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Around 42,000 years ago, anatomically modern humans appeared in Western Europe to the detriment of indigenous Neanderthal groups. It is during this period that new techno-cultural complexes appear, such as the Châtelperronian that extends from northern Spain to the Paris Basin. The Grotte du Renne (Arcy-sur-Cure) is a key site for discussing the biological identity of its makers. This deposit has yielded several Neanderthal human remains in its Châtelperronian levels. However, the last inventory of the paleoanthropological collection attributed to this techno-complex allowed the identification of an ilium belonging to a neonate (AR-63) whose morphology required a thorough analysis to assess its taxonomic attribution. Using geometric morphometrics, we quantified its morphology and compared it to that of 2 Neanderthals and 32 recent individuals deceased during the perinatal period to explore their morphological variation. Our results indicate a morphological distinction between the ilia of Neanderthals and anatomically modern neonates. Although AR-63 is slightly outside recent variability, it clearly differs from the Neanderthals. We propose that this is due to its belonging to an early modern human lineage whose morphology differs slightly from present-day humans. We also explore different hypotheses about the presence of this anatomically modern neonate ilium among Neanderthal remains. [ABSTRACT FROM AUTHOR]

Pierolapithecus catalaunicus (~12 million years ago, northeastern Spain) is key to understanding the mosaic nature of hominid (great ape and human) evolution. Notably, its skeleton indicates that an orthograde (upright) body plan preceded suspensory adaptations in hominid evolution. However, there is ongoing debate about this species, partly because the sole known cranium, preserving a nearly complete face, suffers from taphonomic damage. We 1) carried out a micro computerized tomography (CT) based virtual reconstruction of the Pierolapithecus cranium, 2) assessed its morphological affinities using a series of two-dimensional (2D) and three-dimensional (3D) morphometric analyses, and 3) modeled the evolution of key aspects of ape face form. The reconstruction clarifies many aspects of the facial morphology of Pierolapithecus. Our results indicate that it is most similar to great apes (fossil and extant) in overall face shape and size and is morphologically distinct from other Middle Miocene apes. Crown great apes can be distinguished from other taxa in several facial metrics (e.g., low midfacial prognathism, relatively tall faces) and only some of these features are found in Pierolapithecus, which is most consistent with a stem (basal) hominid position. The inferred morphology at all ancestral nodes within the hominoid (ape and human) tree is closer to great apes than to hylobatids (gibbons and siamangs), which are convergent with other smaller anthropoids. Our analyses support a hominid ancestor that was distinct from all extant and fossil hominids in overall facial shape and shared many features with Pierolapithecus. This reconstructed ancestral morphotype represents a testable hypothesis that can be reevaluated as new fossils are discovered. [ABSTRACT FROM AUTHOR]

Objectives: Both morphometric and proteomic studies have revealed the close relationship of Homo antecessor with Neanderthals and H. sapiens. Considering this relationship, we aim to characterize the Early Pleistocene Atapuerca‐Gran Dolina (TD6) maxillary premolars to test if their pattern of enamel thickness is shared with Neanderthals or H. sapiens. Materials and Methods: We employed microcomputed tomography to estimate 2D and 3D tissue proportions in seven H. antecessor maxillary premolars, belonging to two individuals: H1 and H3, and compared them to a sample of extinct and extant Homo populations of African, Asian and European origin (n = 52). Results: Our results reveal a different pattern of enamel thickness between the Atapuerca‐Gran Dolina two individuals. While TD6‐H1 possesses thin‐enameled crowns, with a clear affinity with Neanderthals, TD6‐H3 exhibits the thick pattern, a trait shared with the majority of fossil hominins and H. sapiens. Discussion: This work provides new data on upper premolar enamel thickness in H. antecessor. By documenting both a thin and a thick pattern of enamel thickness in the TD6 sample, we warn about the taxonomic utility of this feature in the characterization of isolated remains. We suggest that the thin enamel condition would have emerged during the Early Pleistocene and it became the most frequent and typical condition in Neanderthals. Possible causes for the pattern observed in TD6 include sexual dimorphism or presence of two populations in the sample; however, population variability is the most plausible explanation with a character expression intermediate between those of Neanderthals and other members of the genus Homo. This interpretation is compatible with the phylogenetic position of H. antecessor close to the ancestor of Neanderthals and H. sapiens. [ABSTRACT FROM AUTHOR]

El Cerro de los Batallones is one of the most important fossil sites of the Miocene. The quantity and preservation of its fossil remains have allowed us to describe many new mammal species. One of these is Hispanomys moralesi, a derived species of the Tribe Cricetodontini, a wide group of rodents with great importance in the faunal assemblage during the Miocene. Unlike most fossil micromammals in this fossil site, we have obtained skull remains that preserve never recorded anatomical structures in this time interval. Due to the fragility of the material, it could only be studied in detail using X-ray micro-computed tomography (µ-CT Scan). With this technique, we obtained three-dimensional models of 12 skulls of Hispanomys moralesi, in which different osteological parts have been described. In this way, we compared the morphological variation of the skull with other fossil cricetids and concluded that Batallones' species is an opisthodont rodent, with a relatively elongated rostrum, a robust base of zygomatic arches and big-sized tooth rows. Opisthodont incisor characterises omnivore and herbivore diets in rodents. Therefore, these new findings of Hispanomys moralesi could allow us to describe the dietary affinities of this extinct species. [ABSTRACT FROM AUTHOR]

