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Stegana (Steganina) nigripes Zhang & Chen, 2015 (Fig. 20A–C) Stegana (Steganina) nigripes Zhang & Chen, 2015: 135. Remarks. Since the terminalia of the only male specimen was partly damaged (probably eaten by a cockroach) and undescribed, the specimens collected from Yingjiang, Yunnan showed no other significant morphological differences with the type material of S. (S.) nigripes from Menghai, Yunnan. They were conservatively identified as conspecific. Description. Female terminalia (Fig. 20A–C): Epiproct and cercus with numerous setae, lacking pubescence (Fig. 20A). Eighth sternite about 2 times as wide as long, with equally distributed numerous prensisetae (Fig. 20B). Hypoproct with numerous setae and much pubescence (Fig. 20C). Specimens examined. CHINA: 1♂ (SCAU, No. 125409, DNA #1985), 1♀ (SCAU, No. 125408, DNA #1981), Tongbiguan, Yingjiang, Yunnan, 24°37'04''N, 97°40'03''E, 19.v.2016, alt. 1310m, 13.v.2016, ex tussock, J. Huang; 1♀ (SCAU, No. 125410, DNA #1981), Botanic Garden, Ruili, Yunnan, 24°04'19''N, 97°49'08''E, alt. 1170m, 22.v.2016, ex tussock, Y.Q. Liu. Distribution. China (Yunnan).

Stegana (Steganina) serratoprocessata Chen & Chen, 2009 Stegana (Steganina) serratoprocessata Chen & Chen, 2009a: 499. Specimens examined. CHINA: 3♂, 3♀ (SCAU, Nos. 125530–35, DNA #1045), Maoershan, Xing’an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 22–24.ix.2006, ex tree trunks, J.J. Jiang, F. Zhao; 3♂, 3♀ (SCAU, Nos. 125536–41, DNA #1949, 1950), Jiuwanshan, Huanjiang, Guangxi, 25°18'48''N, 108°38'02''E, alt. 1500m, 14.ix.2015, ex tree trunks, Y.Q. Liu; 1♂ (SCAU, No. 125542, DNA #2612), Shengtangshan, Jinxiu, Guangxi, 23°58'23''N, 110°06'45''E, alt. 1300m, 31.vii.2017, ex tree trunk, H.W. Chen; 1♂ (SCAU, Nos. 125543, DNA #599), Wuliangshan, Jingdong, Yunnan, 24°20'40''N, 100°48'01''E, alt. 2200m, 27.vii.2006, ex tree trunk, J.J. Gao; 2♂, 1♀ (SCAU, Nos. 125544–46, DNA #2699, 2731), Botanic Garden, Ruili, Yunnan, 24°04'19''N, 97°49'08''E, alt. 1170m, 22.iv.2018, ex tree trunks, H.W. Chen, L. Gong, Y. Lin; 1♂ (SCAU, No. 125547, DNA #2352), Menglai, Cangyuan, Yunnan, 23°13'20''N, 99°15'04''E, alt. 1240m, 30.x.2017, ex tree trunk, B.X. Li; 13♂, 9♀, Yixiang, Pu’er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m, 11♂, 5♀ (SCAU, Nos. 125548–63, DNA #1415, 1445, 1442, 2191) on 13.v.2012, ex tree trunks, H.W. Chen, J.J. Gao, 2♂, 4♀ (SCAU, Nos. 125564–69, DNA #1902, 2560, 2561) on 28– 30.iii.2013, ex tussocks, J.J. Gao; 1♂ (Nos. 125570, DNA #997), Tongmai, Bomi, Xizang, 30°06'00''N, 95°05'00''E, alt. 1000m, 10.x.2010, ex tree trunk, L. Wu. Distribution. China (Guangdong, Hainan, Guangxi *, Yunnan *, Xizang *)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 54-55, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Chen, X. P. & Chen, H. W. (2009 a) Four new species of the subgenus Steganina (Diptera: Drosophilidae) from southern China. Annales Zoologici, 59, 495 - 501. https: // doi. org / 10.3161 / 000345409 X 484874"]}

Stegana (Steganina) silvestrella Zhang & Chen, sp. nov. (Figs. 13G, 13H, 61) Diagnosis. This species is similar to S. (S.) flavipedes sp. nov. in the shape of male terminalia (Figs. 38, 61), but can be distinguished from the latter by having the pregonite blurry or absent (Fig. 61C, D); the aedeagus with numerous minute, serrated processes distally (Fig. 61C, D). Description. Male and female: First flagellomere yellow. Clypeus yellow. Mesonotum brown medially, brownish yellow laterally (Fig. 13G). Katepisternum yellowish (Fig. 13H). Scutellum dark brown in male, brownish yellow in female, black near distal margin (Fig. 13G). Halter mostly yellowish (Fig. 13H). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites brown medially, yellow laterally (Fig. 13G, H); sternites yellow, about 1.8 times as wide as long. Male terminalia (Fig. 61): Epandrium with ca. 9 setae on dorsal to posterior portions per side (Fig. 61A). Surstylus with some setulae near distal margin, lacking serrated processes and pubescence (Fig. 61A, B). Cercus with much pubescence (Fig. 61A). Hypandrium roundly expanded anteromedially (Fig. 61C, D). Aedeagus tubular, distally expanded and rounded, lacking pubescence (Fig. 61C, D). Aedeagal sheath lacking serrated process (Fig. 61C, D). Subepandrial sclerite slightly half-rounded (Fig. 61G). Measurements and indices. BL = 2.63 mm in holotype (2.80–3.40 in 2♀ paratypes), ThL = 1.33 mm (1.33 in ♀), WL = 2.13 mm (2.00– 2.47 in ♀), WW = 1.00 mm (1.00– 1.13 in ♀), arb = 4/3 (4–5/3–4), avd = 1.00 (0.80–1.14), adf = 1.25 (1.00–1.40), flw = 2.42 (2.01–2.50), FW/HW = 0.45 (0.39–0.53), ch/o = 0.09 (0.08–0.11), prorb = 1.38 (0.83–1.33), rcorb = 0.75 (0.50–0.78), vb = 0.67 (0.60–0.80), dcl = 0.46 (0.46–0.48), presctl = 0.72 (0.70–0.82), sctl = 2.34 (2.11–2.50), sterno = 0.80 (0.60–1.11), orbito = 2.67 (2.12–2.67), dcp = 0.28 (0.25–0.28), sctlp = 1.86 (1.75–2.50), C = 1.67 (1.56–2.00), 4c = 1.29 (1.13–1.50), 4v = 1.91 (1.43–2.01), 5x = 2.43 (2.01–2.60), ac = 10.02 (9.00–12.01), M = 0.76 (0.73–0.84), C3F = 0.59 (0.64–0.87). Type materials. Holotype ♂ (SCAU, No. 125657, DNA #2752), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 700m, 25.iv.2007, ex tree trunk, H.W. Chen. Paratypes: 2♀ (SCAU, Nos. 125658, 59, DNA #2562), Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt. 1320m, 10.v.2016, ex tree trunks, J. Huang, Y.L. Wang. Distribution. China (Yunnan). Etymology. From the Latin word " silvestris " (= arboreal), referring to the arboreal Steganina flies., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 96-98, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) cucullata Li & Chen, sp. nov. (Figs. 7C, 7D, 33) Diagnosis. This species closely resembles S. (S.) eurystoma Wang, Gao, & Chen, 2013 in the shape of male terminalia (as fig. 3 in Wang et al. 2013, Fig. 33), but can be distinguished from the latter by having the pregonite blurry or absent (Fig. 33C, D, J, K); the aedeagus lacking serrated process near distal margin (Fig. 33C, D, J, K); the aedeagal sheath distally slightly strong and dorsad curved (Fig. 33E, F, L, M). Remarks. The type specimens collected from Guangxi and Sichuan slightly differ from those from Chongqing in the surstylus lacking serrated process, with 1 peg-like tooth near distal margin (Fig. 33B, I), the cercus lacking pubescence (Fig. 33A, H), and the aedeagus with several minute, serrated processes distally and pubescence distoventrally (Fig. 33C, D, J, K). They were conservatively identified as conspecific. Description. Male and female: First flagellomere brown. Clypeus yellow. Mesonotum dark brown, with brownish yellow longitudinal stripes submedially and laterally (Fig. 7C). Katepisternum yellowish (Fig. 7D). Scutellum dark brown (Fig. 7C). Halter mostly grey (Fig. 7D). Fore femur with a row of 4 or 5 sturdy setae on distal part of posteroventral surface. Abdominal tergites dark brown (Fig. 7C, D); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 33): Epandrium with ca. 14 setae on dorsal to posterior portions per side (Fig. 33A, H). Surstylus with numerous setulae near distal margin (Fig. 33A, B, H, I). Hypandrium roundly expanded anteromedially (Fig. 33C, D, J, K). Pregonite with 1 or 2 sensilla (Fig. 33C, J, K). Aedeagus spoon-shaped, distally broadened (Fig. 33C, D, J, K). Subepandrial sclerite slightly vaulted, broadened distally (Fig. 33G, N). Measurement and indices. BL = 3.64 mm in holotype (2.35–3.15 in 5♂, 2.40–4.00 in 5♀ paratypes), ThL = 1.75 FW/HW = 0.37 (0.30–0.48), ch/o = 0.27 (0.18–0.36), prorb = 0.71 (0.76–1.38), rcorb = 0.53 (0.60–0.90), vb = 0.38 (0.50–0.78), dcl = damaged (0.36–0.75), presctl = 0.52 (0.31–0.78), sctl = 1.52 (1.56–2.13), sterno = 0.56 (0.55–1.17), orbito = 2.50 (1.20–2.40), dcp = 0.43 (0.15–0.34), sctlp = 1.64 (1.54–2.00), C = 1.71 (1.68–2.05), 4c = 1.24 (0.95–2.40), 4v = 1.88 (1.48–2.33), 5x = 1.57 (0.72–1.91), ac = 10.50 (7.33–12.67), M = 0.65 (0.53–0.82), C3F = 0.53 (0.54–0.72). Type materials. Holotype ♂ (SCAU, No. 121109, DNA #546), CHINA: Fengtongzhai, Baoxing, Sichuan, 30°32'42''N, 102°54'24''E, alt. 1700m, 19.ix.2005, ex tree trunk, H.W. Chen. Paratypes: 1♀ (SCAU, No. 121110), other data same as holotype; 5♂, 3♀ (SCAU, Nos. 125595–603, DNA #2609), 18.vii.2017, L. Gong, Y. Lin, Y.L. Wang, other data same as holotype; 1♂ (SCAU, No. 125604, DNA #983), Maoershan, Xing'an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 24.ix.2006, ex tree trunk, J.J. Jiang; 1♂ (SCAU, No. 125605, DNA #1782), Simianshan, Jiangjing, Chongqing, 28°37'19''N, 106°23'50''E, alt. 1108m, 7.vii.2014, ex tree trunk, Y.Q. Liu. Distribution. China (Guangxi, Chongqing, Sichuan). Etymology. From the Latin word " cucullatus " (= cucullate), referring to the spoon-shaped aedeagus.

