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Recent studies have documented benefits of small, prescribed fire and wildfire for grassland-dependent wildlife, such as lesser prairie-chickens (

Grazing management recommendations often sacrifice the intrinsic heterogeneity of grasslands by prescribing uniform grazing distributions through smaller pastures, increased stocking densities, and reduced grazing periods. The lack of patch-burn grazing in semi-arid landscapes of the western Great Plains in North America requires alternative grazing management strategies to create and maintain heterogeneity of habitat structure (e.g., animal unit distribution, pasture configuration), but knowledge of their effects on grassland fauna is limited. The lesser prairie-chicken (Tympanuchus pallidicinctus), an imperiled, grassland-obligate, native to the southern Great Plains, is an excellent candidate for investigating effects of heterogeneity-based grazing management strategies because it requires diverse microhabitats among life-history stages in a semi-arid landscape. We evaluated influences of heterogeneity-based grazing management strategies on vegetation structure, habitat selection, and nest and adult survival of lesser prairie-chickens in western Kansas, USA. We captured and monitored 116 female lesser prairie-chickens marked with very high frequency (VHF) or global positioning system (GPS) transmitters and collected landscape-scale vegetation and grazing data during 2013-2015. Vegetation structure heterogeneity increased at stocking densities = 40%. Probability of use was positively affected by increasing pasture area. A quadratic relationship existed between growing season deferment and probability of use. We found that 70% of nests were located in grazing units in which grazing pressure was

Development of systematic methods for conservation planning has improved effectiveness and efficiency of implementing such plans. The lesser prairie-chicken (Tympanuchus pallidicinctus) is a grouse species of conservation concern native to the southwestern Great Plains of the United States. Recent lesser prairie-chicken conservation planning has involved identifying ecologically important areas but has not incorporated economic data into prioritization of areas to target for conservation management. We used the program Marxan to develop a decision-support tool for managers in Kansas to prioritize tracts for improving lesser prairie-chicken habitat quality and increasing habitat availability. We developed three different conservation scenarios and evaluated the tradeoffs among multiple planning objectives in these scenarios. We incorporated population targets from an existing conservation plan and agricultural economic data to help select land with maximum ecological value and minimum economic productivity to prioritize for lesser prairie-chicken conservation. We compared potential conservation plans and incorporated a post hoc connectivity model to test potential for individuals to travel among habitat patches in these plans during dispersal events. We found that different conservation scenarios led to different solutions, though differences varied by ecoregion. Potential solutions for all scenarios contained habitat patches not currently included in existing conservation plans and had high connectivity potential. These results provide context for spatial prioritization of lesser prairie-chicken habitat management in Kansas. Application of this approach to species of conservation interest could help managers incorporate socioeconomic factors into planning methods and identify important tracts for conservation currently overlooked by existing planning methods.

Projected twenty first century increases in temperature and precipitation intensity in the U.S. Great Plains may alter playa wetland hydroperiods. Our objective was to identify favorable germination conditions for a common moist-soil grass, Barnyardgrass (Echinochloa crusgalli L.), by evaluating emergence and growth response to various environmental conditions specific to the Northern (Nebraska) and Southern (Texas) range of playas. We used a temperature-controlled growth chamber experiment to evaluate emergence and growth response of Barnyardgrass to three main effects: (i) weekly temperatures representing historical and future conditions under a moderate emissions scenario, (ii) dry, moist, and saturated soil moisture conditions, and (iii) various seed bank densities. In Nebraska samples, projected future temperatures reduced emergence percentage by up to 20%, but increased emergence percentage by up to 15% for Texas samples. For Nebraska samples, plants were 9.6 cm taller under field capacity moisture compared to saturated moisture. Texas plant height was driven by temperature, where historical conditions produced plants that were 13 cm shorter than future warm conditions. These effects may be exacerbated in natural settings over time and when inter-specific competition exists; thus, temperature, soil moisture, and seed bank densities may be important considerations when planning for playa management in future climate conditions.

The Rainwater Basin in south-central Nebraska once supported a complex network of ~ 12,000 spatially-isolated playa wetlands, but ~ 90% have been lost since European settlement. Future losses are likely and expected reductions in connectivity could further isolate populations, increasing local extinction rates of many wetland species. However, to what extent future losses will affect wildlife likely depends on the role of lost wetlands in maintaining connectivity. We compared the current Rainwater Basin network to future wetland loss scenarios to assess minimum, mean, and maximum effects of losses on network connectivity for a range of wildlife taxa. We used network models to rank wetlands by their functionality and relative importance in maintaining connectivity. We then removed 10–50% of high-ranked, low-ranked, or random subsets of wetlands and assessed connectivity of the remaining network. A 10% loss of highly-ranked wetlands substantially decreased connectivity for species with dispersal capabilities

Temperate‐breeding, or resident, Canada geese were once extirpated in Kansas, USA, but currently provide abundant viewing and hunting opportunities. Kansas Department of Wildlife, Parks, and Tourism (KDWPT) began reintroducing geese in 1980 with a goal of re‐establishing a breeding population. Successful reintroductions led to translocating flocks to regions with no previous records of nesting geese; however, KDWPT continues to translocate individuals from nuisance flocks in urban areas to rural reservoirs to reduce human conflicts with urban geese. Our goal was to determine the effects of such translocations on survival and recovery of adult, sub‐adult, and juvenile temperate‐breeding Canada geese. We used Brownie dead‐recovery models in Program MARK to compare survival and recovery probabilities between translocated and nontranslocated (normal wild) Kansas‐banded Canada geese for 2012–2017. Model‐estimated annual survival differed between status (normal wild S^ = 0.761, 95% CI 0.734–0.785; translocated S^ = 0.598, 95% CI 0.528–0.665). Recovery probability differed between normal and translocated adults (normal wild f^ = 0.074, 95% CI = 0.069–0.078; translocated f^ = 0.138, 95% CI = 0.120–0.158) and juveniles (normal wild f^ = 0.067, 95% CI = 0.059–0.075; translocated f^ = 0.250, 95% CI = 0.199–0.310). Recovery probability did not differ between status in the sub‐adult age class (normal wild f^ = 0.126, 95% CI = 0.115–0.137; translocated f^ = 0.090, 95% CI = 0.055–0.144). Translocation is a viable management option to successfully reduce survival and increase recovery probability of urban nuisance geese in Kansas.

