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Guiodytes inexpectatus Tian & Zhou sp. nov. urn:lsid:zoobank.org:act: EA3D7117-F8F3-4F45-848A-9E3879ABCBF8 Figs 1, 3F, 4F, 9, 10D, 11 Diagnosis This new species is most similar to G. bedosae Tian, 2014, which also occurs in Longzhou County, by the similar character configuration of the head and elytra and the seven-setose labrum present in both species. It is easily separated from the latter species by its much larger body, conspicuously emarginate labrum, the strongly protruding fore angles of the pronotum and the special elytral setiferous chaetotaxy. Etymology The species epithet refers to the unexpected discovery of this new species. The single specimen was encountered by Jiajun Zhou (the second author) after diving in the cave. It is his first discovery of a new cave beetle. Type material Holotype CHINA ��� ♀, broken on the left supra-antennal plate; Guangxi, Longzhou, Xiadong, Zhuzhu Dong; 22��24���10.75��� N, 106��38���52.36��� E; alt. 170 m; 18 Oct. 2020; J. Zhou leg.; SCAU. Description Female MEASUREMENTS. Length: 7.5 mm; width: 2.4 mm. Habitus as in Fig. 9. BODY. Moderately depigmented, concolorous brown. HEAD. Stout (Fig. 3F), much longer than wide from apex of mandible, HLm/HW=1.44, or slightly shorter than wide when measured from apex of labrum, HLl/HW=0.97; clypeus strongly transverse and with anterior margin slightly, median part moderately concave, bisetose at base, frontoclypeal sulcus well marked, clypeal wings not projecting, nearly at same level as clypeus, distinctly separated from clypeus and supra-antennal plates by notches; supra-antennal plates well developed, reflexed margined, strongly convex, smooth and glabrous; frons and vertex moderately convex, with transverse rugosity; with two setiferous pores, situated at posterior end of supra-antennal plates and before neck constriction, respectively, frontal furrows distinct, wide, slightly divergent towards middle of frons, strongly divergent posteriorly; frontal carinae blunt, wide, parallel; eyes completely lacking; genae well developed, not parallel-sided, sides tumid; neck constriction distinct, wide; labrum transverse, frontal margin conspicuously emarginate as a semicircular incision, seven-setose along frontal part, with six cilia on both sides; mandibles short and stout. Palpomeres ensiform, glabrous, apical segments of maxillary palpomere about twice as long as penultimate one, labial palpomeres subequal; labial palpomere two bisetose on inner margin; ligula unisetose at apex; labial suture well marked; mentum well developed, with two pairs of setae, one pair situated beneath median tooth, the other at base near lateral margin, with two large and deep concavities near base; median tooth simple, blunt at apex, lateral lobes wide, gently and obliquely truncated; basal foveae slightly convex; submentum narrow, quadrisetose. Antennae filiform, elongated, reaching elytral base; scapus unisetose subapically, pedicellus smooth and glabrous, pubescent from antennomere three; antennomere three slightly longer than pedicellus, then gradually decreasing in length towards penultimate segment, which is slightly shorter than terminal segment. Underside of head rough. THORAX. Pronotum peltate (Fig. 3F), much wider than head, PW/HW =1.47, slightly shorter than wide, PL/PW=0.90; disc moderately convex; anterior margin unbordered, indistinctly convex in median portion, margin with a row of cilia throughout, anterior transverse line marked with punctation, fore angles strongly protruding; widest at end of second third, gently and gradually narrowed anteriorly, with conspicuous notch before posterior angle, posterior angle large, with two conspicuously toothed projections between posterior angle and basal constriction; with two lateral setiferous pores, anterior seta at about anterior fifth, posterior one slightly before notch at posterior angle; reflexed lateral margin slightly crenulate, lateral channel broad throughout, narrower before anterior seta, wider between anterior and posterior lateral setae, ending at notch of posterior angle; basal constriction wide, unbordered, with narrow transverse carina; median line engraved, wide, not joining basal constriction. Peduncle short, scutellum small. Prosternum and propleura smooth, with sparse punctures. ABDOMEN. Abdominal ventrites more or less punctured; ventrite seven with two pairs of subapical setae, widely separated at each side. WINGS. Elytra elongate ovate, strongly convex, wider than pronotum, EW/PW= 1.29, much longer than wide, EL/EW =1.67; widest at about one third from apex, slightly contracted anteriad and moderately and regularly contracted posteriad; base with a pair of setiferous punctures at base of stria one, intervals two and three each with a pair of small tubercles at base; shoulders broadly obtuse, with two large teeth on each side; apex pointed; sides distinctly crenulate from base to apical third, then indistinctly continued towards apex; elytral striae broad and well marked, with large and isolated punctures, only striae six and seven ending before reaching basal margin of elytron; intervals slightly convex; intervals six and seven carinate at base, joining and continuing into humeral tooth, interval eight carinate throughout, joining with seven before reaching base; arrangement of elytral setiferous chaetotaxy as in Fig. 3F: nine and six foveolate setiferous dorsal pores on interval three and five, respectively, pores nearly regularly arranged; with series of setiferous pores in marginal channel, uninterrupted and small, with several large pores bearing much longer setae. Hind wings reduced. LEGS. Moderately elongated; proleg stout, profemur moderately dilated, smooth, with a long seta and several short ones; protibia well developed, with distinct and complete carina dorsally, sulcus distinct, quadridentate; lateral teeth blunt at apex, lowest lateral much shorter than others, upper two stoutensiform; subapical spur elongate-ensiform, blunt at apex, shorter and more slender than uppermost lateral teeth; protarsi slender, tarsomere longer than tarsomeres two to four combined; meso- and metalegs slender, with width of tarsomeres in both narrower than in proleg; mesotibia gradually dilated towards apex, with an elongated tuber-like subapical spur, longer than wide and furnished with an isometric seta. GENITALIA (Fig. 10D). Similar to those of G. cavicola (Fig. 10B), gonosubcoxite IX stout, with one long seta and two short setae at median portion of outer margin; gonocoxite IX curved and slender, apex very sharp, with three dorsal and three ventral setae, plus two short and sturdy ensiform setae on outer margin. Male Unknown. Distribution China (Guangxi: Longzhou County) (Fig. 1). Known only from Zhuzhu Dong cave (Fig. 11). The cave is not easy to access due to the extremely narrow entrance as well as the vertical depth of three meters after the entrance. Then, it reaches a relatively wider and flat passage. But the ventilation of the passage is inadequate, causing the collector to experience symptoms of shortness of breath and chest distress. The single specimen was found on the muddy surface near the bank of an underground river, which is 50 meters from the entrance., Published as part of Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi & Faille, Arnaud, 2021, Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini), pp. 135-154 in European Journal of Taxonomy 774 on pages 148-152, DOI: 10.5852/ejt.2021.774.1537, http://zenodo.org/record/5567094, {"references":["Tian M. Y. 2014. New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae). Zootaxa 3861 (4): 355 - 362. https: // doi. org / 10.11646 / zootaxa. 3861.4.5","Tian M. Y. 2013. Occurrence of troglobitic clivinines in China (Insecta: Coleoptera: Carabidae). Journal of Cave and Karst Studies 75 (2): 113 - 120. https: // doi. org / 10.4311 / 2011 LSC 0226"]}

