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Teleostean evolution produced enormous variation in tooth morphology. This variation is related tothe exploration and exploitation of very diverse dietary niches. Among extant fishes, several taxa of the order Tetraodontiformes (e.g. pufferfishes, porcupinefishes and ocean sunfishes) have evolved highly specialized beak-like tooth structures. Here we provide the first description of tetraodontiform crushing beaks from Argentina. A single complete premaxillary beak recovered from the early Miocene Gaiman Formation in Chubut Province is assigned to Molidae based on its premaxillaries being completely fused to one another along the midline, general morphology, and size. Moreover, due to the putative presence of rows of teeth fused to the thick and massive bone structure, the specimen might belong to the genus Ranzania Nardo, 1840. Several premaxillary beaks recovered in the late Miocene Paraná Formation of Entre Ríos Province are assigned to Tetraodontidae Bonaparte, 1832 based on the presence of long, slender, rod-like, and parallel dental units; premaxillae not fused in the midline but articulated by interlocking emarginations, and a few small and one large trituration dental units. Argentinian tetraodontiform fishes inhabited the warm-temperate Paranense and Patagoniense Seas (marine transgressions developed in southeastern South America) during the early-late Miocene; these specimens are the southernmost fossil record of the group worldwide. Tetraodontiforms are extremely rare in recent Argentinian marine waters. The new fossil records agree with the higher sea water surface temperatures suggested both locally and worldwide for the Miocene.

We describe isolated teleostean teeth found in no association with the jaw bone. The specimens have been recovered in Late Cretaceous marine deposits of the Vivian Formation in the Peruvian Sub-Andean Region. The deposition sequence from where the teeth come is interpreted as a shallowing-upward sequence of low salinity. The fish material is identified as Enchodus aff. E. gladiolus based on the presence of a small but well-developed post-apical barb, an anterior cutting edge, the crown is symmetrical in cross-section, have a sigmoidal profile, and bears strong ridges (=striations). The Peruvian material differs from the typical E. gladiolus teeth in having a faintly serrated anterior cutting edge which is absent in most specimens referred to E. gladiolus. We also highlight that taxonomic assignments made based on isolated teeth must be taken with care. Despite scarce, the material recovered denotes that the marine units of Peru can give valuable information about the Pacific fish fauna during the Late Cretaceous.

Although the Cretaceous is characterized by a rich fish diversity, Cretaceous continental fishes from Gondwana are poorly known and comparatively scarce. Among these fishes, the family Pleuropholidae is only known by a few species relatively poorly preserved, from the Middle Jurassic to Lower Cretaceous of Europe, Africa, North America, and South America. In this paper, two new species of the pleuropholid new genus Zurupleuropholis are described, Z. quijadensis gen. et sp. nov. and Z. decollavi gen. et sp. nov. The new fishes were recovered in the Lower Cretaceous lacustrine Lagarcito Formation of central-west Argentina. This taxon constitutes a relevant finding considering that the representation of the family Pleuropholidae is rare worldwide. Zurupleuropholis gen. nov. appears to be the youngest known member of Pleuropholidae, and it represents the second record of the family in South America and the first record in the Cretaceous of the continent. Fil: Giordano, Paula Guillermina. Universidad Nacional de San Luis. Facultad de Ciencias Físico Matemáticas y Naturales. Departamento de Geología; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Succar, Cecilia Andrea. Boston College; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Codorniú Dominguez, Laura Susana. Universidad Nacional de San Luis. Facultad de Ciencias Físico Matemáticas y Naturales. Departamento de Geología; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Cione, Alberto Luis. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Arratia, Gloria. University of Kansas; Estados Unidos

The material reported herein comes from the Lower Cretaceous Agrio Formation (Neuquén Basin, west-central Argentina). These fishes dwelled in a Pacific Ocean embayment in southwestern Gondwana before the establishment of the Andean Cordillera. The material consists of an incomplete dorsal fin spine of the widespread Hybodontoidea clade (from upper Valanginian beds) and a partial tooth of one of the earliest known lamniforms (from upper Hauterivian beds)., Facultad de Ciencias Naturales y Museo

