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Athlia rustica Erichson, 1835 Figs. 48���53, 72. Original combination. Athlia rustica Erichson, 1835: 267 Type locality. ��� Chile ��� (Erichson 1835). Type series. Lectotype (Figs. 48���53) at ZMHB labeled a) ���25008��� (typeset), b) ���Type��� (typeset, orange label), c) ��� rustica / Er. / Chile Meyen.��� (handwritten, green label), d) ���SYNTYPUS / Athlia / rustica Erichson, 1835 / labelled by MNHUB 2009��� (typeset, orange label), e) ��� ATHLIA / RUSTICA / ERICHSON, 1835 / LECTOTYPE / A.B.T. SMITH��� (handwritten and typeset, red label). Lectotype here designated. Four paralectotypes at ZMHB labeled a) ���Type��� (typeset, orange label), b) ���Hist.-Coll. (Coleoptera) / Nr. 25008 / Athlia / rustica Erichs. / Chili, Meyen / Zool. Mus. Berlin��� (typeset), c) ���SYNTYPUS / Athlia / rustica Erichson, 1835 / labelled by MNHUB 2009��� (typeset, orange label), d) ��� ATHLIA / RUSTICA / ERICHSON, 1835 / PARALECTOTYPE / A.B.T. SMITH��� (handwritten and typeset, yellow label). A lectotype is designated to preserve the stability of the nomenclature by selecting a single specimen as the name-bearing type for this species. Since Erichson (1835) did not explicitly state how many specimens he examined, it is impossible to know the exact composition of the type series. Distribution. Fig. 72. Chile: IV Regi��n de Coquimbo, V Regi��n de Valpara��so, Regi��n Metropolitana de Santiago, VI Regi��n del Libertador General Bernardo O���Higgins, VII Regi��n del Maule, VIII Regi��n del Biob��o, IX Regi��n de la Araucan��a (based on specimens from AMNH, AVEC, BMNH, CASC, CMNC, CNCI, FMNH, FSCA, INHS, LEMQ, MNHN, MNNC, UMCE, UMSP, USNM). One 19th century specimen is labeled ���Valdivia��� (BMNH) but we consider this record erroneous. The record from IX Regi��n de la Araucan��a is based on a single collecting event from ��� 30 km NE Villarrica���, which might need further verification., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on pages 294-295, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270."]}

Athlia freyi Martínez, 1974 Figs. 24–27, 67. Original combination. Athlia freyi Martínez, 1974: 347. Type locality. “ Brasil, estado de São Paulo, Serra da Bocaina, 1500 m ” (Martínez 1974). Type series. Holotype at Georg Frey Collection, Naturhistorisches Museum Basel, Switzerland (not examined). One paratype at CMNC was examined. Distribution. Fig. 67. Brazil: São Paulo (based on specimens from CMNC)., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on page 287, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Martinez, A. (1974) Nuevas consideraciones sobre el Genero Athlia Erichson, 1835 (Col., Scarabaeidae, Sericinae). Entomologische Arbeiten aus dem Museum G. Frey, 25, 343 - 353."]}

Athlia rotundata Guti��rrez, 1949 Figs. 44���47, 71. Original combination. Athlia rotundata Guti��rrez, 1949: 12. Type locality. ��� Chile: Prov. Valpara��so, Conc��n ��� (Guti��rrez 1949). Type series. Holotype, allotype, and nine paratypes at Museo de Zoolog��a, Universidad de Concepci��n, Concepci��n, Chile (Cekalovic & Artigas 1969). Two paratypes were examined from CMNC and FMNH. Distribution. Fig. 71. Chile: V Regi��n de Valpara��so (based on specimens from CASC, CMNC, FMNH, UMCE)., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on page 293, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Gutierrez, R. (1949) Notas sobre Scarabaeidae neotropicos (Coleoptera Lamellicornia). Anales de la Sociedad Cientifica Argentina, 148, 9 - 35.","Cekalovic, T. & Artigas, J. N. (1969) Catalogo de los tipos de Insecta depositados en la coleccion del Departamento de Zoologia de la Universidad de Concepcion (INCO) (Diciembre, 1968). Boletin de la Sociedad de Biologia de Concepcion, 41, 111 - 133."]}

