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Cangshanaltica javana sp. nov. Fig.s 5, 8D Type locality: Indonesia, Java Barat, Gunung Tangkuban Perahu. Type material. HOLOTYPE (male, MHNG): (1) INDONESIE — East Java 30 km N. Bandung, volcan Tangkuban Prahu, J. Robert 10.VIII.85. ◉ Paratypes (1 male MHNG, 1 male NMPC) same labels as holotype; (2 female MHNG, 1 female NMPC): JAVA Volcan Tangkuban Prahu env 30 km N. Bandung J. Robert 7. VIII. 84. Differential diagnosis. The species can be easily distinguished from all known species by its genitalic morphology (median lobe of aedeagus pointed with sharp parallel margins on dorsal side, spermathecal duct long, U shaped). Additionally, the species can be distinguished by the key below. Description. Habitus. Body ovate, convex, 2.4 mm long, 1.7 mm wide, 1.1 mm high. Dorsal surfaces dark blackish-brown without metallic luster. Ventral side brown. Head. Nearly hypognathous. Labrum light brown, deeply incised; clypeus black. Labial and maxillary palpi light brown. Frons wide, brown, flat, frontal ridge absent. Suprafrontal sulcus well-developed, frontal calli feebly projecting, flat, feebly delimited posteriorly. Orbit narrow, orbital and supraorbital sulci deep. Vertex very finely punctured with shallow scattered punctures. Antennae reaching basal ¼ of elytra. Antennomeres I–IV and XI light yellowish-brown, antennomeres V–X brown. Antennomeres V–XI gradually widening, flat. Thorax. Pronotum wide, convex, punctured with small scattered punctures. Anterior pronotal margins feebly projecting anteriorly, but forming lobes. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra convex, about ¼ wider than maximal width of pronotum, irregularly punctured with distinct punctures twice as large as those on pronotum. Epipleura extremely widened in basal parts of elytra, nearly as wide as half of pronotal width. Procoxal cavity widely open posteriorly. Anterior metaventral process completely covering the mesoventrite between mesocoxae, as a deeply excavated “horseshoe-like” structure. Legs light brown, metafemora darkened. Metatibiae slightly curved laterally, with a long apical spine. Metatarsomere I elongated. Abdomen. Abdominal ventrite I as long as ventrites II and III combined. Anterior process of abdominal ventrite I reaching space between metacoxae, bulbose, forming a ventrally projecting elevated ridge. Genitalia. Median lobe of aedeagus long, slender, pointed apically, slightly curved in lateral view. Dorsal side of median lobe of aedeagus with sharp parallel margins narrowing in the middle, ventral side not narrowed. Spermathecal pump as long as receptacle, with digitiform apical part; receptacle twice as wide as pump, almost parallel-sided. Spermathecal duct long, U-shaped, reaching the apical 1/3 of the receptacle. Etymology. The species is named after the island of Java, where the type series was collected.

Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge & Yang, 2013 Figs 2, 8H Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge & Yang, 2013: 16 Type locality: China, Yunnan prov., Dali, Cangshan (Ẽ山). Type series: Holotype — IZAS, paratypes: IZAS, USNM. Type material examined. PARATYPE (1 male, USNM): China: Yunnan, env. Dali, Cangshan Mountains, 3478 m, 19 July 2011, 25°41.100′ N, 100°06.228′ E, moss, Konstantinov, Chamorro and Volkovitsh; (2) Paratype Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge, Yang, sp. nov., 2011. Additional material examined. 1 female (UABG): CHINA (Yunnan) Dali Bai Aut. Pref., Jizu Shan, summit plateau, 37 km NE Dali 3150 m 25°58′30″N, 100°21′36″E (mixed forest, sifted from litter, moss) 5. IX. 2009 D. W. Wrase [28]., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on pages 58-59, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Konstantinov, A., Chamorro, M. L., Prathapan, K. D., Ge, S. - Q. & Yang, X. - K. (2013) Moss-inhabiting flea beetles (Coleoptera: Chrysomelidae: Galerucinae: Alticini) with description of a new genus from Cangshan, China. Journal of Natural History, 47 (37 - 38), 2459 - 2477. https: // doi. org / 10.1080 / 00222933.2012.763068"]}

Cangshanaltica luzonica Damaška, Mohagan and Fikáček, 2020 Fig. 8E Cangshanaltica luzonica Damaška, Mohagan and Fikáček, 2020: 127 Type locality: Philippines, Luzon, Sagada. Type series: Holotype — MHNG, paratypes: MHNG, NMPC. Remarks. The species is known only from the type series., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on page 57, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Damaska, A. F., Mohagan, D. J. & Fikacek, M. (2020) Moss-inhabiting flea beetles in the Philippines (Coleoptera, Chrysomelidae, Alticinae). ZooKeys, 960, 125 - 142. https: // doi. org / 10.3897 / zookeys. 960.54011"]}

Cangshanaltica ogatai Takemoto & Suenaga, 2021 Cangshanaltica ogatai Takemoto & Suenaga, 2021: 146 Type locality: Japan, Okinawa, Iriomote isl. Type series: Holotype —SEHU; paratypes: HSPC, TTPC. Remarks. Cangshanaltica ogatai is the only species described, so far, which has an elytral pattern with multiple spots. This coloration is uncommon among Cangshanaltica species distributed in continental Asia. Species with a similar color pattern are distributed also in Taiwan, however, we hesitate to either assign them to C. ogatai or describe them as a new species (see comments below).

Cangshanaltica guanxiensis sp. nov. Figs 4, 8C Type locality: China, Guanxi, Panxie Gou. Type material. HOLOTYPE (male, NMPC): (1) CHINA: Guangxi, Pangxie Gou, 1.3 km S of Longsheng Hot Spring, 25°53.0′N 110°12.6′E, 385 m 13.iv.2013, Fikáček, Hájek & Růžička lgt. (2) sifting of leaf litter and leaf accumulations along stony stream in evergreen lowland subtropical forest [sifting sample 13]; (3) VOUCHER SPECIMEN DNA collection No. AFD-125 Albert F. Damaška coll. Differential diagnosis. The species can be distinguished from all other known Cangshanaltica species by its coloration—elytra ochraceous with one darkened spot in the center of each elytron. DNA sequences. 18S: MZ312403.1; 16S: MZ312387.1; Cox1: MZ303694.1, MZ220536.1 (barcode). All sequences were published by Damaška et al., 2021. Description. Habitus. Body round, convex, 2 mm long, 1.7 mm wide, 0.9 mm high. Dorsal and ventral surfaces ochrebrown, each elytron with a darkened spot in the middle. Head. Nearly hypognathous, triangular. Labrum and clypeus blackish brown, vertex brown.Frontal ridge absent; frons shiny, impunctate, with bunches of setae below antennae. Frontal calli nearly absent. Suprafrontal sulcus shallow, orbital and supraorbital sulci developed. Eyes round. Antennae reaching basal 1/3 of elytra. Antennomeres I–VI and XI light brown, antennomeres V–X darkened. Antennomeres I and II bulbose, wide; antennomeres VII–XI elongate, as long or longer as antennomeres III and IV combined.Antennomere VII bearing a slight distal protrusion. Thorax. Pronotum brownish ochraceous, wide, convex, impunctate. Anterior margins of pronotum lobate, projecting, forming distinct “flap-like” structures towards head; posterior pronotal edges dull. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra convex, brownish ochraceous with a darkened spot in the central part of each elytron, irregularly punctured. Epipleura distinctly widened in basal part. Anterior metaventral process completely covering the mesoventrite between mesocoxae, as a sharply pentagonal, deeply excavated “horseshoe-like” structure. Legs brownish ochraceous. Metatibiae slightly curved, apical metatibial spine short. Metatarsomere I slightly elongated. Abdomen. Abdominal ventrite I longer than ventrite II, but not as long as ventrites II and III combined.Anterior process of abdominal ventrite I reaching space between metacoxae, bulbose, forming a sharp, ventrally projecting elevated ridge. Genitalia. Median lobe of aedeagus long but wide, parallel-sided in dorsal view, slightly curved in lateral view. Apex dull, slightly widening posteriorly. Female unknown. Remarks. The only known specimen is apparently not fully sclerotized—the coloration of a mature adult is probably somewhat darker. Despite that, we are describing the species, as all morphological characters are welldeveloped, and available DNA markers clearly indicate its isolated phylogenetic position among other studied Cangshanaltica species (Damaška et al. 2021). Etymology. The species name is derived from Guanxi, the name of the Chinese province where the holotype was collected.

