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Genus Habrochiton Semenov Tjan-Shansky, 1907 Figs 1C–D, 10, 15, 26A, 27E Habrochiton Semenov-Tjan-Shansky, 1907: 179–181. Type species Habrochiton vernus Semenov-Tjan-Shansky, 1907, by original designation. Species included Habrochiton vernus Semenov-Tjan-Shansky, 1907, H. primaeveris Semenov-Tjan-Shansky, 1907. Diagnosis Body (Fig. 1C–D) brown, slender, elongate-oval, completely covered with moderately dense short (H. vernus) or long (H. primaeveris) longitudinally striated scales, sparse scale-like and simple setae (Fig. 10). Body length 7–12 mm. Eyes small circular, moderately convex in dorsal view; surface behind eyes concealed under anterior margin of pronotum. Pronotum weakly transverse (1.4 × as wide as long). Anterolateral angles not expressed. Prosternum without transverse triangular depression along anterior margin. Prosternal process short and narrow (length 1.7 × width), not raised above procoxae, not protruding. Prosternum very short, near 2 × as long as longitudinal diameter of one procoxa. Elytra elongate-oval (H. vernus) or triangular, almost wedge-shaped (H. primaeveris). Scutellar shield completely (H. vernus) or partially concealed (H. primaeveris) by elytral base; surface around scutellar shield without triangular depression. Transverse length of one metacoxa less than intercoxal process of abdominal ventrite 1 (Fig. 1D). Trochanters with very short dense setae. Meso- and metafemora curved outward along elytral lateral vertical side. Pro- and metatibiae weakly curved outward, protibiae (Figs 26A, 27E) widened at apex, outer margin of protibia projecting into short rounded process at apex with one (H. primaeveris) or several acutely angulate and closely located (H. vernus) short spines; outer margin armed with very short tubercles with long and thin spines and additionally with long and sparse setaesetae (Fig. 27E). Protibial terminal spurs strongly elongated, extending to base of protarsomere 5, mesotibial terminal spurs extending to base of mesotarsomere 3, metatibial terminal spurs extending to midlength of metatarsomere 1. Outer and inner protibial terminal spurs subequal in length, inner meso- and metatibial terminal spurs longer than outer one; spurs flattened, outer spur moderately narrowed to apex, acutely angulate at apex. Protarsi laterally not flattened, covered with short strong setae. Meso- and metatarsi flattened from sides, covered with long fine setae dorsally and shorter and stronger setae ventrally. Tarsal claws elongated, thin, weakly curved outward. Male genitalia Male genitalia identical to those in Apatopsis. Ovipositor (Fig. 15J–L) Very short, paraproct slightly longer than coxite. Paraproct with thin baculi, indistinct at apical third. Coxite lobe baculi short, more sclerotized in apical part, with additional sclerotized projection on ventral side; lobe II very narrow (Fig. 15L); lobe III slightly longer, conical, without additional projecting apical processes; lobe IV membranous, conical, rounded at apex, densely pubescent with long setae. Female genital ducts Destroyed during preparation. Distribution China (Taklamakan Desert)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 14-17, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184.","Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382.","Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102."]}

Habrobates vejisovi Kelejnikova, 1977 Material examined Paratypes TURKMENISTAN • 2 ♀♀; “Репетек, 23–24.IV.1974, КуЗнецов” [Repetek]; “ 22-IV-74, гол. барх.” [bare barchans dunes]; “Paratypus, ♀, Habrobates vejisovi Kelejn 19.II.1977 ”; 23–24 Apr. 1974; N.Ya. Kuznetsov leg.; ZIN • 1♀; same collection data as for preceding; HNHM. Additional material TURKMENISTAN • 1 ♂; Karakum, Repetek; 27 Jan. 1957; N.G. Skopin leg.; ZIN • 7 ♂♂, 6 ♀♀; same collection data as for preceding; 6 Mar. 1958; G.S. Medvedev leg.; ZIN • 1 ♂, 1 ♀; same collection data as for preceding; 12 Apr. 1972; Begov leg.; ZIN • 2 ♂♂, 3 ♀♀; same collection data as for preceding; 16 Apr. 1980; B.A. Korotyaev leg.; ZIN. Distribution Turkmenistan (Karakum Desert, east)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 23, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425

Platyope proctoleuca proctoleuca Fischer von Waldheim, 1820 Fig. 6A Material examined KAZAKHSTAN • 1♂; Zaysan valley, Aygy-kum; 10 Jun. 1968; N.G. Skopin leg.; ZIN. RUSSIA • 1 ♂, 4 ♀♀; “Sibir.”; ZIN • 2 ♂♂, 3 ♀♀; “Sibir. occ.”; ZIN. Distribution Russia (Altai: Kulunda Steppe). Kazakhstan (Zaysan Depression, Irtysh River and Black Irtysh River valleys). This species was listed only for Zaysan Depression in Kazakhstan in the Palaearctic Catalogue (Iwan et al. 2020), but Medvedev (1990) recorded P. proctoleuca proctoleuca also for Russian Altai and the Irtysh valley in Kazakhstan., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 45-46, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow.","Iwan D., Lobl I., Bouchard P., Bousquet Y., Kaminski M. J., Merkl O., Ando K. & Schawaller W. 2020. Family Tenebrionidae Latreille, 1802. In: Iwan D. & Lobl I. (eds) Catalogue of Palaearctic Coleoptera. Vol. 5: Tenebrionoidea: 104 - 476. Brill, Leiden. https: // doi. org / 10.1163 / 9789004434998","Medvedev G. S. 1990. Key to the darkling beetles of Mongolia. Proceedings of the Zoological Institute 220. Zoological Institute of the USSR, Leningrad. [In Russian.]"]}