The family Sciaenidae comprises about 300 species. The black drum Pogonias cromis was the only valid species of the genus. Herein, Pogonias courbina Lacepède 1803 is rede-scribed based on morphological and molecular evidence and a neotype is designated. Pogonias courbina is distinguished by the following characters: the occurrence of characteristic thickening of the dorsal spines VII to XI in all specimens larger than 250 mm SL; all pterygiophores in the dorsal-fin laminar, thin; anal-fin pterygiophores slender excluded those of spines; lateral projections of gas bladder with few finger-like projections; genetic distance between both species 1%; exclusive occurrence of characters in six informative sites of COI (58 G; 214 G; 328 A; 331 A; 553C; 580 G). The method Automatic Barcode gap Discovery detected gaps in nucleotid distance congruent with the NJ, MP, and ML tree analysis. Also, advertisement calls are three times shorter in duration in P. courbina than in P. cromis. In addition, two monophyletic groups for P. cromis and P. courbina appear in trees obtained with different methodologies, emphasizing the absence of shared haplotypes. A gap of about 8000 km occurs in the distribution of both species along coastal areas of the Atlantic Ocean. Fil: Azpelicueta, María de las Mercedes. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. Área Zoología; Argentina Fil: Delpiani, Sergio Matías. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Mar del Plata. Instituto de Investigaciones Marinas y Costeras. Universidad Nacional de Mar del Plata. Facultad de Ciencias Exactas y Naturales. Instituto de Investigaciones Marinas y Costeras; Argentina Fil: Cione, Alberto Luis. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. Departamento Paleontología; Argentina Fil: Oliveira, Claudio. Universidade Estadual Paulista Julio de Mesquita Filho; Brasil Fil: Marceniuk, Alexandre Pires. Universidade Estadual Paulista Julio de Mesquita Filho; Brasil Fil: Díaz de Astarloa, Juan Martín. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Mar del Plata. Instituto de Investigaciones Marinas y Costeras. Universidad Nacional de Mar del Plata. Facultad de Ciencias Exactas y Naturales. Instituto de Investigaciones Marinas y Costeras; Argentina

Catfish bones from Tortonian (Miocene) freshwater beds of central Argentina are here identified as pertaining to a new species of the tropical pimelodid genus Phractocephalus. The new species differs from the other recent and fossil species of the genus in skull, pectoral girdle and spine characters. The material was found in different localities near the city of Paraná, Entre Ríos Province. The bearing horizon is the so-called “Conglomerado osífero” which constitutes the lowermost beds of the fluvial Ituzaing o Formation. The aquatic vertebrate fauna occurring in the bearing bed shows a similar generic composition to several northern South American Miocene units where Phractocephalus remains were found. This report extends the range of Phractocephalus more than 2000 km to the South. The record is in agreement with higher global temperatures and putative ample hydrographic connections of the river basins in the Paran a area with the Amazon basin until at least the early late Miocene., Facultad de Ciencias Naturales y Museo

The Upper Cretaceous record of lungfishes from South America is abundant but represented mainly by tooth plates and associated jaw bones. A new dipnoan species from the Upper Cretaceous of the La C...

An ichthyofauna recovered from the L opez de Bertodano Formation at Units 9 (uppermost Maastrichtian) and 10 (lowermost Danian) and the Sobral Formation (Danian), in Marambio (Seymour) Island in the James Ross Basin is described and analyzed herein. A review of previously described taxa based on the new material and several new fish records for the L opez de Bertodano Formation is provided, including the youngest record of Enchodus and the first Cretaceous evidence of Pachycormiformes.We also identify the first Paleocene fishes for the continent. The new information and the reinterpretation of previous Cretaceous records show that there was no decline in fish diversity until the CretaceousePaleogene (K/ Pg) boundary in the area and extinction event appears to have been rapid. Finally, we find that the distribution of some chondrichthyans and teleosts in the James Ross Basin appears to be climatically determined., Facultad de Ciencias Naturales y Museo, Consejo Nacional de Investigaciones Científicas y Técnicas

The marine deposits of the Vaca Muerta Formation (Tithonian–Berriasian) houses one of the most diverse Late Jurassic ichthyofaunas of Gondwana. However, most of the specimens remain undescribed. Jurassic fishes have been recovered from several localities at Neuquen Province (i.e., Picun Leufu, Plaza Huincul, Cerro Lotena, Portada Las Lajas, Los Catutos, and Arroyo Covunco) but also from Mendoza Province (i.e., La Valenciana, Los Molles, and Arroyo del Cajon Grande). Presently, the fish fauna of Los Catutos, near Zapala city (Neuquen Province), has yielded the highest number of specimens, which are taxonomically and morphologically diverse. At Los Catutos locality, the Vaca Muerta Formation is represented by the Los Catutos Member, which is considered the only lithographic limestones known in the Southern Hemisphere. Here, we review the Tithonian fish faunas from the Vaca Muerta Formation. During Late Jurassic times, the actual Argentinian territory could have been a morphological diversification center, at least for some actinopterygian groups. The apparently lower species diversity recorded in marine Jurassic ichthyofaunas of Argentina (and some Gondwanan countries) in comparison with Chilean and European fish faunas could be related to the fish paleontological research history in Gondwana and the low number of detailed studies of most of specimens recorded.

