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Knowledge of drought variability and their possible mechanisms during the past hundred years is still limited in the mountainous region of south-central Tibetan Plateau (TP). In this study, a long-term tree-ring width chronology dating back to 1190 CE was combined using 328 increment cores from the Nagqu region. Based on the relationships between this tree-ring width chronology and climate data, we reconstructed May–June self-calibrated Palmer Drought Severity Index (scPDSI) for the past 821 years (1190–2010 CE). Additional comparisons with other available precipitation or drought reconstructions were conducted. We further investigated the influence of the South Asian summer monsoon (SASM) on the drought variability in our study region. Results indicated that our tree-ring width chronology contained stable drought signal in the early summer season (May–June). During the past 821 years, the longest dry and wet periods lasted for 116 and 90 years, respectively, based on a 21-year Fast Fourier transform filter. Specifically, longer than ten years’ dry periods prevailed during 1211–1245 CE, 1280–1358, 1421–1471, 1500–1571, 1580–1598, 1650–1691, 1782–1807 and 1867–1982; while wet intervals occurred in 1190–1210 CE, 1246–1279, 1359–1420, 1472–1499, 1599–1649, 1692–1781, 1808–1866 and 1983–2010. Generally consistent dry and wet intervals across the southern TP were found by comparisons with other available datasets during their common periods. Interestingly, we detected an unstable influence of the SASM on the May–June drought variability in our study region, at least for the past three and a half centuries. This study therefore gives a new perspective of drought variability as well as their relationships with the SASM over a long-term period on the south-central TP. [ABSTRACT FROM AUTHOR]

Climate change has a significant impact on natural ecosystems, particularly on floodplain forests that are among the most transformed ecosystems in the world. The climate sensitivity of dominant species is likely to play a key role in determining the susceptibility of flooded forests to climate changes. Here, we use dendrochronological approaches and local climate records from 1880 to 2015 to assess the response in pedunculate oak (Quercus robur L.) trees growing in a floodplain of the Dnipro River near Kyiv to climatic variables. Correlation analysis reveals the strongest positive association of the Q. robur tree-ring width chronology with May-June precipitation, May-June temperature, and May self-calibrating Palmers drought severity indices (scPDSI). The moving-window correlation analysis points to positive association with the scPDSI after the 1950s, when local river regulation was implemented. The positive correlation with current March precipitation is the least expected change in the oak growth-to-climate relationship that occurred in the aftermath of human alterations in the local river and regional climate changes. This study discusses the probable ecological consequences and ecophysiological mechanisms of observable climate-to-growth relationships and their temporal stability. [ABSTRACT FROM AUTHOR]

Density is an important wood mechanical property and an indicator of xylem architecture and hydraulic conductivity. It can be influenced by forest management and climate. We studied the impact of thinning and climate variables on annual stem radial growth (ring width and ring density, and their earlywood and latewood components) in two contrasting Scots pine (Pinus sylvestris L.) stands in northern Spain (one continental, one Mediterranean). At each site, three thinning regimes (control or T0, removing 20% basal area or T20, and removing 30% or T30) were randomly applied to nine plots per site (three plots per treatment) in 1999. Thinning was repeated at the Mediterranean site in 2009 (increasing thinning intensity in T30 to 40%). Eight trees per plot were cored in spring 2014. Second thinning at the Mediterranean site and first thinning at the continental site generally caused significantly wider ring (RW), earlywood (EW) and latewood (LW) widths, although no differences between T20 and T30/40 were found, supporting in part the common observation that radial growth is enhanced following thinning as competition for water and nutrients is reduced. At the Mediterranean site, values of latewood density (LD) and maximum density (Dmax) relative to pre-thinning conditions were significantly lower in T0 than in T30. However, at the continental site, relative changes of ring density (RD) and LD were significantly higher in T0 than in T20 and T30. Climate significantly affected not only RW but also RD, with significant RD drops during or right after unusually warm-dry years (e.g., 2003, 2011), which were characterized by LD reductions between 5.4 and 8.0%. Such RD decreases were quickly followed by recovery of pre-drought density values. These results indicate trees temporarily reduce LD as a way to enhance hydraulic conductivity during dry summers. However, climate effects on wood density were site-dependent. We also detected that the thinning effect was not intense enough to prevent drought-induced changes in wood density by altering water availability, but it could help to reduce wood properties fluctuations and therefore maintain more homogeneous wood mechanic features. [ABSTRACT FROM AUTHOR]

A dendrochronological profile was generated from Chinese pines ( Pinus tabulaeformis Carr.) in the Qianshan Mountains in northeastern China. Based on correlation analyses, the pattern of precipitation from March to June ( P ) was reconstructed using a simple linear model, which explained 42.7% of the total variance in observed precipitation from 1951 to 2012. The reconstructed P series revealed a consistently increasing trend in precipitation during the twentieth century in the Qianshan Mountains. The reconstructed data showed trends that were similar to those in the variation in trends for March-June precipitation observed at the Shenyang station, the reconstructed January-May precipitation trends in Shenyang City, and the reconstructed average June-September relative humidity for Yiwulü Mountain. The reconstructed data also showed good agreement with the droughts reported in historical documents and recorded by meteorological stations in Liaoning. Spatial correlation analyses show that the reconstructed data reflect the variability in precipitation that occurs over much of northeastern China. In addition, our reconstruction showed a significant periodicity. The significant correlations between the reconstructed P and the El Niño-Southern Oscillation (ENSO), Pacific Decadal Oscillation (PDO) and sunspot numbers indicate that precipitation variability in the Qianshan Mountain region is probably driven by extensive atmosphere-sea interactions and solar activities. [ABSTRACT FROM AUTHOR]

We describe the physiological responses of boreal conifers to climate change for the past 112 years using ring-width and carbon isotope ratio (δ13C) chronologies at six forest sites in northern Eurasia and Canada. Responses differed among regions, depending on their climatic and/or geographic characteristics. Tree radial growth decreased over the past 52 years in central eastern Siberia with the higher rate of summer temperature increase than other regions, as indicated by the negative correlation between radial growth and summer temperature, but increased in northern Europe and Canada. Changes in tree-ring δ13C indicated that recent climatic conditions have induced stronger drought stress for trees from central eastern Siberia than for those from other regions. The observed tree growth trends were compared to those simulated using a dynamic global vegetation model. Although the modeled annual net primary production (NPP) for trees generally exhibited similar decadal variation to radial growth, simulations did not show a recent decrease in tree growth, even in central eastern Siberia. This was probably due to an overestimation of the sensitivity of modeled tree NPP to precipitation. Our results suggest that the tree NPP forecasted under the expected future increases in temperature and average precipitation might be overestimated, especially in severely dry regions such as central eastern Siberia. [ABSTRACT FROM AUTHOR]

Age, genetics and social status of trees affect their sensitivity to environmental factors, and information about such effects is needed for comprehensive assessment of growth potential. Climatic sensitivity of radial increment (i.e., tree-ring width) of introduced European beech ( Fagus sylvatica L.) of different generations and social status, growing in its northeasternmost stands in Europe, was studied by dendroclimatological methods. At present, the studied stands occur outside of the natural distribution area of the species, providing opportunity to study adaptability and potential growth of beech in novel environments under changing climate. The sensitivity of radial growth to climatic factors was modulated by the generation and social status (size) of trees. The first generation trees, which were propagated from the material transferred from the northern Germany, were highly sensitive to climatic factors and showed wide spectrum of responses. The dominant trees were particularly sensitive to June precipitation, indicating sensitivity to water deficit in summer. The suppressed trees were mainly sensitive to temperature in the dormant period. Tree-ring width of the second generation trees, which were propagated from the first generation stands, was mainly affected by water deficit in summer, yet the local factors, modulated the mechanisms of response. In one stand, tree-ring width was affected by conditions during the formation of tree-ring, indicating direct influence of weather conditions on xylogenesis. In the other stand, tree-ring width was correlated to weather conditions in the preceding year, suggesting influence via carbohydrate reserves. The effect of social status on climatic sensitivity in the second generation stands was considerably weaker, likely due to the natural and anthropogenic selection of the material best adapted for local conditions. The effect of climatic factors on radial growth of beech has shifted during the 20th century. The effect of autumn temperature has weakened, likely due to warming; the effect of factors related to water deficit in summer has intensified that could be related to both, changes in climate and ageing. The observed climate-growth relationships suggested that conditions in winter have become suitable for beech, yet careful selection of sites/regions with appropriate hydrological conditions appear necessary to counteract the increasing effect of water deficit, hence to ensure productivity of future beech sites in Latvia. [ABSTRACT FROM AUTHOR]