VI. Stegana (Steganina) shirozui species group (26 spp.) alianya Zhang & Chen, 2021; China (Yunnan). ctenaria Nishiharu, 1979; Russia (Far East), China (Jilin, Liaoning, Taiwan), Japan (Hokkaido, Honshu, Kyushu), Korea. danbaensis Chen & Chen, 2012; China (Sichuan). dianensis Chen & Chen, 2012; China (Yunnan). diodonta Zhang & Chen, 2021; China (Yunnan). glabra Chen & Chen, 2012; China (Shaanxi *, Hubei *, Sichuan *, Guizhou, Yunnan). jiajinshanensis Chen, Gao, & Chen, 2009; China (Sichuan, Yunnan). jianfenglingensis Chen, Gao, & Chen, 2009; China (Hainan, Guangxi *). maoershanensis Chen, Gao, & Chen, 2009; China (Guangxi). masanoritodai Okada & Sidorenko, 1990; China (Jilin, Liaoning *), Russia (Far East), Japan (Hokkaido, Honshu, Kyushu). montana Chen & Chen, 2012; China (Yunnan). multidentata Chen, Gao, & Chen, 2009; China (Hubei, Sichuan *). oligochaeta Chen & Chen, 2012; China (Yunnan). parvispina Chen & Chen, 2012; China (Shaanxi). pianmaensis Chen & Chen, 2012; China (Yunnan). protuberans Chen & Chen, 2012; China (Yunnan). qinlingensis Chen, Gao, & Chen, 2009; China (Shaanxi, Sichuan). shennongi Chen, Gao, & Chen, 2009; China (Hubei). shirozui Okada, 1971; China (Taiwan). tentaculifera Chen & Chen, 2012; China (Hubei). unidentata Takada, 1968; Russia (Far East), China (Jilin, Liaoning, Taiwan), Korea, Japan (Hokkaido, Honshu, Kyushu). wuyishanensis Chen, Gao, & Chen, 2009; China (Jiangxi, Fujian, Yunnan). xanthosticta Chen, Gao, & Chen, 2009; China (Hubei). xishuangbanna Chen & Chen, 2012; China (Yunnan). zebromyia Zhang & Chen, 2021; China (Yunnan). zopheria Zhang & Chen, 2021; China (Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 105-106, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Nishiharu, S. (1979) Three new species of Drosophilidae from Japan (Diptera). Kontyu, Tokyo, 47, 38 - 43.","Chen, X. P. & Chen, H. W. (2012) Ten new species of the Stegana (Steganina) shirozui species group (Diptera, Drosophilidae) from China. Zootaxa, 3333 (1), 24 - 37. http: // dx. doi. org / 10.11646 / zootaxa. 3333.1.2","Okada, T. (1971) A revision and taxometric analysis of the genus Stegana Meigen of Japan and adjacent countries (Diptera, Drosophilidae). Mushi, 45, 81 - 99.","Takada, H. (1968) Drosophila survey of Hokkaido, XXVI. Descriptions of three new species of Drosophilidae from Japan. Journal of the Faculty of General Education, Sapporo University, 1, 119 - 127."]}

Stegana (Steganina) flabella Li & Chen, sp. nov. (Figs. 8C, 8D, 37) Diagnosis. This species closely resembles S. (S.) euryphylla Chen & Chen, 2009 in the shape of male terminalia (as figs. 14–20 in Chen & Chen 2009, Fig. 37), but can be distinguished from the latter by having the aedeagus distomedially and subdistolaterally lacking much pubescence (Fig. 37). Description. Male and female: First flagellomere black. Clypeus yellow. Mesonotum brown to dark brown medially, yellow to brownish laterally (Fig. 8C). Katepisternum yellowish (Fig. 8D). Scutellum dark brown (Fig. 8C). Halter mostly brownish (Fig. 8D). Fore femur with 3 or 4 long setae on distal part of posteroventral surface. Abdominal tergites dark brown to black (Fig. 8C, D); sternites brown, slightly wider than long. Male terminalia (Fig. 37): Epandrium with ca. 11 setae on dorsal to posterior portions per side (Fig. 37A). Surstylus with minute, serrated processes and numerous setulae on outer surface, 1 peg-like tooth near distal margin (Fig. 37A, B). Cercus lacking pubescence (Fig. 37A). Hypandrium expanded anteromedially (Fig. 37C, D). Pregonite small, with 3 sensilla (Fig. 37C, D). Aedeagus distally broadened and with numerous minute, serrated processes, with a row of serrated processes laterally and some pubescence distoventrally (Fig. 37C, D). Aedeagal sheath distally dorsad curved and with several minute, serrated processes (Fig. 37E, F). Subepandrial sclerite rounded distally (Fig. 37G). Measurements and indices. BL = 2.37 mm in holotype (2.19–2.39 in 3♂, 2.34–2.41 in 3♀ paratypes), ThL = 1.09 mm (1.09–1.12 in ♂, 0.95–1.21 in ♀), WL = 2.12 mm (2.03–2.13 in ♂, 1.91–2.32 in ♀), WW = 0.95 mm (0.87–0.97 in ♂, 0.84–1.01 in ♀), arb = 6/4 (6–7/4–5), avd = 0.71 (0.71–0.86), adf = 1.46 (1.05–1.34), flw = 1.85 (1.67–2.05), FW/HW = 0.44 (0.38–0.42), ch/o = 0.32 (0.25–0.38), prorb = 0.83 (0.68–1.32), rcorb = 0.67 (0.59– 0.92), vb = 0.51 (0.50–0.71), dcl = 0.76 (0.53–0.81), presctl = 0.65 (0.55–0.78), sctl = 1.96 (1.61–2.13), sterno = 0.71 (0.55–1.39), orbito = 2.00 (1.50–2.08), dcp = 0.29 (0.27–0.42), sctlp = 1.39 (1.39–1.82), C = 1.99 (1.71–2.13), 4c = 1.25 (1.00–1.59), 4v = 2.25 (1.57–2.37), 5x = 1.34 (1.32–1.75), ac = 8.44 (8.87–12.29), M = 0.71 (0.59–0.96), C3F = 0.60 (0.47–0.61). Type materials. Holotype ♂ (SCAU, No. 125710, DNA #2659), CHINA: Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 670m, 25.iv.2017, ex tree trunk, H.W. Chen. Paratypes: 3♂, 3♀ (SCAU, Nos. 125711–16, DNA #2691), other data same as holotype. Distribution. China (Yunnan). Etymology. From the Latin word " flabellum " (= knife), referring to the distally broadened aedeagus.

Stegana (Steganina) izu Sidorenko, 1997 (Fig. 24) Stegana (Steganina) izu Sidorenko, 1997b: 167. Remarks. The male specimens from Ryukyu Islands, Japan slightly differ from the type materials from Izu Islands, Japan in the distal shape of aedeagus and aedeagal sheath (as figs. 4–7 in Sidorenko 1997b, Fig. 24C–F). They were conservatively identified as conspecific. Description. Female terminalia (Fig. 24H–J): Epiproct with several setae, lacking pubescence (Fig. 24H). Cercus with numerous setae and much pubescence (Fig. 24H). Eighth sternite about 1.5 times as wide as long, glabrous on basal 2/3, with some setae near posterior margin (Fig. 24I). Hypoproct with numerous setae and much pubescence (Fig. 24J). Specimens examined. CHINA: 9♀ (SCAU, Nos. 125427–35, DNA #1893, 1977–79), Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 660m, 21–25.iv.2007, ex tree trunks, H.W. Chen, J.J. Gao; JAPAN: 2♂ (SCAU, Nos. 125806, 07, DNA #1872), Iriomote Island, Ryukyu Islands, 24°20'31''N, 123°49'20''E, alt. 330m, 14,18. vi.1999, ex tree trunks, H.W. Chen. Distribution. China (Taiwan, Yunnan*), Japan (Honshu, Ryukyu Islands*)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 51-52, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Sidorenko, V. S. (1997 b) New Asian species and new records of the genus Stegana Meigen (Diptera, Drosophilidae). II. Subgenera Pseudostegana Okada and Steganina Wheeler. Annales de la Societe entomologique de France, New Series, 33, 165 - 172."]}

I. Stegana (Steganina) biprotrusa species group (5 spp.) acantha Wu, Gao, & Chen, 2010; China (Hainan). ancistrophylla Wu, Gao, & Chen, 2010; China (Guangdong, Guangxi, Guizhou *, Yunnan *). langufoliacea Wu, Gao, & Chen, 2010; China (Taiwan, Guangxi, Yunnan*). otocondyloda Wu, Gao, & Chen, 2010; China (Yunnan). iaspidea Zhang & Chen, sp. nov.; China (Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 104, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) jianfenglingensis Chen, Gao, & Chen, 2009 Stegana (Steganina) jianfenglingensis Chen, Gao, & Chen in Chen et al. 2009: 1914. Specimens examined. CHINA: 1♂ (SCAU, Nos. 125426, DNA #2656), Nonggang, Longzhou, Guangxi, 22°32'26''N, 106°43'18''E, alt. 230m, 23.viii.2004, ex tree trunk, H.W. Chen. Distribution. China (Hainan, Guangxi *)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 49, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) zhuoma Cui & Chen, sp. nov. (Figs. 14G, 14H, 65D–F) Diagnosis. This species is similar to S. (S.) zaduo sp. nov. in the shape of female terminalia (Fig. 65), but can be distinguished from the latter by having the yellowish katepisternum brown on anterior corner (Fig. 14H); the 8th sternite nearly squared, with several setae and much pubescence on distolateral 1/3 (Fig. 65E). Description. Female: First flagellomere brown. Clypeus black. Mesonotum black medially, brown laterally (Fig. 14G). Scutellum