Large-scale declines of grassland ecosystems in the conterminous United States since European settlement have led to substantial loss and fragmentation of lesser prairie-chicken (Tympanuchus pallidicinctus) habitat and decreased their occupied range and population numbers by ∼85%. Breeding season space use is an important component of lesser prairie-chicken conservation, because it could affect both local carrying capacity and population dynamics. Previous estimates of breeding season space use are largely limited to one of the four currently occupied ecoregions, but potential extrinsic drivers of breeding space use, such as landscape fragmentation, vegetation structure and composition, and density of anthropogenic structures, can show large spatial variation. Moreover, habitat needs vary greatly among the lekking/prelaying, nesting, brood-rearing, and postbreeding stages of the breeding season, but space use by female lesser prairie-chickens during these stages remain relatively unclear. We tested whether home range area and daily displacement (the net distance between the first and last location of each day) of female lesser prairie-chickens varied among ecoregions and breeding stages at four study sites in Kansas and Colorado, U.S.A., representing three of the four currently occupied ecoregions. We equipped females with very-high-frequency (VHF) or Global Positioning System (GPS) transmitters, and estimated home range area with kernel density estimators or biased random bridge models, respectively. Across all ecoregions, breeding season home range area averaged 190.4 ha (±19.1 ha se) for birds with VHF and 283.6 ha (±23.1 ha) for birds with GPS transmitters, whereas daily displacement averaged 374.8 m (±14.3 m). Average home range area and daily displacement of bird with GPS transmitters were greater in the Short-Grass Prairie/ Conservation Reserve Program Mosaic and Sand Sagebrush Prairie Ecoregions compared to sites in the Mixed-Grass Prairie Ecoregion. Home range area and daily displacement were greatest during lekking/prelaying and smallest during the brood-rearing stage, when female movements were restricted by mobility of chicks. Ecoregion- and breeding stage-specific estimates of space use by lesser prairie-chickens will help managers determine the spatial configuration of breeding stage-specific habitat on the landscape. Furthermore, ecoregion- and breeding stage-specific estimates are crucial when estimating the amount of breeding habitat needed for lesser prairie-chicken populations to persist.

The predominant grazing-management practice of the Kansas Flint Hills involves annual prescribed burning in March or April with postfire grazing by yearling beef cattle at a high stocking density from April to August. There has been a dramatic increase in sericea lespedeza (Lespedeza cuneata [Dumont] G. Don) coincident with this temporally focused use of prescribed fire in the Flint Hills region. The species is an aggressive invader and a statewide noxious weed in Kansas. Control has generally been attempted using repeated herbicide applications. This approach has not limited proliferation of sericea lespedeza and resulted in collateral damage to nontarget flora and fauna. Alternative timing of prescribed fire has not been evaluated for its control. Our objectives for this 4-yr experiment were to (1) document the effects of prescribed burning during early April, early August, or early September on vigor of sericea lespedeza, standing forage biomass, and basal cover of native graminoids, forbs, and shrubs and (2) measure responses to fire regimes by grassland bird and butterfly communities. Whole-plant dry mass, basal cover, and seed production of sericea lespedeza were markedly less (P < 0.01) in areas treated with prescribed fire in August or September compared with April. Forage biomass did not differ (P ≥ 0.43) among treatments when measured during July; moreover, frequencies of bare soil, litter, and total basal plant cover were not different (P ≥ 0.29) among treatments. Combined basal covers of C4 grasses, C3 grasses, annual grasses, forbs, and shrubs also did not differ (P ≥ 0.11) between treatments. Densities of grasshopper sparrow (Ammodramus savannarum), dickcissel (Spiza americana), and eastern meadowlark (Sturnella magna) were not negatively affected (P > 0.10) by midsummer or late-summer fires relative to early-spring fires. There were no differences (P > 0.10) in densities of grassland-specialist butterfly species across fire regimes. Under the conditions of our experiment, prescribed burning during summer produced no detrimental effects on forage production, desirable nontarget plant species, grassland birds, or butterfly communities but had strong suppressive effects on sericea lespedeza. Additional research is warranted to investigate how to best incorporate late-summer prescribed fire into common grazing-management practices in the Kansas Flint Hills.

The predominant grazing-management practice of the Kansas Flint Hills involves annual prescribed burning in March or April with postfire grazing by yearling beef cattle at a high stocking density from April to August. There has been a dramatic increase in sericea lespedeza (Lespedeza cuneata [Dumont] G. Don) coincident with this temporally focused use of prescribed fire in the Flint Hills region. The species is an aggressive invader and a statewide noxious weed in Kansas. Control has generally been attempted using repeated herbicide applications. This approach has not limited proliferation of sericea lespedeza and resulted in collateral damage to nontarget flora and fauna. Alternative timing of prescribed fire has not been evaluated for its control. Our objectives for this 4-yr experiment were to (1) document the effects of prescribed burning during early April, early August, or early September on vigor of sericea lespedeza, standing forage biomass, and basal cover of native graminoids, forbs, and shrubs and (2) measure responses to fire regimes by grassland bird and butterfly communities. Whole-plant dry mass, basal cover, and seed production of sericea lespedeza were markedly less (P < 0.01) in areas treated with prescribed fire in August or September compared with April. Forage biomass did not differ (P ≥ 0.43) among treatments when measured during July; moreover, frequencies of bare soil, litter, and total basal plant cover were not different (P ≥ 0.29) among treatments. Combined basal covers of C4 grasses, C3 grasses, annual grasses, forbs, and shrubs also did not differ (P ≥ 0.11) between treatments. Densities of grasshopper sparrow (Ammodramus savannarum), dickcissel (Spiza americana), and eastern meadowlark (Sturnella magna) were not negatively affected (P > 0.10) by midsummer or late-summer fires relative to early-spring fires. There were no differences (P > 0.10) in densities of grassland-specialist butterfly species across fire regimes. Under the conditions of our experiment, prescribed burning during summer produced no detrimental effects on forage production, desirable nontarget plant species, grassland birds, or butterfly communities but had strong suppressive effects on sericea lespedeza. Additional research is warranted to investigate how to best incorporate late-summer prescribed fire into common grazing-management practices in the Kansas Flint Hills.