A specimen of a remarkable new catfish genus and species was collected in middle/late Miocene marine beds of the Puerto Madryn Formation at the base of the marine cliff of the sea lion colony area near Puerto Pirámide, southern coast of Península Valdés, northeastern Patagonia, Argentina. Siluriforms (catfishes) constitute a most important monophyletic ostariophysan group of mainly freshwater fishes that occurs in almost all continents but it is especially diverse in South America. Catfishes are presently distributed in tropical to temperate areas and a small number of species are marine or amphibiotic. The new catfish shows many primitive features for catfishes in the maxilla, autopalatine, hyal elements, and Weberian apparatus. The genus is clearly distinguished by four autapomorphies: sand clock–shaped autopalatine, posterior limb of autopalatine widening strongly, post-articular arm of autopalatine longer, and a metapterygoid longer than broad. One tree was obtained both under equal and implied weighting with the following topology: a basal polytomy in the Siluriformes formed by Diplomystidae, Bachmanniidae, Kooiichthys and the Siluroidei. The new species appears to have been a marine or amphibiotic taxon: it was collected in beds considered to represent the Maximum Flooding Horizon of the transgression that deposited the Puerto Madryn Formation. The coast at this moment was at approximately 90 km to the west. According to faunistic evidence, the sea was warm temperate.

Vallone, E.R., Vezzosi, R.I. & Cione, A.L. February 2017. First fossil fish (Teleostei, Siluriformes) from the Late Pleistocene of Santa Fe Province, Argentina. Alcheringa 00, 000–000. ISSN 0311-5518. The record of fossil fish from the Pleistocene of Argentina is poor. Here we describe the first ichthyofauna from Late Pleistocene riverbank beds in the Salado River of Santa Fe Province, Argentina. The material consists of isolated pectoral and dorsal fin spines, together with skull fragments. Four species-level taxa referable to three families can be identified: Pterodoras granulosus (Doradidae), Pimelodus cf. maculatus and Pimelodus cf. albicans (Pimelodidae) and cf. Hypostomus sp. (Loricariidae). Specimens attributed to Pterodoras granulosus and Pimelodus maculatus represent a minimum age for origin of these taxa. The Salado River assemblage includes the richest record of Pleistocene catfishes yet documented from southern South America. Evelyn Romina Vallone [evelynvallone@conicet.gov.ar] and Raúl Ignacio Vezzosi [vezzosiraul@gmail.com], Laboratorio de Paleontología de Vertebrados, CICYTTP-CONICET, Materi y España, (3105) Diamante, Entre Ríos, Argentina; Alberto Luis Cione [acione@museo.fcnym.unlp.edu.ar], División Paleontología de Vertebrados. Museo de La Plata. Paseo del Bosque s/n, (1900) La Plata, Argentina. Fil: Vallone, Evelyn Romina. Provincia de Entre Ríos. Centro de Investigaciones Científicas y Transferencia de Tecnología a la Producción. Universidad Autónoma de Entre Ríos. Centro de Investigaciones Científicas y Transferencia de Tecnología a la Producción. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Santa Fe. Centro de Investigaciones Científicas y Transferencia de Tecnología a la Producción; Argentina Fil: Vezzosi, Raúl Ignacio. Provincia de Entre Ríos. Centro de Investigaciones Científicas y Transferencia de Tecnología a la Producción. Universidad Autónoma de Entre Ríos. Centro de Investigaciones Científicas y Transferencia de Tecnología a la Producción. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Santa Fe. Centro de Investigaciones Científicas y Transferencia de Tecnología a la Producción; Argentina Fil: Cione, Alberto Luis. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina

Isolated teeth of a new species of the rare batomorph genus Hypolophodon were found in Paleocene beds of the Lefipan Formation (Maastrichtian-Paleocene) of southern Argentina. The new species H. patagoniensis differs from the type species H. sylvestris in the absence of an occlusal tranverse crest on the crown of unworn teeth and of a broad and rounded central uvula on the lingual face of the crown; and in the presence of a lower boundary of the enameloid smooth on all crown faces, many expanded basoapically striae on all sides of the root, and a deep root. It differs from the Eocene species H. dockery in the larger size, the less widely separated root branches, the presence of root foraminae or striae, the absence of uvula, and the deeper crown. The new species material, the two teeth from Jaguel Formation, and an indeterminate tooth of Hypolophodon from the Lower Cenozoic of Chile represent the sole records of the genus in the Southern Hemisphere. The shark fauna helped to date the bearing beds.

