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15. Folsomides centralis (Denis, 1931) Type locality. Costa Rica: Orijuaco and San José (Denis 1931: 111). Records from Colombia. Locality 1 (New record). Distribution. Pantropical (Mari-Mutt & Bellinger 1990, 1996; Mari-Mutt et al. 1997 –2023; Bellinger et al. 1996 –2023). Examined material. 1 male and 1 female on slides and 1 specimen in alcohol (UNIANDES, INPA): Locality 1, 2400 m, 28.ix.2022, manual collect in branch with moss, DRR Fernandes coll., Published as part of Cipola, Nikolas Gioia, 2023, An updated catalogue of the Collembola (Hexapoda) from Colombia and a perspective for unexplored richness, pp. 499-520 in Zootaxa 5293 (3) on page 507, DOI: 10.11646/zootaxa.5293.3.4, http://zenodo.org/record/7961407, {"references":["Denis, J. R. (1931) Collemboles de Costa Rica avec une contribution au species de l'ordre. Contributo alla conoscenza de Microgenton di Costa Rica, 2, 69 - 170.","Mari-Mutt, J. A. & Bellinger, P. F. (1990) A catalog of the Neotropical Collembola: Including Nearctic areas of Mexico. Sandhill Crane Press, Gainsville, Florida, 237 pp.","Mari-Mutt, J. A. & Bellinger, P. F. (1996) Supplement to the catalog of the Neotropical Collembola. Caribbean Journal of Science, 32, 166 - 175.","Mari-Mutt, J. A., Bellinger, P. F. & Janssens, F. (1997 - 2023) Checklist of the Collembola: Supplement to the Catalog of the Neotropical Collembola. Available from: http: // www. collembola. org / publicat / neotrcat. htm (accessed 18 September 2022)","Bellinger, P. F., Christiansen, K. A. & Janssens, F. (1996 - 2023) Checklist of the Collembola of the World. Available from: http: // www. collembola. org. (accessed 9 March 2022)"]}

Folsomides subvinosus Arbea & Kahrarian, 2015 – Folsomides subvinosus Arbea & Kahrarian, 2015 a: 285. KS: Arbea & Kahrarian 2015a, b. Ecology and habitat in Iran: soil and leaf litter in oak forest and grassland. General distribution: species endemic to Iran (Arbea & Kahrarian 2015a,b)., Published as part of Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1) on page 51, DOI: 10.11646/zootaxa.5275.1.1, http://zenodo.org/record/7898948, {"references":["Kahrarian, M. (2015) Study on population dynamics, biodiversity, fauna and habitat suitability modeling of springtails (Apterygota: Collembola) in Kermanshah provience. Ph. D. Thesis, Islamic Azad University, Arak, 91 pp. [in Persian with English summary]"]}

Folsomides marchicus (Frenzel, 1941) – Proisotoma marchica Frenzel, 1941: 321. KS: Kahrarian et al. 2012; Amiri & Kahrarian 2015; Arbea & Kahrarian 2015a, b; Kahrarian 2015, 2019; LO: Moradi et al. 2018, 2019; IL: Azizi et al. 2019; Bazgir & Mirab-balou 2019; Shayanmehr et al. 2019; Heydari et al. 2021; KU: Ahmadi et al. 2023. Ecology and habitat in Iran: soil and leaf litter of oak forest and grassland. General distribution: Palaearctic (Fjellberg 2007)., Published as part of Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1) on page 51, DOI: 10.11646/zootaxa.5275.1.1, http://zenodo.org/record/7898948, {"references":["Frenzel, G. (1941) Eine neue Collembolenart: Proisotoma marchica nov. spec. Markische Tierwelt, 4, 321 - 324.","Kahrarian, M., Nikpy, A. & Mohammadi Noor, L. (2012) Preliminary checklist of the Collembolan fauna in Kermanshah, Sahneh and Harsin counties (Kermanshah: Iran) with two new records for Iranian fauna. Pakistan Entomologist, 34 (1), 27 - 30.","Kahrarian, M. (2015) Study on population dynamics, biodiversity, fauna and habitat suitability modeling of springtails (Apterygota: Collembola) in Kermanshah provience. Ph. D. Thesis, Islamic Azad University, Arak, 91 pp. [in Persian with English summary]","Moradi, T. Vafaei-Shoushtari, R., Kahrarian, M. & Mohseni-Amin, A. (2018) The study on Springtails in west part of Iran with new records for Iranian fauna. Journal of Entomological Research, 3 (10), 43 - 53.","Moradi, T. Vafaei-Shoushtari, R., Kahrarian, M. & Mohseni-Amin, A. (2019) Species Diversity of Springtails (Hexapoda: Collembola) in Four Different Ecosystems of Lorestan Province, Iran. Taxonomy and Biosystematics, 11 (41), 101 - 116. https: // doi. org / 10.22108 / tbj. 2020.118851.1097","Shayanmehr, M., Mirab-balou, M., Yoosefi Lafooraki, E. & Kaprus', I. (2019) Introduction to Springtails (Arthropoda: Hexapoda) fauna from Ilam province (western Iran) with a new record from Sminthuridae. Journal of Insect Biodiversity and Systematics, 5 (4), 399 - 410. https: // doi. org / 10.52547 / jibs. 5.4.399","Heydari, M., Eslaminejad, P., Kakhki, F. V., Mirab-balou, M., Omidipour, R., Zema, D. A., Ma, C. & Lucas-Borja, M. E. (2021) Spatio-temporal heterogeneity differently drives the diversity of various trophic guilds of mesofauna in semi-arid oak forests. Trees, 35, 171 - 187. https: // doi. org / 10.1007 / s 00468 - 020 - 02025 - 3","Fjellberg, A. (2007) The Collembola of Fennoscandia and Denmark. Part II: Entomobryomorpha and Symphypleona. Fauna Entomologica Scandinavica, 42, 1 - 264. https: // doi. org / 10.1163 / ej. 9789004157705. i- 265"]}

Folsomides angularis (Axelson, 1905) – Isotoma angularis Axelson, 1905: 791. GI: Daghighi 2012; Daghighi et al. 2013a, b; Yoosefi Lafooraki et al. 2020b; KU: Ahmadi 2021; Ahmadi et al. 2023. Ecology and habitat in Iran: soil. General distribution: probably widely distributed in the Holarctic (Potapov 2001)., Published as part of Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1) on page 50, DOI: 10.11646/zootaxa.5275.1.1, http://zenodo.org/record/7898948, {"references":["Daghighi, E. (2012) Fauna of Collembola (Insecta: Apterygota) from Rasht and its regions. M. Sc. Thesis, University of Guilan, Guilan, 97 pp. [in Persian with English summary]","Daghighi, E., Hajizadeh, J., Hosseini, R. & Moravvej, A. (2013 a) A checklist of Iranian Collembola with six new records from family Isotomidae (Collembola: Isotomidae). Entomofauna, 11, 149 - 156.","Yoosefi Lafooraki, E., Hajizadeh, J., Shayanmehr, M. & Hosseini, R. (2020 b) Isotomidae (Collembola) from northern Iran with description of a new species of Isotomodes Linnaniemi. Journal of Asia-Pacific Biodiversity, 13, 545 - 553. https: // doi. org / 10.1016 / j. japb. 2020.08.012","Potapov, M. (2001) Isotomidae. In: Dunger, W. (Ed.), Synopses on Paleartic Collembola. Vol. 3. Abdhandlungen und Berichte des Naturkundemuseums G ˆ rlitz, 73 (2), pp. 1 - 603."]}

Folsomides halshinicus Arbea & Kahrarian, 2015 – Folsomides halshinicus Arbea & Kahrarian, 2015 a: 283. KS: Arbea & Kahrarian 2015a, b; KU: Ahmadi 2021; Ahmadi et al. 2023. Ecology and habitat in Iran: soil and leaf litter of oak forest and grassland. General distribution: species possibly endemic to Iran (Arbea & Kahrarian 2015a)., Published as part of Mayvan, Mahmood Mehrafrooz, Greenslade, Penelope & Sadeghi-Namaghi, Hussein, 2023, An annotated checklist of the Collembola (Hexapoda) from Iran, pp. 1-101 in Zootaxa 5275 (1) on page 51, DOI: 10.11646/zootaxa.5275.1.1, http://zenodo.org/record/7898948, {"references":["Kahrarian, M. (2015) Study on population dynamics, biodiversity, fauna and habitat suitability modeling of springtails (Apterygota: Collembola) in Kermanshah provience. Ph. D. Thesis, Islamic Azad University, Arak, 91 pp. [in Persian with English summary]"]}