Key to species of Athlia (based on Saylor 1946b; Gutiérrez 1949; Martínez 1967, 1974) 1. Metathoracic wings reduced and non-functional (slightly lift an elytron to check). Central Chile (VIII Región del Biobío to IX Región de la Araucanía) (Fig. 69)..................................................... Athlia plebeja Burmeister - Metathoracic wings full sized and functional................................................................ 2 2. Pronotum with basal angles evenly rounded, not protruding or angulate (Figs. 30, 34, 46)............................ 3 - Pronotum with basal angles protruding or angulate (Figs. 13, 17, 22, 26, 42, 52).................................... 5 3. Length less than 10 mm. Metatarsomere 1 noticeably shorter than metatarsomere 2 (Fig. 32). Southern Argentina (Neuquén to Chubut, Fig. 69)................................................................... Athlia parvissima Saylor - Length greater than 10 mm. Metatarsomere 1 equal to or slightly shorter than metatarsomere 2 (Fig. 28)................. 4 4. Labrum weakly curved, weakly elevated medially; lateral border of clypeus slightly reflexed; dorsal surface tan to chestnut brown. Central Chile (V Región de Valparaíso, Fig. 79)................................... Athlia rotundata Gutiérrez - Labrum evenly curved and elevated medially; lateral border of clypeus strongly reflexed; dorsal surface light brown to light tan. Southern Argentina (Neuquén to Chubut) and southern Chile (Región de Aisén, Fig. 68).......... Athlia giaii Martínez 5. Labrum not reflexed, on the same plane as clypeal surface. Southeastern Brazil (Santa Catarina, Fig. 67).................................................................................................... Athlia brasilica Saylor - Labrum reflexed above the plane of clypeal surface........................................................... 6 6. Pronotum surface with dense, even punctures (Figs. 22, 42, 52)................................................. 7 - Pronotum surface with sparse, sometimes unevenly distributed punctures (Figs. 13, 26).............................. 9 7. Elytra evenly covered with setae; pronotum with long setae along lateral edges. Central Chile (IV Región de Coquimbo to IX Región de la Araucanía, Fig. 72)........................................................ Athlia rustica Erichson - Elytra glabrous medially and sparsely setose laterally and apically; pronotum with long setae along lateral edge but mainly in apical half........................................................................................... 8 8. Elytra length less than twice the length of head and pronotum combined Northern and central Argentina (Jujuy to Río Negro, Fig. 67).............................................................................. Athlia bruchi Moser - Elytra length greater than twice the length of head and pronotum combined. Central Chile (IV Región de Coquimbo to Región Metropolitana de Santiago, Fig. 70)................................................ Athlia problematica Gutiérrez 9. Length greater than 12 mm; apical and basal pronotal angles well developed (Fig. 26). Southern Brazil (São Paulo, Fig. 67)...................................................................................... Athlia freyi Martínez - Length less than 12 mm; apical and basal pronotal angles weakly developed (Fig. 13). Northeastern Argentina (Buenos Aires, Fig. 67)............................................................................ Athlia bollei Martínez

Athlia bollei Martínez, 1955 Figs. 11–14, 67. Original combination. Athlia bollei Martínez, 1955: 418. Type locality. “ Argentina, provincial de Buenos Aires, Partido de Tandill (Serra-nía)” (Martínez 1955). Type series. Holotype at Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Buenos Aires, Argentina (not examined). Distribution. Fig. 67. Argentina: Buenos Aires (based on specimens from CMNC)., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on page 286, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Martinez, A. (1955) Una nueva especie de Athlia de la Argentina (Col. Scarab. Melolonth. Sericinae). Revista Ecuatoriana de Entomologia y Parasitologia, Guayaquil, 2, 417 - 423."]}