Cangshanaltica castanea (Gruev, 1985) comb. nov. Figs 3, 8A Taizonia castanea Gruev, 1985: 126 Ivalia gruevi Nadein, 2013: 396 (nom. nov. for castanea Gruev, not Samuelson) Type locality: China: Da-laen-saen (type locality identity is discussed below). Type material examined. HOLOTYPE (male, BMNH): (1) Holotype, (2) G. C. Champion Coll. B. M. 1927- 409, (3) Da-laen-saen, China. J. J. Walker, (4) HOLOTYPE; (5) Taizonia castanea Gruev sp. n.; ◉ PARATYPES (1 male; 4 female, BMNH): (1) Da-laen-saen, China. J. J. Walker, (2) PARATYPE. Additional material examined. 6 spec. (4 male, 2 female, BMNH): (1) G. C. Champion Coll. B. M. 1927-409; (2) Da-laen-saen, China. J. J. Walker; (3) Taizonia castanea Gruev det. A. S. Konstantinov, 2003; 9 spec. (9 male, BMNH): (1) Nyew-tew I., China, J. J. W [alker]; (2) ◉ 4 spec. (3 male, 1 female, NMPC, ADPC): (1) CHINA — Zhejiang prov., Lishui pref., Lu’aocun (炉岙村); Monkey Valley (Ñšẽ); 27°54′55.2″N 119°11′55.4″E; 1100 m; 11. v. 2019; A. F. Damaška lgt.; (2) VOUCHER SPECIMEN DNA collection No. AFD-235; 236; 282 283; 284 Albert F. Damaška coll.; (3) Cangshanaltica cf. castanea (spec. complex), A. F. Damaška det. 2022 ◉ 2 spec. (2 male, SZPT): (1) Hangzhou Wuyun Mt. (杭州五云山), on moss, under tree root, 2021-III-25, leg. Wei Wang (Ɨ ‼). Redescription. Habitus. Body oval-rounded, convex, holotype 1.6 mm long, 1.2 mm wide, 0.9 mm high. Dorsal surfaces chestnut brown without metallic luster. Ventral surfaces light brown. Head. Nearly hypognathous, triangular, chestnut brown. Frontal calli flat, feebly developed, distinctly delimited anteriorly, feebly delimited posteriorly. Frontal ridge wide, flat, not projecting; interantennal space as wide as diameter of eye. Supraantennal, suprafrontal, orbital and supraorbital sulci shallow but developed. Orbit narrow. Frontal ridge, orbit and frons impunctate. Antennae with 11 antennomeres, short, reaching anterior ¼ of elytra. Antennomeres I and II large, wide, light yellowish brown; antennomere III slightly elongated, antennomeres IV–VI similar in length, slender, antennomere VI wider than V. Antennomere VII short, with a distinct distal protrusion; antennomeres VIII–XI gradually widening, forming a long and slender antennal club. Thorax. Pronotum brown, wide, convex, feebly punctate by scattered shallow punctures.Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Posterior edges of pronotum sharp. Elytra convex, wide, feebly irregularly punctured. Maximal width of elytra 1/3 wider than maximal width of pronotum. Procoxal cavity widely open posteriorly.Anterior metaventral process completely covering mesoventrite between mesocoxae, developing a pentagonal “horseshoe-like” structure. Legs light brown. Metatibiae nearly straight, not visibly curved laterally. Metatarsomere I elongated. Abdomen and genitalia. Abdominal ventrite I as long as ventrites II and III combined. Median lobe of aedeagus variable in width, and in sharpness of the apex (Fig. 3G,H). Apex pointed, arrow-like. Median lobe widened from apex to basal part. Basal orifice pyriform, reaching ¼ of the median lobe of aedeagus length. Spermatheca slender, duct without coils. Vaginal palpi widely separated apically. Generic placement. Cangshanaltica castanea had been described in Taizonia by Gruev (1985), a genus subsequently synonymized with Ivalia by Duckett et al. (2006). Due to homonymy with Ivalia castanea (Samuelson, 1966), a new name Ivalia gruevi was proposed for the species by Nadein (2013). Our placement of Gruev’s species in Cangshanaltica makes Nadein’s replacement name redundant (ICZN Art 59.4). We move the species to Cangshanaltica based on the following characters: (1) antennomere VII bearing a slight distal protrusion; (2) metaventrite bearing an anterior, horseshoe-like process reaching mesocoxae and covering the mesoventrite; (3) anterolateral pronotal setiferous pore placed in the middle of the pronotal margin; and (4) round, ovate, and convex body shape. Cangshanaltica castanea is similar to other Chinese Cangshanaltica species distributed in lowlands— C. sprynari, and C. fuanensis. The species can be distinguished using the key below. Type locality identity and distribution. The type locality “ China: Da-laen-saen” named by Walker has been discussed by authors who have revised J. Walker’s Chinese material (Schawaller & Aston 2017; Smetana 2009). According to their findings, the most likely type locality of C. castanea is China, Zhejiang prov., Ningbo pref., Dalei Shan (kDZ山), cca 29.5928 N, 121.2930 E. The second Walker locality in the BMNH material is “Nyew-tew I.” This locality refers most likely to the Nantian Dao Island (Éffl岛) in Zhejiang province, named “Nyew-tew Island” in 19th century Western maps, likely after the island’s peak Niutou Shan (牛Ë山). The second type locality should therefore be: China, Zhejiang prov., Ningbo pref., Nantian Dao (Éffl岛); cca 29.1361 N, 121.9388 E. Additional specimens assigned to the species complex of C. castanea were collected recently in Lu’aocun (炉岙村) in the Lishui prefecture, and in Hangzhou, both in Zhejiang province, China. Species identity and variability. Cangshanaltica castanea was described based on a type series collected by J. J. Walker. Besides the type series, we examined additional Walker material from the type locality (Da Laen Saen) and its surroundings (Nyew-tew I.). We also collected fresh DNA-grade specimens in southern Zhejiang —they strongly resemble several specimens from the C. castanea type series. All the specimens have similar morphology of external characters and female genitalia, however, the morphology of median lobe of aedeagus is quite variable among all specimens, even within the type material (fig. 3 G–H). Specimens also differ in elytral punctuation (in the holotype and most of Walker’s specimens, elytra have very feeble punctures, in several of Walker’s specimens and all the fresh specimens from Lishui, punctures are shallow but distinctly visible, forming a pitted elytral relief, similar to the surface of a golf ball), and size (males are up to 1.9 mm long). A similar species in external morphology is C. fuanensis Ruan, Konstantinov & Damaška, 2022, which differs in having a more slender median lobe of aedeagus, but is strongly divergent from the Lishui specimens based on DNA phylogenetic analyses (Damaška et al. 2021). This strong difference in both mitochondrial and nuclear DNA markers among the two morphologically similar species suggests that the C. castanea species complex can consist of more species which are similar in morphology but phylogenetically isolated from each other. Due to the lack of any DNA-grade material from the two historical collecting sites in Zhejiang, we decided not to treat any of the specimens with different genitalia morphology as distinct species. However, the variability of median lobe of aedeagus in C. castanea in our understanding is apparently larger than in other Cangshanaltica species. The species definition of C. castanea may therefore represent a species complex. A careful analysis of DNA and morphology based on fresh material from more localities in Zhejiang and around (including Walker’s sites) is the only way to yield enough information to resolve the C. castanea species complex.

Cangshanaltica ogatai Takemoto & Suenaga, 2021 Cangshanaltica ogatai Takemoto & Suenaga, 2021: 146 Type locality: Japan, Okinawa, Iriomote isl. Type series: Holotype —SEHU; paratypes: HSPC, TTPC. Remarks. Cangshanaltica ogatai is the only species described, so far, which has an elytral pattern with multiple spots. This coloration is uncommon among Cangshanaltica species distributed in continental Asia. Species with a similar color pattern are distributed also in Taiwan, however, we hesitate to either assign them to C. ogatai or describe them as a new species (see comments below)., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on page 59, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Takemoto, T. & Suenaga, H. (2021) Four New Species of the Genus Ivalia Jacoby and a New Species of Cangshanaltica Konatantinov, Chamorro, Prathapan, Ge et Yang (Coleoptera, Chrysomelidae, Galerucinae) from the Ryukyus, Southwestern Japan. Elytra, 11, 129 - 153."]}

Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge & Yang, 2013 Figs 2, 8H Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge & Yang, 2013: 16 Type locality: China, Yunnan prov., Dali, Cangshan (Ẽ山). Type series: Holotype — IZAS, paratypes: IZAS, USNM. Type material examined. PARATYPE (1 male, USNM): China: Yunnan, env. Dali, Cangshan Mountains, 3478 m, 19 July 2011, 25°41.100′ N, 100°06.228′ E, moss, Konstantinov, Chamorro and Volkovitsh; (2) Paratype Cangshanaltica nigra Konstantinov, Chamorro, Prathapan, Ge, Yang, sp. nov., 2011. Additional material examined. 1 female (UABG): CHINA (Yunnan) Dali Bai Aut. Pref., Jizu Shan, summit plateau, 37 km NE Dali 3150 m 25°58′30″N, 100°21′36″E (mixed forest, sifted from litter, moss) 5. IX. 2009 D. W. Wrase [28].