222. Folsomides centralis (Denis, 1931) Proisotoma centralis Denis, 1931: 111. Folsomides centralis (Denis): Yoshii & Suhardjono 1992: 11. Distribution in Vietnam. Central Highlands: Lâm Đồng (Bidoup Núi Bà national park). Southeast: Đồng Nai (Cát Tiên national park). Mekong River Delta: Kiên Giang (Hòn Chông). Authorship. EC: Anichkin et al. 2007; Shveenkova 2011. CI: Deharveng et al. 2009; Deharveng et al. 2020. Type localities. Orijuaco and San José (Costa Rica). General distribution. Pantropical species., Published as part of Nguyen, Anh T. T., Thuy, Kim N. & Arbea, Javier I., 2022, Catalogue of Vietnamese springtails (Hexapoda, Collembola), pp. 1-177 in Zootaxa 5147 (1) on page 94, DOI: 10.11646/zootaxa.5147.1.1, http://zenodo.org/record/6606048, {"references":["Denis, J. R. (1931) Collemboles de Costa Rica avec une contribution au species de l'ordre. Bollettino del Laboratorio di zoologia generale e agraria della R. Scuola superiore d'agricoltura in Portici, 25, 69 - 170.","Yoshii, R. & Suhardjono, Y. R. (1992) Collembolan Fauna of Indonesia and its Vicinities II: Collembola of Irian Java and Maluku Island. Azao, Acta Zoologica Asiae Orientalis, 2, 1 - 52.","Anichkin, A. E., Belyaeva, N. V., Dovgobrod, I. G., Shveenkova, Y. B. & Tiunov, A. V. (2007) Soil Microarthropods and Macrofauna in Monsoon Tropical Forest of Cat Tien and Bi Dup-Nui Ba national Parks, Southern Vietnam. Biology Bulletin, 34 (5), 498 - 506. [♦] https: // doi. org / 10.1134 / S 1062359007050135","Shveenkova, Y. B. (2011) Springtail communities (Collembola, Hexapoda). In: Tiunov A. V. (Ed.), Structure and Functions of Soil Communities of a Monsoon Tropical Forest (Cat Tien National Park, southern Vietnam). KMK Scientific Press, Moscow, 131 - 147. [♦]","Deharveng, L., Bedos, A., Le, C. K., Le, C. M. & Tryong, Q. T. (2009) Endemic arthropods of the Hon Chong hills (Kien Giang), an unrivaled biodiversity heritage in Southeast Asia. In: Le, C. K., Truong, Q. T. & Ly, N. S. (Eds.), Beleaguered hills: managing the biodiversity of the remaining karst hills of Kien Giang, Vietnam. Nha Xuat Ban Nong Nghiep, Ho Chi Minh City, Vietnam, 31 - 57. [♦]","Deharveng, L., Bedos, A. & Lukic, M. (2020). The genus Folsomides in Hon Chong hills, Vietnam (Collembola: Isotomidae). Raffles Bulletin of Zoology, 35, 32 - 47. [♦]"]}

Folsomides portucalensis da Gama, 1961 sensu Fjellberg 1993 Folsomides variabilis ssp. portucalensis da Gama, 1961: 24. Subisotoma variabilis psammophila Loksa & Vogojević, 1970: 128. MATERIAL EXAMINED. — Spain • 2 ♀; SSD-7, slides 07 and 10; Ortuño et al. leg.; MZNA. REMARK Only two specimens have appeared and in a single trap. The species seems to have a European distribution (de Jong et al. 2014)., Published as part of Baquero, Enrique, Jordana, Rafael & Ortuño, Vicente M., 2021, Distinctive Collembola Communities in the Mesovoid Shallow Substratum: Entomobryomorpha of the Sierra de Guadarrama National Park (Central Spain), pp. 37-78 in Zoosystema 43 (3) on page 46, DOI: 10.5252/zoosystema2021v43a3, http://zenodo.org/record/4487162, {"references":["GAMA M. M. 1961. - Nouvelle contribution pour l'etude des Collemboles du Portugal continental. Memorias e estudos do Museu zoologico da Universidade de Coimbra 269: 1 - 43.","FJELLBERG A. 1993. - Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavia 23: 453 - 473.","LOKSA I. & BOGOJEVIC J. 1970. - Einige interessante Collembolen- Arten aus der Sandwuste von Deliblat, Jugoslawien. Opuscula Zoologica, X, 1: 125 - 142.","DE JONG Y. ET AL. 2014. - Fauna Europaeaall European animal species on the web. Biodiversity Data Journal 2: e 4034. https: // doi. org / 10.3897 / BDJ. 2. e 4034."]}

Folsomides cariocus sp. nov. Figs 2 ̅13 Type material. Holotype: Female: mounted on slide Nº 334 CM / MNRJ, Southeast Brazil, City of Rio de Janeiro, 30.V.2019, M.C. de Mendonça, G. Queiroz, T. Silveira, and A.C. Neves coll. Paratypes 1 specimen on slides Nº 225 CM / MNRJ, 18.II.2019; 1 specimen on slide Nº 228a CM / MNRJ, 18.II.2019; 5 specimens on slide Nº 228b CM / MNRJ, 18. II.2019, M.C. de Mendonça, G. Queiroz, T. Silveira, and A.C. Neves coll., at the same locality. All specimens were deposited in the Museu Nacional / Universidade Federal do Rio de Janeiro, Brazil. Type locality: Bed with compacted clay soil covered by undergrowth and no litter, where bee Epicharis analis Lepeletier, 1841 (Apidae) make roles for reproduction at Botanical Garden, State of Rio de Janeiro, Brazil (Fig. 1). Coordinates: 21°42’37.20’’S; 43°53’46.36’’O. Description. Total holotype body length: 0,84 mm. Paratypes body length: 0,62mm– 0,82 mm. Habitus slender and cylindrical, typical of the genus. Color of body white. Cuticle smooth, with primary granulation. Tergites of thorax and abdomen with 3,3/2,2,2,2,4 S-chaetae and 1,1/0,0,1 S-microchaetae formula. Eyes missing. PAO long and narrow (22 µm) with slight constriction in the anterior middle part and three chaetae on the posterior region. Close to the postantennal organ, one semicircular area without convexity on external part, without pigment and with a finer texture than the surrounding area (Figs. 2) Cephalic chaetae subequal, the anterior and posterior are a little longer (Fig. 3) Antennae slightly shorter than the head. Ratio antennae: head: = 1:1,4. Ant. IV with one subapical small organite on dorsal side, protected by curved chaeta and about eight sensilla (Fig. 4). Ant III with about 18 ordinary chaetae, two S-chaetae S3 and S4 (3µm); two guard S-chaetae S2 and S5 (7 µm) and with one short dorso-lateral sensillum S1 (3µm). Ant. II with about 16 ordinary chaetae, one ventro-external sensillum (5 µm) distally and 2 basal microchaetae, 1 dorsal and 1 ventral (Fig. 5). Ant. I with 11 ordinary chaetae, two subequal sensilla, one thicker (7µm) and one thinner (5 µm) on ventral side and two basal microchaetae, one dorsal and one ventral (Fig. 6). Ratio antennal segments: I: II: III: IV = 1: 1,5: 1,6: 2,7. Labral chaetae formula 2/5,5,4, anterior row inserted on papillae. Maxillary outer lobe with bifurcated palp and three sublobal hairs (Fig. 7). Labium with 3 proximal, 4 basomedial and 5 basolateral chaetae. Labial palps complete. Ventral head with 3+3 post-labial chaetae. Body chaetae smooth and subequal; some longer laterally. Th II with 5+5 axial mesochaetae, 1+1 lateral macrochaetae (32 µm); 2+2 lateral largest sensilla, (11 µm), 1+1 smaller (5 µm) and 1+1 dorsal sensilla (12 µm); Th. III with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (30 µm), 2+2 lateral sensilla (11 µm) and 1+1 dorsal sensilla (12 µm) Thorax without ventral chaetae. Abd. I with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (30 µm) little longer than dorsal chaetae (29 µm), 1+1 lateral sensilla and 1+1 dorsal sensilla (12 µm) (Fig. 8). Abd. II with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (32 µm), 1+1 lateral sensilla situated almost on ventral side (11 µm) and 1+1 dorsal sensilla (12 µm). Abd. III with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (30 µm), 1+1 dorsal sensilla (12 µm), 1+1 lateral sensilla (12 µm) and 1+1 lateral microsensilla (5 µm) both situated on latero-ventral side. Abd. IV with 4+4 axial mesochaetae, 1+1 lateral macrochaetae (31µm) and 1+1 dorsal macrochaetae (29 µm), 1+1 dorsal sensilla (15 µm) and 1+1 lateral sensilla situated on latero-ventral side (12 µm). Dorsal sensilla on Abd I̅IV situated in mid-tergal position and dorsal sensilla on Abd. IV situated behind to the dorsal macrochaeta. Abd. V with 2+2 axial mesochaetae and one unpaired axial mesochaeta, 1+1 lateral macrochaeta (34 µm) and 1+1 dorsal macrochaeta (34 µm), 3+3 dorsal sensilla (20 µm) and 1+1 ventral (18 µm). Abd. VI some macrochaetae (35–40 µm) distributed among few mesochaetae, three unpaired chaetae (a0: 30 µm: m0: 38 µm: p0: 20 µm) (Fig. 9). Appendages. Claws simple, without teeth (18 µm). Unguiculi reduced (4 µm) with a basal lamella and a short apical filament. Tita I, II, III with 20, 20, 21–22 chaetae, Femora I, II, III with 11, 11, 13 chaetae. Trochanter I, II, III with 8, 8, 6 chaetae. Tita I and II with unpaired chaetae B 4/5. Tita III with 7 chaetae in distal whorl with undifferentiated tenent chaetae (Fig. 10, 11). Ventral tube with 3+3 laterodistal and two posterior chaetae. Tenaculum with 3+3 teeth and no chaetae on corpus. Anterior furcal subcoxae with 5 chaetae, one longer than others and posterior subcoxae with 3 chaetae, one longer than others (Fig. 12). Manubrium (60µm) with 7+7 chaetae on posterior side and no chaeta on anterior side. Dens (30µm) with 3+3 anterior chaetae and no chaetae on posterior side. Mucro (10µm) bidentate, separated from dens (Fig.13). Ratio manubrium:dens:mucro as 1:3:6. Females genital plates with 1+1 microchaetae on anterior region. Etymology. The name of the new species cariocus derives from the word ‘’carioca’’ originated from the Tupi indigenous language that means house of white man and actually refers to people born in the city of Rio de Janeiro.

