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Andrena (Longandrena) longiceps Morawitz, 1895 Andrena longiceps F. Morawitz, 1895: 65, ♀; Turkmenistan [Ahal Region], Germab (ZISP). Lectotype designated by A. Z. Osytshnjuk (Osytshnjuk et al. 2008) (Fig. 3). Male first description: Popov 1960: 247. Female. Body black, its length (vertex to T6) 7.5–8.5 mm (Figs 8, 9). Head: viewed frontally length approximately equal to width (L = 2.3–2.4 mm, W = 2.3–2.4 mm) (Fig. 10) or weakly longer or wider; lectotype body length 8.5 mm, its head L = 2.4 mm, W = 2.4 mm. Paraocular and supraclypeal areas above antennal socket level densely striate-wrinkled and sparsely punctate with dense long erect silver-white hairs; frons and vertex strongly shagreened, weakly punctate with short thin hairs; facial fovea shallow, moderately broad, dorsally occupying more than half ocellocular distance, narrowed below to level of antennal socket, covered with short dense very dark desaturated yellow pubescence. Compound eyes weakly convex, moderately narrow, 2.7 times longer than their maximum width; inner margin of compound eye curved only in upper third. Vertex widely concave; distance between middle and lateral ocelli equalling 2/3 of ocellus diameter (Fig. 11). Clypeus elongate and domed, black, polished, shiny, smoothed except basal part, with very sparse scattered rough hair-bearing punctures, hair long, plumose semi-adjacent, and along epistomal suture more densely and shallowly punctate with very short hairs, punctures separated by 0.5–2 puncture diameters; clypeus with basal half laterally weakly shagreened; apical part impunctate, extreme apex of clypeus impressed, laterally with elongate angles (Fig. 10). Genae as wide as compound eye, in upper part with dense adjacent long silvery-white hairs shorter near compound eyes; occiput covered with dense long flexed pubescence. Hypostomal carina clearly developed in lower part of hypostomal area and very weakly developed in upper part; lateral extremity of hypostomal carina with strongly developed subgenal coronet. Subantennal sutures only faintly discernible (Fig. 12). Malar area translucent, centrally 5 times shorter than width (Fig. 13). Antennae dark above, 3 rd –10 th flagellomeres light brown (in lectotype) or lighter yellow-brown (in Iranian specimens) beneath; first flagellomere slightly longer than 2 nd and 3 rd segments together (Fig. 14). Mandibles moderately short, their apical half brownish-red, lower margin with long yellow-white hairs, inner tooth small. Process of labrum widely trapezoidal, 2 times wider than long, shiny, densely transversely wrinkled basally and centrally, and strongly wrinkled laterally, with deeply emarginated apex bearing tuft of golden-yellow thick setae and with similar tufts on laterally lower parts; labrum apical margin ventrally with dense fringe of golden-yellow thick setae (Fig. 15). Galea shiny, strongly shagreened, impunctate, its apical half covered with tubercles; galeal fringe with short setae (Fig. 16); maxillary palpus equal 2/3 length of galea (Fig. 18). Glossa strongly pointed and very long, more than 16 times longer than its widest part, and more than 2 times longer than labial palpus (Fig. 17); paraglossae well developed (Fig. 19). Mesosoma: W (between tegulae) = 1.6 mm, scutum smooth and shining, without shagreen, sparsely, coarsely and irregularly punctate, centrally polished, strongly shiny with single hair-bearing punctures with moderately short erect silver-white plumose hairs; laterally near tegulae and posteriorly with shorter, denser pale gray pubescence (Fig. 20). Mesepisternum and metanotum with strongly granular shagreenation and with long erect silver-white plumose pubescence. Propodeal triangle very shiny, impunctate, weakly shagreened with larger mesh than on propodeum laterally, centrally weakly concave, basally wrinkled; posterior and lateral surfaces of propodeum strongly shagreened, finely reticulate, in upper part with very short dirty gray pubescence, in lower half with very dense long propodeal corbicula of silvery-white hairs (Fig. 21). Tegulae light yellow, translucent, dark brown at base; wings hyaline, venation and stigma yellow-brown, nervulus interstitial (in lectotype and one of Iranian specimens) or weakly antefurcal. Legs dark-brown, with dense, short silvery-white hairs, flocculus well developed; scopa with unbranched hairs. Inner metatibial spurs weakly curved along entire length, not widened, apical part straight (Fig. 22), 2 nd –5 th tarsal segments strongly narrowed basally, tarsal claws with pronounced inner teeth (Fig. 25). Metasoma black (L = 3.9 mm; W = 2.4 mm), terga shining, weakly shagreened, irregularly shallowly punctate, marginal parts of T1–T4 weakly depressed, yellow-white translucent with wide bands of dense silver-white hairs, moderately narrowed medially and on T1 weakly interrupted, terga basal part with short sparse gray-yellow setae, laterally with long, curved semi-erect silvery-white hairs; prepygidial and pygidial fimbriae long goldish-yellow and light brownish, covered with longer white-gray hairs dorsally (Fig. 29). Pygidial plate strongly convex and wrinkled centrally with flat margins (Fig. 30). Sterna dark brown, strongly shagreened, superficially, densely and finely punctate, apical margins transparent with very sparse bands of plumose white hairs, basal parts with short gray-yellow setae (Fig. 31). Male. Body black, its length (vertex to T7) = 7–8 mm. Similar to female in body shape, sculpturing, and coloration of pubescence (Figs 32–33). Head: slightly longer than its width (L = 2.0–2.1; W = 1.9–2.0 mm), shiny and wrinkled except smoothed clypeus, malar and lower part of paraocular areas, with silvery-white long sparse hair (Fig. 34). Compound eyes weakly convex, wider than in females, 2.3 times longer than their maximum width; inner margin of compound eye curved only in upper third. Vertex widely concave. Distance between middle and lateral ocelli equaling 4/5 of ocellus diameter. Clypeus elongate, polished, shiny with very weakly depressed apical margin, much weaker than in female, colored yellow with exception of two small lateral dark triangular marks, and black narrow strips along epistomal suture, with very sparse hair-bearing punctures, and laterally on apical part with short, dense silvery-white pubescence. Lower part of paraocular area also yellow, polished, shiny. Malar area longer than in females, 4 times as wide as long, basally near compound eyes with sparse short silvery-white hairs (Fig. 35). Process of labrum wide trapezoidal, 2 times wider than its length, shiny, polished, smooth except for narrow, wrinkled basal part (Fig. 36). Mandibles moderately short, not pointed, not reaching process of labrum, with long sparse hairs on lower margin, preapical tooth very weakly developed (Fig. 37). Both subantennal sutures inconspicuous. Scapus slightly curved, with sparse erect silvery-white hairs. Antennae moderately short, 1 st flagellomere twice as long as wide at apex, equal to 2 nd and 3 rd taken together, 2 nd 0.75 times shorter than its width and 1.5 times shorter than 3 rd, 3 rd –10 th – 1.25 times longer than their width, and 11 th – twice as long as wide; 2 nd –11 th flagellomeres light yellowbrown beneath (Fig. 38). Galea shining and wavy shagreened with individual superficial punctures, almost impunctate (Fig. 39). Glossa very long, 20 times longer than its greatest width (Fig. 40); paraglossae well developed. Mesosoma: W (between tegulae) = 1.5 mm, black; pronotum weakly shagreened, shiny; scutum smoothed, polished, very shiny, coarsely punctate, punctures separated by several puncture diameters, with sparse long plumose white pubescence, except for almost impunctate and hairless central part; laterally near tegulae pubescence shorter, denser, and with darker dirty gray hairs; anterior part of scutum very weakly shagreened, shining; central dorsal part of scutellum polished, strongly shining, almost impunctate, hairless, lateral and posterior parts sparsely punctate and covered with long erect silver-white pubescence (Fig. 41). Mesepisternum coarsely shagreened, finely punctate by hair-bearing punctures with long dense silvery-white hairs (Fig. 42). Propodeal triangle marked by more polished shiny and coarse mesh, basally with long rugosity extended to medial part (Fig. 43); propodeum lateral parts slightly duller with sparse, long, erect plumose white hairs. Tegulae translucent white-yellow, darker brown basally; wings hyaline, venation with light amber coloration, stigma darker, nervulus interstitial (in two of Iranian specimens) or weakly antefurcal. Legs dark brown, hind trochanter and femur with sparse long white hairs, tibia and basitarsus with dense short light hairs. Inner hind tibial spur not broadened, light yellow. Hairs of last tarsal segments white with faint light golden tints. Metasoma: L = 3.8 mm; W = 2.3 mm (Figs 44, 45). Terga black, polished, shiny, weakly punctured; T1–T4 marginal parts slightly depressed, lightened yellow-white with long dense white hair bands, widely interrupted on Т1, its basal part with