Guiodytes yueliangensis Huang & Tian sp. nov. urn:lsid:zoobank.org:act: 54A7DD47-8EAB-4A9A-9C49-2E6383B4842A Figs 1, 3E, 4E, 5D, 7–8 Diagnosis This new species belongs to the group with distinctly projecting clypeal wings and is close to G. deharvengi Tian, 2014, which also occurs in the same karst area in Huanjiang County, by the similar character configuration of head and elytra, but it is easily separated from the latter species by the missing central pore on the frons of the head and the flattened neck constriction. In addition, the pronotum, with the anterior margin almost not emarginate, is much longer, its median line is much narrower, and the anterior transverse line is finely developed. Etymology The species epithet is derived from Yueliang Shan (Moon Mountain in Chinese), a famous karst landscape with a naturally-formed moon-like cavern passing through it (Fig. 8A). The type locality, Huang Dong, is one of the caves of this landscape. Type material Holotype CHINA • ♂, broken on the left supra-antennal plate; Guangxi, Huanjiang, Shuiyuan, Sancai, Neiwen, Huang Dong; 24°50′05.74″ N, 108°05′1.86″ E; alt. 354 m; 6 Jun. 2017; S. Huang leg.; SCAU. Description Male MEASUREMENTS. Length: 9.3 mm; width: 2.9 mm. Habitus as in Fig. 7. BODY. Well-pigmented, concolorous dark brown. HEAD. Stout (Fig. 3E), from apex of mandible slightly longer than wide, HLm/HW= 1.06, or shorter than wide when measured from apex of labrum, HLl/HW = 0.82; clypeus strongly transverse and moderately emarginate, surface of median part moderately convex, bisetose at base, with flattened clypeofrontal sulcus, clypeal wings fused, divided from supra-antennal plates by distinct notches; supra-antennal plates rounded and well developed, reflexed margined, strongly convex, smooth and glabrous; frons and vertex fairly convex; laterally with two setiferous pores, situated at posterior end of supra-antennal plates and at level of neck constriction, respectively; frontal furrows deep, conspicuously widened, subparallel at middle of frons, divergent posteriorly; frontal carinae blunt but well marked, not parallelsided; eyes completely lacking; genae well developed, anteriorly conically raised laterally then sinuated, sides nearly vertically truncated; neck constriction distinct, relatively wide; labrum transverse, slightly emarginate, slightly wider than clypeus, seven-setose, ciliate laterally; mandibles of moderate size. Palpomeres ensiform, glabrous, apical segments of both maxillary and labial palpomeres much longer than penultimate ones, respectively; labial palpomere two bisetose on inner margin; ligula unisetose at apex; labial suture well marked, deeply and widely furrowed in median portion; mentum well developed, with two pairs of setae, one pair situated beneath mental tooth, posterior one localized in centre of two large, deep concavities restricted at base near lateral margin; median tooth simple, pointed at apex, lateral lobes wide, obliquely truncated; submentum narrow, quadrisetose. Antennae filiform, elongate, reaching over base of pronotum; scapus unisetose subapically, pedicellus smooth and glabrous, pubescent from antennomere three; pedicellus slightly longer than antennomere three, then gradually decreasing in length towards penultimate segment, antennomeres five to ten subelongate, penultimate segment slightly shorter than terminal one. Underside of head rough. THORAX. Pronotum peltate (Fig. 3E), much wider than head, PW/HW=1.25, slightly shorter than wide, PL/PW=0.96; disc strongly convex and with noticeable rugosity; anterior margin unbordered, slightly convex in median portion, margin barely emarginate, anterior transverse line fine and traceable, fore angles protruding; widest slightly behind middle, gently curved and gradually narrowed anteriorly, more evidently contracted towards posterior angles; with distinct teeth at posterior angles, lateral margin between posterior angles and basal constriction with one conspicuous and one minute toothed projections before basal constriction; with two lateral setiferous pores, anterior one at about apical quarter, posterior one at basal angle, just before tooth at posterior angle; reflexed lateral margin nearly smooth, lateral channel with uneven surface, narrower before anterior seta, wider between anterior and posterior lateral setae, interrupted by tooth of posterior angle; basal constriction wide, with noticeable transverse carina; median line deeply furrowed but faint when joined with transverse line in front. Peduncle short, scutellum small. Prosternum and propleura smooth, with dense and isodiametric punctuation. ABDOMEN. Abdominal ventrites also densely punctured; ventrites four to six each with a pair of setae; ventrite seven with two pairs of subapical setae, widely separated at each side. WINGS. Elytra elongate ovate, wider than pronotum, EW/PW= 1.29, much longer than wide, EL/ EW=1.84; strongly convex, widest at middle, gently contracted anteriad and posteriad; base with pair of basal setiferous pores located at beginning of stria one, joined with an additional tubercle at base of interval two; shoulders broadly obtuse, with two teeth on each side; apex pointed; sides distinctly crenulated from base to apical third, then very feebly marked towards apex; elytral striae punctatestriate, narrow and deep, with small and isolated punctures, only striae six and seven ending before reaching basal margin of elytron; intervals distinctly convex; intervals seven and eight joined near base, then joined to interval six at base, intervals six to eight carinated near base; arrangement of elytral setiferous chaetotaxy as in Fig. 4E: five foveolate setiferous pores present on interval three; marginal channel with uninterrupted series of small setiferous pores and several large pores bearing much longer setae. Hind wings reduced. LEGS. Legs moderately elongated; proleg stout, profemur moderately dilated, smooth, with several sparsely distributed setae; protibia well developed, with distinct and complete carina dorsally, sulcus indistinct, quadridentate; lateral teeth blunt at apex, lowest lateral tooth unobtrusive, upper three stoutensiform and moderately protruded anteriorly toward apex; subapical spur elongate-ensiform, blunt at apex, much longer and more slender than uppermost lateral teeth; protarsi slender, tarsomere one very long, much longer than tarsomeres two to four combined; meso- and meta-legs slender, with width of tarsomeres in both narrower than in proleg; mesotibia gradually dilated towards apex, with elongated tuber-like subapical spur, distinctly longer than wide and furnished with isometric seta. GENITALIA (Fig. 5D). Moderately sclerotized; aedeagus similar to that of G. deharvengi Tian, 2014 (Fig. 5A); median lobe strongly arcuated ventrally, elongated and blunt at apex; copulatory piece irregular, elongated, strongly sclerotized in basal and frontal part; parameres asymmetrical, large one with three setae, small one with four setae. Female Unknown. Distribution China (Guangxi: Huanjiang County) (Fig. 1). Known only from Huang Dong cave (Fig. 8B). The cave is still preserved as a natural environment and the entrance opens at the middle of the hill. The cave is not well explored and its length remains unknown. The passages in the cave are complicated, and most of them are moist and wet. The single beetle specimen was collected in a wet place about 100 m from the entrance (Fig. 8C)., Published as part of Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi & Faille, Arnaud, 2021, Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini), pp. 135-154 in European Journal of Taxonomy 774 on pages 144-148, DOI: 10.5852/ejt.2021.774.1537, http://zenodo.org/record/5567094, {"references":["Tian M. Y. 2014. New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae). Zootaxa 3861 (4): 355 - 362. https: // doi. org / 10.11646 / zootaxa. 3861.4.5"]}

Guiodytes weii Huang & Faille sp. nov. urn:lsid:zoobank.org:act: E0CD76FD-36A6-48D6-BDEE-21A62C3FDABD Figs 1–2, 3D, 4D, 5C, 6 Diagnosis This new species is most similar to G. deharvengi Tian, 2014, which also occurs in Huanjiang County, by the similar character configuration of the head and elytra. It is easily separated from the latter species by the clypeal wings not projecting anteriorly. Moreover, its body shape is much smaller and more slender, the elytral stria punctures are larger and sparser, and intervals four and five are carinated near the base. Etymology This new species is dedicated to Mr. Guofu Wei (Center of World Natural Heritage, Huanjiang) for his support during our biospeleological investigations. Type material Holotype CHINA • ♂; Guangxi, Huanjiang, Jiale, Dapo Dong; 25°06′06.78″ N, 108°31′12.48″ E; alt. 211 m; 5 Dec. 2016; A. Faille leg.; SCAU. Description Male MEASUREMENTS. Length: 4.5 mm; width: 1.1 mm. Habitus as in Fig. 2. BODY. Slightly depigmented, concolorous brown. HEAD. Stout (Fig. 3D), from apex of mandible much longer than wide, HLm/HW=1.41, or slightly longer than wide when measured from apex of labrum, HLl/HW =1.06; distinctly narrower than pronotum, HW/ PW=0.67; clypeus with fused wings transverse, moderately convex medially, bisetose at base, anterior margin slightly bisinuate, bordered, clypeal wings not protruding anteriorly; supra-antennal plates well developed, gently rounded, reflexed, margined, strongly convex, smooth and glabrous; frons and vertex moderately convex, with longitudinal pore at middle; laterally with two setiferous pores, situated at posterior end of supra-antennal plates and at level of neck constriction, respectively; frontal furrows deep, wide, distinctly divergent posteriorly; frontal carinae distinct, slightly convergent to neck constriction; frontoclypeal sulcus not deep but distinct; eyes completely lacking; genae well developed, subparallelsided, sides nearly vertically truncated; neck constriction distinct, slightly stepped; labrum transverse, slightly wider than clypeus, seven-setose, ciliate laterally, straight at anterior margin; mandibles of moderate size. Palpomeres ensiform, glabrous, apical segments of both maxillary and labial palpomeres much longer than penultimate ones, respectively; labial palpomere bisetose on inner margin; ligula unisetose at apex; labial suture well marked, deeply and widely furrowed in median portion; mentum well developed, with two pairs of setae, one pair situated beneath mental tooth, the other at base near lateral margin, and with two large and deep concavities near base; median tooth simple, blunt at apex, lateral lobes wide, gently and obliquely truncated; submentum narrow, quadrisetose. Antennae filiform, rather short, reaching posterior angles of pronotum; scapus unisetose subapically, pedicellus smooth and glabrous, pubescent from antennomere three; pedicellus slightly longer than antennomere three; antennomeres five to ten subelongate, terminal segment slightly longer than penultimate. Underside of head rough. THORAX. Pronotum peltate (Fig. 3D), much wider than head, slightly longer than wide, PL/PW=1.06; disc smooth, moderately convex; anterior margin slightly concave, beaded in median line; fore angles slightly protruding; widest at beginning of posterior third, gently and gradually narrowed anteriorly, strongly contracted towards posterior angles; posterior angles with obtuse teeth, lateral margin between posterior angles and basal constriction with two conspicuously toothed projections; with two pairs of lateral setiferous pores, anterior one at about anterior quarter, posterior one at basal angle and distinctly removed from channel; reflexed lateral margin with slight notches, lateral channel narrow before anterior seta, distinctly widened between anterior and posterior lateral setae, ending before posterior seta; basal constriction wide, basal carina narrow; median line engraved, deep, wide, distinctly joining basal constriction, surpassing anterior transverse line without joining. Peduncle short, scutellum small. Prosternum and propleura smooth, with dense and isodiametric abdominal punctuation. ABDOMEN. Ventrites also densely punctured, ventrites four to six each with a pair of setae; ventrite seven with two pairs of subapical setae, widely separated at each side. WINGS. Elytra elongate ovate, wider than pronotum, EW/PW= 1.27, much longer than wide, EL/ EW=1.86; strongly convex; widest behind middle, gently contracted anteriad and posteriad; base finely bisinuate, with a pair of additional tubercles followed by a basal setiferous puncture located at base of interval two; shoulders broadly obtuse, with sharp tooth; apex of elytra pointed; sides distinctly crenulated from base to apical quarter; elytral striae punctate-striate, wide and deep, with large and isolated punctures, only striae six and seven ending before reaching basal margin of elytron; intervals distinctly convex; intervals one to four unbordered at base; intervals seven and eight joined near base, then joined to interval six at base, intervals six to eight carinated throughout, intervals four and five carinated near base; arrangement of elytral setiferous chaetotaxy as in Fig. 4D, five foveolate setiferous pores on interval three; marginal channel with uninterrupted series of small setiferous pores and several large pores bearing much longer setae. Hind wings reduced. LEGS. Moderately elongated; proleg stout, profemur distinctly dilated, smooth, with a long seta and two shorter ones; protibia well developed, with distinct and complete carina dorsally, sulcus indistinct, quadridentate; lateral teeth blunt at apex, lowest lateral one much shorter than others, upper two stoutensiform; subapical spur elongate-ensiform, blunt at apex, shorter and more slender than uppermost lateral teeth; protarsi slender, tarsomere one longer than tarsomeres two to four combined; meso- and meta-legs slender, with width of tarsomeres in both narrower than in proleg; mesotibia gradually dilated towards apex, with an elongated, tuber-like subapical spur, longer than wide and furnished with an isometric seta. GENITALIA (Fig. 5C). Moderately sclerotized; aedeagus similar to those of G. deharvengi Tian, 2014 (Fig. 5A) and G. cavicola Tian, 2013 (Fig. 5B), median lobe gently arcuated ventrally, whereas strongly arcuated in G. deharvengi, or evidently bisinuate in G. cavicola, blunt at apex; parameres asymmetrical, one much longer and broader than the other, both parameres with three long setae at apex. Female Unknown. Distribution China (Guangxi: Huanjiang County) (Fig. 1). Known only from Dapo Dong cave. This cave opens at the bottom of a hill near a country road in the village of Jiale. The entrance is narrow, but accessible (Fig. 6A). It is about 90 m long with a short and narrow side passage inside. A large part of the passage is dry, but there are some moist places. The single blind beetle specimen was discovered on the moist wall about 30 m from the entrance at the left side of the main passage. Other syntopic cave animals were observed during the investigation in the cave, for instance, a flatworm and a bat.(Fig. 6B, D)., Published as part of Huang, Sunbin, Zhou, Jiajun, Tian, Mingyi & Faille, Arnaud, 2021, Three new species of the subterranean genus Guiodytes from Guangxi, China (Coleoptera: Carabidae: Clivinini), pp. 135-154 in European Journal of Taxonomy 774 on pages 137-144, DOI: 10.5852/ejt.2021.774.1537, http://zenodo.org/record/5567094, {"references":["Tian M. Y. 2014. New records and a new species of the cavernicolous genus Guiodytes Tian, 2013 from Guangxi, China (Coleoptera: Carabidae: Scaritinae). Zootaxa 3861 (4): 355 - 362. https: // doi. org / 10.11646 / zootaxa. 3861.4.5","Tian M. Y. 2013. Occurrence of troglobitic clivinines in China (Insecta: Coleoptera: Carabidae). Journal of Cave and Karst Studies 75 (2): 113 - 120. https: // doi. org / 10.4311 / 2011 LSC 0226"]}

