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Crossipalpus gersoni Beard and Seeman sp. nov. (Figs 25���31) Type material examined. Holotype female ex. stems of Allocasuarina luehmannii (Casuarinaceae), AUSTRALIA: Queensland, near Mount Slopeaway, on the old Marlborough-Sarina Road, 22 �� 52 ��� 16 ��� S 149 �� 50 ��� 28 ��� E, 19 March 2005, coll. J.J. Beard and P.I. Forster (QM). Paratypes. 14 females, 6 males, 4 deutonymphs, 3 protonymphs, and 4 larvae, same data as holotype (QM, ANIC, USNM). Diagnosis. Dorsal setae short, broadly lanceolate, strongly barbed; dorsal setae subequal in length except c 1, d 1, e 1, h 1 obviously shorter. Palpal segments as wide as long. Genua I���II with seta d, without l������; tarsi I���IV without seta tc������. Solenidia of male much thicker and longer than in female. Female (n = 15). Dorsum. (Fig. 25 a) Body measurements: distance between setae v 2 -h 1 275���325 [325]; sc 2 - sc 2 88���110 [110]; other measurements: v 2 -v 2 19���22 [22], sc 1 -sc 1 69���89 [89], c 1 -c 1 29���37 [37], c 2 -c 2 105���125 [125], c 3 -c 3 125���145 [145], d 1 -d 1 23���28 [28], d 2 -d 2 89���110 [110], d 3 -d 3 105���130 [130], e 1 - e 1 6���17 [6], e 2 - e 2 92���120 [120], e 3 - e 3 83���100 [100], f 3 -f 3 71���89 [89], h 1 -h 1 14���18 [17], h 2 -h 2 49���58 [58]. Gnathosoma completely concealed beneath prodorsum (Fig. 26). Anterior margin of prodorsum rounded. Seta v 2 inserted dorsally, though its setal base is sometimes partially concealed by an anterior fold (Fig. 25 a). Prodorsal shield weakly developed with papillate-rugose sculpturing laterally, longitudinal grooves medially. Opisthosomal shield weakly developed with rugose to lineate pattern, becoming weakly rugose medially between setae d 1 -e 1. Lateral cuticle surrounding shields rugose-papillate. Most dorsal setae broadly lanceolate, barbed; medial opisthosomal setae shorter than lateral setae, palmate. Setal lengths: v 2 13���17 [17], sc 1 15���19 [19], sc 2 15���21 [21], c 1 8���11 [11], c 2 13���15 [15], c 3 15���22 [22], d 1 5���9 [9], d 2 12���14 [14], d 3 13���18 [18], e 1 4 ���7 [7], e 2 14 ���17 [17], e 3 14 ���17 [17], f 3 15���18 [18], h 1 5���7 [7], h 2 13���17 [17]. Palps. (Figs 25 b, 26) Setal formula 0, 0, 0, 1, 3 (1 s+ 2 e). Tibial seta 6���8 [6] long; tarsal eupathidia 6 long, 5���6 [5] long; solenidion 5���6 [6] long. Vent er. (Figs 26, 27 a) Cuticle with fine transverse striae between setae 1 a - 3 a, longitudinal between setae 3 a - 4 a, then convex for 20���30, striae longitudinal posterior to setae 4 a; fine striae become coarse lateral to genital area. Genital setae inserted in more-or-less transverse row, g 1 inserted slightly posterior to level of g 2. Genital shield smooth, poorly developed, ca. 15���20 [16] long, 35���40 [38] wide; anal setae ps 1���2 inserted medially on anal plates in longitudinal line; seta ps 3 absent. Coxal setae fine, except 2 c heavily barbed; setae ag 1, g 1���2, ps 1���2 lanceolate. Setal lengths: 1a 44 ��� 55 [44], 1 b 20���26 [25], 2 b 13���19 [19], 2 c 14���17 [17], 3a 35 ��� 50 [50], 3 b 12���22 [22], 4a 40 ��� 46 [45], 4 b 13���17 [17], ag 1 9���11 [11], g 1 12���15 [15], g 2 12���14 [13], ps 1 7���11 [11], ps 2 8���10 [10]. Spermatheca. (Fig. 27 b) Spermathecal tube long, distinctly broad near external opening, becoming narrow and convoluted distally, maximum 2 wide, ca. 100 long. Thickened part of tube 52���56 long. Spermatheca vesicle not visible. Genital opening between setae ps 2. Legs. (Fig. 28) Setal formula for legs I���IV (coxae to tarsi) 1 - 1-3 - 1-4 - 8 (1), 2 - 1-3 - 1-4 - 8 (1), 1-2 - 2 - 0-3 - 4, 1 - 1 - 1 - 0-3 - 4. Tarsi I and II each with 1 antiaxial solenidion ��" (7���8 [8] long) and 2 eupathidia p��'-p��" (6���7 [7] long, 6���8 [8] long). Leg setation as in Table 1 except: coxae I without 1 c; genua I���II with d, without l ��� and l������; tarsi I���IV without tc������. Setae v' added to tr IV. MALE (6 paratypes). Dorsum. ( Figs 29, 30 a) Body measurements: distance between setae v 2 -h 1 195���225, sc 2 -sc 2 76���81; other measurements: v 2 -v 2 11���13, sc 1 -sc 1 56���63, c 1 -c 1 25���28, c 2 -c 2 78���87, c 3 -c 3 99���110, d 1 -d 1 10���14, d 2 -d 2 68���71, d 3 -d 3 79���87, e 1 - e 1 7���8, e 2 - e 2 67���71, e 3 - e 3 61���65, f 3 -f 3 52���58, h 1 -h 1 8���10, h 2 -h 2 30���40. Gnathosoma not concealed beneath prodorsum. Anterior margin of prodorsum smooth, weakly convex. Prodorsal shield weakly developed. Opisthosoma with mesonotal shield (appearing as a pair of shields) on which at least setae c 1, d 1 and d 2 inserted, with weak rugose-lineate pattern; and pygidial shield with at least setae e 1, e 3, f 3, h 2 inserted, with oblique-longitudinal weakly lineate pattern; setae h 1 often under posterior extension of pygidial shield (Fig. 30 a); shields separated by transverse striae; cuticle laterad mesonotal shield papillate; several minute pores visible on shields. Setal lengths: v 2 13���15, sc 1 13���17, sc 2 15���18, c 1 9���11, c 2 10���12, c 3 12���17, d 1 8���9, d 2 10���12, d 3 12���16, e 1 5, e 2 13 ���15, e 3 12 ���16, f 3 13���14, h 1 6���7, h 2 12���14. Palps. (Fig. 29) Palps similar to female. Tibial seta 8 long; tarsal eupathidia 5���6, 6 long; solenidion 7 long. Venter. (Figs 30 b, c) All striae transverse, becoming coarse on opisthogaster and weak around setae ag 1. Coxal setae fine, except 2 c narrowly lanceolate. Setae ag 1, g 2 thick, barbed; g 1, ps 2 thin, barbed; setae ps 1 modified to form thick blades (sexually dimorphic) (Figs 29; 30 a, b). Setal lengths: 1a 33 ��� 40, 1 b 23���25, 2 b 18���20, 2 c 10���15, 3 a 42���45, 3 b 10���12, 4 a 36���45, 4 b 10��� 14, ag 1 7���10, g 1 7���10, g 2 10���11, ps 1 13���16, ps 2 6. Aedeagus. (Figs 30 