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A new osmylid neuropteran insect Archaeosmylidia fusca gen. et sp. nov. is described from the Middle Jurassic locality of Daohugou (Inner Mongolia, China). Its forewing venation differs from that of other hitherto known osmylids by a set of plesiomorphic features. This genus is considered here as representing a basal group of Osmylidae. The Permian-Triassic family Archeosmylidae comprises the genera Archeosmylus, Babykamenia, and Lithosmylidia. Archaeosmylidia and Archeosmylidae share the few-branched CuP, the absence of zigzag vein pattern, and the scarcity of the crossveins in the radial space. We estimate that Osmylidae might have originated in the Triassic from some "archeosmylid-like" ancestor. [ABSTRACT FROM AUTHOR]

Two new species of the genus Sinosmylites Hong are described from the Middle Jurassic locality at Daohugou (Inner Mongolia, China): S. fumosus sp. n. and S. rasnitsyni sp. n. This is the oldest known occurrence of the family Berothidae. The berothid affinity of this genus is confirmed by examination of the hind wing venation characteristic of the family. The Late Triassic family Mesoberothidae may represent an early group of Berothidae. [ABSTRACT FROM AUTHOR]