Earophanta pilosissima (Reitter, 1895) Earophanta loudoni Semenov, 1903a: 11 (original description). Material examined Holotype of Earophanta loudoni TURKMENISTAN • ♂; goldish circle; “Уч- Аджи 6.III.01 бар. Г. Лоудон” [Uch-Adzhi – now Bagtyýarlyk in Mary Prov.]; “ Platyope Loudoni m, Typ un. (♂?) XI.02 A. Semenow det.”; “coll. Semenov-Tian-Shansky”; “Holotype”; 6 Mar. 1901; Baron G. Loudon leg.; ZIN. Additional material AFGHANISTAN • 4 specs; Maymana Prov., Andkhoy; alt. 350–400 m; 17 Mar. 1964; O. Jakeš leg.; ZIN. TURKMENISTAN • 1 spec.; Berkarar [Ravnina railway station]; 8 Mar. 1908; Baron G. Loudon leg.; ZIN • 11 specs; Repetek; 6–14 Mar. 1958; inter-barchan depression with black saxaul; G.S. Medvedev leg.; ZIN • 3 specs; Karakum channel, Nichki; 25 Apr. 1968; G.S. Medvedev leg.; ZIN • 3 specs; Repetek; 19 Feb. 1973; V.G. Kaplin leg.; ZIN • 1 ♂, 1 ♀; Karakum channel, Chaloy; 10 Apr. 1983; E.A. Khachikov leg.; CMN. Distribution Turkmenistan, Afghanistan. Notes Earophanta loudoni was described from a single specimen and was synonymised by Medvedev (2005). The specimens from Turkmenistan, which were standing in the collection of ZIN as E. loudoni, differ from the specimens from Afghanistan (determined by Z. Kaszab as E. pilosissima) by much more pubescent elytra and visibly smaller smooth tubercles on the elytra. A larger series of specimens is necessary to establish the status of these populations., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 55, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov A. P. 1903 a. Coleoptera novae faunae turanicae. Revue russe dEntomologie 3: 8 - 13.","Medvedev G. S. 2005. On connections of the sand desert faunas of Tenebrionidae (Coleoptera) of Middle Asia, Iran, and Afghanistan. In: Konstantinov A. S., Tishechkin A. K. & Penev L. (eds) Contribution to Systematics and Biology of Beetles. Papers Celebrating the 80 th Birthday of Igor Konstantinovich Lopatin: 299 - 314. Pensoft, Moscow."]}

Earophanta tomentosa (Semenov, 1893) Material examined Lectotype designated here according to Articles 61.1 and 74.1 of ICZN (1999) UZBEKISTAN • 1 ♂; “Kisil-Kum, mer. Sarybai-Bulysch, Glasunov, 1892” [Kyzylkum Desert]; “ Platyope tomentosa m. ♂ Typ. Aes. II.93 ”; “ Holotypus ”; 1892; D.K. Glasunov leg.; ZIN. Paralectotypes UZBEKISTAN • 1♀; “Trkst. mnt. NURATA, Temir-Kauk, Glasunov, 1892” [Turkestan mountains]; “ Platyope tomentosa m. ♀ Typ. II.93, A. Semenow det.”; “ Allotypus ”; 1892; D.K. Glasunov leg.; ZIN • 1♀: “ Kisil-Kum, mer. Kalma-Tai, Glasunov, 1892” [Kyzylkum Desert]; “ Platyope tomentosa m. ♀ Typ. Aes. II.93 ”; “Paratypus; 1892; D.K. Glasunov leg.; ZIN. Distribution Uzbekistan (Kyzyl Kum Desert)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 57, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264.","ICZN. 1999. International Code of Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. The International Trust for Zoological Nomenclature. London, UK. Available from http: // iczn. org [accessed 9 Mar. 2022]."]}

Earophanta collaris (Fischer von Waldheim, 1844) Fig. 7A Material examined KAZAKHSTAN • 2 ♂♂, 1 ♀; Kapchagay, Kokbastau; 27 Jun. 1952; P. Ler leg.; ZIN • 4 ♂♂, 2 ♀♀; near Ili; 22 Jun. 1950; N.G. Skopin leg.; ZIN • same collection data as for preceding; 6 Jul. 1954; ZIN • 1 ♂, 2 ♀♀; Ili River; 8 Jun. 1952; P. Ler leg.; ZIN • 1 ♀; Kapchagay; 15 Apr. 1962; G. Nikolaev leg.; ZIN. CHINA • 45 ♂♂, 70 ♀♀; Yining [Qulja]; 1879; A. Regel’ leg.; ZIN. Distribution South Eastern Kazakhstan, China (Xijiang)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 52-53, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425

Platyope mongolica Faldermann, 1835 Material examined CHINA • 1 ♂; Manchuria; 10–28 May 1905; A.V. Serebryan leg.; ZIN. MONGOLIA • 5 ♂♂, 10 ♀♀; Khovd [Kobdo]; 16 Jun. 1911; K.V. Yurganova leg.; ZIN • 2 ♂♂; near Orog Lake, Gobi Desert; 30 May 1926; M.A. Kozlov leg.; ZIN • 1 ♂; Böön Tsagaan Lake; 6 Jun. 1962; Tsendsuren leg.; ZIN • 15 ♂♂, 23 ♀♀; 55 km SE of Airag [Hara-Ayrag]; 12 Jun. 1971; I.V. Emel’yanov leg.; ZIN • 16 ♂♂, 13 ♀♀; 12 km NE of Mandakh; 15 Jun. 1971; I.M. Kerzhner leg.; ZIN • 15 ♂♂, 23 ♀♀; 60 km ESE of Manlay; 15 Jun. 1971; G.S. Medvedev leg.; ZIN • 30 ♂♂, 25 ♀♀; 25 km ESE of Manlay; 16 Jun. 1971; G.S. Medvedev leg.; ZIN • 3 ♂♂, 8 ♀♀; 60 km SSE of Manlay; 17 Jun. 1971; M.A. Kozlov leg.; ZIN • 9 ♂♂, 5 ♀♀; SW Tahilga-Ula; 15 Jun. 1973; G.S. Medvedev leg.; ZIN • 26 ♂♂, 38 ♀♀; 10 km SW of Huh-Mor’t; 21 Aug. 1971; K.V. Arnol’di leg.; ZIN • 8 ♂♂, 3 ♀♀; Bayankhongor, 60 km NNE of Orog Lake; 4 Jul. 1973; G.S. Medvedev leg.; ZIN. Distribution Russia (Tuva), Mongolia, China (Jilin, Inner Mongolia, Liaoning, Ningxia)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 48, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425