Plusiocampa (Stygiocampa) dalmatica Condé, 1959 Figs 205, 208 Plusiocampa dalmatica Condé, 1959: 48, fig. 2. Diagnosis (Condé 1959; Bareth 1988; Condé & Bareth 1996) Troglomorphic species. Antennae with 41–47 antennomeres; numerous sheet-shaped complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1+1 ma, 3+3–4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 1+1 la, 2+2 lp macrosetae; long, barbed notal macrosetae; long, barbed notal clothing setae. No ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergite III with 0+0–1+1 (1+0) la macrosetae; urotergite IV with 1+1 la, 0+1 post macrosetae; urotergite V with 1+1 la, 4+4 (3+4) post macrosetae; urotergites VI–VII with 1+1 la, 4+4 post macrosetae; urotergite VIII with 5+5–6+6 post macrosetae; abdominal segment IX with 8+8– 9+9 (10+9) post macrosetae. Urosternite I with 14+15–21+20, urosternites II–VII with 9+9–12+12, urosternite VIII with 2+2–3+3 macrosetae. Male with lateral expansions of posterior margin occupied by an area of glandular g 1 setae in adults (no expansion in juveniles and little expansion in young males), subcylindrical appendages without glandular a 2 setae; female with subcylindrical appendages with glandular a 1 setae. Remarks The studied material (see Table S 2 in Supplementary Material) allowed us to make additions to the diagnosis (Coll. AS). Habitat and distribution Subterranean species well distributed around the northwestern Dinaric Mountains, found in 18 caves (Figs 205, 208)., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 97-98, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1959. Un singulier diploure cavernicole de Dalmatie. Fragmenta Balcanica 2 (20): 165 - 172.","Bareth C. 1988. Campodes cavernicoles de la Venetie orientale (Frioule, Venetie Julienne) et de la Calabre (Insecta, Aprterygota Diplura). Atti e Memorie della Commissione Grotte \" Eugenio Boegan \" 27: 45 - 55.","Bareth C. & Conde B. 1996. Le complexe de Plusiocampa pouadensis (Denis) dans les grottes de la region pyreneenne. Memoires de Biospeologie 23: 127 - 131."]}

Plusiocampa (Plusiocampa) dolichopoda inguscioni Bareth, 2002 Plusiocampa dolichopoda inguscioni Bareth, 2002: 122, figs 1–2. Differential diagnosis against the type form (Bareth 2002) Pronotum with 3+3 lateral posterior macrosetae, instead of 2+ 2 in the type form. Habitat and distribution Subterranean subspecies inhabiting one cave, Grotte Scaloria, Manfredonia, Gargano, Italian Peninsula, near the type locality., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 45, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Bareth C. 2002. Campodeidea de la Grotte Scaloria (Gargano, Italia), Diplura, Campodeidae. Bulletin de l'Academie lorraine des Sciences 41: 3 - 4."]}

Plusiocampa (Plusiocampa) rhea Sendra sp. nov. urn:lsid:zoobank.org:act: 79F00B32-CCCA-4195-BB1F-52D3B2225C20 Fig. 124; Table 9 Etymology This species is named in honour of Rhea, mother of the Greek god Zeus and daughter of Gaea, the Earth goddess. She gave birth and cared for her son Zeus in the Ideon Andron Cave located in Mount Psiloritis, the highest mountain in Crete, where this new species was found. Material examined Holotype GREECE • 1 ♂; Crete, Anogeia, Ideon Antron Cave (Zeus Cave) at twilight zone area, near entrance; 1174 m a.s.l.; 22 Mar. 2006; Alberto Sendra leg.; MZB 2019-1033. Paratypes GREECE – Crete • 1 ♂; Pombia; 211 m a.s.l.; Alberto Sendra leg.; MSS habitat; Coll. AS • 1 ♂; Sissarha; 601 m a.s.l.; MSS habitat; Alberto Sendra leg.; Coll. AS. Description BODY. Body length 3.8–4.9 mm. Epicuticle smooth under optical microscope; body with thin, middlesized clothing, covered or not by a few thin barbs. HEAD. Three intact antennae in holotype and paratypes, with 30, 32 and 33 antennomeres, slightly shorter than body length (Table 9). Small coniform sensillum of third antennomere located in ventral position between c and d macrosetae. Central antennomeres 1.5× as long as wide, apical antennomere 1.6× as long as wide. Cupuliform organ occupying of total length of apical antennomere, with 8–9 complex olfactory chemoreceptors. Gouge sensilla 30 ⅕ – 32 µm long in a single distal whorl of 12–16 sensilla on each medial and distal antennomere. Frontal process plain, with non-tubercular setae and macrosetae with barbs along half to two-thirds; macrosetae along line of insertion of antennomere and x setae with similar lengths (a/i/p/x with relative lengths of 36/42/35/ 52 in holotype). Suboval labial palps with latero-external sensillum larger than sensillum of third antennomere, with two guard setae, up to 12 setae on anterior border and up to 180 neuroglandular setae. THORAX. Distribution of thoracic macrosetae (Fig. 124): pronotum with 1+1 ma, 2+2 la 3,4 and 2+2 sla 1,2 , 2+2 lp 1,3 (with additional 1+1 extra sma near the sagittal plane); mesonotum with 1+1 ma, 3+3 la 1,2,3 , 3+3 lp 1,2,3 , 1+1 mp; metanotum with 1+1 ma, 3+3 lp 1,2,3 , 1+1 mp. All notal macrosetae long, robust, covered by thin barbs along distal third or half; submacrosetae sla 1,2 and sma thinner and shorter than notal macrosetae, but with same large base; marginal setae longer than clothing setae, covered by thin barbs along distal three-fourths. Legs elongated, pretarsus of metathoracic legs slightly overpassing end of abdomen (Table 9). Femur I without dorsal macrosetae but with one long, barbed ventral macroseta; femora II–III with one long, barbed dorsal and one long, barbed ventral macroseta. Tibia I with one short, barbed ventral macroseta; tibiae II–III with two short, barbed ventral macrosetae. Calcars with 2–3 rows of long barbs from base to tip, smooth on one narrow side along calcar. Dorsal and lateral subapical tarsal setae smooth. Slightly unequal claws (posterior claw 1.15–1.2× as long as anterior one); posterior claw without backward overhang; lateral crests well developed. Pretarsal processes long, setiform, reaching extremities of claws. ABDOMEN. Distribution of abdominal macrosetae on tergites: 1+1 post 1 on I–III (0+1 la on III); 1+1 la, 2+2 post 1,2 on IV; 1+1 la, 5+5 post 1–5 on V–VII; 7+7 post on VIII; 9+9 post on abdominal segment IX. All post urotergal macrosetae thick, long and covered by thin barbs along distal four-fifths; la urotergal macrosetae shorter than post macrosetae, covered by barbs along distal half. Urosternite I with 9+9, urosternites II–VII with 5+5; urosternite VIII with 2+2 macrosetae; all urosternal macrosetae robust and large, covered by long barbs along distal three-fourths to four-fifths. Apical, subapical and ventromedial setae with abundant long barbs. CERCI. One incomplete cercus on 4 mm long paratype with six articles in addition to basal article, 3.8 mm long; their articles show whorls of long macrosetae covered by thin barbs along distal twothirds, combined with whorls of smooth, thin setae, shorter than macrosetae, in addition to one apical whorl of short setae with a few distal barbs. GENITAL ORGANS. Male urosternite I with up to 160 glandular g 1 setae arranged in up to five rows with slightly enlarged subcylindrical appendages, each bearing up to 19 glandular a 1 setae. Phyletic affinities, habitat and distribution Plusiocampa (P.) rhea sp. nov. has 3+3 lateral posterior macrosetae on the mesonotum and metanotum, a feature only found in P. (P.) pouadensis (Condé 1948c; Sendra & Escolà 2004). Nevertheless, P. (P.) rhea sp. nov. has medial posterior mesonotal and metanotal macrosetae, which are absent in P. (P.) pouadensis. Furthermore, P. (P.) pouadensis is a troglomorphic species from the Eastern Pyrenees, whereas P. (P.) rhea sp. nov. has been found in hypogean habitats, in soil, MSS and the twilight zones of caves at three locations on Crete., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 76-78, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1948 c. Addition a la faune des campodeides cavernicoles de France. Bulletin de la Societe des Sciences de Nancy, new series 7 (2): 45 - 54.","Sendra A. & Escola O. 2004. Campodeides cavernicoles du nord-est de la peninsule Iberique. Revue suisse de Zoologie 111 (4): 889 - 914. Available from https: // biodiversitylibrary. org / page / 41227849 [accessed 13 Nov. 2019]."]}