long, sparse, erect white hairs; T2–T4 basal parts with sparse fine white-yellow setae. Sterna dark brown, medial part of S2–S5 strongly shagreened, sparsely punctate, punctures more distinct than on terga, with short thin yellow-brown setae on basal part; marginal parts lightened yellow-white with sparse complete white hair bands. S7 moderately narrow, its margin part centrally with almost rectangular ventrally flat projection with strongly bent edges and two dense tufts of long white plumose hair apically (Figs 46, 47). S8 of irregular rhombic shape with broadly rounded base, marginal parts around densely covered by long, plumose yellow-white hairs (Figs 48, 49). Genitalia elongate, almost twice as long as wide (Figs 50–56). Gonocoxites with very long dorsal lobes, gonostyli narrowed in apical half and very curved, their apices strongly diverging laterally, penis valves very dilated and flattened in basal half, and strongly narrowed and pointed apically; volsella rounded, moderately large. Distribution. Turkmenistan, Uzbekistan, Central Kazakhstan (Popov 1960; Pisanty et al. 2021), Iran (first record) (Figs 1, 2). Floral visitation. This species has been recorded on the flowers of Astragalus sp. (Fabaceae), Lycium sp. (Solanaceae), Tamarix sp. (Tamaricaceae), and Cirsium sp. (Asteraceae) (Popov 1960, and this study). Flight period. The representatives of the species start flying from mid-April in the southern parts of its distribution range, and from mid-May in the northern areas, where females have been observed until mid-July (Popov 1960)., Published as part of Radchenko, Vladimir G., Allahverdi, Mohammad & Fekrat, Lida, 2021, Revision of the mining bee subgenus Andrena (Longandrena) (Hymenoptera: Apoidea: Andrenidae), pp. 489-515 in Zootaxa 5032 (4) on pages 494-501, DOI: 10.11646/zootaxa.5032.4.2, http://zenodo.org/record/5497060, {"references":["Osytshnjuk, A., Romasenko, L., Banaszak, J. & Motyka, E. (2008) Andreninae of the Central and Eastern Palaearctic. Part 2. Polish Entomological Monographs. Vol. 5. Polish Entomological Society, Poznan ', Bydgoszcz, 233 pp.","Popov, V. B. (1960) New and little-known species of bees (Hymenoptera, Apoidea) from Turkmenia. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, Leningrad, 27, 247 - 263. [in Russian]","Pisanty, G., Richter, R., Martin, T., Dettman, J. & Cardinal, S. (2021) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 107151. [in press] https: // doi. org / 10.1016 / j. ympev. 2021.107151"]}

Andrena (Longandrena) nativa Osytshnjuk 1984 Andrena nativa Osytshnjuk, 1984: 93; ♀; Turkmenistan, 100 km NE Kyzyl-Arvat, [Serdar], Kirpili (ZISP) (Fig. 4). Female (holotype and paratype). Body length (vertex to T6) 7.0–7.5 mm (Figs 96, 97). Head and mesosoma with blue-green metallic sheen, metasoma black. Body mainly with silver-white to gray-white pubescence, in some places with yellow tint due to color of collected pollen. Head: viewed frontally of almost equal width and length (L = 2.1, W = 2.2 mm), very slightly narrowed downwards, almost square, densely longitudinally striated on upper and central part of paraocular areas and on frons, and with inconspicuous hair-bearing punctures (Fig. 99). Lower paraocular area shining, polished, sparsely punctate, punctures small and separated by 2–4 puncture diameters. Facial fovea wide, depressed, especially in lower part, with white pubescence, occupying 2/3 ocellocular distance (Fig. 102), ventrally reaching level of clypeus base. Space between antennal scapes smoother and more densely punctate; supraclypeal area striate-rugose, almost impunctate. Compound eyes weakly convex, slightly wider than in other females of subgenus, 2.4 times longer than their maximum width, inner margin of compound eye curved only in upper half; distance between central and lateral ocelli almost equal to ocellus diameter. Head frontally with more or less long, dense erect hairs, and on genal area with semi-appressed ones. Clypeus slightly elongated, wide, almost hexagonal, centrally broadly concave, polished, strongly shining with very scattered isolated punctures bearing long semi-appressed white hair; lateral margins of clypeus in its basal half and narrow part of its base weakly shagreened, with scattered punctures. Narrow apical margin and widely elongated lateral angles of clypeus strongly depressed, at base of depression and apical half of clypeus laterally with long dense semi- appressed white hairs. Vertex strongly narrowed along posterior margin, widely concave, coarsely shagreened, with sparse punctures, except for smoothed, polished, shiny triangular area located between ocelli; distance between apical margin of vertex and lateral ocellus almost equal to ocellus diameter. Gena upper part moderately broad, equaling width of compound eye, significantly narrowed at lower part; gena surface shagreened, shiny, with sparse hair-bearing punctures (Fig. 100). Malar area short, 5 times wider than its length (Fig. 104). External subantennal sutures clearly discernible (Fig. 106). Mandibles basally black, their apical half light-red with almost undeveloped preapical teeth (Fig. 105); lower margin of mandibles with fringe of long sparse white hair. Process of labrum large, trapezoidal (Fig. 107), 2 times shorter than its width, transversely wrinkled in basal part, smoothed, polished and strongly shining in apical half, depressed margins of process with deep hair-bearing punctures; labrum apically with a cavity that is visible only ventrally (Fig. 108), with dense tuft of short golden-yellow hairs, broadly rounded with slightly elongated lateral corners and with continuous fringe of long thick golden-yellow hairs. Glossa narrow and very long (Fig. 103), more than 15 times longer than its base width, much longer than labial palpus. Galea long, weakly shagreened, with very sparse small superficial punctures bearing thin short setae, and along the lower margin with a fringe of long sparse hair, denser and longer at galea apex (Fig. 101); maxillary palpus slightly longer than galea. Antennae dark above, and flagellum red-yellow beneath (Fig. 98). Mesosoma: W (between tegulae) = 1.6 mm. Pronotum without lateral carinae. Scutum strongly shining, polished, except for weakly shagreened narrow anterior part, with irregularly scattered punctures, separated by 0.5–4 puncture diameters, centrally with very sparse punctures bearing long semi-erect whitish hairs (Fig. 109). Scutellum centrally polished, strongly shiny, almost impunctate, hairless, and on margins clearly shagreened, punctate with dense white erect pubescence. Mesepisternum entirely shagreened with small superficial punctures bearing long, dense plumose hairs. Propodeum finely reticulate, propodeal triangle with shinier and coarser reticulation, well-marked only on posterior half (Fig. 110). Propodeal corbicula incomplete with sparse long weakly flexed hairs; floccus well developed with long more strongly flexed hairs. Tegulae light-yellow, transparent; wings hyaline; venation and stigma yellow; forewing nervulus strongly antefurcal. Tibial scopa with long sparse intermixed simple and weakly plumose silver-white hairs, inner part of tibia with long plumose and weakly flexed hairs; metatibial inner spurs widened almost along entire length, strongly curved at apex (Fig. 24); 2 nd –5 th tarsal segments of all legs widened, yellow-red (Fig. 28); midbasitarsus curved laterally, central inner part of this segment with more short and dark pubescence (Fig. 111). Metasoma: L = 4 mm, W (in widest part) = 2.3 mm, black, ovoid (Figs 113, 114). Terga weakly convex, shining; T1 smoothed, polished, strongly shiny on basal part, weakly shagreened before apical part, with irregularly scattered small punctures, separated by 2–3 puncture diameters, in some parts more sparsely punctate, punctures denser before narrow impunctate apical margin, here separated by one puncture diameter; T2–T4 centrally weakly shagreened, more densely and evenly punctate, punctures separated by 1–2 puncture diameters; T3–T4 weakly and shallowly punctate; apical margins of all terga strongly depressed, lightened, transparent, densely and weakly punctate at base of marginal part. Tergal marginal zones with wide bands of dense and long appressed white hairs narrowly interrupted at T1, apical hairbands on T2–T4 broad, occupying more than 1/3 of tergum length, basal parts of these terga centrally with sparse short semi-appressed white hairs, and with semi-erect hairs laterally. Pygidial plate flat, triangular, rounded at apex, without lateral grooves (Fig. 112). Pygidial and prepygidial fimbriae yellowish-white. Sterna more strongly shagreened and more densely punctate than terga; basal parts of S2–S5 centrally impunctate, hairless, on S3 strongly convex; marginal parts of S2–S5 enlightened, transparent with long appressed thin white hairs (Fig. 111). Male: unknown. Studied material: 2 ♀ (holotype and paratype): 1 ♀ Turkmenistan, Kirpili, 100 km NE Kyzyl-Arvat [Serdar] 27.iv.[1]953, [on flowers of] Astragalus, leg. E. Arens (holotype – ZISP; Fig. 4); 1 ♀ ibid, 27.iv.[1]953, leg. E. Arens (paratype – SIZK; Fig. 7). Another female paratype specified in the description and was stored in SIZK (Osytshnjuk 1984): Turkmenistan, Uzboy, Alty-Kuyu, 28.iv.1952, leg. O. Kryzhanovsky, probably lost. Distribution. The species is known only by the original description from Turkmenistan (Fig. 2). Flight period. Late April. Floral visitation. Females were collected at the flowers of Astragalus (Fabaceae)., Published as part of Radchenko, Vladimir G., Allahverdi, Mohammad & Fekrat, Lida, 2021, Revision of the mining bee subgenus Andrena (Longandrena) (Hymenoptera: Apoidea: Andrenidae), pp. 489-515 in Zootaxa 5032 (4) on pages 507-508, DOI: 10.11646/zootaxa.5032.4.2, http://zenodo.org/record/5497060, {"references":["Osytshnjuk, A. Z. (1984) Four new species of the genus Andrena (Hymenoptera, Andrenidae) from Middle Asia. In: Savchenko, E. N. (Ed.), Taxonomy and Zoogeography of Insects. A Collection of Papers. Naukova Dumka, Kiev, pp. 89 - 94. [in Russian]"]}

