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In southern Africa, the Howiesons Poort is one of the earliest technocomplexes in which small blades and bladelets were systematically produced and retouched into formal tools. While the presence of small lithic artefacts in an assemblage is welldocumented, little is known about production pathways. This paper reports on the analysis of the lithic assemblage from Horizon 6/7 of Charles Keller's excavations of Montagu Cave, South Africa, to evaluate how raw material and decision-making affects the size of stone tools in Howiesons Poort assemblages. It addresses the following questions: how were small blades produced? Was it a deliberate choice or a constraint imposed by raw material availability? Results show branching modes of production for large and small blanks beginning with raw material acquisition. The independent production of small blanks is reinforced by the presence of cores on flakes in the assemblage. However, a relatively standardised core reduction process was used consistently, regardless of core size. The main technological and typological features observed in Horizon 6/7 are in line with other known Howiesons Poort assemblages. Considered as a whole, these observations highlight the consistency of Howiesons Poort blade technology and the deliberate production of small blanks and tools. [ABSTRACT FROM AUTHOR]

South Africa is arguably one of the most studied regions in Stone Age research. There are, however, considerable differences in research intensity with respect to different regions and time periods. While KwaZulu-Natal is an epicenter for Middle Stone Age (MSA) research, the Late Pleistocene LSA record is largely understudied in this region. Here we present a lithic assemblage from the site Umbeli Belli near Scottburgh dated to 17.8 ± 1.5ka BP. The lithic analysis of the GH 3 assemblages revealed both gradual and abrupt changes within this stratigraphic horizon, indicating relatively short-term changes in material cultural traditions. A comparison with other Robberg sites in the wider surroundings highlights the regional variability of the Robberg techno-complex and indicates potential directions for future research. [ABSTRACT FROM AUTHOR]

We explore the correspondence between changing palaeoenvironments, patterns of site use, and lithic technology at the rock shelter site Klein Kliphuis (South Africa) across the interval 65–55 000 years before present. This period coincides with the termination of Marine Isotope Stage (MIS) 4, and the disappearance of an iconic late Pleistocene archaeological unit known as the Howiesons Poort. Wood charcoals indicate sufficient soil moisture around Klein Kliphuis throughout the Howiesons Poort to support diverse tree species at a time when site occupation was relatively intense. At least some fuelwood‐gathering in this period may have been undertaken to support heat treatment of silcrete, which was the dominant lithology in tool production. A coherent set of changes occurs across the MIS 4/3 transition: occupational intensity declines, tracked by declining diversity of fuelwood species, an increase in the proportion of charcoals from shrubs and small flowering plants, lower prevalence of silcrete, and less heat treatment. While declining soil moisture is implied, there appears to be a significant change in behaviour relating to site usage, whereby foraging for dense fuelwoods was replaced by the construction of 'fast fires' that may reflect briefer visits and increased forager mobility in early MIS 3. [ABSTRACT FROM AUTHOR]

South Africa is one of the best-studied regions in terms of Stone Age research in the last few decades. Considerable progress has been made, especially for the Middle Stone Age (MSA). Recently the late Pleistocene Later Stone Age (LSA) has come back into focus. However, there are still large understudied areas such as northeastern South Africa. Here we present the first data from an archaeological site containing late Pleistocene occupations associated with the Robberg techno-complex in this region. Iron Pig rock shelter provides a well-dated sequence spanning from >16000 cal BP to <9000 cal BP. A lithic analysis of the Robberg occupations of the lowermost layers 5 and 6 provided here implies gradual temporal shifts in technology indicating short-term changing traditions. A comparative review of other LSA sites in the wider region suggests considerable research gaps and the necessity of intensified work in this area. [ABSTRACT FROM AUTHOR]