Plusiocampa (Plusiocmapa) solerii Silvestri, 1932 Plusiocmpa solerii Silvestri, 1932: 72, figs viii–ix. Diagnosis (Silvestri 1933a; Sendra et al. 2006) Non-troglomorphic species. Antennae with 22–25 antennomeres; frontal process with non-tubercular setae. Pronotum with 1+1 ma, 4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 1+1 la, 2+2 lp macrosetae; long, barbed notal macrosetae. Two ventral tibial macrosetae; slightly unequal claws (1.15) with large lateral crests, posterior claw with no backward overhang. Urotergites I–III with 1+1 post macrosetae; urotergite IV with 1+1 la, 2+2 post macrosetae; urotergites V–VII with 1+1 la, 4+4 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 macrosetae; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male unknown; female with long subcylindrical appendages with glandular a 1 setae., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 82, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Silvestri F. 1932. Campodeidae (Thysanura) de Espana (primera parte). Eos 8: 115 - 164.","Silvestri F. 1933 a. Nuovi contributi alla conoscenza della fauna delle isole italiane dell'Egeo. Bolletino del Laboratorio di Zoologia generale e agraria in Portici 27: 61 - 111.","Sendra A., Satar A. & Montagud S. 2006. Premiere contribution a la faune de Diploures Campodeides de la Peninsule d'Anatolie, Turquie (Diplura: Campodeidae). Revue suisse de Zoologie 113 (3): 693 - 709. Available from https: // biodiversitylibrary. org / page / 41228213 [accessed 13 Nov. 2019]."]}

Vandelicampa tisserandi Condé, 1955 Vandelicampa tisserandi Condé, 1955c: 407, figs 3–4. Diagnosis (Condé 1955c) Troglomorphic species. Antennae with 29–38 antennomeres; eight to ten olfactory chemoreceptors in cupuliform organ. Long, barbed notal macrosetae; sparse, smooth, long notal clothing setae. One dorsal femoral macroseta; two or three ventral tibial macrosetae; subequal claws (1.04) with large lateral crests, posterior claw without a backward overhang; laminar pretarsus processess with a sharp ending covered with long ventral barbs. Urotergites I–III with 1+1 post macrosetae; urotergite IV with 1+1 la, 3+3 post macrosetae; urotergites V–VII with 3+3 (2+2) la, 4+4 post macrosetae; urotergite VIII with 6+6 post macrosetae; urosternite I with 8+8, urosternites II–VII with 6+6, urosternite VIII with 2+2 macrosetae. Male with subcylindrical appendages of first urosternite similar to those of female. Habitat and distribution Subterranean species found in three caves in the karst areas of Lebanon (Fig. 210)., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 109, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1955 a. Materiaux pour une monographie des Diploures Campodeides. Memoires du Museum national d'histoire naturelle, serie A, Zoologie 12: 1 - 202.","Conde B. 1955 c. Mision Henri Coiffait au Liban (1951). 4. Protoures et Diploures Campodeides. Biospeologica 75: 397 - 412."]}

Patrizicampa Condé, 1962 Patrizicampa Condé, 1962: 199. Diagnosis (Condé 1962) Without dorsal femoral macrosetae or ventral tibial macrosetae; abundant macrosetae on all urosternites, including 3+3 macrosetae on urosternite VIII. Its apomorphic feature, the shape of the laminar pretarsal lateral process, is unique among diplurans. It starts with a large laminar lateral process, well barbed on the ventral side and ending in a thin, smooth, pointed process., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 12, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1962. Campodeides des grottes de Sardaigne (Insecta Diplura). Atti VII Congresso nazionale di Speleologia, Memoria III: 199 - 202."]}

Plusiocampa (Plusiocampa) isterina Condé, 1993 Plusiocampa isterina Condé, 1993a: 736, figs 1a, 2, 3a, 4, 6a. Diagnosis (Condé 1993a, 1996) Non-troglomorphic species. Antennae with 23 antennomeres; four simple olfactory chemoreceptors in cupuliform organ; slightly protruding frontal process with some slightly tubercular setae. Pronotum with 1+1 ma, 3+3 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 1+1 (2+1) la, 2+2 lp macrosetae; long, thin, barbed notal macrosetae; short, smooth clothing setae. Two ventral tibial macrosetae; unequal claws (1.29) with large lateral crests, posterior claw with a backward overhang; pretarsal processes short, with a few sparse barbs. Urotergites I–II with 1+1 post macrosetae; urotergite III with 2+2 post macrosetae; urotergites IV–V with 1+1 la, 5+5 post macrosetae; urotergites VI–VII with 2+2 la, 5+5 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7, urosternites II– VII with 5+5, urosternite VIII with 2+2 macrosetae. Male unknown; subcylindrical female appendages bearing glandular a 1 setae., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 64, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1993 a. Une lignee danubienne du genre Plusiocampa (Diploures Campodeides). Revue suisse de Zoologie 100 (3): 735 - 745. Available from https: // biodiversitylibrary. org / page / 41223871 [accessed 13 Nov. 2019].","Conde B. 1996. Diploures Campodeides de la Pestera de la Movile (Movile Cave), Dobroudja meridionales (Roumanie). Revue suisse de Zoologie 103 (1): 101 - 114. Available from https: // biodiversitylibrary. org / page / 41267657 [accessed 13 Nov. 2019]."]}

Plusiocampa (Plusiocampa) tinoamorei Sendra & Nicolosi, 2019 Plusiocampa (Plusiocampa) tinoamorei Sendra & Nicolosi in Sendra et al., 2019: 298, figs 1–22, tables 1–2.. Diagnosis (Sendra et al. 2019) Troglomorphic species. Antennae with 37–41 antennomeres. Very slight protruded frontal process with slightly differentiated smooth macrosetae. Pronotum with 1+1 ma, 4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp, 1+1 mp macrosetae; metanotum with 1+1 ma, 1+1 la, 2+2 lp, 1+1 mp macrosetae; long, barbed notal macrosetae; body with a small number of thin clothing setae with thin distal barbs. Two or three ventral tibial macrosetae; slightly unequal claws (1.08–1.12) with large lateral crests, posterior claw with backward overhang. Urotergites I–III with 1+1 post macrosetae; urotergite IV with 1+1 la, 3+3 post macrosetae; urotergites V–VII with 1+1 la, 5+5 post macrosetae; urotergite VIII with 6+6 (6+5) post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Stylar setae with a few thin barbs. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; appendages of female slightly thinner than those of male. Habitat and distribution Subterranean species found in three caves near Siracusa, southeastern Sicily., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 95, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Sendra A., Nicolosi G. & Amore E. 2019. Subterranean Campodeidae fauna from Sicily Island (Diplura); its biogeographical interest with the description of a new species of Plusiocampa. Zootaxa 4567 (2): 379 - 386. https // doi. org / 10.11646 / zootaxa. 4567.2.10","Sendra A. & Wagnell C. 2019. The cave-dwelling dipluran (Diplura, Campodeidae) on the edge of the Last Glacial Maximum in Vancouver Island caves, North America (Canada). Subterranean Biology 29: 59 - 77. https: // doi. org / 10.3897 / subtbiol. 29.31467"]}

Plusiocampa (Plusiocampa) dargilani (Moniez, 1894) Campodea dargilani Moniez, 1894: 82. Campodea staphylinus – Viré 1897: 7, figs 1–2 (not Westwood, 1852). Troglocampa dargilani – Denis 1930 pars: 231, figs 25–28. Diagnosis (Denis 1923; Condé 1946; Bareth 2006) Troglomorphic species. Antennae with 40–48 antennomeres; eight to nine complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1+1 ma, 4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 2+2 la, 2+2–1+1 lp macrosetae; thin, poorly barbed notal macrosetae; sparse long, thin, barbed notal clothing setae. No ventral tibial macrosetae; unequal claws with large lateral crests, posterior claw with a backward overhang. Urotergite III with 1+1 la macrosetae; urotergite IV with 1+1 la, 1+1 post macrosetae; urotergites V–VII with 1+1 la, 3+3–4+4 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 11+11, urosternites II–VII with 9+9, urosternite VIII with 3+3 macrosetae. Male with a narrow area of glandular g 1 setae; subtrapezoidal appendages with an area of glandular a 2 setae; female with subcylindrical appendages with glandular a 1 setae. Spermatozoid fascicles 60–65 µm in diameter and 25–30 µm wide, wheels with 2½–3 turns of 650 µm long and 13 µm diameter filament. Remarks The study of one female from Puech Cave, Grands Causses, France (see Table S 2 in Supplementary Material) has allowed us to supplement the former description, including regarding the presence of only 1+1 lp metanotal macrosetae. Habitat and distribution Subterranean species, well-spread around the Grands Causses, where it is found in eight caves, living alongside the troglobite P. (P.) balsani., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 43, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Moniez R. 1894. Especes nouvelles de thysanoures trouvees dan la grotte de Dargilan. Revue biologique du Nord de la France 6: 81 - 86. Available from https: // biodiversitylibrary. org / page / 10877325 [accessed 2 Jan. 2020].","Vire M. A. 1897. Le Campodea staphylinus Westwood, et ses varietes cavernicoles (C. cookei Packard; C. dargilani Moniez; C. nivea Joseph; C. erebophila Amann). Bulletin du Museum national d'histoire naturelle 3: 89 - 95.","Denis J. R. 1930. Sur la faune francaise des Apterygotes, XIe note: Diploures avec tableau de determination des especes francaises. Bulletin de la Societe zoologique de France 55: 19 - 41.","Denis J. R. 1923. Notes sur les Apterygotes. Ordre: Diplura C. B. Annales de la Societe entomologique de France 92: 231 - 236.","Bareth C. 2006. Les Campodes de France (Diploures Campodeides). Memoires de Biospeologie, hors serie: 2 - 108."]}