Phthiria notiosa Gharali, Mogbeli & Fekrat sp. nov. (Figs. 1���4) Type material. Holotype (male), 1 female & 1 male paratypes (dried by HMDS) and 20 female & 10 male paratypes (in alcohol), Kerman, Hanza, Bondare-Hanza N 57��12��� 39���, E 19��19��� 29���, 2767 m a.s.l., white pan traps, leg. Sajjad Mogbeli, 10.vi.2009 (IRIPP); 10 female and 10 male paratypes, same data as holotype (ZMHB), 74 female and 22 male paratypes, same data as holotype (BG). Diagnosis. Phthiria notiosa sp. nov. is morphologically similar to Ph. vagans Loew, 1846, but in the female, it is easily distinguished by the black scutellum (yellow with a small black mark in Ph. vagans) and in the male, Phthiria notiosa sp. nov. is easily distinguished from the other species of Phthiria belonging to the group with short prong of first flagellomere (see Gharali & Evenhuis, 2017 for grouping) by having two longitudinal dusted stripes from the anterior to the posterior margin of mesonotum. Description. Holotype male. Body length: 4 mm, wing length: 5 mm. Head (Fig. 1b, c) completely black; frons black, gray dusted, with long white hairs; parafacials completely black, with long white hairs; mentum gray dusted, with sparse long white hairs; ocellar tubercle prominent black, with a few white hairs; eyes in contact for about 2.0 times length of frons, contact sector of eyes medially depressed, facets of upper half of eyes enlarged; antennae (Fig. 1c) completely black, scape quadrate, wider than long, pedicel round, first flagellomere long, about 5 times greatest width, dorsal prong about 1.5 times length of transparent style; palpi black, proboscis black, long, 2.5 times head length. Thorax (Fig. 1d, e). Mesonotum black with two dusted longitudinal stripes from anterior margin to posterior margin of mesonotum, with sparse long white hairs marginally, length of hairs about equal to combined length of two basal antennal segments; scutellum black, gray dusted with long dense white hairs marginally; pleura completely black, gray dusted, anepisternum and katepisternum with long white hairs, laterotergite with few white hairs, halter stem and knob white. Wing. Hyaline, uniformly microtrichose; veins blackish brown; costa reaches vein CuA1+A1; R 4+5 originates before level of m-m crossvein; crossvein r-m beyond middle of discal cell; anal cell closed before wing margin, stalked; anal lobe broad. Leg (Fig. 1a). Black; coxae and femur basally with long white hairs; tibia with a few black bristles; tarsal segments ventrolaterally with dense black bristles; empodium white, as long as claw. Abdomen (Fig. 1a). Tergites and sternites black with sclerotized black spots, sternites with long white hairs, tergites laterally dusted triangularly. Terminalia. Hypopygium black; epandrium triangular in dorsal view, with small invagination apically (Fig. 2c); gonocoxites ovoid, fused medially narrowly, apicomesal process as long as gonostylus, with small bristles; gonostylus oval, apically curved, pointed (Figs. 2b, c); basal aedeagal process narrow in dorsal view with two lateral rami basally, lateral aedeagal apodemes narrow (Fig. 2a); aedeagus apically curved (Fig. 2d). Female. Head. (Figs. 3a, b, f) Occiput black with eye margins yellow; frons black medially, almost yellow laterally, ocellar triangle black; ocelli an equilateral triangle; oral margin black narrowly, with short scattered white hairs; antennae completely black, scape slightly longer than wide; pedicel quadrate; first antennal flagellomere long, with short dorsal prong; style transparent, shorter than antennal prong; insertion of antennae yellow above; mentum black medially; palpi black, with a few short white hairs; proboscis long, 2.5 times head length, black. Thorax (Fig. 3c, b). Mesonotum glossy black, bare, with two grey dusted stripes extended from anterior margin to posterior margin of mesonotum, laterally yellow, postpronotum white, scutellum black, bare; pleura black except following: anteroventral corner of anepisternum, upper margin of katepisternum and meron; lower margin of anepimeron and laterotergite; halter stem and knob white. Leg. Coxae black with scattered white hairs, femora black, tibiae black with small black bristles, tarsi black, empodium yellow, as long as claw. Wing (Fig. 4a). Hyaline, uniformly microtrichose; veins blackish brown; costa reaches vein CuA1+A1; R 4+5 originates before level of m-m crossvein; crossvein r-m beyond middle of discal cell; anal cell closed, stalked; anal lobe wide. Abdomen (Fig. 3e). Tergites glossy black with posterior margins yellow; sternites black, bare. Terminalia (Figs. 4b���c). Furca consisting of two parallel bars; common spermathecal duct short, membranous; basal ducts membranous; sperm pump narrower than apical ducts, striated, with apical valve absent, apical ducts sclerotized, curved; basal sac membranous, round, spermathecal reservoir well sclerotized, cone-shaped, slightly curved, about 3 times capsule length. Distribution. This species is currently known only from Bondar-Hanza village belonging to Hanza city. This city is in a small part of Kerman province in the south of Iran. Specimens were collected by using white pan traps scattered in a mountainous area (Fig. 5) dominated mostly by plant in Asteraceae (Artemisia sieberi Besser, Hertia intermedia Kuntz, and Astragalus fasciculifolius Boiss) and Apiaceae (Ferulago angulate (Schlecht), and Ferula oopoda (Boiss. et Buhse). Etymology. The name of new species, Phthiria notiosa Gharali, Mogbeli & Fekrat sp. nov. is derived from the Greek �������������� meaning ���southern��� referring to the distribution of the new species being confined to the southern parts of Iran. Taxonomic remarks. Engel (1933) was the first who presented a key to almost all known species of the genus Phthiria in the Palaearctic Region. Using Engel���s key, the male specimens of the new species run to Ph. vagans ssp. pallescens , but Phthiria notiosa Gharali, Mogbeli & Fekrat sp. nov. is distinguished by two longitudinal dust stripes (absent in Ph. vagans ssp. pallescens) and the color of halter knob, which is completely yellow (with a brown spot dorsally in Ph. vagans ssp. pallescens). The female specimens of Phthiria notiosa Gharali, Mogbeli & Fekrat sp. nov. run to couplet 23 where Ph. incisa and Ph. vagans are separated from each other based on the color of scutellum. This new species is also separated from both those species by the color of scutellum, being completely black in the new species, as opposed to the brown with a central yellow spot (Ph. incisa) and almost completely yellow (Ph. vagans)., Published as part of Moghbeli, Sajjad, Gharali, Babak & Fekrat, Lida, 2021, Phthiria notiosa sp. nov., a new bombyliid species of the tribe Phthiriini (Diptera Bombyliidae, Phthiriinae) from Kerman province in the south of Iran, pp. 378-386 in Zootaxa 4990 (2) on pages 379-385, DOI: 10.11646/zootaxa.4990.2.11, http://zenodo.org/record/5026012, {"references":["Loew, H. (1846) Fragmente der Kenntniss der europaischen Arten einiger Dipteren Gattungen. Linnaea Entomologica, 1, 319 - 530.","Gharali, B. & Evenhuis, N. L. (2017) Phthiria vaganoides sp. nov. (Bombyliidae, Phthiriinae, Phthiriini) from Northern Iran with the first record of Phthiria fulva Meigen, 1804 reared from a lepidopteran and description of the pupa. Zootaxa, 4300 (1), 055 - 070. https: // doi. org / 10.11646 / zootaxa. 4300.1.3","Engel, O. (1933) Bombyliidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. IV (3). E. Schweizerbart, Stuttgart, pp. 97 - 144."]}