Speleothems are a recognized source of palaeoclimatic information, but their value as a source of signals from human activities in caves with an archaeological record has rarely been explored. Previous studies of speleothems in the Sierra de Atapuerca karst system (Burgos, northern Spain) revealed an important human fossil record, provided information about human activities in and around these caves, and the impacts on their natural environment. The present study reports the results of molecular characterization of darkcoloured laminae from the stalagmites Ilargi (Galería de las Estatuas) and GS1, GS2, and GS3 (Galería del Silo), by pyrolysis-GC-MS (Py-GC-MS) and thermally assisted hydrolysis and methylation (THM-GC-MS). The features of the organic matter demonstrate the presence of 1) a dominant aliphatic fraction probably from in situ bacterial and ex situ plant-derived lipids, 2) black carbon (from soot and/or charcoal), 3) polysaccharides and N-rich moieties (probably from bat guano and microbial sources), and 4) a signal of terpenoid derivatives that may originate from the overlying limestone (kerogen) and extant gymnosperm resin (soils) or cyanobacteria (cave). Some plant-derived lignin may be present as well but was not identified unambiguously. It is concluded that this approach allows identifying multiple general sources of organic matter which can help understand speleothem formation processes, and evidence of soot deposition could be clearly linked to human activities. [ABSTRACT FROM AUTHOR]

The Middle Pleistocene Sima de los Huesos (SH) site has yielded more than 7.500 human fossil remains belonging to a minimum of 29 individuals. Most of these individuals preserve either the complete mandibular molar series or at least the first (M1) and second (M2) molars. The inhibitory cascade mathematical model was proposed by Kavanagh et al. (Nature, 449, 427–433 [2007]) after their experimental studies on the dental development of murine rodent species. The activator–inhibitor mechanism of this model has shown its ability for predicting evolutionary size patterns of mammalian teeth, including hominins. The main aim of this study is to test whether the size molar patterns observed in the SH hominins fit the inhibitory cascade model. With this purpose, we have measured the crown area of all SH molars in photographs, using a planimeter and following techniques used and well contrasted in previous works. Following one of the premises of the inhibitory cascade model, we expect that the central tooth (M2 in our case) of a triplet would have the average size of the two outer teeth. The absolute difference between the observed and the expected values for the M2s ranges from 0.23 to 8.46 mm2 in the SH sample. In terms of percentage, the difference ranges between 0.25% and 10.34%, although in most cases, it is below 5%. The plot of the estimated M3/M1 and M2/M1 size ratios obtained in the SH hominins occupies a small area of the theoretical developmental morphospace obtained for rodent species. In addition, the majority of the values are placed near the theoretical line which defines the relationship predicted by the inhibitory cascade model in these mammals. The values of the slope and intercept of the reduced major regression obtained for the SH individuals do not differ significantly from those obtained for rodent species, thus confirming that the size of the molars of the SH hominins fits the inhibitory cascade model. We discuss these results in terms of dental development. Despite the promising results in the SH sample, we draw the attention to the fact that most Early Pleistocene Homo specimens exhibit a pattern (M1 < M2 > M3), which is outside the expected theoretical morphospace predicted by the inhibitory cascade model. The shift from the M1 < M2 < M3 size relationship observed in early hominins (including H. habilis) to the M1 > M2 > M3 size relationship, which is predominant in modern humans, includes sequences that depart from predictions of the inhibitory cascade model. Additional studies are required to understand these deviations. [ABSTRACT FROM AUTHOR]

The variability observed in the growing Middle Pleistocene hominin fossil record of Europe continues to trigger much debate on taxonomic issues and the biological processes that gave rise to Neanderthals. Here we present a metric and morphological comparative study of the dental samples recovered from the sites of Arago (southeast France) and Sima de los Huesos (SH) in the Sierra de Atapuerca (northern Spain). These sites are key to providing answers to these questions since they have yielded the largest hominin samples so far recovered for this time period. Despite the geographical proximity of the two sites and the contemporaneity of their hominin assemblages, we have observed remarkable metric and morphological differences between the teeth at Arago and SH. Whereas the SH teeth present an almost morphological identity with European Neanderthals, the Arago teeth exhibit a combination of plesiomorphic as well as some Neanderthal-derived features. In addition, the Arago crown dimensions are remarkably larger than those from SH, the differences being statistically significant for most variables. We hypothesize that during the Middle Pleistocene the European continent was settled at different points in time by hominin groups coming from Southwest Asia, probably from a common mother population evolving in this latter region. These first settlers can be identified by their more plesiomorphic morphology, whereas the most recent settlers are closer in appearance to Neanderthals. In addition, genetic processes such as isolation, genetic drift, directional adaptation or hybridization would have given rise to the puzzle we observe in the current fossil record. • This report compares the two largest dental samples form the European Middle Pleistocene. • The results highlight the differences and similarities between these samples. • We conclude that during the Middle Pleistocene at least two genealogies live in Europe during this geological period. [ABSTRACT FROM AUTHOR]

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In the eastern Iberian Peninsula, the archaeological site of Cuesta de la Bajada (Teruel, Spain) has produced some of the earliest evidence of Middle Paleolithic stone tool traditions together with evidence of equid and cervid carcasses defleshed by hominins. Based on the numerical age of 317-240 ka derived from OSL, ESR and AAR dating methods for the lower part of the Cuesta de la Bajada sedimentological sequence (level CB3), as well as the biochronological inferences for the small and large mammal associations, the site can be attributed to Marine Isotope Stage (MIS) 8 or the end of MIS 9. As amphibians and reptiles have precise environmental and climatic requirements and do not differ at species level from the extant herpetofauna of the Iberian Peninsula, they can contribute to the reconstruction of the landscape and climate. In this paper, the fossil amphibians and squamate reptiles from Cuesta de la Bajada are studied for the first time. The mutual ecogeographic range and habitat weighting methods were applied to the herpetofaunal assemblages to estimate quantitative data for the landscape and climate reconstructions. The climate is shown to have been colder and wetter than today in the interior of eastern Spain, with mean annual temperature up to 2.5 °C lower and mean annual precipitation slightly higher than at present. The monthly climatic reconstruction shows differences in the distribution of precipitation over the course of the year, with more abundant precipitation from late autumn to spring (i.e. from October to May), and more or less similar precipitation to today during the summer months (July and August). This suggests there was stronger rainfall seasonality between winter and summer than currently occurs. The paleoenvironmental reconstruction based on the herpetofaunal assemblage depicts a poorly forested (15–20%) patchy landscape with a large representation of dry meadows and scrubland habitats together with well-evidenced aquatic habitats. These reconstructions are consistent with other proxies recovered at Cuesta de la Bajada (pollen, small and large mammals) as well as other European MIS 8–9 paleoclimatic records. We can thus correlate levels CB2 and CB3 with the later part of MIS 8 (265-257 ka) or with a humid fluctuation within the MIS9b (303-290 ka). It is also possible to provide a new description of the environmental and climatic conditions that occurred in inner Spain during a cold period of the late Middle Pleistocene. [ABSTRACT FROM AUTHOR]