Key message: This paper provides the first interannual variability of PDSI series in Central Inner Mongolia, China. Abstract: A Pinus tabulaeformis tree-ring-width chronology in Baotou, China, was developed. Based on the chronology, we reconstructed the local interannual variability of the Palmer Drought Severity Index (PDSI) from the previous September to current July (PDSI) for the past 278 years. The reconstruction explained 41.3% of the variance (40.3% after adjusting the degrees of freedom) of the actual interannual variability of the PDSI during the calibration period (1951-2012 AD). The spectral analyses revealed 3.5-, 2.6-, 2.3-, 2.1- and 2-year quasi-cycles at a 95% confidence level and 7.9-, 7.1- and 5.2-year quasi-cycles at a 90% confidence level. The PDSI reconstruction series was consistent with several reconstructed climatic series based on the tree-ring chronology in neighboring regions. The PDSI exhibits excellent regional representativeness, which indicates the consistency of the drought variation of the central part of the monsoon margin area in north China, whereas PDSI can represent the drought history of the central part of the north monsoon margin area. In addition, significant relationships were found between the PDSI series and the Arctic Oscillation, North Atlantic Oscillation, El Niño-Southern Oscillation and Pacific Decadal Oscillation series, which indicates that the PDSI variation in the study area was influenced by a combination of these factors. [ABSTRACT FROM AUTHOR]

In order to investigate the effects of climate factors on the tree-ring width of Quercus acutissima in the rocky mountain area of northern China, correlation analysis and response function analysis were used to determine the relationship between the tree-ring width of Quercus acutissima and climate factors. The results showed that: 1. The residual (RES) chronology of the Quercus acutissima forest contained richer environmental information than standard (STD) chronology and was suitable for response analysis. 2. Tree-ring width had a general positive correlation with the average monthly temperature, a significantly positive correlation (P<0.05) with the April temperature, and a highly significantly positive correlation (P<0.01) with the May temperature of the current year. Tree-ring width had a positive correlation with precipitation from March to June in the current year, especially a significantly positive (P<0.05) correlation with May and June precipitation, and a negative correlation with precipitation from September of the previous year to February of the current year; however, the correlation coefficients with precipitation were basically lower than the coefficients with temperature basically 3. Temperature and humidity index were the main climate factors that influenced the tree-ring width of Quercus acutissima, followed by precipitation. [ABSTRACT FROM AUTHOR]

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Dendrochronological studies have revealed several environmental signals of many oak species supporting the suitability of these species for climate reconstruction (Haneca et al. 2009). Despite the large number of studies, limited information is available on the influence of climatic condition and soil moisture on the growth characteristics of Turkey oak (Q. cerris). Although soil moisture is probably the most limiting factor on growth of most oak species, relevant studies in Hungary primarily focused on the effect of temperature and rainfall, and ignored the impact of soil moisture. The combined analysis of climatic factors and soil moisture would reveal the importance of the influencing factors and would point out the anatomical patterns and environmental signals of a given area. In the current study, we analyzed the dendrochronological characteristics of a grove in Western Hungary (Central Transdanubian Region), and correlated tree anatomical patterns with the mean monthly temperatures, monthly precipitation totals, and modeled soil moisture at five different depths. The grove is exposed to environmental stress due to the local soil and climatic factors. Our findings indicate that the most pronounced environmental signal stored in the oaks is the change of soil moisture that exposes a direct impact on tree growth. [ABSTRACT FROM AUTHOR]

We present the longest high-elevation tree-ring width dataset in the Mediterranean reaching back to the 6th century CE. The network includes 101 living and 92 relict Pinus heldreichii Christ trees from four differently exposed sites in the 2100–2200 m a.s.l. elevation range of Mt. Smolikas in the Pindus Mountains in Greece. Though the sites were all sampled within a distance of <1 km, inter-site correlations are surprisingly low (r 1550–2014 = 0.65–0.87), indicating site exposure might affect tree-ring formation. We here explore the consequence of exposure differences on the climate signals in an eastern Mediterranean treeline ecotone. Temporally stable growth/climate relationships reveal similar seasonal patterns among the four sites, but differences in signal strength. P. heldreichii growth at Mt. Smolikas is significantly controlled by temperature in April (r 1951–2014 = 0.33–0.50) and precipitation in June-July (r 1951–2014 = 0.23–0.42), which emphasizes the overall importance of an early growth onset and subsequent moisture conditions. The association between stem growth and April climate is strongest in the South-facing stand, supporting the significance of higher insolation rates at this thermally privileged site. Strongest summer precipitation signals are found in the NE-facing stand, where trees seem to benefit least from an early growth onset and where reduced meltwater supply may enhance the dependency on early summer precipitation. The significance of spring temperature on tree growth in all four sites constrains the emergence of a distinct summer precipitation signal in the Mt. Smolikas high elevation ecotone. Exploration of the site-specific influences on a new millennium-long tree-ring width dataset is an important step towards an improved understanding of long-term climate variability in the Eastern Mediterranean. Site-related differences in climate sensitivity in the high-elevation tree-ring network at Mt. Smolikas indicate that both temperature and precipitation during different seasons could potentially be reconstructed if distinct site exposures (S versus NE) are considered. [ABSTRACT FROM AUTHOR]

ABSTRACT Understanding long-time scale of drought variability is vital to the hydrological management and agriculture in dryland. Based on two sites of Schrenk spruce ( Picea schrenkiana), a robust tree-ring chronology was developed and employed to study the drought variability in central Tianshan Mountains. The result shows that the chronology significantly positively correlates with precipitation in previous December, April, May and June in current year, and significantly negatively correlates with temperature in April, May, July and August in current year. Highest correlation was found between tree-ring width and both April-June standardized precipitation evapotranspiration index ( SPEI) and Palmer drought severity index, indicating that Schrenk spruce's growth strongly responds to the current summer drought. We further reconstruct SPEI for the period 1820-2005, and the series are verified both with the statistical methods and other nearby reconstructions. Multi-taper method spectral and wavelet analyses reveal the significant low- and high-frequency cycles (26-36, 8.5, 6.0, 5.7 and 3.4 years). Drought has alternatively varied in the past two centuries with four apparently dry periods and wet periods, and the driest period occurs from 1870 to 1890 and the wettest period occurs from 1842 to 1855 in the central Tianshan Mountains. Some extreme drought events are detected, such as 1885, 1918, 1944-1945 and 1951. Significant correlations of reconstructed SPEI and precipitation and temperature indicate that the source of air masses and the water vapour transport paths lies in the northwestern direction of our study area and are controlled by westerly circulation. This SPEI reconstruction provides a long-term drought history in the central Tianshan Mountains and can serve as scientific basis for water resources management in this semi-arid region. [ABSTRACT FROM AUTHOR]