The Cradle of Humankind World Heritage site in South Africa is renowned for its karstic cave systems, which have yielded extensive fossil and stone tool assemblages dating to the Plio-Pleistocene period. The archaeological record from these caves has provided significant evidence for the evolution of lithic technology in southern Africa. Open-air sites have also been documented in this region, although they rarely receive scientific attention due to their lack of spatial and stratigraphic contexts. However, archaeological research has demonstrated that lithic production strategies were organized through landscape-scale constraints including the availability of raw materials and the energetic costs of their transport. As such, the study of open-air Stone Age sites in the Cradle of Humankind can potentially offer insight into the relationships between raw material procurement and reduction processes in this region. Here, we present a case study from the open-air locality of Elandsdrift Farm, which preserves a palimpsest of Earlier and Middle Stone Age lithic materials. While the chronological and spatial contexts of this locality present significant interpretative challenges, the Elandsdrift Farm lithic materials are not without scientific value. A landscape archaeological approach is employed to explore the influence of raw material transport on tool-making. Trends in core reduction intensity and flaking efficiency suggest that landform ruggedness and distances traveled to procure raw materials significantly influenced lithic production strategies throughout the Early to Middle Pleistocene. Despite the contextual challenges presented by open-air localities, landscape archaeological methods can inform the economic aspects of lithic scatters in this region, which cannot be easily inferred from archaeological assemblages in cave contexts. [ABSTRACT FROM AUTHOR]

Klipfonteinrand 1 (KFR1) is a foundational but poorly documented Middle Stone Age (MSA) site located in the south west of South Africa. Originally excavated by John Parkington in 1969 but undated for more than 50 years, the MSA component of the sequence formed a part of Thomas Volman's influential culture-historic technocomplex scheme. Renewed excavations identified four distinct MSA stratigraphic units at the site, the oldest two of which date to ∼85 ka and ∼156 ka. This paper presents an analysis of ∼4500 artefacts from the oldest units to test the viability of Volman's scheme and its derivatives. Coeval assemblages from other regional sites are also reviewed to test alternative models of regional technological variability driven by demographic dynamics in MIS 6 and MIS 5. We find that assemblages from neither of the deepest two stratigraphic units at KFR1 map perfectly to prevailing technocomplex schemes for southern Africa, and that they are also distinct from nearby assemblages of comparable age. Ultimately, KFR1 provides an important data point furthering the identification of variable technological adaptations during an important phase in the evolution of human behaviour. • Re-excavation and reassessment of Klipfonteinrand 1's Early Middle Stone Age units. • Validity testing of the relevancy of Thomas Volman's technocomplex scheme. • We find that the deepest two units do not map perfectly to Volman's determination. • Basal units from Klipfonteinrand 1 are distinct from coeval assemblages. • Typologically based assessments result in generalisations and mask variation. [ABSTRACT FROM AUTHOR]

Howiesons Poort (HP) sites, over the past decades, have provided exceptional access to anthropogenic remains that are enhancing our understanding of early modern human behaviour during the Middle Stone Age in southern Africa. Here, we analyse the technological and typological trends in the lithic record that form part of these behaviours, based on the HP sequence recently excavated at Klipdrift Shelter, located on the southern Cape coast of South Africa. This study contributes to enhance knowledge on the mechanisms of changes that occurred during the transition to the post-HP. Despite patterns of continuity observed, notably for core reduction methods, the seven successive lithic assemblages show significant changes in the typological characteristics and raw material selection but also in the relative importance of blade production over time. However, these changes are not necessarily synchronic and occur either as gradual processes or as abrupt technological shifts. Consequently, we cross-examine the association between the lithic phasing and other anthropogenic remains within the HP sequence at Klipdrift Shelter. We explore the implications of these patterns of changes in terms of cultural behaviours and population dynamics during the HP and we highlight the relationship between the different phases of the HP sequence at Klipdrift Shelter and those from other South African HP sites. [ABSTRACT FROM AUTHOR]

In the past few decades, a diverse array of research has emphasized the precocity of technically advanced and symbolic practices occurring during the southern African Middle Stone Age. However, uncertainties regarding the regional chrono-cultural framework constrain models and identification of the cultural and ecological mechanisms triggering the development of such early innovative behaviours. Here, we present new results and a refined chronology for the Pietersburg, a techno-complex initially defined in the late 1920’s, which has disappeared from the literature since the 1980’s. We base our revision of this techno-complex on ongoing excavations at Bushman Rock Shelter (BRS) in Limpopo Province, South Africa, where two Pietersburg phases (an upper phase called ‘21’ and a lower phase called ‘28’) are recognized. Our analysis focuses on the ‘28’ phase, characterized by a knapping strategy based on Levallois and semi-prismatic laminar reduction systems and typified by the presence of end-scrapers. Luminescence chronology provides two sets of ages for the upper and lower Pietersburg of BRS, dated respectively to 73±6ka and 75±6ka on quartz and to 91±10ka and 97±10ka on feldspar, firmly positioning this industry within MIS5. Comparisons with other published lithic assemblages show technological differences between the Pietersburg from BRS and other southern African MIS5 traditions, especially those from the Western and Eastern Cape. We argue that, at least for part of MIS5, human populations in South Africa were regionally differentiated, a process that most likely impacted the way groups were territorially and socially organized. Nonetheless, comparisons between MIS5 assemblages also indicate some typological similarities, suggesting some degree of connection between human groups, which shared similar innovations but manipulated them in different ways. We pay particular attention to the end-scrapers from BRS, which represent thus far the earliest documented wide adoption of such tool-type and provide further evidence for the innovative processes characterizing southern Africa from the MIS5 onwards. [ABSTRACT FROM AUTHOR]