Folsomides cariocus sp. nov. Figs 2 ��13 Type material. Holotype: Female: mounted on slide N�� 334 CM / MNRJ, Southeast Brazil, City of Rio de Janeiro, 30.V.2019, M.C. de Mendon��a, G. Queiroz, T. Silveira, and A.C. Neves coll. Paratypes 1 specimen on slides N�� 225 CM / MNRJ, 18.II.2019; 1 specimen on slide N�� 228a CM / MNRJ, 18.II.2019; 5 specimens on slide N�� 228b CM / MNRJ, 18. II.2019, M.C. de Mendon��a, G. Queiroz, T. Silveira, and A.C. Neves coll., at the same locality. All specimens were deposited in the Museu Nacional / Universidade Federal do Rio de Janeiro, Brazil. Type locality: Bed with compacted clay soil covered by undergrowth and no litter, where bee Epicharis analis Lepeletier, 1841 (Apidae) make roles for reproduction at Botanical Garden, State of Rio de Janeiro, Brazil (Fig. 1). Coordinates: 21��42���37.20������S; 43��53���46.36������O. Description. Total holotype body length: 0,84 mm. Paratypes body length: 0,62mm��� 0,82 mm. Habitus slender and cylindrical, typical of the genus. Color of body white. Cuticle smooth, with primary granulation. Tergites of thorax and abdomen with 3,3/2,2,2,2,4 S-chaetae and 1,1/0,0,1 S-microchaetae formula. Eyes missing. PAO long and narrow (22 ��m) with slight constriction in the anterior middle part and three chaetae on the posterior region. Close to the postantennal organ, one semicircular area without convexity on external part, without pigment and with a finer texture than the surrounding area (Figs. 2) Cephalic chaetae subequal, the anterior and posterior are a little longer (Fig. 3) Antennae slightly shorter than the head. Ratio antennae: head: = 1:1,4. Ant. IV with one subapical small organite on dorsal side, protected by curved chaeta and about eight sensilla (Fig. 4). Ant III with about 18 ordinary chaetae, two S-chaetae S3 and S4 (3��m); two guard S-chaetae S2 and S5 (7 ��m) and with one short dorso-lateral sensillum S1 (3��m). Ant. II with about 16 ordinary chaetae, one ventro-external sensillum (5 ��m) distally and 2 basal microchaetae, 1 dorsal and 1 ventral (Fig. 5). Ant. I with 11 ordinary chaetae, two subequal sensilla, one thicker (7��m) and one thinner (5 ��m) on ventral side and two basal microchaetae, one dorsal and one ventral (Fig. 6). Ratio antennal segments: I: II: III: IV = 1: 1,5: 1,6: 2,7. Labral chaetae formula 2/5,5,4, anterior row inserted on papillae. Maxillary outer lobe with bifurcated palp and three sublobal hairs (Fig. 7). Labium with 3 proximal, 4 basomedial and 5 basolateral chaetae. Labial palps complete. Ventral head with 3+3 post-labial chaetae. Body chaetae smooth and subequal; some longer laterally. Th II with 5+5 axial mesochaetae, 1+1 lateral macrochaetae (32 ��m); 2+2 lateral largest sensilla, (11 ��m), 1+1 smaller (5 ��m) and 1+1 dorsal sensilla (12 ��m); Th. III with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (30 ��m), 2+2 lateral sensilla (11 ��m) and 1+1 dorsal sensilla (12 ��m) Thorax without ventral chaetae. Abd. I with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (30 ��m) little longer than dorsal chaetae (29 ��m), 1+1 lateral sensilla and 1+1 dorsal sensilla (12 ��m) (Fig. 8). Abd. II with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (32 ��m), 1+1 lateral sensilla situated almost on ventral side (11 ��m) and 1+1 dorsal sensilla (12 ��m). Abd. III with 3+3 axial mesochaetae, 1+1 lateral macrochaetae (30 ��m), 1+1 dorsal sensilla (12 ��m), 1+1 lateral sensilla (12 ��m) and 1+1 lateral microsensilla (5 ��m) both situated on latero-ventral side. Abd. IV with 4+4 axial mesochaetae, 1+1 lateral macrochaetae (31��m) and 1+1 dorsal macrochaetae (29 ��m), 1+1 dorsal sensilla (15 ��m) and 1+1 lateral sensilla situated on latero-ventral side (12 ��m). Dorsal sensilla on Abd I��IV situated in mid-tergal position and dorsal sensilla on Abd. IV situated behind to the dorsal macrochaeta. Abd. V with 2+2 axial mesochaetae and one unpaired axial mesochaeta, 1+1 lateral macrochaeta (34 ��m) and 1+1 dorsal macrochaeta (34 ��m), 3+3 dorsal sensilla (20 ��m) and 1+1 ventral (18 ��m). Abd. VI some macrochaetae (35���40 ��m) distributed among few mesochaetae, three unpaired chaetae (a0: 30 ��m: m0: 38 ��m: p0: 20 ��m) (Fig. 9). Appendages. Claws simple, without teeth (18 ��m). Unguiculi reduced (4 ��m) with a basal lamella and a short apical filament. Tita I, II, III with 20, 20, 21���22 chaetae, Femora I, II, III with 11, 11, 13 chaetae. Trochanter I, II, III with 8, 8, 6 chaetae. Tita I and II with unpaired chaetae B 4/5. Tita III with 7 chaetae in distal whorl with undifferentiated tenent chaetae (Fig. 10, 11). Ventral tube with 3+3 laterodistal and two posterior chaetae. Tenaculum with 3+3 teeth and no chaetae on corpus. Anterior furcal subcoxae with 5 chaetae, one longer than others and posterior subcoxae with 3 chaetae, one longer than others (Fig. 12). Manubrium (60��m) with 7+7 chaetae on posterior side and no chaeta on anterior side. Dens (30��m) with 3+3 anterior chaetae and no chaetae on posterior side. Mucro (10��m) bidentate, separated from dens (Fig.13). Ratio manubrium:dens:mucro as 1:3:6. Females genital plates with 1+1 microchaetae on anterior region. Etymology. The name of the new species cariocus derives from the word ������carioca������ originated from the Tupi indigenous language that means house of white man and actually refers to people born in the city of Rio de Janeiro., Published as part of Mendon��a, Maria Cleide De & Neves, Ana Carolina Da Rocha, 2020, New Brazilian species of Folsomides Stach (Collembola: Isotomidae), pp. 443-450 in Zootaxa 4896 (3) on pages 444-445, DOI: 10.11646/zootaxa.4896.3.5, http://zenodo.org/record/4384010, {"references":["Lepeletier, A. L. M. (1841) Histoire Naturelle des Insectes. Hymenopteres. Vol. 2. Roret, Paris, 680 pp."]}

The research about composition and structure of Collembola community was conducted at TKA company area, west Sumatra. Collembola were collected at oil palm plantation, conservasion forest and forest edge sites (between conservation forest and plantation area) by survey method with systematic random sampling for collembolla in litter and soil. A total of four Collembola species that belonging to 2 orders, 3 families, 4 genera and 57 individuals was collected. The density of Collembola in litter at conservation forest site was higher than density of Collembola at oil palm plantation site meanwhile the density of Collembola in soil at conservation forest site was higher than density of Collembola in the soil at palm plantation site. Relative density of Isotomiella sp. in litter at oil palm plantation site (100%) was higher than relative density of Folsomides sp. at conservation forest site (5,5%) as well as in the soil. The highest diversity index of Collembola was found in litter of conservation forest (1,18). The highest similarity index was found between conservation forest and forest edge (85,71%) meanwhile the lowest similarity index was found between conservation forest and oil palm plantation (40%).

We present the catalogue of species of Collembola of the Colecao de Referencia de Fauna de Solo at the Universidade Estadual da Paraiba (CRFS/ UEPB), including 6 holotypes of species of the Isotogastruridae, Lepidocyrtidae, Neanuridae, and Sminthuridae families. Four species are newly recorded in Brazil: Arrhopalites caecus (Tullberg, 1871) Borner, 1906; Lepidocyrtus neofasciatus Wray, 1948; Folsomides famarensis Fjellberg, 1993; Sminthurides aquaticus (Bourlet, 1842) Folsom, 1896; and 1 genus, Neelus Folsom, 1896. Resumo Apresentamos o catalogo de especies de Collembola da Colecao de Referencia de Fauna de Solo da Universidade Estadual da Paraiba (CRFS / UEPB), com seis holotipos de especies das familias Isotogastruridae, Lepidocyrtidae, Neanuridae e Sminthuridae. Quatro novos registros de especies foram feitos para o Brasil: Arrhopalites caecus (Tullberg, 1871) Borner; Lepidocyrtus neofasciatus Wray, 1948; Folsomides famarensis Fjellberg, 1993; Sminthurides aquaticus (Bourlet, 1842) Folsom, 1896; e um genero, Neelus Folsom, 1896. View this article in BioOne