Subtribe Habrobatina Nabozhenko & S. Chigray subtrib. nov. Type genus Habrobates Semenov, 1903. Diagnosis Body entirely covered with dense scales. Pronotum and elytra without large tubercles or strong sculpture, or only with small sparse tubercles. Eyes rounded or weakly oval, dorsoventrally placed above genal level. Legs thin, slender, long, not fossorial, with long sparse setae. Ovipositor very short and weakly sclerotized, transverse or of subequal length and width; paraproct very short, often with reduced or very short baculi; proctiger strongly widened, with very wide weakly sclerotized baculi; lobes 4 of coxite membranous, rounded, with long, very dense hair brush. Spiculum ventrale very short, strongly widened and without common stem. Aedeagus with strongly sub-angularly curved apical piece. Etymology From the Ancient Greek ‘ἁβρός’ (‘habros’ = delicate or splendid) and ‘βατίς ’(‘bates’ is plural inflection of ‘batis’ = a plant, samphire; genitive = ‘batis’; stem = ‘bat’). Genera included Apatopsis Semenov, 1891, Habrobates Semenov, 1903, Habrochiton Semenov-Tjan-Shansky, 1907, Dietomorpha Kühnelt, 1957. Distribution and bionomics The representatives of the new subtribe are distributed in the deserts of Central and South Asia (Turkmenistan, Iran, Afghanistan, Pakistan, China): Karakum, Dasht-e Lut, Dasht-e Kewir, Registan, Taklamakan. Species of the mentioned genera inhabit non-fixed barchan sand dunes. Comparison The subtribe Habrobatina Nabozhenko & S. Chigray subtrib. nov. can be distinguished from the nominotypical one by the following key: Key to the subtribes of Pimeliini Latreille, 1802 1. Combination of characters: eyes located dorsoventrally, above genal level; body completely covered with scales; ovipositor very short and weakly sclerotized, with membranous rounded and densely pubescent apical lobes of coxite; spiculum ventrale very wide and short, without or with very short common stem. Apical piece of aedeagus strongly sub-angularly curved............................................. subtribe Habrobatina Nabozhenko & S. Chigray subtrib. nov. – Combination of characters: eyes located dorsoventrally above genal level and body without scales (Platyope genus group) or eyes located laterally on genal level (body can be without or rarely with scales); ovipositor long, with fossorial, bare, strongly sclerotized apical lobes of coxite. Apical piece of aedeagus weakly or moderately (not sub-angularly) curved....subtribe Pimeliina Latreille, 1802, Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 5-6, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382.","Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184.","Kuhnelt W. 1957. Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Die Tenebrioniden Irans. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch- Naturwissenschaftliche Klasse 166 (2): 65 - 102."]}

Earophanta serrata (Semenov, 1893) Figs 7C–D, 13E–F, 22, 23H, 24E, 25B, 26G, 28C Material examined Lectotype designated here TURKMENISTAN • 1 ♂; “ АШхабад, 12.IV.28, E. Шестоперов” [Ashgabat]; “ Earophanta serrata m. ♂ A. Semenov-Tian-Shansky det. IV.31 ”; “Paratypus”; 12 Apr. 1928; E.L. Shestoperov leg.; ZIN. Additional material TURKMENISTAN • 2 ♂♂, 2 ♀♀; “TransCaspi, Eylandt.”; ZIN • 6 ♂♂, 10 ♀♀; Kushka River; 4 May 1908; I. Ivanov leg.; ZIN • 3 ♂♂, 4 ♀♀; same collection data as for preceding; 22 Apr. 1957; G.S. Medvedev leg.; ZIN • 2 ♀♀; Nokhur; 22 Jul. 1926; Moritz leg.; ZIN • 3 ♂♂, 3 ♀♀; Ashgabat; 12 March 1937; Ya.P. Vlasov leg.; ZIN • 1 ♂, 1 ♀; same collection data as for preceding; 25 Mar. 1964; L. Freyberg leg.; ZIN • 3 ♀♀; near Ashgabat 20 km NO; 27 Mar. 1952; Il’ichev leg.; ZIN • 2 ♂♂, 4 ♀♀; 20 km NW of Ashgabat; 27 Mar. 1952; O.L. Kryzhanovsky leg.; ZIN • 2 ♀♀; Badhyz State Nature reserve; 11 Apr. 1952; Z. Yudina leg.; ZIN • 1 ♂, 1 ♀; May 1986; V.A. D’yakov leg.; CMN • 2 ♂♂, 1 ♀; Badhyz State Nature reserve; Kepele cordon; 11 Apr. 1990; A. Napolov leg.; CMN • 2 ♂♂; 40 km W of Yasga [Yaskhan]; 22 Apr. 1952; O.L. Kryzhanovsky leg.; ZIN • 13 ♂♂, 7 ♀♀; Badhyz, 8 km of Galaymor; 28 Mar. 1957; G.S. Medvedev leg.; ZIN • 6 ♂♂, 3 ♀♀; Badhyz, 12 km W of Galaymor; 21 Mar. 1957; G.S. Medvedev leg.; ZIN • 3 ♂♂, 2 ♀♀; same collection data as for preceding; 9 Apr. 1957; ZIN. Distribution Turkmenistan., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 52, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264."]}