Plusiocampa (Plusiocampa) grandii Silvestri, 1933 Plusiocampa grandii Silvestri, 1933b: 1, figs i–ii. Diagnosis (Silvestri 1933b; Condé 1954a) Troglomorphic species. Antennae with 33–38 antennomeres. Pronotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 lp, 1+1 mp macrosetae; metanotum with 1+1 ma, 1+1 la, 2+2 lp, 1+1 mp macrosetae; long, barbed notal macrosetae; abundant clothing setae. Three ventral tibial macrosetae; unequal claws (1.25) with large lateral crests, posterior claw with a backward overhang. Urotergites I–II with 1+1 post macrosetae; urotergite III with 1+1 la, 1+1 post macrosetae; urotergite IV with 1+1 la, 5+5 post macrosetae; urotergites V–VII with 2+2 la, 5+5 post macrosetae; urotergite VIII with 6+6 post macrosetae; urosternite I with 7+7, urosternites II–VII with 6+6, urosternite VIII with 2+2 macrosetae. Male unknown; female with subcylindrical appendages bearing glandular a 1 setae., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 51, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Silvestri F. 1933 a. Nuovi contributi alla conoscenza della fauna delle isole italiane dell'Egeo. Bolletino del Laboratorio di Zoologia generale e agraria in Portici 27: 61 - 111.","Silvestri F. 1933 b. Descrizione di una nuova specie cavernicola di Campodeidae (Thysanura Entothropha) del Trentino. Bollettino del Laboratorio di Entomologia agraria de Bologna 6: 1 - 4.","Conde B. 1954 a. Ordnung: Entotropha (Diplura). In: Franz H. (ed.) Der Nordost im Spiegel ihrer Landtierwelt: 644 - 649. Universitatsverlag Wagner, Innsbruck, Austria."]}

Plusiocampa (Plusiocampa) gadorensis Sendra, 2001 Plusiocampa gadorensis Sendra, 2001: 73, figs 1–4, tables i–ii. Diagnosis (Sendra 2001) Troglomorphic species. Antennae with 46–52 antennomeres; 15–18 complex olfactory chemoreceptors in cupuliform organ; small protruding frontal process with a few slightly tubercular setae. Pronotum with 1+1 ma, 2+2–3+3 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 (2+3) lp macrosetae; metanotum with 1+1 ma, 2+2 la, 2+2 (2+3) lp macrosetae; long, barbed notal macrosetae; short notal clothing setae. Two or three ventral tibial macrosetae; slightly unequal claws (1.15) with large lateral crests, posterior claw with no backward overhang. Urotergite II with 0+0–1+1 (0+1) post macrosetae; urotergite III with 0+0–1+1 la, 1+1–2+2 (0+1) post macrosetae; urotergite IV with 1+1 (1+2) la, 3+3– 4+4 (4+5) post macrosetae; urotergite V with 1+1 (1+2) la, 4+4–5+5 (4+5) post macrosetae; urotergite VI with 1+1 (1+2) la, 5+5 (4+5) post macrosetae; urotergite VII with 1+1 la, 5+5 (4+5) post macrosetae; urotergite VIII with 6+6 (6+5) post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7–8+8 (8+9), urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male with an area of glandular g 1 setae, subcylindrical appendages with glandular a 1 setae; female and male appendages identical. Habitat and distribution Subterranean species endemic to the isolated karst of Sierra de Gádor, in the Peni-Baetic System, southeastern Iberian Peninsula. It is known from seven natural caves and one mine., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 50-51, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Sendra A. 2001. Dipluros campodeidos (Diplura: Campodeidae) de las grutas almerienses (Almeria, Espana). Zoologica Baetica 12: 71 - 82."]}

Plusiocampa (Plusiocampa) sardiniana Condé, 1981 Plusiocampa sardiniana Condé, 1981b: 593, fig. 1b. Plusiocampa provincialis ? – Condé 1953: 36. Diagnosis (Condé 1953 b, 1981 b, 1983) Troglomorphic species. Antennae with 33–38 antennomeres; protruding frontal process with tubercular setae. Pronotum with 1+1 ma, 4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp, 1+1 mp macrosetae; metanotum with 1+1 ma, 2+2 la, 2+2 lp, 1+1 mp macrosetae. Two ventral tibial macrosetae; slightly unequal claws (1.15) with large lateral crests, posterior claw with a backward overhang. Urotergites I–III with 2+2 post macrosetae; urotergite IV with 1+1 (0+1) la, 5+5 post macrosetae; urotergite V with 1+1 (1+2) la, 5+5 post macrosetae; urotergite VI with 2+2 (1+2) la, 5+5 post macrosetae; urotergite VII with 2+2–3+3 la, 5+5 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; thick clothing setae on abdominal segments IX–X; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male without glandular g 1 setae or glandular a 2 setae, with the exception of 1+ 1 g 1 and 2–3 a 2 in one specimen; female with subcylindrical appendages with glandular a 1 setae. Habitat and distribution Subterranean species found only in the San Giovanni Caves, Sardinia., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 79, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1981 a. Description d'un Campodeide enigmatique de Sardaigne (Insecta: Diplura). Revue suisse de Zoologie 88 (1): 95 - 99. Available from https: // biodiversitylibrary. org / page / 41323335 [accessed 13 Nov. 2019].","Conde B. 1981 b. Un nouveau Plusiocampa troglobie du massif de la Sainte-Baume (Diploures Campodeides). Revue suisse de Zoologie 88 (3): 589 - 594. Available from https: // biodiversitylibrary. org / page / 41337647 [accessed 13 Nov. 2019].","Conde B. 1953 a. Diagnoses de nouvelles Campodeides Corses. Bulletin du Museum d'histoire naturelle, 2 ° serie 25: 562 - 567.","Conde B. 1953 b. Campodeides de la grotte de San Giovanni (Sardaigne). Notes biospeologiques 8: 33 - 37.","Conde B. 1983. Le genre Plusiocampa dans les grottes sardes (Diploures Campodeides). Revue suisse de Zoologie 90 (4): 899 - 905. Available from https: // biodiversitylibrary. org / page / 41311753 [accessed 13 Nov. 2019]."]}

Plusiocampa (Plusiocampa) dallai Bareth & Condé, 1984 Plusiocampa dallai Bareth & Condé, 1984: 138, fig. 3. Diagnosis (Bareth & Condé 1984) Troglomorphic species. Antennae with 32 antennomeres; 8 complex olfactory chemoreceptors with an atypical multi-sheet shape in cupuliform organ; protruding frontal process with tubercular setae. Pronotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 1+1 la, 2+2 lp macrosetae; long, barbed notal macrosetae; thin, barbed notal clothing setae. Three ventral tibial macrosetae; unequal claws (1.25) with large lateral crests, posterior claw with a backward overhang. Urotergites I–III with 1+1 post macrosetae; urotergite IV with 1+1 la, 5+5 post macrosetae; urotergites V–VII with 2+2 la, 5+5 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male unknown, female with subcylindrical appendages with glandular a 1 setae. Habitat and distribution Subterranean species only known from Grotta di Montenero, in the Gargano Promontory part of the Apulia Carbonate Platform, Italian Peninsula., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 42-43, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Bareth C. & Conde B. 1984. Nouveaux Plusiocampa cavernicoles d'Italie continentale (Diplura Campodeidae). Bollettino della Societa entomologica italiana 116 (8 - 10): 132 - 147."]}

Patrizicampa sardoa Condé, 1962 Figs 1–2 Patrizicampa sardoa Condé, 1962: 199, figs a–b. Diagnosis (Condé 1962) Troglomorphic species. Cupuliform organ with 10–14 complex olfactory chemoreceptors. Protruding frontal process with non-tubercular setae. Pronotum with 1+1 ma, 4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 3+3 la, 2+2 lp macrosetae; long, barbed notal macrosetae; long clothing setae covered with thin barbs. Subequal elbow-like claws (1.1) with large lateral crests, posterior claw with a backward overhang. Urotergite III with 1+1 la, 0+0–1+1 post macrosetae; urotergite IV with 2+2 la, 1+1–3+3 post macrosetae; urotergite V with 2+2–3+3 la, 2+2–4+4 post macrosetae; urotergites VI–VII with 3+3 la, 4+4 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8–8+9 post macrosetae. Urosternite I with 15+15– 16+16, urosternites II–VII with 7+7–8+8, urosternite VIII with 3+3 macrosetae. Male without glandular g 1 setae, slightly subtrapezoidal appendages with glandular a 1 and a 2 setae; females with subcylindrical appendages and an apical area with glandular a 1 setae. Remarks (Figs 1–2) The study of many specimens collected in 6 new caves (see Table S 2 in Supplementary Material) allowed us to complete Condé’s diagnosis, as well as to supplement it with some minor taxonomic details. Medial antennomeres with 10–12 gouge sensilla 40–50 µm long. End of metathoracic leg reaches end of abdomen; calcars covered with barbs; subapical setae with thin barbs almost from base up to twothirds of length. Stylus with an apical seta with a few thin barbs and a large, thick basal tooth, medial and subapical setae covered with thin barbs. Male appendages with up to 60 glandular a 2 setae and 20 a 1 glandular setae (Fig. 1); female appendages with up to 18 a 1 glandular setae. Spermatozoid fascicle wheels with 2 turns of 140–160 µm long and up to 4 µm wide filament (Fig. 2). Habitat and distribution Subterranean species found in eight caves in eastern Sardinia (Fig. 210)., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 12-13, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1962. Campodeides des grottes de Sardaigne (Insecta Diplura). Atti VII Congresso nazionale di Speleologia, Memoria III: 199 - 202.","Conde B. 1956 a. Campodeides cavernicoles de Grece. Notes biospeologiques 11: 7 - 12."]}

Plusiocampa (Plusiocampa) suspiciosa Condé & Mathieu, 1957 Plusiocampa suspiciosa Condé & Mathieu, 1957: 467, fig. xi, 45–52. Diagnosis (Condé & Mathieu 1957) Non-troglomorphic species. Antennae with 20–21 antennomeres; four olfactory chemoreceptors in cupuliform organ; non-protruding frontal process with non-tubercular setae. Pronotum with 1+1 ma, 3+3–2+1 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp, 1+1 mp macrosetae; metanotum with 1+1 ma, 2+2 la, 2+2 lp, 1+1 mp macrosetae; long, barbed notal macrosetae; sparse smooth notal clothing setae. Two or three ventral tibial macrosetae; subequal small claws with small lateral crests, posterior claw with no backward overhang. Urotergites I–II with 2+2 post macrosetae; urotergite III with 4+4 post macrosetae; urotergites IV with 2+2 la, 5+5 post macrosetae; urotergites V–VII with 3+3 la, 5+5 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male unknown; female with subcylindrical to conical appendages with glandular a 1 setae. Habitat and distribution The locatity of this species is still not clear, as a label confusion may have occurred between two localities, one endogean habitat in France (Tarn-et-Garonne, La Capelle-Livran, Caylus) and a small cave in Sicily (Scaletta Zanclea, south of Messina).

