least in the short–medium term. On the other hand, increasing drought and disturbance risks will cause adverse effects. These negative impacts are very likely to outweigh positive trends in southern and eastern Europe. From west to east, the drought risk increases. In the Mediterranean regions productivity is expected to decline due to strongly increased droughts and fire risks. Adaptive capacity consists of the inherent adaptive capacity of trees and forest ecosystems and of socio-economic factors determining the capability to implement planned adaptation. The adaptive capacity in the forest sector is relatively large in the Boreal and the Temperate Oceanic regions, more constrained by socio-economic factors in the Temperate Continental, and most limited in the Mediterranean region where large forest areas are only extensively managed or unmanaged. Potential impacts and risks are best studied and understood with respect to wood production. It is clear that all other goods and services provided by European forests will also be impacted by climate change, but much less knowledge is available to quantify these impacts. Understanding of adaptive capacity and regional vulnerability to climate change in European forests is not well developed and requires more focussed research efforts. An interdisciplinary research agenda integrated with monitoring networks and projection models is needed to provide information at all levels of decision making, from policy development to the management unit. [Copyright &y& Elsevier]

This article reviews major impacts of climate change on agriculture and forestry. [ABSTRACT FROM AUTHOR]

A study was carried out to clarify the role of the fungus Chalara fraxinea in decline of Fraxinus excelsior, which is observed on a large scale in central and northern Europe with high incidence of tree mortality. The aims of this work were: (i) to check for the presence of C. fraxinea in various tissues of declining F. excelsior by agar culture isolations and by direct analysis of plant tissues using molecular techniques (DNA extraction, ITS-PCR, cloning, ITS sequencing and T-RFLP); (ii) to study fungal communities inhabiting tissues with symptoms; and (iii) to test the pathogenicity of C. fraxinea to F. excelsior. Chalara fraxinea was isolated from 93% of stem cankers, 91% of necrotic leaf stalks, 27–28% of bark wounds and 30% of visually healthy leaf stalks. Molecular analyses of necrotic leaves, leaf stalks and bark revealed the presence of 25 different fungal taxa, 14 of which were detected in all three types of tissue sample. Chalara fraxinea was the second most common species (61% of samples), and only Cryptococcus foliicola occurred more often (70%). All eight of the tested C. fraxinea isolates induced necroses in bark and cambium on each of 86 inoculated trees, and all controls remained healthy. Average length of necroses caused by different C. fraxinea strains varied from 4·2 to 8·9 cm, but the differences were statistically insignificant. Instead, differences in resistance of individual trees to C. fraxinea were observed. [ABSTRACT FROM AUTHOR]

Despite a large literature that addresses the gendered structure of forestry occupations in Europe and the USA, relatively little attention has been paid to these issues in Canada. In this paper, it is argued that policy makers and academics have used outdated statistics about forestry employment to shape policies and programs of government and industry. Use of these data has been to the general disadvantage of women working in the forest industry and in forestry communities. This paper draws mainly on three studies, spanning the mid-1990s to the mid-2000s. Each study relied on secondary sources such as Census data, government and industry documents, as well as primary sources such as interviews or questionnaires. On Vancouver Island, British Columbia, analysis of land reallocation policies and transition strategies for displaced workers in the 1990s did not consider women as forestry workers or consider how job losses and opportunities for retraining might vary between men and women. Consequently, gender-neutral transition strategies were more accessible for men than for women. A study of female forestry workers in Saskatchewan revealed there was very little knowledge about employment conditions for women in forestry. Women reported that they missed out on training and advancement opportunities because they lacked networks, childcare support or other benefits that might encourage them to improve their circumstances. Finally, a survey of forest sector advisory committees across the country revealed a very low rate of participation by women, even though those who did participate had some significantly different viewpoints about forestry than their male counterparts. In combination, these efforts suggest that improvements in the understanding of forestry employment would benefit forestry workers, both male and female, and ensure that all workers made lasting contributions to the industry. [ABSTRACT FROM AUTHOR]