The formation of the Isthmus of Panama stands as one of the greatest natural events of the Cenozoic, driving profound biotic transformations on land and in the oceans. Some recent studies suggest that the Isthmus formed manymillions of years earlier than the widely recognized age of approximately 3 million years ago (Ma), a result that if true would revolutionize our understanding of environmental, ecological, and evolutionary change across the Americas. To bring clarity to the question of when the Isthmus of Panama formed, we provide an exhaustive review and reanalysis of geological, paleontological, and molecular records. These independent lines of evidence converge upon a cohesive narrative of gradually emerging land and constricting seaways,withformationof theIsthmus of Panama sensustricto around 2.8 Ma. The evidence used to support an older isthmus is inconclusive, and we caution against the uncritical acceptance of an isthmus before the Pliocene., Facultad de Ciencias Naturales y Museo

A penguin humerus constitutes the most recent record of the widely distributed genus Palaeospheniscus in the Atlantic Ocean. Penguins are unknown in late Miocene and Plicene beds in the region, whereas they have been reported from the occidental coasts of the continent. The gap in the record is occasioned mainly by the dearth of beds of this age. The present finding confirms that a high diversity still existed in the middle Miocene. Fil: Acosta Hospitaleche, Carolina Ileana Alicia. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina Fil: Cione, Alberto Luis. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina

Metaceratodus wollastoni, an Australian species, was reported from Late Cretaceous beds of Patagonia in 1997. Later, three new species (Ceratodus wichmanni, Ptychoceratodus kaopen and Ptychoceratodus cionei), based on scarce material, were described. Finally, two of these species were referred to genus Ferganoceratodus. After examining much more abundant and better-preserved material, we suggest that neither the occurrence of the species Metaceratodus wollastoni nor that of genera Ptychoceratodus and Ferganoceratodus in the Cretaceous of South America is supported. In this paper, we consider that C. wichmanni and P. cionei are synonyms and we reassign the three putative species to genus Metaceratodus under two new combinations: M. kaopen comb. nov. and M. wichmanni comb. nov. Both differ from the other species of the genus because specimens have occlusal pits over most of the occlusal surface and a different tooth plate occlusal profile, and most have four ridges in the lower and upper tooth plates. M. wichmanni differs from M. kaopen in several features. M. kaopen is known from the late Santonian–early Campanian of Anacleto Formation at Río Negro province and M. wichmanni from late Campanian–early Maastrichtian units of Chubut, Río Negro, Neuquén and Mendoza provinces, Argentina. The occurrence of Metaceratodus in southern South America corroborates a close biogeographical relationship with Australia in the latest Cretaceous. Fil: Cione, Alberto Luis. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Gouiric Cavalli, Soledad. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina

Lamnid teeth close to but more plesiomorph than Carcharodon carcharias were collected in the late Miocene beds of the Parana Formation in the central eastern Argentina. We propose, as other authors, that some species formerly assigned to Isurus or to Cosmopolitodus should be included in Carcharodon. Some workers suggested that C. carcharias originated by phyletic evolution in the Pacific basin. The teeth from Parana could pertain to a new species of Carcharodon already known from Peru. In contrast with the Pacific basin, Carcharodon sp. was sympatric with the wide toothed species Carcharodon plicatilis in central Argentina. This is the oldest record of Carcharodon in the southern Atlantic.

Fil: Perez, Leandro Martin. Consejo Nacional de Investigaciones Cientificas y Tecnicas; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. Division Paleozoologia Invertebrados; Argentina

The first lungfish tooth plate from the Las Flores Formation, Chubut, southern Argentina, is described. This is the youngest ceratodontid known from the continent. In Africa, ceratodonts disappeared in the Eocene. Afterwards, they are only known from Australia until their extinction during the Pleistocene. The Las Flores tooth plate also represents the southernmost lungfish known since the Coniacian (early Late Cretaceous).