Dietomorpha gonzalesi S. Chigray & Nabozhenko sp. nov. urn:lsid:zoobank.org:act: 391D6624-EE07-4DC5-AE48-811D8A915504 Fig. 3B–G Etymology The new species is named after David Gonzales (Saint-Laurent-du-Pape, France), who made a great contribution to the Tenebrionidae of the New and Old Worlds. Material examined Holotype PAKISTAN • ♀; Balochistan, Khuzdar; May 2009; Hafiz Akhtar leg.; V.2009; coll. David Gonzales; MNHP. Paratypes PAKISTAN • 1 spec.; Webb Ware Quetta, Balochistan [handwritten]; “ Dietomorpha sp. n. det. Kaszab ”; HNHM • 1 ♀; WEBB WARE/ QUETTA [Printed]; “2355 / 14” [Printed: Indian Museum accession number?]; “ D. pardella obliterata n. [handwriting of K.G. Blair] det. K.G. Blair [printed]”; Dietomorpha pardalis V. obliterata Blair in litt. 1973. N. Skopin [handwriting of Skopin]; “Brit. Mus. [printed] 197[printed]3- 38. [handwritten]”; “MANUSCRIPT NAME” [red frame and text, printed]; BMNH • 1 ♀; same labels, but “2352/ 14”; BMNH. Notes on paratypes (Maxwell Barclay, personal communication) Ernest Vredenburg (the donor of the specimens, Superintendant of the Geological Survey of India) and Captain Frank C. Webb Ware (the collector of the paratypes, at Quetta, ‘Political Assistant’ at Chagai and who was working on the Sistan-Quetta caravan route) went to Quetta together. It is likely that the beetles were collected at Quetta in 1899, given the Indian Museum registration in 1914, and eventually given the BMNH registration in 1973. Description Female Body oval, covered by creamy scales, with striped pattern of dark spots on pronotum and elytra; scales lanceolate, longitudinally striated. Anterior margin of epistoma widely emarginated. Epistoma and frons sparsely covered with erected, yellowish setae. Genae rounded. Eyes large, circular, convex in dorsal view; surface behind eyes covered by anterior margin of pronotum. Ratio of head width across eyes to interocular distanceon frons is 1.3. Antennae long, with ultimate antennomere extending elytral base. Ratio of length (width) of 2 nd –11 th antennomeres: 0.5(0.5), 2.4(0.5), 0.9(0.4), 0.8(0.5), 0.8(0.5), 0.8(0.5), 0.7(0.8), 0.7(0.8), 0.2(0.5), 0.3(0.4). Head covered with fine, sparse spinose granules (granule diameter 2–3 × less than intergranular spaces). Pronotum transverse (1.7 × as wide as long), widest before middle, where it 1.2 × as wide as head, with two dark, broad fields and pale spot in middle; lateral margins at base together with contiguous part of prothoracic hypomera with glabrous and smooth black spot. Ratio of width of pronotum at anterior margin to its maximum width and to width at base: 8: 8.5: 8. Anterior margin and base of pronotum widely weakly emarginated at middle. Lateral margins rounded, emarginated at base. Anterolateral angles not expressed, posterolateral angles obtuse, widely rounded. Disc of pronotum transversely convex, with two triangular depressions at base. Apical half and middle of pronotum covered with projecting spine-like spinose tubercles (mainly on sides of pronotal disc). Prosternal process narrow, elongated (3× as long as wide), not raised between procoxae, slightly protruding beyond procoxae. Elytra weakly elongate (1.2× as wide as long), 3.4 × as long and 1.8× as wide as pronotum, 2.1× as wide as head. Each elytron with five brown, longitudinal stripes on creamy background. Elytra without humeral ribs, with four longitudinal rows of spine-like tubercles: one row on each lateral vertical side, one humeral row and four discal rows. Apex of elytra with dense tubercles. Scutellar shield not concealed by base of elytra; surface around scutellar shield with triangular depression. Metepisterna, meso- and metaventrites with fine sparse spinose granules. Transverse length of one metacoxa subequal to intercoxal process of abdominal ventrite 1. Abdominal ventrites covered with fine, sparse granules (granule diameter 1.5–3 × as short as intergranular spaces). Trochanters with elongated, sparse, yellowish setae. Meso- and metafemora curved outward. Protibiae sharply widened at apex, outer margin with row of short, acutely angulate spines and several sparse, long setae; spines closely spaced at apex. Outer and inner margins with row of long, thin setae. Tibial terminal spurs flattened, straight, acutely angulate at apex (except for metatibial terminal spurs), widened to apex and rounded apically. Pro- and mesotibial terminal spurs moderately elongated, extending base of tarsomere 2, metatibial terminal spurs extending to midlength of metatarsomere 1. Meso- and metatibiae slightly curved outward, with similar spines and dorsal setae as on protibiae. All tarsi flattened from sides, protarsi only with spines, meso- and especially metatarsi with dense, long setae dorsally and spines ventrally. Tarsal claws large, thin, visibly curved outward. Length ratio for femur, tibia, and tarsus: 6.5: 4: 2 in fore leg, 7.5: 5.5: 3.5 in middle leg, 8.5:8: 6 in hind leg. Body length 12 mm, width 7.5 mm. Comparative diagnosis This new species differs from D. pardalis by the more elongated body, the slightly protruding prosternal process (strongly protruding beyond procoxae in D. pardalis), the striped pattern (spotted in D. pardalis), the longitudinal rows of spine-like setae on elytra (the tubercles located only in spots in D. pardalis), the absence of elytral humeral ribs and not concealed scutellar shield., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 27-29, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425