The oldest evidence of a human presence in western Europe is currently documented at the late Early Pleistocene sites of Barranco León and Fuente Nueva 3 (Guadix-Baza Basin, SE Spain), dated respectively to 1.4 and 1.2 Ma. Understanding the climatic variability that these hominins encountered and coped with is of outstanding importance for placing their activities in an ecological context and understanding their capacity to adapt to changing conditions. Glacial-interglacial variability during this period was considerably less intense than during later phases of the Pleistocene. To date, however, no quantitative estimates are available for this climate variability in Early Pleistocene hominin sites, and no 'cold' reconstructions directly associated with the early hominin occupations of Europe have been performed either. Here stratigraphically constrained quantitative climatic reconstructions are provided for the sites of Barranco León and Fuente Nueva 3, using an improvement on the Mutual Ecogeographic Range method by projecting the niche envelope of the extant representatives of Ophisaurus sp. We ascertain differences in temperature and rainfall between the different layers of the two sites, in line with previous reconstructions that revealed warm, humid periods ('interglacial') as well as more temperate but drier periods ('glacial''), consistent with Early Pleistocene climate cyclicity. Our new estimates suggest that late Early Pleistocene hominins, though conditioned to some degree by climatic factors, were able to deal with changing climatic and environmental conditions ('interglacial' and 'glacial'') in the southwestern extremity of the European continent. [Display omitted] • The oldest human evidence in western Europe is currently documented at 1.4 Ma. • Understanding the climatic variability that these hominins encountered is relevant. • Paleoclimatic data are provided for an archaeopaleontological sequence. • The Early Pleistocene hominins were able to cope with changing climatic conditions. [ABSTRACT FROM AUTHOR]

It is eighteen years since the human fossils recovered from the TD6 level of the Gran Dolina cave site, in Sierra de Atapuerca (Burgos, northern Spain) were assigned to a new hominin species, Homo antecessor . This review summarizes the main results obtained from the study of these fossils during this period. The increase of the African and Eurasian fossil record, as well as the application of new methodological approaches, has led to competing interpretations about its hypothetical phylogenetic position and possible evolutionary scenarios. At present, we can argue that this species is defined by a unique mosaic of primitive traits for the Homo clade, a certain number of derived features present in modern humans, a significant suite of derived features shared with Neandertals and their ancestors in the European Middle Pleistocene (in particular with the Atapuerca-Sima de los Huesos hominins), and some derived features shared with the Chinese Middle Pleistocene hominins. From this evidence, we suggest that a speciation event could have occurred in Africa/Western Eurasia, originating a new Homo clade. Homo antecessor , most probably dated to the MIS 21, could be a side branch of this clade placed at the westernmost region of the Eurasian continent. [ABSTRACT FROM AUTHOR]

The sites of Barranco León D (BL-D) and Fuente Nueva 3 (FN-3) in the Guadix-Baza Basin (Granada, Spain), together with the site of Sima del Elefante in the Sierra de Atapuerca (Burgos, Spain), constitute one of the oldest records of the earliest hominid population in the European continent west of Dmanisi (Georgia, Lesser Caucasus). In the Guadix-Baza Basin, evidence of human occupation has been found to date in the form of lithic industry (Mode 1) and cut marks in large-mammal fossil remains (mainly of hippopotamuses and elephants), and recently a human tooth considered as the oldest in Europe has been discovered. Although in the case of Sima del Elefante there is unanimity among the scientific community regarding the chronology of the unit in which the hominid remains were found (Unit TE9c, 1.22 Ma), there is continuing debate on the chronology of the sites of the Guadix-Baza Basin (FN-3 and BL-D). This applies especially to BL-D, as the numerical datings published for this site have a very high error range (1.4 ± 0.38 Ma). In this paper, the chronology of these two sites is determined using as a marker the morphological and morphometric changes undergone byMimomys saviniin its first lower molar (m1) over the course of its evolutionary history. It has been possible to confirm that the oldest human presence in the Guadix-Baza Basin and at Sima del Elefante (Atapuerca) share a similar chronology, dated to between 1.1 and 1.4 Ma. Apparently, the oldest site with human remains in Europe is seen to be BL-D, dated to 1.26 ± 0.13 Ma, followed by Level TE9c, dated to 1.22 ± 0.16 Ma, and FN-3, dated to 1.20 ± 0.12 Ma. [ABSTRACT FROM AUTHOR]