Over the last 20 years, climate change has become an increasing concern for scientists, public opinions and policy makers. Due to the pervasive nature of its impacts for many important aspects of human life, climate change is likely to influence and be influenced by the most diverse policy or management choices. This is particularly true for those interventions affecting agriculture and forestry: they are strongly dependent on climate phenomena, but also contribute to climate evolution being sources of and sinks for greenhouse gases (GHG). This paper offers a survey of the existing literature assessing cost-effectiveness and efficiency of greenhouse gas mitigation strategies or the effects of broader economic reforms in the agricultural and forestry sectors. The focus is mainly on European countries. Different methodological approaches, research questions addressed and results are examined. The main findings are that agriculture can potentially provide emissions reduction at a competitive cost, mainly with methane abatement, while carbon sequestration seems more cost-effective with appropriate forest management measures. Afforestation, cropland management and bioenergy are less economically viable measures due to competition with other land use. Mitigation policies should be carefully designed either to balance costs with expected benefits in terms of social welfare. Regional variability is one of the main drawbacks to fully assess the cost-effectiveness of different measures. Integration of models to take into account both social welfare and spatial heterogeneity seems to be the frontier of the next model generation. [Copyright &y& Elsevier]

Abstract: Silvoarable agroforestry could promote use of trees on farms in Europe, but its likely effect on production, farm profitability, and environmental services is poorly understood. Hence, from 2001 to 2005, the Silvoarable Agroforestry for Europe project developed a systematic process to evaluate the biophysical and economic performance of arable, forestry, and silvoarable systems in Spain, France, and The Netherlands. A biophysical model called “Yield-SAFE” was developed to predict long-term yields for the different systems and local statistics and expert opinion were used to derive their revenue, costs, and pre- and post-2005 grant regimes. These data were then used in an economic model called “Farm-SAFE” to predict plot- and farm-scale profitability. Land equivalent ratios were greater than one, showing Yield-SAFE predicted that growing trees and crops in silvoarable systems was more productive than growing them separately. Pre-2005 grants in Spain and The Netherlands penalised silvoarable systems, but post-2005 grants were more equitable. In France, walnut and poplar silvoarable systems were consistently the most profitable system under both grant regimes. In Spain, holm oak and stone pine silvoarable systems were the least profitable system under pre-2005 grants, but only marginally less profitable than arable systems under post-2005 grants. In The Netherlands, low timber values and the opportunity cost of losing arable land for slurry manure application made silvoarable and forestry systems uncompetitive with arable systems under both grant regimes. [Copyright &y& Elsevier]

Abstract: Scenarios with either 3% or 5% of the productive forest stock in Western Europe set-aside for conservation are compared to a baseline case. The latter case is further compared to cases where the whole of Europe and North America join the 5% conservation program. The scenarios are created by employing a global forest sector model, EFI-GTM. The aggregate conservation impacts seem relatively low, mainly because international trade counterbalances regional shortages and because roundwood harvests are well below the annual forest growth in many European countries. Conservation increases the forest owners’ wood sales income. The 5% set-aside raises roundwood prices in Western Europe by 4% and decreases harvests by 3% from the base case levels during 2010–2020. The production of pulp and mechanical forest products then decrease by 1.4 and 1.7%, respectively. The paper market is affected least. The leakage impact of forest protection is most pronounced in Russian roundwood harvests. Simultaneous conservation in the whole of Europe raises the roundwood costs of the Western European forest industry, but it does not prevent the leakage of roundwood harvests to Russia. Conservation in North America has a low impact on the European roundwood market. [Copyright &y& Elsevier]

AbstractLake Peipsi is the fourth largest lake in Europe and the largest international lake in Europe. Lake Peipsi is shallow, eutrophic, biologically very productive and is bordered by many wetland areas along its coasts. Large peat deposits in the Lake Peipsi basin are protected under national legislation. The lake is most important as a resource for recreation and fisheries. The fish stock in Lake Peipsi is one of the richest in Europe. The most critical environmental issues for the Peipsi watershed are eutrophication and unregulated fishing in the lake. During the Soviet period, large collective farms dramatically accelerated eutrophication. Despite the drop in fertilizer use during the last 10 years and a decrease in the number of domestic animals in the watershed, the impact of the earlier agricultural pollution is still significant. Untreated sewage has also been dumped in the Narva River tributary as a result of an international dispute concerning water supply and wastewater treatment. People living in the Lake Peipsi region have faced traumatic changes as a result of the re-establishment of the border between Russia and Estonia in 1992. The establishment of an international border on the waters of the lake has resulted in fishermen losing access to many of their traditional fishing areas. Political and economic transformations of the last 10 years have seriously interrupted information exchange between different institutions and organizations concerned with the health of the lake. The Centre for Transboundary Cooperation facilitates international communication and cooperation between the Estonian and Russian governments and promotes public participation in environmental, social and economic development and decision-making. The centre is in the process of guiding the development of a watershed management plan for the Lake Peipsi watershed and a vision for watershed development that is truly sustainable. [ABSTRACT FROM AUTHOR]