Mantichorula semenowi Reitter, 1889 Figs 4C–D, 11E–F, 12A–B, 19, 23E, 24C, 26D, 27F–G Material examined Lectotype designated here CHINA • 1 ♂; “Kan-ssu 1886 G. Patanin”; “Typus Mantichorula Semenovi Rtt. Coll. Reitter ”; “ Mantichorula Semenovi m. 1889”; 1886; G.N. Potanin leg.; HNHM. Paralectotypes CHINA • 1 spec.; “Kan-ssu 1886 G. Patanin”; “Cotypus Mantichorula Semenovi Rtt. Coll. Reitter ”; “ Mantichorula Semenovi m. 1889”; 1886; G.N. Potanin leg.; HNHM • 1 spec.; “Ordos 1884 Patanin 5.VIII ”; “Cotypus Mantichorula Semenovi Rtt. Coll. Reitter ”; “ Mantichorula m. semenovi m. 1888”; 5Aug. 1884; G.N. Potanin leg.; HNHM. Additional material CHINA – Inner Mongolia Province • 3 ♂♂, 3 ♀♀; Ordos; 1884; G.N. Potanin leg.; ZIN • 5 ♂♂, 8 ♀♀; Alashan, Tain-Ula; 16–19 Apr. 1908; M.A. Kozlov leg.; ZIN • 15 ♂♂, 9 ♀♀, Sogo-Nur; 9 May 1926; M.A. Kozlov leg.; ZIN. – Gansu Province • 1 ♂; Gansu; 1886; G.N. Potanin leg.; ZIN • 1 ♂; same collection data as for preceding; 3–6 Jul. 1908; M.A. Kozlov leg.; ZIN • 2 ♂♂, 5 ♀♀; Gan River; 25 Apr. 1990; Martin leg.; ZIN. MONGOLIA • 1 ♂; Tengri-elisu Desert; 3 Sep. 1901; M.A. Kozlov leg.; ZIN • 4 ♂♂, 6 ♀♀; Gojczo; 1–2 Apr. 1908; M.A. Kozlov leg.; ZIN • 4 ♂♂, 4 ♀♀; Dyn-yuan-in, Utzur-huduk; 20–22 Apr. 1908; M.A. Kozlov leg.; ZIN • 6 ♂♂, 4 ♀♀; Bain-huduk; 2–4 Apr. 1909; M.A. Kozlov leg.; ZIN • 4 ♂♂, 2 ♀♀; Hayry-huduk; 22–23 May 1909; M.A. Kozlov leg.; ZIN • 2 ♂♂, 1 ♀; Bordzon-Gobi 80 km SSE of Nomgona; 5–8 Aug. 1967; V.F. Zaytsev leg.; ZIN. Distribution Mongolia, China (Inner Mongolia, Xinjiang, Gansu, Ningxia, Shaanxi provinces)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 33, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710."]}

Habrobates gabrieli (Schuster, 1935) comb. nov. Figs 2, 26B Material examined Lectotype designated here IRAN • 1 spec. (sex unknown); “ Kawiria n. g. Gabrieli n. sp. Schuster”; “Pers Halwan” [Alwand or Halwan River]; “TYPUS”; “ gabrieli; Pfaundler”; NMB. Paralectotypes IRAN • 1 spec.; “ Pers. G. Kawir S. ” [Dasht-e Kavir Desert]; “4”; “COTYPUS”; NMB • 1 spec.; “ Kawiria gabrieli ”; “Pers Halwan” [Alwand or Halwan River]; “COTYPUS”; “Paratypus 1935 Kawiria Gabrieli Schuster ”; HNHM. Additional material IRAN • 2 specs; Yazd, Riq-e Dion, 32 km of Chah Malek 33°46ʹ N, 52°42ʹ E versus Chupunum 33°31ʹ N, 54°19ʹ E, alt. 800 m; 16 May 1975; W. and F.C. Rechinger leg.; HNHM • 1 spec.; Yazd, Sudrand, Kavir desert, between Tabas and Anarak; 15 May 1975; W. and F.C. Rechinger leg.; HNHM • 1 spec.; “ Persia Centr.”; 1948; N. Alexandrov leg; ZMMU. Distribution Iran (Dasht-e Kavir Desert).

Platyope granulata Fischer von Waldheim, 1820 Fig. 5I Material examined KAZAKHSTAN • 32 ♂♂, 25 ♀♀; Kumtobe sands; 19 Jun. 1910; G.G. Jacobson leg.; ZIN • 2 ♂♂, 4 ♀♀; Zaysan, Aygyrkum; 8 Jun. 1958; N.G. Skopin leg.; ZIN • 2 ♂♂, 4 ♀♀; same collection data as for preceding; 9 Jun. 1958; ZIN • 2 ♂♂, 4 ♀♀; same collection data as for preceding; 10 Jun. 1958; ZIN • 1 ♂; Karatal sands; 16 Jun. 1964; P.Yu. Gorbunov leg.; ZIN. RUSSIA • 2 ♂♂; “Sibir.”; ZIN. Distribution Eastern Kazakhstan (new record for the country), China (Xinjiang)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 45, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow."]}