Numerous studies have shown how closely nation-building accompanies research into human prehistory. While these studies primarily focus on the period before 1945, the example of the Spanish site Atapuerca demonstrates that the strong link between hominid fossils and national identity still exists in the twenty-first century. The article argues that there are different ways of appropriating prehistoric human remains. One may distinguish the concept of 'biological continuity' in which the fossils represent some kind of ancestor from the concept of 'scientific nationalism'. The latter consists in the pride in the scholarly achievement and international recognition of 'our own' scientists. The Atapuerca project was crucial in overcoming the Spanish 'inferiority complex' with respect to scientific prowess. In Atapuerca we may even discern a third concept: the marketing of origins in order to lure tourists to the site - nationalism enterprised-up. The Spanish researchers created a multifold 'popularization industry' and forged a close alliance with the national media. Thus, this article will try to explain how Atapuerca turned within less than ten years from a little known archaeological site to the - however imaginary - starting point of Spanish history at the turn of the millennium. [ABSTRACT FROM AUTHOR]

The article discusses research into 17 cranial specimens of Middle Pleistocene Neandertal hominins from the Sima de los Huesos site in Atapuerca, Spain, as of 2014. Topics include the dating of the fossils to around 430,000 years ago, examination of cranial, facial, and dental features, and speculation into Neandertal evolution.

This article reinforces the hypothesis that humans were well established in Europe by the time of the Jaramillo subchron (1.07–0.98 Ma). The lithic industry found in the Untermassfeld site demonstrates human occupation at 50° N in Europe. This evidence extends human occupation beyond the Mediterranean, as do other sites such as Kozarnika Cave (1.6–1.4 Ma), Pont de Lavaud (1.1 Ma), and Happisburgh 3 and Pakefield at the Matuyama–Bruhnes limit. Demonstrating the presence of humans at Untermassfeld has important implications regarding the human settlement of northern latitudes. Its stone tool assemblage shares the same open-air alluvial situation, chronological proximity, Epivillafranchian faunal association and Mode 1 technology as that of Vallparadís, which allows us to hypothesise that human groups with the same technology and acting as predators spread throughout the continent from the Jaramillo subchron onwards. These groups moved from already inhabited areas in the Mediterranean such as the Iberian Peninsula (the Trinchera Elefante-TE9 and Orce sites), and/or from the Near East along coastal routes around the Black Sea, along the Danube and/or along the Rhône-Saône-Rhine route (“Belfort Gap”) to the north-western coastline of Europe. The finds from Untermassfeld as well as the probably slightly more recent archaeological sites of Happisburgh 3 (England) and Dorn-Dürkheim 3 (Germany) suggest that landscapes in latitudes around 50° N were already part of the initial colonisation period of Europe in the late Early Pleistocene using Mode 1 technologies. The latter sites bridge the time period between Untermassfeld and the early Middle Pleistocene of mid-latitude Europe, as is indicated by Pakefield in Britain, and may indicate that north-western and central parts were at least intermittently recolonised from southern Europe after periods of full glaciation. This model is supported by the absence of early Mode 2 lithic industry in Europe in the second half of the Early Pleistocene which would have accompanied recolonising populations if the primary population source of Europe came from south-west Asia. Thus, the fact that Mode 1 lithic industry remains present during this period argues against the depopulation of Europe at this time. [ABSTRACT FROM AUTHOR]

This article discusses the press coverage and controversies surrounding the discovery of a fossilized hominid named the Hombre de Orce, in Spain, over the period from 1983-2007. The author comments on the developments and advancements in the field of paleoanthropology during this period and describes the role of the Institut de Paleontología de Sabadell (Institute of Paleontology of Sabadell) in Barcelona, Spain, in the discovery of and research into the Hombre de Orce. Particular focus is given to the coverage of the event in the newspaper "El País."

Abstract: In this paper we present an experimental study of bone modifications caused by human chewing during consumption. Traditionally, tooth marks have been attributed to non-human actors. However, ethnoarchaeological as well as previous experimental work has shown that humans can make many modifications during chewing. Our results suggest that the range of damage is as extensive as that most likely to be produced by carnivores. This damage includes furrowing, scooping-out, crenulated and saw-toothed edges, longitudinal cracking, crushing, peeling and tooth marks. In this paper we present a description of the types of damage observed in the experimental sample. Some of this damage shows parallels between the experimental modifications and archaeological assemblages from Pleistocene and Holocene deposits at the Sierra de Atapuerca sites (Burgos, Spain). The repetition of morphologies allowed us to attribute some of the damage to tooth marks made by human chewing. [Copyright &y& Elsevier]

This paper presents a synthesis of the combined Uranium-Series/ESR age results obtained on fossil teeth from several Early Pleistocene archaeological and/or palaeontological Spanish localities: Atapuerca Gran Dolina TD-6, Fuente Nueva-3, Barranco León, Venta Micena and Vallparadís. While the age results show some scattering, they are nevertheless encouraging and provide a significant contribution to the understanding of the first hominin settlements in Western Europe. These “old” samples are mainly characterised by high 230Th/234U activity ratios that may sometimes exceed secular equilibrium, and by high Uranium concentrations (several tens of ppm or more). The modelling of U-uptake is therefore really crucial for the age calculation process. In the case of the combined US–ESR approach, this uptake is derived from the U-series data measured in dental tissues, which seem to be strongly dependant on the hydro-geological context. Consequently, when dating “old” samples, a special attention has to be paid to the accuracy and the meaning of the U-series data collected from the dental tissues, which may be influenced by several factors such as spatial heterogeneity and recent U-uptake overprint. Some rapid scanning procedures should be developed in order to pre-identify the samples that are the most suitable for ESR dating. [ABSTRACT FROM AUTHOR]