Globally, migration phenologies of numerous avian species have shifted over the past half-century. Despite North American waterfowl being well researched, published data on shifts in waterfowl migration phenologies remain scarce. Understanding shifts in waterfowl migration phenologies along with potential drivers is critical for guiding future conservation efforts. Therefore, we utilized historical (1955–2008) nonbreeding waterfowl survey data collected at 21 National Wildlife Refuges in the mid- to lower portion of the Central Flyway to summarize changes in spring and autumn migration phenology. We examined changes in the timing of peak abundance from survey data at monthly intervals for each refuge and species (or species group;n= 22) by year and site-specific temperature for spring (Jan–Mar) and autumn (Oct–Dec) migration periods. For spring (n= 187) and autumn (n= 194) data sets, 13% and 9% exhibited statistically significant changes in the timing of peak migration across years, respectively, while the corresponding numbers for increasing temperatures were 4% and 9%. During spring migration, ≥80% of significant changes in the timing of spring peak indicated advancements, while 67% of significant changes in autumn peak timing indicated delays both across years and with increasing temperatures. Four refuges showed a consistent pattern across species of advancing spring migration peaks over time. Advancements in spring peak across years became proportionally less common among species with increasing latitude, while delays in autumn peak with increasing temperature became proportionally more common. Our study represents the first comprehensive summary of changes in spring and autumn migration phenology for Central Flyway waterfowl and demonstrates significant phenological changes during the latter part of the twentieth century.

Spatially distributed populations often rely on large‐scale processes for long‐term population stability. These processes are driven by individuals moving across the landscape through long‐distance dispersal movements. However, as landscapes are continually altered by anthropogenic development, increased fragmentation and avoidance behavior can affect landscape permeability and limit dispersal. Lesser prairie‐chickens (Tympanuchus pallidicinctus) are a species of concern that have lost significant portions (>90%) of their historic distribution in the Southern Great Plains of the United States and are currently being impacted by continued anthropogenic development. Using GPS telemetry locations of 346 lesser prairie‐chickens across their entire geographic distribution, we identified 184 different long‐distance movements that drive population connectivity. We used empirical cumulative distribution functions to create a selection–avoidance–neutral curve and estimated the spatial scale of response to anthropogenic features (i.e., towers and windmills, large transmission and smaller distribution powerlines, oil wells, roads, and fences) during these movements. In addition, we tested for behavioral differences between movement types (e.g., exploratory loops vs. long‐distance movements between home ranges) and for regional differences in response among study areas. We found that during long‐distance movements, lesser prairie‐chickens generally avoided all anthropogenic feature types we tested despite some variation in the reported response distance among study areas. However, they avoided the tallest features (i.e., towers and windmills and transmission powerlines) at much greater distances in comparison with the shorter features in our analysis. Our results show that long‐distance movements are likely affected by responses to functional landscape fragmentation through increased development of anthropogenic features in important connectivity zones. As our estimated response distances during long‐distance movements varied in comparison with previously reported response distances during other behavioral states (e.g., breeding or nesting), using long‐distance or dispersal specific movement data may be more appropriate when asking questions related to connectivity across the landscape.

Loss and degradation of grasslands in the Great Plains region have resulted in major declines in abundance of grassland bird species. To ensure future viability of grassland bird populations, it is crucial to evaluate specific effects of environmental factors among species to determine drivers of population decline and develop effective conservation strategies. We used threshold models to quantify the effects of land cover and weather changes in "lesser prairie‐chicken" and "greater prairie‐chicken" (Tympanuchus pallidicinctus and T. cupido, respectively), northern bobwhites (Colinus virginianus), and ring‐necked pheasants (Phasianus colchicus). We demonstrated a novel approach for estimating landscape conditions needed to optimize abundance across multiple species at a variety of spatial scales. Abundance of all four species was highest following wet summers and dry winters. Prairie chicken and ring‐necked pheasant abundance was highest following cool winters, while northern bobwhite abundance was highest following warm winters. Greater prairie chicken and northern bobwhite abundance was also highest following cooler summers. Optimal abundance of each species occurred in landscapes that represented a grassland and cropland mosaic, though prairie chicken abundance was optimized in landscapes with more grassland and less edge habitat than northern bobwhites and ring‐necked pheasants. Because these effects differed among species, managing for an optimal landscape for multiple species may not be the optimal scenario for any one species., We quantified the effects of land cover and weather changes in Galliformes in Kansas. We found evidence of nonlinear, threshold effects of land cover change in abundance of all four focal species, suggesting that the presence of both grassland and cropland is important in the landscape to simultaneously maintain these populations.

Agricultural landscapes are the leading edge in the advancement of sustainability and climate change adaptation. The purpose of this study is to endogenize culture as shaped by natural-cultural feedback into individuals’ decision-making processes on sustainability policy support. We present an agent-based model in which an adaptive cultural decision-rule quantifies the probability of an agent deciding to support a wildlife area policy for the Smoky Hill River Watershed (SHRW) in Kansas, USA. By using an ABM to examine the watershed as a coupled natural and human system, we learned that agents would adopt a new behavior, voting for the policy, if the cultural conditions were right, with high levels of beliefs and norms for freshwater and its biota. Our results indicate that individuals in the SHRW are not engaged in caring for fish, plants, and bird richness in their rivers and playas with few individuals supporting the policy in the naive cultural setting (8.9 % of simulated population). However, enough agents would support the policy under a lower cultural threshold (40.7 % of simulated population). Our results show that sustainability policies need to account for the local culture to gain support, and if a policy is culturally meaningful, it does not need to be cheap. For an agricultural landscape, such as those commonly found in the Central Great Plains, this study presents new levers for policymakers on the conditions needed to help assemble popular support for sustainability policies.

The Rainwater Basin region in south-central Nebraska supports a complex network of spatially-isolated wetlands that harbor diverse floral and faunal communities. Since European settlement, many wetlands have been lost from the network, which has increased distances among remaining wetlands. As a result, populations of wildlife species with limited dispersal capabilities may have become isolated and face greater local extinction risks. We compared the pre-European settlement and current extent of the Rainwater Basin network to assess the effects of wetland losses on network connectivity for a range of maximum dispersal distances. We constructed network models for a range of maximum dispersal distances and calculated network metrics to assess changes in network connectivity and the relative importance of individual wetlands in regulating flow. Since European settlement, the number of wetlands in the Rainwater Basin has decreased by > 90%. The average distance to the nearest neighboring wetland has increased by 150% to ~ 1.2 km, and the dispersal distance necessary to travel throughout the whole network has increased from 3.5 to 10.0 km. Last, relative importance of individual wetlands depended on the maximum dispersal distance. Which wetlands to preserve to maintain connectivity might therefore depend on the dispersal capabilities of the species or taxa of interest. To preserve a broad range of biodiversity, conservation efforts should focus on preserving dense clusters of wetlands at fine spatial scales to maintain current levels of network connectivity, and restoring connections between clusters to facilitate long-range dispersal of species with limited dispersal capabilities.