Genus Phthiria Meigen Phthiria Meigen, 1803: 268. Type species: Bombylius pulicarius Mikan, 1796, by monotypy. Ptimia Rafinesque, 1815: 221 (unjustified emendation of Phthiria Meigen, 1820). Type species: Bombylius pulicarius Mikan, 1796, automatic. Diagnosis. The family Bombyliidae is divided in two artificial groups, Homoeophthalmae and Tomophthalmae, based on postcranial features. Phthiria belongs to the Homoeophthalmae and the subfamily Phthiriinae, in this subfamily the genus Phthiria is easily recognized from all other genera by the following characters: Metapleuron with micropubescence, first flagellomere with short and sparse hairs, costal vein ending at anal vein, male terminalia rotated 180��, basistylus without mesal lobe bearing spines, apicomesal lobe well developed, and female spermathecal reservoir curved apically (Evenhuis, 1990; Greathead & Evenhuis, 2001)., Published as part of Moghbeli, Sajjad, Gharali, Babak & Fekrat, Lida, 2021, Phthiria notiosa sp. nov., a new bombyliid species of the tribe Phthiriini (Diptera Bombyliidae, Phthiriinae) from Kerman province in the south of Iran, pp. 378-386 in Zootaxa 4990 (2) on page 379, DOI: 10.11646/zootaxa.4990.2.11, http://zenodo.org/record/5026012, {"references":["Meigen, J. W. (1803) Versuch einer neuen GattungsEintheilung der europaischen zweiflugligen Insekten. Magazin fur Insektenkunde, 2, 259 - 281.","Mikan, J. C. (1796) Monographia Bombyliorum Bohemiae iconibus illustrata. J. Herrl, Pragae [Prague], 59 + [1] pp.","Rafinesque, C. S. (1815) Analyse de la nature ou tableau de l'univers et des corps organises. Le nature est mon guide, et linneus mon maitre. Privately published, Palermo, 224 pp. https: // doi. org / 10.5962 / bhl. title. 106607","Meigen, J. W. (1820) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Zweiter Theil. F. W. Forstmann, Aachen, x + 363 pp.","Evenhuis, N. L. (1990) Systematics and evolution of the genera of the subfamilies Usiinae and Phthiriinae of the world (Diptera: Bombyliidae). Entomonograph, 11, 1 - 72.","Greathead, D. J. & Evenhuis, N. L. (2001) Annotated keys to the Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) of Africa. African Invertebrates, 47, 105 - 244."]}

Empidideicus jirofti Gharali, Mogbeli & Fekrat sp. nov. Material examined. Holotype female (dried by HMDS)and 15 females and 20 males paratypes (preserved in alcohol, 1 female and 1 male paratypes dissected); IRAN, Kerman Province, Jiroft City, Esfandagheh Village, 57��28���21.07���E, 28��44���08.67���N, 1255m, 30 May 2017, leg. S. Mogbeli. Type depository. Holotype and 5 female and 5 male paratypes in MTD, 5 female and 5 male paratypes in BPBM, the other paratypes in BG. Diagnosis. Empidideicus jirofti Gharali, Mogbeli & Fekrat sp. nov. is differentiated from all other species of the genus from Iran by the combination of following characters: Frons yellow with a quadrate brown spot below ocellus, occiput brown with a small yellow spot; mesonotum brown with two admedian yellow stripes; scutellum brown with yellowish tinge basomedially; furca U-shaped with two process on lateral arms; reservoir globular with shallow invagination; gonocoxae with two triangular plate including perpendicular pointed process. Description. Male (Fig. 2a). Lengths: Body: 0.9���1.3 mm. Wing: 0.8���1.2 mm. Head (Fig. 2 a���b). Head ovoid, longer than high; eyes dichoptic, separated at vertex by about 1.6 times between lateral ocelli; vertex blackish brown; occiput blackish brown, with tiny yellow spot in upper half; frons slightly depressed medially, yellow with quadrate brown spot just below median ocellus; antennae set in deep pocket; Scape yellow, subrectangular, wider than long, Pedicel yellow, subglobular, slightly wider than long, first flagellomere light brown, elliptical, length two-time greatest width, about three times (3.2) second flagellomere in length, second flagellomere light brown, cylindrical, apically with tiny hyaline stylus (Fig.1a); face yellow; mentum yellow medially brown laterally; labrum stiff, pointed apically, brown, about 1.2 head high in length; labium brown, 1.7 head high in length, pointed apically. Thorax (Fig. 2 a���c). Mesonotum completely brown with two admedian yellow stripes coming from anterior margin of mesonotum to level of postalar calli; postpronotal lobe, postalar calli, lateral margin from postpronotal to postalar calli yellow; prescutellar area completely blackish brown; scutellum brown medially with yellowish tinge; pleura predominantly dark brown except following: upper margins of katepisternum and meron and dorsolateral corner of anepisternum and posterior margin of anepimeron; halter stem and knob white. Legs. Femora brown, distally narrowly yellow; tibia yellow; two basal segment of tarsi yellow, remainder brown. Wing (Fig. 1b). Hyaline, costa ends slightly beyond vein R 4+5, Sc ends much before costa; br and bm cells confluent, discal cell absent, M 1 curve; M 2 straight; M 1+2 0.7 times M 2 in length; anal cell open for width subequal to r-m cross vein; veins M 1, M 2,Cua 1 translucent. Abdomen. Tergites almost brown, posterior and lateral margins narrowly yellow; sternites brown, posterior margins narrowly yellow; pleura completely white. Male genitalia (Fig. 3). Epandrium (Fig. 3b) subrectangular, laterally with two processes; cercus triangular, large; gonocoxites (Fig. 3c) connected narrowly medially, subtriangular in dorsal view, apical margin round less sclerotized than other parts, with two sclerotized triangular plates including perpendicular pointed spike, medially with narrow apically curved sclerotized appendage; gonostyli subtrapezoidal with round margin including long bristles; distiphallus (Fig. 3a) wide, short, lateral aedeagal apodeme slightly curved, longer than tumefied basiphallus; apical aedeagal apodeme large with two large rectangular rami basally. Female (Fig. 2 b���c). Similar to male except larger in size and with larger yellow spot on lateral margin of occiput and pleural sclerites lighter. Female genitalia (Fig. 4). Furca U-shaped, lateral arms with two inwardly directed processes subapically (Fig. 4c); common spermathecal duct absent; basal spermathecal duct (Fig. 4a) 1.2 times apical duct in length, striated, membranous, slightly widening distally, one tenth of apical part cylindrical, narrow, smooth; apical spermathecal duct narrower than basal duct, tubular, smooth, spermathecal reservoir (Fig. 4b) globular with shallow, cylindrical invagination, depth of invagination one tenth dimeter of reservoir. Remarks. The species of the genus Empidideicus described from Iran have been recently keyed by Gharali et al. (2014). This new species is stymied at couplet 3 because the lateral margin of occiput is not completely yellow or black. Empidideicus jirofti Gharali, Mogbeli & Fekrat sp. nov. is easily separated from the other species by this character. Distribution. The new species is currently known only from Jiroft city (Kerman province) in the south of Iran (Fig. 5). The type locality is a mountainous place covered with plants of the families Rhamnaceae (Ziziphus spp.), Fabaceae (Astragalus spp.), and Gramineae. Etymology. The name of new species, Empidideicus jirofti Gharali, Mogbeli & Fekrat sp. nov., is derived from the word Jiroft, a city in the southern Kerman province, an area of about 18 square kilometers with hot and humid climate, so this city is called the greenhouse city of Iran., Published as part of Moghbeli, Sajjad, Fekrat, Lida & Gharali, Babak, 2020, Empidideicus jirofti sp. nov. (Diptera: Mythicomyiidae: Empidideicinae), a new microbombyliid from Kerman province, southern Iran, with a key to species of Empidideicus from Iran, pp. 352-360 in Zootaxa 4722 (4) on pages 353-359, DOI: 10.11646/zootaxa.4722.4.4, http://zenodo.org/record/3609589, {"references":["Gharali, B., Majnon Jahromi, B., Evenhuis, N. L. & Fallahzadeh, M. (2014) First record of the genus Empidideicus Becker (Diptera: Mythicomyiidae, Empidideicinae) from southern Iran with description of a new species, Entomologist's Monthly Magazine, 151, 27 - 34."]}

Harmonia axyridis (Pallas, 1773) Figs 1���4, 6, 14���16, 20 Coccinella axyridis Pallas, 1773. Material examined. IRAN: GILAN PROVINCE: Rudsar County, Rudsar, 37��8���11.739���N, 50��16���55.037���E, vi.2016, 2 spec., ex Corylus sp., Z. M. Haghghadam leg., A. Biranvand det. ; Rasht County, Rasht, Melat Park, 37��17���27.151���N, 49��35���47.65���E, iii.���ix.2016 + 2017, 85 ♂♂, 230 ♀♀, ex Corylus sp., Hibiscus sp., Lagerstroemia sp., Nerium sp., Populus, sp., Prunus sp., Robinia sp., Z. M. Haghghadam & M. Mardani-Talaee leg., A. Biranvand & O. Nedvěd det.; Fuman County, Fuman, Shahr Park, 37��13���51.45���N, 49��19���6.134���E, viii.2017, 1 spec., ex Lagerstroemia sp., Z. M. Haghghadam leg., A. Biranvand det.; Langarud County, Langarud, 37��11���33.1���N, 50��09���09.2���E, vii.2017, 5 spec., ex Citrus sp., Z. M. Haghghadam leg., A. Biranvand det.; Lahijan County, Lahijan, 37��11���22.652���N, 50��1���10.229���E, viii.2017, 2 spec., ex Quora sp., Z. M. Haghghadam leg., A. Biranvand det. MAZANDARAN PROVINCE: Nur County, Chamestan, vii.2017, 1 spec., E. Shafiee & E. Dehghan leg., M. Z. Khormizi det.; Ramsar County, Sadat Shahr, 36��53���21.449���N, 50��41���45.658���E, vii.2017, 1 spec., E. Shafiee & E. Dehghan leg., M. Z. Khormizi det. SAUDI ARABIA: RIYADH PROVINCE: Diriyah, 24��48���43.39���N 46��31���17.27���E, v.2005, 1 spec. (forma conspicua), S. A. Turkestani leg., A. N. Al Ansi det. TURKEY: BILECIK PROVINCE: Bilecik Central District, Pelit��z��, 40��11���6.9756������N, 29��58���5.5704���E, xi.2014, ix.���xi.2015, iii.