181 results on '"Beverton–Holt model"'
Search Results
2. Model‐based offline reinforcement learning for sustainable fishery management.
- Author
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Ju, Jun, Kurniawati, Hanna, Kroese, Dirk, and Ye, Nan
- Subjects
- *
PARTIALLY observable Markov decision processes , *SUSTAINABLE fisheries , *FISHERY policy , *STOCHASTIC systems , *REINFORCEMENT learning - Abstract
Fisheries, as indispensable natural resources for human, need to be managed with both short‐term economical benefits and long‐term sustainability in consideration. This has remained a challenge, because the population and catch dynamics of the fisheries are complex and noisy, while the data available is often scarce and only provides partial information on the dynamics. To address these challenges, we formulate the population and catch dynamics as a Partially Observable Markov Decision Process (POMDP), and propose a model‐based offline reinforcement learning approach to learn an optimal management policy. Our approach allows learning fishery management policies from possibly incomplete fishery data generated by a stochastic fishery system. This involves first learning a POMDP fishery model using a novel least squares approach, and then computing the optimal policy for the learned POMDP. The learned fishery dynamics model is useful for explaining the resulting policy's performance. We perform systematic and comprehensive simulation study to quantify the effects of stochasticity in fishery dynamics, proliferation rates, missing values in fishery data, dynamics model misspecification, and variability of effort (e.g., the number of boat days). When the effort is sufficiently variable and the noise is moderate, our method can produce a competitive policy that achieves 85% of the optimal value, even for the hardest case of noisy incomplete data and a misspecified model. Interestingly, the learned policies seem to be robust in the presence of model learning errors. However, non‐identifiability kicks in if there is insufficient variability in the effort level and the fishery system is stochastic. This often results in poor policies, highlighting the need for sufficiently informative data. We also provide a theoretical analysis on model misspecification and discuss the tendency of a Schaefer model to overfit compared with a Beverton–Holt model. [ABSTRACT FROM AUTHOR]
- Published
- 2025
- Full Text
- View/download PDF
3. Who is afraid of modelling time as a continuous variable?
- Author
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Hanna Kokko
- Subjects
Beverton–Holt model ,evolutionary dynamics ,Gillespie algorithm ,mate searching ,population dynamics ,predator–prey dynamics ,Ecology ,QH540-549.5 ,Evolution ,QH359-425 - Abstract
Abstract Most models of ecological and eco‐evolutionary processes involve creating trajectories of something, be it population densities, average trait values, or environmental states, over time. This requires decision‐making regarding how to represent the flow of time in models. Most ecologists are exposed to continuous‐time models (typically in the form of ordinary differential equations) as part of their training, especially since the famous Lotka‐Volterra predator–prey dynamics are formulated this way. However, few appear sufficiently well trained to produce their own work with continuous‐time models and may lack exposure to the true versatility of available methods. Specifically, knowledge that discrete individuals can be modelled in continuous time using the Gillespie algorithm is not as widespread as it should be. I will illustrate the flexibility of continous‐time modelling methods such that researchers can make informed choices, and not resort to discretizing time as a ‘default’ without a clear biological motivation to do so. I provide three example‐based tutorials: (1) a comparison of deterministic and stochastic dynamics of the Lotka‐Volterra predator–prey model, (2) an evaluation of matelessness in a hypothetical insect population (and of selection to mate more often by either searching more efficiently or by shortening the ‘time out’ after each mating) and (3) within‐season density dependence followed by a birth pulse leading to Beverton‐Holt or Ricker dynamics depending on whether the deaths of conspecifics help reduce the mortality of others or not (compensatory mortality). I highlight properties of the exponential distribution that, while counter‐intuitive, are good to know when deriving expected lifetime reproductive success or other similar quantities. I also give guidance on how to proceed if the so‐called memorylessness assumption does not hold in a given situation, and show how continuous and discrete times can be freely mixed if the biological situation dictates this to be the preferred option. Continuous‐time models can also be empirically fitted to data, and I review briefly the insight this gives into the so‐called ‘do hares eat lynx?’ paradox that has been plaguing the interpretation of the Hudson Bay hare and lynx dataset.
- Published
- 2024
- Full Text
- View/download PDF
4. Who is afraid of modelling time as a continuous variable?
- Author
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Kokko, Hanna
- Subjects
CONTINUOUS time models ,DISTRIBUTION (Probability theory) ,DIFFERENTIAL forms ,BOBCAT ,ORDINARY differential equations - Abstract
Most models of ecological and eco‐evolutionary processes involve creating trajectories of something, be it population densities, average trait values, or environmental states, over time. This requires decision‐making regarding how to represent the flow of time in models. Most ecologists are exposed to continuous‐time models (typically in the form of ordinary differential equations) as part of their training, especially since the famous Lotka‐Volterra predator–prey dynamics are formulated this way. However, few appear sufficiently well trained to produce their own work with continuous‐time models and may lack exposure to the true versatility of available methods. Specifically, knowledge that discrete individuals can be modelled in continuous time using the Gillespie algorithm is not as widespread as it should be.I will illustrate the flexibility of continous‐time modelling methods such that researchers can make informed choices, and not resort to discretizing time as a 'default' without a clear biological motivation to do so. I provide three example‐based tutorials: (1) a comparison of deterministic and stochastic dynamics of the Lotka‐Volterra predator–prey model, (2) an evaluation of matelessness in a hypothetical insect population (and of selection to mate more often by either searching more efficiently or by shortening the 'time out' after each mating) and (3) within‐season density dependence followed by a birth pulse leading to Beverton‐Holt or Ricker dynamics depending on whether the deaths of conspecifics help reduce the mortality of others or not (compensatory mortality).I highlight properties of the exponential distribution that, while counter‐intuitive, are good to know when deriving expected lifetime reproductive success or other similar quantities. I also give guidance on how to proceed if the so‐called memorylessness assumption does not hold in a given situation, and show how continuous and discrete times can be freely mixed if the biological situation dictates this to be the preferred option.Continuous‐time models can also be empirically fitted to data, and I review briefly the insight this gives into the so‐called 'do hares eat lynx?' paradox that has been plaguing the interpretation of the Hudson Bay hare and lynx dataset. [ABSTRACT FROM AUTHOR]
- Published
- 2024
- Full Text
- View/download PDF
5. Global Attractivity for Nonautonomous Delay-Differential Equations with Mixed Monotonicity and Two Delays.
- Author
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El-Morshedy, Hassan and Ruiz-Herrera, Alfonso
- Subjects
- *
DELAY differential equations , *DIFFERENCE equations , *EQUATIONS , *BLOWFLIES - Abstract
In this paper we study the scalar delay differential equation x ′ (t) = α (t) x (t - g 1 (t)) f (a (t) , x (t - g 2 (t))) - β (t) x (t) where f is decreasing in both arguments and the coefficients are positive and bounded. Sufficient conditions for the permanence and global attractivity for a fixed positive solution are derived. We apply our results to nonautonomous variants of Nicholson's blowfly equation and the Beverton–Holt model. [ABSTRACT FROM AUTHOR]
- Published
- 2024
- Full Text
- View/download PDF
6. The shape of density dependence and the relationship between population growth, intraspecific competition and equilibrium population density.
- Author
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Fronhofer, Emanuel A., Govaert, Lynn, O'Connor, Mary I., Schreiber, Sebastian J., and Altermatt, Florian
- Abstract
The logistic growth model is one of the most frequently used formalizations of density dependence affecting population growth, persistence and evolution. Ecological and evolutionary theory, and applications to understand population change over time often include this model. However, the assumptions and limitations of this popular model are often not well appreciated. Here, we briefly review past use of the logistic growth model and highlight limitations by deriving population growth models from underlying consumer–resource dynamics. We show that the logistic equation likely is not applicable to many biological systems. Rather, density‐regulation functions are usually non‐linear and may exhibit convex or concave curvatures depending on the biology of resources and consumers. In simple cases, the dynamics can be fully described by the Schoener model. More complex consumer dynamics show similarities to a Maynard Smith–Slatkin model. We show how population‐level parameters, such as intrinsic rates of increase and equilibrium population densities are not independent, as often assumed. Rather, they are functions of the same underlying parameters. The commonly assumed positive relationship between equilibrium population density and competitive ability is typically invalid. We propose simple relationships between intrinsic rates of increase and equilibrium population densities that capture the essence of different consumer–resource systems. Relating population level models to underlying mechanisms allows us to discuss applications to evolutionary outcomes and how these models depend on environmental conditions, like temperature via metabolic scaling. Finally, we use time‐series from microbial food chains to fit population growth models as a test case for our theoretical predictions. Our results show that density‐regulation functions need to be chosen carefully as their shapes will depend on the study system's biology. Importantly, we provide a mechanistic understanding of relationships between model parameters, which has implications for theory and for formulating biologically sound and empirically testable predictions. [ABSTRACT FROM AUTHOR]
- Published
- 2024
- Full Text
- View/download PDF
7. 东海带鱼的最适可捕规格.