Two biogeographical units are generally recognized in the present shelf area of Argentina: the Magellanian and Argentinian Provinces. The two provinces differ in their fossil record. The evolution of these provinces has been characterized by migrations, extinctions, pseudoextinctions and, perhaps, even speciation events. Marine vertebrate assemblages with some similarities to the Argentinian fauna were already present in the Miocene, whereas no associations similar to those of the Magellanian fauna have been found in South America before the Pleistocene. Two successive major marine transgressions flooded northern Patagonia during the Miocene: the ‘Patagoniense’ (Early Miocene) and the ‘Entrerriense’ (Middle to Late Miocene). We analyse three rich fossil assemblages that were formed during these transgressions. The absence of Magellanian Miocene vertebrate assemblages is consistent with the hypothesis of a more southern distribution of the cold-temperate fauna at that time. In Patagonia, as in other regions, an increased number of living groups appeared from the Lower to Upper Miocene. The Late Miocene aquatic mammals had a modern aspect, and some of the fish species are still living in the South Atlantic Ocean. In this contribution, we stress that warm-temperate fishes and a high diversity of penguins are found together at the base of the Gaiman Formation. We hypothesize that penguins were adapted to live in warmer waters than those of the latest Cenozoic and the Recent. Finally, we recall that many taxa became extirpated because of the global temperature drops of the late Cenozoic. However, some fishes and pinnipeds which were extirpated only in the Atlantic Ocean are discussed here. © 2011 The Linnean Society of London, Biological Journal of the Linnean Society, 2011, 103, 423–440.

The Siluriformes constitutes an important monophyletic ostariophysan group of mainly freshwater fishes that occurs in almost all continents. They are extremely abundant and diverse in South America, where the oldest otophysans and siluriforms occur. We restudied two putative catfish species from the early Eocene lacustrine Tufolitas Laguna del Hunco of northwestern Patagonia, Argentina, concluding that Arius argentinus Dolgopol, 1941, is a junior synonym of Bachmannia chubutensis Dolgopol, 1941. Phylogenetic analyses were performed to show the relationships of B. chubutensis, which appears as the sister group of Diplomystes. Bachmannia chubutensis is one of the most primitive catfishes known and we based on it a new family. The family is clearly diagnosed by the presence of a anterior ventrolateral expansion of cleithrum, short maxilla with two rows of teeth, double-headed autopalatine, accessory tooth plates, very huge teeth, supraoccipital process long and acute, first and fifth vertebrae not j...

Different traces occur on fossil bones and teeth coming from the Early Miocene Gaiman Formation (Patagonia, Argentina). Most traces were attributed to the action of terrestrial and marine predators and scavengers. However, other traces on bones and teeth from this unit and one tooth from the Eocene La Meseta Formation (Antarctica) are attributed to chemical corrosion by lichens in recent times, that is, in a very late diagenetic time. The living lichens and calcium oxalate deposits occurring on the traces and their particular pattern indicates that they were not produced by vegetal roots. The lichens include reproductive structures which allowed a proper determination. A kind of corrosion pattern (Type 1) on bones and teeth from Patagonia is associated to Sarcogyne orbicularis K̈orber, Verrucaria sp. Schrad, and Buellia aff. punctiformis (Hoff.) Massal. The lichen Aspicilia aff. Aquatica produced rounded holes on an Antarctic tooth (Type 2). On the same tooth, the epilithic lichen Caloplaca sp. Th. Fries did not leave any kind of mark on the enameloid. Fil: Acosta Hospitaleche, Carolina Ileana Alicia. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. Departamento Científico de Paleontología de Vertebrados; Argentina Fil: Marquez, Gonzalo Javier. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. Cátedra de Palinología; Argentina Fil: Pérez, Leandro Martín. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. División Paleozoología Invertebrados; Argentina Fil: Rosato, Vilma Gabriela. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. Instituto de Botánica Spegazzini; Argentina Fil: Cione, Alberto Luis. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - La Plata; Argentina. Universidad Nacional de La Plata. Facultad de Ciencias Naturales y Museo. Departamento Científico de Paleontología de Vertebrados; Argentina

Fil: Scanferla, Carlos Agustin. Consejo Nacional de Investigaciones Cientificas y Tecnicas. Oficina de Coordinacion Administrativa Parque Centenario. Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”; Argentina

Several traces of biological interaction were found on penguin bones from the basal levels (Aquitanian) of the Miocene Gaiman Formation in the lower Chubut valley of the Provincia del Chubut, Argentina. The fossil-bearing beds were deposited in littoral to sublittoral environments within sediments of mostly pyroclastic origin. We interpret many traces to have been produced by predators and/or scavengers while the penguins were still in a breeding area. Many bones show cracking marks due to aerial exposure. The material is disarticulated as is usual in recent breeding areas. Potential predators were coeval terrestrial mammals, most probably marsupial carnivores. After a marine transgression, these bones were buried or exposed on the sea bottom where they could be colonized by algae, sponges, cnidarians, and other benthic organisms. We identified sponge borings in several bones. Other traces are interpreted to have been produced by echinoderms feeding on sponges or algae. No evidence of other invertebrate p...