Athlia parvissima Saylor, 1946 Figs. 32���35, 69. Original combination. Athlia parvissima Saylor, 1946: 24. Type locality. ��� Zapala, Neuquen, Argentina ��� (Saylor 1946). Type series. Holotype at CASC labeled a) ���Zapala, / Neuquen ��� (typeset), b) ��� Argentina / Dec 9 1920 ��� (typeset), c) ��� HOLOTYPE / Athlia / PARVISSIMA / L.W.Saylor��� (typeset and handwritten on red paper), d) ���PARVISSIMA / Saylor / DET. / L.W. SAYLOR��� (handwritten and typeset), e) ���L.W. Saylor / Collection��� (typeset), f) ��� California Academy / of Sciences / Type / No. 7923��� (typeset and handwritten), g) ���Southern Neotropical Scarabs / database # AS 2617613 / Athlia parvissima / Saylor, 1946 ♀ / DET: A.B.T. SMITH 2008 ��� (typeset). Distribution. Fig. 69. Argentina: Neuqu��n, R��o Negro, Chubut (based on specimens from CASC, CDFA, CMNC, LEMQ)., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on pages 288-289, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Saylor, L. W. (1946) Studies in the melolonthine scarab beetle genera of the American continents. I. Revision of the genus Athlia. Proceedings of the Entomological Society of Washington, 48, 18 - 25.","Smith, A. B. T. (2008) South American Melolonthinae (Coleoptera: Scarabaeidae) classification and nomenclature: some problems and solutions. Insecta Mundi, 60, 1 - 28."]}

Athlia giaii Mart��nez, 1959 Figs. 28���31, 68. Original combination. Athlia giaii Mart��nez, 1959: 23. Type locality. ��� Neuqu��n, Piedra del Aguila ��� (Mart��nez 1959). Type series. Holotype at Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Buenos Aires, Argentina (not examined). Twenty paratypes at CMNC were examined. Distribution. Fig. 68. Argentina: Neuqu��n, R��o Negro, Chubut (based on specimens from CDFA, CMNC, LEMQ). Mondaca (2011) recorded specimens from Chile: XI Regi��n de Ais��n., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on page 288, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Martinez, A. (1959) Scarabaeoidea neotropica. VI. Algunos Melolonthidae nuevos o poco conocidos (Col.). Neotropica, 5 (16), 23 - 29.","Mondaca, J. (2011) Presencia de Athlia giaii y Myloxena patagonica (Coleoptera: Scarabaeidae: Melolonthinae) en la Patagonia occidental, Chile. Revista Chilena de Entomologia, 36, 43 - 49."]}