The lesser prairie-chicken Tympanuchus pallidicinctus has experienced significant declines in distribution and abundance since the early 1900s. A severe and prolonged drought from 2009 to 2013 resulted in further declines in population numbers and despite improved environmental and habitat conditions since 2013, populations of lesser prairie chickens have shown little improvement. To investigate whether breeding season survival of lesser prairie-chickens in eastern New Mexico could be driving this response, we developed the following objectives: 1) estimate male and female breeding-season survival; 2) determine whether male and female survival varies temporally among lekking, nesting, and brood-rearing periods; and 3) determine cause-specific mortality during the breeding season. We captured and radiocollared 76 lesser prairie-chickens (50 male, 26 female) during spring of 2014 and 2015 and estimated their survival throughout the breeding season (15 March–31 August). Male survival was nearly double that of females in both years (0.79–0.81 and 0.38–0.45, respectively). Males had similar survival across all periods (lekking, postlekking, late summer: 0.89–0.95). Females had the greatest period-specific survival during lekking and brood rearing (0.87 ± 0.08 and 0.85 ± 0.10, respectively) relative to the nesting period (0.58 ± 0.11). Mammalian predation was the primary cause of mortality in both years. Our results indicate that in New Mexico 1) lesser prairie-chicken breeding season survival was consistent with geographically similar studies, 2) females have lower survival during the nesting period, and 3) female lesser prairie-chicken survival was lower than male survival regardless of time period. Management actions that provide and protect high-quality nesting habitat may help ensure that female survival is maximized during the nesting period.

Conservation of small populations is often based on limited data from spatially and temporally restricted studies, resulting in management actions based on an incomplete assessment of the population drivers. If fluctuations in abundance are related to changes in weather, proper management is especially important, because extreme weather events could disproportionately affect population abundance. Conservation assessments, especially for vulnerable populations, are aided by a knowledge of how extreme events influence population status and trends. Although important for conservation efforts, data may be limited for small or vulnerable populations. Integrated population models maximize information from various sources of data to yield population estimates that fully incorporate uncertainty from multiple data sources while allowing for the explicit incorporation of environmental covariates of interest. Our goal was to assess the relative influence of population drivers for the Lesser Prairie-Chicken ...

Within the U.S. portion of the Central Flyway, the U.S. Fish and Wildlife Service manages waterfowl on numerous individual units (i.e., Refuges) within the National Wildlife Refuge System. Presently, the extent of waterfowl use that Refuges receive and the contribution of Refuges to waterfowl populations (i.e., the proportion of the Central Flyway population registered at each Refuge) remain unassessed. Such an evaluation would help determine to what extent Refuges support waterfowl relative to stated targets, aid in identifying species requiring management attention, inform management targets, and improve fiscal efficiencies. Using historic monitoring data (1954–2008), we performed this assessment for 23 Refuges in Texas, New Mexico, Oklahoma, Kansas, and Nebraska during migration and wintering months (October–March). We examined six dabbling ducks and two diving ducks, plus all dabbling ducks and all diving ducks across two periods (long-term [all data] and short-term [last 10 October–March periods]). Individual Refuge use was represented by the sum of monthly duck count averages for October–March. We used two indices of Refuge contribution: peak contribution and January contribution. Peak contribution was the highest monthly count average for each October–March period divided by the indexed population total for the Central Flyway in the corresponding year; January contribution used the January count average divided by the corresponding population index. Generally, Refuges in Kansas, Nebraska, and New Mexico recorded most use and contribution for mallards Anas platyrhynchos. Refuges along the Texas Gulf Coast recorded most use and contribution for other dabbling ducks, with Laguna Atascosa and Aransas (including Matagorda Island) recording most use for diving ducks. The long-term total January contribution of the assessed Refuges to ducks wintering in the Central Flyway was greatest for green-winged teal Anas crecca with 35%; 12–15% for American wigeon Mareca americana, gadwall Mareca strepera, and northern pintail Anas acuta; and 7–8% for mallard and mottled duck Anas fulvigula. Results indicated that the reliance on the National Wildlife Refuge System decreased for these ducks, with evidence suggesting that, for several species, the assessed Refuges may be operating at carrying capacity. Future analyses could be more detailed and informative were Refuges to implement a single consistent survey methodology that incorporated estimations of detection bias in the survey process, while concomitantly recording habitat metrics on and neighboring each Refuge.

Coastal wetlands along the Gulf of Mexico support a wide diversity of wildlife, are important nurseries for sport and commercial fisheries, provide erosion and flood control, and serve many other ecological functions and services. These marshes have been declining in area and degrading at alarming rates since the 1930s. Effective conservation planning is vital to protect these ecosystems, but decision makers often lack knowledge of expected future conditions to strategically target conservation actions. To address this issue, we focus on a species of conservation concern, the mottled duck (Anas fulvigula), that resides year-round in the coastal marshes of the Gulf of Mexico. We used location data collected from radiomarked hen mottled ducks from 2006 to 2011 to create an ensemble model of habitat selection for 2010. We then projected future habitat states using models of sea-level rise and human development. By combining future predictions with our ensemble model, we predict future habitat for mottled ducks through 2100, in 20-year time steps beginning with 2020. Sea-level rise models predicted reductions in coastal marsh habitats and our ensemble model predict corresponding declines in overall habitat quantity and quality for mottled ducks, with the largest rate of habitat loss predicted within the Chenier Plain of Louisiana, USA at 71%. In some areas, particularly the Texas Mid-Coast, USA, future urbanization and human development is expected to reduce the ability of wetland habitat to migrate inland with rising sea-levels. Our results also highlight areas of coastal marsh particularly vulnerable to sea-level rise; and conversely, identify areas most likely to persist into the future that could be targeted for habitat conservation to help mottled ducks persist on the landscape.