black, yellow at tip (Fig. 14G). Halter mostly white (Fig. 14H). Fore femur with a row of 3 long, strong setae on distal part of posteroventral surface. All abdominal tergites dark brown to black (Fig. 14G, H); sternites brown, about 2 times as wide as long. Female terminalia (Fig. 65D–F): Epiproct with several setae, lacking pubescence (Fig. 65D). Cercus and hypoproct with numerous setae and much pubescence (Fig. 65D, F). Eighth sternite glabrous on basal 2/3 (Fig. 65E). Measurements and indices. BL = 2.70 mm in holotype, ThL = 1.45 mm, WL = 2.92 mm, WW = 1.26 mm, arb = 7/5, avd = 0.85, adf = 1.17, flw = 2.22, FW/HW = 0.39, ch/o = 0.25, prorb = 1.02, rcorb = 0.76, vb = 0.74, dcl = 0.48, presctl = 0.57, sctl = damaged, sterno = 0.77, orbito = 1.90, dcp = 0.32, sctlp = 1.45, C = 1.66, 4c = 1.35, 4v = 1.95, 5x = 1.45, ac = 12.89, M = 0.71, C3F = 0.63. Type material. Holotype ♀ (SCAU, No. 125789, DNA #2757), Tongmai, Bomi, Xizang, 30°06'00''N, 95°05'00''E, alt. 2000m, 11.x.2010, ex tree trunk, L. Wang. Distribution. China (Xizang). Etymology. From the word " zhuoma " in Zang language from the Zang nationality living in Xizang, China, referring to women., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 102, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) bimai Cui & Chen, sp. nov. (Figs. 6E, 6F, 29) Diagnosis. This species closely resembles S. (S.) clavispinula Chen & Chen, 2009 in the shape of epandrium, surstylus, cercus (as figs. 1, 2 in Chen & Chen 2009, Fig. 29A, B), and aedeagal sheath in ventral view (as fig. 6 in Chen & Chen 2009, Fig. 29E), but can be distinguished from the latter by having the surstylus with 1 peg-like tooth near distal margin (Fig. 29B); the aedeagus expanded distally in ventral view (Fig. 29D), lacking serrated process in median process (Fig. 29C, D); the subepandrial sclerite slightly X-shaped (Fig. 29G). Description. Male and female: First flagellomere brown. Clypeus yellow. Mesonotum brown medially, yellow laterally (Fig. 6E). Scutellum brown, yellow along margin (Fig. 6E). Katepisternum yellowish (Fig. 6F). Halter mostly grey (Fig. 6F). Fore femur with 3 thin, long setae on distal part of posteroventral surface. Abdominal tergites brown (Fig. 6E, F); sternites brownish, about 2 times as wide as long in male, about 3 times as wide as long in female. Male terminalia (Fig. 29): Epandrium with ca. 15 setae on dorsal to posterior portions per side (Fig. 29A). Surstylus slightly squared, with minute, serrated processes distally and several setulae near distal margin (Fig. 29A, B). Cercus lacking pubescence (Fig. 29A). Hypandrium expanded anteromedially (Fig. 29C, D). Pregonite small, with 3 sensilla (Fig. 29C, D). Aedeagus with some pubescence distoventrally (Fig. 29C). Aedeagal sheath with several minute, serrated processes distally (Fig. 29E, F). Measurements and indices. BL = 2.63 mm in holotype (2.52–2.61 in 2♂, 2.48–2.65 in 2♀ paratypes), ThL = 1.31 mm (1.31–1.34 in ♂, 1.26–1.28 in ♀), WL = 2.32 mm (2.28–2.51 in ♂, 2.31–2.61 in ♀), WW = 1.00 mm (0.97–1.01 in ♂, 0.95–1.08 in ♀), arb = 6/4 (7/4–5), avd = 0.65 (0.84–0.93), adf = 1.12 (1.08–1.20), flw = 2.12 (1.87–2.39), FW/HW = 0.40 (0.36–0.44), ch/o = 0.26 (0.24–0.27), prorb = 0.68 (0.87–1.07), rcorb = 0.71 (0.60– 0.71), vb = 0.60 (0.61–0.89), dcl = damaged (0.55–0.72), presctl = damaged (0.46–0.66), sctl = 2.19 (2.17–2.89), sterno = 0.39 (0.47–0.93), orbito = 1.80 (1.80–1.98), dcp = 0.30 (0.28–0.36), sctlp = 1.78 (1.50–1.73), C = 1.67 (1.72–1.98), 4c = 1.17 (1.22–1.28), 4v = 1.83 (1.88–2.19), 5x = 1.81 (1.44–1.86), ac = 15.17 (9.00–13.14), M = 0.74 (0.61–0.71), C3F = 0.55 (0.60–0.64). Type materials. Holotype ♂ (SCAU, No. 125744, DNA #2706), CHINA: Muyiji Park, Ximeng, Yunnan, 22°37'23''N, 99°36'11''E, alt. 1100m, 16.iv.2018, ex tree trunk, L. Zhu. Paratypes: 2♀ (SCAU, Nos. 125745, 46), other data same as holotype; 2♂ (SCAU, Nos. 125747, 48, DNA #2727), Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 860m, 24.xii.2003, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. From the word " bimai " in Dai language from the Dai nationality living in Yunnan, China, referring to the Water-Sprinkling Festival (new year)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 57-59, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657

