Daniela Ruberti, Gabriele Carannante, Lucia Simone, Francesco Toscano, Ruberti, Daniela, Toscano, F, Carannante, G, Simone, L., PEDLEY, H.M. CARANNANTE G., Carannante, G., and Toscano, F.
Shallow-water foramol limestones have been studied from a locality in the southern Apennines in which outcrop conditions show an excellent overview of the lateral and vertical evolution of rudist bodies and allow their geometry and the dynamic aspects to be reconstructed. The lithofacies suggest open depositional settings characterized by peloidal siltymuddy sediments. Rudists inhabited well-defined sectors of these shelves, giving rise to wide biostromal bodies, and supplied most of the skeletal debris via bioerosion and minor physical breakdown. In particular, the characteristics of rudist lithosomes document the existence of a complex network of channel-like depressions. In such a depositional context, the evolution of rudist lithosomes was controlled by the environmental hydrodynamic conditions. The resulting composite rudist assemblages are characterized by often-toppled individuals, suggesting continuous sediment removal between the organisms. The good preservation of the shells and the common articulation of the valves, however, point to an absence of sustained transport but rather a slight sediment destabilization. The gross lenticular geometry of the shell beds could be related to the above-mentioned patterns of weak, perhaps channelized, pathways. In such a depositional context, rudist colonization on channel margins assumes particular importance as it documents the rudist ability to exploit a wide array of environments, comparable to that of oysters in Recent seas, and reflecting the probable opportunistic nature of rudists. In the last few decades, several studies carried out on the Upper Cretaceous rudist-bearing carbonate sequences of the central-southern Apennines and northwestern Sardinia, Italy, have shown the inadequacy of the tropical, chlorozoan model applied to the depositional environment of these sequences: a ‘temperate-type’, foramol model was suggested by Carannante & Simone (1987) and Carannante et al. (1995, 1997, 1999) based on the nature of the skeletal grains, biological assemblages, lithofacies organization, sedimentary body geometries and responses to sea-level changes. Similar characteristics have been recognized, on a very large scale, in many other Upper Cretaceous rudist-bearing limestones cropping out in the periMediterranean region (cf. Gili et al. 1995; Moro 1997; Moro et al. 2002; Stossel & Bernoulli 2000; among others). Many complex factors controlled the inception, growth and demise of the carbonate factories and the sedimentary bodies they produced were related to the interplay between eustasy and tectonics in the sedimentation basin, plus ecological factors, which determined the delicate biological balance of the primary sediment producer assemblages (see Carannante et al. 1995, 1997; James & Clarke 1997; Weissert et al. 1998). The temperate-type carbonate depositional model led to critical reinvestigation of many Cretaceous carbonate sequences, characterized by the flourishing of opportunistic rudist-dominated communities. Previous studies allowed a possible sedimentary scenario to be reconstructed for the Upper Cretaceous shallow-water depositional areas, pertaining to the southern Tethys belt. According to recent papers, these depositional areas were complexes of unprotected shelves occupied by foramol assemblages that produced loose, diagenetically stable bioclastic debris not involved in significant in situ cementation processes. Both stormand wind-induced currents and waves exercised strong control on the distribution of the shifting biogenic sediments which covered the open seafloor, constituting large coalescing sheets of winnowed fine to coarse skeletal sands (cf. Simone et al. 2003). Rudists spread over all shelf sectors, from more open and external areas to more internal ones, occupying different substrata and providing the bulk of the skeletal component by means of bioerosion processes. They colonized mobile sediments giving rise to complex bodies with peculiar characteristics related to environmental conditions of the different sectors of the shelf. On the basis of detailed sedimentological, taphonomic and palaeontological data, two main rudist-rich depositional settings (‘end-members’) have been recognized in the central-southern Italy Senonian rudist-bearing successions (Carannante et al. 1993, 2001; Simone et al. 2003). The latter were characterized by contrasting current regimes (high energy vs. low energy; cf. Burchette & Wright 1992) which controlled development and growth of the rudist lithosomes (see also analogous observations in Stossel & Bernoulli 2000). The taphonomic and palaeoecological features of the rudist-dominated benthic communities have been analysed in limestones, showing that they were deposited in very different hydrodynamic conditions. Investigations have pointed out the differences in terms of anatomy of the rudist bodies and their relationship with the availability of accommodation space (cf. Simone et al. 2003, for a review). Several papers deal with the rudist-bearing marginal depositional settings (cf. Simone et al. 2003 and references therein) in which more or less frequent episodes of high energy (storm-related events) resulted in a complete reorganization of the depositional bodies and in which only a few of the organisms were able to keep their original position. Moreover, few papers deal with successions pertaining to more internal and/or lowenergy sectors, in which rudist-rich beds rhythmically alternate with finer-grained foraminiferal limestones (cf. Simone et al. 2003 and references therein). In these sequences, dominated by silt/ mud-rich lithotypes, small elevator radiolitids with oligospecific diversity are dominant. In the related depositional settings, the preservation of the original rudist congregations was associated with the in situ maintenance of the bioerosion-derived finer skeletal debris. Rudists in growth position are abundant, although also a large quantity of shells appears toppled with little reworking. They may form laterally continuous biostromal shell beds. Sedimentary structures such as cross-lamination and grading are present only occasionally. The resulting facies are arranged commonly in peritidal/ shallow subtidal cycles in which evidence of subaerial (up to pedogenic modifications on a large and small scale) and, less frequently, submarine exposure is common. Based on detailed observations, in such sedimentary contexts, peculiar areas have been identified corresponding to current pathways in a persistently subtidal setting (Carannante et al. 2003) which cannot be likened to well-described tide channels in tropical contexts. Several papers deal with temperate, high-energy, recent and ancient depositional systems (cf. James & Clarke 1997 for a review). Low-energy or protected, temperate, shallow-water systems have been described poorly and have come to international attention only recently (cf. Choi & Simo 1998; Gomez-Perez et al. 1998; Lukasik et al. 2000; Carannante et al. 2003; Pufahl et al. 2004). This paper sets out to describe such a peculiar shallow-water environment from late Cretaceous (post-Cenomanian) shallow-water foramol open shelves. A locality was selected in the southern Apennines in which outcrop conditions show an excellent overview of the lateral and vertical evolution of rudist bodies and allow the geometry and the dynamic aspect to be reconstructed.