Abstract: Systematic excavations carried out since 1989 at Bolomor Cave have led to the recovery of four Pleistocene human fossil remains, consisting of a fibular fragment, two isolated teeth, and a nearly complete adult parietal bone. All of these specimens date to the late Middle and early Late Pleistocene (MIS 7-5e). The fibular fragment shows thick cortical bone, an archaic feature found in non-modern (i.e. non-Homo sapiens) members of the genus Homo. Among the dental remains, the lack of a midtrigonid crest in the M1 represents a departure from the morphology reported for the majority of Neandertal specimens, while the large dimensions and pronounced shoveling of the marginal ridges in the C1 are similar to other European Middle and late Pleistocene fossils. The parietal bone is very thick, with dimensions that generally fall above Neandertal fossils and resemble more closely the Middle Pleistocene Atapuerca (SH) adult specimens. Based on the presence of archaic features, all the fossils from Bolomor are attributed to the Neandertal evolutionary lineage. [Copyright &y& Elsevier]

In this report, we present a morphometric comparative study of two Early Pleistocene humeri recovered from the TD6 level of the Gran Dolina cave site in Sierra de Atapuerca, northern Spain. ATD6-121 belongs to a child between 4 and 6 years old, whereas ATD6-148 corresponds to an adult. ATD6-148 exhibits the typical pattern of the genus Homo, but it also shows a large olecranon fossa and very thin medial and lateral pillars (also present in ATD6-121), sharing these features with European Middle Pleistocene hominins, Neandertals, and the Bodo Middle Pleistocene humerus. The morphology of the distal epiphysis, together with a few dental traits, suggests a phylogenetic relationship between the TD6 hominins and the Neandertal lineage. Given the older geochronological age of these hominins (ca. 900 ka), which is far from the age estimated by palaeogenetic studies for the population divergence of modern humans and Neandertals (ca. 400 ka), we suggest that this suite of derived 'Neandertal' features appeared early in the evolution of the genus Homo. Thus, these features are not 'Neandertal' apomorphies but traits which appeared in an ancestral and polymorphic population during the Early Pleistocene. Am J Phys Anthropol 2012. © 2012 Wiley Periodicals, Inc. [ABSTRACT FROM AUTHOR]

Considerations of Neandertal geographical variation have been hampered by the dearth of remains from Mediterranean Europe and the absence there of sufficiently complete associated postcrania. The 2006 and 2007 excavation of an articulated partial skeleton of a small adult female Neandertal at the Sima de las Palomas, Murcia, southeastern Spain (Sima de las Palomas 96) provides substantial and secure information on body proportions among southern European Neandertals, as well as further documenting the nature of Neandertal biology in southern Iberia. The remains exhibit a suite of cranial, mandibular, dental, and postcranial features, of both Neandertals and archaic Homo generally, that distinguish them from contemporary and subsequent early modern humans. Its lower limbs exhibit the robustness of later Pleistocene Homo generally, and its upper limbs conform to the pattern of elevated robustness of the Neandertals. Its body proportions, including relative clavicular length, distal limb segment lengths, and body mass to stature indicators, conform to the "cold-adapted" pattern of more northern Neandertals. Palomas 96 therefore documents the presence of a suite of "Neandertal" characteristics in southern Iberia and, along with its small body size, the more "Arctic" body proportions of other European Neandertals despite the warmer climate of southern Iberia during marine isotope stage 3. [ABSTRACT FROM AUTHOR]

The Sima de las Palomas, southeastern Spain, has yielded a series of Neandertal postcranial remains, including immature and mature isolated elements and the fragmentary partial skeleton of a young adult (Palomas 92). The remains largely conform to the general late archaic/Neandertal morphological pattern in terms of humeral diaphyseal shape, pectoralis major tuberosity size and pillar thickness, ulnar coronoid process height, manual middle phalangeal epiphyseal breadth, manual distal phalangeal tuberosity shape and breadth, femoral diaphyseal shape, and probably body proportions. Palomas 92 contrasts with the Neandertals in having variably gracile hand remains, a more sellar trapezial metacarpal 1 facet, more anteroposteriorly expanded mid-proximal femoral diaphysis, and less robust pedal proximal phalanges. The Palomas Neandertals contrast with more northern European Neandertals particularly in various reflections of overall body size. Am J Phys Anthropol, 2011. © 2010 Wiley-Liss, Inc. [ABSTRACT FROM AUTHOR]

Abstract: Bolomor Cave, located in Tavernes de la Valldigna (Valencia, Spain), contains a sedimentary deposit composed of seventeen stratigraphic levels ranging from MIS 9 to MIS 5e. This extensive sequence allows us to ask different questions about how the human groups of this chronological period related to their environment. We report several different strategies by which animal resources were obtained from level XII faunal assemblage (MIS 6). These practices range from scavenging to the complex hunting techniques, such as multiple predation. This practice is inferred in the case of horses (Equus ferus) on the basis of: (1) the proportional representation of skeletal elements (elements with high marrow content, such as stylopodials and zeugopodials); (2) age at death of these animals (from infantile to advanced adult); (3) the position and function of cutmarks and toothmarks (mainly oblique and longitudinal incisions on limb bones diaphyses); (4) archaeostratigraphic distribution of their remains in the assemblage; and (5) presence of refits among the bones located at this archaeostratigraphical line. Evidence of anthropogenic processing of small prey (Oryctolagus cuniculus and Cygnus olor) is also identified at level XII. These discoveries imply the development of techniques different from those used to capture large and medium ungulates. From this perspective, a variety of strategies to obtain animal food can be documented at Bolomor Cave. This phenomenon can be related to the highly adaptive subsistence strategies of these hominids, who could take advantage of the benefits offered by a diverse and rich environment. In general, the aim of this study is to provide data about the subsistence strategies of human groups from Bolomor Cave and contribute to knowledge about the human groups’ way of life of in the late Middle Pleistocene. [ABSTRACT FROM AUTHOR]