c, d) Narrow, sclerotised, tapering to a point, 55���61 long. Membranous duct runs from inside aedeagus, becoming indistinguishable. Legs. (Fig. 29) Setal formula same as female. Tarsi I and II each with 1 antiaxial solenidion ��" (ta I 9���10 long, ta II 9 long) and 2 eupathidia p��'-p��" (ta I 5���6, 6 long; ta II 6, 6 long). Solenidia much thicker and longer than in female. DEUTONYMPH (4 paratypes). Dorsum. Body measurements: distance between setae v 2 -h 1 215���250, sc 2 -sc 2 71���87; other measurements: v 2 -v 2 18���22, sc 1 -sc 1 57���67, c 1 -c 1 22, c 2 -c 2 76���90, c 3 -c 3 97���113, d 1 -d 1 15���20, d 2 - d 2 56���68, d 3 -d 3 77���94, e 1 - e 1 7���11, e 2 - e 2 68���81, e 3 - e 3 62���78, f 3 -f 3 52���62, h 1 -h 1 11���13, h 2 -h 2 35���39. Prodorsal shield weakly developed with oblique-longitudinal striations. Opisthosoma with setae c 1, d 1 and d 2 on paired, weak platelets. Striae transverse, becoming convex posteriorly. Setal lengths: v 2 12���18, sc 1 16���17, sc 2 16���20, c 1 11���13, c 2 14���17, c 3 13���16, d 1 8���11, d 2 13���17, d 3 14���17, e 1 5, e 2 14 ���17, e 3 14 ���15, f 3 13���15, h 1 6���8, h 2 13���16. Palps. Palps similar to adult. Tibial seta 6 long; tarsal eupathidia 3, 4 long, solenidion 4 long. Venter. Cuticle with transverse striae anteriorly, longitudinal between setae 3 a- 4 a, then transverse, becoming concave, to ag 1, then transverse and coarse to posterior margin. Coxal setae fine, except 2 c barbed; setae ag 1 narrowly lanceolate, g 1 barbed. Setal lengths: 1a 35 ��� 40, 1 b 12���16, 2 b 11���17, 2 c 12���14, 3 a 26���36, 3 b 10���15, 4 a 27���30, 4 b 10���15, ag 1 8��� 10, g 1 8���12, ps 1 6���8, ps 2 6���8. Legs. Setal formula for legs I���IV (coxae to tarsi) 1 - 1-3 - 1-4 - 8 (1), 2 - 1-3 - 1-4 - 8 (1), 1- 2 - 2 - 0-3 - 4, 1 - 0-1 - 0-3 - 4. Tarsi I and II each with 1 antiaxial solenidion ��" (ta I 4���6 long, ta II 4���5 long) and 2 eupathidia p��'-p��" (5, 5���6 long). Leg setation as in adult except: tr IV without seta v ���. Setae v' added to tr I���III. PROTONYMPH (3 paratypes). Dorsum. Body measurements: distance between setae v 2 -h 1 158���181, sc 2 -sc 2 64���70; other measurements: v 2 -v 2 16���19, sc 1 -sc 1 52���57, c 1 -c 1 16���24, c 2 -c 2 63���76, c 3 -c 3 97���99, d 1 -d 1 16���17, d 2 -d 2 49���51, d 3 -d 3 67���73, e 1 - e 1 6, e 2 - e 2 64���67, e 3 - e 3 57���61, f 3 -f 3 43���44, h 1 -h 1 22���24, h 2 -h 2 7���9. Prodorsal shield weakly developed. Opisthosoma with setae c 1, d 1 and d 2 on paired, weak platelets. Striae of idiosoma similar to deutonymph. Setal lengths: v 2 13���15, sc 1 14���15, sc 2 14���15, c 1 12���13, c 2 14���16, c 3 13���14, d 1 11, d 2 13���14, d 3 14, e 1 4 ���5, e 2 12, e 3 14 ���15, f 3 9���13, h 1 4���5, h 2 9���15. Palps. Palps similar to deutonymph except solenidion 3 long. Venter. Same as deutonymph. Coxal setae fine, except 2 b, ag 1 barbed. Setal lengths: 1a 36, 1 b 11���13, 2 b 13���16, 3 a 30, 3 b 9, ag 1 7, ps 1 5, ps 2 5���6. Setae 2 c, 4 a, 4 b, g 1, g 2 absent. Legs. Setal formula for legs I��� IV (coxae to tarsi) 1 - 0-3 - 0-4 - 8 (1), 1 - 0-3 - 0-4 - 8 (1), 1 - 1-2 - 0-3 - 4, 0- 0-1 - 0-3 - 3. Tarsi I and II each with 1 antiaxial solenidion ��" (4 long) and 2 eupathidia p��'-p��" (5, 4 ��� 5 long). Leg setation as in deutonymph except: seta 2 c absent; seta 4 b absent; tr I���III without seta v ���; ge I���II without seta d; ta IV without setae tc ���. Setae l' added to tr III. LARVA (4 paratypes). Dorsum. (Fig. 31) Body measurements: distance between setae v 2 -h 1 130���150, sc 2 -sc 2 55���60; other measurements: v 2 -v 2 13���15, sc 1 -sc 1 43���49, c 1 -c 1 12���16, c 2 -c 2 51���55, c 3 -c 3 84���85, d 1 -d 1 13���15, e 1 - e 1 4���5, e 2 - e 2 57���62, e 3 - e 3 38���44, f 3 -f 3 11���13, h 1 -h 1 5���7, h 2 -h 2 10���13. Prodorsal shield weakly formed, 58��� 60 long, 61���66 wide, with few irregular striations. Opisthosomal shields absent; coarse, irregular transverse striae becoming oblique posteriorly. Setal lengths: v 2 11���14, sc 1 12���13, sc 2 14���15, c 1 10, c 2 12���14, c 3 11, d 1 9���11, d 2 11���14, d 3 11���13, e 1 3 ���4, e 2 11 ���15, e 3 11 ���12, f 3 11���13, h 1 4���5, h 2 11���14. Palps. (Fig. 31 a) Palps same as deutonymph. Tibial seta 5���6 long; tarsal eupathidia 2, 3 long, solenidion 2���3 long. Venter. (Fig. 31 b) Striation same as deutonymph. All setae fine. Setal lengths: 1a 17 ��� 27, 1 b 9���13, 3 a 25���33, ps 1 4���5, ps 2 4���5. Legs. (Fig. 31 c) Setal formula for legs I���III (coxae to tarsi) 1 - 0-3 - 0-4 - 7 (1), 0- 0-3 - 0-4 - 7 (1), 0- 0-2 - 0-3 - 3. Tarsi I and II each with 1 antiaxial solenidion ��" (3 long) and 2 eupathidia p��'-p��" (ta I 4, 4 long; ta II 5, 5 long). Leg setation as in protonymph except: seta 2 b absent; seta 3 b absent; tr I���III nude; ta I���III without seta tc ���. Etymology. It is with great pleasure that we name this species for our colleague and friend Prof. Uri Gerson, in recognition of his acarological works, especially those on the Tegopalpinae. Remarks. Crossipalpus gersoni sp. nov. was found on Bull-Oak Allocasuarina luehmannii in Eucalyptus fibrosa woodland on red soil, with vine thicket in small patches. This species is similar to Cr. raveni sp. nov., as they both have seta d present on ge I���II, but Cr. gersoni has spatulate setae d on femora and genua I���II (lanceolate in Cr. raveni), and posterior ventral setae are thick, broadly lanceolate and strongly barbed (thin, setiform and weakly barbed in Cr. raveni)., Published as part of Beard, Jennifer J., Seeman, Owen D. & Bauchan, Gary R., 2014, Tenuipalpidae (Acari: Trombidiformes) from Casuarinaceae (Fagales), pp. 1-157 in Zootaxa 3778 (1) on pages 36-44, DOI: 10.11646/zootaxa.3778.1.1, http://zenodo.org/record/251337