Isolated shark and actinopterygian teeth and scales were found in marine levels of the El Jarillal Formation (Early Permian), which crops out near the Cerro Vizcacha, Provincia de Mendoza, Argentina. Scarce previous Paleozoic vertebrate remains are known from Argentina and none was described from the Permian. Only tetrapod traces were reported before. However, vertebrates (especially fishes) are relatively abundant in other South American countries such as Brazil, Bolivia, Uruguay, Colombia, and Venezuela, from where “agnathan”, placoderm, acanthodian, chondrichthyan, actinopterygian, dipnoan, crossopterygian, and tetrapod remains are known. In this paper we report the southernmost Paleozoic vertebrates body remains record and the first description of a hybodontid tooth from the Paleozoic of South America. Hybodontiforms were previously known in South America by fin spines, placoid scales and one putative undescribed tooth. We also give the first report South American shark dermal denticles traditionally ascribed to the form genus Petrodus , which probably belongs to a hybodontoid shark. Finally, several “basal actinopterygians” scales and teeth are assigned to morphotypes found in other South American Permian Formations. We did not found other fish taxa already known from late Paleozoic beds of the continent such as xenacanthiform sharks, lungfishes or coelacanthiforms. As an appendix, we summarize the Permian fish record of the continent.

Megapiranha paranensis from the Upper Miocene of Argentina is described based on a large, partially toothed premaxilla as a new genus and species of serrasalmid fish (pacus and piranhas) and is diagnosed and distinguished from other serrasalmids based on the following unique combination of characters: seven premaxillary teeth with the first four arranged in a shallow, zig-zag row, and third tooth shaped similarly to the fourth and fifth teeth; large, triangular, unicuspid crowns with finely serrated cutting edges. The phylogenetic position of Megapiranha was determined by parsimony analysis of morphological characters. The resulting analysis recovered Megapiranha as sister to the piranha clade (Pygopristis, Pygocentrus, Pristobrycon, Serrasalmus) and is supported by two synapomorphies: (1) teeth triangular in labial view with well-developed cutting edges, and (2) serrations along both sides of tooth cutting edges. The pattern of tooth placement exhibited by the fossil Megapiranha is intermediate between the double-row condition of pacus and the single-row condition of piranhas, and suggests how the double row of teeth may have been rearranged into a single row in the evolution of piranhas.

The Great American Biotic Interchange (GABI) between North and South America is one of the most important events in the history of land mammals. The interchange occurred in several phases during more than nine million years. We here analyze the chronology and dynamics of the GABI, the evolution of some South American mammalian groups through time, and the Quaternary mammalian extinctions. As the GABI was a complex process, we divide it into ProtoGABI and GABI 1 to 4. In our concept, the extinction of the megafauna by the gatherer/hunters that entered South American during the latest Pleistocene is a part of the GABI. The putative scarce frequency of extinct mammals in archeological sites is discussed. The evolutive relevance of the GABI is reflected in that half of the species living in South America had a North American ancestry. A final process, not included in the GABI, is the remarkable alteration of ecosystems by modern man. Presently, the composition and distribution of almost all autochthonous land mammal faunas are changing dramatically. Moreover, frequently they are replaced with domesticated and/or wild exotic species. Facultad de Ciencias Naturales y Museo

The biotic interchange between the Americas occurred in several phases during more than nine million years. In this book, we focus on mammals of southern South America, where the most important and richest localities with fossil vertebrates of Late Miocene–Holocene Age were reported. We here provide basic information about taxonomic composition, biostratigraphy, climate evolution, continental tectonics, and biogeography for better understanding the GABI. Furthermore, we analyze the chronology and dynamics of the GABI, the evolution of some South American mammalian groups through time, and the Quaternary mammalian extinctions. As the GABI was a complex process, we divide it into ProtoGABI and GABI 1 to 4.

Jurassic fishes from the Southern Hemisphere are poorly known in comparison with those of the Northern Hemisphere. The Tithonian (Late Jurassic) deposits of Argentina have yielded a rich and taxonomically diverse ichthyofauna. However, these fishes have been only partially described and need to be revised or studied. In this paper, the putative ichthyodectiform actinopterygian Notodectes argentinus is redescribed. The holotype comes from the Vaca Muerta Formation, in the southwestern part of Mendoza Province, Argentina. It consists of an incomplete, partially articulated skull in which some of the bones are preserved in three dimensions. The specimen has a unique character combination that allows us not only to diagnose the genus Notodectes but also confirm that it is a member of the actinopterygian order Pachycormiformes. At present, Notodectes represents the first endemic pachycormiform from the Southern Hemisphere and the first specimen found in Mendoza Province (northwestern region of the Neuqu en Basin). Also, Notodectes is the first fastswimming and clearly ichthyophagous pachycormid reported from the Jurassic of South America. Fil: Gouiric Cavalli, Soledad. Universidad Nacional de la Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina Fil: Cione, Alberto Luis. Universidad Nacional de la Plata. Facultad de Ciencias Naturales y Museo. División Paleontología Vertebrados; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina

(2006). The earliest Tertiary therian mammal from South America. Journal of Vertebrate Paleontology: Vol. 26, No. 2, pp. 505-510.