Cangshanaltica marginata sp. nov. Figs 6, 8F Type locality: China, Yunnan, 24°29′31″N 98°52′57″E Type material. HOLOTYPE (male, DEIM): (1) CHINA (Yunnan), Dehong Dai Aut. Pref., mount. range 31 km E Luxi, 2280 m, 24°29′31″N / 98°52′58″E (grassland/pasture, under shrubs, moss/litter, sifted) 3. VI. 2007 D. W. Wrase [19A]; (2) HOLOTYPE Cangshanaltica marginata sp. nov., Damaška, Ruan, Fikáček des. 2020. ◉ Paratypes (1 male, 2 female UABG, 1 male, 1 female NMPC, 1 female ADPC) have same labels as holotype. Differential diagnosis. The species can be distinguished from all other known Cangshanaltica species by its coloration—elytra black with fulvous margin. Additionally, it can be distinguished from all other known species by the shape of the aedeagal apex (widened with a narrow tip). Description. Habitus. Body oblong-ovate, convex, holotype 1.8 mm long, 1.2 mm wide, 1 mm high. Head and pronotum brown, elytra black with elytral suture brown, basal and humeral margins of elytra with wide yellow spots, lateral elytral margins and apices of elytra yellow. Ventral side of head and prothorax light yellowish brown, ventral side of mesothorax, metathorax and abdomen brown. Head. Nearly hypognathous. Labrum pitchy black, widely, but not deeply incised. Maxillary and labial palpi brown. Clypeus and frons light brown, feebly pilose, frontal ridge absent. Suprafrontal sulcus shallow, frontal calli almost indistinct. Orbital and supraorbital sulci developed but shallow. Orbit and space around eyes slightly darker than vertex and frons in some specimens. Vertex shiny, almost impunctate. Eyes round. Antennae reaching basal ¼ of elytra. Antennomeres I–V and XI light brown, antennomeres VI–X dark brown. Antennomere VII slightly protruded. Thorax. Pronotum wide, convex, impunctate. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra slightly, irregularly punctured. Humeral part of lateral elytral margin and basal elytral margin with two wide light brownish yellow spots; lateral elytral margin with a wide yellowish-brown stripe; pleura brown. Elytral apices yellow. Procoxal cavity widely open posteriorly; prosternal intercoxal process elevated ventrally. Anterior metaventral process completely covered by mesoventrite between mesocoxae, with a deeply excavated, upturned U-shaped “horseshoe-like” structure. Legs brown. Tibiae feebly but distinctly pilose. Metatibiae slightly curved laterally, with a long apical spine. Metatarsomere I not strongly elongated. Abdomen. Ventrites slightly pilose. Abdominal ventrite I with a bulbose anterior process reaching the space between metaxoxae, with a slightly elevated ridge. Abdominal ventrite V slightly paler than ventrites I–IV. Genitalia. Median lobe of aedeagus short, wide and slightly curved in lateral view; in ventral view slightly widened towards apex in basal 2/3, apical 1/3 of median lobe of aedeagus narrowing, triangular, apex narrowly rounded. Spermatheca slender; spermathecal pump short, receptacle elongate-oval, spermathecal duct shortly Ushaped, without coils, ending in ½ of the receptacle. Vaginal palpi short, joined basally, V-shaped, slightly separated apically. Tignum slender. Etymology. The species is named after its specific elytral color pattern—black with yellow lateral margin (marginata —with notable margin).

Cangshanaltica mindanaoensis Damaška, Mohagan and Fikáček, 2020 Fig. 8G Cangshanaltica mindanaoensis Damaška, Mohagan and Fikáček, 2020: 129 Type locality: Philippines, Mindanao, Mt. Hamiguitan. Type series: Holotype — NMPC; paratypes: NMPC, ADPC, USNM. Remarks. The species is described from moss-sifting samples in Mt. Hamiguitan, Mindanao, Philippines. A probable conspecific specimen is known from Mt. Musuan, Mindanao. In Damaška et al. (2021) the species was sister to all other Cangshanaltica species were included in their analyses (C. siamensis, C. fuanensis, C. guanxiensis, C. castanea, and C. nr. ogatai from Taiwan)., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on page 58, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Damaska, A. F., Mohagan, D. J. & Fikacek, M. (2020) Moss-inhabiting flea beetles in the Philippines (Coleoptera, Chrysomelidae, Alticinae). ZooKeys, 960, 125 - 142. https: // doi. org / 10.3897 / zookeys. 960.54011","Damaska, A. F., Konstantinov, A., Lee, C. - F., Ruan, Y., Mohagan, D. J. & Fikacek, M. (2021) Molecular phylogeny of mossinhabiting flea beetles from the Chabria group (Coleoptera: Chrysomelidae: Alticini) reveals multiple colonizations and radiations in Taiwan. Systematic Entomology, 46 (4), 915 - 925. https: // doi. org / 10.1111 / syen. 12502"]}

Cangshanaltica marginata sp. nov. Figs 6, 8F Type locality: China, Yunnan, 24°29′31″N 98°52′57″E Type material. HOLOTYPE (male, DEIM): (1) CHINA (Yunnan), Dehong Dai Aut. Pref., mount. range 31 km E Luxi, 2280 m, 24°29′31″N / 98°52′58″E (grassland/pasture, under shrubs, moss/litter, sifted) 3. VI. 2007 D. W. Wrase [19A]; (2) HOLOTYPE Cangshanaltica marginata sp. nov., Damaška, Ruan, Fikáček des. 2020. ◉ Paratypes (1 male, 2 female UABG, 1 male, 1 female NMPC, 1 female ADPC) have same labels as holotype. Differential diagnosis. The species can be distinguished from all other known Cangshanaltica species by its coloration—elytra black with fulvous margin. Additionally, it can be distinguished from all other known species by the shape of the aedeagal apex (widened with a narrow tip). Description. Habitus. Body oblong-ovate, convex, holotype 1.8 mm long, 1.2 mm wide, 1 mm high. Head and pronotum brown, elytra black with elytral suture brown, basal and humeral margins of elytra with wide yellow spots, lateral elytral margins and apices of elytra yellow. Ventral side of head and prothorax light yellowish brown, ventral side of mesothorax, metathorax and abdomen brown. Head. Nearly hypognathous. Labrum pitchy black, widely, but not deeply incised. Maxillary and labial palpi brown. Clypeus and frons light brown, feebly pilose, frontal ridge absent. Suprafrontal sulcus shallow, frontal calli almost indistinct. Orbital and supraorbital sulci developed but shallow. Orbit and space around eyes slightly darker than vertex and frons in some specimens. Vertex shiny, almost impunctate. Eyes round. Antennae reaching basal ¼ of elytra. Antennomeres I–V and XI light brown, antennomeres VI–X dark brown. Antennomere VII slightly protruded. Thorax. Pronotum wide, convex, impunctate. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra slightly, irregularly punctured. Humeral part of lateral elytral margin and basal elytral margin with two wide light brownish yellow spots; lateral elytral margin with a wide yellowish-brown stripe; pleura brown. Elytral apices yellow. Procoxal cavity widely open posteriorly; prosternal intercoxal process elevated ventrally. Anterior metaventral process completely covered by mesoventrite between mesocoxae, with a deeply excavated, upturned U-shaped “horseshoe-like” structure. Legs brown. Tibiae feebly but distinctly pilose. Metatibiae slightly curved laterally, with a long apical spine. Metatarsomere I not strongly elongated. Abdomen. Ventrites slightly pilose. Abdominal ventrite I with a bulbose anterior process reaching the space between metaxoxae, with a slightly elevated ridge. Abdominal ventrite V slightly paler than ventrites I–IV. Genitalia. Median lobe of aedeagus short, wide and slightly curved in lateral view; in ventral view slightly widened towards apex in basal 2/3, apical 1/3 of median lobe of aedeagus narrowing, triangular, apex narrowly rounded. Spermatheca slender; spermathecal pump short, receptacle elongate-oval, spermathecal duct shortly Ushaped, without coils, ending in ½ of the receptacle. Vaginal palpi short, joined basally, V-shaped, slightly separated apically. Tignum slender. Etymology. The species is named after its specific elytral color pattern—black with yellow lateral margin (marginata —with notable margin)., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on pages 57-58, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144

Cangshanaltica sprynari Damaška & Aston, 2019 Fig. 8J Cangshanaltica sprynari Damaška & Aston, 2019: 153 Type locality: China, Hong Kong, Lantau. Type series: Holotype: NMPC; paratypes: NMPC, ADPC, AFCD, USNM, SYSU. Remarks. The species is only known from the type series., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on page 60, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Damaska, A. F. & Aston, P. (2019) Leaf litter and moss-inhabiting flea beetles of Hong Kong (Coleoptera: Chrysomelidae: Alticini). Acta Entomologica Musei Nationalis Pragae, 59, 151 - 161. https: // doi. org / 10.2478 / aemnp- 2019 - 0013"]}