���viii.2016, 55 spec., ex Cydonia sp., Juglans sp., Malus sp., Medicago sp., Morus sp., Phaseolus sp., Prunus sp., weeds, insidebuildings, D. Şenal leg.et det.; Osmaneli District, Sarıyazı village, 40��27���19.188���N, 29��56���22.56���E, xi.2014, xi.2015, vi.2016, 10 spec., ex Abelmoschus sp., Cydonia sp., Paliurus sp., Solanum sp., Triticum sp., D. Şenal leg.et det.; Yenipazar District, 2.5 km NWYenipazar, G��lpazarı Yenipazar Yolu, 40��11���30.048���N, 30��30���5.184���E, vi.���xi.2015, vi.���x.2016, 46 spec., ex Helianthus sp., Medicago sp., Paliurus sp., Prunus sp., Pyrus sp., Triticum sp., Vitis sp., D. Şenal leg. et det.; S��ğ��t District, K��re, 40��5���14.892���N, 30��9���2.7936���E, vii.���xi.2015, vi.���viii.2016, 20 spec., ex Abelmoschus sp., Capsicum sp., Cucurbita sp., Juglans sp., Medicago sp., Mentha sp., Phaseolus sp., Prunus sp., Solanum sp., weeds, D. Şenal leg.et det.; Pazaryeri District, Arapdede, 39��59���40.92���N, 29��52���44.328���E, viii.���xi.2015, vi.���viii.2016, 25 spec., ex Humulus sp., Juglans sp., Malus sp., Medicago sp., Pinus sp., D.Şenal leg.et det.; Boz��y��k District, 1.6 km NW ��ok��apınar, ��ok��apınar Ecek��y Yolu, 39��46���19.2���N, 29��58���3.9���E, ix.2015, viii.2016, 4 spec., ex Medicago sp., Quercus sp., D. Şenal leg. et det.; G��lpazarı District, 4.2 km SW G��lpazarı, Reşadiye Mahallesi, 40��15���53.2764���N, 30��16���26.5044���E, viii.���xi.2015, vii.2016, 25 spec., ex Helianthus sp., Malus sp., Prunus sp., weeds, D. Şenal leg. et det.; İnhisar District, Karaağa�� Mahallesi, Mihalgazi ��� İnhisar Yolu, 40��2���42.36���N, 30��24���9.684���E, vii.���xi.2015, vi.2016, 67 spec., ex Capsicum sp., Paliurus sp., Punica sp., Solanum sp., weeds, D. Şenal leg. et det. Notes. The specimen of H. axyridis from Saudi Arabia was already collected in 2005 but remained unidentified until now. As a result, this is the first formal report of H. axyridis from Saudi Arabia. The first publishedreports of this species in Turkey were from 2011, collected in the European part of the country, in Tekirdağ Province (AYSAL & KIVAN 2014). Other recent collections were made in several localities in the provinces of Bartın (TOPER KAYGIN & SOBUTAY KAPTAN 2017), ��anakkale (BAŞTUĞ & KASAP 2015), D��zce (��ZTEMIZ & YAYLA 2018), Isparta (OĞUZOĞLU 2017), Tekirdağ (AYSAL & KIVAN 2014), and Yozgat (TIFTIK��I 2017). Asingle specimen has been found in Nevşehir Province, in the vicinityof G��reme (BUKEJS & TELNOV 2015). This was suggested to be the first report of H. axyridis for Turkey but it was only collected in July 2013, whereas several specimens from Tekirdağ Province were collected in 2011���2012 (AYSAL & KIVAN 2014). In Iran, H. axyridis was for the first time collected in April 2016, from the city of Rasht in Gilan Province (MARDANI-TALAEE et al. 2019). Similar to what these authors found, we collected many adults from various host plants. Moreover, H. axyridis was reported from Israel based on a single specimen (forma succinea) collected in 2017 from Kibbutz Netzer Sereni (MIENIS 2017). All known records of H. axyridis from the Middle East are summarized in Fig. 20., Published as part of Biranvand, Amir, Nedv��d, Old��ich, Tomaszewska, Wioletta, Al Ansi, Amin N., Fekrat, Lida, Haghghadam, Zahra Mojib, Khormizi, Mehdi Zare, Noorinahad, Sara, ��enal, Derya, Shakarami, Jahanshir & Haelewaters, Danny, 2019, The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East region, pp. 163-170 in Acta Entomologica Musei Nationalis Pragae 59 (1) on pages 165-166, DOI: 10.2478/aemnp-2019-0014, http://zenodo.org/record/4552749, {"references":["PONTOPPIDAN E. 1763: Den Danske Atlas, eller Konge-Riget Dannemark. Efter HOy-Kongelig allernaadigst Befalning. Tomus I. A. H. Godiche, Kopenhagen, 723 pp.","AYSAL T. & KIVAN M. 2014: Occurrence of an invasive alien species Harmonia axyridis (Pallas) (Coleoptera: Coccinellidae) in Turkey. Turkiye Entomoloji Bulteni 4: 141 - 146.","TOPER KAYGIN A. & SOBUTAY KAPTAN U. 2017: Coccinellidae (Insecta: Coleoptera) species of Bartin Province. Journal of Bartin Faculty of Forestry 19: 227 - 236.","BASTUG G. & KASAP I. 2015: Faunistic studies on Coccinellidae (Coleoptera) family in the province of Canakkale. Turkish Journal of Biological Control 6: 41 - 50.","OZTEMIZ S. & YAYLA S. 2018: Two new species of Harmonia (Coleoptera: Coccinellidae) from Duzce, Turkey. Munis Entomology & Zoology 13: 318 - 322.","OGUZOGLU S. 2017. Cinara cedri'nin Isparta Orman Bolge Mudurlu g u Sedir Ormanlarinda Yayili s i, Zarari, Biyolojisi ve Do g al Du s manlari. [Distribution, damage, biology and natural enemies of Cinara cedri in Isparta forest regional directorate cedar forests]. Thesis deposited at Department of Forest Engineering, Suleyman Demirel University, Isparta, 116 pp.","TIFTIKCI P. 2017: Studies on Coccinellidae species identified on sugar beet production areas in Yozgat province. Journal of Natural and Applied Sciences 33: 79 - 90 (in Turkish, English abstract).","BUKEJS A. & TELNOV D. 2015: The first record of the invasive lady beetle Harmonia axyridis (Pallas, 1773) (Coleoptera: Coccinellidae) in Turkey. Zoology and Ecology 25: 59 - 62.","MARDANI-TALAEE M., ZIBAEEA., RAHIMI V., KHORMIZI M. Z., MOZAFAR MANSOURI S. & NEDVED O. 2019: Occurrence of invasive ladybeetle Harmonia axyridis (Pallas) (Coleoptera: Coccinellidae) in Iran. Coleopterists Bulletin 73: 1 - 7.","MIENIS H. K. 2017: A first record of the harlequin ladybird Harmonia axyridis from Israel (Coleoptera, Coccinellidae). Natural History and Other Notes 14: 5 - 7."]}

Harmonia quadripunctata (Pontoppidan, 1763) Figs 5, 7���13, 17���19 Coccinella quadripunctata Pontoppidan, 1763; C. albida Gmelin, 1790; C. marginella M��ller, 1776; C. marginepunctata Schaller, 1783; C. notata Olivier, 1791; C. sedecimpunctata Fabricius, 1781. Material examined. IRAN: NORTH KHORASAN PROVINCE: Bojnord County, Bojnurd, Baba Aman Park, 37��29.338���N, 57��26.123���E, vi.2013, 4 ♂♂ 4 ♀♀, (3 ♂♂ 2 ♀♀ ex Hordeum sp.), E. Hamidi leg., O. Nedvěd & C. Canepari det. SEMNAN PROVINCE: Shahrud County, Bastam, 36��30.314���N, 55��00.290���E, v.2014, 2 ♂♂ 1♀, M. Toozandejani leg., A. Biranvand det. TURKEY: BILECIK PROVINCE: Bilecik Central District, Pelit��z��, 40��11���06.9756���N, 29��58���05.5704���E, iv.���xi.2015, vii.2016, 31 spec., ex Malus sp., Pinus sp., Prunus sp., weeds, D. Şenal leg. et det.; Osmaneli District, 2.6 km NE D��zmeşe, 40��23���55.5���N, 29��56���40.092���E, vi.���ix.2015, vi.���viii.2016, 5 spec., ex Olea sp., Paliurus sp., Prunus spp., D. Şenal leg.et det.; İnhisar District, G��ner Orbay Mahallesi, 1.9 km NW İnhisar, 40��03���37.116���N, 30��22���5.088���E, vi.2016, 1 spec., ex Paliurus sp., D. Şenal leg. et det. Notes. Harmonia quadripunctata was originally described from Denmark (PONTOPPIDAN 1763). It is native to the Palearctic Region, distributed especially in Europe and eastern Asia ��� Russia, China, and Korea (BIELAWSKI 1963, 1980; VANDENBERG 1990; PANG et al. 2004; KOV��Ř 2007). However, there aresome reports of thisspecies from other countries of the Palearctic Regionincluding Syria (KHALIL & MOURAD 2006), Israel (HALPERIN et al. 1995), Lebanon and Turkey (G��NTHER 1958, BIELAWSKI 1963). Harmonia quadripunctata was introduced into the USA and reported there since 1924 (VANDENBERG 1990). In southern America, introduced populations have only been reported from Argentina and Chile so far (GONZ��LEZ 2006). The earliest record of H. quadripunctata in Turkey was provided by G��NTHER (1958). Since then, many studies reported H. quadripunctata in 22 of Turkey���s 81 provinces: Ankara (G��NTHER 1958, UYGUN 1981, ��LGENT��RK & TOROS 2000), Kayseri (G��NTHER 1958), Konya (G��NTHER 1958, ŞAHBAZ & UYSAL 2006), Adana, Afyon, Bursa, Denizli (UYGUN 1981), Isparta (UYGUN, 1981, ASLAN & KARACA 2005, KARACA et al. 2006, KAYA BAŞAR & YAŞAR 2011, OĞUZOĞLU 2017), Amasya, Samsun, Tokat (KILI�� & AYKA�� 1989), Erzurum (��ZBEK & ��ETIN 1991), İzmir, Manisa (TEZCAN & UYGUN 2003), Kahramanmaraş (ASLAN & UYGUN 2005), Balıkesir (VURAL VARLI et al. 2013), Artvin (PORTAKALDALI & SATAR 2010), ��anakkale (BAŞTUĞ & KASAP 2015), Bartın (TOPER KAYGIN & SOBUTAY KAPTAN 2017), Yozgat (TIFTIK��I 2017), D��zce (��ZTEMIZ & YAYLA 2018), and Bilecik (this study). In Iran, H. quadripunctata was first reportedfrom North Khorasan Province in 2013 (BIRANVAND et al. 2017). It was subsequently reported from Mazandaran Province, in northern Iran (PAHLAVAN YALI et al. 2017). Here, wepresent the first report from Semnan Province., Published as part of Biranvand, Amir, Nedv��d, Old��ich, Tomaszewska, Wioletta, Al Ansi, Amin N., Fekrat, Lida, Haghghadam, Zahra Mojib, Khormizi, Mehdi Zare, Noorinahad, Sara, ��enal, Derya, Shakarami, Jahanshir & Haelewaters, Danny, 2019, The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East region, pp. 163-170 in Acta Entomologica Musei Nationalis Pragae 59 (1) on pages 166-167, DOI: 10.2478/aemnp-2019-0014, http://zenodo.org/record/4552749, {"references":["PONTOPPIDAN E. 1763: Den Danske Atlas, eller Konge-Riget Dannemark. Efter HOy-Kongelig allernaadigst Befalning. Tomus I. A. H. Godiche, Kopenhagen, 723 pp.","BIELAWSKI R. 1963: Coccinellidae (Coleoptera) iz Bliznego Vostoka. Fragmenta Faunistica (Warszawa) 10: 391 - 398.","BIELAWSKI R. 1980: Die Marienkafer (Coleoptera: Coccinellidae) der Koreanischen Volksdemokratischen Republik. Fragmenta Faunistica (Warszawa) 25: 221 - 254.","VANDENBERG N. J. 1990: First North American records for Harmonia quadripunctata (Pontopiddian) (Coleoptera: Coccinellidae), a lady beetle native to the Palaearctic. Proceedings of the Entomological Society of Washington 92: 407 - 410.","PANG H., PANG X. F., REN S. X. & ZENG T. 2004: Biodiversity and their utilization of Coccinellidae in China. Guangdong Science and Technology Press, Guangzhou, 168 pp.","KOVAR I. 2007: Family Coccinellidae Latreille 1807. Pp. 568 - 631. In: LOBL I. & SMETANAA. (eds): Catalogue of Palaearctic Coleoptera. Volume 4. Elateroidea, Derodontoidea, Bostrichoidea, Lymexyloidea, Cleroidea, Cucujoidea. Apollo Books, Stenstrup, 935 pp.","KHALIL N. & MOURAD A. R. 2006: A preliminary survey of ladybirds (Coccinellidae) on coniferales in south of Syria. Bassel Al-Assad Journal for Engineering Sciences 22: 147 - 165.","HALPERIN J., MERKL O. & KEHAT M. 1995: An annotated list of the Coccinellidae (Coleoptera) of Israel and adjacent areas. Phytoparasitica 23: 127 - 137.","GUNTHER V. 1958: Ergebnisse der zoologischen Expedition des National Museums in Prag nach der Turkei. 