Crossipalpus Smiley, Frost and Gerson, 1996 Type species. Crossipalpus verticillatae Smiley, Frost and Gerson, 1996, by original designation. Diagnosis. All life stages: dorsal opisthosoma with 12 pairs of thinly to broadly lanceolate setae; c 2, d 2, and e 2 present; setae f 2 absent; setae e 2 close to marginal position; setae h 2 similar in size and form to other dorsal setae; palps 5 -segmented, setal formula 0, 0, 0, 1, 3 (1); anterior margin of prodorsum smoothly rounded, without projections/notches; ventral plate absent; 2 pairs of pseudanal setae (ps 1���2) on weakly developed anal plates. Adult female: anterior margin of prodorsum without projections, smoothly rounded, completely concealing the gnathosoma; genital plate membranous, weakly developed; metapodal plates not developed; coxae I without 1 c; trochanters I���IV 1 - 1 -2- 1; femora I���IV 3 - 3 - 2 - 1; genua I���IV 1 - 1 -0-0 (ge I���II with either l������ or d present); tibiae I���IV 4 - 4 - 3 - 3; tarsi I���IV with or without tc������. Setae v ��� added to tr IV in adult; v ��� added to tr I���III in deutonymph; l ��� added in protonymph. Solenidia of male much thicker and longer than those of female. Species. Four species: Cr. gersoni, Cr. muellerianae, Cr. raveni, Cr. verticillatae. Hosts and distribution. Casuarinaceae, Australia. Remarks. All known Crossipalpus species have the setal formula for genua I���IV 1 - 1 -0-0. In Cr. muellerianae and Cr. verticillatae, the seta present on ge I���II is l������, but in the two new species, Cr. gersoni and Cr. raveni, the seta is d. This difference is an example of how setal counts alone can conceal useful information that only chaetotaxy can present., Published as part of Beard, Jennifer J., Seeman, Owen D. & Bauchan, Gary R., 2014, Tenuipalpidae (Acari: Trombidiformes) from Casuarinaceae (Fagales), pp. 1-157 in Zootaxa 3778 (1) on page 22, DOI: 10.11646/zootaxa.3778.1.1, http://zenodo.org/record/251337, {"references":["Smiley, R. L., Frost, W. E. & Gerson, U. (1996) A revision of the subfamily Tegopalpinae, with the description of two new genera and five new species (Acari: Tenuipalpidae). International Journal of Acarology, 22 (3), 167 - 180. http: // dx. doi. org / 10.1080 / 01647959608684092"]}

Crossipalpus raveni Beard and Seeman sp. nov. (Figs 32���37) Type material examined. Holotype female ex. Woolly Oak Allocasuarina inophloia (Casuarinaceae), AUSTRALIA: Queensland, Moonie Highway, 90 km W Moonie, 27 �� 56 ��� 57 ��� S 149 �� 31 ��� 19 ��� E, 0 5 May 2007, coll. J.J. Beard and P.I. Forster (QM; BRI voucher PIF 32483). Paratypes. 9 females, 5 males, and 4 larvae, same data as holotype (QM, ANIC, USNM). Diagnosis. Dorsal setae lanceolate, strongly barbed; setae e 1 shortest. Palpal segments as wide as long. Genua I���II with seta d, without l������; tarsi I���IV without tc������. Solenidia of male much thicker and longer than in female. FEMALE (n = 10). Dorsum. (Fig. 32 a) Body measurements: distance between setae v 2 -h 1 310���365 in 9 paratypes [355]; sc 2 -sc 2 100���110 [110]; other measurements: v 2 -v 2 22���24 [24], sc 1 -sc 1 86���98 [98], c 1 -c 1 15���22 [22], c 2 -c 2 100���115 [115], c 3 -c 3 125���145 [135], d 1 -d 1 14���21 [21], d 2 -d 2 85���95 [94], d 3 -d 3 110���120 [120], e 1 - e 1 14���18 [16], e 2 -e 2 100���110 [110], e 3 - e 3 89���100 [95], f 3 -f 3 70���80 [73], h 1 -h 1 12���17 [13], h 2 -h 2 35���54 [48]. Gnathosoma completely concealed beneath prodorsum. Anterior margin of prodorsum rounded. Prodorsal shield weakly developed with oblique grooves laterally, longitudinal grooves medially. Opisthosomal shield weakly developed with oblique-longitudinal lineate grooves, becoming transverse between setae d 1 -e 1. Lateral cuticle surrounding shields smooth. Dorsal setae lanceolate, barbed; setae e 1 shorter than other setae. Setal lengths: v 2 21��� 25 [22], sc 1 20���25 [20], sc 2 20���22 [20], c 1 15���19 [15], c 2 14���20 [16], c 3 17���21 [18], d 1 12���19 [15], d 2 14���21 [18], d 3 17���21 [18], e 1 7 ���9 [8], e 2 17 ���24 [19], e 3 17 ���24 [20], f 3 18���23 [21], h 1 9���12 [11], h 2 21���24 [21]. Palps. (Fig. 32 b) Setal formula 0, 0, 0, 1, 3 (1 s+ 2 e). Tibial seta 9���11 [9] long; tarsal eupathidia 6 [6], 5 [5] long; solenidion 4���5 [4] long. Venter. (Fig. 33 a) Cuticle with fine transverse striae between setae 1 a - 3 a, longitudinal between setae 3 a - 4 a, then convex for 20���30 ��m, striae longitudinal posterior to setae 4 a; fine striae become coarse lateral to genital area. Genital setae inserted in more-or-less transverse row, g 1 inserted slightly posterior to level of g 2. Genital shield smooth, poorly developed; anal setae ps 1���2 inserted medially on anal plates in longitudinal line; seta ps 3 absent. Coxal setae fine, 2 c barbed; setae ag 1, g 1���2, ps 1���2 fine, with few barbs. Setal lengths: 1a 64 ��� 83 [64], 1 b 18���30 [21], 2 b 18���25 [18], 2 c 18���21 [19], 3a 35 ��� 66 [35], 3 b 16���22 [17], 4a 42 ��� 92 [45], 4 b 14���26 [15], ag 1 14���18 [14], g 1 20���24 [21], g 2 18���23 [19], ps 1 11���16 [14], ps 2 10���13 [13]. Spermatheca. (Fig. 33 b) Spermathecal tube long, becoming narrow and sometimes convoluted distally, maximum 2 wide, ca. 80 long. Spermatheca vesicle not oval-shaped, 4 long, 2 wide. Genital opening between setae ps 2. Legs. (Fig. 34) Setal formula for legs I���IV (coxae to tarsi) 1 - 1-3 - 1-4 - 8 (1), 2 - 1-3 - 1-4 - 8 (1), 1-2 - 2 - 0-3 - 4, 1 - 1 - 1 - 0-3 - 4. Tarsi I and II each with 1 antiaxial solenidion ��" (5���6 long) and 2 eupathidia p��'-p��" (7���8 [7] long). Leg setation as in Table 1 except: coxae I without 1 c; genua I���II with d, without l������; tarsi I���IV without tc������. Setae v' added to tr IV. MALE (5 paratypes). Dorsum. (Fig. 35) Body measurements: distance between setae v 2 -h 1 245���265, sc 2 -sc 2 83���91; other measurements: v 2 -v 2 13���17, sc 1 -sc 1 69���74, c 1 -c 1 12���17, c 2 -c 2 85���94, c 3 -c 3 96���112, d 1 -d 1 11���13, d 2 -d 2 69���74, d 3 -d 3 78���87, e 1 - e 1 11���17, e 2 - e 2 73���81, e 3 - e 3 69���77, f 3 -f 3 57���66, h 1 -h 1 7���10, h 2 -h 2 35���43. Gnathosoma not concealed beneath prodorsum. Anterior margin of prodorsum smooth, weakly convex. Prodorsal shield weakly developed. Opisthosoma with mesonotal shield (appearing as a pair of shields) on which at least setae c 1, d 1 and d 2 inserted, with weak rugose-lineate pattern; and pygidial shield with at least setae e 1, e 3, f 3, h 2 inserted, with oblique-longitudinal weakly lineate pattern; setae h 1 often under posterior overhang of pygidial shield; shields separated by transverse striae; cuticle laterad mesonotal shield smooth with few papillations; several minute pores visible on shields. Setal lengths: v 2 19���24, sc 1 16���19, sc 2 15���20, c 1 9���14, c 2 11���16, c 3 15���17, d 1 7��� 13, d 2 9���15, d 3 12���18, e 1 6 ���8, e 2 14 ���17, e 3 15 ���18, f 3 15���19, h 1 7���10, h 2 15���19. Palps. Palps similar to female. Tibial seta 8���10 long; tarsal eupathidia 6���7, 5 ��� 6 long; solenidion 7���8 long, swollen. Venter. (Fig. 36 a) All striae transverse, becoming coarse posteriorly and weak around setae ag 1. Coxal setae fine, except 2 c barbed. Setae ag 1 barbed; g 1, g 2, ps 2 fine; setae ps 1 modified to form thick blades (sexually dimorphic). Setal lengths: 1a 47 ��� 52, 1 b 19���27, 2 b 19���24, 2 c 13���21, 3 a 46���60, 3 b 17���21, 4 a 37���48, 4 b 15���18, ag 1 13���16, g 1 11���15, g 2 15���20, ps 1 15���16, ps 2 8���11. Aedeagus. (Fig. 36 b) Narrow, sclerotised, tapering to a point, 67���70 long. Membranous duct runs from inside aedeagus, becoming indistinguishable. Legs. (Fig. 35) Setal formula same as female. Tarsi I and II each with 1 antiaxial solenidion ��" (ta I 9���11 long, ta II 9���10 long) and 2 eupathidia p��'-p��" (ta I 6���7, 5��� 6; ta II 7, 7���8). Solenidia much thicker and longer than those in female. DEUTONYMPH and PROTONYMPH. Unknown. LARVA (4 paratypes). Dorsum. (Fig. 37) Body measurements: distance between setae v 2 -h 1 170���180, sc 2 -sc 2 61���64; other measurements: v 2 -v 2 15���16, sc 1 -sc 1 51���55, c 1 -c 1 8���9, c 2 -c 2 58���60, c 3 -c 3 84���87, d 1 -d 1 11���13, e 1 - e 1 5���6, e 2 - e 2 53���54, e 3 - e 3 40���44, f 3 -f 3 34���35, h 1 -h 1 5���7, h 2 -h 2 13���17. Prodorsal shield obsolete. Opisthosomal shields absent; sparse irregular transverse striae anteriorly. Setal lengths: v 2 24���27, sc 1 12���16, sc 2 15���17, c 1 17��� 22, c 2 15���17, c 3 10���11, d 1 15���20, d 2 14���18, d 3 11���12, e 1 4 ���8, e 2 15 ���17, e 3 14 ���17, f 3 16���19, h 1 5���6, h 2 18���19. Palps. Palps same as adult. Tibial seta 6���7 long; tarsal eupathidia 3, 4 long, solenidion 2 long. Venter. Transverse striae, becoming oblique around anal region. All setae fine. Setal lengths: 1a 21 ��� 28, 1 b 13���16, 3 a 16���30, ps 1 4���6, ps 2 4���6. Legs. (Fig. 37) Setal formula for legs I���III (coxae to tarsi) 1 - 0-3 - 0-4 - 7 (1), 0- 0-3 - 0-4 - 7 (1), 0- 0-2 - 0-3 - 3. Tarsi I and II each with 1 antiaxial solenidion ��" (3 long) and 2 eupathidia p��'-p��" (4���5 long). Leg setation as in Table 1 except genua I���III without l ���. Trochanters I���III nude. Etymology. It is with great pleasure that we name this species for our colleague and friend Dr Robert Raven, in recognition of his support for our work and his immense contribution to Arachnology. Remarks. Crossipalpus raveni sp. nov. is similar to Cr. gersoni sp. nov., as they both have seta d present on ge I���II, but Cr. raveni has lanceolate setae d on femora and genua I���II (spatulate in Cr. gersoni), and posterior ventral setae are thin, setiform and weakly barbed (broadly lanceolate and strongly barbed in Cr. gersoni)., Published as part of Beard, Jennifer J., Seeman, Owen D. & Bauchan, Gary R., 2014, Tenuipalpidae (Acari: Trombidiformes) from Casuarinaceae (Fagales), pp. 1-157 in Zootaxa 3778 (1) on pages 44-51, DOI: 10.11646/zootaxa.3778.1.1, http://zenodo.org/record/251337

Crossipalpus verticillatae Smiley, Frost and Gerson, 1996 (Figs 19���24) Crossipalpus verticillatae Smiley, Frost and Gerson, 1996: 174, figs 16���18. Type material examined. Holotype female ex. Drooping She-Oak Allocasuarina verticillata (Casuarinaceae), AUSTRALIA: South Australia, 10 km WNW Nairne, 0 9 September 1995, coll. W.E. Frost (WINC; 1 male paratype, deutonymph on same slide). Paratypes. 1 male, 1 deutonymph, same slide as holotype (WINC); 1 female, same data as holotype (WINC); 3 females, 1 male, and 1 deutonymph, ex. same host (tree # 258) [feeding on bark of branchlets], AUSTRALIA: South Australia, Glen Osmond, Waite Institute Arboretum, 11 January 1996, coll. W.E. Frost (WINC; 1 slide); 1 female, 1 deutonymph, 4 protonymphs, and 3 larvae ex. same host [feeding on bark of branchlets], AUSTRALIA: South Australia, 8 km south of Harrogate, 34.57 S 139.01 E, 10 November 1995, W.E. Frost (WINC; one slide); 1 pharate female, 1 male, 3 deutonymphs, and 1 protonymph, same data (except 5 km south of Harrogate) (WINC; one slide). Non-type material examined. 