ABSTRACT High mountains are some of the most vulnerable regions to climate change and therefore a matter of global concern. Here, the climatic growth factors of conifers and their course in time and space along an elevational gradient in the northwestern Himalayan part of India were studied. Increment cores of Juniperus semiglobosa and Cedrus deodara (xeric species), and Abies pindrow and Picea smithiana (mesic species) were collected from thirteen sites. Tree-ring width and maximum latewood density were measured and cross-dated. The time-series were standardized and site- and species-level chronologies were built. Static and moving bootstrap correlation and response functions between the tree-ring chronologies and monthly climatic variables were computed. The largest climate changes in the region were increasing winter and early-spring temperatures and decreasing monsoon precipitation. The growth of all species was negatively correlated with pre-monsoon temperature, as the higher temperatures probably increased evapotranspiration and caused water deficit. The phenomenon was most pronounced in May, but also in June for the Juniperus at the northernmost, highest, and driest sites. The pre-monsoon temperature signal of the drought-prone Juniperus and Cedrus endured, while the signal of the climate-susceptible Abies and Picea at the mesic sites was unstable. Namely, the May temperature signal of Picea became significant since the second half of the last century whereas the signal of Abies shifted from May to April. This apparently related to the earlier onset of spring due to the accelerated warming in the region. Besides, maximum latewood density of Picea and Abies negatively correlated with May and June temperature, respectively. Additionally, the Cedrus benefitted from winter precipitation and the Abies and Picea from pre-monsoon rainfall. Counterintuitively, we detected no direct effect of monsoon precipitation decrease on the conifers because their growth was driven by pre-monsoon conditions, which changed only slightly. [ABSTRACT FROM AUTHOR]

The 2003 and 2012 summer seasons were among the warmest and driest of the last 200 years over southeastern Europe, and in particular in the Karst region (northeastern Italy). Starting from winter-spring 2013, several black pines (Pinus nigra J.F. Arnold) suffered crown die-back. Declining trees occurred nearby individuals with no signs of die-back, raising hypotheses about the occurrence of individual-specific hydraulic strategies underlying different responses to extreme drought. We investigated possible processes driving black pine decline by dendrochronological and wood anatomical measurements, coupled with analysis of tree-ring carbon (δ13C) and oxygen (δ18O) isotopic composition in healthy trees (H) and trees suffering die-back (D). Die-back trees showed higher growth rates than H trees at the beginning of the last century, but suffered important growth reduction following the dry summers in 2003 and 2012. After the 2012 drought, D trees produced tracheids with larger diameter and greater vulnerability to implosion than H ones. Healthy trees had significantly higher wood δ13C than D trees, reflecting higher water-use efficiency for the surviving trees, i.e., less water transpired per unit carbon gain, which could be related to lower stomatal conductance and a more conservative use of water. Relatively high δ18O for D trees indicates that they were strongly dependent on shallow water sources, or that they sustained higher transpiration rates than H trees. Our results suggest that H trees adopted a more conservative water-use strategy under drought stress compared with D trees. We speculate that this diversity might have a genotypic basis, but other possible explanations, like different rooting depth, cannot be ruled out. [ABSTRACT FROM AUTHOR]

Key message: In the center of the species' distribution range in Germany, European beech stands show continued climate-change-related growth decline since the 1980s at low elevations, but growth increase at high elevations. Abstract: Contradicting reports exist about the climate-change sensitivity of European beech ( Fagus sylvatica), showing either a sensitive response of radial growth to dry and hot summer episodes and long-term growth decline with recent warming, or apparent insensitivity of growth and a remarkable potential for post-stress recovery. With a dendroecological study along an altitudinal transect in the center of the species' distribution range in south-western Germany, we analyzed the climate response of radial growth with variation in altitude (110-1230 m) and associated temperature (10.6-3.5 °C MAT) and precipitation change (755-1788 mm year MAP). Climate sensitivity analysis showed that annual stem increment was strongly limited at low elevations (110-300 m) by low precipitation in April/May, but by low summer temperatures at 1230 m. At intermediate elevation (640 m), indications of both moisture and temperature limitation were found. The differences in climate sensitivity were linked to contrasting long-term growth trends. At 110-300 m, radial growth has continually decreased since about the 1980s, while it has increased at 1230 m. Our results from the four stands suggest for the study region that the abiotic control of beech radial growth switches from moisture to temperature limitation at a threshold situated between 160 and 235 mm of precipitation in April/May (which corresponds to 200 and 313 mm of precipitation in June-August), in accordance with dendroecological results from other Central European lowland regions. This indicates that, with further warming and drying of the climate, beech may suffer in lowland and lower montane regions of Central Europe from reduced vitality and productivity, whereas it may profit from warming in montane to upper montane elevation. We conclude that climate-change-related growth decline is more widespread in the center of the species' distribution range than previously thought, which is highly relevant for forestry planning. [ABSTRACT FROM AUTHOR]

The Kashmir region in the western Himalayas is located in a transition zone between areas dominated by the South Asian Summer Monsoon (SASM) and the North Atlantic Oscillation (NAO). Currently being primarily influenced by westerly disturbances (WDs), the area is important to decipher teleconnections between these two important circulation systems for the assessment of past climate variability. We evaluated climate-growth relationships of Abies pindrow (Royle ex D. Don) and reconstructed April to June (AMJ) self-calibrated Palmer drought severity index (scPDSI) for the south Kashmir region during the period 1643–2016 CE. Our reconstructed scPDSI revealed a long wet phase during 1650–1816 CE, indicating the impact of the Little Ice Age (LIA) over the region, followed by prominent drier post-LIA episodes. The mid 18th century (1730–1760 CE) was the wettest period in the past four centuries, whereas the period 1817 to 1865 CE marked the driest phase. These phases are consistent with other precipitation reconstructions from the WD-dominated western and Trans-Himalayan regions, but inconsistant with summer precipitation reconstructions from the SASM-dominated Himalayan regions. A significant positive correlation between our scPDSI reconstruction and the North Atlantic Oscillation (NAO) for the wet phase of the LIA suggests that the NAO remained dominant in modulating the winter and spring precipitation at the study region. During the 19th and 20th centuries, scPDSI was either weakly or negatively correlated with the NAO index, indicating the influence of other atmospheric circulation systems in driving the spring/summer precipitation in the study area. This study, augmented with other moisture records, contributes to analyze the temporal and spatial extent of moisture variability in a regional perspective. • Abies at westerly dominated Kashmir Himalaya are sensitive to spring moisture. • A 373-yrs of drought reconstruction revealed moist Little Ice Age and dry post-LIA. • Radiative forcings due to volcanic eruption affected the annual growth of Abies. • North Atlantic Oscillations influence winter/spring precipitation at study region. • Strong teleconnection with NINO 3.4 in last four decades due to ocean warming. [ABSTRACT FROM AUTHOR]

[Display omitted] • A new annual runoff was reconstructed since 1672 in Moqu river based on tree ring. • Moqu river was undergoing the most protracted low flow period over the past 348 years. • Runoff reconstruction shows close linkages with climate pattern and Indian summer monsoon. The runoff of Moqu river in Zoige, one of most important water conservation ecological function areas in the upper reaches of the Yellow River, located in the north-east of the Tibetan Plateau, is highly sensitive to climate change. However, the current understanding of the long-term changes in the hydrology of this rives is still fragmentary. In this study, we created a regional tree-ring width chronology from two sample sites in Moqu River Basin and analysed the relationships between tree-ring width, climate, and runoff. We found significant positive correlations between tree-ring width and runoff in multiple months, the highest correlation coefficient occurred with annual runoff from previous July to current June (r = 0.617, 1981–2013). Based on this highly significant (p < 0.01) relationship, we reconstructed runoff variations of Moqu River for the period 1672 to 2019. The reconstructed runoff showed unusual prolonged and frequent low flow periods in recent decades. We found that the continuous low precipitation and the high temperature in the context of global warming are the main reasons for the abnormal runoff in recent decades. We assume that a weakening of the Indian summer monsoon (ISM) circulation system may be the key large-scale factor that caused the unusually prolonged low flow. With global warming, the weakening of ISM intensity leads to reduced water vapor transport from the tropical Indian Ocean to High Asia, resulting in less precipitation and low runoff events in Zoige. Our reconstruction improves the understanding of regional runoff variability over multiple timescales, and also provides a scientific basis for the protection, development, and utilization of water resources related to ongoing global warming. [ABSTRACT FROM AUTHOR]