This paper describes the late Pleistocene early microlithic at Putslaagte 8 (PL8) rockshelter in the eastern Cederberg of the Western Cape. Focus is placed on: 1) characterising the pre-Robberg Early Later Stone Age (ELSA) and Robberg at the site, and 2) determining if and how the ELSA anticipates the Robberg. Results suggest that the ELSA in the eastern Cederberg is associated with the production of hornfels blades with a shift in production emphasis towards smaller silcrete blades during the Robberg. Blade production systems and modes of delivery, however, are quite different in the two periods. Bipolar technology forms only a minor component of the ELSA technology and there is minimal use of quartz despite its presence in the landscape. This contrasts with the ELSA at sites on the opposite side of the Cape Fold Mountain Range and in the north of the subcontinent where bipolar technology and a preference for quartz appear to be defining features. The late Pleistocene sequence at PL8 is particularly significant for helping to characterise the pre-Robberg in the southwest of southern Africa - an area where the nature of the ELSA is currently under-defined. [ABSTRACT FROM AUTHOR]

Lithic material recovered from excavations at Holkrans rock shelter, a late Holocene, Later Stone Age site in the Vredefort Dome, South Africa, indicates forager occupation from at least 2000 years ago, and culture contact with farmer groups in the centuries around 1500 AD. A technological analysis of cryptocrystalline silicate (CCS) cores and bladelets indicates marked change in knapping strategy between the pre-ceramic (PC) and ceramic (C) phases, with bipolar knapping of opaline prevalent in the C phase compared to a predominance of freehand knapping on dolomitic chert in the PC phase. The study demonstrates the usefulness of the technological approach as a tool in the study of contact-period LSA lithic assemblages in southern Africa. [ABSTRACT FROM AUTHOR]

The assemblage of stone tools from P. Beaumont's Excavation 1 at Wonderwerk Cave, Northern Cape Province, South Africa, provides a unique stratified sequence covering a large part of the Earlier Stone Age. A combination of cosmogenic burial age and paleomagnetic age dating provides limited chronometric constraint on this sequence. The Wonderwerk sequence provides evidence for the development of Earlier Stone Age technology in southern Africa that parallels the sequence known from East Africa. This paper presents a technological discussion of biface technology at Wonderwerk Cave as well as an overview of the associated lithic assemblage. [ABSTRACT FROM AUTHOR]

While the majority of research on the Middle Stone Age (MSA) in southern Africa has been conducted in the southern and western Cape, studies of the east coast of South Africa have become increasingly important due to the existence of well-stratified sites such as Sibudu. Because of the scarcity of comparable localities, however, we still know little about the spatial and temporal variability of MSA lithic technology in this region. We therefore chose to expand our research focus to other, lesser-known sites in the eastern part of South Africa. One such site is Holley Shelter which was excavated by Gordon Cramb between 1950 and I960. Since its archaeological material was only studied in a cursory manner, we conducted a detailed technological study of the MSA lithic artefacts from Cramb's excavations, including attribute analysis and examination of reduction sequences. Our first aim was to assess the degree of potential mixing and recovery bias among the lithic material. We then characterised the different assemblages and investigated their diachronic variation throughout the occupation sequence. In order to obtain a rough age estimate of the so far undated sequence of Holley Shelter, we compared its lithic technology to other MSA sites in the eastern part of South Africa. Our results indicate three different phases of MSA occupation that vary in terms of raw material composition, core reduction, and tool manufacture. The assemblages are characterised by a blade and point technology that mostly derives from platform cores as well the highest proportions of splintered pieces reported from a southern African MSA site. The sequence does not feature Later Stone Age (LSA), Howieson's Poort, Still Bay or final MSA industries. Compared to other sites in the general region, the assemblages are most similar to lithic technology post-dating the Howieson's Poort, suggesting that the occupations fall broadly into the earlier part of MIS 3. [ABSTRACT FROM AUTHOR]