12 females, 1 male, 1 pharate male, 1 deutonymph, and 1 protonymph ex. A. verticillata, AUSTRALIA: South Australia, Adelaide, Athelston, Maryvale Road, Wadmore Park, 34 �� 52 ��� 51 ��� S 138 �� 42 ���09��� E, 21 April 2009, coll. J.J. Beard and C.J. Burwell (QM, USNM) (BRI voucher BRI [AQ 818801]). Diagnosis. Dorsal setae all broadly lanceolate, strongly barbed and subequal in length except c 1, d 1, e 1, h 1 distinctly short; prodorsal setae v 2 subequal in length with sc 1. Palpal segments as wide as long. Genua I���II with seta l������, without d; tarsi I���IV without seta tc������. FEMALE (n = 18). Dorsum. (Fig. 19 a) Body measurements: distance between setae v 2 -h 1 310���330 [330]; sc 2 - sc 2 105���110 [110]; other measurements: v 2 -v 2 20���25 [20], sc 1 -sc 1 87���93 [90], c 1 -c 1 15���25 [19], c 2 -c 2 120���130 [125], c 3 -c 3 140���150 [140], d 1 -d 1 17���19 [19], d 2 -d 2 95���105 [95], d 3 -d 3 120���130 [120], e 1 - e 1 13���25 [21], e 2 -e 2 110���115 [110], e 3 - e 3 98���105 [98], f 3 -f 3 78���89 [78], h 1 -h 1 12���19 [12], h 2 -h 2 52���61 [52]. Gnathosoma completely concealed beneath prodorsum; anterior margin of prodorsum rounded; setae v 2 often inserted ventrally under anterior margin of prodorsum (similar to Fig. 14). Prodorsal shield weakly developed with broadly separated longitudinal central folds and oblique lateral folds; cuticle with fine network of folds laterally. Narrow band of fine transverse striae anterior to c 1 -c 1. Opisthosoma with pair of weakly developed mesonotal shields capturing setae c 1, c 2, d 1, d 2; weakly developed pygidial shield capturing e 1, e 3, f 2���3 and h 1���2 setae, with weak oblique folds; band of transverse folds between d 1 -d 1 and e 1 -e 1; paired minute pores present between c 1 -c 2, d 1 -d 2, laterad e 1. Lateral cuticle surrounding shields wrinkled. Most dorsal setae broadly lanceolate and barbed; setae c 1, d 1, e 1, h 1 much shorter than other dorsal setae. Setal lengths: v 2 15���20 [15, 19], sc 1 17���20 [19, 20], sc 2 19���22 [21, 22], c 1 6���7 [6, 7], c 2 13���17 [13, 16], c 3 19���21 [19. 21], d 1 6���8 [7, 8], d 2 14���18 [14, 15], d 3 19���24 [20, 21], e 1 6 ���8 [6, 7], e 2 17 ���22 [20, 21], e 3 20 ���23 [20, 21], f 3 20���22 [20, 22], h 1 7���10 [7, 8], h 2 19���22 [19, 22]. Palps. (Fig. 19 b) Setal formula 0, 0, 0, 1, 3 (1 s+ 2 e). Tibial seta 8���10 [10] long; tarsal eupathidia 4���5 [5] long (thin, tapered), 5���6 [6] long (blunt); solenidion 5���6 [5]. Vent er. (Fig. 20) Ventral cuticle of gnathosoma to setae 1 a transverse; striae longitudinal between 1 a- 1 a; striae between 1 a- 3 a transverse, between 3 a- 4 a longitudinal; transverse striae 4 a to just posterior 4 a, becoming longitudinal to g 1���2; longitudinal striae laterad genital region. Genital setae not widely separated (25), inserted in more-or-less transverse row, g 1 inserted slightly posterior to level of g 2. Genital shield weakly developed, cuticle on genital flap between g setae with longitudinal to oblique striae; anal setae ps 1���2 inserted in longitudinal line along inner edge of anal plates; seta ps 3 absent. Coxal, genital and anal setae fine and weakly barbed, except 2 c thicker and more strongly barbed; setae 1 a, 3 a, 4 a long, finely tapered, often broken and difficult to determine full length. Setal lengths: 1a 52 ��� 59 [54, 58], 1 b 22���25 [22, 24], 2 b 21���25 [22, 24], 2 c 15���19 [15, 16], 3a 45 ��� 68 [65, 68], 3 b 21���24 [21, 23], 4a 46 ��� 52 [48, 51], 4 b 20���24 [22, 24], ag 1 14���17 [14, 16], g 1 21���25 [22, 23], g 2 18���22 [20, 21], ps 1 12���13 [12, 13], ps 2 11���14 [11, 14]. Spermatheca. Spermatheca with a long coiled tube, with some granular texture along its length; tightly coiled and multi-branched distally. Distal vesicle not visible. Legs. (Fig. 21) Setal formula for legs I���IV (coxae to tarsi) 1 - 1-3 - 1-4 - 8 (1), 2 - 1-3 - 1-4 - 8 (1), 1-2 - 2 - 0-3 - 4, 1 - 1 - 1 - 0-3 - 4. Tarsi I and II each with 1 antiaxial solenidion ��" (6���7 [6] long) and 2 eupathidia p��'-p��" (7���8 long). Leg setation as in Table 1 except: coxae I without 1 c; genua I���II with l", without d; tarsi I���IV without tc������. Setae v' added to tr IV. MALE (n = 5). Dorsum. (Fig. 22) Body measurements: distance between setae v 2 -h 1 230���250, sc 2 -sc 2 87���91; other measurements: v 2 -v 2 12���18, sc 1 -sc 1 61���66, c 1 -c 1 14���21, c 2 -c 2 96���105, c 3 -c 3 110���117, d 1 -d 1 8���12, d 2 -d 2 74���79, d 3 -d 3 90���96, e 1 - e 1 11���13, e 2 - e 2 77���80, e 3 - e 3 74���77, f 3 -f 3 61���67, h 1 -h 1 6���10, h 2 -h 2 38���42. Gnathosoma fully exposed, not concealed by anterior margin of prodorsum (Fig. 22). Four weak dorsal shields developed, separated by bands of transversely striate soft cuticle. Prodorsal shield similar to female. Mesonotal shield with 2 weak shields, weakly wrinkled or folded. Pygidial shield with weak longitudinal-oblique wrinkles or folds. Lateral cuticle surrounding shields with some weak papillae. Most dorsal setae short, lanceolate, barbed; setae c 1, d 1 and e 1 much smaller than other setae. Setal lengths: v 2 14���18, sc 1 13���15, sc 2 16���20, c 1 5���6, c 2 9���12, c 3 12���14, d 1 5��� 7, d 2 9���13, d 3 13���15, e 1 5 ���8, e 2 10 ���15, e 3 14 ���16, f 3 14���16, h 1 7, h 2 14���15. Palps. (Fig. 22) Palps similar to female, except solenidion greatly swollen; solenidion 7���8 long, eupathidia 6���8 long (blunt), 5���6 long (thin, tapered). Venter. (Fig. 23 a) Cuticle with mostly transverse striae, with band of broadly separated transverse folds just posterior to leg IV; broadly separated transverse folds between ag and g 1���2. Pair of large rounded, weakly developed shields laterad setae ag. Coxal setae fine. Setae 1 a, 3 a, 4 a elongate, finely tapered, often broken. Setae ag 1, g 1���2, ps 2 weakly barbed; setae ps 1 modified, obviously thicker and stronger than ps 2. Setal lengths: 1a 37 ��� 49, 1 b 16���22, 2 b 15���22, 2 c 15���20, 3 a 34���68, 3 b 16���23, 4 a 31���63, 4 b 13���21, ag 1 11���13, g 1 11���14, g 2 12���17, ps 1 15���17, ps 2 7���10. Aedeagus. (Fig. 23 a) Narrow, sclerotised, finely tapered distally, 81���92. Membranous