Large spatial and between-tree variability has recently been observed in the response of boreal forests to ongoing climate change, spanning from growth stimulation by increasing temperatures to drought limitation. To predict future responses of boreal forests, it is necessary to disentangle the drivers modulating the temperature-growth interaction. To address this issue, we established two inventory plots (at a treeline and closed-canopy forest) and assembled site chronologies in Picea glauca stands at the transition between boreal forest and tundra in Northern Quebec, Canada. In addition to site chronologies, we established a set of chronologies containing, for each year, exclusive subsets of tree-rings with specific cambial age (young/old), tree dimensions (small/large) and tree social status (dominant/suppressed). All chronologies were correlated with climatic data to identify the course of climatic conditions driving variability in tree-ring widths. Our results show that the growth of P. glauca correlates significantly with summer temperature in tree-ring formation years and during up to two prior summers. Tree-ring width is positively influenced by summer temperatures in tree-ring formation year and two years prior to tree-ring formation. In addition, climate-growth correlations indicate a negative effect of summer temperature one year before tree-ring formation at the closed-canopy forest site. The pattern of climate-growth correlations is tightly synchronized with previously published patterns of climate-reproduction correlations of P. glauca , suggesting a growth-reproduction trade-off as a possible factor modulating the response of boreal forests to summer temperatures. Climatic signal does not differ between pairs of chronologies based on subsets of cambial ages, stem dimensions or tree competition status at the treeline site. However, the response to summer temperatures one year before tree-ring formation is significant only in mature (old, large and dominant) individuals at the closed-canopy site. The inverse pattern of temperature-growth correlations during a sequence of three years challenges predictions of how boreal forests respond to climate change.

Variability of width, maximum density and stable isotopes (δ13C) in tree-rings of Scots pine (Pinus sylvestris L.) were studied in Northern Caucasus. Statistically sufficient agreements between ring width chronologies allow to construct composite chronology for the Elbrus area. Absence or low correlation between indices of the ring width and maximum density chronology point out different climatic signal. The influence of temperature and precipitation on these tree-ring parameters was also analyzed. The ring width of pine at the upper tree limit in the Baksan valley correlates positively with the June and July precipitation (r=0.3; 0.3; 0.4, p

Background and Purpose: This paper explore the growth-climate relationships in total ring width chronologies of silver fir (Abies alba Mill.) and Bosnian pine (Pinus heldreichii Christ). The objective of this study is to quantify the climate influence on radial growth of both species. The relationships between climate and ring widths were analyzed using extreme growing years (called pointer years), simple correlations and response functions analysis (bootstrapped coefficients). The objectives of this study were: (1) to define the pattern of climatic response of each species, (2) to highlight the influence of local ecological conditions on tree's growth, and (3) to compare the response of silver fir and Bosnian pine to climate. Responses of total ring width to climate were estimated by establishing the mean relationship between growth and climate through simple correlations analysis and bootstrapped response functions. The response to climatic variability was also assessed by analyzing pointer years which correspond to abrupt changes in growth pattern and revealing the tree-growth response to extreme climatic events. For the period 1908-2008 the mean sensitivity (MS) of total ring width chronology for Bosnian pine (0.209) was higher than silver fir (0.169) suggesting that Bosnian pine is more sensitive to climate (pointer years were more frequent in ring width chronology of Bosnian pine than in silver fir ring width chronology). The high values of first-order autocorrelations for Bosnian pine (0.674) indicated a strong dependence of current growth on the previous year’s growth. Pointer years analysis underlined the high sensitivity to spring temperatures and precipitation for both species. Radial growth for both species depends strongly on spring climate variables (temperatures and precipitation) which play a significant role particularly for earlywood production. Material and Methods: We selected 12 silver fir trees and 15 Bosnian pine trees and took two 5 mm cores per tree perpendicular to the slope. Each core was mounted and sanded following standard dendrochronological procedures. Annual radial growth was measured with a measuring system LINTAB (Rinntech-Germany) where tree-ring widths were recorded using TSAPWin 0.55 software. Each tree-ring width series was crossdated using visual comparisons and statistical parameters like: Cross Date Index-CDI (Schmidt 1987), an index of synchronization estimated by TSAPWin derived from GLK% and t value (Baillie and Pilcher 1973). The value of CDI> 10 (Rinntech 2003) were considered as significant. The ARSTAN software was used to remove age trends in the ring width data and build site chronology. Indices were calculated as ratios between the actual and fitted values. Response from both species to climate was correctly shown by pointer years analysis, Pearson’s correlation coefficients and DENDROCLIM (2002) software. Results and Conclusion: We compiled for each specie a mean chronology of radial growth with good replication. The chronology length of Silver fir is 64 years with mean tree-ring width 3.59 mm (0.506), while the chronology length of Bosnian pine is 104 years with mean tree-ring width 2.44 mm (± 0.482). The Bosnian pine chronology was more sensitive than silver fir chronology (sensitivity 0.209 >0.169). Response of both species to climate was good. That was verified from pointer years analysis, Simple Pearson’s correlation coefficients and DENDROCLIM (2002) software. Pointer year analysis showed that spring temperatures and precipitation are the most important factors that enhance radial growth for both species. The pointer years 1953 and 1955 appear to be the most geographically extended pointer years throughout of Europe. The negative pointer year 1953 was identified in growth of silver fir by Serre Bachet 1986, and positive pointer year 1955 was also observed for silver fir in France. Both species have reacted strongly to several pointer years but bosnian pine was more affected and more sensitive to climate than silver fir. For A. alba, high temperatures and plenty rainfalls during the first part of growing season are the keys for production of earlywood). The response of Bosnian pine growth to climate was quite different than response of silver fir. Response functions analysis showed that precipitation during September is important for latewood production (growing season ends on October). While low temperatures during winter (January) and especially frosts cause substantial growth reduction, delaying the growth starting during spring. Earlier studies has shown that photosynthesis is possible for A. alba in winter, where high temperatures could play an important role in improving carbohydrate storage and growth at following year. For species grown under a Mediterranean climate high temperatures and low precipitation during growing season may cause water stress, which is the main limiting factor for tree growth. Although the drying season lasts in the study area for 2 months we did not note any sign of defoliation or needle yellowing in standing trees.

Influence of growth conditions to the wood structure was researched on three species of conifers from various topoecological conditions in forest-steppe zone of Khakassia. By correlation analysis of climatic factors (average temperatures and precipitation amounts on ten-day periods) of growth seasons and structure parameters of wood rings (tree ring width, maximum density, radial size of tracheids and thickness of cell walls) was revealed that at different conditions of growth significant transformation of climatic signal depending on the specific features of tree species takes place. It is shown also that the main target for external signal is cambial zone; subsequent processes of cells differentiation and their result are less depending on external conditions. Nevertheless, cells parameters and particularly their combination at trees from different topoecological conditions allow to receive additional and more detailed information about climatic features of growth season and to reveal key intervals of season, when climatic factors can be evaluated quantitatively.