This paper presents the first detailed analysis of the lithic industry from El Sidrón Cave. Previously we have published other works concerning Anthropology and Palaeogenetics. The lithic assemblage contains 415 artefacts coming from both unofficial and official excavations and is largely flake-based. The raw material derives from the immediate cave environment, and is mostly chert, with quartzite in a lesser proportion. To date, 67 artefacts have been refitted, including some tools and cores. The spatial distribution of the refitted lithic artefacts confirms, on the one hand, the unity of the assemblage and, on the other, its secondary context. [ABSTRACT FROM AUTHOR]

A well-preserved 11.8-million-years-old lower face attributed to the seminal taxon Dryopithecus fontani (Primates, Hominidae) from the Catalan site ACM/C3-Ae of the Hostalets de Pierola area (Vallès-Penedès Basin, Catalonia, NE Spain) is described. The new data indicate that D. fontani is distinct at the genus level from Late Miocene European taxa previously attributed to Dryopithecus, which are here reassigned to Hispanopithecus. The new facial specimen also suggests that D. fontani and the Middle Miocene Pierolapithecus catalaunicus are not synonymous. Anatomical and morphometric analyses further indicate that the new specimen shows a combination of lower facial features-hitherto unknown in Miocene hominoids-that resembles the facial pattern of Gorilla, thus providing the first nondental evidence of gorilla-like lower facial morphology in the fossil record. Considering the current evidence, the gorilla-like facial pattern of D. fontani is inferred to be derived relative to previously known stem hominids, and might indicate that this taxon is either an early member of the Homininae or, alternatively, a stem hominid convergent with the lower facial pattern of Gorilla. The biogeographic implications of both alternatives are discussed. This new finding in the Hostalets de Pierola section reinforces the importance of this area for understanding the elusive question of the Middle Miocene origin and early radiation of great apes. [ABSTRACT FROM AUTHOR]

Abstract: The archaic lithic industries of Barranco León and Fuente Nueva sites in Orce in Andalusia and the stone tools of the Vallonnet french site was studied by petroarcheological analyses and reveal the sources exploited by the hominids. The localisation of these sources give data to define the territory exploited. These three examples provide the opportunity to assess the raw material selectivity, the behaviour and the territory occupation system of the first inhabitants of Europe. These lithic industries are composed by local raw material collected very close to the site or even on the site. These sources were probably an important argument on choosing the occupation site. [Copyright &y& Elsevier]

Middle Paleolithic fossil human remains from the Sima de las Palomas in southeastern Iberia (dated to ≤43,000-40,000 calendar years before present) present a suite of derived Neandertal and/or retained ancestral morphological features in the mandibular symphysis, mandibular ramus, dental occlusal morphology, and distal hand phalanx. These traits are combined with variation in the mandibular corpus, discrete dental morphology, tooth root lengths, and anterior dental size that indicate a frequency difference with earlier Iberian and more northern European Neandertals. The Palomas Neandertals therefore confirm the late presence of Neandertals associated with the Iberian persistence of the Middle Paleolithic, but suggest microevolutionary processes and/or population contact with contemporaneous modern humans to the north. [ABSTRACT FROM AUTHOR]

Abstract: We present the description of a new mandibular specimen, ATD6-113, recovered in 2006 from the TD6 level of the Gran Dolina cave site in Sierra de Atapuerca, northern Spain. A detailed study of the lithostratigraphy of the top sequence of this level, the section from where all human remains have been recovered so far, is also presented. We have observed that the hominin stratum, previously defined as Aurora Stratum, represents a condensed deposit of at least six layers, which could not be distinguished in the test pit made in 1994–95. Therefore, the human fossil remains were probably deposited during a discrete and undetermined time period. The new mandibular fragment exhibits a very similar morphology to that of the most complete specimen, ATD6-96, which was recovered in 2003 from a different layer. This suggests that both specimens represent the same biological population. The two mandibles, as well as the small mandibular fragment ATD6-5 (which constitutes part of the holotype of Homo antecessor), present a morphological pattern clearly derived with regard to that of the African early Homo specimens usually included in H. habilis and H. rudolfensis, the mandibles D211 and D2735 from Dmanisi, and most of the early Pleistocene mandibles from Sangiran. The TD6 mandibles also exhibit some derived features with regard to the African early Pleistocene specimens included in H. ergaster (or African H. erectus). Thus, the TD6 hominins seem to represent a lineage different from other African and Asian lineages, although some (metric in particular) similarities with Chinese middle Pleistocene mandibles are noted. Interestingly, none of the apomorphic mandibular features of the European middle and early late Pleistocene hominins are present in the TD6 mandibles. [Copyright &y& Elsevier]

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Copyright of Archivo de Prehistoria Levantina is the property of Museu de Prehistoria de Valencia and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)

Presents information on a study which examined the paleomagnetic data on the Middle Pleistocene hominid-bearing strata in the Sima de los Huesos, North Spain. Methodology and geological setting of the study; Results and discussion of the study; Conclusions.

Focused on a study that analyzed the buccal striation pattern in teeth samples from Homo heidelbergensis hominid fossils recovered from Sima de los Huesos, Spain. Inferences about dietary behavior from analysis; Results and discussion; Conclusions.

Presents the analysis of an almost complete human male pelvis, Pelvis 1, from the Middle Pleistocene site of Sima de los Huesos in Sierra de Atapuerca, Spain. Other fossils from this site, and their dating and classification; Details about the pelvis; How a female pelvis would differ; Body mass of the male; Encephalization quotients compared with Neanderthals and modern humans.

Studies interproximal grooving in the hominid dentitions recovered from the Sima de los Huesos Middle Pleistocene cave site of the Sierra de Atapuerca in Spain. Observations of interproximal wear grooves in maxillary and mandibular posterior teeth; Similarity of wear grooves to those in other fossil hominids.