The MSA/LSA transition is a major shift in the African archaeological record, but questions on its beginning remain debated. In southern Africa, most sites suggest an origin of LSA technology after about 30.000 years BP. The single exception is Border Cave situated at the border between South Africa and Eswatini, with surprisingly old dates of ∼43.000 BP associated with an LSA-like bipolar quartz assemblage. While many researchers now consider Border Cave to represent the origin of the LSA in southern Africa, these findings lack proper contextualization with regional lithic and chronometric data. Here we pursue the question whether Border Cave provides firm evidence for the source of LSA technology that later spread to the rest of southern Africa. To test between different hypotheses, we provide new chronometric and lithic data from the site of Sibebe, situated in the highveld of Eswatini only 100 km distant to Border Cave, and contextualize these results with nearby localities. Eswatini represents an ideal study area as it features many excavated sites but remains heavily understudied, rarely appearing in comparative MSA/LSA research. Our analyses at Sibebe identify two distinct groups of MSA lithic assemblages dated to between 43.000 and 27.000 cal BP by C14, overlain by a late Holocene LSA industry. The latest MSA expression at Sibebe features finely shaped unifacial and bifacial points without trends towards LSA technology. A comparative review of sites in Eswatini, South Africa, and Mozambique confirms similar MSA industries post-dating 30.000 BP but finds no evidence for LSA technologies before ∼27.000 BP. These findings suggest that Border Cave cannot mark the origin of the much later LSA in southern Africa, but rather signifies an early set of locally specific behavioral innovations that appeared and disappeared again. These findings have important implications for the spread of new technologies and our understanding of cultural evolutionary trajectories in southern Africa and beyond. • New lithic technological and chronometric data on Sibebe shows the persistence of Middle Stone Age technologies beyond 30.000 cal BP in southern Africa. • Our analysis places the MSA/LSA transition in southern Africa after 30.000 cal BP and refute the hypothesis that Border Cave constitutes the origin of Later Stone Age technologies already at ∼43.000 cal BP. • There is a significant gap of several thousand years between Middle and Later Stone Age occupations in Eswatini. [ABSTRACT FROM AUTHOR]

Abstract: New excavations at the Middle Stone Age (MSA) open-air site of Hoedjiespunt 1 (HDP1) on the west coast of South Africa advance our understanding of the evolution of coastal adaptations in Homo sapiens. The archaeological site of HDP1 dates to the last interglacial and consists of three phases of occupation, each containing abundant lithic artifacts, shellfish, terrestrial fauna, ostrich eggshell and pieces of ground ocher. The site provides an excellent case study to analyze human behavioral adaptations linked to early exploitation of marine resources. Here we reconstruct human activities through a detailed study of the lithic assemblages, combining analyses of the reduction sequences, artifact attributes and quartz fracturing. These methods provide insights into raw material procurement, lithic reduction sequences, site use and mobility patterns, and foster comparison with other MSA coastal sites. The main characteristics of the lithic assemblages remain constant throughout the use of the site. Quartz dominates silcrete and other raw materials by almost four to one. Knappers at HDP1 produced different forms of flakes using multiple core reduction methods. Denticulates represent the most frequent tool type. The assemblages document complete, bipolar and hard hammer reduction sequences for the locally available quartz, but highly truncated reduction sequences with many isolated end products for silcrete, a material with a minimum transport distance of 10–30km. This observation suggests that well provisioned individuals executed planned movements to the shoreline to exploit shellfish. Our excavations at HDP1 furthermore demonstrate the simultaneous occurrence of flexible raw material use, anticipated long-distance transport, systematic gathering of shellfish and use of ground ocher. The HDP1 lithic assemblages document a robust pattern of land-use that we interpret as a stable adaptation of modern humans to coastal landscapes as early as MIS 5e. [Copyright &y& Elsevier]