Platyope leucogramma (Pallas, 1781) Figs 4E–F, 5 AC, 12C–F, 20, 23F, 24D, 26H, 28D Material examined KAZAKHSTAN • 6 ♂♂, 4 ♀♀; Bol’shiye Barsuki; 28 Apr. 1904; V.A. Dubyansky leg.; ZIN • 4 ♂♂, 3 ♀♀; same collection data as for preceding; 20 May 1906; L. Berg leg.; ZIN • 2 ♂♂, 1 ♀; near Temir; 1 May 1905; B.P. Uvarov leg.; ZIN • 1 ♀; Temir, Kumkuduk; 30 May 1908; D.N. Borodin leg.; ZIN • 1 ♀; same collection data as for preceding; B.P. Uvarov leg.; ZIN • 8 ♂♂, 6 ♀♀; near Shalkar [Chelkar], Bol’shiye Barsuki; N.V. Androsov leg.; ZIN • 8 ♂♂, 4 ♀♀; Petrovo, Embulatovka River; 2–3 June 1949; K.G. Romadina leg.; ZIN • 5 ♂♂, 2 ♀♀; Yanvarcevo, right bank of Ural River; 23 Jun. 1949; L.V. Arnol’di leg.; ZIN • 2 ♂♂, 1 ♀; sands of Atyrau [= Guryev]; 3 May 1951; L.V. Arnol’di leg.; ZIN • 1 ♂, 1 ♀; same collection data as for preceding; K.G. Romadina leg.; ZIN • 3 ♂♂, 6 ♀♀; Kzyl-Chachyl; 9 Jun. 1952; L.V. Arnol’di leg.; ZIN • 1 ♂, 2 ♀♀; Karaagach 60 km NW of Zhanaarka; 19 May 1962; L.V. Arnol’di leg.; ZIN • 4 ♂♂, 7 ♀♀; sands 12 km N of Kensu; 28 May 1962; L.V. Arnol’di leg.; ZIN • 4 ♂♂, 2 ♀♀; Zaysan; ZIN • 4 specs; Atyrau Region, Akkiztogay; 9 Jun. 1980, 20 June 1980; E.A. Khachikov leg.; CMN. RUSSIA • 8 ♂♂, 9 ♀♀; “Sibir. occ.” [Eastern Siberia]; ZIN • 2 ♂♂, 3 ♀♀; Volgograd Region, Sarepta; 13 Jun. 1891; A.K. Becker leg.; ZIN • 1 ♂, 3 ♀♀; same collection data as for preceding; G.L. Suvorov leg.; ZIN • 3 ♂♂, 2 ♀♀; Volgograd Region, Volzhsky; 30 Apr. 2016; M.V. Mokrousov leg.; ZIN • 2 ♀♀; Kalmykia, Artezian, 26 Apr.–2 May 1998; I.V. Shokhin leg. ZMSFU • 1 spec.; Kalmykia, Artezian, Kuma River bank; 18 May 2013; I.V. Shokhin leg.; CMN • 3 ♂♂, 2 ♀♀; Astrakhan Region, Selitrennoye; 2 Feb. 1910; Chernavin leg.; ZIN • 2 specs; Rostov Region, Nizhnekundryuchenskaya; 1 May 1997; CMN • 11 specs; Rostov Region, Tsymlansk sands near Nizhnegnutov; 7–10 May 1998; M.V. Nabozhenko, D.A. Dubovikoff, L.V. Markitan leg.; ZMSFU, CMN. Distribution South of the European part of Russia (Rostov, Stavropol, Volgograd, Astrakhan regions, Kalmykia, Chechnya, Dagestan,), North Kazakhstan from Volga River to Zaysan Lake., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 43-45, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264."]}

Apatopsis grombczewskii Semenov, 1891 Figs 1A, 8A–D, 14, 23A, 26F, 27A–B Apatopsis conradti Semenov, 1891: 370 (original description). Syn. resurr. Material examined Lectotype of Apatopsis grombczewskii designated here CHINA • 1 ♂; “M. Keрieй и Нieй, 8.III.90, Громбчевскiй” [Xinjiang: between Keriya and Niya]; 8 Mar. 1890; B.L. Grombchevsky leg.; “ Apatopsis Grombczewskii m. Typ. ♂ XI.02. A. Semenov det.”; ZIN. Paralectotype of Apatopsis grombczewskii CHINA • 1 ♀; same collection data as for lectotype; ZIN • 1 ♀; “М. Хотаном и Керiей (бл. последн.), Х.89, Пѣвцов” [Xinjiang: between Hotan and Keriya, near the latter]; Oct. 1889; M.V. Pevtsov leg.; ZIN. Lectotype of Apatopsis conradti designated here CHINA • 1 ♀; “M. Keрieй и Нieй, 8.III.90, Громбчевскiй.” [Xinjiang: between Keriya and Niya]; 8 March 1890; B.L. Grombchevsky leg.; ZIN. Additional material CHINA • 1 ♂; “Kara-Kosсh” [Karakax County], “Pudshia” [Puji]; 36°55′ N, 79°24′ E; alt. 2450 m; 21 Feb. 1890; S. Conradt leg.; ZIN • 3 ♂♂, 1 ♀; same collection data as for preceding; B.L. Grombchevsky leg.; ZIN • 1 ♂; between Keriya and Niya; 8 Mar. 1890; B.L. Grombchevsky leg.; ZIN • 15 ♂♂, 19 ♀♀; Yasumun; 8 Mar. 1890; B.L. Grombchevsky leg.; ZIN. Distribution China (Xijiang, Taklamakan Desert)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 14, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382."]}

Platyope ordossica Semenov-Tjan-Shansky, 1907 Fig. 6B Material examined Holotype CHINA • ♂; “Ordos 1884 G. Patanin” [Inner Mongolia]; “ Platyope ordossica A. Semenow det.”; “Coll. Semenov-Tian-Shansky”; “Holotype”; 1884; G. Patanin leg.; ZIN. Additional material CHINA • 4 ♂♂, 5 ♀♀; Alashan, 15 May 1908; M.A. Kozlov leg.; ZIN • 4 ♂♂, 6 ♀♀, Dyn-Kou; 28–30 May 1908; M.A. Kozlov leg.; ZIN • 9 ♂♂, 7 ♀♀; Alashan, Dyn-yuan-in; 17–22 Jun. 1908; M.A. Kozlov leg.; ZIN • 2 ♂♂, 1 ♀; Southern Alashan, Bain-huduk; 2–4 Apr. 1909; M.A. Kozlov leg.; ZIN • 10 ♂♂, 5 ♀♀; near Linzhou; 24 Apr. 1909; M.A. Kozlov leg.; ZIN. Distribution China (Gansu, Ningxia, Inner Mongolia)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 48, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184."]}