F. parvulus Stach, 1922 – Folsomides parvulus Stach, 1922: 17 – Folsomides americanus Denis, 1931: 126 – Folsomides parvus Folsom in Mills, 1934: 43 NT Hammer 1953 as parvus; Christiansen & Bellinger 1980 as americanus; Skidmore 1995 as americanus; Christiansen & Bellinger 1998; Babenko & Fjellberg 2006 BC Cannings 2010 AB Berg & Pawluk 1984; Skidmore 1995; Lindo 2014 all as americanus MB Hammer 1953 as parvus; Mills & Sinha 1971; Christiansen & Bellinger 1980 as americanus; Skidmore 1995 as americanus; Christiansen & Bellinger 1998; Porco et al. 2014 ON Broadbent & Tomlin 1980 as parvus QC Therrien et al. 1999, 1999a; Chagnon et al. 2000; Rochefort 2006 General distribution: cosmopolitan., Published as part of Babenko, Anatoly, Stebaeva, Sophya & Turnbull, Matthew S., 2019, An updated checklist of Canadian and Alaskan Collembola, pp. 1-125 in Zootaxa 4592 (1) on page 58, DOI: 10.11646/zootaxa.4592.1.1, http://zenodo.org/record/2656929, {"references":["Stach, J. (1922) Apterygoten aus dem nordwestlichen Ungarn. Annales Musei Nationalis Hungarici, 19, 1 - 75.","Denis, J. R. (1931) Contributo alla conosenza del \" MICROGENTON \" di Costa Rica. II. Collemboles de Costa Rica avec une contribution au species de l'ordre. Bollettino del Laboratorio di zoologia generale e agraria della R. Scuola superiore d'agricoltura in Portici, 25, 69 - 170.","Mills, H. B. (1934) Monograph of the Collembola of Iowa. Ames, Collegiate Press Inc., 143 pp. https: // doi. org / 10.31274 / rtd- 180813 - 15077","Hammer, M. (1953) Investigations on the microfauna of Northern Canada. II. Collembola. Acta Arctica, 6, 108 pp.","Christiansen, K. & Bellinger, P. (1980) The Collembola of North America, North of the Rio Grande, 1 st edition. Part 1. Poduridae & Hypogastruridae, Part 2. Onychiuridae & Isotomidae, Part 3. Entomobryidae. Grinnell College, Grinnell, Iowa, 1 - 1042.","Skidmore, R. E. (1995) Checklist of Collembola (Insecta: Apterygota) of Canada and Alaska. Proceedings of the Entomological Society of Ontario, 126, 45 - 76.","Christiansen, K. A. & Bellinger, P. (1998) The Collembola of North America, North of the Rio Grande, 2 nd edition. Grinnell College, Grinnell, Iowa, 1518 pp.","Babenko, A. & Fjellberg, A. (2006) Collembola septentrionale. A catalogue of springtails of the Arctic regions. KMK Scientific Press Ltd, Moscow, 190 pp.","Cannings, R. A. (2010) Checklist of the Collembola: Nomina Collembola Canada British Columbia. Available from: http: // www. collembola. org / publicat / collbc. pdf (Accessed May 2018)","Berg, N. W. & Pawluk, S. (1984) Soil mesofaunal studies under different vegetative regimes in North Central Alberta. Canadian Journal of Soil Science, 64, 209 - 223. https: // doi. org / 10.4141 / cjss 84 - 022","Lindo, Z. (2014) Springtails (Hexapoda: Collembola) of the Prairie Grasslands of Canada. In: Carcamo, H. & Giberson, D. J. (Eds), Arthropods of Canadian Grasslands, Volume 3. Biodiversity and Systematics, Part 1. Chapter 6. Biological Survey of Canada, Ottawa, Canada, 191 - 198. https: // doi. org / 10.3752 / 9780968932162. ch 6","Mills, J. T. & Sinha, R. N. (1971) Interactions between a springtail, Hypogastrura tullbergi, and soil-borne fungi. Journal of Economic Entomology, 64 (2), 398 - 401. https: // doi. org / 10.1093 / jee / 64.2.398","Porco, D., Skarzynski, D., Decaens, T., Hebert, P. D. N. & Deharveng, L. (2014) Barcoding the Collembola of Churchill: a molecular taxonomic reassessment of species diversity in a sub-Arctic area. Molecular Ecology Resources, 14 (2), 249 - 261. https: // doi. org / 10.1111 / 1755 - 0998.12172","Broadbent, A. B. & Tomlin, A. D. (1980) Effects of carbofuran on the soil microarthropod community in a cornfield. In: Dindal, D. L. (Ed.), Soil Biology as Related to Land Use Practices. Proceedings of the VII th International Colloquium of Soil Zoology, Syracuse. Environmental Protection Agency, Washington, DC, pp. 82 - 93.","Therrien, F., Chagnon, M. & Hebert, C. (1999) Biodiversity of Collembola in sugar maple (Aceraceae) forests. The Canadian Entomologist, 131, 613 - 628. https: // doi. org / 10.4039 / Ent 131613 - 5","Therrien, F., Chagnon, M. & Hebert, C. (1999 a) Liste des especes de Collemboles du Quebec. Bulletin de l'entomofaune, 21, 8 - 11.","Chagnon, M., Hebert, Ch., & Pare, D. (2000) Community structures of Collembola in sugar maple forests: relations to humus type and seasonal trends. Pedobiologia, 44, 148 - 174. https: // doi. org / 10.1078 / S 0031 - 4056 (04) 70035 - 9","Rochefort, S. (2006) Impact de differents types d'entretien de pelouses sur l'abondance et la diversite des Arthropodes, et potentiel des graminees endophytiques dans la lutte aux insectes ravageurs. Ph. D. Thesis, Universite Laval, Quebec, 153 pp. Available from: htttp: // www. amicus. collectionscanada. gc. ca (accessed 1 May 2019)"]}

Folsomides parvulus Stach, 1922 Folsomides parvulus Stach, 1922: 17. CITATIONS FROM ALGERIA OUTSIDE COLLO. — Cassagnau (1963: 198, as Folsomides parvus Folsom, 1934); Thibaud & Massoud (1980: 517, as F. parvus); Hamra-Kroua & Cancela da Fonseca (2009: 36). DISTRIBUTION. — Cosmopolitan (Potapov 2001). REMARK Rather frequent in our samples., Published as part of Bendjaballah, Mohamed, Zoughailech, Abdelmalek, Brahim-Bounab, Hayette, Hamra-Kroua, Salah, Bedos, Anne & Deharveng, Louis, 2018, Annotated checklist of the springtails (Hexapoda: Collembola) of the Collo massif, northeastern Algeria, pp. 389-414 in Zoosystema 40 (16) on page 403, DOI: 10.5252/zoosystema2018v40a16, http://zenodo.org/record/4335964, {"references":["CASSAGNAU P. 1963. - Les Collemboles d'Afrique du Nord. Etude de quelques especes du Nord Constantinois. Bulletin de la Societe d'Histoire naturelle de Toulouse 98 (1 / 2): 197 - 206.","THIBAUD J. M. & MASSOUD Z. 1980. - Etude des Collemboles de certains milieux du Maroc et considerations biogeographiques sur la faune du Maghreb. Revue suisse de Zoologie 87 (2): 513 - 548. https: // doi. org / 10.5962 / bhl. part. 85530","POTAPOV M. 2001. - Synopses on Palaearctic Collembola, Volume 3. Isotomidae. Abhandlungen und Berichte des Naturkundemuseums 73 (2): 1 - 602."]}

Folsomides angularis (Axelson, 1905) Isotoma angularis Axelson, 1905: 791. Folsomides angularis – Kseneman 1937: 109. NEW DATA. — First record of the species for Algeria. DISTRIBUTION. — Holarctic region (Potapov 2001). REMARK Very rare in our samples, only recorded in moss from Siouane., Published as part of Bendjaballah, Mohamed, Zoughailech, Abdelmalek, Brahim-Bounab, Hayette, Hamra-Kroua, Salah, Bedos, Anne & Deharveng, Louis, 2018, Annotated checklist of the springtails (Hexapoda: Collembola) of the Collo massif, northeastern Algeria, pp. 389-414 in Zoosystema 40 (16) on page 403, DOI: 10.5252/zoosystema2018v40a16, http://zenodo.org/record/4335964, {"references":["POTAPOV M. 2001. - Synopses on Palaearctic Collembola, Volume 3. Isotomidae. Abhandlungen und Berichte des Naturkundemuseums 73 (2): 1 - 602."]}

In Nederland zijn natuurlijke en semi-natuurlijke graslanden relatief slecht onderzocht op de aanwezigheid van springstaarten. In de ons omringende landen zijn deze biotopen vaak rijk aan specifieke soorten die alleen daar voorkomen. Naar verwachting kunnen in dit milieu veel nieuwe soorten voor onze fauna gevonden worden, vooral op nutriëntarme, zandige en droge ondergrond. Recentelijk zijn twee soorten uit het genus Folsomides in graslanden in het zuiden van ons land aangetroffen. Het betreft hier een nieuw genus voor de Nederlandse fauna. In dit artikel worden het genus en de twee soorten voorgesteld, beschrijven we de habitat en geven we infor

Folsomides xerophilus Fjellberg, 1993 Folsomides xerophilus Fjellberg, 1993: 462. DISTRIBUTION. — Sud-Estde la France. Cassagnau 1961., Published as part of Thibaud, Jean-Marc, 2017, Catalogue des collemboles de France, pp. 297-436 in Zoosystema 39 (3) on page 357, DOI: 10.5252/z2017n3a1, http://zenodo.org/record/5496272, {"references":["FJELLBERG A. 1993. - Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica 23 (4): 453 - 473. https: // doi. org / 10.1163 / 187631292 X 00245","CASSAGNAU P. 1961. - Ecologie du sol dans les Pyrenees centrales. Hermann, Paris 1 - 235."]}

