Leptoderma ospesca n. sp. (Fig. 1) Holotype. USNM 421478, 197 mm SL; El Salvador, Central America, Eastern Pacific Ocean, research vessel B/ O Miguel Oliver station MOP 11���94, trawl on soft-bottom from 12 �� 53 ' 15.72 " N, 90 �� 9 ' 30.6 " W to 12 �� 53 ' 22.9194 " N, 90 �� 7 ' 56.28 " W, at 1368���1406 m; December 11, 2010; D. Ross Robertson. Diagnosis. Leptoderma ospesca differs from L. affinis, L. macrops and L. retropinna by having a distinct gap between the posterior ends of the dorsal and anal fins and the origins of the upper and lower procurrent caudal-fin rays. From L. lubricum, the new species differs by lacking dermal papillae along the lateral line, by having pectoral fins low on body (vs. high) with 6 fin rays (vs. 7���8), and by having fewer caudal-fin rays (16, vs. 17���18). From L. macrophthalmum, the new species differs by having a longer pre-dorsal length (54.9 % SL, vs. 49.3 %), fewer pectoral-fin rays (6, vs. 8), fewer pelvic-fin rays (5, vs. 8) and longer dorsal- and anal-fin rays (vs. shorter). Leptoderma ospesca also differs from all congeneric species by having fewer pre-ural vertebrae (60, vs. 64���84). Description. Measurements and counts of L. ospesca and comparative data are presented in Table 1. Body naked, relatively long, subcylindrical anteriorly and laterally compressed posteriorly, with greatest body depth at a distance of 24.3 % SL from snout; dermal papillae absent from head, along lateral line and dorsum; lateral line pores inconspicuous. Head length (HL) almost twice greatest body depth. Nostrils with large longitudinal openings, partly covered by a dermal flap, located nearer to eye than to snout tip. Mandibular pores 4, supraorbital pores 3, suborbital pores 4 and preopercular pores 3. Snout blunt, relatively large, its length 30.3 % of HL, shorter than eye. Mouth subterminal. Teeth minute to small, conical and well separated, present in one row on premaxilla and one on dentary; no teeth on palate. Supramaxilla slender, curved. Upper jaw relatively short, barely reaching front border of eye, its length 33.0% of HL. Lower jaw ending under orbit. Eyes large, 37.5 % of HL, directed outward and slightly upward, with upper margin protruding above dorsal profile of head. Interorbital space relatively large, 17.2 % of HL. Postorbital length 32.2 % of HL. Maximal head width (behind eyes) 34.5 % of HL. Pseudobranchiae small. Pectoral fins inserted relatively low on body, uppermost fin ray at level of dorsal corner of gill opening, slightly below midbody. Pelvic fins abdominal, length 71.1 % of pectoral-fin length. Pelvic-fin base located just in front of anus. Genital papilla short, located between anus and anal-fin origin. Anal-fin origin well in advance of dorsal-fin origin; length of dorsal-fin base 60.4 % of length of anal-fin base. Dorsal- and anal-fin rays long, longer rays greater than half body depth at level of anus. Both dorsal and anal fins separated from caudal fin. Procurrent caudal-fin rays numerous, extending far forward on caudal peduncle. Dorsal and anal procurrent caudal sections about equal in length. Caudal fin small, forked, with 8 principal rays in each lobe. Color in life. Head, eye, and anterior part of body bluish black, turning gradually lighter brownish towards posterior part of body (Fig. 1 A). TABLE. 