Key to species of Athlia (based on Saylor 1946b; Gutiérrez 1949; Martínez 1967, 1974) 1. Metathoracic wings reduced and non-functional (slightly lift an elytron to check). Central Chile (VIII Región del Biobío to IX Región de la Araucanía) (Fig. 69)..................................................... Athlia plebeja Burmeister - Metathoracic wings full sized and functional................................................................ 2 2. Pronotum with basal angles evenly rounded, not protruding or angulate (Figs. 30, 34, 46)............................ 3 - Pronotum with basal angles protruding or angulate (Figs. 13, 17, 22, 26, 42, 52).................................... 5 3. Length less than 10 mm. Metatarsomere 1 noticeably shorter than metatarsomere 2 (Fig. 32). Southern Argentina (Neuquén to Chubut, Fig. 69)................................................................... Athlia parvissima Saylor - Length greater than 10 mm. Metatarsomere 1 equal to or slightly shorter than metatarsomere 2 (Fig. 28)................. 4 4. Labrum weakly curved, weakly elevated medially; lateral border of clypeus slightly reflexed; dorsal surface tan to chestnut brown. Central Chile (V Región de Valparaíso, Fig. 79)................................... Athlia rotundata Gutiérrez - Labrum evenly curved and elevated medially; lateral border of clypeus strongly reflexed; dorsal surface light brown to light tan. Southern Argentina (Neuquén to Chubut) and southern Chile (Región de Aisén, Fig. 68).......... Athlia giaii Martínez 5. Labrum not reflexed, on the same plane as clypeal surface. Southeastern Brazil (Santa Catarina, Fig. 67).................................................................................................... Athlia brasilica Saylor - Labrum reflexed above the plane of clypeal surface........................................................... 6 6. Pronotum surface with dense, even punctures (Figs. 22, 42, 52)................................................. 7 - Pronotum surface with sparse, sometimes unevenly distributed punctures (Figs. 13, 26).............................. 9 7. Elytra evenly covered with setae; pronotum with long setae along lateral edges. Central Chile (IV Región de Coquimbo to IX Región de la Araucanía, Fig. 72)........................................................ Athlia rustica Erichson - Elytra glabrous medially and sparsely setose laterally and apically; pronotum with long setae along lateral edge but mainly in apical half........................................................................................... 8 8. Elytra length less than twice the length of head and pronotum combined Northern and central Argentina (Jujuy to Río Negro, Fig. 67).............................................................................. Athlia bruchi Moser - Elytra length greater than twice the length of head and pronotum combined. Central Chile (IV Región de Coquimbo to Región Metropolitana de Santiago, Fig. 70)................................................ Athlia problematica Gutiérrez 9. Length greater than 12 mm; apical and basal pronotal angles well developed (Fig. 26). Southern Brazil (São Paulo, Fig. 67)...................................................................................... Athlia freyi Martínez - Length less than 12 mm; apical and basal pronotal angles weakly developed (Fig. 13). Northeastern Argentina (Buenos Aires, Fig. 67)............................................................................ Athlia bollei Martínez, Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on pages 285-286, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Saylor, L. W. (1946) Studies in the melolonthine scarab beetle genera of the American continents. I. Revision of the genus Athlia. Proceedings of the Entomological Society of Washington, 48, 18 - 25.","Gutierrez, R. (1949) Notas sobre Scarabaeidae neotropicos (Coleoptera Lamellicornia). Anales de la Sociedad Cientifica Argentina, 148, 9 - 35.","Martinez, A. (1967) El genero Athlia Erichson (Col. Scarabaeidae, Sericinae). Entomologische Arbeiten aus dem Museum G. Frey, 18, 327 - 372.","Martinez, A. (1974) Nuevas consideraciones sobre el Genero Athlia Erichson, 1835 (Col., Scarabaeidae, Sericinae). Entomologische Arbeiten aus dem Museum G. Frey, 25, 343 - 353."]}

Athlia Erichson, 1835 Athlia Erichson, 1835: 266. Type species Athlia rustica Erichson, 1835 by monotypy. Synonym. Rivera Germain, 1903: 392. Type species Athlia plebeja Burmeister, 1855 by monotypy. Redescription. Length 6.5–17.0 mm. Elongate and robust, widest at about elytral declivity; uniformly pale to medium yellowish brown, or brown or reddish brown; dorsal surface dull, sometimes with a faint, glaucous coating on pronotal and elytral surfaces, or weakly shiny, sometimes faintly iridescent, and with or without short, scattered, and appressed setae. Head: labrum wide, broadly emarginate and evenly reflexed anteriorly, and completely fused to clypeus with labroclypeal suture more-or-less distinct and terminating in small, marginal notches. Mentum longer than wide, narrowly emarginate in front, with sides gradually converging posteriorly. Antennae with 9 antennomeres, club with 3 short, thick lamellae. Pronotum: convex, wider than long, widest medially or slightly behind middle. Elytra: disc of each with four weak and narrow costae, interstriae broad and confusedly punctate. Metathoracic wings fully developed or reduced in size (brachypterous). Abdomen: with lateral carinae on ventrites. Legs: protibia with three apical teeth on outer margin, spur present and shorter than protarsomere 1. Claws symmetrical and incised, with short, bisetose onychium in between. Metatibial apices with spurs set in notch, longest spur reaching half to three-quarter point of metatarsomere 1. Male genitalia: symmetrical parameres simple in form, not split or with strong sinuations, and somewhat shorter than phallobase. Composition. All seven known species of Athlia are endemic to the southern half of South America. Remarks. Saylor (1946b) and Martínez (1967, 1974) previously reviewed this genus but a detailed taxonomic inventory of all specimens in collections is needed. We predict that new species of Athlia will be found in southern Brazil and northern Argentina, where endemism is high and specimens are poorly represented in collections., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on page 285, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Erichson, W. F. (1835) Neue Sudamerikanische Kafergattungen aus der familie der Blatterhorner. Archiv fur Naturgeschichte, 1, 256 - 270.","Germain, P. (1903) Apuntes entomologicos. Anales de la Universidad. Republica de Chile, 112 - 113, 391 - 445.","Burmeister, H. C. C. (1855) Handbuch der Entomologie. Fierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Friedr. Enslin, Berlin, x + 569 pp.","Saylor, L. W. (1946) Studies in the melolonthine scarab beetle genera of the American continents. I. Revision of the genus Athlia. Proceedings of the Entomological Society of Washington, 48, 18 - 25.","Martinez, A. (1967) El genero Athlia Erichson (Col. Scarabaeidae, Sericinae). Entomologische Arbeiten aus dem Museum G. Frey, 18, 327 - 372.","Martinez, A. (1974) Nuevas consideraciones sobre el Genero Athlia Erichson, 1835 (Col., Scarabaeidae, Sericinae). Entomologische Arbeiten aus dem Museum G. Frey, 25, 343 - 353."]}

