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We present a high‐resolution 2‐D P‐wave velocity model from a 225‐km‐long active seismic profile, collected over ~60–75 Ma central Atlantic crust. The profile crosses five ridge segments separated by a transform and three nontransform offsets. All ridge discontinuities share similar primary characteristics, independent of the offset. We identify two types of crustal segment. The first displays a classic two‐layer velocity structure with a high gradient Layer 2 (~0.9 s−1) above a lower gradient Layer 3 (0.2 s−1). Here, PmP coincides with the 7.5 km s−1 contour, and velocity increases to >7.8 km s−1 within 1 km below. We interpret these segments as magmatically robust, with PmP representing a petrological boundary between crust and mantle. The second has a reduced contrast in velocity gradient between the upper and lower crust and PmP shallower than the 7.5 km s−1 contour. We interpret these segments as tectonically dominated, with PmP representing a serpentinized (alteration) front. While velocity‐depth profiles fit within previous envelopes for slow‐spreading crust, our results suggest that such generalizations give a misleading impression of uniformity. We estimate that the two crustal styles are present in equal proportions on the floor of the Atlantic. Within two tectonically dominated segments, we make the first wide‐angle seismic identifications of buried oceanic core complexes in mature (>20 Ma) Atlantic Ocean crust. They have a ~20‐km‐wide "domal" morphology with shallow basement and increased upper crustal velocities. We interpret their midcrustal seismic velocity inversions as alteration and rock‐type assemblage contrasts across crustal‐scale detachment faults. Key Points: We identify that magmatic and tectonic modes of slow Atlantic crustal accretion are distinct and equal, based on seismic characteristicsThe structure of crustal discontinuities is controlled by the accretion mode of neighboring segments, independent of ridge offset and orderTwo buried oceanic core complexes in mature Atlantic Ocean crust are identified, based on their dimensions and seismic structure [ABSTRACT FROM AUTHOR]

North America's ~1.1‐Ga failed Midcontinent Rift (MCR) is a striking feature of gravity and magnetic anomaly maps across the continent. However, how the rift affected the underlying lithosphere is not well understood. With data from the Superior Province Rifting Earthscope Experiment and the USArray Transportable Array, we constrain three‐dimensional seismic velocity discontinuity structure in the lithosphere beneath the southwestward arm of the MCR using S‐to‐P receiver functions. We image a velocity increase with depth associated with the Moho at depths of 33–40 ± 4 km, generally deepening toward the east. The Moho amplitude decreases beneath the rift axis in Minnesota and Wisconsin, where the velocity gradient is more gradual, possibly due to crustal underplating. We see hints of a deeper velocity increase at 61 ± 4‐km depth that may be the base of underplating. Beneath the rift axis further south in Iowa, we image two distinct positive phases at 34–39 ± 4 and 62–65 ± 4 km likely related to an altered Moho and an underplated layer. We image velocity decreases with depth at depths of 90–190 ± 7 km in some locations that do not geographically correlate with the rift. These include a discontinuity at depths of 90–120 ± 7 km with a northerly dip in the south that abruptly deepens to 150–190 ± 7 km across the Spirit Lake Tectonic Zone provincial suture. The negative phases may represent a patchy, frozen‐in midlithosphere discontinuity feature that likely predates the MCR and/or be related to lithospheric thickness. Key Points: S‐to‐P receiver functions constrain a Moho, crustal underplating and midlithosphere discontinuities beneath the Midcontinent RiftThe Moho and crustal underplating directly beneath the rift axis vary in character from north to southMidlithospheric discontinuities predate the rifting event and seem unperturbed by rift‐related activity [ABSTRACT FROM AUTHOR]