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Galve (Teruel, Spain) is not a single fossil site but an ensemble of more than 50 localities with Mesozoic terrestrial vertebrate remains of Tithonian-Lower Barremian age. Galve fossil sites belong to the Maestrazgo basin (Central Iberian Range), Gaive Subbasin, and appear in four geological formations: Higueruelas Formation (Tithonian), Villar del Arzobispo Formation (upper Tithonian-middle Berriasian), El Castellar Formation (uppermost Hauterivianlowermost Barremian) and Camarillas Formation (lower Barremian). Most sites contain bony remains, but there are also paleoichnological and paleoological sites, with dinosaur and other reptilian tracks and eggshells. Mammals and dinosaurs are the best-known Gaive vertebrates, but some studies on sharks, bony fishes, amphibians, squamates and crocodiles have been published. Some groups, like turtles and pterosaurs, have not practically been studied yet. Galve is the type locality of severa1 taxa: the shark Lonchidion microselachos, the amphibian Galverpeton ibericum, the dinosaur Aragosaurus ischiaticus, the mammals Galveodon nannothus, Lavocatia alfambrensis, Eobaatar hispanicus, Parendotherium herreroi, Spalacotherium henkeli and Pocamus pepelui, and the dinosaur eggshell Macroolithus turolensis. In this paper we revise the knowledge on Galve vertebrates, and update the faunal list of its fossil sites, taking into account, for the first time, the stratigraphic distribution of the taxa.En Galve (Teruel) hay más de 50 localidades con restos de vertebrados continentales en las Formaciones Higueruelas (Titónico), Villar del Arzobispo (Titánico superior-Berriasiense medio), El Castellar (Hauteriviense terminal-Barremiense basal) y Camarillas (Barremiense inferior). Por tanto, «Galve» no es un único yacimiento de vertebrados mesozoicos, sino una localidad con numerosos yacimientos de vertebrados del intervalo Titónico-Barremiense, que geológicamente pertenecen a la Cuenca cretácica inferior del Maestrazgo (Cordillera Ibérica Central), Subcuenca de Galve. La mayor parte de los yacimientos contienen restos óseos, pero también son abundantes los yacimientos paleoicnológicos y paleoológicos. Los vertebrados mejor conocidos son los mamíferos y dinosaurios, aunque también hay estudios sobre los tiburones, peces óseos, anfibios, escamosos y cocodrilos. Algunos grupos, como tortugas y pterosaurios, permanecen prácticamente sin estudiar. Galve es la localidad tipo de varios taxones de vertebrados: el tiburón Lonchidion microselachos, el anfibio Galverpeton ibericum, el dinosaurio Aragosaurus ischiaticus, los mamíferos Galveodon nannothus, Lavocatia alfambrensis, Eobaatar hispanicus, Parendotherium herreroi, Spalacotherium henkeli y Pocamus pepelui, y la cáscara de huevo de dinosaurio Macroolithus turolensis. Es este trabajo se revisa el estado de conocimientos sobre los vertebrados de Galve, y se actualiza la lista faunística de sus yacimientos, siendo la primera vez que se realiza teniendo en cuenta la distribución estratigráfica de los taxones.