Tree growth varies closely with high–frequency climate variability. Since the 1930s detrending climate data prior to comparing them with tree growth data has been shown to better capture tree growth sensitivity to climate. However, in a context of increasingly pronounced trends in climate, this practice remains surprisingly rare in dendroecology. In a review of Dendrochronologia over the 2018–2021 period, we found that less than 20 % of dendroecological studies detrended climate data prior to climate-growth analyses. With an illustrative study, we want to remind the dendroecology community that such a procedure is still, if not more than ever, rational and relevant. We investigated the effects of detrending climate data on climate–growth relationships across North America over the 1951–2000 period. We used a network of 2536 tree individual ring-width series from the Canadian and Western US forest inventories. We compared correlations between tree growth and seasonal climate data (Tmin, Tmax, Prec) both raw and detrended. Detrending approaches included a linear regression, 30-yr and 100-yr cubic smoothing splines. Our results indicate that on average the detrending of climate data increased climate–growth correlations. In addition, we observed that strong trends in climate data translated to higher variability in inferred correlations based on raw vs. detrended climate data. We provide further evidence that our results hold true for the entire spectrum of dendroecological studies using either mean site chronologies and correlations coefficients, or individual tree time series within a mixed-effects model framework where regression coefficients are used more commonly. We show that even without a change in correlation, regression coefficients can change a lot and we tend to underestimate the true climate impact on growth in case of climate variables containing trends. This study demonstrates that treating climate and tree-ring time series "like-for-like" is a necessary procedure to reduce false negatives and positives in dendroecological studies. Concluding, we recommend using the same detrending for climate and tree growth data when tree-ring time series are detrended with splines or similar frequency-based filters. • Climate data are rarely detrended prior to dendroecological analyses. • We remind the community of the relevance of detrending climate data in dendro-ecological studies. • Detrending climate data on average strengthened climate–growth correlations. • The stronger the climate trend the stronger the possible correlation change. • Even with no correlation change, regression coefficients can vary substantially. [ABSTRACT FROM AUTHOR]

This study concerns dendrochronology and dendroclimatology of yew growing in Poland. The yew (Taxus baccata L.), a long-lived, slow-growing tree, is regarded as a species threatened with extinction. The eastern boundary of its range transects Poland. The analyses were performed on 34 yew populations which are protected as parts of nature reserves, as nature monuments, or which are planned to be protected. Samples were collected using Pressler borer from a total of 774 trees yielding 1307 profiles. Classical dating techniques (including cross-dating method) were subsequently used to reconstruct 34 local chronologies. Average tree-ring width was 0.84 mm and ranged from 0.27 to 1.47 mm. For most stands, the age of the yew trees is overestimated. The studied populations most frequently are 100–200 years old, and the oldest yew trees in Poland are growing in the KS population (age estimated at 500–600 years). Average tree-ring width was found to be strongly dependent on the chronology length/tree age. The 674 pointer years calculated for the local chronologies served as a basis for determining pointer years for the entire study area. Negative years include: 1862, 1865, 1917, 1927, 1940, 1947, 1956, 1963, 1969, 1976, 1979, 1993, 1996, and 2003. Positive years are: 1884, 1914, 1916, 1965, 1977, 1988, 1997, and 2007. Analysis of weather conditions in those years demonstrated a strong relationship between tree-ring width and air temperature in winter, pre-spring, and early spring. Higher-than-average temperatures during those seasons correlate positively with yew tree-ring width. Response function analysis performed for local and regional chronologies point to a dominant role of air temperature in December of the year preceding growth and in January, February, and March of the current year (linear relationships). June precipitation is an additional factor affecting tree-ring width in some areas of northern and northwestern Poland: the higher the precipitation, the wider the tree-rings. The results obtained, particularly those concerning growth-climate relationships and dendroclimatic regionalisation, can be used in the on-going programme of yew restitution in Poland. • Yew trees growing in Poland, on most stands, are less than 200 years old, but their age is often overestimated. The oldest yew population in Poland is 500–600 years old. • The yew is a slow-growing species – average tree-ring width is only 0.84 mm. • Tree-ring width in yew trees depends on winter and early spring air temperature (cold winters and late onset of spring cause a decrease in tree-ring width). [ABSTRACT FROM AUTHOR]

High-resolution long-term temperature reconstructions in subtropical southeast China (SSC) are very scarce, yet indispensable for the comprehensive understanding of climate change in China, even in East Asia. We reconstructed the first previous growth-season temperature in the Sanqingshan Mountains (SQS), southeast China since 1806 based on tree-ring width data. The reconstruction accounts for 56.4 % of the total variance in the instrumental record over 1954-2009. Unlike the Northern Hemispheric warming during recent two centuries, the reconstruction captured a slowly cooling trend from 1806 to 1980, followed by a rapid warming afterward. 2003-2009 was the warmest period in the reconstruction. 1970-2000 was colder than the last stage of the Little Ice Age (LIA). Most of the warm and cold periods in this reconstruction could be found in the tree-ring based temperature reconstructions of vicinity area, indicating that the temperature variations in SSC were almost synchronous at least at decadal scale. This regional coherence of temperature variation was further confirmed by the spatial correlation patterns with the CRU TS3.22 grid dataset. A strong positive relationship between the temperature over SQS region and sea surface temperature (SST) over the North Pacific Ocean (NP) has been noted, suggesting that SST variations over NP and the related Pacific Decadal Oscillation significantly influenced the temperature variability over SSC. To better understand the climate variability during the LIA and the regional differences in temperature variations over SQS and northern Hemisphere, long data sets from more diverse areas of southern China are needed. [ABSTRACT FROM AUTHOR]

The formation of tree-rings is closely related to climate variation. This paper establishes the tree-ring chronology of Pinus massoniana for a period of 36 years and examines the relationship between the tree-ring chronology and climatic conditions based on tree-ring width samples from three sites on the northeast slopes of Yangming Mountain. This data is used to study the relationship between the tree-ring width of a young tree and climatic conditions in a moist hilly region of southern China and to understand the general pattern of climate variation and its effects on tree growth in the past in this region. The results indicate that changes in tree-ring widths in these sites are closely related to local climatic conditions. There is a significant positive correlation between the radial growth of Pinus massoniana and the mean temperature of the current year from January to May (coefficient of correlation, R, is 0.596, P<0.01) and the mean temperature of the previous year from June to July (R is 0.639, P<0.01). The radial growth of Pinus massoniana is negatively correlated with the total precipitation of the previous year from January to March and November to December (R is -0.46, P<0.05) and negatively correlated with the total precipitation of the previous year from June to July (R is -0.582, P<0.05). The effect of precipitation on the radial growth of Pinus massoniana is known as the 'hysteresis effect'. [ABSTRACT FROM AUTHOR]

A tree-ring-width chronology was developed from Pinus tabulaeformis aged up to 333 years from central Inner Mongolia, China. The chronology was significantly correlated with the local Palmer Drought Severity Index (PDSI). We therefore reconstructed the first PDSI reconstruction from March to June based on the local tree ring data from 1680 to 2012 AD. The reconstruction explained 40.7 % of the variance (39.7 % after adjusted the degrees of freedom) of the actual PDSI during the calibration period (1951-2012 AD). The reconstructed PDSI series captured the severe drought event of the late 1920s, which occurred extensively in northern China. Running variance analyses indicated that the variability of drought increased sharply after 1960, indicating more drought years, which may imply anthropogenic related global warming effects in the region. In the entire reconstruction, there were five dry periods: 1730-1814 AD, 1849-1869 AD, 1886-1942 AD (including severe drought in late 1920s), 1963-1978 AD and 2004-2007 AD; and five wet periods: 1685-1729 AD, 1815-1848 AD, 1870-1885 AD, 1943-1962 AD and 1979-2003 AD. Conditions turned dry after 2003 AD, and the PDSI from March to June (PDSI) captured many interannual extreme drought events since then, such as 2005-2008 AD. The reconstruction is comparable to other tree-ring-width-based PDSI series from the neighboring regions, indicating that our reconstruction has good regional representativeness. Significant relationships were found between our PDSI reconstruction and the solar radiation cycle and the sun spot cycle, North Atlantic Oscillation, the El Niño-Southern Oscillation, as well as the Pacific Decadal Oscillation. Power spectral analyses detected 147.0-, 128.2-, 46.5-, 6.5-, 6.3-, 2.6-, 2.2- and 2.0-year quasi-cycles in the reconstructed series. [ABSTRACT FROM AUTHOR]