Cangshanaltica javana sp. nov. Fig.s 5, 8D Type locality: Indonesia, Java Barat, Gunung Tangkuban Perahu. Type material. HOLOTYPE (male, MHNG): (1) INDONESIE — East Java 30 km N. Bandung, volcan Tangkuban Prahu, J. Robert 10.VIII.85. ◉ Paratypes (1 male MHNG, 1 male NMPC) same labels as holotype; (2 female MHNG, 1 female NMPC): JAVA Volcan Tangkuban Prahu env 30 km N. Bandung J. Robert 7. VIII. 84. Differential diagnosis. The species can be easily distinguished from all known species by its genitalic morphology (median lobe of aedeagus pointed with sharp parallel margins on dorsal side, spermathecal duct long, U shaped). Additionally, the species can be distinguished by the key below. Description. Habitus. Body ovate, convex, 2.4 mm long, 1.7 mm wide, 1.1 mm high. Dorsal surfaces dark blackish-brown without metallic luster. Ventral side brown. Head. Nearly hypognathous. Labrum light brown, deeply incised; clypeus black. Labial and maxillary palpi light brown. Frons wide, brown, flat, frontal ridge absent. Suprafrontal sulcus well-developed, frontal calli feebly projecting, flat, feebly delimited posteriorly. Orbit narrow, orbital and supraorbital sulci deep. Vertex very finely punctured with shallow scattered punctures. Antennae reaching basal ¼ of elytra. Antennomeres I–IV and XI light yellowish-brown, antennomeres V–X brown. Antennomeres V–XI gradually widening, flat. Thorax. Pronotum wide, convex, punctured with small scattered punctures. Anterior pronotal margins feebly projecting anteriorly, but forming lobes. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra convex, about ¼ wider than maximal width of pronotum, irregularly punctured with distinct punctures twice as large as those on pronotum. Epipleura extremely widened in basal parts of elytra, nearly as wide as half of pronotal width. Procoxal cavity widely open posteriorly. Anterior metaventral process completely covering the mesoventrite between mesocoxae, as a deeply excavated “horseshoe-like” structure. Legs light brown, metafemora darkened. Metatibiae slightly curved laterally, with a long apical spine. Metatarsomere I elongated. Abdomen. Abdominal ventrite I as long as ventrites II and III combined. Anterior process of abdominal ventrite I reaching space between metacoxae, bulbose, forming a ventrally projecting elevated ridge. Genitalia. Median lobe of aedeagus long, slender, pointed apically, slightly curved in lateral view. Dorsal side of median lobe of aedeagus with sharp parallel margins narrowing in the middle, ventral side not narrowed. Spermathecal pump as long as receptacle, with digitiform apical part; receptacle twice as wide as pump, almost parallel-sided. Spermathecal duct long, U-shaped, reaching the apical 1/3 of the receptacle. Etymology. The species is named after the island of Java, where the type series was collected., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on pages 55-56, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144

Cangshanaltica sprynari Damaška & Aston, 2019 Fig. 8J Cangshanaltica sprynari Damaška & Aston, 2019: 153 Type locality: China, Hong Kong, Lantau. Type series: Holotype: NMPC; paratypes: NMPC, ADPC, AFCD, USNM, SYSU. Remarks. The species is only known from the type series.

Cangshanaltica fuanensis Ruan, Konstantinov and Damaška, 2022 Fig. 8B Cangshanaltica fuanensis Ruan, Konstantinov and Damaška, 2022: 29 Type locality: China, Fujian, Fuan. Type series: Holotype — IZAS; paratypes: IZAS, SZPT, ADPC. Remarks: The species was described originally by Ruan et al. (2020), however, the description did not comply with the ICZN, and the name has been considered as nomen nudum. Therefore, it has been described correctly by Ruan et al. (2022). The species is known only from the type series (30 specimens). It is very similar to the C. castanea species complex (from which it can be separated by using the key below) in having brown, ellipticalrounded body. However, Damaška et al. (2021) revealed that C. fuanensis is more closely related to the Taiwanese Cangshanaltica species complex, rather than to the C. castanea s. lat. from Zhejiang to which it is most similar., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on page 54, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Ruan, Y., Konstantinov, A. S. & Damaska, A. F. (2022) Description of the moss-eating flea beetle Cangshanaltica fuanensis sp. nov. (Coleoptera, Chrysomelidae, Galerucinae, Alticini) from China. Journal of Insect Biodiversity, 034, 28 - 32. https: // doi. org / 10.12976 / JIB / 2022.34.2.1","Ruan, Y., Konstantinov, A. S. & Damaska, A. F. (2020) The biology and immature stages of the moss-eating flea beetle Cangshanaltica fuanensis sp. nov. (Coleoptera, Chrysomelidae, Galerucinae, Alticini), with description of a fan-driven high-power Berlese funnel. Insects, 11, 1 - 27. https: // doi. org / 10.3390 / insects 11090571","Damaska, A. F., Konstantinov, A., Lee, C. - F., Ruan, Y., Mohagan, D. J. & Fikacek, M. (2021) Molecular phylogeny of mossinhabiting flea beetles from the Chabria group (Coleoptera: Chrysomelidae: Alticini) reveals multiple colonizations and radiations in Taiwan. Systematic Entomology, 46 (4), 915 - 925. https: // doi. org / 10.1111 / syen. 12502"]}

Cangshanaltica castanea (Gruev, 1985) comb. nov. Figs 3, 8A Taizonia castanea Gruev, 1985: 126 Ivalia gruevi Nadein, 2013: 396 (nom. nov. for castanea Gruev, not Samuelson) Type locality: China: Da-laen-saen (type locality identity is discussed below). Type material examined. HOLOTYPE (male, BMNH): (1) Holotype, (2) G. C. Champion Coll. B. M. 1927- 409, (3) Da-laen-saen, China. J. J. Walker, (4) HOLOTYPE; (5) Taizonia castanea Gruev sp. n.; ◉ PARATYPES (1 male; 4 female, BMNH): (1) Da-laen-saen, China. J. J. Walker, (2) PARATYPE. Additional material examined. 6 spec. (4 male, 2 female, BMNH): (1) G. C. Champion Coll. B. M. 1927-409; (2) Da-laen-saen, China. J. J. Walker; (3) Taizonia castanea Gruev det. A. S. Konstantinov, 2003; 9 spec. (9 male, BMNH): (1) Nyew-tew I., China, J. J. W [alker]; (2) ◉ 4 spec. (3 male, 1 female, NMPC, ADPC): (1) CHINA — Zhejiang prov., Lishui pref., Lu’aocun (炉岙村); Monkey Valley (Ñšẽ); 27°54′55.2″N 119°11′55.4″E; 1100 m; 11. v. 2019; A. F. Damaška lgt.; (2) VOUCHER SPECIMEN DNA collection No. AFD-235; 236; 282 283; 284 Albert F. Damaška coll.; (3) Cangshanaltica cf. castanea (spec. complex), A. F. Damaška det. 2022 ◉ 2 spec. (2 male, SZPT): (1) Hangzhou Wuyun Mt. (杭州五云山), on moss, under tree root, 2021-III-25, leg. Wei Wang (Ɨ ‼). Redescription. Habitus. Body oval-rounded, convex, holotype 1.6 mm long, 1.2 mm wide, 0.9 mm high. Dorsal surfaces chestnut brown without metallic luster. Ventral surfaces light brown. Head. Nearly hypognathous, triangular, chestnut brown. Frontal calli flat, feebly developed, distinctly delimited anteriorly, feebly delimited posteriorly. Frontal ridge wide, flat, not projecting; interantennal space as wide as diameter of eye. Supraantennal, suprafrontal, orbital and supraorbital sulci shallow but developed. Orbit narrow. Frontal ridge, orbit and frons impunctate. Antennae with 11 antennomeres, short, reaching anterior ¼ of elytra. Antennomeres I and II large, wide, light yellowish brown; antennomere III slightly elongated, antennomeres IV–VI similar in length, slender, antennomere VI wider than V. Antennomere VII short, with a distinct distal protrusion; antennomeres VIII–XI gradually widening, forming a long and slender antennal club. Thorax. Pronotum brown, wide, convex, feebly punctate by scattered shallow punctures.Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Posterior edges of pronotum sharp. Elytra convex, wide, feebly irregularly punctured. Maximal width of elytra 1/3 wider than maximal width of pronotum. Procoxal cavity widely open posteriorly.Anterior metaventral process completely covering mesoventrite between mesocoxae, developing a pentagonal “horseshoe-like” structure. Legs light brown. Metatibiae nearly straight, not visibly curved laterally. Metatarsomere I elongated. Abdomen and genitalia. Abdominal ventrite I as long as ventrites II and III combined. Median lobe of aedeagus variable in width, and in sharpness of the apex (Fig. 3G,H). Apex pointed, arrow-like. Median lobe widened from apex to basal part. Basal orifice pyriform, reaching ¼ of the median lobe of aedeagus length. Spermatheca slender, duct without coils. Vaginal palpi widely separated apically. Generic placement. Cangshanaltica castanea had been described in Taizonia by Gruev (1985), a genus subsequently synonymized with Ivalia by Duckett et al. (2006). Due to homonymy with Ivalia castanea (Samuelson, 1966), a new name Ivalia gruevi was proposed for the species by Nadein (2013). Our placement of Gruev’s species in Cangshanaltica makes Nadein’s replacement name redundant (ICZN Art 59.4). We move the species to Cangshanaltica based on the following characters: (1) antennomere VII bearing a slight distal protrusion; (2) metaventrite bearing an anterior, horseshoe-like process reaching mesocoxae and covering the mesoventrite; (3) anterolateral pronotal setiferous pore placed in the middle of the pronotal margin; and (4) round, ovate, and convex body shape. Cangshanaltica castanea is similar to other Chinese Cangshanaltica species distributed in lowlands— C. sprynari, and C. fuanensis. The species can be distinguished using the key below. Type locality identity and distribution. The type locality “ China: Da-laen-saen” named by Walker has been discussed by authors who have revised J. Walker’s Chinese material (Schawaller & Aston 2017; Smetana 2009). According to their findings, the most likely type locality of C. castanea is China, Zhejiang prov., Ningbo pref., Dalei Shan (kDZ山), cca 29.5928 N, 121.2930 E. The second Walker locality in the BMNH material is “Nyew-tew I.” This locality refers most likely to the Nantian Dao Island (Éffl岛) in Zhejiang province, named “Nyew-tew Island” in 19th century Western maps, likely after the island’s peak Niutou Shan (牛Ë山). The second type locality should therefore be: China, Zhejiang prov., Ningbo pref., Nantian Dao (Éffl岛); cca 29.1361 N, 121.9388 E. Additional specimens assigned to the species complex of C. castanea were collected recently in Lu’aocun (炉岙村) in the Lishui prefecture, and in Hangzhou, both in Zhejiang province, China. Species identity and variability. Cangshanaltica castanea was described based on a type series collected by J. J. Walker. Besides the type series, we examined additional Walker material from the type locality (Da Laen Saen) and its surroundings (Nyew-tew I.). We also collected fresh DNA-grade specimens in southern Zhejiang —they strongly resemble several specimens from the C. castanea type series. All the specimens have similar morphology of external characters and female genitalia, however, the morphology of median lobe of aedeagus is quite variable among all specimens, even within the type material (fig. 3 G–H). Specimens also differ in elytral punctuation (in the holotype and most of Walker’s specimens, elytra have very feeble punctures, in several of Walker’s specimens and all the fresh specimens from Lishui, punctures are shallow but distinctly visible, forming a pitted elytral relief, similar to the surface of a golf ball), and size (males are up to 1.9 mm long). A similar species in external morphology is C. fuanensis Ruan, Konstantinov & Damaška, 2022, which differs in having a more slender median lobe of aedeagus, but is strongly divergent from the Lishui specimens based on DNA phylogenetic analyses (Damaška et al. 2021). This strong difference in both mitochondrial and nuclear DNA markers among the two morphologically similar species suggests that the C. castanea species complex can consist of more species which are similar in morphology but phylogenetically isolated from each other. Due to the lack of any DNA-grade material from the two historical collecting sites in Zhejiang, we decided not to treat any of the specimens with different genitalia morphology as distinct species. However, the variability of median lobe of aedeagus in C. castanea in our understanding is apparently larger than in other Cangshanaltica species. The species definition of C. castanea may therefore represent a species complex. A careful analysis of DNA and morphology based on fresh material from more localities in Zhejiang and around (including Walker’s sites) is the only way to yield enough information to resolve the C. castanea species complex., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on pages 50-52, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Gruev, B. (1985) Two new taxa of Alticinae from China (Coleoptera: Chrysomelidae). Entomological Review of Japan, 40 (2), 125 - 127.","Nadein, K. S. (2013) Ivalia Jacoby-a flea beetle genus new to Australia (Coleoptera: Chrysomelidae: Galerucinae). Zootaxa, 3669 (3), 384 - 400. https: // doi. org / 10.11646 / zootaxa. 3669.3.11","Konstantinov, A., Chamorro, M. L., Prathapan, K. D., Ge, S. - Q. & Yang, X. - K. (2013) Moss-inhabiting flea beetles (Coleoptera: Chrysomelidae: Galerucinae: Alticini) with description of a new genus from Cangshan, China. Journal of Natural History, 47 (37 - 38), 2459 - 2477. https: // doi. org / 10.1080 / 00222933.2012.763068","Duckett, C. N., Prathapan, K. D. & Konstantinov, A. S. (2006) Notes on identity, new synonymy and larva of Ivalia Jacoby (Coleoptera: Chrysomelidae) with description of a new species. Zootaxa, 68, 49 - 68. https: // doi. org / 10.11646 / zootaxa. 1363.1.4","Schawaller, W. & Aston, P. (2017) Two new species of the genus Laena Dejean (Coleoptera: Tenebrionidae: Lagriinae) from Hong Kong. Zootaxa, 4344 (1), 169 - 173. https: // doi. org / 10.11646 / zootaxa. 4344.1.10","Smetana, A. (2009) Contributions to the knowledge of the \" Staphylinus - complex \" (Coleoptera: Staphylinidae: Staphylinini) of China. Part 19. The genus Ocypus Leach, 1819, subgenus Pseudocypus Mulsant & Rey, 1876. Section 3. Acta Entomologica Musei Nationalis Pragae, 49, 683 - 694.","Ruan, Y., Konstantinov, A. S. & Damaska, A. F. (2022) Description of the moss-eating flea beetle Cangshanaltica fuanensis sp. nov. (Coleoptera, Chrysomelidae, Galerucinae, Alticini) from China. Journal of Insect Biodiversity, 034, 28 - 32. https: // doi. org / 10.12976 / JIB / 2022.34.2.1","Damaska, A. F., Konstantinov, A., Lee, C. - F., Ruan, Y., Mohagan, D. J. & Fikacek, M. (2021) Molecular phylogeny of mossinhabiting flea beetles from the Chabria group (Coleoptera: Chrysomelidae: Alticini) reveals multiple colonizations and radiations in Taiwan. Systematic Entomology, 46 (4), 915 - 925. https: // doi. org / 10.1111 / syen. 12502"]}