Galve (Teruel, Spain) is not a single fossil site but an ensemble of more than 50 localities with Mesozoic terrestrial vertebrate remains of Tithonian-Lower Barremian age. Galve fossil sites belong to the Maestrazgo basin (Central Iberian Range), Gaive Subbasin, and appear in four geological formations: Higueruelas Formation (Tithonian), Villar del Arzobispo Formation (upper Tithonian-middle Berriasian), El Castellar Formation (uppermost Hauterivianlowermost Barremian) and Camarillas Formation (lower Barremian). Most sites contain bony remains, but there are also paleoichnological and paleoological sites, with dinosaur and other reptilian tracks and eggshells. Mammals and dinosaurs are the best-known Gaive vertebrates, but some studies on sharks, bony fishes, amphibians, squamates and crocodiles have been published. Some groups, like turtles and pterosaurs, have not practically been studied yet. Galve is the type locality of severa1 taxa: the shark Lonchidion microselachos, the amphibian Galverpeton ibericum, the dinosaur Aragosaurus ischiaticus, the mammals Galveodon nannothus, Lavocatia alfambrensis, Eobaatar hispanicus, Parendotherium herreroi, Spalacotherium henkeli and Pocamus pepelui, and the dinosaur eggshell Macroolithus turolensis. In this paper we revise the knowledge on Galve vertebrates, and update the faunal list of its fossil sites, taking into account, for the first time, the stratigraphic distribution of the taxa. En Galve (Teruel) hay más de 50 localidades con restos de vertebrados continentales en las Formaciones Higueruelas (Titónico), Villar del Arzobispo (Titánico superior-Berriasiense medio), El Castellar (Hauteriviense terminal-Barremiense basal) y Camarillas (Barremiense inferior). Por tanto, «Galve» no es un único yacimiento de vertebrados mesozoicos, sino una localidad con numerosos yacimientos de vertebrados del intervalo Titónico-Barremiense, que geológicamente pertenecen a la Cuenca cretácica inferior del Maestrazgo (Cordillera Ibérica Central), Subcuenca de Galve. La mayor parte de los yacimientos contienen restos óseos, pero también son abundantes los yacimientos paleoicnológicos y paleoológicos. Los vertebrados mejor conocidos son los mamíferos y dinosaurios, aunque también hay estudios sobre los tiburones, peces óseos, anfibios, escamosos y cocodrilos. Algunos grupos, como tortugas y pterosaurios, permanecen prácticamente sin estudiar. Galve es la localidad tipo de varios taxones de vertebrados: el tiburón Lonchidion microselachos, el anfibio Galverpeton ibericum, el dinosaurio Aragosaurus ischiaticus, los mamíferos Galveodon nannothus, Lavocatia alfambrensis, Eobaatar hispanicus, Parendotherium herreroi, Spalacotherium henkeli y Pocamus pepelui, y la cáscara de huevo de dinosaurio Macroolithus turolensis. Es este trabajo se revisa el estado de conocimientos sobre los vertebrados de Galve, y se actualiza la lista faunística de sus yacimientos, siendo la primera vez que se realiza teniendo en cuenta la distribución estratigráfica de los taxones.

Se describen cinco grandes contramoldes ornitópodos encontrados en Galve durante una campaña de prospección. Algunas de estas icnitas ornitópodas son las mayores del Cretácico Inferior del Este de la Península Ibérica. Los contramoldes confirman que había ornitópodos grandes en esta cubeta durante el Barremiense. Las icnitas, que también proporcionan información sobre el movimiento del pie durante la pisada, se atribuyen al cf. Iguanodontipus, siendo la segunda vez que se cita este tipo de huella en la provincia de Teruel, We describe five large ornithopod natural casts found in Galve (Teruel, Spain) during an exploration campaign. Some of these ornithopod footprints are the largest of the Lower Cretaceous in the east of the Iberian Peninsula. The casts support that there were big ornithopods in the Galve Basin during the Barremian age. The footprints, wich also provide information on the movement of the foot during the gait, are attributed to cf. Iguanodontipus. It is the second time that this type of track cites in the Teruel province

Important sites of direct and indirect dinosaur fossils can see in the Galve area. This paper shows the sauropod footprint structures found in a new ichnite outcrop in the Cabezo de San Cristobal. The tracksite is situtated in a sandstone of the Camarillas Formation (lower Barremian). It is the first time of the sauropod footprints are described in this Formation and therefore the latest from the Teruel province. In the mostly prints, the shaft base is a dead zone composed by a broken fragment of the sandstone level intruded several decimetres in deeper argillaceous sediments. Four of the five sauropod tracksites from Galve were made in a moment of waterlogged sediment in agreement with the cautious dinosaurs hypothesis

Important sites of direct and indirect dinosaur fossils can see in the Galve area. This paper shows the sauropod footprint structures found in a new ichnite outcrop in the Cabezo de San Cristobal. The tracksite is situtated in a sandstone of the Camarillas Formation (lower Barremian). It is the first time of the sauropod footprints are described in this Formation and therefore the latest from the Teruel province. In the mostly prints, the shaft base is a dead zone composed by a broken fragment of the sandstone level intruded several decimetres in deeper argillaceous sediments. Four of the five sauropod tracksites from Galve were made in a moment of waterlogged sediment in agreement with the cautious dinosaurs hypothesis

The association of dinosaurs of the Villar del Arzobispo Formation in Galve (Teruel, Aragon, Spain) represents a «jurassic association» by the abundance of primitive clades like basal neosauropods and the scarcity of ornithopods. This fact could indicate the persistence of the Jurassic dinosaur faunas in this part of Laurasia, at least up to the Berriasian. Anyway the absence of stegosaurians could be interpreted as the first evidence of the turnover that would come in the Cretaceous faunas. Our data indicate that the climatic change did not suddenly affected to the dinosaurs of the Tethyan province. On the contrary, the turnover to the modern faunas took place gradually at least during the Berriasian