Loricariid and pimelodid-like siluriforms—along with undetermined percomorph—are described for the first time from Patagonia. Vertebrate fossils, including fish, mammals and birds, were found in beds attributed to the top of the Puerto Madryn Formation. These levels supposedly corresponded to the “Rionegrense marino” of former authors. Mammals occurring in the site suggest a Huayquerian age for the fossiliferous beds. The base of the Huayquerian was dated at about 9 Ma and the top is younger than 6.5 Ma. Radioisotopic dating in the marine shell beds of the Puerto Madryn Formation ranges from 11 to 9 Ma. Consequently, the section described here appears to be younger than the typical Puerto Madryn Formation from which it is separated by an unconformity. The section is correlated with the type Rio Negro Formation from northern Patagonia, which also includes Huayquerian fossils. Freshwater fishes were previously unknown in beds younger than the middle Miocene in southern South America. This is also the southernmost record of loricariid fishes. The association of aquatic continental and terrestrial vertebrates indicates for the first time in Peninsula Valdes beds of freshwater origin. The evidence apported by fossils is also in agreement with global climate trends. The local extinction of loricariids in Patagonia possibly occurred much later than the time of deposition.

Geographic variation in tooth row counts among sand tiger sharks Carcharias taurus (Chondrichthyes), from the SW Atlantic, NW Atlantic and the East China Sea is analyzed in this paper. We found significant differences between sand tigers from the SW Atlantic (Southern Hemisphere population) and each of the other two (Northern Hemisphere) regions in the number of upper lateral tooth rows, and between individuals from the SW Atlantic and the East China Sea in the total number of upper tooth rows. Sand tiger sharks from the two Northern Hemisphere populations did not differ in any of the studied variables. Our results agree with comparisons of vertebral counts between sand tiger sharks from Southern and Northern Hemispheres. Both lines of evidence suggest that Southern and Northern Hemisphere populations of C. taurus were isolated to a larger extent than populations of the Northern Hemisphere. The fossil record of the genus Carcharias begins in the Early Cretaceous and C. taurus is certainly known since the Late Miocene. During the Miocene, the Tethys Sea separating northern and southern land masses was still present and it provided a continuous temperate shallow sea that could allow dispersal of sand tiger sharks along Northern Hemisphere seas. Independent observations on the distribution and evolutionary history of the genera Myripristis, Neoniphon, Sargocentron and Aphanius, and genetic studies on the temperate shark genus Mustelus that indicate a close relationship between the Indo-Pacific M. manazo and the Mediterranean M. asterias suggest that this hypothesis is plausible and deserves to be tested.

During the latest Pleistocene-earliest Holocene, South American terrestrial vertebrate faunas suffered one of the largest (and probably the youngest) extinction in the world for this lapse. Megamammals, most of the large mammals and a giant terrestrial tortoise became extinct in the continent, and several complete ecological guilds and their predators disappeared. This mammal extinction had been attributed mainly to overkill, climatic change or a combination of both. We agree with the idea that human overhunting was the main cause of the extinction in South America. However, according to our interpretation, the slaughtering of mammals was accomplished in a particular climatic, ecological and biogeographical frame. During most of the middle and late Pleistocene, dry and cold climate and open areas predominated in South America. Nearly all of those megamammals and large mammals that became extinct were adapted to this kind of environments. The periodic, though relatively short, interglacial increases in temperature and humidity may have provoked the dramatic shrinking of open areas and extreme reduction of the biomass (albeit not in diversity) of mammals adapted to open habitats. Many populations were surely close to a minimum level of population viability. During the longer glacial periods, mammals populations recovered. This alternation of low and high biomass of mammals from open and closed areas is what we refer to as the Zig-Zag. During the present interglacial, humans entered South America and broke the Zig-Zag when killed all the megamammals and almost all the large mammals during their less favourable periodic lapse.

The first record of the knifejaw family Oplegnathidae in the Atlantic Ocean and in South America is reported. It comes from the lowermost beds of the Early Miocene Gaiman Formation at the lower Rio Chubut valley, central-eastern Patagonia. The family Oplegnathidae does not occur in the Atlantic today, but it was widespread in comparison to other knifejaw fishes, such as scarids and odacids. Several aquatic vertebrates were extirpated from the southern Atlantic Ocean in the Late Neogene. This record establishes a minimal age (Early Miocene) for the extirpation of the family Oplegnathidae in the Atlantic Ocean.