Private forest owners are the main forest ownership group within Europe, and important conservation values have been found on their land. Yet, small plot sizes, societal heterogeneity, and structural changes impede developing and implementing effective conservation programs in private forests. We present a systematic literature review focusing on small-scale private forest owners and their perspectives on nature conservation by synthesizing research approaches, social-ecological drivers, and policy recommendations. Conservation perspectives were positively related to female gender, higher levels of education, formalized forest management, an active relation to the forest, and ecological values of the property. In contrast, high age, rural orientation, economic forest management factors, large parcel size, and economic and sentimental property values negatively influenced conservation perspectives. Applying a natural resource conflict management framework, we synthesized recommendations covering three dimensions: substance, procedure, relationship. Considering perspectives of small-scale private forest owners in current forestry decision-making has great potential to strengthen sustainable forest management that integrates nature conservation and resource use., (© 2021. The Author(s).)

In certain indigenous Picea abies Karst stands in the Thuringian Forest (Middle­Europe), the incidence of butt rot and die-back of saplings by Armillaria mellea (Vahl ex Fr.) Karst is conspicuously low in spite of a general spread of this fungus on stumps of clear-fellings. The majority of 43 isolates of A. mellea recovered from 13 highland infection centres were unable to form the brown type of rhizomorphs (BR) in soil but could form the white type rhizomorphs (WR) on sterile substrates. In contrast, 17 isolates from lowland hardwood stands of average disease incidence produced both types of rhizomorphs on both substrates. Results of inoculation of exercised stem and root sections of Picea abies and Fagus sylvatica L. and saplings of P. abies associated with soils of different fertility levels indicated that the low disease index of certain P. abies stands is caused by the dominance of low-pathogenicity strains of A. mellea which lack the ability to form BR and penetrate non-injured bark of roots. Moreover, the poor podzolic highland soil types may contribute more to increase the survival rate and resistance of transplanted P. abies saplings than to reduce the spread of A. mellea. [ABSTRACT FROM AUTHOR]

The spatial and temporal dynamics of the forest cover can be captured using remote sensing data. Forest masks are a valuable tool to monitor forest characteristics, such as biomass, deforestation, health condition and disturbances. This study was carried out under the umbrella of the EC H2020 MySustainableForest (MSF) project. A key achievement has been the development of supervised classification methods for delineating forest cover. The forest masks presented here are binary forest/non-forest classification maps obtained using Sentinel-2 data for 16 study areas across Europe with different forest types. Performance metrics can be selected to measure accuracy of forest mask. However, large-scale reference datasets are scarce and typically cannot be considered as ground truth. In this study, we implemented a stratified random sampling system and the generation of a reference dataset based on visual interpretation of satellite images. This dataset was used for validation of the forest masks, MSF and two other similar products: HRL by Copernicus and FNF by the DLR. MSF forest masks showed a good performance (OAMSF = 96.3%; DCMSF = 96.5), with high overall accuracy (88.7–99.5%) across all the areas, and omission and commission errors were low and balanced (OEMSF = 2.4%; CEMSF = 4.5%; relBMSF = 2%), while the other products showed on average lower accuracies (OAHRL = 89.2%; OAFNF = 76%). However, for all three products, the Mediterranean areas were challenging to model, where the complexity of forest structure led to relatively high omission errors (OEMSF = 9.5%; OEHRL = 59.5%; OEFNF = 71.4%). Comparing these results with the vision from external local stakeholders highlighted the need of establishing clear large-scale validation datasets and protocols for remote sensing-based forest products. Future research will be done to test the MSF mask in forest types not present in Europe and compare new outputs to available reference datasets. [ABSTRACT FROM AUTHOR]

Integrated forest management (IFM) can help reconcile critical trade-offs between goals in forest management, such as nature conservation and biomass production. The challenge of IFM is dealing with these trade-offs at the level of practical forest management, such as striving for compromises between biomass extraction and habitat retention. This paper reviews some of the driving factors that influence the integration of nature conservation into forest management. The review was conducted in three steps – a literature review, an expert workshop and an expert-based cooperative analysis. Of 38 driving factors identified, three were prioritised by more of the participants than any of the others: two are socio-cultural factors, identity (how people identify with forest) as well as outreach and education, and one is economic – competitiveness in forest value chains. These driving factors correspond to what are considered in the literature as enablers for IFM. The results reveal that targeted, group-oriented, adaptive and innovative policy designs are needed to integrate nature conservation into forest management. Further, the results reveal that a " one-size-fits-all " governance approach would be ineffective, implying that policy instruments need to consider contextually specific driving factors. Understanding the main driving factors and their overall directions can help to better manage trade-offs between biodiversity conservation and biomass production in European forests. Image 1 • Integrated Forest Management may help to reconcile critical trade-offs in forest management. • 38 driving factors identified as important for integrating nature conservation into forest management. • Several driving factors correspond to what can be considered as enablers of Integrated Forest Management. • Policy need to account for contextually specific driving factors to integrate nature conservation. • Integrative approaches demonstrate the need for targeted, group-oriented and adaptive policy designs. [ABSTRACT FROM AUTHOR]