22. Coleoptera Coccinellidae. Acta Entomologica Musei Nationalis Pragae 32: 19 - 36.","GONZALEZ G. 2006: Los Coccinellidae de Chile. Permanent electronic publication available at http: // www. coccinellidae. cl (Accessed on 23 December 2017).","UYGUN N. 1981: Taksonomische Untersuchungen uber die Coccinelliden Fauna (Coleoptera) der Turkei. Cukurova University, Faculty of Agriculture Res. Publ. no. 157 / 48. Adana, Turkey.","ULGENTURK S. & TOROS S. 2000: Preliminary studies on parasitoids and predators of Diaspididae (Homoptera: Coccoidea) species on park plants. Journal of Agricultural Science 6: 106 - 110.","SAHBAZ A. & UYSAL M. 2006: The predators and parasitoids of the aphid species (Homoptera: Aphididae) on poplars in Konya province of Turkey. Selcuk Journal of Agriculture and Food Sciences 20: 119 - 125.","ASLAN B. & KARACA I. 2005: Fruit tree aphids and their natural enemies in Isparta region, Turkey. Journal of Pest Science 78: 227 - 229.","KARACA I., KARSAVURAN Y., AVCI M., DEMIROZER O., ASLAN B., SOKELI E. & BULUT H. S. 2006: Faunistic studies on Coleoptera species of Isparta province (Turkey). Suleyman Demirel University, Journal of Science and Technology Institute 10: 180 - 184.","KAYA BASAR M. & YASAR B. 2011: Determination of ladybird species (Coleoptera: Coccinellidae) on fruit trees in Isparta, Turkey. Turkish Journal of Entomology 35: 519 - 534.","OGUZOGLU S. 2017. Cinara cedri'nin Isparta Orman Bolge Mudurlu g u Sedir Ormanlarinda Yayili s i, Zarari, Biyolojisi ve Do g al Du s manlari. [Distribution, damage, biology and natural enemies of Cinara cedri in Isparta forest regional directorate cedar forests]. Thesis deposited at Department of Forest Engineering, Suleyman Demirel University, Isparta, 116 pp.","KILIC M. & AYKAC M. K. 1989: Karadeniz bolgesi seftali bahcelerindeki zararlilarla mucadelenin yonetimi uzerinde arastirmalar. [Investigations on the pest management in peach orchards in the black sea region of Turkey]. Bitki Koruma Bulteni 29: 211 - 241 (in Turkish, English summary).","OZBEK H. & CETIN G. 1991: Contribution to the fauna Coccinellidae (Coleoptera) from eastern Anatolia along with some new records from Turkey. Turkish Journal of Entomology 15: 193 - 202.","TEZCAN S. & UYGUN N. 2003: Evaluation of the Coccinellidae (Coleoptera) fauna of ecologically managed cherry orchards in Izmir and Manisa provinces of Turkey. Turkish Journal of Entomolgy 27: 73 - 79.","ASLAN M. M. & UYGUN N. 2005: The aphidophagous coccinellid (Coleoptera: Coccinellidae) species in Kahramanmaras, Turkey. Turkish Journal of Zoology 29: 1 - 8.","VURAL VARLI S., SAKIN G. & ONCUL ABACIGIL T. 2013: The Coccinellidae (Coleoptera) species collected by different trap methods in olive orchards of Edremit Bay's (Balikesir / Turkey). Turkish Bulletin of Entomology 3: 151 - 160.","PORTAKALDALI M. & SATAR S. 2010: Research on Coccinellidae (Coleoptera) fauna in Artvin and Rize province. Bulletin of Plant Protection 50: 89 - 99.","BASTUG G. & KASAP I. 2015: Faunistic studies on Coccinellidae (Coleoptera) family in the province of Canakkale. Turkish Journal of Biological Control 6: 41 - 50.","TOPER KAYGIN A. & SOBUTAY KAPTAN U. 2017: Coccinellidae (Insecta: Coleoptera) species of Bartin Province. Journal of Bartin Faculty of Forestry 19: 227 - 236.","TIFTIKCI P. 2017: Studies on Coccinellidae species identified on sugar beet production areas in Yozgat province. Journal of Natural and Applied Sciences 33: 79 - 90 (in Turkish, English abstract).","OZTEMIZ S. & YAYLA S. 2018: Two new species of Harmonia (Coleoptera: Coccinellidae) from Duzce, Turkey. Munis Entomology & Zoology 13: 318 - 322.","BIRANVAND A., NEDVED O., SHAKARAMI J., FEKRAT L., HAMI- DI E., VOLF M. & HANLEY G. A. 2017: The ladybeetle community (Coleoptera: Coccinellidae) in North East of Iran. Baltic Journal of Coleopterology 17: 49 - 67.","PAHLAVAN YALI K., PASHAI RAD S., KHORMIZI M. Z., HAGH- GHADAM Z. M., HEIDARI LATIBARI M., HANLY G. 2017: Research on Coccinellidae (Coleoptera) fauna in Mazandarn province, Iran. Journal of Biological Control 31: 123 - 127."]}

Harmonia Mulsant, 1846 = Leis Mulsant, 1850; Ballia Mulsant, 1853; Callineda Crotch, 1871; Stictoleis Crotch, 1874; Ptychanatis Crotch, 1874; Rhopaloneda Timberlake, 1943 (sensu ŚLIPIŃSKI 2007). Diagnosis. Length 4���8 mm. Elytra moderately to strongly convex, glabrous; elytral color pattern variable. Clypeus anteriorly straight between lateral projections. Antenna composed of 11 antennomeres, slightly shorter than the width of head capsule, with club moderately compact, composed of three antennomeres (Fig. 9). Terminal maxillary palpomere securiform (Fig. 11). Pronotal disc evenly convex, transverse with narrowly upturned and/or slightly thickenedexternal borders. Prothoracichypomeronwithout fovea near anterior angles; prosternal process without or withdistinctlateral carinae extendinganteriorly (Figs 6���7). Anteriormarginof mesoventriteweaklyfoveate mediallyto straight. Elytral lateral marginsnarrowly raised; epipleuron not foveate. Apices of middle and hindtibiae without spurs (Fig. 8). Abdominalpostcoxallinesnot recurved, incomplete laterally (Fig. 12). Penis stout, consists of basal sclerite and additionalapical piece. Ovipositor withcoxitesclub-handle type; infundibulum present; spermatheca C-shaped., Published as part of Biranvand, Amir, Nedv��d, Old��ich, Tomaszewska, Wioletta, Al Ansi, Amin N., Fekrat, Lida, Haghghadam, Zahra Mojib, Khormizi, Mehdi Zare, Noorinahad, Sara, ��enal, Derya, Shakarami, Jahanshir & Haelewaters, Danny, 2019, The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East region, pp. 163-170 in Acta Entomologica Musei Nationalis Pragae 59 (1) on page 164, DOI: 10.2478/aemnp-2019-0014, http://zenodo.org/record/4552749, {"references":["SLIPINSKI A. 2007: Australian ladybird beetles (Coleoptera: Coccinellidae): their biology and classification. Australian Biological Resources Study, Canberra, 286 pp."]}

Neoheegeria astragali sp.n. Female microptera. Body and legs dark brown (Fig. 1), fore tibia and fore tarsi yellow, mid and hind tarsi paler than their tibia, antennal segments III���V light brown, VI darker (Fig. 5), major setae including sub-basal wing setae light brown; fore wing clear, weakly shaded around sub-basal setae (Fig. 2). Head longer than wide, weakly transversely striate but without sculpture between ocelli; post ocular setae finely acute, extending beyond hind margin of eye; maxillary stylets long and slender, retracted to eyes (Fig. 11) (in a few specimens including holotype this is retracted to basal half of head length (Fig. 12)), almost one third of head width apart, maxillary bridge well developed; mouth cone relatively pointed. Antennae 8-segmented, segment III and IV with 3 and 4 sense cones respectively; segment VIII narrowed at base but not constricted (Fig. 5). Pronotum with 5 pairs of well-developed pointed major setae (anteromarginals, anteroangulars, midlaterals, epimerals, and posteroangulars). Epimeral sutures complete (Fig. 11). Prosternal basantra and ferna developed, ferna wider than long. Fore tarsal tooth absent or a minute fore tarsal tooth present (Figs 3, 8). Mesopraesternum weakly joined medially. The mesonotum has a short postero-median cleft. Metanotum very weakly sculptured, median setae arising on anterior half of sclerite (Fig. 14). Metathoracic sternopleural sutures extend posteriorly from the midcoxal cavities (Fig. 15). Fore wings with three sub-basal setae situated in a triangle. Pelta usually D- shaped, with weak sculpture (Fig. 24), campaniform sensilla present at posterior; abdominal tergites II���VII with two pairs of wing-retaining setae (Fig. 19); tergite VII with two campaniform sensilla close together (Fig. 23); segment IX with seta S1 and S2 acute, smaller than tube; tube short, length less than twice basal width (Fig. 22), shorter than head; anal setae about as long as tube or a little longer. Measurements of holotype micropterous female ( in microns): Body distended length 2344, Head