Key to described species of Cangshanaltica Konstantinov, Chamorro, Prathapan, Ge & Yang 1 Pronotum and elytra differently colored or elytra with a color pattern (Figs 4,6,7).................................. 2 - Pronotum and elytra unicolorous......................................................................... 5 2 Elytra black or dark blackish-brown with yellowish spots in basal and humeral parts and a yellowish brown stripe along epipleura; elytral apices yellowish brown (Median lobe of aedeagus widened towards apex; Fig. 6)... C. marginata sp. nov. - Elytra with a dark spot or spots.......................................................................... 3 3 Each elytron brownish ochraceous with a dark central spot (Antennomeres VII—XI distinctly elongated; Fig. 4)........................................................................................... C. guanxiensis sp. nov. - Elytra yellow with multiple dark spots.................................................................... 4 4 Elytra with 6 dark spots.................................................. C. ogatai Takemoto & Suenaga, 2021 - Elytra with 4 dark spots (Fig. 7)......................... A species complex related to C. ogatai distributed in Taiwan. 5 Dorsum without, or with a feeble metallic luster............................................................ 6 - Dorsum with a strong blue or green metallic luster......................................................... 11 6 Length greater than 1.5 mm in males, 1.7 mm in females..................................................... 7 - Body minute, length less than or equal to 1.5 mm in males, 1.7 mm in females.................................... 9 7 Dorsum brown....................................................................................... 8 - Dorsum black with feeble greenish metallic luster (median lobe of aedeagus gradually narrowing towards apex; Fig. 2)...................................................... C. nigra Konstantinov, Chamorro, Prathapan, Ge & Yang, 2013 8 Dorsum chestnut brown; antennae flattened,; median lobe of aedeagus elongate and slender with sharp parallel margins on dorsal side, pointed; antennomere XI distinctly paler than X (Fig. 5)................... Cangshanaltica javana sp. nov. - Dorsum dark brown; median lobe of aedeagus shorter, parallel-sided without distinct sharp margins, apex blunt; antennomere XI only slightly paler than X, antennae almost unicolorous chestnut brown... C. siamensis Damaška & Konstantinov, 2016. 9 Body circular; median lobe of aedeagus with blunt apex; spermathecal receptacle short.................................................................................................... C. sprynari Damaška & Aston, 2019 - Body oval; median lobe of aedeagus with apex pointed; spermathecal receptacle oval-elongate...................... 10 10 Median lobe of aedeagus much slenderer in apical third, then gradually narrowing towards apex, basal part wide and bulbose........................................................... C. fuanensis Ruan, Konstantinov & Damaška, 2022 - Median lobe of aedeagus slightly narrowing towards apex, almost parallel-sided, apex arrow-like, basal part not strongly widened (Fig 3)................................................................. C. castanea (Gruev, 1985) 11 Median lobe of aedeagus narrower, narrowing towards apex; spermathecal duct with coils; metallic luster violet-blue; elytra almost impunctate; tibial spur length same or less than length of metatarsomere II.............................................................................................. C. luzonica Damaška, Mohagan & Fikáček, 2020 - Median lobe of aedeagus wider, parallel-sided; spermathecal duct without coils; elytra irregularly but distinctly punctate; metallic luster greenish; tibial spur longer than metatarsomere II... C. mindanaoensis Damaška, Mohagan & Fikáček, 2020, Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on page 62, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Takemoto, T. & Suenaga, H. (2021) Four New Species of the Genus Ivalia Jacoby and a New Species of Cangshanaltica Konatantinov, Chamorro, Prathapan, Ge et Yang (Coleoptera, Chrysomelidae, Galerucinae) from the Ryukyus, Southwestern Japan. Elytra, 11, 129 - 153.","Konstantinov, A., Chamorro, M. L., Prathapan, K. D., Ge, S. - Q. & Yang, X. - K. (2013) Moss-inhabiting flea beetles (Coleoptera: Chrysomelidae: Galerucinae: Alticini) with description of a new genus from Cangshan, China. Journal of Natural History, 47 (37 - 38), 2459 - 2477. https: // doi. org / 10.1080 / 00222933.2012.763068","Damaska, A. F. & Aston, P. (2019) Leaf litter and moss-inhabiting flea beetles of Hong Kong (Coleoptera: Chrysomelidae: Alticini). Acta Entomologica Musei Nationalis Pragae, 59, 151 - 161. https: // doi. org / 10.2478 / aemnp- 2019 - 0013","Ruan, Y., Konstantinov, A. S. & Damaska, A. F. (2022) Description of the moss-eating flea beetle Cangshanaltica fuanensis sp. nov. (Coleoptera, Chrysomelidae, Galerucinae, Alticini) from China. Journal of Insect Biodiversity, 034, 28 - 32. https: // doi. org / 10.12976 / JIB / 2022.34.2.1","Gruev, B. (1985) Two new taxa of Alticinae from China (Coleoptera: Chrysomelidae). Entomological Review of Japan, 40 (2), 125 - 127.","Damaska, A. F., Mohagan, D. J. & Fikacek, M. (2020) Moss-inhabiting flea beetles in the Philippines (Coleoptera, Chrysomelidae, Alticinae). ZooKeys, 960, 125 - 142. https: // doi. org / 10.3897 / zookeys. 960.54011"]}