Siluriformes constitute an important monophyletic group of mainly freshwater fishes (Fink and Fink, 1996). Presently, there are about 35 families with over 2,000 species in all continents except Antarctica (Lundberg, 1993; Nelson, 1994); however, at least in the Eocene, they also inhabited Antarctica as well (Grande and Eastman, 1986). A small number of catfishes (most of the ariids and many of the plotosids) are marine and some species of other families can enter brackish waters (Nelson, 1994). Pre-Cenozoic catfishes are extremely rare outside of South America. One of us (G.V.R.P.) has been studying the vertebrate assemblages from the late Cretaceous inter-trappean beds of peninsular India for several years (e.g., Prasad, 1987; Prasad and Sahni, 1988; Prasad and Rage, 1991, 1995; Rage and Prasad, 1992; Prasad and Cappetta, 1993; Prasad et al., 1994; Prasad and Godinot, 1994; Prasad and Khajuria, 1996; Krause et al., 1997; Khajuria and Prasad, 1998). Recently, the authors found a catfish spine from an intertrappean bed exposed near the village Naskal, Andhra Pradesh (Fig. 1). The record is important because it demonstrates that ostariophysans inhabited India during the Mesozoic Era. In this paper we describe the spine, and comment about the environment and the Cretaceous catfish record in other continents. The catfish locality lies about 70 km west of Hyderabad and 2 km northeast of Naskal village, Rangareddi District, Andhra Pradesh, India (Fig. 1). This area is covered by flat-topped hills of Deccan volcanic flows with an altitude of 475 to 700 m above mean sea level. Nine volcanic flows have been delineated in the study area, of which …

Ten oblong aligned depressions in the Rio Cuarto area (provincia de Cordoba, Argentina) were supposed to be the result of very-low-angle Holocene meteoroid impacts. However, we consider that authors that studied the structures did not demonstrated their extraterrestrial origin. We suggest that an eolian origin for the structures of Rio Cuarto is more likely. Actually, these landforms integrate large systems of similar deflation/accumulation geoforms aligned according to predominant winds during different periods.

A date of 4300 ± 90 BP for extinct megafauna in Argentina is discussed. The fossil remains come from the Guerrero Member (area) of the Luján Formation near the city of Luján, Buenos Aires Province. The age of the top of the Guerrero Member is constrained by more than 60 radiocarbon dates obtained from the overlying Río Salado Member of Luján Formation, Las Escobas Formation, and Puesto Callejón Viejo Soil, most of them older than 4300 BP. In view of its low collagen content, the 14C measurement of bone sample from Luján should not be accepted uncritically. Because of the poor bone preservation and the possible introduction of "young" contaminants that were not completely eliminated, the 14C date of 4300 ± 90 BP is not reliable. Both biostratigraphic and 14C dating evidence indicates that the date of 4300 BP for the last representative of extinct megafauna in South America is unsupported., Facultad de Ciencias Naturales y Museo

The so-called Pampean sediments are the uppermost sedimentary levels of the Chacopampean basin. They record the sequence of geologic changes that occurred during the late Cenozoic.In the Pampean area, the most important environmental and faunistic changes occurred close to the end of the Matuyama Chron and beginning of the Brunhes Chron. The relatively wet climate recorded in the region at the end of Matuyama Chron turned to drier and colder conditions during the beginning of the Brunhes Chron, which also seems to be characterized by an important volcanic activity in the Cordillera de los Andes. An important faunistic turnover (between Tolypeutes pampaeus and Megatherium americanum biozones) coincides with lithostratigraphic changes (between the Ensenada and Buenos Aires formations). The change of polarity Matuyama–Brunhes is observed below this boundary.

An interdisciplinary study of a section in a quarry near La Plata sheds new light on the geologic, climatic, and biologic evolution of the northeastern Pampean area of Argentina. The stratigraphic succession is composed of seven mainly eolian levels, each one including a soil and separated by a disconformity. Two geosols defined in northern Buenos Aires Province (Hisisa and El Tala) are identified in the Ensenada Formation. The boundary between the Ensenada and Buenos Aires formations is defined by a conspicuous disconformity which overlies the El Tala Geosol. In the section, the boundary between the Tolypeutes pampaeus (Ensenadan) and Megatherium americanum (lower Lujanian) biozones coincides with that of Ensenada and Buenos Aires formations. The boundary between the Matuyama and Brunhes zones of polarity occurs in the upper part of the Ensenada Formation above the Hisisa Geosol. According to geologic evidence, most of the succession was deposited under semiarid to arid climate.