duct runs from inside aedeagus for about 40 then disappearing. Legs. (Fig. 23 b) Legs similar to female, except solenidia obviously swollen. Tarsi I and II each with 1 antiaxial solenidion ��" (10���11 long) and 2 eupathidia p��'-p��" (7���9, 7 ��� 8 long). DEUTONYMPH (n = 8). Dorsum. Body measurements: distance between setae v 2 -h 1 250���295, sc 2 -sc 2 84��� 100; other measurements: v 2 -v 2 17���23, sc 1 -sc 1 70���87, c 1 -c 1 17���23, c 2 -c 2 91���112, c 3 -c 3 119���136, d 1 -d 1 15���22, d 2 -d 2 70���93, d 3 -d 3 86���106, e 1 - e 1 12���14, e 2 - e 2 79���99, e 3 - e 3 72���90, f 3 -f 3 65���76, h 1 -h 1 14���16, h 2 -h 2 41���52. Dorsal shields with patterns similar to adult female. Setal lengths: v 2 21���26, sc 1 17���22, sc 2 19���24, c 1 5���7, c 2 14��� 19, c 3 16���22, d 1 5���6, d 2 16���21, d 3 18���22, e 1 5 ���7, e 2 19 ���25, e 3 20 ���24, f 3 19���24, h 1 6���8, h 2 16���23. Palps. Palps same as adult. Tibial seta 8���9 long; tarsal eupathidia 4���5 long (thin, tapered), 5���7 long (thick, blunt); solenidion 3��� 4 long. Venter. Cuticle with longitudinal striae between 1 a- 1 a; transverse striae between 1 a- 4 a, with mixed striae 4 a- 4 a; transverse striae just posterior to 4 a becoming longitudinal to setae g 1; transverse striae between g 1 -ps 1, with longitudinal striae laterad genital region. Coxal setae fine and finely barbed, except setae 2 c thicker and more heavily barbed. Setal lengths: 1a 31 ��� 52, 1 b 12���22, 2 b 12���21, 2 c 19���22, 3 a 32���57, 3 b 13���16, 4 a 20���42, 4 b 12���18, ag 1 13���16, g 1 12���19, ps 1 9���13, ps 2 10���15. Legs. Setal formula for legs I���IV (coxae to tarsi) same as adult female except tr IV nude: 1 - 1-3 - 1-4 - 8 (1), 2 - 1-3 - 1-4 - 8 (1), 1-2 - 2 - 0-3 - 4, 1 - 0-1 - 0-3 - 4. Tarsi I and II each with 1 antiaxial solenidion ��" (4 long) and 2 eupathidia p��'-p��" (ta I 6, 6 long; ta II 5���6, 5��� 6 long). Leg setation as in adult except: tr IV without seta v ���. Setae v' added to tr I���III. PROTONYMPH (n = 6). Dorsum. Body measurements: distance between setae v 2 -h 1 190���250, sc 2 -sc 2 75��� 91; other measurements: v 2 -v 2 13���17, sc 1 -sc 1 60���76, c 1 -c 1 5���15, c 2 -c 2 85���105, c 3 -c 3 101���130, d 1 -d 1 10���12, d 2 - d 2 61���77, d 3 -d 3 66���97, e 1 - e 1 7���12, e 2 - e 2 68���88, e 3 - e 3 51���71, f 3 -f 3 41���54, h 1 -h 1 8���12, h 2 -h 2 22���30. Prodorsal shield weakly developed, with similar cuticle pattern to female. Anterior opisthosoma with 2 pairs of weakly developed platelets, between c 1 -c 2 and d 1 -d 2; platelets with few transverse folds. Setae in rows E, F, H inserted on weakly formed pygidial shield, with few oblique folds. Striae of idiosoma similar to deutonymph. Setal lengths: v 2 22���24, sc 1 14���18, sc 2 17���20, c 1 5���6, c 2 12���17, c 3 13���16, d 1 4���5, d 2 17���18, d 3 15���19, e 1 4 ���7, e 2 19 ���22, e 3 16 ��� 19, f 3 16���19, h 1 5���7, h 2 17���18. Palps. Palps similar to deutonymph except tibial seta 6���8 long; tarsal eupathidia 3���4 long (thin, tapered), 4���5 long (thick, blunt); solenidion 3���4 long. Venter. Cuticle between 1 a- 1 a with longitudinal striae; striae 1 a and leg III with transverse striae; cuticle between leg III to IV with longitudinal striae; cuticle between legs IV transverse to oblique near ag; cuticle between ag-ag mixed; cuticle laterad genital-anal region longitudinal. Setal lengths: 1a 25 ��� 54, 1 b 13���24, 2 b 15���16, 3 a 22���49, 3 b 10���14, ag 1 9���13, ps 1 7���9, ps 2 7��� 9. Setae 2 c, 4 a, 4 b, g 1, g 2 absent. Legs. Setal formula for legs I���IV (coxae to tarsi) 1 - 0-3 - 0-4 - 8 (1), 1 - 0-3 - 0-4 - 8 (1), 1 - 1-2 - 0-3 - 4, 0- 0-1 - 0-3 - 3. Tarsi I and II each with 1 antiaxial solenidion ��" (3���4 long) and 2 eupathidia p��'-p��" (ta I with both 5���6 long; ta II 5���6, 4��� 6 long). Leg setation as in deutonymph except: seta 2 c absent; seta 4 b absent; tr I��� III without seta v ���; ge I���II without seta l������; ta IV without setae tc ���. Setae l' added to tr III. LARVA (n = 3). Dorsum. (Fig. 24 a) Body measurements: distance between setae v 2 -h 1 135���180, sc 2 -sc 2 74��� 79; other measurements: v 2 -v 2 13���15, sc 1 -sc 1 56���60, c 1 -c 1 8���10, c 2 -c 2 72���80, c 3 -c 3 104���108, d 1 -d 1 9���10, d 2 -d 2 49���51, d 3 -d 3 70���71, e 1 - e 1 7���8, e 2 - e 2 62���65, e 3 - e 3 49���65, f 3 -f 3 33���41, h 1 -h 1 6���9, h 2 -h 2 18���28. Dorsal shields not developed; dorsal cuticle with few weak transverse folds. Lateral setae lanceolate, obviously barbed; central setae short, weakly barbed. Setal lengths: v 2 22���25, sc 1 14���16, sc 2 19���21, c 1 4���6, c 2 12���14, c 3 13���14, d 1 4���7, d 2 14���16, d 3 15���19, e 1 5, e 2 13 ���19, e 3 15 ���18, f 3 16���19, h 1 5���6, h 2 15���20. Palps. (Fig. 24 a) Palps same as deutonymph. Tibial seta 5���6 long; tarsal eupathidia 3, 4 long; solenidion 2���3 long. Venter. All setae fine. Setal lengths: 1a 23 ��� 33, 1 b 11���19, 3 a 20���26, ps 1 4���7, ps 2 4���6. Legs. (Fig. 24 b) Setal formula for legs I���III (coxae to tarsi) 1 - 0-3 - 0-4 - 7 (1), 0- 0-3 - 0-4 - 7 (1), 0- 0-2 - 0-3 - 3. Tarsi I and II each with 1 antiaxial solenidion ��" (2���3 long) and 2 eupathidia p��'-p��" (5 long). Leg setation as in protonymph except: seta 2 b absent; seta 3 b absent; tr I���III nude; ta I���III without seta tc ���. Remarks. This species is very similar to Cr. muellerianae but can be readily distinguished by lacking setae tc'' on tarsi I���IV (present in Cr. muellerianae), the stouter palpal segments (finer in Cr. muellerianae), lanceolate setae h 1 (more palmate in Cr. muellerianae), and irregular dorsal reticulation (linear in Cr. muellerianae)., Published as part of Beard, Jennifer J., Seeman, Owen D. & Bauchan, Gary R., 2014, Tenuipalpidae (Acari: Trombidiformes) from Casuarinaceae (Fagales), pp. 1-157 in Zootaxa 3778 (1) on pages 29-36, DOI: 10.11646/zootaxa.3778.1.1, http://zenodo.org/record/251337, {"references":["Smiley, R. L., Frost, W. E. & Gerson, U. (1996) A revision of the subfamily Tegopalpinae, with the description of two new genera and five new species (Acari: Tenuipalpidae). International Journal of Acarology, 22 (3), 167 - 180. http: // dx. doi. org / 10.1080 / 01647959608684092"]}