Abstract: This paper considers the transition from Howiesons Poort to post-Howiesons Poort technologies at the rock shelter site of Klein Kliphuis, South Africa. The transition at this site is shown to be gradual, with incremental changes in material selection and in the size and shapes of flakes and cores. Implements which appear to blend characteristics of those distinctive of the earlier and later industries appear briefly at the mid-point of the transition. The results suggest that there is unlikely to have been an occupational hiatus between the Howiesons Poort and post-Howiesons Poort. Explanations for the Howiesons Poort phrased in terms of population expansion and contraction are also difficult to support. Technological changes at this time may relate to environmental variation, though the limited nature of terrestrial archives documenting changes in resource productivity necessitate that any such suggestion be made with caution. [Copyright &y& Elsevier]

George Leith's investigations at Cape St Blaize Cave in the late 1880s laid the foundations for an influential set of studies on the Middle Stone Age in and around Mossel Bay. Those studies, and particularly the descriptions of the lithics from Cape St Blaize, have figured prominently in discussions of Middle Stone Age lithic variation in South Africa. The lithics from Cape St Blaize form the basis of the Mossel Bay, which has been variously defined as a culture, industry, variant, technique, and sub-stage. We review past and present usage of the Mossel Bay lithic variant and conclude that our understanding is fraught with ambiguity and lack of precision due to the selective nature of the original collection and the lack of a complete published description and analysis of the type-site. We then report on the first complete study of the Cape St Blaize lithic sample and compare it to more recent, complete, and well-dated samples from nearby Pinnacle Point. The Cape St Blaize collection differs from those at Pinnacle Point in several important ways, despite the relative proximity. To correctly define the Mossel Bay, the original type-site needs to be reinvestigated with better stratigraphic control, unbiased samples, and radiometric dating. [ABSTRACT FROM AUTHOR]

Abstract: It has been suggested that many behavioral innovations, said to appear during the late Middle Stone Age in sub-Saharan Africa, facilitated the expansion of anatomically modern humans from Africa and the Near East into Europe at about 50 kyr; the process eventually led to the replacement of Neanderthals by modern humans and the emergence of the Upper Paleolithic. However, assemblages in this time range are little known in South Africa. In fact, the transition from Middle to the Later Stone Age in Southern Africa is controversial. The early appearance in South Africa of many innovations, such as sophisticated knapping techniques (e.g. the use of soft hammer or indirect percussion in blade production, of composite tools, of microlithic and bladelet technologies) remains to be established through technological analysis. We present here the first results of a project designed to carry out detailed technological studies of several lithic assemblages in South Africa and France dated to the transition period. At this time we have completed the study of a post-Howiesons Poort assemblage from the rock shelter site of Sibudu. The >2 m deep stratigraphic sequence of Sibudu extends from Howiesons Poort at its base to final Middle Stone Age, directly under Iron Age layers. We have analyzed in detail layer RSP (ca. 53 kyr, 1 m above the Howiesons Poort levels) which has provided a large assemblage of several thousand stone artifacts. Compared to published MSA assemblages this industry is unusual for the very high proportions of retouched pieces (15%). The technology is not very elaborate and there is no strong standardization of the end-products. There are no flakes of predetermined shapes; retouch is used to modify irregular flakes to obtain desired edges. Knapping of flakes and blades is done by hard hammer; soft hammer is used only for retouching tools. Interestingly the older Howiesons Poort blades were produced on the same raw materials by soft hammer. Raw material (hornfels and dolerite) was procured from distances of less than 20 km. Unifacial points are the dominant type and there is strong evidence of hafting and use as spear armatures. Detailed comparisons with Middle Paleolithic assemblages of Western Europe show that the late Middle Stone Age technology in South Africa is very similar to that of the Middle Paleolithic; in fact we see no fundamental differences between the two entities, as far as lithic technology is concerned. Implications for the Out of Africa hypothesis are discussed. [Copyright &y& Elsevier]