- Author
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袁帆, 朱文斌, 王忠明, 朱凯, 周永东, and 徐汉祥
- Abstract
Copyright of Chinese Journal of Applied Ecology / Yingyong Shengtai Xuebao is the property of Chinese Journal of Applied Ecology and its content may not be copied or emailed to multiple sites or posted to a listserv without the copyright holder's express written permission. However, users may print, download, or email articles for individual use. This abstract may be abridged. No warranty is given about the accuracy of the copy. Users should refer to the original published version of the material for the full abstract. (Copyright applies to all Abstracts.)
- Published
- 2023
- Full Text
- View/download PDF
8. Economic Dynamics of Russia: Approach Based on the Solow-Swan Model
- Author
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Tolmachev, M. N., Latkov, A. V., Mitrofanov, A. Y., Barashov, N. G., Howlett, Robert J., Series Editor, Jain, Lakhmi C., Series Editor, Solovev, Denis B., editor, Savaley, Viktor V., editor, Bekker, Alexander T., editor, and Petukhov, Valery I., editor
- Published
- 2021
- Full Text
- View/download PDF
9. Derivation and dynamics of discrete population models with distributed delay in reproduction.
- Author
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Streipert SH and Wolkowicz GSK
- Subjects
- Animals, Mathematical Concepts, Reproduction physiology, Population Dynamics statistics & numerical data, Models, Biological
- Abstract
We introduce a class of discrete single species models with distributed delay in the reproductive process and a cohort dependent survival function that accounts for survival pressure during that delay period. These delay recurrences track the mature population for species in which individuals reach maturity after at least τ and at most τ+τ
M breeding cycles. Under realistic model assumptions, we prove the existence of a critical delay threshold, τ˜c . For given delay kernel length τM , if each individual takes at least τ˜c time units to reach maturity, then the population is predicted to go extinct. We show that the positive equilibrium is decreasing in both τ and τM . In the case of a constant reproductive rate, we provide an equation to determine τ˜c for fixed τM , and similarly, provide a lower bound on the kernel length, τ˜M for fixed τ such that the population goes extinct if τM ≥τ˜M . We compare these critical thresholds for different maturation distributions and show that if all else is the same, to avoid extinction it is best if all individuals in the population have the shortest delay possible. We apply the model derivation to a Beverton-Holt model and discuss its global dynamics. For this model with kernels that share the same mean delay, we show that populations with the largest variance in the time required to reach maturity have higher population levels and lower chances of extinction., Competing Interests: Declaration of competing interest The authors, declare that they have no conflicts of interest., (Copyright © 2024 Elsevier Inc. All rights reserved.)- Published
- 2024
- Full Text
- View/download PDF
10. An alternative delayed population growth difference equation model.
- Author
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Streipert, Sabrina H. and Wolkowicz, Gail S. K.
- Abstract
We propose an alternative delayed population growth difference equation model based on a modification of the Beverton–Holt recurrence, assuming a delay only in the growth contribution that takes into account that those individuals that die during the delay, do not contribute to growth. The model introduced differs from a delayed logistic difference equation, known as the delayed Pielou or delayed Beverton–Holt model, that was formulated as a discretization of the Hutchinson model. The analysis of our delayed difference equation model identifies a critical delay threshold. If the time delay exceeds this threshold, the model predicts that the population will go extinct for all non-negative initial conditions. If the delay is below this threshold, the population survives and its size converges to a positive globally asymptotically stable equilibrium that is decreasing in size as the delay increases. We show global asymptotic stability of the positive equilibrium using two different techniques. For one set of parameter values, a contraction mapping result is applied, while the proof for the remaining set of parameter values, relies on showing that the map is eventually componentwise monotone. [ABSTRACT FROM AUTHOR]
- Published
- 2021
- Full Text
- View/download PDF
11. Age structured discrete-time disease models with demographic population cycles
- Author
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P. van den Driessche and Abdul-Aziz Yakubu
- Subjects
adults ,beverton-holt model ,juveniles ,population cycles ,ricker model ,Environmental sciences ,GE1-350 ,Biology (General) ,QH301-705.5 - Abstract
We use juvenile-adult discrete-time infectious disease models with intrinsically generated demographic population cycles to study the effects of age structure on the persistence or extinction of disease and the basic reproduction number, $\mathcal {R}_{0} $. Our juvenile-adult Susceptible-Infectious-Recovered (SIR) and Infectious-Salmon Anemia-Virus (ISA $v) $ models share a common disease-free system that exhibits equilibrium dynamics for the Beverton-Holt recruitment function. However, when the recruitment function is the Ricker model, a juvenile-adult disease-free system exhibits a range of dynamic behaviours from stable equilibria to deterministic period k population cycles to Neimark-Sacker bifurcations and deterministic chaos. For these two models, we use an extension of the next generation matrix approach for calculating $\mathcal {R}_{0} $ to account for populations with locally asymptotically stable period k cycles in the juvenile-adult disease-free system. When $\mathcal {R}_{0} \lt 1 $ and the juvenile-adult demographic system (in the absence of the disease) has a locally asymptotically stable period k population cycle, we prove that the juvenile-adult disease goes extinct whenever $\mathcal {R}_{0} \lt 1 $. Under the same period k juvenile-adult demographic assumption but with $\mathcal {R}_{0} \gt 1 $, we prove that the juvenile-adult disease-free period k population cycle is unstable and the disease persists. When $\mathcal {R}_{0} \gt 1 $, our simulations show that the juvenile-adult disease-free period k cycle dynamics drives the juvenile-adult SIR disease dynamics, but not the juvenile-adult ISAv disease dynamics.
- Published
- 2020
- Full Text
- View/download PDF
12. Influence of technological progress and renewability on the sustainability of ecosystem engineers populations
- Author
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Guilherme M Lopes and José F Fontanari
- Subjects
discrete dynamical systems ,human-nature dynamics ,beverton-holt model ,ecosystem engineers ,population collapse ,Biotechnology ,TP248.13-248.65 ,Mathematics ,QA1-939 - Abstract
Overpopulation and environmental degradation due to inadequate resource-use are outcomes of human's ecosystem engineering that has profoundly modified the world's landscape. Despite the age-old concern that unchecked population and economic growth may be unsustainable, the prospect of societal collapse remains contentious today. Contrasting with the usual approach to modeling human-nature interactions, which are based on the Lotka-Volterra predator-prey model with humans as the predators and nature as the prey, here we address this issue using a discrete-time population dynamics model of ecosystem engineers. The growth of the population of engineers is modeled by the Beverton-Holt equation with a density-dependent carrying capacity that is proportional to the number of usable habitats. These habitats (e.g., farms) are the products of the work of the individuals on the virgin habitats (e.g., native forests), hence the denomination engineers of ecosystems to those agents. The human-made habitats decay into degraded habitats, which eventually regenerate into virgin habitats. For slow regeneration resources, we find that the dynamics is dominated by rounds of prosperity and collapse, in which the population reaches vanishing small densities. However, increase of the efficiency of the engineers to explore the resources eliminates the dangerous oscillatory patterns of feast and famine and leads to a stable equilibrium that balances population growth and resource availability. This finding supports the viewpoint of growth optimists that technological progress may avoid collapse.