Cangshanaltica guanxiensis sp. nov. Figs 4, 8C Type locality: China, Guanxi, Panxie Gou. Type material. HOLOTYPE (male, NMPC): (1) CHINA: Guangxi, Pangxie Gou, 1.3 km S of Longsheng Hot Spring, 25°53.0′N 110°12.6′E, 385 m 13.iv.2013, Fikáček, Hájek & Růžička lgt. (2) sifting of leaf litter and leaf accumulations along stony stream in evergreen lowland subtropical forest [sifting sample 13]; (3) VOUCHER SPECIMEN DNA collection No. AFD-125 Albert F. Damaška coll. Differential diagnosis. The species can be distinguished from all other known Cangshanaltica species by its coloration—elytra ochraceous with one darkened spot in the center of each elytron. DNA sequences. 18S: MZ312403.1; 16S: MZ312387.1; Cox1: MZ303694.1, MZ220536.1 (barcode). All sequences were published by Damaška et al., 2021. Description. Habitus. Body round, convex, 2 mm long, 1.7 mm wide, 0.9 mm high. Dorsal and ventral surfaces ochrebrown, each elytron with a darkened spot in the middle. Head. Nearly hypognathous, triangular. Labrum and clypeus blackish brown, vertex brown.Frontal ridge absent; frons shiny, impunctate, with bunches of setae below antennae. Frontal calli nearly absent. Suprafrontal sulcus shallow, orbital and supraorbital sulci developed. Eyes round. Antennae reaching basal 1/3 of elytra. Antennomeres I–VI and XI light brown, antennomeres V–X darkened. Antennomeres I and II bulbose, wide; antennomeres VII–XI elongate, as long or longer as antennomeres III and IV combined.Antennomere VII bearing a slight distal protrusion. Thorax. Pronotum brownish ochraceous, wide, convex, impunctate. Anterior margins of pronotum lobate, projecting, forming distinct “flap-like” structures towards head; posterior pronotal edges dull. Anterolateral pronotal setiferous pore placed in the middle of lateral pronotal margin. Elytra convex, brownish ochraceous with a darkened spot in the central part of each elytron, irregularly punctured. Epipleura distinctly widened in basal part. Anterior metaventral process completely covering the mesoventrite between mesocoxae, as a sharply pentagonal, deeply excavated “horseshoe-like” structure. Legs brownish ochraceous. Metatibiae slightly curved, apical metatibial spine short. Metatarsomere I slightly elongated. Abdomen. Abdominal ventrite I longer than ventrite II, but not as long as ventrites II and III combined.Anterior process of abdominal ventrite I reaching space between metacoxae, bulbose, forming a sharp, ventrally projecting elevated ridge. Genitalia. Median lobe of aedeagus long but wide, parallel-sided in dorsal view, slightly curved in lateral view. Apex dull, slightly widening posteriorly. Female unknown. Remarks. The only known specimen is apparently not fully sclerotized—the coloration of a mature adult is probably somewhat darker. Despite that, we are describing the species, as all morphological characters are welldeveloped, and available DNA markers clearly indicate its isolated phylogenetic position among other studied Cangshanaltica species (Damaška et al. 2021). Etymology. The species name is derived from Guanxi, the name of the Chinese province where the holotype was collected., Published as part of Damaška, Albert František, Ruan, Yongying & Fikáček, Martin, 2022, The genus Cangshanaltica (Coleoptera: Chrysomelidae: Alticinae): overview, new species, and notes on species complexes, pp. 49-64 in Zootaxa 5219 (1) on pages 54-55, DOI: 10.11646/zootaxa.5219.1.2, http://zenodo.org/record/7408144, {"references":["Damaska, A. F., Konstantinov, A., Lee, C. - F., Ruan, Y., Mohagan, D. J. & Fikacek, M. (2021) Molecular phylogeny of mossinhabiting flea beetles from the Chabria group (Coleoptera: Chrysomelidae: Alticini) reveals multiple colonizations and radiations in Taiwan. Systematic Entomology, 46 (4), 915 - 925. https: // doi. org / 10.1111 / syen. 12502"]}

CANGSHANALTICA KONSTANTINOV ET AL., 2013 (FIG. 4) Type species: Cangshanaltica nigra Konstantinov et al., 2013. Synonymy: No generic synonyms. Phylogenetic position: Cangshanaltica has recently been revealed as a member of the Chabria group, together with the moss-inhabiting genus Ivalia and the leaf-surfaceliving genera Chabria Jacoby, 1887, Chabriosoma Chen, 1934 and Sutrea Baly, 1876 (DamaŠka et al., 2020). In our present study, the sister-group of Cangshanaltica is Ivalia (pp = 0.95). However, a larger phylogenetic analysis of the Chabria group revealed Cangshanaltica as a sister-group of a clade containing mostly winged genera, namely Chabria, Chabriosoma, Sutrea and Parathrylea Duvivier, 1892 (DamaŠka et al., 2020). Revisions: The genus was recently described (Konstantinov et al., 2013) as monotypic, with additional species described since (DamaŠka & Konstantinov, 2016; Konstantinov et al., 2016; DamaŠka & Aston, 2019; Ruan et al., 2020; Takemoto & Suenaga, 2021). Diversity and distribution: Seven species have been described; descriptions of four additional species are in preparation (DamaŠka et al., in prep.). The genus is distributed in the Himalayan foothills (China and Thailand), the Philippines, the Chinese coast from Hong Kong to Zhejiang, in Taiwan (DamaŠka et al., 2021) and in the Ryukyu Archipelago (Takemoto & Suenaga, 2021). Morphological characteristics: Body generally ovate to round in dorsal view, strongly convex in lateral view, head nearly hypognathous. Antennal calli indistinct. Antennae with 11 antennomeres; apical antennomeres tend to be widened. Antennomere 7 bears a distal protrusion. A remarkably similar protrusion was recently discovered in a flightless galerucine Prathapanius fortis Viswajyothi & Clark, 2020. Pronotum short, convex, with an anterolateral setiferous pore placed nearly in the midlength of the lateral pronotal margin. Procoxal cavity open posteriorly. Mesoventrite entirely covered by the anterior process of the metaventrite in the area between the mesocoxae. The surface of the anterior metaventral process is strongly modified with distinctly projecting lateral margin, forming a specific ‘horseshoe-like’ shape. Elytra convex, usually covered with scattered feeble punctures. Legs robust, metatibiae slightly or strongly curved. First metatarsomere elongated. First abdominal ventrite with an elevated ridge, surrounded by deep punctures, exceeding as an anterior abdominal process between metacoxae. Aedeagus usually simple, long, with somewhat arrow-like apex. Spermathecal duct without coils. Vaginal palpi fused basally. Ecology: Unlike other moss-inhabiting flea beetle genera, the ecology of Cangshanaltica is well documented. The montane species (C. mindanaoensis DamaŠka et al., 2020, C. nigra and C. siamensis DamaŠka & Konstantinov, 2003) live in moss cushions in montane cloud forests or close to the tree line. Cangshanaltica sprynari DamaŠka & Aston, 2019 and C. fuanensis Ruan et al., 2020, on the contrary, were found at low elevations. Individuals of C. sprynari were documented to be active only for few days during the year, usually after rains, walking over substrate covered by thin layers of mosses (instead of inside of moss cushions). Cangshanaltica nigra is documented to feed on tissues of Hypnum Hedw. moss, as documented by the dissection of its gut content; C. sprynari feeds on Fissidens Hedw., which was also demonstrated by dissection (Konstantinov et al., 2013; DamaŠka & Aston, 2019). Ruan et al. (2020) described the larva and ecology of C. fuanensis, including the length of the larval development and egg-laying ecology. Remarks: Externally, Cangshanaltica is s i m i l a r t o I v a l i a, s h a r i n g m o s t o f i t s g e n e r a l morphological features, i.e. round and convex body, curved metatibiae and the horseshoe-like anterior metaventral process. The main diagnostic characters between Cangshanaltica and Ivalia are: the anterolateral pronotal setiferous pore situated nearly in the midlength of the pronotal margin (in Ivalia, it is placed anteriorly, usually with an anterior lobe placed before the pore); seventh antennomere is modified, bearing a distal protrusion (in Ivalia, seventh antennomere does not bear such a protrusion). Members of Ivalia are also usually more ovate (Cangshanaltica is usually more rounded) and less convex in body shape., Published as part of Damaška, Albert František, Konstantinov, Alexander & Fikáček, Martin, 2022, Multiple origins of moss-inhabiting flea beetles (Coleoptera: Chrysomelidae): molecular phylogeny, overview of genera and a new genus from Africa, pp. 647-676 in Zoological Journal of the Linnean Society 196 on pages 659-660, DOI: 10.1093/zoolinnean/zlab112, http://zenodo.org/record/7196247, {"references":["Konstantinov A, Chamorro ML, Prathapan KD, Ge SQ, Yang XK. 2013. Moss-inhabiting flea beetles (Coleoptera: Chrysomelidae: Galerucinae: Alticini) with description of a new genus from Cangshan, China. Journal of Natural History 47: 2459 - 2477.","Damaska AF, Mohagan DJ, Fikacek M. 2020. Mossinhabiting flea beetles in the Philippines (Coleoptera, Chrysomelidae, Alticinae). ZooKeys 960: 125 - 142.","Damaska A, Konstantinov A. 2016. A new species of Cangshanaltica Konstantinov et al., a moss-inhabiting flea beetle from Thailand (Coleoptera: Chrysomelidae: Galerucinae: Alticini). Zootaxa 4107. 93 - 97.","Damaska AF, Aston P. 2019. Leaf litter and moss-inhabiting flea beetles of Hong Kong (Coleoptera: Chrysomelidae: Alticini). Acta Entomologica Musei Nationalis Pragae 59: 151 - 161.","Ruan Y, Konstantinov AS, Damaska AF. 2020. The biology and immature stages of the moss-eating flea beetle Cangshanaltica fuanensis sp. nov. (Coleoptera, Chrysomelidae, Galerucinae, Alticini), with description of a fan-driven high-power Berlese funnel. Insects 11: 1 - 27.","Takemoto T, Suenaga H. 2021. Four new species of the genus Ivalia Jacoby and a new species of Cangshanaltica Konatantinov, Chamorro, Prahtapan, Ge et Yang (Coleoptera, Chrysomelidae, Galerucinae) from the Ryukyus, southwestern Japan. Elytra 11: 129 - 153.","Damaska AF, Konstantinov A, Lee CF, Ruan Y, Mohagan DJ, Fikacek M. 2021. Molecular phylogeny of mossinhabiting flea beetles from the Chabria group (Coleoptera: Chrysomelidae: Alticini) reveals multiple colonizations and radiations in Taiwan. Systematic Entomology 46: 915 - 925."]}