Stegana (Steganina) hirticlavata Cui & Chen, sp. nov. (Figs. 9G, 9H, 44) Diagnosis. This species is similar to S. (S.) jianqinae Zhang, Tsaur, & Chen, 2014 in the shape of male terminalia (as fig. 8 in Zhang et al. 2014, Fig. 44), but can be distinguished from the latter by having the first flagellomere yellow on basal 2/3, black on distal 2/3 (Fig. 9H); the pregonite fused basally, with much long pubescence distally (Fig. 44C, D). Description. Male and female: Face mostly yellow, black near lower margin; facial carina absent. Clypeus yellow. Mesonotum yellow, with blurry longitudinal stripes (Fig. 9G). Katepisternum yellowish (Fig. 9H). Scutellum yellow, brown along margin in male (Fig. 9G). Halter mostly yellow (Fig. 9H). Fore femur with a row of 3 strong setae on distal part of posteroventral surface. Abdominal tergites brownish (Fig. 9G, H); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 44): Epandrium with 4 setae near dorsal portion per side (Fig. 44A). Surstylus with numerous setulae on outer surface and some pubescence medially (Fig. 44A, B). Cercus with much pubescence (Fig. 44A). Hypandrium expanded anteromedially (Fig. 44C, D). Aedeagus expanded distally, with a pair of rod-like processes (Fig. 44C, D). Aedeagal sheath slightly saddle-shaped (Fig. 44E, F). Subepandrial sclerite slightly quadrate (Fig. 44G). Measurements and indices. BL = 2.64 mm in holotype (2.54 in 1♂, 2.52–2.88 in 5♀ paratypes), ThL = 1.33 mm (1.53 in ♂, 1.31–1.57 in ♀), WL = 2.65 mm (2.56 in ♂, 2.32–2.85 in ♀), WW = 1.28 mm (1.40 in ♂, 1.21–1.34 in ♀), arb = 7/5 (7–8/5–6), avd = 0.82 (0.74–0.80), adf = 1.31 (1.13–1.56), flw = 2.33 (2.17–2.72), FW/HW = 0.34 (0.34–0.39), ch/o = 0.17 (0.12–0.22), prorb = 0.88 (0.82–1.01), rcorb = 0.74 (0.63–0.77), vb = 0.82 (0.56–0.86), dcl = 0.44 (0.35–0.68), presctl = 0.45 (0.45–0.73), sctl = 1.84 (1.74–2.00), sterno = 0.74 (0.70–0.93), orbito = 2.00 (1.74–2.04), dcp = 0.14 (0.14–0.29), sctlp = 1.75 (1.31–1.98), C = 1.74 (1.83–1.95), 4c = 1.06 (0.97–1.10), 4v = 1.39 (1.28–1.55), 5x = 1.31 (1.17–1.27), ac = 13.00 (8.17–15.43), M = 0.52 (0.43–0.53), C3F = 0.70 (0.59–0.69). Type materials. Holotype ♂ (SCAU, No. 125571, DNA #1970), CHINA: Baihualing, Baoshan, Yunnan, 25°17'50''N, 98°48'12''E, alt. 1700m, 21.vi.2013, ex tussock, H.W. Chen. Paratypes: 4♀ (SCAU, Nos. 125572–75, DNA #2177), Mengdong, Cangyuan, Yunnan, 23°10'08''N, 99°13'52''E, alt. 1320m, 6.v.2016, ex tree trunks, J. Huang, Y.Q. Liu, Y.L. Wang, L. Zhu; 1♀ (SCAU, No. 125577, DNA #1890), Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m, 28.v.2015, ex tree trunk, H.W. Chen; 1♂ (SCAU, No. 125576, DNA #2181), Qimaba, Lüchun, Yunnan, 22°49'08''N, 102°17'25''E, alt. 1280m, 3.viii.2016, ex tree trunk, H.W. Chen. Distribution. China (Yunnan). Etymology. A combination of the Latin words '' hirtus '' (= hirsute) and '' clavatus '' (= rod-like), referring to the pregonite distally with long pubescence., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 75-76, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Zhang, Y., Tsaur, S. C. & Chen, H. W. (2014) Survey of the genus Stegana Meigen (Diptera, Drosophilidae) from Taiwan, with"]}