Plusiocampa (Plusiocampa) festae Silvestri, 1932 Plusiocampa festae Silvestri, 1932: 75, figs x–xi. Plusiocampa festae coi Silvestri, 1932: 78. Diagnosis (Silvestri 1933a; Condé 1984a) Slightly troglomorphic species. Antennae with 25 and 35 antennomeres. Non-protruding frontal process with non-tubercular setae. Pronotum with 1+1 ma, 3+3 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp, 1+1 mp macrosetae; metanotum with 1+1 ma, 2+2 lp, 1+1 mp macrosetae; long, barbed notal macrosetae; abundant notal clothing setae. Two (rarely three) ventral tibial macrosetae; subequal claws (1.1) with large lateral crests; posterior claw with no backward overhang. Urotergites I–II with 1+1 post macrosetae; urotergite III with 1+0 la, 1+1 post macrosetae; urotergite IV with 1+1 la, 2+2– 3+3 post macrosetae; urotergites V–VII with 1+1 la, 5+5 post macrosetae; urotergite VIII with 7+7 post macrosetae; abdominal segment IX with 9+9 post macrosetae; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male with an area of glandular g 1 setae and enlarged appendages with glandular a 1 setae. Remarks The studied material (see Table S 2 in Supplementary Material) from MSS habitats was collected from inland Greece. Three specimens from Évia show variability in the number of post macrosetae on the third urotergite; they have 2+1, 2+2 and 3+3 post macrosetae, showing the dubious value of the subspecies P. (P.) festae coi. In addition, a juvenile from Montes Orno, Crete, has also been studied and it matches in all chaetotaxic characters with P. (P.) festae festae. In conclusion, we suggest that P. (P.) festae coi should be synonymized with the nominate subspecies. Habitat and distribution Soil-dwelling species previously found on Karpathos and Leros, islands of the Dodecanese Archipelago; also collected in endogean environments at two sites on Crete and at a locality near Izmir, Anatolia, Turkey. It seems to inhabit MSS, where it has also been found at 14 localities in continental Greece., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 49-50, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Silvestri F. 1932. Campodeidae (Thysanura) de Espana (primera parte). Eos 8: 115 - 164.","Silvestri F. 1933 a. Nuovi contributi alla conoscenza della fauna delle isole italiane dell'Egeo. Bolletino del Laboratorio di Zoologia generale e agraria in Portici 27: 61 - 111.","Conde B. 1984 a. Diploures Campodeides (Insectes) de Grece (1 ere note). Revue suisse de Zoologie 91 (1): 173 - 201. Available from https: // biodiversitylibrary. org / page / 41303604 [accessed 13 Nov. 2019]."]}

Plusiocampa (Plusiocampa) lindbergi Condé, 1956 Fig 119 Plusiocampa lindbergi Condé, 1956a: 7, figs 1–2. Diagnosis (Condé 1956a) Slightly troglomorphic species. Antennae with 34–37 antennomeres; five olfactory chemoreceptors in cupuliform organ of a possibly regenerated antenna; non-protruding frontal process with nontubercular setae. Pronotum with 1+1 ma, 4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 2+2 lp macrosetae; long, large, barbed notal macrosetae; smooth clothing setae. Two ventral tibial macrosetae; subequal claws (1.05–1.1) with middle-sized lateral crests, posterior claw with no backward overhang. Urotergites I–III with 1+1 post macrosetae; urotergite IV with 1+1 la, 2+2 post macrosetae; urotergites V–VII with 1+1 la, 4+4 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male with glandular g 1 setae, enlarged subcylindrical appendages with glandular a 1 setae; female with slender, subcylindrical appendages. Remarks The study of one male adult of 4.6 mm body length collected from the MSS habitat in Xànthi, Gérakas, Greece (see Table S 2 in Supplementary Material) allows the addition of information on the urosternites of the male. It has a narrow area of 96 glandular g 1 setae, arranged in 2–3 rows; enlarged and subcylindrical appendages with a distal area of up to 60 glandular a 1 setae (Fig. 119). Habitat and distribution Subterranean species known from Saint-Jean-le-Prodrome Cave (Spilaio Pelade) and from the MSS habitat near Xánthi, both on the Balkan Peninsula, in the Macedonian and Greek regions., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 70-71, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1956 a. Campodeides cavernicoles de Grece. Notes biospeologiques 11: 7 - 12."]}

Plusiocampa (Dydimocampa) euxina Condé, 1996 Plusiocampa euxina Condé, 1996: 102, figs 1–6. Diagnosis (Condé 1996) Troglomorphic species. Antennae with 44–45 antennomeres; eight to ten complex olfactory chemoreceptors in cupuliform organ; slightly protruding frontal process with a few anterior tubercular setae. Pronotum with 1+1 ma, 1+1 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 2+2 la, 2+2 lp macrosetae; metanotum with 1+1 ma, 1+2 la, 2+2 lp macrosetae; long, barbed notal macrosetae; sparse short, smooth notal clothing setae. Three ventral tibial macrosetae; unequal claws (1.25) with large lateral crests, posterior claw with a small backward overhang; short lateral processes. Urotergites I–III with 1+1 post macrosetae; urotergite IV with 1+1 la, 4+4 post macrosetae; urotergite V with 2+2 la, 4+4 post macrosetae; urotergites VI–VII with 2+2 la, 5+4 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 9+9 post macrosetae; urosternite I with 8+7, urosternites II–VII with 6+6, urosternite VIII with 2+2 macrosetae. Male without glandular g 1 setae, enlarged subcylindrical appendages with a large area of glandular a 1 setae; adult female unknown. Habitat and distribution Subterranean species inhabiting the Movile Cave, Southern Dubrudja, Romania., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 21, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1996. Diploures Campodeides de la Pestera de la Movile (Movile Cave), Dobroudja meridionales (Roumanie). Revue suisse de Zoologie 103 (1): 101 - 114. Available from https: // biodiversitylibrary. org / page / 41267657 [accessed 13 Nov. 2019]."]}

Plusiocampa (Plusiocampa) bonadonai bonadonai Condé, 1948 Plusiocampa bonadonai Condé, 1948c: 50, figs 2–3. Diagnosis (Condé 1948c, 1949, 1950; Condé & Poivre 1982; Bareth 2006) Troglomorphic species. Antennae with 31–37 antennomeres; 7–8 complex olfactory chemoreceptors in cupuliform organ. Pronotum with 1+1 ma, 4+4 la, 2+2 lp macrosetae; mesonotum with 1+1 ma, 3+3 la, 2+2 lp, 1+1 mp macrosetae; metanotum with 1+1 ma, 1+1 la, 2+2 lp, 1+1 mp macrosetae; long, barbed notal macrosetae; sparse notal clothing seate with a few distal barbs. Two or three ventral tibial macrosetae; subequal claws (1.05) with large lateral crests, posterior claw with a small backward overhang. Urotergites I–III with 1+1 post macrosetae; urotergites IV–VII with 1+1 la, 5+5 post macrosetae; urotergite VIII with 6+6 post macrosetae; abdominal segment IX with 8+8 post macrosetae; urosternite I with 7+7, urosternites II–VII with 5+5, urosternite VIII with 2+2 macrosetae. Male with abundant glandular g 1 setae, subcylindrical appendages like female appendages, with glandular a 1 setae. Habitat and distribution Subterranean species found in 11 caves and at one edaphic site in the Île Saint-Honorat caves around the Western Alps, in the mountain reliefs near the Mediterranean coast in the Provence Alps, France., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 32, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1948 a. Les Campodeides cavernicoles du Jura. Bulletin de l'Association speleologique de l'Est 1 (1): 1 - 6.","Conde B. 1948 c. Addition a la faune des campodeides cavernicoles de France. Bulletin de la Societe des Sciences de Nancy, new series 7 (2): 45 - 54.","Conde B. 1949. Plusiocampa provincialis n. sp. cavernicole de Provence (Diploures Campodeides). Bulletin de la Societe linneenne de Lyon (Oct. 1949): 166 - 169.","Conde B. 1950. Description d'un Campodeide cavernicole de Lombardie. Doriana (Annali del Museo Civico di Storia Naturale \" G. Doria \") 1 (3): 1 - 4.","Conde B. & Poivre C. 1982. Plusiocampa bonadonai Conde au Valais (Insecta, Diplura). Revue suisse de Zoologie 89 (1): 167 - 176. Available from https: // biodiversitylibrary. org / page / 41324017 [accessed 13 Nov. 2019].","Bareth C. 2006. Les Campodes de France (Diploures Campodeides). Memoires de Biospeologie, hors serie: 2 - 108."]}