Folsomides angularis (Axelson, 1905) * Isotoma angularis Axelson, 1905: 791. Folsomides angularis – Arbea 2015: 413. DISTRIBUTION. — Bas-Rhin. Strasbourg (Barra & Poinsot-Balaguer 1978, 1982, 1983). — Belgnaoui & Barra 1988. Bouches-du-Rhône. Plateaudu Petit Arbois (Barra & Poinsot- Balaguer 1978 1983). — Le Petit Arbois au nord de Marseille et Cadarache (Poinsot-Balaguer & Barra 1982). Finistère. Laboratoire de Roscoff (Delamare Deboutteville 1943, Proisotoma angularis). Haute-Garonne. Toulouse (Cassagnau 1954d, 1962a, P. angularis). — Toulouse, jardin botanique (Poinsot-Balaguer & Barra 1982). Pyrénées-Atlantiques. Col du Somport (Cassagnau 1954b). Pyrénées-Orientales. Mas Cournette (Poinsot-Balaguer & Barra 1982). Sud-Ouestdela France. Cassagnau 1961 (Subisotoma angularis). Tarn, Aude. Montagne Noire (Cassagnau 1965). — Bonnet etal. 1972. Tarn. Sidobre (Izarra 1969, S. angularis). Var. Massif des Maures (Poinsot-Balaguer & Kabakibi 1987). TAXONOMIE. — Poinsot-Balaguer & Barra 1982 (redescription). ÉCOLOGIE. — Cassagnau 1965 (écologie, sol); Bonnet etal. 1972 (écologie, muscicoles); Poinsot-Balaguer & Kabakibi 1987. PHYSIOLOGIE. — Barra & Poinsot-Balaguer 1978, 1983 (anhydrobiose)., Published as part of Thibaud, Jean-Marc, 2017, Catalogue des collemboles de France, pp. 297-436 in Zoosystema 39 (3) on page 356, DOI: 10.5252/z2017n3a1, http://zenodo.org/record/5496272, {"references":["AXELSON W. M. 1905 a. - Zur Kenntnis der Apterygotenfauna von Tvarminne. Festschrift Palmen 15: 1 - 46.","ARBEA J. I. 2015. - Revision de las especies del grupo de Folsomides angularis (Axelson, 1905), con description de dos nuevas especies para la fauna europea y del norte, de Africa (Collembola, Isotomidae). Arquivos Entomoloxicos 13: 413 - 428.","BELGNAOUI S. & BARRA J. - A. 1988. - Anhydrobiosis in the Collembola Folsomides angularis. Cytochemical investigations of glycogen status and glycogen phosphorylase activity. Biological Structures and Morphogenesis 1: 147 - 153.","POINSOT- BALAGUER N. 1978. - Contribution a l'etude des Collemboles de Corse (2 e note). Annales de la Societe entomologique de France 14: 185 - 190.","DELAMARE DEBOUTTEVILLE CL. 1943. - Note faunistique sur les Collemboles de France. Bulletin de la Societe entomologique de France 10: 149 - 152.","CASSAGNAU P. 1954 d. - Collemboles de France et d'Espagne. I. Isotomidae. Vie et Milieu 4: 613 - 624.","CASSAGNAU P. 1962 a. - Faune francaise des Collemboles. XII: Les Collemboles des jardins de Toulouse. Bulletin de la Societe d'Histoire naturelle de Toulouse 97: 226 - 232.","CASSAGNAU P. 1954 b. - Faune francaise des Collemboles. (V). Collemboles recoltes en Corse par Miss T. Clay. The Proceedings of the Royal Entomological Society of London 23: 239 - 241.","CASSAGNAU P. 1961. - Ecologie du sol dans les Pyrenees centrales. Hermann, Paris 1 - 235.","CASSAGNAU P. 1965. - Ecologie edaphique de la Montagne Noire basee sur les groupements de Collemboles. Revue d'Ecologie et de Biologie du sol 2: 339 - 375.","BONNET L., CASSAGNAU P. & IZARRA D. - C. 1972. - Etude ecologique des Collemboles muscicoles du Sidobre (Tarn). III. Repartition des especes en fonction des biotopes. Bulletin de la Societe d'Histoire naturelle de Toulouse 108: 263 - 279.","IZARRA D. C. DE 1969. - Etude ecologique des Collemboles muscicoles du Sidobre (Tarn) I. - Le milieu et sa faune. Revue d'Ecologie et de Biologie du sol 6: 447 - 460."]}

Folsomides portucalensis Gama, 1961 sensu Fjellberg 1993 Folsomides variabilis portucalensis Gama, 1961: 24. Folsomides portucalensis – Fjellberg 1993: 461. DISTRIBUTION. — Bouches-du-Rhône. Provenceet Camargue (Poinsot 1974b). — Cadarache (Poinsot-Balaguer 1976d). — Plateau du Petit Arbois (Barra & Poinsot-Balaguer 1977). Hérault. Balaruc-les-Bains (Deharveng 1977, 1979). Corse. Poinsot-Balaguer 1978. CHÉTOTAXIE, MORPHO- ANATOMIE. — Deharveng 1977 (chétotaxie). ÉCOLOGIE. — Poinsot-Balaguer 1976d (irradiation). PHYSIOLOGIE. — Barra & Poinsot-Balaguer 1977 (anhydrobiose)., Published as part of Thibaud, Jean-Marc, 2017, Catalogue des collemboles de France, pp. 297-436 in Zoosystema 39 (3) on pages 356-357, DOI: 10.5252/z2017n3a1, http://zenodo.org/record/5496272, {"references":["GAMA M. M. DA 1961. - Nouvelle contribution pour l'etude des Collemboles du Portugal continental. Memorias e estudos do Museu zoologico da Universidade de Coimbra 269: 1 - 43.","FJELLBERG A. 1993. - Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica 23 (4): 453 - 473. https: // doi. org / 10.1163 / 187631292 X 00245","POINSOT N. 1974 b. - Biologie et ecologie mediterraneenne. Revue de biologie et d'ecologie mediterraneenne 1: 113 - 126.","DEHARVENG L. 1977. - Etude chetotaxique des Collemboles Isotomidae. Bulletin du Museum national d'Histoire naturelle 455: 597 - 619.","BONNET L., CASSAGNAU P. & DEHARVENG L. 1979. - Recherche d'une methodologie dans l'analyse de la rupture des equilibres biocenotiques: applications aux Collemboles edaphiques des Pyrenees. Revue d'Ecologie et de Biologie du sol 16: 373 - 401."]}

Folsomides petiti (Delamare Deboutteville, 1948) sensu Fjellberg 1993 * Subisotoma petiti Delamare Deboutteville, 1948b: 178. Folsomides petiti – Fjellberg 1993: 463. DISTRIBUTION. — Bouches-du-Rhone. Boisdes Riègesen Camargue (Delamare Deboutteville 1948b, Poinsot 1972b). — Camargue, Fiélouse (Poinsot 1972b, 1974b)., Published as part of Thibaud, Jean-Marc, 2017, Catalogue des collemboles de France, pp. 297-436 in Zoosystema 39 (3) on page 356, DOI: 10.5252/z2017n3a1, http://zenodo.org/record/5496272, {"references":["DELAMARE DEBOUTTEVILLE CL. 1948 a. - Sur la presence du genre Acherontiella Absolon dans les grottes de l'Ardeche et du Gard. Notes biospeologiques 3: 49 - 56.","FJELLBERG A. 1993. - Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica 23 (4): 453 - 473. https: // doi. org / 10.1163 / 187631292 X 00245","DELAMARE DEBOUTTEVILLE CL. 1948 b. - Contributions a l'etude biologique de la Camargue. Collemboles nouveaux du sol des Bois des Rieges. Bulletin du Museum d'Histoire naturelle de Marseille 8: 177 - 182.","POINSOT N. 1972 b. - Etude systematique et ecologique des Collemboles Isotomidae de Provence. Annales de la Societe entomologique de France 8: 669 - 691.","POINSOT N. 1974 b. - Biologie et ecologie mediterraneenne. Revue de biologie et d'ecologie mediterraneenne 1: 113 - 126."]}

Folsomides sp. Source: Deharveng & Bedos (1988). Distribution in Thailand. Surat Thani, Phang Nga (P). Habitat: cave., Published as part of Jantarit, Sopark, Bedos, Anne & Deharveng, Louis, 2016, An annotated checklist of the Collembolan fauna of Thailand, pp. 301-360 in Zootaxa 4169 (2) on page 328, DOI: 10.11646/zootaxa.4169.2.4, http://zenodo.org/record/265801, {"references":["Deharveng, L. & Bedos, A. (1988) 7 - Le sud: provinces de Ranong, Chumphon, Surat Thani et Phangnga. In: Association Pyreneenne de Speleologie (Ed.), Expeditions Thai 87 - Thai 88, Rapport Speleologique et Scientifique. Association Pyreneenne de Speleologie, Toulouse, pp. 79 - 114."]}

93. Folsomides centralis Denis, 1931 * Source: Deharveng & Bedos (1993b), Bedos (1994). Distribution in Thailand. Chiang Mai (N), no detail (P). Habitat: at low altitude in secondary forest of Doi Inthanon. Distribution outside Thailand. Pan-tropical., Published as part of Jantarit, Sopark, Bedos, Anne & Deharveng, Louis, 2016, An annotated checklist of the Collembolan fauna of Thailand, pp. 301-360 in Zootaxa 4169 (2) on page 327, DOI: 10.11646/zootaxa.4169.2.4, http://zenodo.org/record/265801, {"references":["Deharveng, L. & Bedos, A. (1993 b) Factors influencing diversity of soil Collembola in a tropical mountain forest (Doi Inthanon, Northern Thailand). In: Paoletti, M. G., Foissner, W. & Coleman, D. (Eds.), Soil Biota Nutrient Cycling and Farming Systems. Lewis, London, pp. 91 - 111."]}