Recent excavations at Rose Cottage Cave, located in the Free State, South Africa, have revealed both a transitional assemblage, dated to ca. 20,000 bp, and a final Middle Stone Age (MSA) assemblage, dated to ca. 28,000 bp. Preliminary analysis of these assemblages was undertaken to determine if the current European model of a “cultural revolution” for the emergence of the Upper Palaeolithic can be applied to the southern African evidence. Examination of these assemblages revealed that differences occurred between the transition in Europe and that in southern Africa in both the chronology of the transition and the degree to which this complete cultural package is linked to the emergence of Late Stone Age (LSA) technology. The methods of lithic production, the chronology of the MSA/LSA transition, and the associated behavioral characteristics were examined and results indicated that the technological change which occurred during the MSA/LSA transition was not a dramatic innovation in technology but, rather, a shift in the emphasis of production from a level of technology already in place and demonstrate a level of continuity between the MSA and the LSA. As a gradual occurrence, the MSA/LSA transition does not seem to fit the time frame for the European Upper Palaeolithic; it both occurs at a later period and takes longer to transpire. In addition, the origins of symbolic use of lithics appear to lie within the MSA, indicating that a more complex set of behavioral adaptation was occurring in the late Pleistocene in southern Africa, and that the MSA/LSA transition in this region does not adequately conform to the model of a revolutionary shift in behavior and technology that is proposed for the Middle/Upper Palaeolithic transition in Europe [ABSTRACT FROM AUTHOR]

The Palaeo-Agulhas Plain (PAP), when exposed, presented Middle Stone Age (MSA) foragers at Pinnacle Point (PP) on the South Coast of South Africa with new sources of raw materials to make stone tools. Sea-level fluctuations and the changing size of the Paleo-Agulhas Plain throughout the Pleistocene PP record ∼165 ka to 50 ka would have altered the availability of different resources, thus potentially forcing new raw material procurement strategies. The relative frequencies of raw material throughout the PP sequence shows that frequencies of raw material types did change, especially after 90 ka. What caused these changing frequencies is debated and centers on whether targeted procurement of specific raw materials was the cause, or if simple raw material availability and abundance due to the changing environmental context in conjunction with opportunistic procurement drove such shifts. The application of a neutral model of stone raw material procurement presented here evaluates whether random walk in the region surrounding the PP site during different coastline configurations (Marine Isotope Stage 6, 5, and 4) explains the observed shifts in raw material usage. Put differently, did opportunistic acquisition of raw materials during random walk in these different environments cause the observed raw material pattern? Model simulations and a sensitivity analysis provide no convincing evidence that observed raw material frequencies at PP resulted from opportunistic acquisition during random walk. • Incorporates marine geophysics survey data with agent-based modeling simulations of foraging behavior. • Applies a neutral model of stone tool raw material procurement to a landscape with three different modeling states. • Study focuses on the stone tool raw material record of the Late Pleistocene locality of Pinnacle Point, South Africa. • Finds little support for random walk explaining the raw material pattern during three different Marine Isotope Stages. [ABSTRACT FROM AUTHOR]

The oldest recognized artifacts at the Swartkrans cave hominid-bearing site in South Africa have long been known to occur in the Lower Bank of Member 1, now dated with the cosmogenic nuclide burial method to ca. 1.8-2.19 Ma. However, the affinities of this industry have been debated due to small sample size. In this paper we present newly excavated material from the Lower Bank retrieved since 2005 in the Swartkrans Paleoanthropological Research Project. The sample is now large enough to confirm its affinity with the Oldowan industrial complex. The assemblage is highly expedient and core reduction strategies are largely casual. Although freehand flaking is present, the bipolar technique is most significant, even in non-quartz raw materials. The Swartkrans assemblage shows some significant contrasts with the Sterkfontein Oldowan, ca. 2.18 Ma, which can be explained by its closer proximity to raw material sources, its somewhat different geographic context, and its more expedient nature. The Swartkrans Oldowan now provides us with the first good indication of Oldowan variability in southern Africa, where only two sizeable assemblages have thus far been discovered. Comparisons are made with other sites across Africa that help to place this variability within our overall understanding of the Oldowan industrial complex., (Copyright © 2018 Elsevier Ltd. All rights reserved.)

The later Pleistocene archaeological record of southernmost Africa encompasses several Middle Stone Age industries and the transition to the Later Stone Age. Through this period various signs of complex human behaviour appear episodically, including elaborate lithic technologies, osseous technologies, ornaments, motifs and abstract designs. Here we explore the regional archaeological record using different components of lithic technological systems to track the transmission of cultural information and the extent of population interaction within and between different climatic regions. The data suggest a complex set of coalescent and fragmented relationships between populations in different climate regions through the late Pleistocene, with maximum interaction (coalescence) during MIS 4 and MIS 2, and fragmentation during MIS 5 and MIS 3. Coalescent phases correlate with increases in the frequency of ornaments and other forms of symbolic expression, leading us to suggest that population interaction was a significant driver in their appearance., (Copyright © 2014 Elsevier Ltd. All rights reserved.)