Plusiocampa (Plusiocampa) hoffmanni Sendra & Paragamian sp. nov. urn:lsid:zoobank.org:act: 7C4D40A0-3013-48A3-B3DF-529E78F99DDC Figs 59–94, 204; Tables 6–8 Etymology This species is named in honour of Luc Hoffmann (1923–2016), an extraordinary man, a naturalist and philanthropist, who helped create many international conservation organisations and supported conservation research and actions in the Mediterranean and elsewhere. Material examined Holotype GREECE • ♀; Crete, Marathos nomos Iraklion, Spilaio Doxa (n. 1065); 490 m a.s.l.; 11 May 2005; Fulvio Gasparo leg.; MZB. Paratypes GREECE – Crete • 2 ♂♂, 2 ♀♀; same collection data as for holotype but 2 Jun. 2002; Coll. AS • 1 ♀; Zoniana nomos Rethymon, Spilaio Sfento Trypa (n. 701); 630 m a.s.l.; 11 Apr. 2003; Kaloust Paragamian leg.; Coll. AS • 6 ♀♀; Krousonas, Malevizi, Krousaniotiko Livadi, Varathro Stou Bokon ton Poro; 1Jul. 1990; Kaloust Paragamian leg.; Coll. AS • 1 ♀; Mylopotamos, Livadia, Varathro Sipouli; 24 Mar. 1990; Kaloust Paragamian leg.; Coll. AS • 1 ♂, 2 ♀♀; Malevizi, Gonies, Varathro Psistraki; 20 Nov. 2005; Kaloust Paragamian and S. Paragamian leg.; Coll. AS. Description BODY. Body length: 4.8 mm (juvenile), 4.7 and 5.4 mm (males from Spilio Doxa), 5.9 and 6.4 mm (females from Spilio Doxa), and 7.4 mm (male from Sfentoni Trypa). Epicuticle smooth under optical microscope, but reticulate at high magnification (Figs 68–69); body with thin, long clothing, smooth or with a few thin distal barbs on dorsal side and well barbed on ventral side, legs and urosternal appendages. HEAD. Antennae with 44–53 antennomeres; central antennomeres 2.5–3× as long as wide, apical antennomere 2.5× as long as wide (Fig. 61). Large cupuliform organ occupying ¼ of total length of apical antennomere, with 8–10 complex olfactory chemoreceptors (Figs 59–60). Small coniform sensillum of third antennomere located in ventral position between c and d macrosetae. Thin and long gouge sensilla, 38–50 µm long, in a single distal whorl of 10–14 sensilla on each medial and distal antennomere (Figs 62–64). Protruding frontal process with tubercular setae and macrosetae with thin barbs. Three macrosetae along line of insertion of antennomere and x setae relatively long, covered from distal third to fourth (Figs 65–66). THORAX. Thoracic macrosetal distribution (Fig. 67): pronotum with 1+1 ma, 4+4 la 1,2,3,4 , 2+2 lp 2,3 ; mesonotum with 1+1 ma, 3+3 la 1,2,3 , 2+2 lp 2,3 , 1+1 mp; metanotum with 1+1 ma, 0+0, 0+1, 1+1, 1+2 la or 0+1, 1+1 sla, 2+2 lp 2,3 , 1+1 mp. All medial posterior macrosetae in a forward position. All long, thin notal macrosetae with very thin barbs along distal third or half; marginal setae longer than clothing setae, covered by a few thin distal barbs. Legs elongated, metathoracic legs overpassing end of abdomen at beginning of tarsus. Tibia longest part of leg, 1.11–1.30× as long as tarsus and 1.21–1.37× as long as femur (Table 7). Ratios between lengths of tibia, tarsus and femur seem more or less fixed (Table 7). Femora II–III with one long, thin dorsal macroseta and one shorter ventral macroseta. Tibiae I–III with two or three ventral macrosetae surrounded by thin, short barbs. Calcars surrounded from base almost to tip by thin, short barbs. Metatarsus with a pseudo-articulation on distal third. Typical double rows of thick, completely barbed setae along ventral side of tarsus. Dorsal and lateral subapical tarsal setae completely covered by thin barbs nearly from base almost to tip. Subequal to slightly unequal claws (posterior claw 1.05–1.15× as long as anterior one), with large lateral crests well developed on posterior claw prolongated backwards. Ventral side of claws with longitudinal ridges separated by a surface of micro-hemispherical structures. Pretarsal processes setiform, with abundant, very short proximal barbs (Figs 71–82). ABDOMEN. Distribution of abdominal macrosetae on tergites: 1+1 post 1 on I–II; 1+1–0+0 la, 1+1 post 1 on III; 1+1 la, 3+3–4+4 post 1,2,4,5 on IV; 1+1 la, 5+5 post 1–5 on V–VII, 7+7 post on VIII; 8+8–9+9 post on abdominal segment IX. Urosternite I with 7+7–9+9, urosternites II–VII with 5+5–7+7, urosternite VIII with 2+2 macrosetae; all urosternal macrosetae robust, large, covered by long barbs along distal half to four-fifths (Figs 87–88). Stylus with subapical setae longer and more barbated, in fact completely surrounded by short, abundant barbs, more numerous on apical and ventromedial setae (Figs 92–93); large eversible vesicles (Figs 92, 94). Two complete cerci, 2.37 and 1.71× as long as body; this ratio higher in an incomplete 14.50 mm long cercus of 5.4 mm long male (Table 8); up to 11 articles, covered with numerous disorganized, long, thin macrosetae with thin barbs along distal half, in addition to a distal whorl of longer, thicker macrosetae on each article plus an apical whorl of barbed, thin, short clothing setae (Fig. 70). SECONDARY SEX CHARACTERS. Female urosternite I with subcylindrical appendages, each bearing up to 18 glandular a 1 setae in a distal area (Figs 89–91). Male urosternite I (Figs 83–86) with a posterior area of up to 160 glandular g 1 setae in larger male from Sfedoni Trypa (absent in smaller male); with thick, large, subcylindrical appendages, much thicker than female appendages, each bearing up to 100 glandular a 1 setae in a large distal area in two largest adults. Spermatozoid fascicles with a thick spiral filament. Phyletic affinities, habitat and distribution The presence of medial posterior macrosetae on the mesonotum and metanotum relates P. (P.) hoffmanni sp. nov. to almost half of the species in the subgenus Plusiocampa, all living in the European region, including most of the Mediterranean islands. Among them, P (P) festae is probably the closest related species due to the distribution pattern of their notal macrosetae; in spite of the fact that both species live in Crete (Condé 1984a; Silvestri 1933a), they occupy different habitats, P. (P.) hoffmanni sp. nov. in deep subterranean habitats, P. (P.) festae in soil and subterranean superficial habitats. In addition to this ecological difference, there are several important features that allow them to be separated. Plusiocampa (P.) hoffmanni sp. nov. has troglomorphic traits, with longer appendages than P. (P.) festae and more antennomeres and cercal articles, with more complex and more numerous olfactory chemoreceptors within the cupuliform organ. In P. (P.) hoffmanni sp. nov., the claws are slightly unequal to subequal, but with large lateral crests and a backward prolongation of the posterior claws. Additionally, P. (P.) hoffmanni sp. nov. shows a protruding frontal process covered by tuberculated setae, 8+8 macrosetae on the first urosternite and 6+6 on the second to seventh urosternites. Some of these features also separate P. (P.) hoffmanni sp. nov. from P. (P.) rybaki, a poorly described troglomorphic species from the southern Balkan Peninsula (Condé 1956). Plusiocampa (P.) hoffmanni sp. nov. has been collected from the deep zone of 14 caves in the numerous karst areas of Crete (Fig. 204).

Plusiocampa (Plusiocampa) bonadonai lanzai Condé, 1961 Plusiocampa bonadonai lanzai Condé, 1961: 399. Diagnosis differential against the type form (Condé 1961; Condé & Poivre 1982) Troglomorphic subspecies.Antennae with 27–33 antennomeres. Urotergite III with 2+2 post macrosetae. No adult males have been observed, and the young males from Valais do not have glandular g 1 setae, without glandular a 2 setae. Habitat and distribution Subterranean subspecies found in a cave of Valais (Switzerland) as well as in two caves in Brescia and Toscana (Italy), around the Western Alps., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on page 32, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1961. Stations et variation de Plusiocampa magdalenae Conde et de P. bonadonai Conde (Diploures Campodeides). Annales de Speleologie 16 (4): 387 - 399.","Conde B. & Poivre C. 1982. Plusiocampa bonadonai Conde au Valais (Insecta, Diplura). Revue suisse de Zoologie 89 (1): 167 - 176. Available from https: // biodiversitylibrary. org / page / 41324017 [accessed 13 Nov. 2019]."]}

Paratachycampa hispanica Bareth & Condé, 1981 Figs 196–201, 206 Paratachycampa hispanica Bareth & Condé, 1981: 776, figs 1a–c, 2a–c, 3a–c, 4a–c, 5a, c, e. Diagnosis (Bareth & Condé 1981; Sendra 1988; Sendra & Escolà 2004) Troglomorphic species. Epicuticle smooth. Antennae with 34–40 elongated antennomeres; 12–16 complex olfactory chemoreceptors in cupuliform organ. Protruding frontal process with abundant tubercular setae. Pronotum with 1+1 ma, 2+2 la, 1+1 lp macrosetae; mesonotum with 1+1 ma, 1+1 la macrosetae; metanotum with 1+1 ma, 1+1 la macrosetae; long, thin notal macrosetae with a very few barbs; long, thin smooth clothing setae. Subequal, elbow-like claws with large lateral crests, posterior claw with a backward overhang; laminar, barbed ventral pretarsal processes. Urotergites III–VII with 1+1 la macrosetae; urotergite VIII with 4+4 post macrosetae; abdominal segment IX with 7+7 post macrosetae; urosternite I with 22+22, urosternites II–VII with 7+7, urosternite VIII with 2+2 macrosetae. Male without glandular g 1 setae; overdeveloped appendages with a large area of glandular a 2 setae and an apical area of glandular a 1 setae; subcylindrical female appendages with glandular a 1 and a 2 setae. Remarks The studied material (see Table S 2 in Supplementary Material) allowed us to add taxonomic details revealed by SEM microscopy. Gouge sensilla abundant, as long as 28 µm, with thick transversal stripes with alternating rows of pores between them (Figs 196–197). Subequal claws with well-developed lateral crests (Figs 198–199). Lateral processes of pretarsus with S-shape, its ventral side covered with long, hooked barbs (Figs 200–201). Habitat and distribution Subterranean species found in nine caves throughout the coastal karst reliefs in the east of the Iberian Peninsula (Fig. 206).