Crossipalpus muellerianae Smiley, Frost and Gerson, 1996 (Figs 13���18) Crossipalpus muellerianae Smiley, Frost and Gerson, 1996: 174, figs 19���20. Type material examined. Holotype female ex. Slaty She-Oak Allocasuarina muelleriana (Casuarinaceae) [feeding on bark of branchlets], AUSTRALIA: South Australia, Flinders Ranges, St Mary���s Peak, at 750 m, 3 October 1995, coll. W. Frost (WINC; on same slide as 7 paratype specimens). Paratypes. 2 females, 1 pharate female, 1 deutonymph, 3 protonymphs, 1 pharate larva, and 1 larva, same data as holotype (WINC; 2 slides, including holotype slide); deutonymph, same data as holotype, on slide with 2 females of Me. delfinadae (WINC). Non-type material examined. 9 females, 3 pharate females, 3 deutonymphs, 1 pharate protonymph, and 1 larva ex. A. muelleriana, AUSTRALIA: South Australia, Adelaide, Athelston, Maryvale Road, Wadmore Park, 34 �� 52 ��� 49 ��� S 138 �� 42 ��� 10 ��� E, 21 April 2009, coll. J.J. Beard and C.J. Burwell (QM, USNM) (BRI voucher BRI [AQ 818802]). Diagnosis. Dorsal setae all thinly lanceolate and weakly barbed. Prodorsal setae v 2 obviously longer than setae sc 1. Palpal segments elongated, thin. Genua I���II with seta l ������ without d; tarsi I���IV with seta tc������. FEMALE (n = 12). Dorsum. (Fig. 13 a) Body measurements: distance between setae v 2 -h 1 335���370 [345]; sc 2 - sc 2 93���105 [105]; other measurements: v 2 -v 2 21���36 [29], sc 1 -sc 1 79���89 [88], c 1 -c 1 14���24 [24], c 2 -c 2 81���96 [93], c 3 -c 3 115���130 [130], d 1 -d 1 19���27 [24], d 2 -d 2 69���87 [85], d 3 -d 3 99���115 [105], e 1 - e 1 13���18 [14], e 2 - e 2 93���105 [97], e 3 - e 3 82���94 [85], f 3 -f 3 63���76 [72], h 1 -h 1 12���17 [12], h 2 -h 2 41���52 [48]. Gnathosoma completely concealed beneath prodorsum. Anterior margin of prodorsum rounded; setae v 2 often inserted ventrally on anterior margin of prodorsum (Fig. 14). Prodorsal shield weakly developed with longitudinal to oblique folds and lines. Some transverse folds between dorsal shields, just anterior to c -setae. Opisthosomal shield weakly developed, mostly with oblique or longitudinal lines and weak folds; short series transverse folds across width of shield just anterior e 1 -e 1; small areas of smooth cuticle between setae c 1 -c 1 and d 1 -d 1 and between d 1 -d 2. Lateral cuticle surrounding shields mostly smooth, with few coarse striae and weak folds. Most dorsal setae thin and barbed; setae v 2, sc 2, e 3, f 2, f 3, and h 2 longer and thicker than other dorsal setae; setae h 1 short, palmate to lanceolate. Setal lengths: v 2 21��� 26 [23, 25], sc 1 8���13 [12], sc 2 11���20 [18, 19], c 1 6���8 [7, 8], c 2 6���11 [9], c 3 10���16 [13, 14], d 1 4���7 [5, 6], d 2 6���9 [7, 9], d 3 10���15 [12, 13], e 1 4 ���8 [6, 7], e 2 10 ���15 [14, 15], e 3 17 ���22 [20], f 3 20���24 [22, 23], h 1 5���10 [7, 8], h 2 20��� 24 [22, 23]. Palps. ( Figs 13 b, 14) Setal formula 0, 0, 0, 1, 3 (1 s+ 2 e). Tibial seta 9���12 [10] long; tarsal eupathidia 4��� 7 [5] long (thin, tapered; could be seta), 5���8 [6, 7] long (thick, blunt); solenidion 4���6 [5, 6] long. Venter. (Figs 14, 15 a) Ventral cuticle of anterior prodorsal margin with coarse striae forming triangular pattern (Fig. 14); cuticle between setae 1 a - 3 a with broadly separated transverse striae; fine longitudinal striae between setae 3 a to just posterior setae 4 a; narrow band of transverse striae ca. 10 wide posterior 4 a, then fine longitudinal striae to ag; oblique striae between ag -ag forming inverted ���v��� shape; coarse striae laterad genital region, longitudinal to oblique. Genital setae widely separated, inserted on posterior corners of genital flap in more-or-less transverse row, g 1 inserted slightly posterior to level of g 2. Genital shield weakly developed, cuticle on genital flap between g setae with mixed oblique striae (g 1 -g 1 33���36) anal setae ps 1���2 inserted in longitudinal line along inner edge of anal plates; seta ps 3 absent (Fig. 15 a). Coxal, genital and anal setae fine, except 2 c thicker, weakly barbed; setae 1 a, 3 a, 4 a long, finely tapered, often broken and difficult to determine full length. Setal lengths: 1a 48 ��� 82 [48, 57], 1 b 19���32 [22, 23], 2 b 17���29 [21, 26], 2 c 18���24 [18, 19], 3a 38 ��� 81 [38], 3 b 17���25 [22], 4a 40 ��� 78 [54], 4 b 17���28 [25, 28], ag 1 10���16 [11], g 1 19���26 [24], g 2 18���25 [23], ps 1 9���14 [14], ps 2 9���14 [13]. Spermatheca. (Fig. 15 b) Spermathecal tube long, distinctly broad near external opening, becoming narrow and convoluted distally, maximum 2 wide, ca. 110 long. Spermatheca vesicle 10���13 long, 2���3 wide proximally, 3���5 wide distally. Genital opening between setae ps 2 and posterior margin of genital flap. Legs. (Fig. 16) Setal formula for legs I���IV (coxae to tarsi) 1 - 1-3 - 1-4 - 9 (1), 2 - 1-3 - 1-4 - 9 (1), 1-2 - 2 - 0-3 - 5, 1 - 1 - 1 - 0-3 - 5. Tarsi I and II each with 1 antiaxial solenidion ��" (ta I 5���7 [6] long, ta II 4���6 [6] long) and 2 eupathidia p��'-p��" (6���7 [6] long). Leg setation as in Table 1 except: coxae I without 1 c; genua I���II with l", without d; tarsi I���IV with tc". Setae v' added to tr IV. MALE. Unknown. DEUTONYMPH (n = 3). Dorsum. (Fig. 17) Body measurements: distance between setae v 2 -h 1 285���305, sc 2 - sc 2 83���91; other measurements: v 2 -v 2 18���21, sc 1 -sc 1 66���75, c 1 -c 1 14���16, c 2 -c 2 72���89, c 3 -c 3 107���109, d 1 -d 1 16���18, d 2 -d 2 64���78, d 3 -d 3 86���99, e 1 - e 1 8���11, e 2 - e 2 72���92, e 3 - e 3 71���78, f 3 -f 3 59���62, h 1 -h 1 12���15, h 2 -h 2 42���46. Prodorsal shield similar to female, but with fewer folds, with central longitudinal folds and sublateral oblique folds. Opisthosoma with widely separated transverse folds between c 1 and e 1, and distinct oblique folds between c 2 -c 3; paired minute pores present between c 1 -c 2, d 1 -d 2 and laterad e 1. Setal lengths: v 2 29���35, sc 1 6���10, sc 2 13���20, c 1 3���7, c 2 4���7, c 3 8���12, d 1 3���5, d 2 4���7, d 3 9���11, e 1 3 ���6, e 2 9 ���15, e 3 20 ���24, f 3 22���26, h 1 4���6, h 2 22���25. Palps. Palps same as adult. Tibial seta 8���9 long; tarsal eupathidia 3���5 long (thin, blunt), 5���6 long (thick, blunt); solenidion 3���5 long. Venter. Cuticle with striae similar to adult female. Coxal, genital and anal setae fine. Setal lengths: 1a 35 ��� 49, 1 b 16���19, 2 b 12���18, 2 c 17���20, 3 a 34���40, 3 b 13���20, 4 a 35���41, 4 b 14���17, ag 1 8���11, g 1 10���17, ps 1 7���8, ps 2 7���10. Legs. Setal formula for legs I���IV (coxae to tarsi) same as adult female except tr IV nude: 1 - 1-3 - 1-4 - 9 (1), 2 - 1-3 - 1-4 - 9 (1), 1-2 - 2 - 0-3 - 5, 1 - 0-1 - 0-3 - 5. Tarsi I and II each with 1 bulbous antiaxial solenidion ��" (3-4 long) and 2 eupathidia p��'-p��" (5���6 long). Leg setation as in adult except: tr IV without seta v ���. Setae v' added to tr I���III. PROTONYMPH (n = 3). Dorsum. Body measurements: distance between setae v 2 -h 1 195���200, sc 2 -sc 2 73��� 75; other measurements: v 2 -v 2 18���19, sc 1 -sc 1 57���60, c 1 -c 1 11���13, c 2 -c 2 62���64, c 3 -c 3 93���100, d 1 -d 1 12���14, d 2 - d 2 56���58, d 3 -d 3 74���77, e 1 - e 1 6���8, e 2 - e 2 65���68, e 3 - e 3 51���55, f 3 -f 3 41���43, h 1 -h 1 9���11, h 2 -h 2 23���25. Prodorsal shield similar to female, but with fewer and much weaker folds. Setal lengths: v 2 24���25, sc 1 6���7, sc 2 11���14, c 1 11���13, c 2 5���7, c 3 9���11, d 1 4���5, d 2 5���6, d 3 7���10, e 1 2 ���4, e 2 10 ���12, e 3 14 ���17, f 3 16���18, h 1 4���5, h 2 18���21. Palps. Palps same as adult. Tibial seta 8 long; tarsal eupathidia both blunt 4, 5; solenidion 3���4 long. Venter. Cuticle with striae similar to adult. Coxal, genital and anal setae fine. Setal lengths: 1a 28 ��� 39, 1 b 16���18, 2 b 15���16, 3 a 17���26, 3 b 11���14, ag 1 8, ps 1 6���7, ps 2 7���8. Legs. Setal formula for legs I���IV (coxae to tarsi) 1 - 0-3 - 0-4 - 9 (1), 1 - 0-3 - 0-4 - 9 (1), 1 - 1-2 - 0-3 - 5, 0- 0-1 - 0-3 - 3. Tarsi I and II each with 1 antiaxial solenidion ��" (ta I 3���4 long, ta II 3 long) and 2 eupathidia p��'-p��" (5 long). Leg setation as in deutonymph except: seta 2 c absent; seta 4 b absent; tr I���II without seta v ���; ge I���II without seta l������; ta IV without setae tc ���, tc������. Setae l' added to tr III. LARVA (n = 2). Dorsum. (Fig. 18) Body measurements: distance between setae v 2 -h 1 140���160, sc 2 -sc 2 61��� 71; other measurements: v 2 -v 2 11���15, sc 1 -sc 1 47���55, c 1 -c 1 10���11, c 2 -c 2 55���58, c 3 -c 3 82���90, d 1 -d 1 13���15, d 2 -d 2 43���49, d 3 -d 3 59���61, e 1 - e 1 7���9, e 2 - e 2 49���57, e 3 - e 3 36���50, f 3 -f 3 21���28, h 1 -h 1 5���9, h 2 -h 2 10���15. Prodorsum with central area smooth to finely striate; lateral cuticle with coarse longitudinal to oblique striae. Opisthosomal cuticle with mostly transverse striae, with some longitudinal striae posteriorly. Setal lengths: v 2 15���24, sc 1 5���7, sc 2 9���13, c 1 4���5, c 2 4���6, c 3 7���8, d 1 3���5, d 2 4���8, d 3 6���7, e 1 3 ���4, e 2 6 ���8, e 3 10 ���13, f 3 11���13, h 1 4���5, h 2 15���17. Palps. (Fig. 18) Palps same as adult. Tibial seta length 6���8; tarsal eupathidia 3���4 long (blunt), 4���5 long (blunt); solenidion 2���3 long. Venter. Striae transverse to anal area, becoming longitudinal and coarse around anal area. All setae fine. Setal lengths: 1a 21 ��� 37, 1 b 10���18, 3 a 22���33, ps 1 6���7, ps 2 5���8. Legs. (Fig. 18) Setal formula for legs I���III (coxae to tarsi) 1 - 0-3 - 0-4 - 7 (1), 0- 0-3 - 0-4 - 7 (1), 0- 0-2 - 0-3 - 3. Tarsi I and II each with 1 bulbous antiaxial solenidion ��" (2���3 long) and 2 eupathidia p��'-p��" (4���5 long). Leg setation as in protonymph except: seta 2 b absent; seta 3 b absent; tr I���III nude; ta I���III without seta tc ���, tc������. Remarks. Smiley et al. (1996) remarked that this species was unique with the adult female having tarsi I���IV 6 (1)- 6 (1)- 6 - 5. This is incorrect and the true count is 9 (1)- 9 (1)- 5 - 5. Individuals were found wedged within pits and crevices on the bark of stems of the host (Fig. 17). This species is very similar to Cr. verticillatae but can be readily distinguished from it by the presence of setae tc'' on tarsi I���IV (absent in Cr. verticillatae), the finer palpal segments (stout in Cr. verticillatae), palmate setae h 1 (lanceolate setae h 1 in Cr. verticillatae), and irregular dorsal reticulation (linear in Cr. verticillatae)., Published as part of Beard, Jennifer J., Seeman, Owen D. & Bauchan, Gary R., 2014, Tenuipalpidae (Acari: Trombidiformes) from Casuarinaceae (Fagales), pp. 1-157 in Zootaxa 3778 (1) on pages 22-29, DOI: 10.11646/zootaxa.3778.1.1, http://zenodo.org/record/251337, {"references":["Smiley, R. L., Frost, W. E. & Gerson, U. (1996) A revision of the subfamily Tegopalpinae, with the description of two new genera and five new species (Acari: Tenuipalpidae). International Journal of Acarology, 22 (3), 167 - 180. http: // dx. doi. org / 10.1080 / 01647959608684092"]}