The lesser prairie-chicken (Tympanuchus pallidicinctus), a species of conservation concern with uncertain regulatory status, has experienced population declines over the past century. Most research on lesser prairie-chickens has focused on the breeding season, with little research conducted during the nonbreeding season, a period that exerts a strong influence on demography in other upland game birds. We trapped lesser prairie-chickens on leks and marked them with either global positioning system (GPS) satellite or very high frequency (VHF) transmitters to estimate survival and home-range size during the nonbreeding season. We monitored 119 marked lesser prairie-chickens in 3 study areas in Kansas, USA, from 16 September to 14 March in 2013, 2014, and 2015. We estimated home-range size using Brownian Bridge movement models (GPS transmitters) and fixed kernel density estimators (VHF transmitters), and female survival using Kaplan–Meier known-fate models. Average home-range size did not differ between sexes. Estimated home-range size was 3 times greater for individuals fitted with GPS satellite transmitters (x¯ = 997 ha) than those with VHF transmitters (x¯ = 286 ha), likely a result of the temporal resolution of the different transmitters. Home-range size of GPS-marked birds increased 2.8 times relative to the breeding season and varied by study area and year. Home-range size was smaller in the 2013–2014 nonbreeding season (x¯ = 495 ha) than the following 2 nonbreeding seasons (x¯ = 1,290 ha and x¯ = 1,158 ha), corresponding with drought conditions of 2013, which were alleviated in following years. Female survival (Sˆ) was high relative to breeding season estimates, and did not differ by study area or year (Sˆ = 0.73 ± 0.04 [SE]). Future management could remain focused on the breeding season because nonbreeding survival was 39–44% greater than the previous breeding season; however, considerations of total space needs would benefit lesser prairie-chickens by accounting for the greater spatial requirements during the nonbreeding season. © 2017 The Wildlife Society.

Understanding the relative influence of environmental and intrinsic effects on populations is important for managing and conserving harvested species, especially those species inhabiting changing environments. Additionally, climate change can increase the uncertainty associated with management of species in these changing environments, making understanding factors affecting their populations even more important. Coastal ecosystems are particularly threatened by climate change; the combined effects of increasing severe weather events, sea level rise, and drought will likely have non-linear effects on coastal marsh wildlife species and their associated habitats. A species of conservation concern that persists in these coastal areas is the mottled duck (Anas fulvigula). Mottled ducks in the western Gulf Coast are approximately 50% below target abundance numbers established by the Gulf Coast Joint Venture for Texas and Louisiana, USA. Although evidence for declines in mottled duck abundance is apparent, specific causes of the decrease remain unknown. Our goals were to determine where the largest declines in mottled duck population were occurring along the system of Texas Gulf Coast National Wildlife Refuges and quantify the relative contribution of environmental and intrinsic effects on changes to relative population density. We modeled aerial survey data of mottled duck density along the Texas Gulf Coast from 1986–2015 to quantify effects of extreme weather events on an index to mottled duck density using the United States Climate Extremes Index and Palmer Drought Severity Index. Our results indicate that decreases in abundance are best described by an increase in days with extreme 1-day precipitation from June to November (hurricane season) and an increase in drought severity. Better understanding those portions of the life cycle affected by environmental conditions, and how to manage mottled duck habitat in conjunction with these events will likely be key to persistence of the species under future environmental conditions. © 2017 The Wildlife Society.

Speyeria idalia is a prairie specialist that has experienced dramatic population declines throughout its range. Speyeria idalia is nearly extirpated from the eastern portion of its former range; however, populations within Kansas are relatively stable. We made several previously undescribed field observations of late-instar larvae and post-diapause female S. idalia in northeastern Kansas during 2014–2016. We report finding late-instar larvae at locations that were burned within weeks of detection. The observations of larvae shortly following a burn suggests that S. idalia larvae are capable of surviving fire and contradicts our current knowledge of this species. Additionally, we describe a feeding behavior characteristic of late-instar larvae. Larvae observed in the field and lab stripped leaves of host plants leaving only stems. This strip-style feeding behavior provided unique feeding evidence that was valuable to detecting the presence of larvae in the field. Finally, we documented larvae and...

The mottled duck (Anas fulgivula) is a non-migratory duck dependent on coastal habitats to meet all of its life cycle requirements in the Western Gulf Coast (WGC) of Texas and Louisiana, USA. This population of mottled ducks has experienced a moderate decline during the past 2 decades. Adult survival has been identified as an important factor influencing population demography. Previous work based on band-recovery data has provided only annual estimates of survival. We assessed seasonal patterns of female mottled duck survival from 2009 to 2012 using individuals marked with satellite platform transmitter terminals (PTTs). We used temperature and movement sensors within each PTT to indicate potential mortality events. We estimated cumulative weekly survival and ranked factors influential in patterns of mortality using known-fate modeling in Program MARK. Models included 4 predictors: week; hunting and non-hunting periods; biological periods defined as breeding, brooding, molt, and pairing; and mass at time of capture. Models containing hunt periods, during and outside the mottled duck season, comprised essentially 100% of model weights where both legal and illegal harvest had a negative influence on mottled duck survival. Survival rates were low during 2009–2011 (12–38% annual rate of survival), when compared with the long-term banding average of 53% annual survival. During 2011, survival of female mottled ducks was the lowest annual rate (12%) ever documented and coincided with extreme drought. Management actions maximizing the availability of wetlands and associated upland habitats during hunting seasons and drought conditions may increase adult female mottled duck survival. © 2017 The Authors. Journal of Wildlife Management Published by Wiley Periodicals, Inc. on behalf of The Wildlife Society.

Since the beginning of the 20th century, the overall occupied range of the lesser prairie-chicken (Tympanuchus pallidicinctus) has declined by 84% commensurate with population trends. Much of this decline has been attributed to the loss and fragmentation of native grasslands throughout the lesser prairie-chicken range. However, quantification of changes in land cover in the distribution of the lesser prairie-chicken is lacking. Our objectives were to (1) document changes in the areal extent and connectivity of grasslands in the identified lesser prairie-chicken range in Kansas, USA, (>60% of extant lesser prairie-chicken population) from the 1950s to 2013 using remotely sensed data and (2) assess the potential of the Conservation Reserve Program (U.S. Department of Agriculture Program converting cropland to permanent cover; CRP) to mitigate grassland loss. Digital land cover maps were generated on a decadal time step through spectral classification of LANDSAT images and visual analysis of aerial photographs (1950s and 1960s). Landscape composition and configuration were assessed using FRAGSTATS to compute a variety of landscape metrics measuring changes in the amount of grassland present as well as changes in the size and configuration of grassland patches. With the exception of a single regional portion of the range, nearly all of the grassland converted to cropland in the lesser prairie-chicken range of Kansas occurred prior to the 1950s. Prior to the implementation of CRP, the amount of grassland decreased 3.6% between the 1950s and 1985 from 18,455 km2 to 17,788 km2. Since 1985, the overall amount of grassland in the lesser prairie-chicken range has increased 11.9% to 19,898 km2 due to implementation of CRP, although the area of grassland decreased between 1994 and 2013 as CRP contracts were not renewed by landowners. Since 1986 grassland in Kansas became more connected and less fragmented in response to the CRP. While the CRP has been successful in increasing grassland quantity and connectivity throughout the lesser prairie-chicken range in Kansas, offsetting loss of grassland since the 1950s, abundance and occupied range of lesser prairie-chickens has declined since the 1980s, suggesting that habitat quality is the principal factor influencing population demography of the species. Although the CRP is contributing to conservation actions for lesser prairie-chickens, efforts to improve habitat quality throughout the range of the lesser prairie-chicken are likely necessary to meet management goals. Continuation of the CRP faces an uncertain future in the face of rising commodity prices, energy development, and reduction in program scope, leaving open the possibility that these areas that have created habitat for lesser prairie-chickens could be lost.