length (width across cheeks) 257 (208), post ocular setae 69. Pronotum, length 160; median width 305; major setae, anteromarginal 32, anteroangulars 36, midlaterals 43, epimerals 63, posteroangulars 40. Mesonotal lateral setae 27. Fore wing, length 330; sub-basal setae S1 36, S2 40, S3 59. Tergite IX setae S1 85, S2 86. Tube length (basal width) 122 (68). Antennal segments I���VIII length 30, 52, 58, 62, 63, 56, 57, 43. Female macroptera. Colour and structure very similar to microptera. Fore wings with 7���10 duplicated cilia. Measurements of a macropterous female (in PPSU) (in microns): Body distended length 2182, Head length (width across cheeks) 234 (224), post ocular setae 63. Pronotum, length 154; median width 311; major setae, anteromarginals 30, anteroangulars 33, midlaterals 33, epimerals 63, posteroangulars 40. Mesonotal lateral setae 28. Fore wing, length 795; sub-basal setae 36, 65, 60. Tergite IX setae S1 81, S2 85. Tube length (basal width) 121 (70). Antennal segments I���VIII length 31, 46, 57, 62, 63, 54, 51, 45. Male macroptera. Generally similar to female but smaller. All antennal segments are almost brown (Fig. 13), fore tarsal tooth well developed (Fig. 4). Fore wings with 7���10 duplicated cilia. The tube is long and pseudovirga divided at the apex. Measurements of a macropterous male (in PPSU) (in microns): Body distended length 1935, Head length (width across cheeks) 221 (196), post ocular setae 63. Pronotum, length 154; median width 294; major setae, anteromarginal 27, anteroangular 30, midlateral 40, epimeral 70, posteroangular 54. Mesonotal lateral seta 23. Fore wing, length 766; sub-basal setae 51, 47, 66. Tergite IX setae S1 143, S2 45. Tube length (basal width) 149 (56). Antennal segments I���VIII length 33, 47, 59, 56, 51, 46, 40, 32. Material studied. Holotype micropterous female, IRAN, Fars province, Sepidan, Barme Firooz Mountain, Astragalus sp., 24.v.2013 (KM 1021) (in NHM). Paratypes: 9 female micropterae, 1 female, 1 male macropterae, same data as holotype (in ANIC and PPSU); 2 female micropterae, 2 female, 1 male macropterae, same place, same plant, 16.v.2014 (KM 1193) (in PPSU); 8 female micropterae, same place, same plant, 12.v.2017 (KM 1627) (in PPSU). Non type specimens: 8 female micropterae (2 in PPSU, 6 in PPFU), IRAN, Khorasan-e-Razavi, Tandooreh National Park, Tivan, same plant, 14.vi.2013, Lida Fekrat. Comments. Among Haplothripini species, the tenth abdominal segment of Neoheegeria species, the tube, is characteristically long, being at least 2 times as long as the median length of the ninth abdominal segment. The new species is thus unique among Neoheegeria species, not only in its short wings but also in having the tube scarcely 1.8 times as long as the ninth abdominal segment. Moreover, three of the five Neoheegeria species have no metathoracic sternopleural sutures, whereas micropterae and macropterae of N. astragali have these sutures long and slender. The presence of metathoracic sternopleural sutures is shared with gigantea, but astragali has smaller body size, shorter tube, and in females antennal segments III���V are clearly paler. The micropterous condition in most specimens distinguishes this new species. N. astragali is also distinct in metanotum sculpture which is almost not reticulated (Fig. 14) in contrast to other species of the genus that have weak reticulation (Fig. 16)., Published as part of Minaei, Kambiz, Fekrat, Lida & Mound, Laurence, 2018, The genus Neoheegeria with a new species from Iran exhibiting wing-dimorphism (Thysanoptera: Phlaeothripidae), pp. 563-570 in Zootaxa 4455 (3) on pages 564-565, DOI: 10.11646/zootaxa.4455.3.12, http://zenodo.org/record/1457385

Neoheegeria dalmatica Schmutz Neoheegeria dalmatica Schmutz, 1909: 344. This is the type species of the genus and is probably the most common species of Neoheegeria in Europe and Asia, including northern and southern Iran (Collins 2007; Minaei et al. 2007). The large number of dalmatica deposited in PPSU indicates that the species is collected from leaves and flowers of various species of Stachys and Phlomis [Lamiaceae]., Published as part of Minaei, Kambiz, Fekrat, Lida & Mound, Laurence, 2018, The genus Neoheegeria with a new species from Iran exhibiting wing-dimorphism (Thysanoptera: Phlaeothripidae), pp. 563-570 in Zootaxa 4455 (3) on page 565, DOI: 10.11646/zootaxa.4455.3.12, http://zenodo.org/record/1457385, {"references":["Schmutz, K. (1909) Zur kenntnis einiger neuer Thysanopterengenera und - spezies. Annalen des Naturhistorischen Museums in Wien, 23, 342 - 347.","Collins, D. W. (2007) Two species of thrips (Thysanoptera) new to Britain, Neoheegeria dalmatica Schmutz and Frankliniella pallida (Uzel), with an updated key to the British species of Frankliniella Karny. British Journal of Entomology and Natural History, 20, 241 - 248."]}

Neoheegeria persica Priesner Neoheegeria persica Priesner, 1954: 54. This species was described from a single female collected on Prangos ferulacea [Apiaceae] in Iran. Subsequently, Minaei et al. (2007) collected this species from various plants including Astragalus sp. and Phlomis sp. More recently, a series of both sexes was collected from Mentha pulegium [Lamiaceae] at Sepidan (Fars province). The only difference between persica and dalmatica is the color of antennal segments III���VI. The fustis in the ninth abdominal segment was reported in persica to be unusually elongate (Minaei et al. 2007), but this is not confirmed by the specimens collected recently on Mentha pulegium., Published as part of Minaei, Kambiz, Fekrat, Lida & Mound, Laurence, 2018, The genus Neoheegeria with a new species from Iran exhibiting wing-dimorphism (Thysanoptera: Phlaeothripidae), pp. 563-570 in Zootaxa 4455 (3) on page 568, DOI: 10.11646/zootaxa.4455.3.12, http://zenodo.org/record/1457385, {"references":["Priesner, H. (1954) On some Thysanoptera from Persia. Annals and Magazine of Natural History, 7, 49 - 57. https: // doi. org / 10.11646 / zootaxa. 3956.1.4"]}

Neoheegeria sinaitica Priesner Neoheegeria sinaitica Priesner, 1934: 278 The type specimens were collected on Verbascum sp. in Sinai, Egypt but another series was collected at the same locality on Phlomis floccosa [Lamiaceae]. Priesner (1965) concluded that the identification of Verbascum might be incorrect. The species is distinguished from other species in the genus by the presence of a ���conspicuous��� or ���rather prominent, slender��� fore tarsal tooth in females (Priesner 1934; 1965). A single female of sinaitica was collected on Astragalus sp. with the new species described above, and this is the first report of sinaitica since its description. The validity of this species remains doubtful, in the absence of further specimens from the area of the type locality., Published as part of Minaei, Kambiz, Fekrat, Lida & Mound, Laurence, 2018, The genus Neoheegeria with a new species from Iran exhibiting wing-dimorphism (Thysanoptera: Phlaeothripidae), pp. 563-570 in Zootaxa 4455 (3) on page 569, DOI: 10.11646/zootaxa.4455.3.12, http://zenodo.org/record/1457385, {"references":["Priesner, H. (1934) Contributions towards a knowledge of the Thysanoptera of Egypt, IX. Bulletin de la Societe Royal Entomologie d'Egypte, 18, 275 - 285.","Priesner, H. (1965) A Monograph of the Thysanoptera of the Egyptian Deserts. Publications de l'Institut Desert Egypte, 13, 1 - 549."]}

Key to the Iranian species of Hyperaspis 1. Pronotum black in the middle, with yellow, orange or red lateral, sometimes also front margins...................... 2 - Pronotum orange or brown with contrasting pattern........................................................ 13 2. Elytra completely black, without spots (Fig. 14); pronotum black with red lateral margins; body length 2.8–3.0 mm; penis guide almost symmetrical, pointed (Fig.1).................................................. H. concolor Suffrian - Elytra black with yellow, orange or red markings........................................................... 3 3. Single pair of red spots in posterior half of elytra (preapical).................................................. 4 - More than one pair of yellow, orange or red markings....................................................... 5 4. Body not uniformly oval, wider posteriorly; pronotum with broad red lateral margins; each elytron with reddish spot (Fig. 23); penis guide with truncate apex and large median tooth (Figs. 9, 10); body length 3.1–5.0 mm........ H. reppensis (Herbst) - Body uniformly oval; pronotum with narrow orange lateral margins; each elytron with orange spot (Fig.15); penis guide broad, shorter than parameres, with oblique apex and small median tooth (Fig. 2); body length 3.2–4.0 mm... H. duvergeri Fürsch 5. Each elytron with two rounded spots, one (discal) in anterior half, one (preapical) in posterior half.................... 6 - Elytra with more complex pattern...................................................................... 