- Published
- 2019
- Full Text
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13. Contributions to nonstationary community theory
- Author
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Peter Chesson
- Subjects
Nonstationary process ,climate change ,threshold exponential model ,Beverton-Holt model ,lottery model ,Environmental sciences ,GE1-350 ,Biology (General) ,QH301-705.5 - Abstract
The study of the role of environmental variation in community dynamics has traditionally assumed that the environment is a stationary stochastic process or a periodic deterministic process. However, the physical environment in nature is nonstationary. Moreover, anthropogenically driven climate change provides a new challenge emphasizing a persistent but frequently ignored problem: how to make predictions about the dynamics of communities when the nonstationarity of the physical environment is recognized. Recent work is providing a path to conclusions with none of the traditional assumptions of environmental stationarity or periodicity. Traditional assumptions about convergence of long-term averages of functions of environmental states can be replaced by assumptions about temporal sums, allowing convergence and persistence of population processes to be demonstrated in general nonstationary environments. These tools are further developed and illustrated here with some simple models of nonstationary community dynamics, including the Beverton-Holt model, the threshold exponential and the lottery model.
- Published
- 2019
- Full Text
- View/download PDF
14. Age structured discrete-time disease models with demographic population cycles.
- Author
-
van den Driessche, P. and Yakubu, Abdul-Aziz
- Subjects
- *
BASIC reproduction number , *COMMUNICABLE diseases - Abstract
We use juvenile-adult discrete-time infectious disease models with intrinsically generated demographic population cycles to study the effects of age structure on the persistence or extinction of disease and the basic reproduction number, R 0 . Our juvenile-adult Susceptible-Infectious-Recovered (SIR) and Infectious-Salmon Anemia-Virus (ISA v) models share a common disease-free system that exhibits equilibrium dynamics for the Beverton-Holt recruitment function. However, when the recruitment function is the Ricker model, a juvenile-adult disease-free system exhibits a range of dynamic behaviours from stable equilibria to deterministic period k population cycles to Neimark-Sacker bifurcations and deterministic chaos. For these two models, we use an extension of the next generation matrix approach for calculating R 0 to account for populations with locally asymptotically stable period k cycles in the juvenile-adult disease-free system. When R 0 < 1 and the juvenile-adult demographic system (in the absence of the disease) has a locally asymptotically stable period k population cycle, we prove that the juvenile-adult disease goes extinct whenever R 0 < 1. Under the same period k juvenile-adult demographic assumption but with R 0 > 1 , we prove that the juvenile-adult disease-free period k population cycle is unstable and the disease persists. When R 0 > 1 , our simulations show that the juvenile-adult disease-free period k cycle dynamics drives the juvenile-adult SIR disease dynamics, but not the juvenile-adult ISAv disease dynamics. [ABSTRACT FROM AUTHOR]
- Published
- 2020
- Full Text
- View/download PDF
15. Habitat heterogeneity mediates effects of individual variation on spatial species coexistence.
- Author
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Chen, Dongdong, Liao, Jinbao, Bearup, Daniel, and Li, Zhenqing
- Subjects
- *
COEXISTENCE of species , *SPATIAL variation , *HABITATS , *COMPETITION (Biology) , *HETEROGENEITY - Abstract
Numerous studies have documented the importance of individual variation (IV) in determining the outcome of competition between species. However, little is known about how the interplay between IV and habitat heterogeneity (i.e. variation and spatial autocorrelation in habitat quality) affects species coexistence at the landscape scale. Here, we incorporate habitat heterogeneity into a competition model with IV, in order to explore the mechanism of spatial species coexistence. We find that individual-level variation and habitat heterogeneity interact to promote species coexistence, more obviously at lower dispersal rates. This is in stark contrast to early non-spatial models, which predicted that IV reinforces competitive hierarchies and therefore speeds up species exclusion. In essence, increasing variation in patch quality and/or spatial habitat autocorrelation moderates differences in the competitive ability of species, thereby allowing species to coexist both locally and globally. Overall, our theoretical study offers a mechanistic explanation for emerging empirical evidence that both habitat heterogeneity and IV promote species coexistence and therefore biodiversity maintenance. [ABSTRACT FROM AUTHOR]
- Published
- 2020
- Full Text
- View/download PDF
16. Concluding Remarks
- Author
-
Newman, K. B., Buckland, S. T., Morgan, B. J. T., King, R., Borchers, D. L., Cole, D. J., Besbeas, P., Gimenez, O., Thomas, L., Robinson, Andrew P., Series editor, Buckland, Stephen T., Series editor, Reich, Peter, Series editor, McCarthy, Michael, Series editor, Newman, K. B., Buckland, S. T., Morgan, B. J. T., King, R., Borchers, D. L., Cole, D. J., Besbeas, P., Gimenez, O., and Thomas, L.
- Published
- 2014
- Full Text
- View/download PDF
17. Contributions to nonstationary community theory.
- Author
-
Chesson, Peter
- Subjects
- *
STATIONARY processes , *STOCHASTIC processes , *DETERMINISTIC processes , *THERMODYNAMIC state variables , *COMMUNITIES - Abstract
The study of the role of environmental variation in community dynamics has traditionally assumed that the environment is a stationary stochastic process or a periodic deterministic process. However, the physical environment in nature is nonstationary. Moreover, anthropogenically driven climate change provides a new challenge emphasizing a persistent but frequently ignored problem: how to make predictions about the dynamics of communities when the nonstationarity of the physical environment is recognized. Recent work is providing a path to conclusions with none of the traditional assumptions of environmental stationarity or periodicity. Traditional assumptions about convergence of long-term averages of functions of environmental states can be replaced by assumptions about temporal sums, allowing convergence and persistence of population processes to be demonstrated in general nonstationary environments. These tools are further developed and illustrated here with some simple models of nonstationary community dynamics, including the Beverton-Holt model, the threshold exponential and the lottery model. [ABSTRACT FROM AUTHOR]
- Published
- 2019
- Full Text
- View/download PDF
18. Variation between individuals fosters regional species coexistence.
- Author
-
Uriarte, María and Menge, Duncan
- Subjects
- *
COEXISTENCE of species , *POPULATION , *COMPETITION (Biology) , *HABITAT partitioning (Ecology) , *SPATIAL variation - Abstract
Abstract: Although individual‐level variation (IV) is ubiquitous in nature, it is not clear how it influences species coexistence. Theory predicts that IV will hinder coexistence but empirical studies have shown that it can facilitate, inhibit, or have a neutral effect. We use a theoretical model to explore the consequences of IV on local and regional species coexistence in the context of spatial environmental structure. Our results show that individual variation can have a positive effect on species coexistence and that this effect will critically depend on the spatial structure of such variation. IV facilitates coexistence when a negative, concave‐up relationship between individuals’ competitive response and population growth rates propagates to a disproportionate advantage for the inferior competitor, provided that each species specialises in a habitat. While greater variation in the preferred habitat generally fosters coexistence, the opposite is true for non‐preferred habitats. Our results reconcile theory with empirical findings. [ABSTRACT FROM AUTHOR]
- Published
- 2018
- Full Text
- View/download PDF
19. Quantifying the nature and strength of intraspecific density dependence in Arctic mosquitoes
- Author
-
Melissa H. DeSiervo, Matthew P. Ayres, and Lauren E. Culler
- Subjects
0106 biological sciences ,Beverton–Holt model ,education.field_of_study ,Ecology ,010604 marine biology & hydrobiology ,fungi ,Population ,15. Life on land ,Biology ,010603 evolutionary biology ,01 natural sciences ,Intraspecific competition ,Predation ,Density dependence ,Habitat ,Population model ,Abundance (ecology) ,education ,Ecology, Evolution, Behavior and Systematics - Abstract
Processes that change with density are inherent in all populations, yet quantifying density dependence with empirical data remains a challenge. This is especially true for animals recruiting in patchy landscapes because heterogeneity in habitat quality in combination with habitat choice can obscure patterns expected from density dependence. Mosquitoes (Diptera: Culicidae) typically experience strong density dependence when larvae compete for food, however, effects vary across species and contexts. If populations experience intense intraspecific density-dependent mortality then overcompensation can occur, where the number of survivors declines at high densities producing complex endogenous dynamics. To seek generalizations about density dependence in a widespread species of Arctic mosquito, Aedes nigripes, we combined a laboratory experiment, field observations, and modeling approaches. We evaluated alternative formulations of discrete population models and compared best-performing models from our lab study to larval densities from ponds in western Greenland. Survivorship curves from the lab were the best fit by a Hassell model with compensating density dependence (equivalent to a Beverton-Holt model) where peak recruitment ranged from 8 to 80 mosquitoes per liter depending on resource supply. In contrast, our field data did not show a signal of strong density dependence, suggesting that other processes such as predation may lower realized densities in nature, and that expected patterns may be obscured because larval abundance covaries with resources (cryptic density dependence). Our study emphasizes the importance of covariation between the environment, habitat choice, and density dependence in understanding population dynamics across landscapes, and demonstrates the value of pairing lab and field studies.