The >2,570,000‐ha Flint Hills ecoregion of Kansas, USA, harbors the largest remaining contiguous tract of tallgrass prairie in North America, a unique system, as the remainder of North America's tallgrass prairie has succumbed to development and conversion. Consequently, the loss and degradation of tallgrass prairie has reduced populations of many North American prairie‐obligate species including the regal fritillary (Speyeria idalia) butterfly. Population abundance and occupied range of regal fritillary have declined >99%, restricting many populations to isolated, remnant patches of tallgrass prairie. Such extensive decline has resulted in consideration of the regal fritillary for protection under the Endangered Species Act. Although it is widely accepted that management practices such as fire, grazing, and haying are necessary to maintain prairie ecosystems, reported responses by regal fritillary to these management regimes have been ambiguous. We tested effects of prescribed fire across short, moderate, and long fire‐return intervals as well as grazing and haying management treatments on regal fritillary density. We also tested the relative influence of habitat characteristics created by these management regimes by measuring density of an obligate host plant (Viola spp.) and canopy cover of woody vegetation, grasses, forbs/ferns, bare ground, and litter. We found density was at least 1.6 times greater in sites burned with a moderate fire‐return interval vs. sites burned with short and long fire‐return intervals. Overall management regardless of fire‐return interval did not have an effect on density. Percent cover of grass had the strongest positive association, while percent cover of woody vegetation had the greatest negative effect on density. Our results indicate that patch‐burning is a viable and perhaps even ideal management strategy for regal fritillary in tallgrass prairie landscapes. Additionally, these results elucidate the importance of fire, particularly when applied at moderate‐return intervals to regal fritillary, and corroborate a growing suite of studies that suggest fire is perhaps not as detrimental to populations of regal fritillary as previously believed.

The magnitude and distribution of lead contamination remain unknown in wetland systems. Anthropogenic deposition of lead may be contributing to negative population-level effects in waterfowl and other organisms that depend on dynamic wetland habitats, particularly if they are unable to detect and differentiate levels of environmental contamination by lead. Detection of lead and behavioral response to elevated lead levels by waterfowl is poorly understood, but necessary to characterize the risk of lead-contaminated habitats. We measured the relationship between lead contamination of wetland soils and habitat use by mottled ducks (Anas fulvigula) on the Upper Texas Coast, USA. Mottled ducks have historically experienced disproportionate negative effects from lead exposure, and exhibit a unique nonmigratory life history that increases risk of exposure when inhabiting contaminated areas. We used spatial interpolation to estimate lead in wetland soils of the Texas Chenier Plain National Wildlife Refuge Complex. Soil lead levels varied across the refuge complex (0.01–1085.51 ppm), but greater lead concentrations frequently corresponded to areas with high densities of transmittered mottled ducks. We used soil lead concentration data and MaxENT species distribution models to quantify relationships among various habitat factors and locations of mottled ducks. Use of habitats with greater lead concentration increased during years of a major disturbance. Because mottled ducks use habitats with high concentrations of lead during periods of stress, have greater risk of exposure following major disturbance to the coastal marsh system, and no innate mechanism for avoiding the threat of lead exposure, we suggest the potential presence of an ecological trap of quality habitat that warrants further quantification at a population scale for mottled ducks.

Sustainability has been at the forefront of the environmental research agenda of the integrated anthroposphere, hydrosphere, and biosphere since the last century and will continue to be critically important for future environmental science. However, linking humans and the environment through effective policy remains a major challenge for sustainability research and practice. Here we address this gap using an agent-based model (ABM) for a coupled natural and human systems in the Smoky Hill River Watershed (SHRW), Kansas, USA. For this freshwater-dependent agricultural watershed with a highly variable flow regime influenced by human-induced land-use and climate change, we tested the support for an environmental policy designed to conserve and protect fish biodiversity in the SHRW. We develop a proof of concept interdisciplinary ABM that integrates field data on hydrology, ecology (fish richness), social-psychology (value-belief-norm) and economics, to simulate human agents' decisions to support environmental policy. The mechanism to link human behaviors to environmental changes is the social-psychological sequence identified by the value-belief-norm framework and is informed by hydrological and fish ecology models. Our results indicate that (1) cultural factors influence the decision to support the policy; (2) a mechanism modifying social-psychological factors can influence the decision-making process; (3) there is resistance to environmental policy in the SHRW, even under potentially extreme climate conditions; and (4) the best opportunities for policy acceptance were found immediately after extreme environmental events. The modeling approach presented herein explicitly links biophysical and social science has broad generality for sustainability problems.

Use of dogs has increased for multiple wildlife research purposes ranging from carnivore scat detection to estimation of reptile abundance. Use of dogs is not particularly novel for upland gamebird biologists, and pointing dogs have been long considered an important research tool. However, recent advances in Global Positioning System (GPS) technology and the development of hierarchical modeling approaches that account for imperfect detection may improve estimates of occupancy and density of cryptic species such as the American woodcock (Scolopax minor; hereafter, woodcock). We conducted surveys for woodcock using a trained pointing dog wearing a GPS collar during the winters of 2010–2011 and 2011–2012 in East Texas, USA. We surveyed 0.5-km-radius circular plots (n = 24; survey sites) randomly placed along secondary roads in Davy Crockett National Forest and on private timber property. Surveys lasted 1.5 hrs and were repeated 3–5 times each winter. We estimated woodcock occupancy and density using multiple modeling approaches at the survey site and forest stand scales within survey sites. Woodcock occupied 88% (21/24) of survey sites and 48% (39/82) of forest stands (i.e., unique cover types) within sites. Using a modified distance sampling technique, we estimated an average density of 0.16 birds/ha (SE = 0.13) throughout both study areas. We describe the first attempt to blend use of pointing dogs with hierarchical modeling approaches to derive estimates of regional diurnal woodcock occupancy and density, and describe relationships between these estimates of abundance and habitat covariates. Although forest stand occupancy estimates had the lowest coefficients of variation, our estimates of density provided the most useful inference of habitat use. Surveys using pointing dogs paired with hierarchical models of occupancy and density may provide a cost-efficient and effective approach to estimate habitat abundance at broad spatial scales.