Stegana (Steganina) nigrolimbata Duda, 1924 Stegana (Steganina) nigrolimbata Duda, 1924: 181; Okada, 1971: 83; Cao & Chen, 2008: 29. Specimens examined. CHINA: 7♂, 4♀ (SCAU, Nos. 125307–17, DNA #1961, 1964), Maolan, Libo, Guizhou, 25°14'45''N, 107°57'31''E, alt. 750m, 19.ix.2015, ex tree trunks, J. Huang, Y.Q. Liu, Y.L. Wang, Y. Zhang. Distribution. China (Taiwan, Guangdong, Hainan, Guangxi, Guizhou*, Yunnan)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 44, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Duda, O. (1924) Beitrag zur Systematik der Drosophiliden unter besonderer Berucksichtigung der palaarktischen u. orientalischen Arten (Dipteren). Archiv f ¸ r Naturgeschichte (A), 90, 172 - 234.","Okada, T. (1971) A revision and taxometric analysis of the genus Stegana Meigen of Japan and adjacent countries (Diptera, Drosophilidae). Mushi, 45, 81 - 99.","Chen, X. P. & Chen, H. W. (2008) The Stegana coleoptrata species group (Diptera: Drosophilidae) from mainland China. Zootaxa, 1891 (1), 55 - 65. https: // doi. org / 10.11646 / zootaxa. 1891.1.5"]}

Stegana (Steganina) tongi Wang, Gao, & Chen, 2011 Stegana (Steganina) tongi Wang, Gao, & Chen in Wang et al. 2011: 510. Specimens examined. CHINA: 1♂, 2♀, Yixiang, Pu’er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m: 1♂ (SCAU, No. 125237, DNA #2601) on 2.x.2011, ex tree trunk, H.W. Chen, 2♀ (SCAU, Nos. 125238, 39, DNA #2192) on 28–30.iii.2013, ex tussocks, J.J. Gao; 1♂, 1♀ (SCAU, Nos. 125240, 41, DNA #2633), Menglun, Mengla, Yunnan, 21°47'29''N, 101°22'59''E, alt. 560m, 19.xii.2017, ex tree trunks, H.W. Chen, L. Gong. Distribution. China (Taiwan, Guangdong, Hainan, Guangxi, Yunnan*)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on page 41, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Wang, J. Q., Gao, J. J. & Chen, H. W. (2011) Stegana castanea species group (Diptera, Drosophilidae) from the Oriental region. Journal of Natural History, 45, 505 - 519. https: // doi. org / 10.1080 / 00222933.2010.534239"]}

Stegana (Steganina) lancang Li & Chen, sp. nov. (Figs. 10E, 10F, 46) Diagnosis. This species is similar to S. (S.) kanmiyai Okada & Sidorenko, 1992 in the shape of male terminalia (as fig. 3E–G in Okada & Sidorenko 1992, Fig. 46A–G), but can be distinguished from the latter by having the brownish yellow body color (Fig. 10E, F); the surstylus with 1 peg-like tooth near distal margin (Fig. 46A, B); the aedeagus lacking pubescence distolaterally (Fig. 46C, D). Description. Male: Clypeus and palpus yellow. Mesonotum brownish yellow (Fig. 10E). Katepisternum yellowish (Fig. 10F). Scutellum brownish yellow, brownish near distal margin (Fig. 10E). Halter mostly grey (Fig. 10F). Fore femur with a row of 3 strong setae on distal part of posteroventral surface. Abdominal tergites brownish yellow (Fig. 10E, F); sternites yellow, about 2 times as wide as long. Male terminalia (Fig. 46A–G): Epandrium with 13 setae on dorsal to posterior portions per side (Fig. 46A). Surstylus with numerous setulae distally and some minute, serrated processes distoventrally (Fig. 46C, D). Aedeagal sheath acute apically, with 1 pair of acute processes subapically (Fig. 46E, F). Subepandrial sclerite vaulted (Fig. 46G). Female terminalia (Fig. 46H–J): Epiproct with several setae on distal 1/3 (Fig. 46H). Cercus and hypoproct with numerous setae and much pubescence (Fig. 46H, J). Eight sternite about 2 times as long as wide, mostly glabrous except with some setae distally (Fig. 46I). Measurements and indices. BL = 2.74 mm in holotype (2.33–2.67 in 5♂, 2.39–2.73 in 4♀ paratypes), ThL = 1.32 mm (1.21–1.31 in ♂, 1.21–1.37 in ♀), WL = 2.48 mm (2.16–2.43 in ♂, 2.27–2.63 in ♀), WW = 0.99 mm (0.95–1.02 in ♂, 0.95–1.05 in ♀), arb = 6/4 (6–8/4–5), avd = 0.88 (0.74–0.89), adf = 1.05 (1.00–1.74), flw = 1.81 (1.56–2.15), FW/HW = 0.47 (0.34–0.43), ch/o = 0.27 (0.22–0.30), prorb = 0.93 (0.77–1.53), rcorb = 0.60 (0.62–0.91), vb = 0.52 (0.46–0.83), dcl = 0.87 (0.59–0.87), presctl = 0.52 (0.60–0.84), sctl = damaged (1.41–1.94), sterno = 0.52 (0.63–1.08), orbito = 1.80 (1.59–1.96), dcp = 0.30 (0.24–0.35), sctlp = 1.67 (1.27–2.19), C = 1.76 (1.65–2.11), 4c = 1.26 (0.96–1.30), 4v = 1.86 (1.44–1.84), 5x = 1.59 (1.26–1.76), ac = 10.67 (8.17–11.00), M = 0.67 (0.55–0.72), C3F = 0.68 (0.57–0.66). Type materials. Holotype ♂ (SCAU, No. 125697, DNA #2637), CHINA: Guanlei, Mengla, Yunnan, 21°38'39''N, 101°09'52''E, alt. 560m, 24.iv.2016, ex tree trunk, J. Huang. Paratypes: 4♀ (SCAU, Nos. 125698–701, DNA #2637), other data same