- Published
- 2021
- Full Text
- View/download PDF
20. Baranov's contributions to the Beverton–Holt model
- Author
-
Trevor J Kenchington
- Subjects
0106 biological sciences ,Beverton–Holt model ,Ecology ,010604 marine biology & hydrobiology ,Aquatic Science ,Oceanography ,010603 evolutionary biology ,01 natural sciences ,Ecology, Evolution, Behavior and Systematics - Abstract
The core of Beverton and Holt’s seminal work of the 1950s had been anticipated by Baranov 30 years earlier, but his contributions, published in Russian, remained poorly known to Anglophone scientists until after 1945. By that time, Russell and Graham had presented parallel ideas, though without Baranov’s mathematics. Limited subsequent acknowledgement of the Russian’s contributions has left an impression that each of his advances was achieved independently in England, hence that they contributed little to the Beverton–Holt model. I here construct a timeline of events linking and separating the Russian and English studies. From it, I argue that both Russell’s presentation of the underlying concepts and Graham’s decision to pursue a mathematical realization of his “Modern Theory” may have been influenced by translations of Baranov’s papers. Moreover, when Holt developed the “simple” Beverton–Holt model during 1946–1947, he certainly drew on Baranov’s exponential model of mortality, though perhaps only via Ricker.
- Published
- 2021
- Full Text
- View/download PDF
21. Sidney Holt on principles for the conservation of wild living resources, whaling in the Antarctic, and the Beverton–Holt stock–recruitment relationship
- Author
-
Marc Mangel
- Subjects
0106 biological sciences ,Fishery ,Beverton–Holt model ,Geography ,Ecology ,010604 marine biology & hydrobiology ,Whaling ,Aquatic Science ,Oceanography ,010603 evolutionary biology ,01 natural sciences ,Ecology, Evolution, Behavior and Systematics ,Stock (geology) - Abstract
I review my interactions with Sidney Holt concerning principles for the conservation of wild living resources, the whaling case between Australia and Japan in the International Court of Justice, and the Beverton–Holt stock–recruitment relationship (BH-SRR). Holt and Lee Talbot published a monograph on principles for conservation in 1977; I lead the publication of an update ∼20 years later. I compare the two versions and discuss Holt’s contributions. Holt was active in the world-wide campaign to cease whaling and in efforts to have the Japanese special permit whaling programme in the Antarctic recognized as violating the moratorium on commercial whaling. I describe my involvement in the case and my interactions with him during oral arguments in the case and when the International Court of Justice rendered its decision that the Japanese programme of special permit whaling contravened the international treaty for the regulation of whaling because it was not for purposes of scientific research. In response to a paper of mine concerning steepness, Holt wrote to me that the BH-SSR is a one-, not two-, parameter function. I explain my current understanding of his reasoning, which involves how we use the SRR in fishery management.
- Published
- 2020
- Full Text
- View/download PDF
22. Dynamics and optimal Harvesting strategy for biological models with Beverton â€'Holt growth
- Author
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Oday Kassim Shalsh and Sadiq Al-Nassir
- Subjects
Beverton–Holt model ,Work (thermodynamics) ,Maximum principle ,General Computer Science ,Applied mathematics ,General Chemistry ,Function (mathematics) ,Stability (probability) ,General Biochemistry, Genetics and Molecular Biology ,Mathematics - Abstract
In this work, the dynamic behavior of discrete models is analyzed with Beverton- Holt function growth . All equilibria are found . The existence and local stability are investigated of all its equilibria.. The optimal harvest strategy is done for the system by using Pontryagin’s maximum principle to solve the optimality problem. Finally numerical simulations are used to solve the optimality problem and to enhance the results of mathematical analysis
- Published
- 2020
- Full Text
- View/download PDF
23. Cumulative effects of incorrect use of pesticides can lead to catastrophic outbreaks of pests.
- Author
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Wang, Xia, Xu, Zihui, Tang, Sanyi, and Cheke, Robert A.
- Subjects
- *
APPLICATION of pesticides , *PERTURBATION theory , *EQUILIBRIUM , *PEST control , *DECISION making - Abstract
Modeling external perturbations such as chemical control within each generation of discrete populations is challenging. Based on a method proposed in the literature, we have extended a discrete single species model with multiple instantaneous pesticide applications within each generation, and then discuss the existence and stability of the unique positive equilibrium. Further, the effects of the timing of pesticide applications and the instantaneous killing rate on the equilibrium were investigated in more detail and we obtained some interesting results, including a paradox and the cumulative effects of the incorrect use of pesticides on pest outbreaks. In order to show the occurrences of the paradox and of hormesis, several special models have been extended and studied. Further, the biological implications of the main results regarding successful pest control are discussed. All of the results obtained confirm that the cumulative effects of incorrect use of pesticides may result in more severe pest outbreaks and thus, in order to avoid a paradox in pest control, control strategies need to be designed with care, including decisions on the timing and number of pesticide applications in relation to the effectiveness of the pesticide being used. [ABSTRACT FROM AUTHOR]
- Published
- 2017
- Full Text
- View/download PDF
24. Almost periodic stochastic Beverton-Holt difference equation with higher delays and with competition between overlapping generations
- Author
-
Paul H. Bezandry
- Subjects
beverton-holt difference equation ,Statistics and Probability ,Beverton–Holt model ,Numerical Analysis ,secondary 34f05 ,Differential equation ,Applied Mathematics ,Quantum Physics ,Overlapping generations model ,time delay ,Competition (economics) ,exponential dichotomy ,QA1-939 ,Applied mathematics ,primary 39a10, 60g07 ,almost periodic sequence ,Mathematics ,Analysis - Abstract
The paper studies the existence of an almost periodic solution of some system of stochastic Beverton-Holt equation with higher delays and with competition between overlapping generations under some reasonable assumptions.
- Published
- 2020
- Full Text
- View/download PDF
25. Age structured discrete-time disease models with demographic population cycles
- Author
-
Abdul-Aziz Yakubu and P. van den Driessche
- Subjects
Adult ,Beverton–Holt model ,Aging ,Time Factors ,Adolescent ,Age structure ,Population Dynamics ,Disease ,Biology ,Communicable Diseases ,Models, Biological ,01 natural sciences ,adults ,Humans ,beverton-holt model ,0101 mathematics ,Age structured ,lcsh:QH301-705.5 ,Ecology, Evolution, Behavior and Systematics ,lcsh:Environmental sciences ,Demography ,lcsh:GE1-350 ,Ecology ,010102 general mathematics ,Age Factors ,Ricker model ,010101 applied mathematics ,Discrete time and continuous time ,lcsh:Biology (General) ,Infectious disease (medical specialty) ,ricker model ,Population cycle ,juveniles ,population cycles ,Disease Susceptibility - Abstract
We use juvenile-adult discrete-time infectious disease models with intrinsically generated demographic population cycles to study the effects of age structure on the persistence or extinction of disease and the basic reproduction number, $\mathcal {R}_{0} $. Our juvenile-adult Susceptible-Infectious-Recovered (SIR) and Infectious-Salmon Anemia-Virus (ISA $v) $ models share a common disease-free system that exhibits equilibrium dynamics for the Beverton-Holt recruitment function. However, when the recruitment function is the Ricker model, a juvenile-adult disease-free system exhibits a range of dynamic behaviours from stable equilibria to deterministic period k population cycles to Neimark-Sacker bifurcations and deterministic chaos. For these two models, we use an extension of the next generation matrix approach for calculating $\mathcal {R}_{0} $ to account for populations with locally asymptotically stable period k cycles in the juvenile-adult disease-free system. When $\mathcal {R}_{0} \lt 1 $ and the juvenile-adult demographic system (in the absence of the disease) has a locally asymptotically stable period k population cycle, we prove that the juvenile-adult disease goes extinct whenever $\mathcal {R}_{0} \lt 1 $. Under the same period k juvenile-adult demographic assumption but with $\mathcal {R}_{0} \gt 1 $, we prove that the juvenile-adult disease-free period k population cycle is unstable and the disease persists. When $\mathcal {R}_{0} \gt 1 $, our simulations show that the juvenile-adult disease-free period k cycle dynamics drives the juvenile-adult SIR disease dynamics, but not the juvenile-adult ISAv disease dynamics.