Habrochiton vernus Semenov Tjan-Shansky, 1907 Figs 1C–D, 10, 15, 26A, 27E Material examined Lectotype designated here CHINA • 1 ♂; “OaЗ. НiЯ, Вост. Турк., 3/ 3 II. 2/ 3.III.-90, Пѣвцов” [Xinjiang: Niya oasis, Eastern Turkestan, 38°01′17″, 82°44′15.4″], “ Habrochiton vernus Sem. 1973 N. Skopin det.”; Feb.–Mar. 1890; M.V. Pevtsov leg.; ZIN. Paralectotype CHINA • 1 ♀; “OaЗ. НiЯ, Вост. Турк., 3/ 3 II. 2/ 3.III.-90, Пѣвцов” [Xinjiang: Niya oasis, Eastern Turkestan, 38°01′17″, 82°44′15.4″], “ Habrochiton vernus Sem. 1973 N. Skopin det.”; Feb.–Mar. 1890; M.V. Pevtsov leg.; “ Habrochiton vernus Typ 11.07 A. Semenow det.”; ZIN. Distribution China (Xijiang, Taklamakan Desert)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 17-18, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425

Earophanta beludzhistana Bogatchev, 1957 Fig. 7E–F Material examined Holotype PAKISTAN • ♂; “Quetta, Rost”; “Holotype”; “ Earophanta anglorossica sp. n., A. Semenow-Tian- Shansky det.”; “ Ear. beludzhistana sp. n. typ A. Bog. A. Bogačev det.”; ZIN. Note This species was originally described from a single specimen and known only from the type locality. Key to genera of Pimeliini with dorso-lateral eyes from Central Asia, Iran, Afghanistan and Pakistan 1. Pronotum without large semispherical or conical tubercles or granules, with fine cuticular microgranules or smooth, sometimes with short spines on surface. Body completely covered with scales. Ovipositor very short and weakly sclerotized, with membranous, rounded and densely pubescent apical lobes; spiculum ventrale wide and short, without or with very short common stem (subtribe Habrobatina)...................................................................................................................... 2 – Pronotum with large semispherical or conical tubercles or granules. Body without scales. Ovipositor long, with well-expressed long paraproct and many sclerotized parts, and fossorial, strongly sclerotized, glabrous apical lobes; spiculum ventrale long, with long common stem and narrow rods (subtribe Pimeliina, the Platyope genus group)................................................................................ 5 2. Elytra dorsally with bronze scales and pubescent with erect, yellow setae, especially dense and long in wide longitudinal depression on sides of each elytron. Protibiae with strong, conical granules on outer margin, each granule with strong spine at apex................................ Apatopsis Semenov, 1891 – Elytra dorsally with reddish-brown or creamy scales, or with combination of creamy and brown scales; sometimes with eight longitudinal lines (elytra all together) of short palesetae, but without dense long setaesetae and wide longitudinal depression on sides. Outer margin of protibiae with very short tubercles bearing long thin spines............................................................................................ 3 3. Prothorax with black, clearly visible, small spines and callus-like smooth shiny areas on dorsolateral sides near base. Scales on elytra form spotted or striped creamy-brown colour pattern................................................................................................................................. Dietomorpha Reymond, 1938 – Prothorax without small, black spines and callus-like smooth shiny areas on dorsolateral sides. Elytra with monochromatic scales, without colour pattern.............................................................. 4 4. Head, pronotum and elytra with reddish scales and white spine-like setae (clearly visible among scale cover); setae form eight longitudinal lines on elytra. Width of intercoxal process of the first abdominal ventrite shorter than width of one metacoxa......................... Habrobates Semenov, 1903 – Head, pronotum and elytra with creamy scales, without spine-like setae. Width of intercoxal process of the first abdominal ventrite broader than width of one metacoxa......................................................................................................................................... Habrochiton Semenov-Tjan-Shansky, 1907 5. Elytra spherical, more than twice as wide as pronotum. Lateral margins of pronotum widely completely emarginated in dorsal view. Prosternum before procoxae long, its length between procoxae and anterior margin 1.6 × as short as transverse diameter of one procoxa. Elytra with alternating longitudinal stripes of pale dense recumbent setae and dark, weakly pubescent lines, without coarse granules or large smooth tubercles................................ Przewalskia Semenov, 1893 – Elytra oval, not more than 1.7 × as wide as pronotum. Lateral margins of pronotum weakly or strongly rounded in dorsal view. Prosternum before procoxae short, its length between procoxae and anterior margin 1.8–14 × as short as transverse diameter of one procoxa. Elytra smooth and bare or with large, smooth granules, tubercles or elevations and pale pubescence between them............... 6 6. Elytra smooth and glabrous, only with very small, sparse granules and one elevated line of slightly larger granules along granulated lateral margin. Pronotum with strongly arcuately emarginated base, posterolateral angles protruding far into impressed base of elytra. Prosternal process very broad, weakly narrowed between procoxae, slightly raised between procoxae and horizontally flattened (lateral view), strongly protruding beyond posterior margin of procoxae to mesoventrite. Protibiae with dense, long, thin spines and setae along outer margin.................... Mantichorula Reitter, 1889 – Elytra with coarse granules or/and smooth tubercles completely or at least on sides and pubescence between them or at least apical striped pubescence (sometimes elytra without pubescence as in Platyope ordossica Semenov-Tjan-Shansky, 1907). Pronotum with straight or very weakly emarginated base, posterolateral angles not protruding. Prosternal process broad, but usually visibly narrowed between procoxae, not raised between procoxae, not protruding or very weakly protruding beyond posterior margin of procoxae, rounded down. Protibiae with teeth on outer margin.......... 7 7. Procoxal cavities externally closed. Protibiae with large teeth or teeth partly merged into thin entire lamella (as in Earophanta beludzhistana Bogatchev, 1957); length of largest teeth is at least one third of width of protibia at apex. Lateral rib of elytron with strong acute “serrate” teeth........................................................................................................ Earophanta Semenov, 1903 – Procoxal cavities externally open, postcoxal bridges discontinuous at middle. Protibiae with small teeth; length of largest teeth one-sixth to one-eighth of width of protibiae at apex. Lateral rib of elytron with large or small, wound or conical and acute, but not serrate tubercles................................................................................. Platyope Fischer von Waldheim, 1820, Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on pages 58-59, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Semenov A. P. 1891. Diagnoses coleopterorum novorum ex Asia centrali et orientali. Horae Societatis Entomologicae Rossicae 25: 262 - 382.","Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184.","Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264.","Reitter E. 1889. Insecta, a cl. G. N. Potanin in China et in Mongolia novissime lecta. XIII. Tenebrionidae. Horae Societatis Entomologicae Rossicae 23: 678 - 710.","Fischer von Waldheim G. 1820. Entomographie de la Russie (Entomographia Imperii Rossici). Auctoritate Societatis Caesareae Mosquensis naturae scrutatorum collecta et in lucem edita. Vol. 1. A. Semen, Moscow."]}

