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Pillar Two of the GSP underpins many of the actions under the other Pillars by addressing the general lack of societal awareness of the importance of soil in people’s lives and the well - being of the planet. In many cases, deficiency in education is the specific underlying cause of unsustainable land management practices, of the general lack of investment (both in education and physical measures to protect soil) and, as importantly, of the widespread political reluctance to adopt short - and long - term measures to preserve and enhance soil conditions. The Plan of Action (PoA) for Pillar 2 consists of six interlinked and interdependent components: policy, investment, education, extension, public awareness and technical cooperation. The development of the PoA was initiated at the European Network Soil Awareness (ENSA) workshop in Aberdeen in September 2013. This was followed by discussions at the 2013 Global Soil Week in Berlin which lead to the establishment of a Working Group to produce a draft plan of action , which was eventually endorsed by t he ITPS in April 2014. The PoA was presented, reviewed and adopted by the Plenary Assembly of the GSP in July 2014. Seven key recommendations are presented. Robust policy frameworks are one means of ensuring the sustainable management and protection of soils. Therefore, governments must be in vited to create or reinforce policies on soil and its protection. Whereas soil protection and management is a long - term aim, most political decisions are governed by short - term ambitions that reflect the lifetime of governments or politicians. Today’s highly urbanised society means that the population is largely detached from food and fibre production issues and lack s a fundamental understanding of the role o f soil in enabling such life - critical services. It is disappointing to observe that society in many parts of the world attaches a greater value to developments in subjects such as particle physics and mobile communication than essential issues such as soil fertility. The PoA calls for a systematic awareness raising campaign in all countries on how soil relates to people’s everyday lives. This can be done through brief and vivid messages, not only as part of the World Soil Day celebrations and during the forthcoming International Year of Soils, but also as a sustained long - term outreach and engagement programme. The PoA recommends a significant increase in investments to support such actions. Education in soil sciences is important and needs to be taken into account by other disciplines. The current soil science community should strive to show synergies with other domains to demonstrate its relevance. Pressure should be brought at all levels to halt the decline in soil science teaching at tertiary level, while boosting professional technical qualifications and support to educationalists, so that soils and agriculture can be more appealing for the younger generations. Soil extension services should interpret and present relevant research - based information to a broader range of stakeholders in an understandable and usable form, including farmer - to - farmer schemes or through initiatives such as the Soils Doctors Programme. The technical approach underlying the extension services should reflect mutually beneficial cooperation rather than from mere transfers from one partner to another. Finally, investments must go hand in hand with awareness o f the importance of soil resources. These investments should develop an effective skills base and entrepreneurship among soil users. The GSP should make full use of such tools as the Healthy Soils Facility to generate and facilitate the collection of financial contributions to the PoA.

Gene expression is a regulated process fueled by ATP consumption. Therefore, regulation must be coupled to constraints imposed by the level of energy metabolism. Here, we explore this relationship both theoretically and experimentally. A stylized mathematical model predicts that activators of gene expression have variable impact depending on metabolic rate. Activators become less essential when metabolic rate is reduced and more essential when metabolic rate is enhanced. We find that in the Drosophila eye, expression dynamics of the yan gene are less affected by loss of EGFR-mediated activation when metabolism is reduced, and the opposite effect is seen when metabolism is enhanced. The effects are also seen at the level of pattern regularity in the adult eye, where loss of EGFR-mediated activation is mitigated by lower metabolism. We propose that gene activation is tuned by energy metabolism to allow for faithful expression dynamics in the face of variable metabolic conditions., Competing Interests: COMPETING INTERESTS The authors declare no competing financial interests.