In the last few years, the occurrence and abundance of tree-related microhabitats and habitat trees have gained great attention across Europe as indicators of forest biodiversity. Nevertheless, observing microhabitats in the field requires time and well-trained staff. For this reason, new efficient semiautomatic systems for their identification and mapping on a large scale are necessary. This study aims at predicting microhabitats in a mixed and multi-layered Mediterranean forest using Airborne Laser Scanning data through the implementation of a Machine Learning algorithm. The study focuses on the identification of LiDAR metrics useful for detecting microhabitats according to the recent hierarchical classification system for Tree-related Microhabitats, from single microhabitats to the habitat trees. The results demonstrate that Airborne Laser Scanning point clouds support the prediction of microhabitat abundance. Better prediction capabilities were obtained at a higher hierarchical level and for some of the single microhabitats, such as epiphytic bryophytes, root buttress cavities, and branch holes. Metrics concerned with tree height distribution and crown density are the most important predictors of microhabitats in a multi-layered forest. [ABSTRACT FROM AUTHOR]

In this paper we examine an example of a conservation conflict that is encountered in Ireland arising from the designation of Special Protection Areas (SPAs) for the Hen Harrier under the European Birds Directive (Directive, 2009/147/EC) and the consequent restrictions that are placed on forestry activities within these SPAs. We examine the causes of the Hen Harrier-forestry conflict; identify what stakeholders believe are the policy instruments and management strategies that may be useful in managing the conflict and finally identify plausible solutions that may be relevant to similar conflicts around multi-functional forests elsewhere in Europe and globally. Semi-structured interviews were conducted with key actors in this conflict. Drawing on Walker and Daniels' conflict triangle theory, a qualitative analysis of the transcripts of these interviews revealed that the conflict between Hen Harrier conservation and forestry in Ireland has a number of deep-rooted dimensions including those relating to the substance (e.g. restrictions on forest management activity in the SPAs), as well as procedural (e.g. lack of stakeholder engagement) and relationship dimensions (e.g. lack of trust). The polarisation of views in this conflict testifies to how entrenched stakeholders can become through lack of communication and trust. The policy instruments that stakeholders identified as having potential to address the conflict include the introduction of incentives/compensation scheme; changes to restrictions; more data and research on Hen Harrier bird surveys; implementation of landscape management models; and better communication and stakeholder engagement. The study highlights that conservation conflicts persist due to the multi-functional nature of forests and also due to repeated mistakes in terms of the lack of engagement with local stakeholders. Increasing the involvement of local actors has important substantive and instrumental benefits including improving the quality of decisions, as well as creating a greater chance of policies being better socially and politically acceptable. The need for more and better capacity-building across EU Member States for statutory and government agencies to learn from one another in terms of how to avoid repeating the same mistakes from one site to another is highlighted. • A forestry and conservation conflict is examined. • The conflict has substance, procedural and relationship dimensions. • Stakeholder hold very polarised views. • Lack of communication and trust are major issues. • Stakeholders share a common interest and goal in terms of conservation and incentives. [ABSTRACT FROM AUTHOR]

The natural range of variation of ecosystems provides reference conditions for sustainable management and biodiversity conservation. We review how the understanding of natural reference conditions of boreal forests in northern Europe has changed from earlier perceptions of even-aged dynamics driven by stand-replacing disturbances towards current understanding highlighting the role of non-stand-replacing disturbances and the resultant complex forest dynamics and structures. We show how earlier views and conceptual models of forest disturbance dynamics, including the influential ASIO model, provide estimates of reference conditions that are outside the natural range of variation. Based on a research synthesis, we present a revised forest reference model incorporating the observed complexity of ecosystem dynamics and the prevalence of old forests. Finally, we outline a management model and demonstrate its use in forest ecosystem management and show how regional conservation area needs can be estimated. We conclude that attaining favourable conservation status in northern Europe's boreal forests requires increasing emphasis on ecosystem management and conservation for old forest characteristics.