Studies the prevalence and chronology of enamel hypoplasias in a hominid dental sample at the Sierra de Atapuerca in Spain. Hypoplasias in permanent dentition; Enamel defect in the decidous tooth; Highest frequency of enamel hypoplasia; Comparison of the prevalence and severity of the hypoplasias of the hominid sample and a large Neandertal sample.

Reports the discovery of Homo antecessor, a species of hominid from the lower Pleistocene cave site of Gran Dolina, Sierra de Atapuerca, Spain. Fully modern midfacial morphology of the fossils; Traits defining the fossil; Other fossils discovered at the site; Where H. antecessor is likely situated on the human evolutionary tree.

Reports on the human remains dating more than 780,000 years as associated with a rich faunal and lithic assemblage in the Pleistocene cave site of Gran Dolina, Sierra de Atapuerca, Burgos, Spain. Micrommal species as representing the late Biharian; Lithic objects; Consideration for the hominid fossils.

The Guadix-Baza Basin, in SE Spain, harbors hominin fossils and lithic artifacts dated to ca. 1.4–1.3 Ma, representing the first hominin habitat in the Iberian Peninsula and possibly in Western Europe. Recent palynological studies have described a high diversity of plant taxa and biomes existing in the basin at the time of hominin presence. However, the relationship between these hominins and their environment has not been fully explored. Two novel methodologies are developed. The first method maps the distribution of the Early Pleistocene vegetation units based on paleobotanical and paleogeographic data. The second method assesses the availability of edible plant parts using a combination of Early Pleistocene and modern taxa lists. The resulting vegetation maps reveal a great diversity of vegetation types. During dry (glacial) periods, the vegetation of the basin was represented mostly by steppes, with the appearance of forested vegetation only in the mountainous regions. During humid (interglacial) periods, Mediterranean woodlands represented the dominant vegetation, accompanied by deciduous and conifer forests in the areas of higher altitude. The lake system present in the basin also allowed for the presence of marshland vegetation. The assessment of the availability of edible plant parts reveals that early Homo could have found a high number of resources in marshland and riparian environments throughout the year. Mediterranean woodlands and deciduous forests also provided numerous edible plant parts. During dry periods, the availability of plant resources decreased heavily, but the prevalence of marshland and riparian vegetation and of forested vegetation in the areas of higher altitude could have sustained hominin communities during harsher climatic periods. However, the disappearance of the lake system and an increase of aridity after the Mid-Pleistocene Transition and during the Middle Pleistocene probably led to an impoverishment of plant resources available to early Homo in the Guadix-Baza Basin. • Maps for glacial and interglacial periods show great vegetation diversity. • Earliest Homo in the Iberian Peninsula had access to multiple sources of plant food. • Riparian and marshland vegetation were an important source of food during glacials. • During Early Pleistocene, the basin could have acted as a glacial refugium for Homo. • The loss of marshlands in the Middle Pleistocene may have affected hominins. [ABSTRACT FROM AUTHOR]

Atapuerca, in the north of Spain, is the archaeological site where the oldest hominid remains within Europe have been found. In 2008 a jaw fragment, corresponding to the symphyseal area, was discovered in the area called the 'Elephant's pit'. Its age has been estimated at 1.2 million years and it is considered to be the oldest human fossil found in Europe and is from the lower Pleistocene. This work analyses the dental and skeletal damage to the specimen, detected in a macroscopic study of possible horizontal and vertical bone loss at the level of support of the remaining teeth. The limited presence of dental scale, the pattern of destruction and the decreased bone density due to increased marrow spaces suggest the presence of possible periodontal disease. [ABSTRACT FROM AUTHOR]

The Early Pleistocene sites of Barranco León and Fuente Nueva 3 (Guadix-Baza Basin, SE Spain) have yielded abundant Oldowan lithic artifacts and one hominin tooth (Homo sp. in level D1 or D2 of Barranco León), today considered to be among the earliest evidence for a hominin presence in Western Europe, at ca. 1.4–1.2 Ma. Here, for the first time, the stratigraphic succession of these two sites are studied more precisely from a palaeoenvironmental point of view, taking into account the different levels of the depositional sequences to analyze the successive fossil assemblages of amphibians and reptiles. Palaeoenvironmental reconstructions are carried out by applying the "habitat weighting" method, which uses the modern distribution by habitat of amphibian and reptile species in order to interpret past landscapes. The successive herpetofaunal assemblages from Barranco León show a certain tendency towards more arid conditions from level D1 to level E, whereas in Fuente Nueva 3 environmental reconstructions reveal oscillating conditions, with a tendency towards more arid conditions in the basal part of the sequence, up to level 5, where the tendency shifts back to more humid conditions. Our results show that the layers included in this study with the highest density of anthropic evidence (such as level 5 in FN 3 and levels D1 and D2 in BL) are situated within the late Early Pleistocene climatic and environmental cyclicity, yielding different environmental conditions: a humid, wooded biotope for BL, and a more open and drier biotope in FN 3. This suggests that the hominins of the late Early Pleistocene, although conditioned to some extent by climatic factors, were able to cope with changing environmental conditions, both "interglacial" and "glacial", in the southwestern extremity of the European continent. • Amphibians and reptiles are good proxies for the environmental reconstructions. • Paleoenvironmental conditions are reconstructed by stratigraphic levels. • The first hominins in Western Europe coped with changing environmental conditions. • The paleoenvironmental changes are in agreement with the climate oscillations. [ABSTRACT FROM AUTHOR]

The article reports on the discovery of the presence of hominids in the Andalusian village of Orce in Spain based on artifacts and fossils from area taken during the International Congress of Human Paleontology in 1995. According to the author, Andalusian village of Orce where the congress met dwells in the southeastern part of the Iberian Peninsula, a region Washington Irving described in 1829 as partaking of the savage and solitary character of Africa. Similarity of the sites to African landscapes, and possible route of the hominid dispersal from Africa are analyzed and discussed.