- Published
- 2020
26. Extending integrated stock assessment models to use non-depensatory three-parameter stock-recruitment relationships
- Author
-
Jason M. Cope and André E. Punt
- Subjects
0106 biological sciences ,Beverton–Holt model ,education.field_of_study ,Stock assessment ,Ecology ,010604 marine biology & hydrobiology ,Maximum sustainable yield ,Population size ,Population ,04 agricultural and veterinary sciences ,Aquatic Science ,01 natural sciences ,Depensation ,040102 fisheries ,Econometrics ,0401 agriculture, forestry, and fisheries ,Environmental science ,Groundfish ,education ,Stock (geology) - Abstract
Stock assessments based on the integrated paradigm often include an underlying stock-recruitment relationship. This, along with estimates of fishery selectivity and biological parameters, allows the biomass and fishing mortality associated with Maximum Sustainable Yield (BMSY and FMSY respectively) to be calculated. However, the estimates of these quantities may differ from the proxies assumed in the harvest control rules that are used to provide management advice. Moreover, the estimated values for BMSY and FMSY are related functionally in population dynamics models based on 2-parameter stock-recruitment relationships such as the commonly used Beverton-Holt or Ricker relationships. Use of 2-parameter stock-recruitment relationships (SRRs) consequently restricts the ability to fully quantify the uncertainty associated with estimating BMSY and FMSY because 2-parameter SRRs restrict the potential range of values for BMSY/B0. In principle, BMSY/B0 and FMSY can be more independent if the stock-recruitment relationship is more general than these 2-parameter SRRs. This paper outlines eleven potential 3-parameter stock-recruitment relationships and evaluates them in terms of whether they are able to match a wide range of specifications for BMSY (expressed relative to unfished spawning stock biomass, B0) and FMSY (expressed relative to natural mortality, M). Of the eleven 3-parameter stock-recruitment relationships considered, the Ricker-Power stock-recruitment relationship is found to best satisfy the characteristics of (a) being able to mimic a wide range of BMSY/B0 and FMSY/M values, (b) not to lead to negative recruitment for biomasses between 0 and B0, and (c) not to lead to increasing recruitment while approaching the limit of zero population size. Bayesian assessments of three example groundfish species off the US west coast (aurora rockfish, petrale sole, and cabezon) are conducted using Simple Stock Synthesis based on the Beverton-Holt and Ricker-Power stock-recruitment relationships to illustrate some of the impacts of allowing for a 3-parameter stock-recruitment relationship.
- Published
- 2019
- Full Text
- View/download PDF
27. Attenuation in the almost periodic Beverton-Holt equation
- Author
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Robert J. Sacker and Cymra Haskell
- Subjects
Beverton–Holt model ,Algebra and Number Theory ,Differential equation ,Discrete dynamics ,Applied Mathematics ,Attenuation ,010102 general mathematics ,Mathematical analysis ,01 natural sciences ,010101 applied mathematics ,Carrying capacity ,Ergodic theory ,0101 mathematics ,Analysis ,Mathematics - Abstract
It is known that the Beverton-Holt equation with periodically varying carrying capacity has a globally attracting solution and the solution exhibits attenuation, i.e. the average of the sol...
- Published
- 2019
- Full Text
- View/download PDF
28. Interplay between strong Allee effect, harvesting and hydra effect of a single population discrete-time system.
- Author
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Jana, Debaldev and Elsayed, Elsayed M.
- Subjects
- *
ALLEE effect , *DENSITY dependence (Ecology) , *POPULATION ecology , *POPULATION density , *BIOLOGICAL extinction - Abstract
The dynamics of a single population with non-overlapping generations can be described deterministically by a scalar difference equation in this study. A discrete-time Beverton-Holt stock recruitment model with Allee effect, harvesting and hydra effect is proposed and studied. Model with strong Allee effect results from incorporating mate limitation in the Beverton-Holt model. We show that these simple models exhibit some interesting (and sometimes unexpected) phenomena such as the hydra effect, sudden collapses and essential extinction. Along with this, harvesting is a socio-economic issue to continue any system generation after generation. Different dynamical behaviors for these situations have been illustrated numerically also. The biological implications of our analytical and numerical findings are discussed critically. [ABSTRACT FROM AUTHOR]
- Published
- 2016
- Full Text
- View/download PDF
29. On the Properties of a Class of Impulsive Competition Beverton–Holt Equations
- Author
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Aitor J. Garrido, Asier Ibeas, Santiago Alonso-Quesada, Izaskun Garrido, and Manuel De la Sen
- Subjects
Beverton–Holt model ,Technology ,QH301-705.5 ,QC1-999 ,Parameterized complexity ,Classification of discontinuities ,impulsive equations ,non-negativity ,Discrete Beverton-Holt equation ,Exponential stability ,Applied mathematics ,Quantitative Biology::Populations and Evolution ,General Materials Science ,competition Beverton–Holt equations ,Logistic function ,Biology (General) ,Instrumentation ,Finite set ,QD1-999 ,Mathematics ,Fluid Flow and Transfer Processes ,Equilibrium point ,Extinction ,Competition ,Process Chemistry and Technology ,Physics ,General Engineering ,Impulsive equations ,difference equations ,boundedness ,Engineering (General). Civil engineering (General) ,discrete Beverton–Holt equation ,impulsive equation ,Computer Science Applications ,Chemistry ,Beverton-Holt equations ,equilibrium points ,TA1-2040 - Abstract
This paper is devoted to a type of combined impulsive discrete Beverton–Holt equations in ecology when eventual discontinuities at sampling time instants are considered. Such discontinuities could be interpreted as impulses in the corresponding continuous-time logistic equations. The set of equations involve competition-type coupled dynamics among a finite set of species. It is assumed that, in general, the intrinsic growth rates and the carrying capacities are eventually distinct for the various species. The impulsive parts of the equations are parameterized by harvesting quotas and independent consumptions which are also eventually distinct for the various species and which control the populations’ evolution. The performed study includes the existence of extinction and non-extinction equilibrium points, the conditions of non-negativity and boundedness of the solutions for given finite non-negative initial conditions and the conditions of asymptotic stability without or with extinction of the solutions. This research was supported by the Spanish Government through grant RTI2018-094336-B-100 (MCIU/AEI/FEDER, UE) and by the Basque Government through grant IT1207-19.
- Published
- 2021
30. Economic Dynamics of Russia: Approach Based on the Solow-Swan Model
- Author
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A. Y. Mitrofanov, N. G. Barashov, M. N. Tolmachev, and Andrey Latkov
- Subjects
Beverton–Holt model ,Solow–Swan model ,Macroeconomics ,Penn World Table ,Economics ,Capital services ,Growth model ,Key features ,Human capital ,Economic dynamics - Abstract
The article discusses the construction of a macroeconomic growth model of the Russian economy, which is a modification of the classical Solow-Swan model. Our model comprises four discrete-time econometric equations. Its two key features are the relationship between capital services and GDP growth rate and phenomenological dependence of capital services on the growth rate of the capital stock. Labor dynamics are described by the Beverton-Holt model. The data used are from the Penn World Table version 9.1 (up to 2017). Based on the model, a forecast of the development of the Russian economy up to 2025 is constructed which was found rather realistic. The constructed model allowed to reach several general conclusions concerning the growth path of the Russian economy, including the limited role of large infrastructure projects as growth “drivers”, while the all-important role of human capital was confirmed.