The mountain forest steppe and taiga in northern Mongolia have experienced a forest decline in area and quality since the end of the last century. Changes in land use, climate, fire frequency and pest occurrence are considered to be the main drivers of this vegetation shift and desertification. Because this region is the source for major rivers, is home to a unique flora and fauna and represents an important source of timber for Mongolia, the ability of different tree species to respond to these changes and regenerate is of increasing interest. Our contribution focuses on the climate-growth relationship of old and young birch trees from two valleys in the Mongolian province of Selenge Aimag. The research site Bugant, located in the Western Khentey Mountains, was the most important logging centre in Mongolia during socialist times. Today, the vegetation is dominated by succession forests of light taiga. The research site Altansumber, on the border of the Sant and Khushat soum, is dominated by light taiga and mountain forest steppe. Traditional nomads who depend on these forests for different reasons inhabit this area. Wood cores were sampled and chronologies of young and old birch trees at Bugant and Altansumber were created. Climate data were obtained from the Eroo station, which is known in the region for its long and reliable climate record. We analysed the climate-growth relationships of the chronologies from 1962 to 2009. At both sites and in both age classes, correlations with temperature were predominantly negative, particularly in April (Bugant, south- and east-facing slopes) and May (Altansumber, north-facing slopes). Precipitation of the late summer of the previous year (August/September) positively correlated with the growth of birch at Altansumber. We assume that the significant negative correlation between winter precipitation (December/January) and the growth of old birches at both sites is due to positive effects of snow cover on the survival rate of herbivorous insect populations. Our results indicate that during the early vegetation period, younger birch trees are more dependent on water availability than older ones. Negative pointer years were characterized by below-average precipitation during the current summer period and above-average spring temperatures. For the old trees, positive pointer years were characterized by above-average summer precipitation. We conclude that water availability is the most crucial factor for the growth of white birch in northern Mongolia. [ABSTRACT FROM AUTHOR]

Millennial long tree-ring records are crucial for better understanding temperature and hydroclimatic variability over the globe. Juniper is one of particularly long-lived species, which can provide more than a thousand-year record, especially in Central Asia. However, there is a lack of dendrochronological series from the Pamir Mountains. Here we report the first 1010-year (AD 1005–2014) juniper tree-ring chronology from the mountain ranges of north-western Tajikistan, the western Pamir-Alay. We present the potential of Juniperus semiglobosa and Juniperus seravshanica in developing millennia-long records. We sampled three study sites at the elevations from 2200 to 3500 m. In general, the climate-growth analyses show that radial growth of the Himalayan pencil juniper is positively correlated with the winter precipitation and spring temperature. At some sites tree rings were also positively correlated with summer temperature. Our findings demonstrate the importance of developing the tree-ring data network for the Pamir-Alay and its potential for reconstruction of hydroclimatic variability over the last thousand years in this region. [ABSTRACT FROM AUTHOR]

High-resolution winter temperature reconstructions in China are rare, yet vital for the comprehensive understanding of past climate change. In the present work, the first winter-half year minimum mean temperature from previous November to current April in northwestern Yichang, South Central China, was reconstructed back to 1875 based on tree-ring material. The reconstruction can explain 55 % of the variance over the calibration period during 1955-2011. The temperature maintained at comparatively low level before 1958, and an abnormal warming was seen since 1959. However, the warming trend stagnated after 2000 AD. 2001-2010 was the warmest decade not only during the instrumental period but also during the whole reconstructed period. The reconstruction indicates good spatial resemblance to other temperatures series in adjacent areas and Northern Hemisphere, yet the recent warming in this study is earlier and more prominent than that of Southeast China. This work also manifests that the winter-half year minimum temperature in study area has good agreement with summer (June-September) maximum temperature variation in Southeast China at decadal scale, except that the winter-half year warming in recent decades is more evident than summer. This reconstruction is not only useful in improving our knowledge of long-term temperature variation but also useful in predicting the tree growth dynamics in the future in the study area. [ABSTRACT FROM AUTHOR]

Knowledge about tree age is critical to forestry, nature conservation and ecological studies. Direct age determination of large diameter trees through increment cores is complicated by various obstacles, primarily because of rot and insufficient borer length. Here, we aim to (1) test the accuracy of various methods (tree-ring width, basal area increment, age-size relationship) for tree age estimation, (2) select the most accurate approach and (3) enable age estimation of large individuals of Quercus robur . This was done through increment cores collected in an alluvial forest in the Czech Republic. We achieved 75 age estimates for each tree, including our novel approach, which reduces the effect of decreasing tree-ring width and increasing basal area increment during tree life. The extrapolation of mean ring width to missing radius generally overestimates the number of missing rings (by up to 27.5% of actual age) and the level of overestimation increases with decreasing partial core length, while the application of basal area increment largely underestimates the age estimation (by up to 20.5%). Thus, to eliminate the over- and underestimation caused by natural tree ring width decrease and basal area increase during the tree senescence and increasing size, we averaged the number of estimated rings by these two methods. This technique obtained the most reliable age estimates, with an error up to 3.5%. Thus it is suggested here that this technique provides a relatively accurate age estimate for trees where it is impossible to directly determine the age; at least for light-demanding species. Moreover, the proposed technique does not require complicated analysis and is not time consuming. However, future research should test the applicability of this technique for tree species with various ecological strategies, i.e. shade-tolerant species. Finally, we estimate that the age of large oaks in our study area does not exceed 400 years. Due to such fast growth, it is possible to keep the continuity of these keystone structures in a given landscape and thus preserve the associated biodiversity. [ABSTRACT FROM AUTHOR]

Tree rings reveal climatic variations through years, but also the effect of solar activity in influencing the climate on a large scale. In order to investigate the role of solar cycles on climatic variability and to analyse their influences on tree growth, we focused on tree-ring chronologies of Araucaria angustifolia and Araucaria araucana in four study areas: Irati and Curitiba in Brazil, Caviahue in Chile, and Tolhuaca in Argentina. We obtained an average tree-ring chronology of 218, 117, 439, and 849 years for these areas, respectively. Particularly, the older chronologies also included the period of the Maunder and Dalton minima. To identify periodicities and trends observable in tree growth, the time series were analysed using spectral, wavelet and cross-wavelet techniques. Analysis based on the Multitaper method of annual growth rates identified 2 cycles with periodicities of 11 (Schwebe cycle) and 5.5 years (second harmonic of Schwebe cycle). In the Chilean and Argentinian sites, significant agreement between the time series of tree rings and the 11-year solar cycle was found during the periods of maximum solar activity. Results also showed oscillation with periods of 2–7 years, probably induced by local environmental variations, and possibly also related to the El-Niño events. Moreover, the Morlet complex wavelet analysis was applied to study the most relevant variability factors affecting tree-ring time series. Finally, we applied the cross-wavelet spectral analysis to evaluate the time lags between tree-ring and sunspot-number time series, as well as for the interaction between tree rings, the Southern Oscillation Index (SOI) and temperature and precipitation. Trees sampled in Chile and Argentina showed more evident responses of fluctuations in tree-ring time series to the variations of short and long periodicities in comparison with the Brazilian ones. These results provided new evidence on the solar activity-climate pattern-tree ring connections over centuries. [ABSTRACT FROM AUTHOR]

A dendroclimatic study was conducted in the treeline ecotone of Barun Valley, eastern Nepal, to determine the tree-ring climate response and ring width trend of Abies spectabilis. A 160-year-old chronology, from 1850 to 2010, was developed from 38 tree-ring samples. No higher growth in recent decades was observed in tree-ring width in this area. The mean temperature of the current year in February and in the combined winter months of December, January, and February showed significant positive correlation with tree-ring width, although no significant correlation was found between tree-ring width and the precipitation pattern of the region. This tree-ring climate response result is different from that in other studies in Nepal, which could be attributed to location and elevation. [ABSTRACT FROM AUTHOR]