Przewalskia dilatata (Reitter, 1887) Figs 4A–B, 11A–D, 18, 23D, 24B, 26E, 28A–B Material examined Lectotype of Platyope dilatata designated here CHINA • 1♂; “Prj 85” [green label]; “(Przewalsky) Nördlichen Tibet”; “Typus Platyope ditatata Rtt. Coll. Reitter ”; “ Platyope dilatata m.”; N.M. Przewalsky leg.; HNHM. Paralectotype CHINA • 1 spec.; “IV” [green label]; “Prj 85”; “Typus Platyope dilatata Rtt. Coll. Reitter ”; HNHM. Additional material CHINA • 10 ♂♂, 7 ♀♀; Niya; 3 Mar. 1890; M.V. Pevtsov leg.; ZIN • 10 ♂♂, 6 ♀♀; same collection data as for paralectotype; 1–20 Apr. 1890; ZIN • 21 ♂♂, 40 ♀♀; between Keriya and Niya; 8 Mar. 1890; B.L. Grombchevsky leg.; ZIN • 1 ♂, 1 ♀; Keriya; 5 Mar. 1890; B.L. Grombchevsky leg.; ZIN • 1 ♀; same collection data as for preceding; 6–11 Apr. 1890; ZIN • 15 ♂♂, 10 ♀♀; Karasay; May 1890; M.V. Pevtsov leg.; ZIN • 7 ♂♂, 15 ♀♀; Niya; 1891; V.I. Robarovsky leg.; ZIN • 1 spec.; Turk. Chin. [Turkestanian China], Keria; 5 Mar. 1890; B.L. Grombchevsky leg.; HNHM • 1 spec.; Sinkiang [Xinjiang]; 5 May 1960; unknown collector; HNHM. Distribution China (Xinjiang: Taklamakan Desert; Qinghai)., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 31, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Reitter E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia Centrali novissime lecta. IX. Tenebrionidae. Horae Societatis Entomologicae Rossicae 21: 355 - 389."]}

Earophanta pubescens Skopin, 1960 Material examined Holotype KAZAKHSTAN • ♂; “Ю.З. ПрибалхаШье, КолШенгил, 25.04.1951, M. Мальховский” [SW Balkhash region, west of Muyunkum Desert, Kolshengil]; “Holotypus, Europhanta pubescens 1959 N. Skopin det.”; 25 Apr. 1951; M. Mal’khovskiy leg.; ZIN. Paratypes KAZAKHSTAN • 2 ♂♂; same geographic labels as for holotype; “Paratypus, Earophanta pubescens 1959, N. Skopin det.”; ZIN. Additional material KAZAKHSTAN • 1 spec.; Muyunkum; 26 Apr. 1954; M. Serkova leg.; “ Earophanta pubescens sp. n. Typ. 1959 N. Skopin ”; “Holotypus 1960 Earophanta pubescens Skopin ” (curator’s label); HNHM • 1 spec.; same collection data as for preceding; “ Earophanta pubescens sp. n. paratyp. 1959 N. Skopin ”; “Paratypus 1960 Earophanta pubescens Skopin ” (curator’s label); HNHM • 6 ♂♂, 1 ♀; 18 km E of Kokozek [Kok-uzek]; 5 Oct. 1961; Kunitsky leg.; ZIN • 1 ♂, 1 ♀; same collection data as for preceding; 8 May 1962; ZIN • 2 ♂♂; Karatal; 30 Jun. 1965; N.G. Skopin leg.; ZIN • 1 ♂; Karatal; 100 km below Ushtobe; 28 Apr. 1965; N.G. Skopin leg.; ZIN • 3 ♂♂, 4 ♀♀; Muyunkum Desert; 23 Mar. 1963; N.G. Skopin leg.; ZIN Distribution Kazakhstan (Balchash region, Muyunkum Desert) (Skopin 1960, 1968). Note Skopin and curators marked two specimens of E. pubescens from HNHM as a holotype and a paratype, respectively. However, Skopin (1960) did not include these specimens in the type series., Published as part of Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V., 2022, A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes, pp. 1-71 in European Journal of Taxonomy 809 on page 57, DOI: 10.5852/ejt.2022.809.1719, http://zenodo.org/record/6419425, {"references":["Skopin N. G. 1960. Neue Tenebrioniden aus Zentralasien. I. Annales Historico-Naturales Musei Nationalis Hungarici 52: 295 - 311."]}