- Published
- 2021
- Full Text
- View/download PDF
31. The Neimark–Sacker Bifurcation and Global Stability of Perturbation of Sigmoid Beverton–Holt Difference Equation
- Author
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Mustafa R. S. Kulenović, E. Pilav, and Connor O’Loughlin
- Subjects
Beverton–Holt model ,education.field_of_study ,Article Subject ,Differential equation ,Population ,Sigmoid function ,Stability (probability) ,Exponential stability ,Modeling and Simulation ,QA1-939 ,Applied mathematics ,Invariant (mathematics) ,education ,Mathematics ,Bifurcation - Abstract
We present the bifurcation results for the difference equation x n + 1 = x n 2 / a x n 2 + x n − 1 2 + f where a and f are positive numbers and the initial conditions x − 1 and x 0 are nonnegative numbers. This difference equation is one of the perturbations of the sigmoid Beverton–Holt difference equation, which is a major mathematical model in population dynamics. We will show that this difference equation exhibits transcritical and Neimark–Sacker bifurcations but not flip (period-doubling) bifurcation since this difference equation cannot have period-two solutions. Furthermore, we give the asymptotic approximation of the invariant manifolds, stable, unstable, and center manifolds of the equilibrium solutions. We give the necessary and sufficient conditions for global asymptotic stability of the zero equilibrium as well as sufficient conditions for global asymptotic stability of the positive equilibrium.
- Published
- 2021
- Full Text
- View/download PDF
32. Optimum capture size of Trichiurus japonicus in the East China Sea.
- Author
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Yuan F, Zhu WB, Wang ZM, Zhu K, Zhou YD, and Xu HX
- Subjects
- Animals, Fisheries, China, Conservation of Natural Resources, Perciformes
- Abstract
Trichiurus japonicus is an economically valuable species in the East China Sea, whose allowable capture size varies with fishing effort in different years. To clarify the optimum capture size for T. japonicus in the East China Sea, we collected samples and data from T. japonicus targeting fishing gears such as trawls, canvas spreader stow nets and longlines from 2016 to 2020. We estimated growth and mortality parameters using the FiSAT II software, and calculated the size limit standards for capture. The results showed that the inflection point of anal length, the critical anal length and the anal length of one-year-old T. japonicus was 382.84, 397.12, and 216.05 mm, respectively. The anal length of maturity was 230.38 mm, and the minimum capture size (anal length) was 219.23 mm. Based on the yield per recruitment analysis using Beverton-Holt model, the current fishery reference points were under overfishing condition ( t
c =0.38 a, F =2.11), and the suggested optimum capture size was 364.64 mm. There would be a sharp decline of T. japonicus catches in the East China Sea if the suggested minimum capture size was substantially higher, which might be non-practical for local fishermen. Therefore, we suggested 220 mm as the capture size limit, which was close to the anal length of one-year-old fish and the anal length of maturity.- Published
- 2023
- Full Text
- View/download PDF
33. Dynamics of a discontinuous discrete Beverton–Holt model.
- Author
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Kocic, V.L. and Kostrov, Y.
- Subjects
- *
DYNAMICAL systems , *OSCILLATIONS , *DIFFERENTIAL equations , *ASYMPTOTIC theory of algebraic ideals , *MATHEMATICAL models , *MATHEMATICS - Abstract
Our aim is to investigate the global asymptotic behaviour, oscillation, periodicity and bifurcation in discontinuous Beverton–Holt type difference equationwherex0>0, functionskandrare discontinuous piecewise constant functionssatisfyingandhis Heaviside function. [ABSTRACT FROM AUTHOR]
- Published
- 2014
- Full Text
- View/download PDF
34. Estimation of biological parameters and yield per recruitment for Coilia nasustaihuensis in Dianshan Lake, Shanghai, China.
- Author
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GAO Chun-xia, TIAN Si-quan, and DAI Xiao-jie
- Abstract
Coilia nasustaihuensis is the most abundant species in Dianshan Lake and plays an important role in the lake ecosystem. From July 2010 to August 2011, a total of 3107 samples of C. nasustaihuensis were collected from Dianshan Lake. Based on length data of these samples, ELEFAN I technique was employed to estimate growth and mortality parameters, and the Beverton-Holt dynamic model was used to evaluate the population dynamics trend for C. nasustaihuensis. Growth of this species was described using a von Bertalanffy model, and the estimated parameters were L
∞ =35. 70 cm, k =0. 54, and t0 =-0. 48 a. The turning point for body mass growth curve of the stock was situated at t =1. 37 a. Natural mortality coefficient M was then estimated using Pauly's empirical equation and found to be 0. 872. Length-converted catch curves were used to estimate the total mortality coefficient Z, which was found to be 2. 121. Accordingly, the fishing mortality coefficient (F) was equal to 1. 249, and the current exploitation rate was 0. 589, suggesting the stock was over-exploited. According to the Beverton-Holt dynamic model, the minimum capture size for C. nasustaihuensis should be 21. 42 cm (age 1. 22 years). [ABSTRACT FROM AUTHOR]- Published
- 2014
35. An alternative delayed population growth difference equation model
- Author
-
Sabrina Streipert and Gail S. K. Wolkowicz
- Subjects
0106 biological sciences ,Beverton–Holt model ,Differential equation ,39A28, 39A30, 39A60 ,Population ,Population Dynamics ,Dynamical Systems (math.DS) ,01 natural sciences ,Models, Biological ,Exponential stability ,Stability theory ,FOS: Mathematics ,Applied mathematics ,Humans ,Contraction mapping ,0101 mathematics ,Logistic function ,Mathematics - Dynamical Systems ,education ,Quantitative Biology - Populations and Evolution ,Population Growth ,Mathematics ,Extinction threshold ,education.field_of_study ,010604 marine biology & hydrobiology ,Applied Mathematics ,010102 general mathematics ,Populations and Evolution (q-bio.PE) ,Agricultural and Biological Sciences (miscellaneous) ,Modeling and Simulation ,FOS: Biological sciences - Abstract
We propose an alternative delayed population growth difference equation model based on a modification of the Beverton-Holt recurrence, assuming a delay only in the growth contribution that takes into account that those individuals that die during the delay, do not contribute to growth. The model introduced differs from existing delay difference equations in population dynamics, such as the delayed logistic difference equation, which was formulated as a discretization of the Hutchinson model. The analysis of our delayed difference equation model identifies an important critical delay threshold. If the time delay exceeds this threshold, the model predicts that the population will go extinct for all non-negative initial conditions and if it is below this threshold, the population survives and its size converges to a positive globally asymptotically stable equilibrium that is decreasing in size as the delay increases. Firstly, we obtain the local stability results by exploiting the special structure of powers of the Jacobian matrix. Secondly, we show that local stability implies global stability using two different techniques. For one set of parameter values, a contraction mapping result is applied, while for the remaining set of parameter values, we show that the result follows by first proving that the recurrence structure is eventually monotonic in each of its arguments., Comment: 11 pages, 3 figures, Appendix: 13 pages
- Published
- 2020
36. On discrete time Beverton-Holt population model with fuzzy environment
- Author
-
Fu Biao Lin, Xiao Ying Zhong, and Qian Hong Zhang
- Subjects
Beverton–Holt model ,Generalization ,Population Dynamics ,Stability (learning theory) ,02 engineering and technology ,Models, Biological ,Fuzzy logic ,Fuzzy Logic ,0502 economics and business ,0202 electrical engineering, electronic engineering, information engineering ,Animals ,Applied mathematics ,Fuzzy number ,Probability ,Mathematics ,Reproduction ,Applied Mathematics ,05 social sciences ,Computational Biology ,General Medicine ,Division (mathematics) ,Computational Mathematics ,Discrete time and continuous time ,Population model ,Modeling and Simulation ,020201 artificial intelligence & image processing ,General Agricultural and Biological Sciences ,Algorithms ,050203 business & management - Abstract
In this work, dynamical behaviors of discrete time Beverton-Holt population model with fuzzy parameters are studied. It provides a flexible model to fit population data. For three different fuzzy parameters and fuzzy initial conditions, according to a generalization of division (g-division) of fuzzy number, it can represent dynamical behaviors including boundedness, global asymptotical stability and persistence of positive solution. Finally, two examples are given to demonstrate the effectiveness of the results obtained.