Reports evidence to suggest that Homo antecessor, a 800,000-year-old human species from the Atapuerca Mountains of Spain, may be ancestral to both modern humans and Neandertals. Report by Rosas et al. on page 1392; The fossils representing the remains of the oldest known Europeans; Where the species fits on the human family tree; Changes to the human evolutionary tree in the 1990s. INSET: Into the pit of human history..

Reports on a paleomagnetic investigation at the Gran Dolina site excavation in Atapuerca, Spain, showing that the sediments containing the discovered human occupation were deposited more that 780,000 years ago, near the time of the Matuyama-Brunhes boundary. Number of oriented samples obtained; Reversed-polarity displayed by magnetization.

The basin of Baza is located in the central area of the Betic Range (SE Spain) with a catchment area of over 4000 km2. It contains a continuous and well preserved lacustrine sedimentary record from the Late Miocene to the Middle Pleistocene. The basin encloses numerous Miocene to Pleistocene mammal fossil sites that include some of the oldest hominin occurrences in Western Europe, found at the Orce sites. The present work features the palynological analysis of sediment samples obtained from the Palominas drill core. The Palominas core cuts thorough fine grained sedimentary units located in the depocenter of the basin, which contain well preserved pollen grains. The palynoflora found in the core is rich in herbaceous pollen, as well as Quercus and Pinus , with other tree taxa appearing in lesser quantities. Spectral analysis reveals strong climatically controlled vegetation successions. The application of the Biomization method on these assemblages allows for a reconstruction of the biome succession. The Early Pleistocene landscape was dominated by a sequence of steppe, evergreen-bushland and mixed-forest biomes. The Coexistence Approach method shows that SE Spain experienced humid periods, during which precipitations were significantly higher than during the modern climate. These humid phases are correlated with warm interglacial periods which featured the development of open forests. Dryer phases and fully open environments are correlated with colder glacial periods. Based on the palynoflora a Calabrian age is suggested for the Palominas core. The flora and climatic patterns identified in Baza are similar to the ones identified in Calabrian records of Southern Italy and Greece. Image 1 • The longest Early Pleistocene pollen record from continental deposits in Spain. • The core could belong to the Calabrian stage, comprising over 400 kyr. • Eleven humid phases are defined, superimposed by long-term climatic trends. • A Mediterranean latitudinal gradient reflects drier and more open conditions in southern parts of Spain, Italy and Greece. [ABSTRACT FROM AUTHOR]

Here we present the descriptive and comparative study of two immature scapulae recovered from the TD6.2 level of the Gran Dolina cave site (Sierra de Atapuerca, Spain) and assigned to Homo antecessor. This is the first time that data on the morphology and dimensions of the scapulae of a European late Early Pleistocene hominin population are provided. Considering the state of development and the linear dimensions, the scapula ATD6-116 could belong to a child of about 2–4 years. The morphology of ATD6-116 clearly departs from that of the Australopithecus afarensis juvenile specimen DIK-1-1, pointing to functional differences in locomotor behavior between Australopithecus and the late Early Pleistocene hominins. The immature scapula ATD6-118 belonged to an immature individual with a development of the scapula equivalent to that of adolescents of recent human populations. The scapulae ATD6-118 and KNM-WT 15000 present a similar state of development. Although the scapula KNM-WT 15000 is clearly larger than ATD6-118, these two specimens share some characteristics such as their relative narrowness and the value of the axilloglenoid and spinoglenoid angles. The glenoid fossa of ATD6-116 show a lateral orientation, whereas in ATD6-118 the glenoid fossa is slightly cranially oriented, but still within the range of variation of modern humans. The glenoid index of both ATD6-116 and ATD6-118 is low in accordance to the values usually observed in other early hominins, thus showing the primitive condition for this feature. Both scapulae show a ventrally placed axillary sulcus. The presence of this primitive feature in ATD-116 confirms that the shape of the axillary border has a genetic basis and it is not related to physical activity. [ABSTRACT FROM AUTHOR]

Fossil bats are common components of Pleistocene palaeontological cave-sites, sometimes appearing in levels with evidence of a human presence. As with other small vertebrates, variations observed in fossil bat palaeocommunities through a stratigraphic sequence may be correlated to palaeoclimatic and palaeoenvironmental fluctuations. As many bat species are typically cave-dwelling animals, however, the bat palaeocommunity may also be affected by hominin presence. The correlation between bat palaeocommunity dynamics and both climatic and anthropic factors is addressed in the present study by analysing their record along the Early and Middle Pleistocene sequence in the Sierra de Atapuerca. A marked deterioration in the Sierra de Atapuerca chiropter faunas occurred from 500 ka on simultaneously with a change in the occupational pattern of the predominant bat species Myotis myotis, which apparently stopped using the shelters as nursery roosts. These changes coincides with a time of extreme cold events succession during glacial periods, but also with evidence of intensive human occupation at some of the sites analysed. Some features of bat assemblages at these levels, such as differences in species composition between synchronous levels, the inability of bat palaeocommunities to recover during intermediate warmer periods, or detected seasonal alternation between hominis and humans, suggest that anthropic disturbance could have play a significant part in the decline of Middle Pleistocene bat palaeocommunities of Atapuerca. • Diversity and richness of Atapuerca bat palaeocommunities decrease from 500 ka on. • From there on, there is no record of Myotis myotis using the caves as nursery roosts. • Bat palaeocommunities impoverishment was partially driven by climate factors. • Myotis myotis occupational pattern change could be due to both climate and anthropic factors. • Hominin presence likely played a part in Atapuerca Middle Pleistocene bats decline. [ABSTRACT FROM AUTHOR]