as holotype; 8♂ (SCAU, Nos. 125702–09, DNA #2638), Wangtianshu, Mengla, Yunnan, 21°36'09''N, 101°35'06''E, alt. 560m, 17.xii.2017, ex tree trunks, H.W. Chen, L. Gong. Distribution. China (Yunnan). Etymology. From the word " lancang " in Dai language from the Dai nationality living in Yunnan, China, referring to the Lancang River., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 79-80, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Okada, T. & Sidorenko, V. S. (1992) New or unrecorded species of Stegana (Steganina) (Diptera, Drosophilidae) from the Old World, especially Japan and Russia. Japanese Journal of Entomology, 60, 415 - 426."]}

Stegana (Steganina) kanmiyai Okada & Sidorenko, 1992 (Fig. 25) Stegana (Steganina) kanmiyai Okada & Sidorenko, 1992: 419. Diagnosis. This species is similar to S. (S.) lancang sp. nov. in the shape of male terminalia (Figs. 25, 46A–G), but can be distinguished from the latter by having the cercus lacking pubescence (Fig. 25A); the aedeagus with numerous minute, serrated processes along anterior margin and pubescence anteromedially (Fig. 25C, D). Specimens examined. CHINA: 2♂ (SCAU, Nos. 125436, 37, DNA #785), Mangshan, Yizhang, Hunan, 24°56'53''N, 112°54'01''E, alt. 1260m, 3.x.2004, ex tree trunk, M.F. Xu; 1♂ (SCAU, No. 125438, DNA #2602), Chebaling, Shixing, Guangdong, 24°43'22''N, 114°31'49''E, alt. 780m, 11.ix.2017, ex tussock, L. Gong; 5♂, 11♀ (SCAU, Nos. 125439–54, DNA #786, 1046, 1184, 2759, 2760), Conghua, Guangdong, 23°43'30''N, 113°48'20''E, alt. 500m, 3♂, 6♀ on 3.xi.2003, 2♂, 5♀ on 10.x.2005, ex tree trunks, H.W. Chen; 1♂, 4♀ (SCAU, Nos. 125455– 59, DNA #457, 2709), Dinghushan, Zhaoqing, Guangdong, 23°10'17''N, 112°32'31''E, alt. 580–670m, ex tree trunks, 5,6. vii.2005, J.J. Jiang, MF Xu; 1♀ (SCAU, No. 125460, DNA #2185), Tianluhu, Guangzhou, Guangdong, 23°13'43''N, 113°25'34''E, alt. 360m, 6.ix.2015, ex tree trunk, J. Huang; 4♂, 7♀ (SCAU, Nos. 125461–71, DNA #1193), Maoershan, Xing'an, Guangxi, 25°51'28''N, 110°25'01''E, alt. 700m, 23.x.2004, ex tree trunks and tussocks, H.L. Cao, H.W. Chen, MF Xu; 1♀ (SCAU, No. 125472, DNA #1962), Mulun, Huanjiang, Guangxi, 25°07'13''N, 108°02'34''E, alt. 500m, 16.ix.2015, ex tree trunk, J. Huang; 3♂, 2♀ (SCAU, Nos. 125473–77, DNA #2621), Guposhan, Hezhou, Guangxi, 24°36'10''N, 111°35'18''E, alt. 670m, 29.vii.2017, ex tree trunks, H.W. Chen; 1♂, 1♀ (SCAU, Nos. 125478, 79, DNA #2610), Shengtangshan, Jinxiu, Guangxi, 23°58'23''N, 110°06'45''E, alt. 1300m, 31.vii.2017, ex tree trunks, H.W. Chen; 1♀ (SCAU, No. 125480, DNA #1466), Bapen, Fusui, Guangxi, 22°32'36'' N, 107°53'18''E, alt. 210m, 18.viii.2004, ex tree trunk, H.W. Chen; 1♂, 5♀ (SCAU, Nos. 125481–86, DNA #796), Emeishan, Leshan, Sichuan, 29°36'14''N, 103°28'56''E, alt. 700m, 5.vii.2007, ex tussocks, H.Z. Cao, J.J. Gao, J.J. Jiang; 1♀ (SCAU, No. 125487, DNA #1896), Baihualing, Baoshan, Yunnan, 25°17'50''N, 98°48'12''E, alt. 1420m, 25.viii.2013, ex tussock, Q.S. Gao; 1♂, 1♀ (SCAU, Nos. 125488, 89, DNA #2351), Menglai, Cangyuan, Yunnan, 23°13'20''N, 99°15'04''E, alt. 1240m, 30.x.2017, ex tree trunks, H.W. Chen, L. Gong; 1♂, 2♀ (SCAU, Nos. 125490–92, DNA #1444), Yixiang, Pu'er, Yunnan, 22°44'19''N, 101°07'01''E, alt. 1400m, 13.v.2012, ex tree trunks, H.W. Chen; 1♂, 1♀ (SCAU, Nos. 125493, 94, DNA #2664), Jinshan, Menglian, Yunnan, 22°19'29''N, 99°35'00''E, alt. 940m, 13,14. iv.2018, ex tree trunks, H.W. Chen. Distribution. China (Hunan *, Taiwan, Guangdong*, Guangxi*, Sichuan*, Yunnan*), Japan (Honshu)., Published as part of WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI & HUANG, JIA, 2023, Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae), pp. 1-109 in Zootaxa 5250 (1) on pages 52-54, DOI: 10.11646/zootaxa.5250.1.1, http://zenodo.org/record/7705657, {"references":["Okada, T. & Sidorenko, V. S. (1992) New or unrecorded species of Stegana (Steganina) (Diptera, Drosophilidae) from the Old World, especially Japan and Russia. Japanese Journal of Entomology, 60, 415 - 426."]}