Lesser prairie-chicken (Tympanuchus pallidicinctus) populations have declined since the 1980s. Understanding factors influencing nest-site selection and nest survival are important for conservation and management of lesser prairie-chicken populations. However, >75% of the extant population is in the northern extent of the range where data on breeding season ecology are lacking. We tested factors influencing fine-scale and regional nest-site selection and nest survival across the northern portion of the lesser prairie-chicken range. We trapped and affixed satellite global positioning system and very high frequency transmitters to female lesser prairie-chickens (n = 307) in south-central and western Kansas and eastern Colorado, USA. We located and monitored 257 lesser prairie-chicken nests from 2013 to 2016. We evaluated nest-site selection and nest survival in comparison to vegetation composition and structure. Overall, nest-site selection in relation to vegetation characteristics was similar across our study area. Lesser prairie-chickens selected nest microsites with 75% visual obstruction 2.0-3.5 dm tall and 95.7% of all nests were in habitat with >= 1 dm and

This chapter discusses an interdisciplinary and transdisciplinary project to understand the interactions of agriculture, climate, and water resources in the Central Great Plains as a coupled natural-human system. We focus on the Smoky Hills Watershed in Kansas, where we gathered socioeconomic, hydrological, and climatic data, along with ecological data on fish species. The project involved substantial stakeholder engagement, which was complicated by post-truth attitudes about climate science and environmental regulation by some groups. We discuss the challenges of team management, stakeholder engagement, and data integration for modeling, notably the incorporation of stakeholder support for environmental policy in the context of extreme climatic events. We conclude by offering a framework for good collaborative practice to manage the complications of crossing boundaries in transdisciplinary research and outreach.

Playa wetlands, the dominant hydrological feature of the semi-arid U.S. High Plains providing critical ecosystem services, are being lost and degraded due to anthropogenic alterations of the short-grass prairie landscape. The primary process contributing to the loss of playas is filling of the wetland through accumulation of soil eroded and transported by precipitation from surrounding cultivated watersheds. We evaluated effectiveness of vegetative buffers surrounding playas in removing metals, nutrients, and dissolved/suspended sediments from precipitation runoff. Storm water runoff was collected at 10-m intervals in three buffer types (native grass, fallow cropland, and Conservation Reserve Program). Buffer type differed in plant composition, but not in maximum percent removal of contaminants. Within the initial 60 m from a cultivated field, vegetation buffers of all types removed >50% of all measured contaminants, including 83% of total suspended solids (TSS) and 58% of total dissolved solids (TDS). Buffers removed an average of 70% of P and 78% of N to reduce nutrients entering the playa. Mean maximum percent removal for metals ranged from 56% of Na to 87% of Cr. Maximum removal was typically at 50 m of buffer width. Measures of TSS were correlated with all measures of metals and nutrients except for N, which was correlated with TDS. Any buffer type with >80% vegetation cover and 30-60 m in width would maximize contaminant removal from precipitation runoff while ensuring that playas would continue to function hydrologically to provide ecosystem services. Watershed management to minimize erosion and creations of vegetation buffers could be economical and effective conservation tools for playa wetlands.

Sand shinnery oak (Quercus havardii) prairies are increasingly disappearing and increasingly degraded in the Southern High Plains of Texas and New Mexico. Restoring and managing sand shinnery oak prairie can support biodiversity, specific species of conservation concern, and livestock production. We measured vegetation response to four treatment combinations of herbicide (tebuthiuron applied at 0.60 kg/ha) and moderate-intensity grazing (50% removal of annual herbaceous production) over a 10-year period in a sand shinnery oak prairie of eastern New Mexico. We compared the annual vegetation response to the historical climax plant community (HCPC) as outlined by the U.S. Department of Agriculture Ecological Site Description. From 2 to 10 years postapplication, tebuthiuron-treated plots had reduced shrub cover with twice as much forb and grass cover as untreated plots. Tebuthiuron-treated plots, regardless of the presence of grazing, most frequently met HCPC. Tebuthiuron and moderate-intensity grazi...

Playa wetlands are the primary habitat for numerous wetland-dependent species in the Southern Great Plains of North America. Plant and wildlife populations that inhabit these wetlands are reciprocally linked through the dispersal of individuals, propagules and ultimately genes among local populations. To develop and implement a framework using network models for conceptualizing, representing and analyzing potential biological flows among 48,981 spatially discrete playa wetlands in the Southern Great Plains. We examined changes in connectivity patterns and assessed the relative importance of wetlands to maintaining these patterns by targeting wetlands for removal based on network centrality metrics weighted by estimates of habitat quality and probability of inundation. We identified several distinct, broad-scale sub networks and phase transitions among playa wetlands in the Southern Plains. In particular, for organisms that can disperse >2 km a dense and expansive wetland sub network emerges in the Southern High Plains. This network was characterized by localized, densely connected wetland clusters at link distances (h) >2 km but