94. Folsomides parvulus Stach, 1922 * Name in source. Folsomides exiguus in Deharveng & Bedos (1988), in Deharveng et al. (1989), in Deharveng & Bedos (1993b) and in Bedos (1994), F. purvus in Takeda (1981) (misspelling), F. parvus in Deharveng (1986). Source: Takeda (1981), Deharveng (1986), Deharveng & Bedos (1988), Deharveng et al. (1989), Deharveng & Bedos (1993b), Bedos (1994), Wiwatwitaya & Takeda (2005). Distribution in Thailand. Chiang Mai (N), Khon Kaen, Nakhon Ratchasima (NE), Surat Thani, Phang Nga (P). Habitat: cave in Surat Thani and Phang Nga provinces; at 700���2500 m. in Doi Inthanon forest; soil s.l. in mixed dry deciduous forest and deforested area at 800 m. in Khon Kaen province (Takeda 1981) and in dry evergreen forest in Nakhon Ratchasima province (Wiwatwitaya & Takeda 2005). Distribution outside Thailand. Cosmopolitan., Published as part of Jantarit, Sopark, Bedos, Anne & Deharveng, Louis, 2016, An annotated checklist of the Collembolan fauna of Thailand, pp. 301-360 in Zootaxa 4169 (2) on page 327, DOI: 10.11646/zootaxa.4169.2.4, http://zenodo.org/record/265801, {"references":["Deharveng, L. & Bedos, A. (1988) 7 - Le sud: provinces de Ranong, Chumphon, Surat Thani et Phangnga. In: Association Pyreneenne de Speleologie (Ed.), Expeditions Thai 87 - Thai 88, Rapport Speleologique et Scientifique. Association Pyreneenne de Speleologie, Toulouse, pp. 79 - 114.","Deharveng, L. & Bedos, A. (1993 b) Factors influencing diversity of soil Collembola in a tropical mountain forest (Doi Inthanon, Northern Thailand). In: Paoletti, M. G., Foissner, W. & Coleman, D. (Eds.), Soil Biota Nutrient Cycling and Farming Systems. Lewis, London, pp. 91 - 111.","Takeda, H. (1981) Effects of shifting cultivation on the soil mesofauna with special reference to collembolan populations in North - East Thailand. Memoirs of the College of Agriculture, Kyoto University, 118, 45 - 60.","Deharveng, L. (1986) 26 - Collemboles. In: Association Pyreneenne de Speleologie (Ed.), Expedition Thai-Maros 85, Rapport Speleologique et Scientifique. Association Pyreneenne de Speleologie, Toulouse, pp. 191 - 198.","Wiwatwitaya, D. & Takeda, H. (2005) Seasonal changes in soil arthropod abundance in the dry evergreen forest of north - east Thailand, with special reference to collembolan communities. Ecological Research, 20 (1), 59 - 70. http: // dx. doi. org / 10.1007 / s 11284 - 004 - 0013 - x"]}

92. Folsomides americanus Denis, 1931 * Name in source. Folsomides sp. in Deharveng et al. (1989). Source: Deharveng et al. (1989), Bedos (1994). Distribution in Thailand. Widespread. Habitat: at 1700���2000 m. in Doi Inthanon forest Distribution outside Thailand. Cosmopolitan. Remark. F. americanus was put in synonymy with F. parvulus by Fjellberg (1993). The two forms in our material were usually not mixed, and are therefore considered as good species., Published as part of Jantarit, Sopark, Bedos, Anne & Deharveng, Louis, 2016, An annotated checklist of the Collembolan fauna of Thailand, pp. 301-360 in Zootaxa 4169 (2) on page 327, DOI: 10.11646/zootaxa.4169.2.4, http://zenodo.org/record/265801, {"references":["Fjellberg, A. (1993) Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica, 23, 453 - 473. http: // dx. doi. org / 10.1163 / 187631292 X 00245"]}

A taxonomic revision of the genus Subisotoma Stach, 1947 is presented. Subisotoma pusilla (Schäffer, 1900), S. tenuis (Dunger, 1982), and S. asiatica (Martynova, 1970) are re-described based on type and fresh material. Eight new species of the genus are described; S. pomorskii sp. nov., S. homonomica sp. nov., S. bisensillata sp. nov., S. guzeriplica sp. nov., S. posteriomollis sp. nov., S. multisensillata sp. nov., S. cruda sp. nov., and S. erratica sp. nov., using material from Eurasia. A key to the known species of Subisotoma is given.

Abrantes, Eduardo Assis, Mendonça, Maria Cleide De (2007): New species and a new record of Isotomidae (Collembola) from the coast of Brazil. Zootaxa 1500 (1): 55-60, DOI: 10.11646/zootaxa.1500.1.2

Folsomides parvulus Stach, 1922 Material examined. Iran, Kermanshah Province, Eslam Abad-e-Gharb County, Sia Khor Village, 34 ��07��� N, 46 �� 36 ��� E, 1442 m a.s.l., grassland soil, 5. IV. 2013, Morteza Kahrarian leg., 5 females and 3 young specimens. Kermanshah County, Bilevar City, cave, V. 2013, Morteza Kahrarian leg., 4 females and 10 young specimens. Distribution. Cosmopolitan, already recorded from Iran (Cox 1982, Yahyapour 2012, Kahrarian et al. 2012, Kahrarian & Arbea 2013)., Published as part of Arbea, Javier I. & Kahrarian, Morteza, 2015, The genus Folsomides Stach (Collembola, Isotomidae) in Kermanshah Province (W Iran) with the description of two new species, pp. 281-290 in Zootaxa 3925 (2) on pages 282-283, DOI: 10.11646/zootaxa.3925.2.9, http://zenodo.org/record/241484, {"references":["Stach, J. (1922) Apterygoten aus dem Nordwestlichen Ungarn. Annales Musei Nationalis Hungarici, 19, 1 - 76.","Palacios-Vargas, J. G. & Villarreal-Rosas, J. (2013) A new Mexican species of Folsomides (Collembola: Isotomidae). Revista Mexicana de Biodiversidad, 84, 480 - 484.","Cox, P. (1982) The Collembola fauna of North and North Western Iran. Entomologist's Monthly Magazine, 118, 39 - 43.","Yahyapour, E. (2012) Faunistic study on Collembola (Insecta: Apterygota) in Sari regions. Master of Science thesis in Agriculture Entomology. Sari Agricultural Science and Natural Resources University. [total page number unkown, in Persian with English abstract]"]}

Folsomides halshinicus sp. nov. Figs 1���2, 6, 9 & 13, Tables 1, 2 & 4 Type locality. Western Iran: Sarfirooz Abdb (Halshi), Paveh County, Kermanshah Province, 34 �� 56 ' N, 46 �� 27 ' E, 1663 m a.s.l. Type material. Holotype female and 9 paratypes (5 females, 4 young specimens), Iran, Kermanshah Province, Sarfirooz Abdb (Halshi), soil and leaf litter of oak jungle, 06.II. 2013, Morteza Kahrarian leg. Holotype and 7 paratypes deposited in IAUK, 2 paratypes in JIAP. Other material, Iran, Kermanshah Province, Eslam Abad-e-Gharb County, Sia Khor Village, 34 ��07��� N, 46 �� 36 ��� E, 1442 m a.s.l., grassland soil, 14.V. 2013, Morteza Kahrarian leg., 1 female and 1 young specimen. Kermanshah County, Char zabar (Kor Kor Mountain), 34 �� 13 ��� N, 46 �� 40 ��� E, 1592 m a.s.l., soil and leaf litter of oak jungle, 15.II. 2013, Morteza Kahrarian leg., 1 female. Sar-e-pol-e-Zahab County, Patagh Area, 34 �� 25 ��� N, 46 ��01��� E, 1404 m a.s.l., soil and leaf litter of oak jungle, 14.V. 2013, Morteza Kahrarian leg., 2 females. Kangavar County, Hassam Abad Village, 34 �� 31 ��� N, 47 �� 55 ��� E, 1480 m a.s.l., Forest Plane (Button wood), 14.XII. 2012, Morteza Kahrarian leg., 9 females. Description. Size: 0.75���1.05 mm females (holotype 0.88 mm). Colour: blue greyish; pigment more concentrated in eye fields. Body long and slender (Fig 1���2). Dorsal chaetae smooth and pointed. SA macrochaeta on Abd IV 0.30���0.35, on Abd V 0.45���0.50 times the tergite length (Fig 6). Antennae shorter than cephalic diagonal (10: 12���13). Ant IV with about 10 dorsal cylindrical sensilla, a dorsoexternal microsensillum and a subapical pit organ. Sensory organ of Ant III with a pair of small microsensilla and two guard sensilla shorter than surrounding chaetae, one on each side. Another microsensillum found in a ventrolateral location. Ant III with 23 common chaetae. Ant II with a small subcylindrical ventral sensillum and 18 common chaetae, and some of the basal ones short and fine. Ant I with two subcylindrical ventral sensilla and 12 common chaetae; one of the basal chaetae smaller than the others. Five omma on each side of head. PAO long and narrow, 4.5���5.5 times as long as an omma, with two posterior chaetae (Fig 9). 2 /5,5,4 labral chaetae. Maxillary palp bifurcate with three sublobal hairs. Dorsal chaetotaxy represented in Figure 1. Axial chaetotaxy of Th II-Abd V have 6,4 /3,3,3,5,2 pairs of mesochaetae respectively; unpaired axial p-chaeta present on Abd IV. Macrochaetotaxy: 1,1 /2,2,3,3,3, M on Abd I and SA on Abd III slightly longer than surrounding chaetae (Table 2). Microsensillar chaetotaxy: 1,0/0,0,1 ms freely exposed on the body surface. Th II-Abd V with 3,3 /2,2,2,2,4 sensilla slender and shorter than surrounding chaetae. The lower pair of sensilla on Abd V is shorter and thicker than the upper pair, set in front of p-row (Fig 6). Chaetotaxy of legs I to III: subcoxae I: 1,1,3; subcoxae II: 1,5,5; coxae: 4,8,9; trochanter: 8,8,6; femora: 13,14,14; tibiotarsi: 20,20, 22 pointed chaetae respectively. Chaetae A 1 ���A 7, B 1 ���B 7, C 1 ���C 7 and x present on tibiotarsus of leg III. Tibiotarsi of legs I and II with single chaeta B 4 / 5. Claw with no teeth on inner edge. Empodial appendage sharp; its length less than half internal edge of claw. Macrochaetae Ordinary L M SA chaetae Th II 27 �� �� 18 Th III 28 �� �� 18 Abd I 31 24 (15) 15 Abd II 33 25 (16) 16 Abd III 37 30 24 19 Abd IV 33 33 31 21 Abd V 31 33 36 19 Ventral chaetotaxy represented in Figure 2. Head with 3 + 3 postlabial chaetae. Ventral tube with 3 + 3 laterodistal chaetae and 2 posterobasal ones. The retinacle with 3 + 3 teeth and one chaeta on corpus. Subcoxa of furca with 5���7 chaetae on anterior and 4 (5) on posterior part. Furca well developed. Manubrium with 8 + 8 posterior and no anterior chaetae. Dens with two posterior and no anterior chaetae (Fig 13). The mucro with two teeth, with a small lateral lamella from base to subapical tooth. Discussion. Following Fjellberg (1993), Folsomides halshinicus sp. nov. belongs to the species group of Folsomides angularis (Axelson, 1905) as it has 5 + 5 omma and dens with mucro. According to reduced number of ms on Th II-Abd III (10 /001) this species is related to the subgroup formed by 13 species. Main differences between F. halshinicus sp. nov. and these species are summarized in Table 4. The new species is most similar to F. chichinautzini. They differ in the body length and the dorsal chaetotaxy; 6 axial chaetae on Abd IV in F. chichinautzini vs 5 in the new species; 12233 macrochaetae on Abd I���V in F. chichinautzini (type 2 of Fjellberg 1993, named type 2 b by us) vs 22333 in the new species (similar to type 2 of Fjellberg 1993 but M on Abd I and SA on Abd III slightly longer than surrounding chaetae, named type 2 a by us). Etymology. The species has been named after the type locality., Published as part of Arbea, Javier I. & Kahrarian, Morteza, 2015, The genus Folsomides Stach (Collembola, Isotomidae) in Kermanshah Province (W Iran) with the description of two new species, pp. 281-290 in Zootaxa 3925 (2) on pages 283-285, DOI: 10.11646/zootaxa.3925.2.9, http://zenodo.org/record/241484, {"references":["Fjellberg, A. (1993) Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica, 23, 453 - 473.","Axelson, W. M. (1905) Einige neue Collembolen aus Finland. Zoologischer Anzeiger, 28, 788 - 794."]}

