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Informal science learning centers (ISLCs) like zoos, aquariums and botanic gardens are trusted community assets and perceived as critical sites for learning outside the classroom for multigenerational learners. Moreover, their role in engaging the public in critical topics, such as climate change is widely recognized. As extreme weather events grow more frequent, building climate resilient social and ecological systems will be necessary. However, how ISLCs facilitate climate resilience is understudied. This paper presents findings from two qualitative case studies of climate resilience projects, through partnerships between community organizations and ISLCs. The results demonstrated the need to prioritize relationship building so that organizations with distinct approaches authentically appreciate each other's work and foster power symmetries. Diverse definitions of 'resilience' emerged, where ecological and social resilience held unique meaning for residents most vulnerable to climate events. Ultimately, the research highlighted how ISLCs and community partners can advance equitable and inclusive climate resilience.