Athlia bruchi Moser, 1924 Figs. 19–23, 67. Original combination. Athlia bruchi Moser, 1924: 121. Type locality. “ Argentina: Cordoba ” (Moser 1924). Type series. Lectotype (Figs. 19–23) at ZMHB labeled a) “Rep ARGENTINA / Prov. Cordoba / 1911 / C. Bruch” (typeset), b) “ Athlia / Bruchi / Type Mos.” (handwritten), c) “Bruchi Mos.” (handwritten), d) “SYNTYPUS / Athlia / bruchi Moser, 1924 / labelled by MNHUB 2009” (typeset, orange label), e) “ ATHLIA / BRUCHI / MOSER, 1924 / LECTOTYPE / A.B.T. SMITH” (handwritten and typeset, red label). Lectotype here designated. A lectotype is designated to preserve the stability of the nomenclature by selecting a single specimen as the namebearing type for this species. Since Moser (1924) did not explicitly state how many specimens he examined, it is impossible to know the exact composition of the type series. Distribution. Fig. 67. Argentina: Buenos Aires, Córdoba, Jujuy, La Pampa, Neuquén, Río Negro, San Luis (based on specimens from CASC, CMNC, ZMHB)., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on pages 286-287, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Moser, J. (1924) Beitrag zur Kenntnis der Melolonthiden (Col.). (XIII). Stettiner Entomologische Zeitung, 84, 114 - 136."]}

Athlia brasilica Saylor, 1946 Figs. 15���18, 67. Original combination. Athlia brasilica Saylor, 1946: 23. Type locality. ���Nova Galicia, S. Catarina, Brazylja ��� (Saylor 1946). Type series. Holotype at CASC labeled a) ���Nowa Galicja / Brazylja,S. Catharina ��� (typeset), b) ��� HOLOTYPE / Athlia / BRASILICA / L.W.Saylor��� (typeset and handwritten on red paper), c) ���BRASILICA / Saylor / DET. / L.W. SAYLOR��� (handwritten and typeset), d) ���L.W. Saylor / Collection��� (typeset), e) ��� California Academy / of Sciences / Type / No. 7922��� (typeset and handwritten). One paratype at CASC (examined). Distribution. Fig. 67. Brazil: Santa Catarina (based on specimens from CASC, CMNC)., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on page 286, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Saylor, L. W. (1946) Studies in the melolonthine scarab beetle genera of the American continents. I. Revision of the genus Athlia. Proceedings of the Entomological Society of Washington, 48, 18 - 25."]}