Folsomides subvinosus sp. nov. Figs 3���4, 7, 10 & 12, Tables 1, 3 & 4 Type locality. Western Iran: Sia Khor Village, Eslam Abad-e-Gharb County, Kermanshah Province, 34 ��07' N, 46 �� 38 ' E, 1442 m a.s.l. Type material. Holotype female and 7 females paratypes, Iran, Kermanshah Province, Sia Khor Village, grassland soil, 14.V. 2013, Morteza Kahrarian leg. Holotype and 5 paratypes deposited in IAUK, 2 paratypes in JIAP. Other material. Iran, Kermanshah Province, Kermanshah County, Char zabar (Kor Kor Mountain), 34 �� 13 ��� N, 46 �� 40 ��� E, 1592 m a.s.l., oak jungle soil and leaf litter, 15.II. 2013, Morteza Kahrarian leg., 12 females and 7 young specimens. Paveh County, Shabankareh Village, 34 �� 52 ��� N, 46 �� 30 ��� E, 1632 m a.s.l., oak jungle soil and leaf litter, 06.II. 2013, Morteza Kahrarian leg., 1 female. Same, but 10.V. 2013, 1 young specimen. Sar-e-pol-e-Zahab County, Patagh Area, 34 �� 25 ��� N, 46 ��01��� E, 1404 m a.s.l., oak jungle soil and leaf litter, 14.V. 2013, Morteza Kahrarian leg., 5 females and 11 young specimens. Kangavar County, Hassam Abad Village, 34 �� 31 ��� N, 47 �� 55 ��� E, 1480 m a.s.l., Forest Plane (Button wood), 14.XII. 2012, Morteza Kahrarian leg., 1 female and 1 young specimen. Description. Size: 0.55���0.80 mm females (holotype 0.75 mm). Colour: pale with diffuse pigment more concentrated in eye fields. Body long and slender (Figs 3���4). Dorsal chaetae smooth and pointed. SA macrochaeta on Abd IV 0.20���0.25, on Abd V 0.50���0.55 times the tergite length (Fig 7). Antennae shorter than cephalic diagonal (10: 12). Ant IV with some 8���10 dorsal cylindrical sensilla, a dorsoexternal microsensillum and a subapical pit organ. Sensory organ of Ant III with a pair of small microsensilla and two guard sensilla shorter than surrounding chaetae, one on each side. Another microsensillum found in a ventrolateral location. Ant III with 22 common chaetae. Ant II with a small subcylindrical ventral sensillum and 16���17 common chaetae, and some of the basal ones short and fine. Ant I with two subcylindrical ventral sensilla and 12 common chaetae; one of the basal chaetae smaller than the others. Five omma on each side of head. PAO long and narrow, 3.5 times as long as an omma, with two posterior chaetae (Fig 10). 2 /5,5,4 labral chaetae. Maxillary palp bifurcate with three sublobal chaetae. Dorsal chaetotaxy represented in Figure 3. Axial chaetotaxy of Th II���Abd V have 6,4 /3,3,3,5,2 pairs of mesochaetae respectively. Macrochaetotaxy: 1,1 /2,2,3,3,3; similar to type 2 of Fjellberg (1993), but M on Abd I and SA on Abd III slightly longer than surrounding chaetae (Table 3). Microsensillar chaetotaxy: 1,0/0,0,1 ms freely exposed on the body surface. Th II���Abd V with 3,3 /2,2,2,2,4 sensilla shorter than surrounding chaetae. The lower pair of sensilla on Abd V is longer and thicker than the upper pair, set in front of p-row (Fig 7). Macrochaetae Ordinary L M SA chaetae Th II 16 �� �� 11 Th III 17 �� �� 11 Abd I 17 16 (11) 11 Abd II 18 16 (11) 11 Abd III 18 17 16 11 Abd IV 18 18 17 11 Abd V 17 21 19 11 mucro Dens Ret. Body Max. M ms Species (1) (2) (3) pigment palp (4) (5) (1) Lateral mucronal lamella. (2) Dens: Number of posterior/anterior chaetae. (3) Retinacle: Number of teeth/basal chaetae. (4) Macrochaetae from Th II to Abd V: 1��� 33 / 33333, 2 a������ 11 / 22333 2 b��� 11 / 12233, 3 ������ 11 / 11133. (5) Number of microsensillae from Th II to Abd III. (6) SA: Ratio SA macrochaeta:Abd IV length. (7) Abd IV: Lower sensilla E���equal, L���longer or S���shorter than upper sensilla, T���thicker or N���not thicker than upper sensilla, situated in front of p-row���m or in the p-row���p. (8) B 4 / 5 chaeta on tibiotarsus I/II. Chaetotaxy of legs I to III: subcoxae I: 1,1,3; subcoxae II: 1,5,5; coxae: 4,9,9; trochanter: 8,8,6; femora: 12,14,14; tibiotarsi: 20,20, 22 pointed chaetae respectively. Chaetae A 1 ���A 7, B 1 ���B 7, C 1 ���C 7 and x present on tibiotarsus of leg III. Tibiotarsi of legs I and II with single chaeta B 4 / 5. Claw with no teeth on inner edge. Empodial appendage sharp; its length less than half internal edge of claw. Ventral chaetotaxy represented in Figure 4. Head with 3 + 3 postlabial chaetae. Ventral tube with 3 + 3 laterodistal chaetae and 2 posterobasal ones. The retinacle with 3 + 3 teeth and one chaeta on corpus. Subcoxa of furca with 6���9 chaetae on anterior and 4���5 on posterior part. Furca well developed. Manubrium with 9 + 9 posterior and no anterior chaetae. Dens with three posterior and one anterior chaeta (Fig 12). The mucro with two teeth, with a small lateral lamella from base to subapical tooth. Discussion. As the previous new species, Folsomides subvinosus sp. nov. belongs to the species group of Folsomides angularis (Axelson, 1905) and also have lost several ms (10 /001). Main differences between F. subvinosus sp. nov. and these species are summarized in Table 4. The new species is most similar to F. vinosus, but easily separated by the dorsal macrochaetotaxy; The new species has 22333 macrochaetae on Abd I���V, but F. vinosus has 11133 abdominal macrochaetae (Fig 3 vs 5); these species also differ in the shape of dorsal sensilla on Abd V, lower sensilla longer than upper ones in F. subvinosus sp. nov. and lower sensilla equal in length to upper ones in F. vinosus. Etymology. Its name reflects the similarity to F. vi no s u s., Published as part of Arbea, Javier I. & Kahrarian, Morteza, 2015, The genus Folsomides Stach (Collembola, Isotomidae) in Kermanshah Province (W Iran) with the description of two new species, pp. 281-290 in Zootaxa 3925 (2) on pages 285-290, DOI: 10.11646/zootaxa.3925.2.9, http://zenodo.org/record/241484, {"references":["Fjellberg, A. (1993) Revision of European and North African Folsomides Stach with special emphasis on the Canarian fauna (Collembola: Isotomidae). Entomologica Scandinavica, 23, 453 - 473.","Axelson, W. M. (1905) Einige neue Collembolen aus Finland. Zoologischer Anzeiger, 28, 788 - 794."]}