1. M��rph��metric and meristic characters ��f six Leptoderma species. Measurements and c��unts n��t attainable (i.e. structure absent ��r inc��nspicu��us) ��r unavailable are represented an en���dash (���). When the number ��f examined specimens used t�� determine the respective measurement ��r c��unt differs fr��m the t��tal, the number ��f examined specimens is indicated in parenthesis. Character L. affinis L. lubricum L. macrops L. macrophthalmum L. ospesca L.retropinna n= 1 2, * n= 21 3, * n= 52 4 n= 1 5, * n= 1 1, * n= 27 6 (mm) 222.5 29.7 ��� 210 141 ��� 210 151 197 28.4 ��� 152 Morphometric characters as % SL ���d��rsal length ��� 55.1 ��� 61.1 47.3 ��� 56.4 (49) 49.3 59.4 55.0��� 58.7 (6) ���anal length ��� 41.0��� 46.5 (20) 35.5 ��� 44.2 (49) 45.7 46.4 40.9 ��� 45.6 (6) ���pelvic length ��� 36.8 ��� 41.8 (20) 31.0��� 43.8 (49) 39.5 38.0 34.4 ��� 38.7 (6) D��rsal-fin base, length ��� 25.3 ��� 28.9 (5) 44.7 ��� 58.1 (14) 31.1 25.5 41.0��� 47.6 (6) D��rsal pr��current caudal-fin secti��n, length ��� 15.0��� 16.6 (5) 3.2-3.6 (3) 15.5 14.1 4.5-4.7 (2) Anal-fin base, length ��� 36.6 ���41.0 (5) 58.6 ��� 73.6 (14) 40.7 42.2 50.0���59.0 (6) ventral pr��current caudal-fin secti��n, length ��� 14.0��� 16.9 (5) 3.4-4.2 (3) 15.1 14.0 4.1-5.3 (3) Pect��ral-fin length ��� 5.0��� 11.4 (5) 6.0��� 8.4 (14) 7.3 7.0 4.9 ��� 5.8 (5) Pelvic-fin length ��� 1.8 ��� 6.9 (5) 3.9 ��� 5.8 (14) 4.8 5.0 4.0��� 4.9 (5) Greatest b��dy depth c. 11.1 11.1 ��� 14.7 (5) 9.8 ��� 12.3 (14) 12.8 13.4 7.8 ��� 9.2 (6) Caudal-peduncle depth ��� 2.0��� 2.8 (5) 0.9 ��� 1.5 (14) 2.1 1.8 1.1 ��� 1.3 (6) Maximal head width (behind eyes) ��� 8.6 ��� 14.9 (20) 7.8 ��� 9.2 (14) 11.3 8.4 6.6 ��� 8.6 (6) length c. 22.2 22.8 ��� 25.7 (5) 16.7 ��� 24.4 (49) 23.2 24.3 18.7 ��� 21.7 (6) Sn��ut length ��� 7.0��� 8.3 (5) 3.8 ���7.0 (49) 7 7.4 5.3 ���7.0 (6) H��riz��ntal ��rbital diameter ��� 7.5 ���9.0 (5) 5.7 ��� 9.4 (49) 9.3 9.1 3.3 ��� 7.2 (6) P��st��rbital length ��� ��� ��� 6.9 7.8 ��� Inter��rbital distance ��� 3.1 ��� 5.1 (5) 1.6 ��� 5.6 (49) 3.4 4.2 3.3 ��� 5.7 (6) Upper-��aw length ��� 5.6 ��� 8.9 (5) 4.6 ��� 7.3 (49) 7.9 8.0 5.3 ��� 7.3 (6) ������continued on the next page vertebrae, hemal (excl. ur��style) ��� 49 ��� 55 (4) ��� 51 ��� ��� vertebrae, t��tal (excl. ur��style) ��� 64 ��� 77 (20) 79 ��� 84 (35) 66 60 77 (1) Dermal papillae ��n lateral line:es:es (5):es;��;��;�� (5) US;M 421478, net al. (1965), nakami? @ukui (2010), net al. (2011), nakami? @ukui (2010), nFIGURE. 1. Leptoderma ospesca n. sp., holotype (USNM 421478), A: preserved entire specimen; B: newly caught entire specimen, arrows indicate position of gaps between dorsal and anal fin rays and upper and lower procurrent caudal rays, respectively; C close-up of head region of newly caught specimen. Note: The structure near the lateral midline above the anus that is visible in Fig. 1 A (but not obvious in Figs. 1 B or C), is a piece of loose skin adhered to the body. Color in alcohol. Head and anterior part of body black, turning gradually lighter grayish towards posterior part of body (Fig. 1 B). Distribution. Known only from the holotype collected off El Salvador, between 12 �� 53 ' 15.72 " N, 90 �� 9 ' 30.6 " W and 12 �� 53 ' 22.9194 " N, 90 �� 7 ' 56.28 " W, Central America, eastern Pacific Ocean, at a depth of 1368���1406 m. Etymology. The specific name ��� ospesca ��� recognizes the contribution of OSPESCA (Organizaci��n del Sector Pesquero y Ac��icola de Centroamerica) to the discovery of deepwater marine fishes in Central America by sponsoring research cruises of the Spanish research vessel B/ O Miguel Oliver on both the Pacific (2010) and Atlantic (2011) coasts of Central America. Suggested common names. Eastern eel-slickhead; Cabezalisa oriental., Published as part of Angulo, Arturo, Baldwin, Carole C. & Robertson, D. Ross, 2016, A new species of Leptoderma Vaillant, 1886 (Osmeriformes: Alepocephalidae) from the Pacific coast of Central America, pp. 493-500 in Zootaxa 4066 (4) on pages 494-498, DOI: 10.11646/zootaxa.4066.4.10, http://zenodo.org/record/266387, {"references":["Abe, T., Marumo, R. & Kawaguchi, K. (1965) Description of a new alepocephalid fish from Suruga Bay. Japanese Journal of Ichthyology, 13, 69 - 72.","Markle, D. F. (1976) Preliminary studies on the systematics of deep-sea Alocephaoidea (Pisces: Salmoniformes). Ph. D. Dissertation, the Faculty of the School of Marine Science, the College of William and Mary in Virginia, 265 pp."]}