10 6. Each elytron with two yellowish spots of almost same size (Fig. 20); penis guide with asymmetric apex and small median tooth (Fig. 6); body length 2.8 mm...................................................... H. persica Duverger (male) - Discal spot smaller than preapical one, orange or red........................................................ 7 7. Penis guide symmetrical, slender, with rounded apex, without median tooth (Fig. 3)........... H. femorata (Motschulsky) - Penis guide asymmetrical, with straight or oblique apex...................................................... 8 8. Penis guide broad with squarely truncate apex and large blunt median tooth (Fig. 8); elytron with discal spot moderately large (Fig. 22).................................................................. H. quadrimaculata Redtenbacher - Penis guide with obliquely truncate apex and small median tooth; elytron with discal spot very small or absent.......... 9 9. Penis guide much shorter than parameres (Fig. 4); spots on elytra orange (Fig. 17)............ H. histeroides (Faldermann) - Penis guide almost as long as parameres (Fig. 11); spots on elytra red (Fig. 24)....................... H. syriaca Weise 10. Each elytron with three spots (anterior, discal, preapical) and narrow lateral stripe joined with at least anterior spots, yellow or orange; penis guide long, slender, with sharp median tooth.................................................. 11 - Each elytron with large irregular, red patch in the middle, often reaching lateral margin, sometimes with small black spot in center of patch; head black; body length 3.0 mm (Fig. 19)............................. H. persica Duverger (female) 11. Head brown; pronotum black with anterior and lateral margins yellowish orange; preapical spot on elytron not confluent with lateral stripe (Fig. 21); body length 2.8–3.0 mm; penis guide as long as parameres and much wider than parameres (Fig. 7).......................................................................................... H. polita Weise - Head black with yellow or yellowish brown part anteriorly and sometimes also laterally; pronotum black with yellowish area laterally; preapical spot on elytron confluent with lateral stripe; penis guide much longer and hardly wider than parameres..................................................................................................... 12 12. Head black except for yellowish brown anterior and lateral margins; elytron with discal spot separate from anterior spot/stripe (Fig. 18); body length 2.0–3.0 mm; basal part of penis guide twice as wide as distal part (Fig. 5)...................................................................................................... H. marmottani (Fairmaire) - Head black, yellow anteriorly; elytron with discal spot connected to anterior spot/stripe (Fig. 26); body length 1.8–3.2 mm... ........................................................................... H. vinciguerrae Capra (female) 13. Elytra brown, with basal, discal, lateral and apical creamy brown spots which are variously confluent and irregularly scattered; pronotum brown with creamy brown apical and lateral margins, with an anchor-like spot basomedially which is connected to the anterior margin (Fig. 25); body length 2.5 mm.............................. H. transversoguttata Weise (female) - Elytra whitish, with black stripe along suture and each elytron with black, elongate discal, comma-like stripe; pronotum orange, darker mediobasally (Fig. 27); body length 1.2–2.7 mm; penis guide slender, rounded apically, with small median tooth (Figs. 12, 13)............................................................. H. vinciguerrae Capra (male)

Mycterothrips zagrosi sp. n. (Figs 2, 4, 5, 8, 9, 12, 14) Female macroptera. Body pale brown; major body setae yellowish brown, those on head pale; antennal segment I yellow, II brown, III yellow, slightly shaded at apex, IV pale brown with base yellow, V���VIII shaded (Fig. 2); legs mainly pale, fore legs a little paler than mid- and hind legs; fore wings and scales uniformly yellowish brown; head with round brown marking between ocelli; mesonotum shaded at anterior margin and mediolaterally; metanotum shaded; abdominal tergites II���VIII with dark ridge along median area between antecostal and medial line, tergites IX���X pale brown. Head wider than long, with three pairs of ocellar setae, pair III situated between posterior ocelli, five pairs of postocular setae present, I longer than others, II is smallest (Fig. 4). Pronotum wider than long, with two pairs of long posteroangular setae and one pair of posteromarginal setae (Fig. 8), with 12���15 discal setae; mesonotum with or without one pair of campaniform sensilla anteromedially (campaniform sensilla are present in holotype), median pair of setae far from posterior margin (Fig. 5); metascutum weakly and irregularly reticulate medially, median pair of setae situated near anterior margin; meso- and metasternum with developed spinula. First vein of fore wings with a median long gap in setal row, with 8���9 basal and 2 distal setae, second vein with 7���10 setae, posterior fringe cilia wavy (Fig. 9); scale with 5 veinal and 1 discal setae. Lateral thirds of abdominal tergites and pleurotergites with ciliate microtrichia on lines of sculpture (Fig. 14); tergite II with 3 lateral marginal setae; tergites VI to VIII with S4 setae developed; tergite VIII with complete posteromarginal comb of fine and long microtrichia, the comb is distinctly wider at base (Fig. 12); tergite IX without campaniform sensilla (Fig. 12); tergite X without a median split; sternites without discal setae and pore plate; median pair of setae of VII situated at posterior margin. Ovipositor well developed. Measurements (holotype female in microns): Body length 1330. Head length (width) 112 (173), ocellar setae III length 30, postocular setae I length 13. Pronotum length (width) 115 (210), posteroangular setae I length 45, setae II length 38. Fore wings length 840. Tergite IX S1 setae length 66, S2 setae length 51, S3 setae length 78. Antennal segments III to VIII length: 46, 42, 40, 50, 14, 18. Male not known. Material studied. Holotype female, IRAN, Fars province, Shiraz, on Amygdalus scoparia, 2.v.2016 (KM 1447). Paratypes: from same locality, date and host as holotype, 1 female; same locality and plant, 5 females, 22.iv.2016 (KM 1429); same locality and plant, 1 female, 5.v.2016 (KM 1453). Non type materials: Sarvestan, 100 km west of Shiraz, same plant, 1 female, 29.iv.2016 (KM 1430). Comments. Mycterothrips z agrosi is unusual in the genus in having just one pair of setae on the posterior margin of the pronotum. However, as pointed out by Alavi et al. (2013), this character should be used with caution due to variation in one species (see also Minaei et al. 2012). Furthermore, the structure of the posteromarginal comb on tergite VIII is unique amongst Mycterothrips species, in that the microtrichia share their bases in M. zagrosi whereas in other members of the genus the comb microtrichia are fully separate from each other. Judging from Masumoto and Okajima (2006) and Alavi et al. (2013) the new species is close to the Japanese species, M. glycines as well as another Iranian species, M. sanubari, in the following character states: abdominal sternites without discal setae, abdominal tergites and laterotergites with ciliate microtrichia along lines of sculpture laterally, and antennal segment VI longer than IV. However, the new species is different from both species in having antennal segment II almost pale (vs. brown in both glycines and sanubaria), abdominal tergite II with three lateral marginal setae (instead of four in both glycines and sanubaria), tergite IX without campaniform sensilla (rarely a single one present) (vs. two pairs in both glycines and sanubaria), tergites VII���VIII with S4 welldeveloped (vs. minute in both glycines and sanubaria). M. zagrosi is also distinguished from glycines by the following characters: antennal segment VI the longest (vs. III the longest in M. glycines), pronotum with less than 20 discal setae (vs. about 40 discal setae in M. glycines), mesonotum with campaniform sensilla anteromedially (vs. absent in M. glycines) and median pair of setae on sternite VII situated at the posterior margin in contrast to in front of posterior margin in M. glycines. Etymology. The new species described here is collected in the Zagros Mountains around Shiraz., Published as part of Minaei, Kambiz, Alichi, Mahmood, Fekrat, Lida, Aleosfoor, Maryam & Alavi, Jalil, 2017, Leaf-feeding Mycterothrips: two new species from southern Iran (Thysanoptera: Thripidae), pp. 195-200 in Zootaxa 4243 (1) on pages 198-199, DOI: 10.11646/zootaxa.4243.1.11, http://zenodo.org/record/398788, {"references":["Alavi, J., Modarres Awal, M., Fekrat, L. & Minaei, K. (2013) The genus Mycterothrips (Thysanoptera: Thripidae) in Iran, with three new species. Zootaxa, 3718 (4), 345 - 356.","Minaei, K., Haftbaradaran, F. & Mound, L. A. (2012) A new Ankothrips species Thysanoptera: Melanthripidae) from Iran with unusually short setae. Zootaxa, 3552, 37 - 42.","Masumoto, M. & Okajima, S. (2006) A revision of and key to the world species of Mycterothrips Trybom (Thysanoptera, Thripidae). Zootaxa, 1261, 1 - 90."]}