- Published
- 2019
- Full Text
- View/download PDF
37. Revisiting Beverton–Holt recruitment in the presence of variation in food availability
- Author
-
Brett T. van Poorten, Carl J. Walters, and Josh Korman
- Subjects
0106 biological sciences ,Beverton–Holt model ,Ecology ,Food availability ,010604 marine biology & hydrobiology ,Foraging ,Aquatic Science ,Biology ,010603 evolutionary biology ,01 natural sciences ,Predation ,Variation (linguistics) ,Carrying capacity ,Juvenile ,Trophic level - Abstract
Understanding density-dependent changes in juvenile survival and growth rates is of great importance because these rates determine recovery rates for imperiled populations and/or sustainable harvest rates. Unfortunately, the mechanisms leading to density dependent survival and growth are among the least understood process in biology and fisheries. Previous work has shown that small fish may vary foraging times to achieve a target growth rate, resulting in the well-known Beverton–Holt recruitment function with variation in food availability affected the initial slope of the recruitment curve. We amend their derivation to show that incorporating fish growth under a variety of evolutionary strategies for balancing foraging time and predation risk still leads to recruitment approximately as expected under the Beverton–Holt recruitment model but that changing food availability affects both the initial slope and maximum recruitment level. We demonstrate that when food availability is known to vary over time, these models often result in a more parsimonious alternative than the standard Beverton–Holt function. Further, Beverton–Holt recruitment is expected when foraging times are adjusted to balance fitness gains from growth against mortality risk. Finally, linking recruitment success to food availability warns that species with high scope for density dependent survival (high compensation ratio or steepness) may be extremely sensitive to changes in available food densities. This work emphasizes the sensitivity of stock-recruitment parameters to food availability and strongly suggests a need to carefully monitor lower trophic levels to better understand and predict dramatic changes in juvenile recruitment and carrying capacity.
- Published
- 2018
- Full Text
- View/download PDF
38. Discrete-time models for releases of sterile mosquitoes with Beverton–Holt-type of survivability
- Author
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Yang Li and Jia Li
- Subjects
0106 biological sciences ,Beverton–Holt model ,Mathematical model ,Applied Mathematics ,General Mathematics ,Numerical analysis ,fungi ,Survivability ,Type (model theory) ,01 natural sciences ,Model dynamics ,010101 applied mathematics ,010601 ecology ,Discrete time and continuous time ,parasitic diseases ,Applied mathematics ,0101 mathematics ,Algebra over a field ,Mathematics - Abstract
In this paper, we formulate discrete-time mathematical models for the interactive wild and sterile mosquitoes. Instead of the Ricker-type of nonlinearity for the survival functions, we assume the Beverton–Holt-type in these models. We consider three different strategies for the releases of sterile mosquitoes and investigate the model dynamics. Threshold values for the releases of sterile mosquitoes are derived for all of the models that determine whether the wild mosquitoes are wiped out or coexist with the sterile mosquitoes. Numerical examples are given to demonstrate the dynamics of the models.
- Published
- 2018
- Full Text
- View/download PDF
39. The Beverton–Holt model with periodic and conditional harvesting
- Author
-
Al-Sharawi, Ziyad, Rhouma, Mohamed Ben Haj, Al-Sharawi, Ziyad, and Rhouma, Mohamed Ben Haj
- Abstract
In this theoretical study, we investigate the effect of different harvesting strategies on the discrete Beverton–Holt model in a deterministic environment. In particular, we make a comparison between the constant, periodic and conditional harvesting strategies. We find that for large initial populations, constant harvest is more beneficial to both the population and the maximum sustainable yield. However, periodic harvest has a short-term advantage when the initial population is low, and conditional harvest has the advantage of lowering the risk of depletion or extinction. Also, we investigate the periodic character under each strategy and show that periodic harvesting drives population cycles to be multiples (period-wise) of the harvesting period.
- Published
- 2020
40. Fitting a non-parametric stock–recruitment model in R that is useful for deriving MSY reference points and accounting for model uncertainty.
- Author
-
Cadigan, Noel G.
- Subjects
- *
MARINE sciences , *FISH population measurement , *FISHERY management , *CONFIDENCE intervals , *STATISTICAL hypothesis testing - Abstract
Cadigan, N. G. 2013. Fitting a non-parametric stock–recruitment model in R that is useful for deriving MSY reference points and accounting for model uncertainty. – ICES Journal of Marine Science, 70:56–67.Modelling the relationship between parental stock size and subsequent recruitment of fish to a fishery is often required when deriving reference points, which are a fundamental component of fishery management. A non-parametric approach to estimate stock–recruitment relationships is illustrated using a simulated example and nine case studies. The approach preserves compensatory density dependence in which the recruitment rate monotonically decreases as stock size increases, which is a basic assumption of commonly used parametric stock–recruitment models. The implications of the non-parametric estimates on maximum sustainable yield (MSY) reference points are illustrated. The approach is used to provide non-parametric bootstrapped confidence intervals for reference points. The efficacy of the approach is investigated using simulations. The results demonstrate that the non-parametric approach can provide a more realistic estimation of the stock–recruitment relationship when informative data are available compared with common parametric models. Also, bootstrap confidence intervals for MSY reference points based on different parametric stock–recruitment models often do not overlap. The confidence intervals based on the non-parametric approach tend to be much wider, and reflect better uncertainty due to stock–recruit model choice. [ABSTRACT FROM PUBLISHER]
- Published
- 2013
- Full Text
- View/download PDF
41. Density dependence in group dynamics of a highly social mongoose, Suricata suricatta.
- Author
-
Bateman, Andrew W., Ozgul, Arpat, Coulson, Tim, and Clutton-Brock, Tim H.
- Subjects
- *
MONGOOSES , *POPULATION dynamics , *ANIMAL social behavior , *RAINFALL , *ANIMAL ecology , *ENVIRONMENTAL impact analysis - Abstract
1. For social species, the link between individual behaviour and population dynamics is mediated by group-level demography. 2. Populations of obligate cooperative breeders are structured into social groups, which may be subject to inverse density dependence (Allee effects) that result from a dependence on conspecific helpers, but evidence for population-wide Allee effects is rare. 3. We use field data from a long-term study of cooperative meerkats ( Suricata suricatta; Schreber, 1776) - a species for which local Allee effects are not reflected in population-level dynamics - to empirically model interannual group dynamics. 4. Using phenomenological population models, modified to incorporate environmental conditions and potential Allee effects, we first investigate overall patterns of group dynamics and find support only for conventional density dependence that increases after years of low rainfall. 5. To explain the observed patterns, we examine specific demographic rates and assess their contributions to overall group dynamics. Although per-capita meerkat mortality is subject to a component Allee effect, it contributes relatively little to observed variation in group dynamics, and other (conventionally density dependent) demographic rates - especially emigration - govern group dynamics. 6. Our findings highlight the need to consider demographic processes and density dependence in subpopulations before drawing conclusions about how behaviour affects population processes in socially complex systems. [ABSTRACT FROM AUTHOR]
- Published
- 2012
- Full Text
- View/download PDF
42. The Beverton-Holt model with periodic and conditional harvesting.
- Author
-
AlSharawi, Ziyad and Rhouma, MohamedB.H.
- Subjects
- *
HARVESTING , *MATHEMATICAL models , *POPULATION , *BIOLOGICAL extinction - Abstract
In this theoretical study, we investigate the effect of different harvesting strategies on the discrete Beverton-Holt model in a deterministic environment. In particular, we make a comparison between the constant, periodic and conditional harvesting strategies. We find that for large initial populations, constant harvest is more beneficial to both the population and the maximum sustainable yield. However, periodic harvest has a short-term advantage when the initial population is low, and conditional harvest has the advantage of lowering the risk of depletion or extinction. Also, we investigate the periodic character under each strategy and show that periodic harvesting drives population cycles to be multiples (period-wise) of the harvesting period. [ABSTRACT FROM AUTHOR]
- Published
- 2009
- Full Text
- View/download PDF
43. Sensitivity of common estimators of management parameters derived from stock–recruit relationships
- Author
-
Cadigan, N.G.