Tree-ring research in the Altai-Sayan Mountains so far only considered a limited number of well-replicated site chronologies. The dendroecological and palaeoclimatological potential and limitations of large parts of south-central Russia therefore remain rather unexplored. Here, we present a newly updated network of 13 larch ( Larix sibirica Ldb.) tree-ring width (TRW) chronologies from mid to higher elevations along a nearly 1000 km west-to-east transect across the greater Altai-Sayan region. All data were sampled between 2009 and 2014. The corresponding site chronologies cover periods from 440 to 860 years. The highest TRW agreement is found between chronologies ≥2200 m asl, whereas the material from lower elevations reveals overall less synchronized interannual to longer-term growth variability. While fluctuations in average June–July temperature predominantly contribute to the growth at higher elevations, arid air masses from Mongolia mainly affect TRW formation at lower elevations. Our results are indicative for the dendroclimatological potential of the Altai-Sayan Mountains, where both, variation in summer temperature and hydroclimate can be robustly reconstructed back in time. These findings are valid for a huge region in central Asia where reliable meteorological observations are spatially scarce and temporally restricted to the second half of the 20th century. The development of new high-resolution climate reconstruction over several centuries to millennia will further appear beneficial for timely endeavors at the interface of archaeology, climatology and history. [ABSTRACT FROM AUTHOR]

It is generally assumed in dendroecological studies that annual tree-ring growth is adequately determined by a linear function of local or regional precipitation and temperature with a set of coefficients that are temporally invariant. However, various researchers have maintained that tree-ring records are the result of multivariate, often nonlinear biological and physical processes. To describe critical processes linking climate variables with tree-ring formation, the process-based tree-ring Vaganov–Shashkin model (VS-model) was successfully used. However, the VS-model is a complex tool requiring a considerable number of model parameters that should be re-estimated for each forest stand. Here we present a new visual approach of process-based tree-ring model parameterization (the so-called VS-oscilloscope) which allows the simulation of tree-ring growth and can be easily used by researchers and students. The VS-oscilloscope was tested on tree-ring data for two species ( Larix gmelinii and Picea obovata ) growing in the permafrost zone of Central Siberia. The parameterization of the VS-model provided highly significant positive correlations ( p < 0.0001) between simulated growth curves and original tree-ring chronologies for the period 1950–2009. The model outputs have shown differences in seasonal tree-ring growth between species that were well supported by the field observations. To better understand seasonal tree-ring growth and to verify the VS-model findings, a multi-year natural field study is needed, including seasonal observation of the thermo-hydrological regime of the soil, duration and rate of tracheid development, as well as measurements of their anatomical features. [ABSTRACT FROM AUTHOR]

At present, there is limited research on the effect of climate change on the growth of trees in the Qinling Mountains, and the effect of altitude has rarely been considered. In order to accurately grasp the impact of climatic factors on the natural forest in this region, the effect of water and temperature conditions on the radial growth of Pinus armandii was studied according to dendroclimatology principles. Response function and multivariate regression methods were applied to study the response of the tree-rings, including earlywood width and latewood width, of P. armandii at three elevations (1600-1800, 1800-2000, 2000-2200 m) to climate change in this region. The results showed that the radial growth of P. armandii at the three elevations contained rich climatic information. The tree-ring widths at the three elevations were significantly negatively correlated with September temperature of the previous year and May temperature of the current year (P < 0.05) and were also negatively correlated with August precipitation of the previous year and May precipitation of the current year. The late-wood width of P. armandii at 1600-1800 m and 1800-2000 m was less sensitive to temperature and precipitation than that of the earlywood. The latewood width of P. armandii at 2000-2200 m was significantly negatively correlated with December temperature of the previous year and May temperature of the current year (P < 0.01), and was significantly and positively correlated with August precipitation of the previous year (P < 0.05). Characteristic analysis of the special years showed that large quantities of precipitation in September and October of the previous year were more likely to result in the formation of a very wide tree-ring. High spring and summer temperatures and less precipitation from April to June more likely resulted in the formation of extremely narrow tree-rings. Therefore, high precipitation in autumn of the previous year was conducive to the growth of P. armandii, whereas high temperature and drought in spring and summer inhibited the growth of P. armandii. [ABSTRACT FROM AUTHOR]

This study was conducted in a drained, exploited, and afforested Baltic bog Bagno Kusowo, located in North West Poland. The study aimed (i) to assess if human activity has a stronger impact on tree-ring width of Pinus sylvestris than climatic conditions in this transformed Baltic bog; (ii) to investigate how much the human modification of the ecosystem has influenced tree growth; (iii) to use this knowledge to reconstruct changes in the ecosystem further back in time, in the study area and its immediate neighbourhood. Wood samples for dendrochronological analyses were collected from 45 trees. Next, using classic dating methods and standard procedures (cross-dating methods, COFECHA program), chronologies were constructed (raw tree-ring width and residual chronologies: de-trended, autocorrelation removed, ARSTAN program). They formed a basis for further analyses: signature years, correlation and response function, as well as percentage growth change. The results of dendroclimatological analyses show weak increment-climate relationships and the analysis of weather conditions in the identified signature years did not detect any unambiguous relations with tree-ring width. However, results of the analyses indicate that the dominant factors affecting tree growth dynamics in the bog are changes in the hydrological system. Moreover, our results show many phases of human impact on environmental changes. Dendrochronological methods, combined with an analysis of old maps and other historical records, allowed us to reconstruct transformations of the ecosystem with a high resolution. [ABSTRACT FROM AUTHOR]

We aim to achieve a mechanistic understanding of the eco-physiological processes in Larix decidua and Pinus mugo var. uncinata growing on north- and south-facing aspects in the Swiss National Park in order to distinguish the short- and long-term effects of a changing climate. To strengthen the interpretation of the δ18O signal in tree rings and its coherence with the main factors and processes driving evaporative δ18O needle water enrichment, we analyzed the δ18O in needle, xylem and soil water over the growing season in 2013 and applied the mechanistic Craig–Gordon model (1965) for the short-term responses. We found that δ18O needle water strongly reflected the variability of relative humidity mainly for larch, while only δ18O in pine xylem water showed a strong link to δ18O in precipitation. Larger differences in offsets between modeled and measured δ18O needle water for both species from the south-facing aspects were detected, which could be explained by the high transpiration rates. Different soil water and needle water responses for the two species indicate different water-use strategies, further modulated by the site conditions. To reveal the long-term physiological response of the studied trees to recent and past climate changes, we analyzed δ13C and δ18O in wood chronologies from 1900 to 2013. Summer temperatures as well as summer and annual amount of precipitations are important factors for growth of both studied species from both aspects. However, mountain pine trees reduced sensitivity to temperature changes, while precipitation changes come to play an important role for the period from 1980 to 2013. Intrinsic water-use efficiency (WUEi) calculated for larch trees since the 1990s reached a saturation point at elevated CO2. Divergent trends between pine WUEi and δ18O are most likely indicative of a decline of mountain pine trees and are also reflected in decoupling mechanisms in the isotope signals between needles and tree-rings. [ABSTRACT FROM AUTHOR]

This study investigated the seasonal change in xylem growth of Japanese red pine ( Pinus densiflora). Wood cores were sampled at 2-week intervals from April to November in 2012 using the microcoring method. Daily increment rates of tracheid number and tree-ring width were compared with seasonal changes in daily mean temperature and photoperiod. Xylem growth started in early to late May and stopped in late October to early November. The maximum daily increment rates of tracheid number and tree-ring width were in early July. The 95 % confidence intervals of the timing of the maximum daily increment rates included the summer solstice (23 June) with the longest photoperiod, but not the warmest day (30 July). The maximum daily increment rate of xylem growth is thought to be controlled by the photoperiod rather than by temperature. The daily mean temperature exceeded 20 °C after the summer solstice, indicating that temperature is not a limiting factor for xylem growth. This study suggests that the timing of maximum daily increment rates of xylem growth of P. densiflora is controlled by the photoperiod. [ABSTRACT FROM AUTHOR]