The association of dinosaurs of the Villar del Arzobispo Formation in Galve (Teruel, Aragon, Spain) represents a «jurassic association» by the abundance of primitive clades like basal neosauropods and the scarcity of ornithopods. This fact could indicate the persistence of the Jurassic dinosaur faunas in this part of Laurasia, at least up to the Berriasian. Anyway the absence of stegosaurians could be interpreted as the first evidence of the turnover that would come in the Cretaceous faunas. Our data indicate that the climatic change did not suddenly affected to the dinosaurs of the Tethyan province. On the contrary, the turnover to the modern faunas took place gradually at least during the Berriasian

The new outerop of Santa Bárbara (Galve, Province of Teruel, Spain, here provisionally considered as Barremian-Aptian in age) has yielded sorne autopodial pieces especially of the hind limbo There are a tibial distal portion, tarsus, metatarsal fragmenta and several phalanges of the three digits, especially of digit IV. Within the family Iguanodontidae, two groups of genera can be recognized, One of them, more progressive, has the tendency towards the disappearance of the first digit and the shortening of the acropodial elements. The material from Santa Bárbara c1early belongs to the last group, and its morphology indicates the genus Iguonodon, Based on morfological criteria it is not possible, up to now, to separate I. bernissartensis from I. mantelli, it can be done only based on size criteria. So, the greatest part of the material of Santa Bárbara is identified as l. cf. mantelli, though this outcrop has yielded two phalanges of the size of I. bernissartensis too.El nuevo yacimiento de Santa Bárbara (Galve, Teruel, considerado provisionalmente como Barremiense-Aptiense) ha proporcionado una serie de elementos apendiculares, especialmente del autópodo posterior. En este material se incluye una porción distal de tibia, tarso, fragmentos metatarsales y diversas falanges, especialmente del cuarto dígito. Parece poder distinguirse, dentro de la familia Iguanodontidae, dos grupos de géneros. Uno de ellos, más progresivo, tiende a la desaparición del dedo uno y al acortamiento de los elementos acropodiales . El material de Santa Bárbara se incluye claramente en este grupo. Su morfología indica Su pertenencia al género Iguanodon. Al nivel actual de información, no parece posible distinguir los autópodos posteriores de I. mantelli e I. bernissartesis en base a criterios morfológicos, sino tan sólo a tamaño. Desde este punto de vista, la mayoría del material de Santa Bárbara se identifica como I. cf. mantelli, aunque el yacimiento ha proporcionado igualmente dos falanges de la talla de I.bernissartesis.

The new outerop of Santa Bárbara (Galve, Province of Teruel, Spain, here provisionally considered as Barremian-Aptian in age) has yielded sorne autopodial pieces especially of the hind limbo There are a tibial distal portion, tarsus, metatarsal fragmenta and several phalanges of the three digits, especially of digit IV. Within the family Iguanodontidae, two groups of genera can be recognized, One of them, more progressive, has the tendency towards the disappearance of the first digit and the shortening of the acropodial elements. The material from Santa Bárbara c1early belongs to the last group, and its morphology indicates the genus Iguonodon, Based on morfological criteria it is not possible, up to now, to separate I. bernissartensis from I. mantelli, it can be done only based on size criteria. So, the greatest part of the material of Santa Bárbara is identified as l. cf. mantelli, though this outcrop has yielded two phalanges of the size of I. bernissartensis too. El nuevo yacimiento de Santa Bárbara (Galve, Teruel, considerado provisionalmente como Barremiense-Aptiense) ha proporcionado una serie de elementos apendiculares, especialmente del autópodo posterior. En este material se incluye una porción distal de tibia, tarso, fragmentos metatarsales y diversas falanges, especialmente del cuarto dígito. Parece poder distinguirse, dentro de la familia Iguanodontidae, dos grupos de géneros. Uno de ellos, más progresivo, tiende a la desaparición del dedo uno y al acortamiento de los elementos acropodiales . El material de Santa Bárbara se incluye claramente en este grupo. Su morfología indica Su pertenencia al género Iguanodon. Al nivel actual de información, no parece posible distinguir los autópodos posteriores de I. mantelli e I. bernissartesis en base a criterios morfológicos, sino tan sólo a tamaño. Desde este punto de vista, la mayoría del material de Santa Bárbara se identifica como I. cf. mantelli, aunque el yacimiento ha proporcionado igualmente dos falanges de la talla de I.bernissartesis.