Long-term population and range declines from habitat loss and fragmentation caused the lesser prairie-chicken (Tympanuchus pallidicinctus) to be a species of concern throughout its range. Current lesser prairie-chicken range in New Mexico and Texas is partially restricted to sand shinnery oak (Quercus havardii; hereafter shinnery oak) prairies, on which cattle grazing is the main socioeconomic driver for private landowners. Cattle producers within shinnery oak prairies often focus land management on shrub eradication using the herbicide tebuthiuron to promote grass production for forage; however, herbicide application alone, and in combination with grazing, may affect nest site selection and nest survival of lesser prairie-chickens through the reduction of shinnery oak and native grasses. We used a controlled, paired, completely randomized design study to assess the influence of grazing and tebuthiuron application and their combined use on nest site selection and nest survival from 2001 to 2010 in Roosevelt County, New Mexico, USA at 2 spatial scales (i.e., treatment and microhabitat) in 4 treatments: tebuthiuron with grazing, tebuthiuron without grazing, no tebuthiuron with grazing, and a control of no tebuthiuron and no grazing. Grazing treatment was a short-duration system in which plots were grazed once during the dormant season and once during the growing season. Stocking rate was calculated each season based on measured forage production and applied to remove ≤25% of available herbaceous material per season. At the treatment scale, we compared nest site selection among treatments using 1-way χ2 tests and nest survival among treatments using a priori candidate nest survival models in Program MARK. At the microhabitat scale, we identified important habitat predictors of nest site selection and nest survival using logistic regression and a priori candidate nest survival models in Program MARK, respectively. Females typically used treatments as expected and we did not detect trends in selection. Nest survival did not differ among treatments. At the microhabitat scale, nest sites had less bare ground (P = 0.001) and greater angles of obstruction (P ≤ 0.001) compared to random sites. There was a high degree of model selection uncertainty among our candidate models at the microhabitat scale and survival estimates were similar among habitat covariates. Results suggest a tebuthiuron application rate of 0.60 kg/ha, short-duration grazing, and a combination of these management techniques were not detrimental to lesser prairie-chicken nest site selection or nest survival. However, intensified management that increases bare ground or reduces overhead cover may negatively affect lesser prairie-chicken nesting habitat and nest survival. © 2016 The Wildlife Society.

To identify factors contributing to the long-term decline of American woodcock, a holistic understanding of range-wide population connectivity throughout the annual cycle is needed. We used band recovery data and isotopic composition of primary (P1) and secondary (S13) feathers to estimate population sources and connectivity among natal, early fall, and winter ranges of hunter-harvested juvenile American woodcock. We used P1 feathers from known-origin pre-fledged woodcock (n = 43) to create a hydrogen δ2Hf isoscape by regressing δ2Hf against expected growing-season precipitation (δ2Hp). Modeled δ2Hp values explained 79% of the variance in P1 δ2Hf values, indicating good model fit for estimating woodcock natal origins. However, a poor relationship (r2 = 0.23) between known-origin, S13 δ2Hf values, and expected δ2Hp values precluded assignment of early fall origins. We applied the δ2Hf isoscape to assign natal origins using P1 feathers from 494 hunter-harvested juvenile woodcock in the United States and Canada during 2010–2011 and 2011–2012 hunting seasons. Overall, 64% of all woodcock origins were assigned to the northernmost (>44°N) portion of both the Central and Eastern Management Regions. In the Eastern Region, assignments were more uniformly distributed along the Atlantic coast, whereas in the Central Region, most woodcock were assigned to origins within and north of the Great Lakes region. We compared our origin assignments to spatial coverage of the annual American woodcock Singing Ground Survey (SGS) and evaluated whether the survey effectively encompasses the entire breeding range. When we removed the inadequately surveyed Softwood shield Bird Conservation Region (BCR) from the northern portion of the SGS area, only 48% of juvenile woodcock originated in areas currently surveyed by the SGS. Of the individuals assigned to the northernmost portions of the breeding range, several were harvested in the southern extent of the wintering range. Based upon this latitudinal winter stratification, we examined whether woodcock employed a leapfrog migration strategy. Using δ2Hf values and band-recovery data, we found some support for this migration strategy hypothesis but not as a singular explanation. The large harvest derivation of individuals from the northernmost portions of the breeding range, and the difference in breeding distributions within each Management Region should be considered in future range-wide conservation and harvest management planning for American woodcock. © 2016 The Wildlife Society.

Playa wetlands in the Great Plains, USA support a wide variety of plant species not found elsewhere in this agriculturally-dominated region due to the ephemeral presence of standing water and hydric soils within playas. If longer dry periods occur due to climate change or if changes in surrounding land use alter sediment accumulation rates and water storage capacity in playas, plant communities could experience decreased diversity, with lasting effects on ecosystem services provided by playas in the Great Plains and at a continental-level in North America. We quantified potential changes in playa wetland plant community composition associated with predicted changes in precipitation and land use in the Great Plains through the end of the 21st century. We conducted two six-month greenhouse experiments mimicking field conditions using intact mesocosms collected from playas in Nebraska and Texas. In the precipitation experiment, treatments derived from historical precipitation observations and three future moderate emissions (CMIP5 RCP4.5) downscaled climate projections were applied to mesocosms. For the land use experiment, treatments were simulated by nitrogen (N) applications to soil ranging from 0 to 100 mg-N L-1 with each precipitation event under historical rainfall patterns, representing increasing and decreasing area in agricultural use in playa watersheds. Plant communities tended to shift toward more native species under projected future climate conditions, but as N runoff increased, native species richness decreased. Agricultural land-use surrounding playas may have a greater effect on wetland plant communities than future alterations to hydrology based on climate change in the Great Plains; thus, efforts to reduce nutrient runoff into playas would likely mitigate loss in ecosystem function in the coming decades. National Science FoundationNational Science Foundation (NSF) [IIA-1355406, IIA-1430427]; USDA-NIFAUnited States Department of Agriculture (USDA) [MO-HANR0007]; Missouri Department of Conservation; University of Missouri; U.S. Fish and Wildlife ServiceUS Fish & Wildlife Service; U.S. Geological SurveyUnited States Geological Survey; Wildlife Management Institute This research was part of the Missouri EPSCoR project, funded by the National Science Foundation under Award #IIA-1355406 and #IIA-1430427. Partial financial support was also provided by USDA-NIFA through Hatch funding (MO-HANR0007). Any opinions, findings, and conclusions or recommendations expressed in this manuscript are those of the authors and do not necessarily reflect the views of the National Science Foundation or USDA-NIFA. The Missouri Cooperative Fish and Wildlife Research Unit is jointly sponsored by the Missouri Department of Conservation, the University of Missouri, the U.S. Fish and Wildlife Service, the U.S. Geological Survey and the Wildlife Management Institute. Any use of trade, firm or product names is for descriptive purposes only and does not imply endorsement by the U.S. Government. This research would not have been possible without the assistance of Eric Simpson, Janith Chandrasoma, and Amanda Burnett. Finally, we would like to thank the School of Natural Resources Writing Workshop group for conceptual and technical edits which greatly improved the quality of this manuscript. Public domain – authored by a U.S. government employee