Age-related alternation in the sensitivity of tree-ring width (TRW) to climate variability has been reported for different forest species and environments. The resulting growth-climate response patterns are, however, often inconsistent and similar assessments using maximum latewood density (MXD) are still missing. Here, we analyze climate signal age effects (CSAE, age-related changes in the climate sensitivity of tree growth) in a newly aggregated network of 692 Pinus sylvestris L. TRW and MXD series from northern Fennoscandia. Although summer temperature sensitivity of TRW ( r All = 0.48) ranges below that of MXD ( r All = 0.76), it declines for both parameters as cambial age increases. Assessment of CSAE for individual series further reveals decreasing correlation values as a function of time. This declining signal strength remains temporally robust and negative for MXD, while age-related trends in TRW exhibit resilient meanderings of positive and negative trends. Although CSAE are significant and temporally variable in both tree-ring parameters, MXD is more suitable for the development of climate reconstructions. Our results indicate that sampling of young and old trees, and testing for CSAE, should become routine for TRW and MXD data prior to any paleoclimatic endeavor. [ABSTRACT FROM AUTHOR]

Tree-ring based climate reconstructions have been critical for understanding past variability and recent trends in climate worldwide, but they are scarce in Australia. This is particularly the case for temperature: only one tree-ring width based temperature reconstruction – based on Huon Pine trees from Mt Read, Tasmania – exists for Australia. Here, we investigate whether additional tree-ring parameters derived from Athrotaxis cupressoides trees growing in the same region have potential to provide robust proxy records of past temperature variability. We measured wood properties, including tree-ring width (TRW), mean density, mean cell wall thickness (CWT), and tracheid radial diameter (TRD) of annual growth rings in Athrotaxis cupressoides , a long-lived, high-elevation conifer in central Tasmania, Australia. Mean density and CWT were strongly and negatively correlated with summer temperatures. In contrast, the summer temperature signal in TRW was weakly positive. The strongest climate signal in any of the tree-ring parameters was maximum temperature in January (mid-summer; JanT max ) and we chose this as the target climate variable for reconstruction. The model that explained most of the variance in JanT max was based on TRW and mean density as predictors. TRW and mean density provided complementary proxies with mean density showing greater high-frequency (inter-annual to multi-year) variability and TRW showing more low-frequency (decadal to centennial-scale) variability. The final reconstruction model is robust, explaining 55% of the variance in JanT max , and was used to reconstruct JanT max for the last five centuries (1530–2010 C.E.). The reconstruction suggests that the most recent 60 years have been warmer than average in the context of the last ca. 500 years. This unusually warm period is likely linked to a coincident increase in the intensity of the subtropical ridge and dominance of the positive phase of the Southern Annular Mode in summer, which weaken the influence of the band of prevailing westerly winds and storms on Tasmanian climate. Our findings indicate that wood properties, such as mean density, are likely to provide significant contributions toward the development of robust climate reconstructions in the Southern Hemisphere and thus toward an improved understanding of past climate in Australasia. [ABSTRACT FROM AUTHOR]

Key message : Intensity and magnitude of the growth-climate relationship depends on juniper species and sites. Juniperus seravschanica at low elevations shows highest potential for April-September drought reconstruction in the Turkestan range (Pamir-Alay), Uzbekistan. Abstract: We present a detailed dendroclimatological study of three juniper species, Juniperus seravschanica Kom. , Juniperus semiglobosa Regel and Juniperus turkistanica Kom., sampled at six sites of different elevation (2100-2700 m a.s.l.), exposition (west and south) and steepness (10°-30°) in the Zaamin National Park, Turkestan range, Pamir-Alay mountain system in eastern Uzbekistan. Simple correlation statistics and redundancy analyses were applied to detect species- and site-specific climate responses during the twentieth century, which were additionally investigated in the high-frequency domain by identifying extreme growth years. Our results show that tree-ring formation of J. seravschanica at our low-elevation site is strongly limited by April to September drought conditions, while J. semiglobosa inherits a weak and variable climate response with respect to elevation. J. turkistanica growth at high altitudes is positively associated with warm spring and summer temperatures. Species-specific growth extremes are triggered by incoming air masses from the Atlantic and Arctic, highlighting the connection of synoptic climate regimes across Eurasia. From a dendroclimatic perspective, J. seravschanica exhibits a high potential for reconstructing past drought and pluvials, but under sustained temperature rise also J. semiglobosa will likely increase its sensitivity to drought. Moreover, J. turkistanica at its distribution limit at the tree line is a suitable proxy of summer temperature. Our findings clearly demonstrate that a careful selection of the site, overall topography and elevation as well as the different juniper species are important for successfully reconstructing past climate in Uzbekistan. [ABSTRACT FROM AUTHOR]

Changing climate is expected to cause range shifts and reduced growth in Norway spruce ( Picea abies (L.) Karst). In order to mitigate these changes, genetic variation between populations can be utilized in selecting alternative tree origins that are better suited to the new conditions. The aim of this study was to examine the intraspecific differences in the climatic drivers of radial growth in Norway spruce. We used tree-ring data from seven Norway spruce provenance experiments in Finland, located in different climatic conditions and including a large variety of provenances. The annual ring-width indices were studied with hierarchical clustering, correlation analysis with climate variables, pointer year analysis and linear models to identify the provenance differences in growth variation and its climatic control, and compare them on a latitudinal gradient. The cluster analysis revealed patterns of provenance differences in growth variation: north European and central European provenances were grouped in separate clusters within sites, although with some exceptions. Largest provenance differences in climate-growth responses were found in relation to winter and spring temperatures. In the southern provenances warm winters were typically associated with faster growth whereas for the northern provenances the correlations varied from non-significant to negative. In addition, the pointer year analysis showed negative growth anomalies only in the southern provenances for years with exceptionally cold winters. These patterns may reflect the physiological differences between the provenances relating to, for example, cold tolerance and the timing of spring phenology. As the climate warming in Europe is predicted to be strongest during the winter months, acknowledging the intraspecific growth responses to climate in Norway spruce becomes increasingly important. [ABSTRACT FROM AUTHOR]

Drought is a key factor affecting forest ecosystem processes at different spatio-temporal scales. For accurately modeling tree functioning – and thus for producing reliable simulations of forest dynamics – the consideration of the variability in the timing and extent of drought effects on tree growth is essential, particularly in strongly seasonal climates such as in the Mediterranean area. Yet, most dynamic vegetation models (DVMs) do not include this intra-annual variability of drought effects on tree growth. We present a novel approach for linking tree-ring data to drought simulations in DVMs. A modified forward model of tree-ring width (VS-Lite) was used to estimate seasonal- and site-specific growth responses to drought of Scots pine ( Pinus sylvestris L.), which were subsequently implemented in the DVM ForClim. Ring-width data from sixteen sites along a moisture gradient from Central Spain to the Swiss Alps, including the dry inner Alpine valleys, were used to calibrate the forward ring-width model, and inventory data from managed Scots pine stands were used to evaluate ForClim performance. The modified VS-Lite accurately estimated the year-to-year variability in ring-width indices and produced realistic intra-annual growth responses to soil drought, showing a stronger relationship between growth and drought in spring than in the other seasons and thus capturing the strategy of Scots pine to cope with drought. The ForClim version including seasonal variability in growth responses to drought showed improved predictions of stand basal area and stem number, indicating the need to consider intra-annual differences in climate–growth relationships in DVMs when simulating forest dynamics. Forward modeling of ring-width growth may be a powerful tool to calibrate growth functions in DVMs that aim to simulate forest properties in across multiple environments at large spatial scales. [ABSTRACT FROM AUTHOR]