Cangshanaltica sprynari sp. nov. (Figs 2 A–I, 7A–E) Ivalia sp. n.: ASTON (2009): 11 (note). Type locality. China, Hong Kong, Lantau Island, Wang Tong. Type material. HOLOTYPE: ♁, ‘ China: Hong Kong – Lantau isl., Wang Tong 8. viii. 2010 Paul Aston lgt. (NMPC). PARATYPES: 1 ♁♁ 2 ♀, same locality label as the holotype and additional label ‘Genus Ivalia det. Döberl 2010’ (1 ♁ USNM, 1 ♀ SYSU, 1 ♀ NMPC); 1 ♀, ‘ China: Hong Kong – Lantau isl., Sunset Peak floating in river below waterfall, 27. ix. 2009 Paul Aston lgt.’ (PCPA); 1♁, ‘ China: Hong Kong – Lantau isl., Wang Tong, on moss; 9.iii.2009.Paul Aston lgt.’ (AFCD); 1♁, ‘ China: Hong Kong – Lantau isl., Wang Tong, on moss; 15. xi. 2008 Paul Aston lgt.’ (ADPC). Differential diagnosis. This new species belongs to the genus Cangshanaltica based on following characters: (1) pronotal anterolateral setiferous pore placed in the middle of the pronotal side margin; and (2) antennomere VII with a distal protrusion. Cangshanaltica sprynari sp. nov. differs from all other species of Cangshanaltica by following characters: (1) round anterior pronotal margins (other species have anterior pronotal margins somewhat sharp), (2) short, broad and strongly diverging vaginal palpi (other species have long, slender and parallel vaginal palpi), and (3) nearly undivided frontal calli (C. nigra Konstantinov et al., 2013 has poorly developed frontal calli, C. siamensis Damaška & Konstantinov, 2016 has distinctly divided triangular frontal calli). The species differs from C. nigra in brown coloration (C. nigra is black), in the shape of the aedeagus (C. nigra has the apex of the aedeagus gradually narrowing, C. sprynari has a narrow apex of the aedeagus, distinctly divided by a visible, abruptly narrowing step), in the shape of vaginal palpi (C. nigra has parallel and long vaginal palpi, vaginal palpi of C. sprynari are short, broad and strongly diverging), development of frontal calli (in C. nigra, frontal calli are nearly invisible) and metatarsomere III long (in C. nigra, the metatarsomere III is very short). The species also differs from C. siamensis in the shape of the aedeagus (C. siamensis has a distinctly pointed apex) and in having a bulbose spermathecal receptacle (C. siamensis has a slender receptacle). Description. Habitus (Figs 1 A–C). Body 1.7 mm long, 1.3 mm wide in maximum, oval-rounded in dorsal view, convex in lateral view, 1 mm high in maximum. Color of both ventral and dorsal body surface dark brown, legs light brown, eyes black. Head nearly hypognathous. Frontal calli present, flat, feebly visible, indistinctly divided, distinctly surrounded ventrally and dorsally by longitudinal impressions. Interantennal space wide. Antennae with 11 antennomeres, antennomeres I–V and XI light brown, antennomeres VI–X darker. Antennomere VII bearing distal protrusion anteriorly.Antennomere I triangular, antennomere II oval, antennomeres III–XI regularly elongate. Maxillary palpi light brown. Frontal ridge broad, flat. Clypeus bearing two groups of three setae each, situated symmetrically on each side. Labrum bilobed, with incision reaching ¼ of labrum length, bearing six large setae in posterior parts and some scattered smaller setae on anterior margin. Thorax. Pronotum convex, twice as wide as long, sparsely covered with shallow punctures, anterior pronotal margins strongly rounded, posterior pronotal edges sharp. Elytra strongly convex with irregular punctation, punctures larger and deeper than those on pronotum. Metathoracic wings and humeral calli absent. Metatibiae curved in dorsal view, pilose, with exterior row of teeth reaching from proximal ¼ of length of metatibia to apex. Metatibial teeth gradually elongate apically. Metatarsus attached on metatibia in deep apical impression surrounded externally by long external teeth and internally by short apical row of long teeth. Metatarsomere I 2–3× longer than II, metatibial apical tooth as long as metatarsomere II. Abdomen with five distinct ventrites. Ventrite I as long as II and III combined, with II slightly longer than III. Longitudinal ridge on ventrite I long, reaching 2/3 of its length. Genitalia. Aedeagus (Figs 2 H–I) long, curved and broad in lateral view, parallel-sided in ventral view, apex abruptly narrowed in step-like fashion in apical eighth of aedeagus. Spermathecal pump long, receptacle bulbose, duct simple, short, directed parallel with the receptacle, reaching ½ of receptacle length (Fig. 2D). Vaginal palpi short, broad, strongly diverging, bearing group of long setae (Fig. 2E). Tignum long, slender, simple (Fig. 2F). Etymology. The species is named in honor of Pavel Špryňar, a Czech botanist and entomologist, who has contributed to Czech entomology by hosting an entomological club for children in Prague. Biology. The specimens were collected at night, walking on the surface of a thin layer of moss covering stones in secondary forest growth and orchard, near Wang Tong village, Lantau, Hong Kong (Figs 7 A–B). Cangshanaltica sprynari sp. nov. appears to be active only at night, they usually begin to occur two hours after dusk (P. Aston frequently visits the locality during the day and early evening, finding no specimens) and only in periods of humidity or rainfall, never found when the moss is totally dry. One specimen was collected, with many other terrestrial beetles, in a stream after being washed away in heavy rainfall. Host plant. One of us (PA) found the beetles feeding on moss Fissidens sp., Fissidentaceae (Figs 7 C–D). Also the gut contained the residua of moss (Fig. 7E)., Published as part of Damaška, Albert F. & Aston, Paul, 2019, Leaf litter and moss-inhabiting flea beetles of Hong Kong (Coleoptera: Chrysomelidae: Alticini), pp. 151-161 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 59 (1) on pages 153-154, DOI: 10.2478/aemnp-2019-0013, http://zenodo.org/record/4505179, {"references":["ASTON P. 2009: Chrysomelidae of Hong Kong Part 2 Subfamily Alticinae. Hong Kong Entomological Bulletin 1: 1 - 13.","KONSTANTINOV A., CHAMORRO M. L., PRATHAPAN K. D., GE S. & YANG X. 2013: Moss-inhabiting flea beetles (Coleoptera: Chrysomelidae: Galerucinae: Alticini) with description of a new genus from Cangshan, China. Journal of Natural History 47: 37 - 41.","DAMASKA A. & KONSTANTINOV A. 2016: A new species of Cangshanaltica Konstantinov et al., a moss-inhabiting flea beetle from Thailand (Coleoptera: Chrysomelidae: Galerucinae: Alticini). Zootaxa 4107: 93 - 97."]}