Athlia problematica Guti��rrez, 1950 Figs. 40���43, 70. Original combination. Athlia problematica Guti��rrez, 1950: 270. Type locality. ��� Chile: Prov. Valpara��so, Conc��n ��� (Guti��rrez 1950). Type series. Holotype, allotype, and eight paratypes at Museo de Zoolog��a, Universidad de Concepci��n, Concepci��n, Chile (Cekalovic & Artigas 1969). Two paratypes at BMNH, one paratype at FMNH, and two paratypes at CMNC were examined. Distribution. Fig. 70. Chile: IV Regi��n de Coquimbo, V Regi��n de Valpara��so, Regi��n Metropolitana de Santiago (based on specimens from AMNH, BMNH, CASC, CMNC, FMNH, UMCE). The IV Regi��n de Coquimbo record is based on a single AMNH specimen collected at Combarbal�� on 11 November 1992 by Rosen, Sharkov, and Snyder. Additional specimens are needed to corroborate the Regi��n de Coquimbo distributional record., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on pages 292-293, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Gutierrez, R. (1950) Notas sobre Scarabaeidae Chilenos (Coleoptera Lamellicornia). Arthropoda, 1 (2 / 4), 267 - 278.","Cekalovic, T. & Artigas, J. N. (1969) Catalogo de los tipos de Insecta depositados en la coleccion del Departamento de Zoologia de la Universidad de Concepcion (INCO) (Diciembre, 1968). Boletin de la Sociedad de Biologia de Concepcion, 41, 111 - 133."]}

Athlia plebeja Burmeister, 1855 Figs. 36–39, 69. Original combination. Athlia plebeja Burmeister, 1855: 125. Subsequent combination. Rivera plebeja (Burmeister): Germain 1903: 392. Type locality. “ Chili ” (Burmeister 1855). Type series. Syntypes presumably at Martin-Luther-Universität, Halle, Germany (not examined). Synonym. Athlia rivera Saylor, 1946: 23. Type locality: “ Chile, Concepcion” (Saylor 1946). Type series: holotype male at CASC labeled a) “ CHILE / Concepcion / P.HerbstS.” (typeset) on upperside and “ 30.x.04.” (handwritten) on underside, b) “Moser determ. / Athlia n.sp. ” (typeset and handwritten), c) “ HOLOTYPE / Athlia / RIVERA / L.W.Saylor” (typeset and handwritten on red paper), d) “ RIVERA / Saylor / DET. / L.W. SAYLOR” (handwritten and typeset), e) “L.W. Saylor / Collection” (typeset), f) “ California Academy / of Sciences / Type / No. 7924” (typeset and handwritten), g) “Southern Neotropical Scarabs / database # AS 2617639 / Athlia plebeja / Burmeister, 1855 ♂ / DET: A.B.T. SMITH 2008 ” (typeset). One paratype at CASC erroneously labeled “ Peru ” (examined). Distribution. Fig. 69. Chile: VII Región del Libertador General Bernardo O’Higgins, VIII Región del Biobío, IX Región de la Araucanía (based on specimens from BMNH, CASC, CMNC, FMNH, MNHN, USNM). Two century-old specimens were also seen labeled “Limache” (CASC) and “ Valparaíso ” (CMNC) but we consider these records as erroneous. The Valparaíso region is far outside the distribution of this species., Published as part of Smith, Andrew B. T. & Evans, Arthur V., 2018, Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America, pp. 279-308 in Zootaxa 4471 (2) on pages 289-292, DOI: 10.11646/zootaxa.4471.2.3, http://zenodo.org/record/1439674, {"references":["Burmeister, H. C. C. (1855) Handbuch der Entomologie. Fierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Friedr. Enslin, Berlin, x + 569 pp.","Germain, P. (1903) Apuntes entomologicos. Anales de la Universidad. Republica de Chile, 112 - 113, 391 - 445.","Saylor, L. W. (1946) Studies in the melolonthine scarab beetle genera of the American continents. I. Revision of the genus Athlia. Proceedings of the Entomological Society of Washington, 48, 18 - 25.","Smith, A. B. T. (2008) South American Melolonthinae (Coleoptera: Scarabaeidae) classification and nomenclature: some problems and solutions. Insecta Mundi, 60, 1 - 28."]}