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Simple Summary: Osmylopsychopidae, an extinct family of Neuroptera, is closely related to the extant Psychopsidae. The sporadic records of this family and limited preservation of fossil specimens have hindered the understanding of the evolution of Osmylopsychopidae. In this paper, one new genus with four new species of Osmylopsychopidae are described from the Middle Jurassic Jiulongshan Formation of northeastern China. These newly discovered osmylopsychopids with a forewing length of ca. 10 mm (implying small body size), clearly distinguished from the traditional sense of Osmylopsychopidae, suggest that Osmylopsychopidae possibly underwent a significant diversification during the Mesozoic. One new genus with four new species sharing the similar character of a forewing length of ca. 10 mm, namely, Minipsychops spasulus gen. et sp. nov., Minipsychops polychotomus sp. nov., Minipsychops densus sp. nov., and Minipsychops unicus sp. nov., are described from the Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia in China. These new insects can be attributed to Osmylopsychopidae for their distinctive configuration of costal space and the venations of RP1 and Cu. Compared with the typically medium to large body sizes of known osmylopsychopids, these new Middle Jurassic taxa of a particularly miniaturized lineage not only enrich the species diversity of Osmylopsychopidae, but also enhance our understanding of the historical evolution of these poorly known lacewings. [ABSTRACT FROM AUTHOR]

Fossils of Osmylinae are rare, currently only one fossil genus of this subfamily is known, Lithosmylus which is recorded from the late Eocene Florissant Formation of USA. Herein, we report a new Osmylinae genus with two new species, Vetosmylus tentus gen. et sp. nov. and Vetosmylus maculosus gen. et sp. nov. from the Middle Jurassic Jiulongshan Formation of Daohugou Village, Inner Mongolia, China, The new genus shares many apomorphic characters with extant osmyline relatives, e.g., forked subcostal veinlets in forewing; dense cross-veins present in radial sector, usually forming one complete outer gradate series in forewing; the region between MA and MP significantly broadened in hind wing, but with only one row of cells, implying a likely morphological stasis during the evolution of Osmylinae. The new genus Vetosmylus, the first and earliest record hitherto of Osmylinae from the Mesozoic, enhances our understanding of the early stages of their evolution. [ABSTRACT FROM AUTHOR]

A well-developed recurrent veinlet is found in the forewing of two species of Nymphidae from the Middle Jurassic locality of Daohugou (Inner Mongolia, China), Liminympha makarkini Ren & Engel and Daonymphes bisulca gen. et sp. n. This is the first record of this trait in the clade comprised of the superfamilies Myrmeleontoidea and Chrysopoidea. We interpret the recurrent veinlet in these species as a remnant of the condition present more basally in the psychopsoid + ithonoid + chrysopoid + myrmeleontoid clade (i.e., as a plesiomorphy). Other venational character states of D. bisulca of interest include the configuration of subcosta anterior (ScA), which is very similar to that of extant Nymphidae. We consider the short ScA terminating on ScP to be an autapomorphy of Neuroptera. [ABSTRACT FROM AUTHOR]

Epipanfilovia oviformis gen. et sp. nov. (Neuroptera: Panfiloviidae) is described from the Middle Jurassic locality of Daohugou (Inner Mongolia, China) based on six specimens. A revised diagnosis of this family is provided. It includes two genera, Panfilovia Makarkin and Epipanfilovia gen. nov.; Osmylogramma Ponomarenko is removed from the family. Panfiloviidae is most closely related to another Jurassic family Grammolingiidae, and both are provisionally assigned to the superfamily Osmyloidea. [ABSTRACT FROM AUTHOR]