- Subjects
- *
FISHERY management , *FISH populations , *RECRUITMENT (Population biology) , *MATHEMATICAL models , *PARAMETER estimation , *MEASUREMENT errors , *ERROR analysis in mathematics , *MATHEMATICAL analysis - Abstract
Abstract: Local influence diagnostics are used to develop a theoretical understanding of the sensitivity of some management parameters derived from the Beverton–Holt and Ricker stock–recruit models using a common estimation method. The diagnostics provide an understanding of how characteristics of the data and model affect estimates. Sensitivity is assessed for the slope at the origin, the maximum recruitment, and the stock size corresponding to 50% of the maximum recruitment. Two types of sensitivity are considered. The first is with respect to the weighting that stock–recruit observation pairs (i.e. cases) are given in estimation. Case-weight local influence equations are developed to provide insights about why estimates change when cases are deleted. The second type of sensitivity is with respect to the stock size measurements. These are commonly assumed to be measured without error when fitting a stock–recruit model, when in fact stock size measurements often have substantial error. Local influence diagnostics are developed to assess the impact of errors in stock size, with a particular focus on measurement errors. The diagnostics are applied to 15 case-studies. [Copyright &y& Elsevier]
- Published
- 2009
- Full Text
- View/download PDF
44. On r-periodic orbits of k-periodic maps.
- Author
-
Beyn, Wolf-Jurgen, Hüls, Thorsten, and Samtenschnieder, Malte-Christopher
- Subjects
- *
COMBINATORIAL dynamics , *DIMENSIONAL preference , *NUMERICAL analysis , *BIFURCATION theory , *DIFFERENCE equations - Abstract
In this paper, we analyze r-periodic orbits of k-periodic difference equations, i.e.[image omitted] and their stability. These special orbits were introduced in S. Elaydi and R.J. Sacker (Global stability of periodic orbits of non-autonomous difference equations and population biology, J. Differ. Equ. 208(1) (2005), pp. 258-273). We discuss that, depending on the values of r and k, such orbits generically only occur in finite dimensional systems that depend on sufficiently many parameters, i.e. they have a large codimension in the sense of bifurcation theory. As an example, we consider the periodically forced Beverton-Holt model, for which explicit formulas for the globally attracting periodic orbit, having the minimal period k = r, can be derived. When r factors k the Beverton-Holt model with two time-variant parameters is an example that can be studied explicitly and that exhibits globally attracting r-periodic orbits. For arbitrarily chosen periods r and k, we develop an algorithm for the numerical approximation of an r-periodic orbit and of the associated parameter set, for which this orbit exists. We apply the algorithm to the generalized Beverton-Holt, the 2D stiletto model, and another example that exhibits periodic orbits with r and k relatively prime. [ABSTRACT FROM AUTHOR]
- Published
- 2008
- Full Text
- View/download PDF
45. The Time Invariance Principle, the absence of ecological chaos, and a fundamental pitfall of discrete modeling
- Author
-
Deng, Bo
- Subjects
- *
LIFE sciences , *POPULATION biology , *ENVIRONMENTAL sciences , *BIOLOGY - Abstract
Abstract: This paper is to show that most discrete models used for population dynamics in ecology are inherently pathological that their predications cannot be independently verified by experiments because they violate a fundamental principle of physics. The result is used to tackle an on-going controversy regarding ecological chaos. Another implication of the result is that all dynamical systems must be modeled by differential equations. As a result it suggests that researches based on discrete modeling must be closely scrutinized and the teaching of calculus and differential equations must be emphasized for students of biology. [Copyright &y& Elsevier]
- Published
- 2008
- Full Text
- View/download PDF
46. On the mechanistic underpinning of discrete-time population models with Allee effect
- Author
-
Eskola, Hanna T.M. and Parvinen, Kalle
- Subjects
- *
POPULATION density , *POPULATION geography , *POPULATION biology , *CANNIBALISM - Abstract
Abstract: The Allee effect means reduction in individual fitness at low population densities. There are many discrete-time population models with an Allee effect in the literature, but most of them are phenomenological. Recently, Geritz and Kisdi [2004. On the mechanistic underpinning of discrete-time population models with complex dynamics. J. Theor. Biol. 228, 261–269] presented a mechanistic underpinning of various discrete-time population models without an Allee effect. Their work was based on a continuous-time resource-consumer model for the dynamics within a year, from which they derived a discrete-time model for the between-year dynamics. In this article, we obtain the Allee effect by adding different mate finding mechanisms to the within-year dynamics. Further, by adding cannibalism we obtain a higher variety of models. We thus present a generator of relatively realistic, discrete-time Allee effect models that also covers some currently used phenomenological models driven more by mathematical convenience. [Copyright &y& Elsevier]
- Published
- 2007
- Full Text
- View/download PDF
47. On the Mechanistic Derivation of Various Discrete-Time Population Models.
- Author
-
Eskola, Hanna T. M. and Geritz, Stefan A. H.
- Subjects
- *
REPRODUCTION , *EXAMPLE , *BEHAVIOR , *EQUILIBRIUM , *POPULATION - Abstract
We present a derivation of various discrete-time population models within a single unifying mechanistic context. By systematically varying the with in year patterns of reproduction and aggression between individuals we can derive various discrete-time population models. These models include classical examples such as the Ricker model, the Beverton-Holt model, the Skellam model, the Hassell model, and others. Some of these models until now lacked a good mechanistic interpretation or have been derived in a different context. By using this mechanistic approach, the model parameters can be interpreted in terms of individual behavior. [ABSTRACT FROM AUTHOR]
- Published
- 2007
- Full Text
- View/download PDF
48. Supply regimes in fisheries.
- Author
-
Nielsen, Max
- Subjects
FISHERY management ,COD fisheries ,NATURAL resources management ,FISHERIES - Abstract
Abstract: Supply in fisheries is traditionally known for its backward bending nature, owing to externalities in production. Such a supply regime, however, exist only for pure open access fisheries. Since most fisheries worldwide are neither pure open access, nor optimally managed, rather between the extremes, the traditional understanding of supply regimes in fisheries needs modification. This paper identifies through a case study of the East Baltic cod fishery supply regimes in fisheries, taking alternative fisheries management schemes and mesh size limitations into account. An age-structured Beverton–Holt based bio-economic supply model with mesh sizes is developed. It is found that in the presence of realistic management schemes, the supply curves are close to vertical in the relevant range. Also, the supply curve under open access with mesh size limitations is almost vertical in the relevant range, owing to constant recruitment. The implications are that the effects on supply following from e.g. trade liberalisation and reductions of subsidies are small in several and probably most fisheries worldwide. [Copyright &y& Elsevier]
- Published
- 2006
- Full Text
- View/download PDF
49. A note on the nonautonomous Beverton-Holt model.
- Author
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Kocic, V. L.
- Subjects
- *
MATHEMATICAL models , *EQUATIONS , *MATHEMATICAL inequalities , *INFINITE processes , *PROOF theory , *MATHEMATICS - Abstract
In this paper, we study the asymptotic behavior of positive solutions of nonautonomous Beverton-Holt equationwhere { K n} is a positive persistent and bounded sequence. In our main result, we proved that all positive solutions { x n} satisfy the inequalityThe obtained result represents the extension of the Cushing-Hensons's conjecture. Applications to the case when { K n} is a positive periodic sequence are given and the alternative proof of the Cushing-Henson's conjecture is also presented. [ABSTRACT FROM AUTHOR]
- Published
- 2005
- Full Text
- View/download PDF
50. On the mechanistic underpinning of discrete-time population models with complex dynamics
- Author
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Geritz, Stefan A.H. and Kisdi, Éva
- Subjects
- *
DISCRETE-time systems , *DIFFERENTIABLE dynamical systems , *CONSUMPTION (Economics) , *STATICS - Abstract
We present a mechanistic underpinning for various discrete-time population models that can produce limit cycles and chaotic dynamics. Specific examples include the discrete-time logistic model and the Hassell model, which for a long time eluded convincing mechanistic interpretations, and also the Ricker- and Beverton–Holt models. We first formulate a continuous-time resource consumption model for the dynamics within a year, and from that we derive a discrete-time model for the between-year dynamics. Without influx of resources from the outside into the system, the resulting between-year dynamics is always overcompensating and hence may produce complex dynamics as well as extinction in finite time. We recover a connection between various standard types of continuous-time models for the resource dynamics within a year on the one hand and various standard types of discrete-time models for the population dynamics between years on the other. The model readily generalizes to several resource and consumer species as well as to more than two trophic levels for the within-year dynamics. [Copyright &y& Elsevier]
- Published
- 2004
- Full Text
- View/download PDF
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