Folsomides parvulus Stach, 1922 Folsomides parvulus Stach, 1922: 17. MATÉRIEL ÉTUDIÉ. — Raména: 3 ex. 20.III.2007; 1 ex. 1.VI.2009; 2 ex. 26.V.2010. Vohémar: 1 ex. 5.VI.2011. Île Sainte-Marie: 1 ex. 25.III.2006. Manakara: 1 ex. 26.VI.2007 (MNHN). DISTRIBUTION. — Espèce cosmopolite, très commune sous les tropiques, présente aussi à Madagascar (Thibaud 2008a) et à Maurice (Thibaud 2008b). ÉCOLOGIE. — Espèce hémiédaphique-guanophile et psammophile. Espèce assez rare, présente dans 4 localités sur 12 de la côte est de Madagascar. Absente dans nos prélèvements sur deux ans à Antalaha., Published as part of Thibaud, Jean-Marc, 2015, Étude des collemboles des sables littoraux de la côte est de Madagascar (interstitiels terrestres), pp. 351-362 in Zoosystema 37 (2) on page 357, DOI: 10.5252/z2015n2a4, http://zenodo.org/record/5156014, {"references":["STACH J. 1922. - Apterygoten aus dem nordwestlichen Ungarn. Annales Musei Nationalis Hungarici 19: 1 - 75.","THIBAUD J. - M. 2008 a. - Les collemboles des sables littoraux de Madagascar. Annales de la Societe entomologique de France 44: 503 - 519.","THIBAUD J. - M. 2008 b. - Les collemboles des sables littoraux de l'ile Maurice. Revue francaise d'Entomologie 30: 71 - 76."]}

Folsomides centralis (Denis, 1931) Proisotoma centralis Denis, 1931: 111. MATÉRIEL ÉTUDIÉ. — Raména: 9 ex. (juv.) 20.III.2007; 11 ex. 1.VI.2009; 8 ex. 26.V.2010 (MNHN). DISTRIBUTION. — Espèce pantropicale, déjà trouvée au Sénégal en interstitiel sableux littoral et signalée à Madagascar (Thibaud 2008a). ÉCOLOGIE. — Espèce hémiédaphique-guanophile-psammophile. Espèce rare à Madagascar où elle ne fut trouvée que dans une localité sur 12. Absente dans nos prélèvements sur deux ans à Antalaha., Published as part of Thibaud, Jean-Marc, 2015, Étude des collemboles des sables littoraux de la côte est de Madagascar (interstitiels terrestres), pp. 351-362 in Zoosystema 37 (2) on page 357, DOI: 10.5252/z2015n2a4, http://zenodo.org/record/5156014, {"references":["DENIS J. R. 1931. - Contributo alla conoscenza del \" Microgenton \" di Costa Rica. Bolletino del Laboratorio di Zoologia generale ed agraria del R. Istituto superiore agrario di Portici 25: 69 - 170.","THIBAUD J. - M. 2008 a. - Les collemboles des sables littoraux de Madagascar. Annales de la Societe entomologique de France 44: 503 - 519."]}

A new species of Folsomides from Baja California Sur is described and illustrated. It is clearly differentiated from F. chichinautzini Kovác et Palacios-Vargas and F. decemoculatus Scherbakov by its smaller size, the different shape of the postantennal organ, the number of sensillae in the antennal segment IV, the number of dental setae and the body chaetotaxy. In addition a key for identification of the American species of Folsomides is provided.

The genus Folsomides Stach, 1922 is redefined and given a narrow definition based on a limited set of consistent morphological characters: Position and distribution of tergal macrosensilla, chaetotaxy of mouth region, tibiotarsi, ventral tube and furca. Eighteen new species are described from the Canary Islands: semiparvulus sp. n., xerophilus sp. n., vinosus sp. n., cumbrosus sp. n., unicus sp. n., terrus sp. n., pocosensillatus sp. n., nigrocellatus sp. n., teno sp. n., oromii sp. n., ononicolus sp. n., graminis sp. n., famarensis sp. n., pinicolus sp. n., intermedius sp. n., tonellus sp. n., supranubius sp. n. and analuisae sp. n. The following European/African species are redescribed: parvulus Stach, portucalensis Gama, angularis (Axelson), cf. petiti Delamare, lawrencei Gers & Deharveng, cf. zairensis Martynova, nanus Ellis and centralis (Denis). Highly discriminate species characters are found in maxillary palp, tibiotarsal chaetotaxy and distribution of tergal microsensilla. A number of species are removed from Folsomides, and Subisotoma Stach, 1947 is given generic rank.

Folsomides semiparvulus Fjellberg, 1993 Material examined. Brazil, Restinga de Maricá, Maricá City, Rio de Janeiro State, 5 specimens in slides Nº 1278 CM / MNRJ, 28.i.2003, 18 specimens in slides Nº 1279 CM / MNRJ, 28.i.2003, L.H. Fernandes & E.A. Abrantes. coll. Collected in sand dunes, first record to Brazil. Remarks. This species, originally described for "Punta del Hidalgo " (Teneriff), later recorded to México (Kováč & Palacios-Vargas 1996), was found together with F. parvulus Stach, 1922 The examination of the material, agree with the original description., Published as part of Abrantes, Eduardo Assis & Mendonça, Maria Cleide De, 2007, New species and a new record of Isotomidae (Collembola) from the coast of Brazil, pp. 55-60 in Zootaxa 1500 (1) on page 58, DOI: 10.11646/zootaxa.1500.1.2, http://zenodo.org/record/5088000, {"references":["Kovac, L. & Palacios-Vargas, J. G. (1996) A survey of Mexican Folsomides (Collembola: Isotomidae) with description of three new species. European Journal of Entomology, 93, 595 - 606."]}

The genus Folsomides Stach, 1922 is represented by four species in the Iranian fauna, two of which are new to the science. Folsomides halshinicus sp. nov. and Folsomides subvinosus sp. nov. are found in Kermanshah Province (Western Iran). The new species belong to the Folsomides angularis (Axelson, 1905) group and they are distinguishable from other congeneric species by the macrochaetotaxy (11/22233 macrochaetae on thoracic II to abdominal V tergites) and the number of chaetae on dens. A table with the diagnostic characters for the identification of Iranian species of Folsomides is given. Some figures of F.vinosus Fjellberg, 1993 from Canary Island was given after study of type specimens.

Key of Folsomides species in the Ibero­balearic fauna (A summary of the main diagnostic characteristics of the species is given in Table 2) 1. Mucro absent (F. nanus ­group).................................................................................................. 2 ­ Mucro present............................................................................................................................ 3 2. With 5 + 5 corneola. Microsensilla: 11 / 111. Dens with 2 dorsal chaetae and 1 ventro­apical chaeta. Retinacle with 3 + 3 teeth and 1 chaeta on the corpus.................................................................................................................................................. F. ayllonensis Simón & Luciáñez, 1990 ­ With 2 + 2 corneola. Microsensilla: 10 /000. Dens with 2 dorsal chaetae and no ventro­apical chaeta. Retinacle with 2 + 2 teeth and 1 chaeta on the corpus....................................................... ................................................................................... F. almanzorensis Luciáñez & Simón, 1991 3. With 1­2 + 1­2 corneoles. Long and very narrow postantennal organ. Microsensillae: 10 /001., Published as part of Arbea, Javier I. & Jordana, Rafael, 2002, The genus Folsomides Stach of the Ibero­balearic fauna, with description of a new species (Collembola, Isotomidae), pp. 1-8 in Zootaxa 55 on page 6, DOI: 10.5281/zenodo.156067

[EN] The collembological fauna from different meadows in the Spanish Sierra de Ayllón is studied. Eighteen species were obtained and two new are here described: Folsomides ayllonensis n. sp. and Isotomodes ibanezi n. sp. Of the remaining, five are endemic: Xenyllogastrura affinis (Steiner), Friesea nevadensis Steiner, Protaphorura spinoidea (Steiner), Cryptopygus pazae (Selga) and Lepidocyrtus lusitanicus Gama; two are Mediterranean: Ceratophysella duodecimoculata Steiner and Cryptopygus ponticus Stach; two Holarctic: Xenyllodes armatus Axelson and Folsomia fimetaria (Linnaeus), and five are cosmopolitan., [ES] Se estudia la fauna colembológica de diversos prados húmedos de la Sierra de Ayllón. Se recolectaron 18 especies y se describen dos nuevas: Folsomides ayllonensis n. sp. e Isotomodes ibanezi n. sp. De las restantes, cinco son endémicas de la Península Ibérica: Xenyllogastrura affinis (Steiner), Friesea nevadensis Steiner, Protaphorura spinoidea (Steiner), Cryptopygus pazae (Selga) y Lepidocyrtus lusitanicus Gama; dos especies presentan un área de distribución mediterránea: Ceratophysella duodecimoculata Steiner y Cryptopygus ponticus Stach; dos son holárticas: Xenyllodes armatus Axelson y Folsomia fimetaria (Linnaeus), las demás especies cosmopolitas.

The genus Folsomides Stach, 1922 is represented by nine species in the Ibero-balearic fauna, one of which is new to the science. Folsomides mediterraneus n. sp. is found in Northern Spain (Navarra and Zaragoza) and Central Spain (Madrid). The new species is related to the Folsomides angularis (Axelson, 1905) group and it is easily distinguishable from other species by the chaetotaxy of microsensilla. A key for the identification of Ibero-balearic species of Folsomides is given.

Folsomides deserticola, a new species of Collembola belonging to the family Isotomidae, is described. It is widely distributed throughout the arid and semi-arid regions of southern Australia.