Background: Lacewings (insect order Neuroptera), known in the fossil record since the Early Permian, were most diverse in the Mesozoic. A dramatic variety of forms ranged in that time from large butterfly-like Kalligrammatidae to minute two- winged Dipteromantispidae. Principal Findings: We describe the intriguing new neuropteran family Parakseneuridae fam. nov. with three new genera and 15 new species from the Middle Jurassic of Daohugou (Inner Mongolia, China) and the Early/Middle Jurassic of Sai-Sagul (Kyrgyzstan): Parakseneura undula gen. et sp. nov., P. albomacula gen. et sp. nov., P. curvivenis gen. et sp. nov., P. nigromacula gen. et sp. nov., P. nigrolinea gen. et sp. nov., P. albadelta gen. et sp. nov., P. cavomaculata gen. et sp. nov., P. inflata gen. et sp. nov., P. metallica gen. et sp. nov., P. emarginata gen. et sp. nov., P. directa gen. et sp. nov., Pseudorapisma jurassicum gen. et sp. nov., P. angustipenne gen. et sp. nov., P. maculatum gen. et sp. nov. (Daohugou); Shuraboneura ovata gen. et sp. nov. (Sai-Sagul). The family comprises large neuropterans with most primitive wing venation in the order indicated by the presence of ScA and AA1+2, and the dichotomous branching of MP, CuA, CuP, AA3+4, AP1+2. The phylogenetic position of Parakseneuridae was investigated using a phylogenetic analysis of morphological scoring for 33 families of extinct and extant Neuropterida combined with DNA sequence data for representatives of all extant families. Parakseneuridae were recovered in a clade with Osmylopsychopidae, Prohemerobiidae, and Ithonidae. Conclusions/Significance: The presence of the presumed AA1+2 in wings of Parakseneuridae is a unique plesiomorphic condition hitherto unknown in Neuropterida, the clade comprising Neuroptera, Megaloptera, Raphidioptera. The relative uncertainty of phylogenetic position of Parakseneuridae and the majority of other families of Neuroptera reflects deficient paleontological data, especially from critical important periods for the order, earliest Triassic and latest Triassic/earliest Jurassic. [ABSTRACT FROM AUTHOR]

Kempyninae are a subfamily of osmylid lacewings whose extant representatives are restricted to the austral continents. Three new osmylids, Jurakempynus sinensis gen. et sp. nov., J. bellatulus gen. et sp. nov., and J. epunctatus gen. et sp. nov. are described from the Middle Jurassic Jiulongshan Formation of Daohugou Village, Inner Mongolia, China. The new genus Jurakempynus is assigned to the subfamily Kempyninae constituting its oldest known occurrence. It is allied to the Australian genus Kempynus, sharing two synapomorphies: extremely expanded MP space and distally forked MP2 in the hindwing. The occurrence of Jurakempynus gen. nov. in China (Northern Hemisphere) implies a much wider geographic range of the Mesozoic osmylid lacewings in comparison to their modern counterparts. [ABSTRACT FROM AUTHOR]

The article discusses a new species and genus of Aetheogrammatidae from the Jurassic of Inner Mongolia in China. It says that the new genus such as Aetheogramma from the younger Yixian deposits shows the notably large extent of the diversity in Late Jurassic-Early Cretaceous neuropteran faunas. According to the author, the genus Kalligrammina Panfilov from the Late Jurassi of Karatau in southern part of Kazakhstan may be classified best in Aetheogramma. The diagnosis, description and holotype of the Aetheogramma kalligrammoides species are also discussed.

Two new species of lacewings are described and illustrated from the Jiulongshan Formation (Middle Jurassic) of Inner Mongolia, China: Leptolingia calonervis sp. nov. and Litholingia ptesa sp. nov. (Grammolingiidae). The species can be distinguished by the following features: the outermost branch of MP2 on the hind wing did not reach the margin of wing in Leptolingia calonervis sp. nov.; the second branch of Rs forked deeply while other branches of Rs forked much later in Litholingia ptesa sp. nov. [ABSTRACT FROM AUTHOR]

Two new fossil species of Kalligrammatidae (Neuroptera) are described from the Middle Jurassic locality at Daohugou, Inner Mongolia, China: Limnogramma hani n. sp. and Limnogramma mongolicum n. sp. Some venational homologies in Neuroptera are discussed briefly. [ABSTRACT FROM AUTHOR]