Cangshanaltica siamensis new species (Figs 1, 2) Description. Habitus. Body length 1.94–2.21 mm, width 1.45–1.62 mm. Body oval in dorsal view, convex in lateral view. Upper side dark brown, without metallic lustre. Abdomen and legs lighter, chestnut brown (Fig. 1). Head. Head nearly hypognathous. Vertex shiny, lacking wrinkles, with small, sparse punctures and few thin, scattered setae. Antennal calli poorly developed, triangular, their borders barely traceable. Supracallinal, midfrontal and suprafrontal sulci shallow, barely visible. Supraorbital sulcus deep and straight. Interantennal space wide. Labrum black. Maxillary palpomeres chestnut brown with yellow apices. Antennae with 11 antennomeres, antennomeres 1–4 yellow, continually darkening, antennomeres 5–10 brown, antennomere 11 lighter, yellowish-brown. Antennomere 7 with distal protrusion anteriorly. Antennomere 2 oval, ca. 1.4 × longer than wide. Antennomeres 3 and 4 very thin, much thinner than antennomeres 1 and 2 each. Thorax. Pronotum convex, in the middle 1.79 times as wide as long, with small and shallow, sparse punctures. Lateral margin with setiferous pore situated nearly in middle. Elytra convex, humeral calli absent. Punctation of elytra fine, but coarser than that of pronotum, punctation confused. Metathoracic wings not developed. Anterior coxal cavity open posteriorly. Third tarsomere of all tarsi long, as long or slightly longer than first tarsomere. Inner side of metatibia with a row of teeth situated between distal 1 / 4 and tibial apex. Metafemur pilose. Metatibial apical spine long, reaching ca. 4 / 5 of first metatarsomere. Abdomen with 5 ventrites. Longitudinal ridge on first abdominal ventrite developed only basally and short, almost invisible. Metacoxal cavities very deep, reaching nearly mid length of the first ventrite. Genitalia. Spermathecal pump long, receptacle narrow, slender. Vaginal palpi slender. Aedeagus long, curved in lateral view, parallel-sided from base nearly to apex, then converging apically. Orificium basale very long, arrow-shaped (Fig. 2). Sexual dimorphism. Males generally smaller than females—male 1.94 mm long and female 2.21 mm long. Females with smoother frons, antennal calli less distinct than in males. Differential diagnosis. We place this species in Cangshanaltica based on the following characters: anterior coxal cavities open posteriorly; setiferous pore placed nearly in middle of lateral pronotal margin; antennomere 7 with a distal anterior protrusion (Fig. 1); male genitalia more or less flat in lateral view with indistinct broad depression ventrally, sides parallel at basal half, gradually narrowing without well-defined denticle at apex (Fig. 2). The new species may be distinguished from C. nigra Konstantinov et. al., 2013 based on the following characters: third tarsomeres or all tarsi long (Fig. 2), as long or slightly longer than first (in C. nigra, third tarsomere is significantly shorter than first, especially on metatarsi), longitudinal ridge on first abdominal ventrite very short, present only basally and hence nearly indistinct (in C. nigra, the ridge is long) (Fig. 2), metatibial apical spine long, reaching ca. 4 / 5 of first metatarsomere (in C. nigra, metatibial apical spine reaches at most 1 / 2 of first metatarsomere), upper side of body dark brown (in C. nigra, the upper side is black with metallic lustre). Cangshanaltica siamensis can be also distinguished by the some features of male genitalia: parallel sides of aedeagus extend further apically, apex only very slightly bent ventrally (in C. nigra the bend is more pronounced). Etymology. The name siamensisis derived from the old name for Thailand—Siam. Type locality and ecology. Specimens of C. siamensis were collected by beating moss from tree trunks in a cloud forest in the summit area of Doi Inthanon, ca. at 2 500 m a.s.l. (Fig. 3). Specimens were collected in semi-dry moss during the dry season. Type material. Holotype male. Labels: (1) THAILAND—Chiang Mai prov., Mae Chaem distr., Doi Inthanon, 18 ° 35 ' 10.36 "N, 98 ° 28 ' 57.18 "E, cloud forest in dry season, dry moss on trees, 6. iv. 2015, Albert Damaška lgt.; (2) HOLOTYPE Cangshanaltica siamensis sp nov., Albert Damaška and Alexander Konstantinov des. 2016. Paratypes have same labels as holotype. Holotype NMPC, paratypes: 1 female NMPC, 3 (1 male, 2 female) USNM, 2 (1 male, 1 female) ADPC.

Cangshanaltica siamensis new species (Figs 1, 2) Description. Habitus. Body length 1.94���2.21 mm, width 1.45���1.62 mm. Body oval in dorsal view, convex in lateral view. Upper side dark brown, without metallic lustre. Abdomen and legs lighter, chestnut brown (Fig. 1). Head. Head nearly hypognathous. Vertex shiny, lacking wrinkles, with small, sparse punctures and few thin, scattered setae. Antennal calli poorly developed, triangular, their borders barely traceable. Supracallinal, midfrontal and suprafrontal sulci shallow, barely visible. Supraorbital sulcus deep and straight. Interantennal space wide. Labrum black. Maxillary palpomeres chestnut brown with yellow apices. Antennae with 11 antennomeres, antennomeres 1���4 yellow, continually darkening, antennomeres 5���10 brown, antennomere 11 lighter, yellowish-brown. Antennomere 7 with distal protrusion anteriorly. Antennomere 2 oval, ca. 1.4 �� longer than wide. Antennomeres 3 and 4 very thin, much thinner than antennomeres 1 and 2 each. Thorax. Pronotum convex, in the middle 1.79 times as wide as long, with small and shallow, sparse punctures. Lateral margin with setiferous pore situated nearly in middle. Elytra convex, humeral calli absent. Punctation of elytra fine, but coarser than that of pronotum, punctation confused. Metathoracic wings not developed. Anterior coxal cavity open posteriorly. Third tarsomere of all tarsi long, as long or slightly longer than first tarsomere. Inner side of metatibia with a row of teeth situated between distal 1 / 4 and tibial apex. Metafemur pilose. Metatibial apical spine long, reaching ca. 4 / 5 of first metatarsomere. Abdomen with 5 ventrites. Longitudinal ridge on first abdominal ventrite developed only basally and short, almost invisible. Metacoxal cavities very deep, reaching nearly mid length of the first ventrite. Genitalia. Spermathecal pump long, receptacle narrow, slender. Vaginal palpi slender. Aedeagus long, curved in lateral view, parallel-sided from base nearly to apex, then converging apically. Orificium basale very long, arrow-shaped (Fig. 2). Sexual dimorphism. Males generally smaller than females���male 1.94 mm long and female 2.21 mm long. Females with smoother frons, antennal calli less distinct than in males. Differential diagnosis. We place this species in Cangshanaltica based on the following characters: anterior coxal cavities open posteriorly; setiferous pore placed nearly in middle of lateral pronotal margin; antennomere 7 with a distal anterior protrusion (Fig. 1); male genitalia more or less flat in lateral view with indistinct broad depression ventrally, sides parallel at basal half, gradually narrowing without well-defined denticle at apex (Fig. 2). The new species may be distinguished from C. nigra Konstantinov et. al., 2013 based on the following characters: third tarsomeres or all tarsi long (Fig. 2), as long or slightly longer than first (in C. nigra, third tarsomere is significantly shorter than first, especially on metatarsi), longitudinal ridge on first abdominal ventrite very short, present only basally and hence nearly indistinct (in C. nigra, the ridge is long) (Fig. 2), metatibial apical spine long, reaching ca. 4 / 5 of first metatarsomere (in C. nigra, metatibial apical spine reaches at most 1 / 2 of first metatarsomere), upper side of body dark brown (in C. nigra, the upper side is black with metallic lustre). Cangshanaltica siamensis can be also distinguished by the some features of male genitalia: parallel sides of aedeagus extend further apically, apex only very slightly bent ventrally (in C. nigra the bend is more pronounced). Etymology. The name siamensisis derived from the old name for Thailand���Siam. Type locality and ecology. Specimens of C. siamensis were collected by beating moss from tree trunks in a cloud forest in the summit area of Doi Inthanon, ca. at 2 500 m a.s.l. (Fig. 3). Specimens were collected in semi-dry moss during the dry season. Type material. Holotype male. Labels: (1) THAILAND���Chiang Mai prov., Mae Chaem distr., Doi Inthanon, 18 �� 35 ' 10.36 "N, 98 �� 28 ' 57.18 "E, cloud forest in dry season, dry moss on trees, 6. iv. 2015, Albert Dama��ka lgt.; (2) HOLOTYPE Cangshanaltica siamensis sp nov., Albert Dama��ka and Alexander Konstantinov des. 2016. Paratypes have same labels as holotype. Holotype NMPC, paratypes: 1 female NMPC, 3 (1 male, 2 female) USNM, 2 (1 male, 1 female) ADPC., Published as part of Dama��ka, Albert & Konstantinov, Alexander, 2016, A new species of Cangshanaltica Konstantinov et al., a moss-inhabiting flea beetle from Thailand (Coleoptera: Chrysomelidae: Galerucinae: Alticini), pp. 93-97 in Zootaxa 4107 (1) on pages 93-95, DOI: 10.11646/zootaxa.4107.1.7, http://zenodo.org/record/259651, {"references":["Konstantinov, A., Chamorro, M. L., Prathapan, K. D., Ge, S. - Q. & Yang, X. - K. (2013) Moss-inhabiting flea beetles (Coleoptera: Chrysomelidae: Galerucinae: Alticini) with description of a new genus from Cangshan, China. Journal of Natural History, 47 (37 - 38), 2459 - 2477."]}