Plusiocampa (Plusiocampa) chiosensis Sendra & Gasparo sp. nov. urn:lsid:zoobank.org:act: B3F3B66C-8C7C-48BF-A841-EAD3DF711C47 Figs 44–46; Tables 3–4 Etymology The name ʻchiosensisʼ is proposed based on the island of Chios, where it occurs in its deep subterranean habitat. Material examined Holotype GREECE • ♀; Chios Island, Sykias; 10 Jun. 2006; Fulvio Gasparo leg.; MZB 2019-1030. Paratypes GREECE • 2 ♂♂; same collection data as for holotype; Coll. AS. Description BODY. Body length 4.4 and 4.9 mm (males, paratypes) and 7.8 mm (female, holotype). Epicuticle smooth under optical microscope; body with sparse thin, long clothing, covered or not by a few thin distal barbs. HEAD. Holotype with two intact antennae of 54 antennomeres, 14.8 mm long. Small coniform sensillum of third antennomere located in ventral position between c and d macrosetae. Central antennomeres 3.8× as long as wide, apical anntenomere 3.6× as long as wide. Small cupuliform organ occupying 1/9 of total length of apical antennomere, with 6–7 complex olfactory chemoreceptors. Thin and long gouge sensilla (44–47 µm long) in a single distal whorl of 10–13 sensilla on each medial and distal antennomere. Frontal process developed, with tubercular setae and macrosetae with thin barbs. Three macrosetae along line of insertion of antennae poorly differentiated, only anterior one with 1–2 thin apical barbs; x setae longer than these macrosetae, with a couple of thin apical barbs (Fig. 44). Suboval labial palps with latero-external sensillum larger than sensillum of third antennomere, with two guard setae, up to 12 setae on anterior border and up to 130 neuroglandular setae. THORAX. Distribution of thoracic macrosetae: pronotum with 1+1 ma, 4+4–3+3 la 1,2,3,4 , 2+2 lp 2,3 (additionally in the three type specimens, 1+1 extra ma near the sagittal plane); mesonotum with 1+1 ma, 1+1 (0+1) la 2 , 2+2 lp 2, 3 , 1+1 mp; metanotum with 1+1 ma, 1+1 lp 2 and 1+1, 0+1 or 0+0 mp. All notal macrosetae long, thin, covered by very thin barbs along distal third up to half; marginal setae longer than clothing setae, covered by thin barbs along distal half up to two thirds; spare clothing setae, with 14, 21 and 23 clothing setae on each heminotum of pronotum, mesonotum and metanotum, respectively (holotype). Legs elongated, metathoracic legs overpassing end of abdomen in middle of tibia; measurements as in Table 3. Femora II–III with one long dorsal macroseta and one longer ventral macroseta. Tibiae I–III with two ventral macrosetae surrounded all along by thin, short barbs. Calcars surrounded from base to tip by thin, short barbs. Dorsal and lateral subapical tarsal setae completely covered by thin barbs almost from base and almost to tip. Subequal claws (posterior claw 1.0–1.05× as long as anterior claw); posterior claw with a small backward overhang; lateral crests well developed, noticeably ridged on ventral side. Pretarsal lateral process setiform, with plenty of very short proximal barbs. ABDOMEN. Distribution of abdominal macrosetae on tergites: 1+1 post 1 on I–III; 1+1 la, 2+2 lp 1,2 on IV; 1+1 la, 5+5 post 1–5 on V–VII, 8+8 post on VIII; 11+11 (11+12) post on abdominal segment IX. All post urotergal macrosetae thick, long, covered by thin barbs along distal four-fifths; la urotergal macrosetae shorter than post macrosetae, covered by barbs along distal half. Urosternite I with 7+7, urosternites II– VII with 5+5, urosternite VIII with 2+2 macrosetae; all urosternal macrosetae robust and large, covered by long barbs along distal half up to four-fifths. Apical subapical and ventro-medial setae completely surrounded by short abundant barbs. CERCI. One complete cercus in holotype with eight articles in addition to basal article, 19.70 mm long (Table 4), 2.53 mm longer than body length; articles show whorls of long macrosetae covered by thin barbs along distal two-thirds, combined with whorls of smooth, thin setae shorter than macrosetae. SECONDARY SEX CHARACTERS. Female urosternite I (Fig. 45) with enlarged subcylindrical appendages, each bearing up to 16 glandular a 1 setae in a distal area. Male urosternite I (Fig. 46) without glandular g 1 setae, with moderated large subcylindrical appendages thicker than those of female, each bearing up to 33 glandular a 1 setae. Phyletic affinities, habitat and distribution Although P. (P.) chiosensis sp. nov. has medial posterior macrosetae on the mesonotum and metanotum (sometimes absent in one or two heminota), as do many previously described species of Plusiocampa s. str., a similar reduction in the number of lateral anterior and lateral posterior macrosetae on the mesonotum and metanotum is present in another troglomorphic species, P. (P.) glagra from continental Greece (Condé 1984a). Plusiocampa (P.) chiosensis sp. nov. also shares a couple of noticeable taxonomical features with P. (P.) glabra in their subequal claws and the number of urosternal macrosetae. Nevertheless, the presence of 1+1 lateral anterior macrosetae on the mesonotum, the more abundant macrosetae on all urotergites, including the eighth abdominal segment, and the absence of glandular g 1 setae in males in P. (P.) chiosensis sp. nov. distinguish it from P. (P.) glabra. Plusiocampa (P.) chiosensis sp. nov. is only known from one cave and occupies the deep subterranean spaces, showing highly troglomorphic features (elongation of appendages, antennomeres and articles, and the strengthening of sensorial equipment of the antennae).

Plusiocampa (Plusiocampa) bonneti condei Sendra & Escolà, 2004 Figs 30–41 Plusiocampa bonneti condei Sendra & Escolà, 2004: 892. Plusiocampa pouadensis bonneti – Condé 1951: 61. Differential diagnosis against the type form (Sendra 1985; Sendra & Escolà 2004) Antennae with 30–43 antennomeres; eight to nine complex olfactory chemoreceptors in cupuliform organ; protruding frontal process with slightly tubercular setae. Spermatozoid fascicle wheels with 2–2½ turns of 300–400 µm long and 7 µm diameter filament. Remarks (Figs 24–35) One male of P. (P.) bonneti bonneti from Cova Mora (San Juan de la Peña, Huesca, Spain) and six specimens of P. (P.) bonneti condei from Cova Toll (Moià, Barcelona, Spain) (see Table S 2 in Supplementary Material) were used for SEM to remark on some features that are difficult to observe under the optical microscope. In P. (P.) bonneti bonneti, the first urosternite shows very long subtrapezoidal appendages with a large area of glandular a 2 setae and a large area of glandular g 1 setae, with the surface covered by barbed clothing setae and well-barbed macrosetae (Figs 24–27). In P. (P.) bonneti condei, the epicuticle varies from completely smooth to gently reticulate (Fig. 35); antennomeres with a whorl of gouge sensilla, 16–18 µm long (Figs 31, 33). Olfactory chemoreceptors with a central column unfolded in several layers with a reticulate surface covered by pores (Fig. 30). Protruding frontal process with slightly tubercular macrosetae and setae (Figs 32, 34). Claws with large lateral crest with ventral longitudinal grooves separated by reticulate surfaces (Figs 36–39). Stylar setae totally covered by short, thin barbs (Fig. 40). Large eversible vesicles (Fig. 41). Habitat and distribution Subterranean subspecies found in 35 caves in the Eastern Pyrenees and the northern part of the Catalan Mediterranean Mountain System.

Plusiocampa (Plusiocampa) pouadensis fouresi Condé, 1955 Plusiocampa pouadensis fouresi Condé, 1955a: 27. Differential diagnosis against the type form (Condé 1955a; Bareth & Condé 1996) Urotergite III with 2+2 post macrosetae; urotergites V–VII with 2+2 la, 6+6 post macrosetae; urotergite VIII with 7+7 post macrosetae. Habitat and distribution Subterranean subspecies inhabiting two caves on the French side of the Eastern Pyrenees., Published as part of Sendra, Alberto, Antić, Dragan, Barranco, Pablo, Borko, Špela, Christian, Erhard, Delić, Teo, Fadrique, Floren, Faille, Arnaud, Galli, Loris, Gasparo, Fulvio, Georgiev, Dilian, Giachino, Pier Mauro, Lukić, Marko, Marcia, Paolo, Miculinić, Kazimir, Nicolosi, Giuseppe, Palero, Ferran, Paragamian, Kaloust, Pérez, Toni, Polak, Slavko, Prieto, Carlos E., Turbanov, Ilya, Vailati, Dante & Reboleira, Ana Sofia P. S., 2020, Flourishing in subterranean ecosystems: Euro-Mediterranean Plusiocampinae and tachycampoids (Diplura, Campodeidae), pp. 1-138 in European Journal of Taxonomy 591 on pages 73-74, DOI: 10.5852/ejt.2020.591, http://zenodo.org/record/3659823, {"references":["Conde B. 1955 a. Materiaux pour une monographie des Diploures Campodeides. Memoires du Museum national d'histoire naturelle, serie A, Zoologie 12: 1 - 202.","Bareth C. & Conde B. 1996. Le complexe de Plusiocampa pouadensis (Denis) dans les grottes de la region pyreneenne. Memoires de Biospeologie 23: 127 - 131."]}