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16. Heteragrion eboratum Donnelly, 1965 Abundant in open sunspots along mid-sized to large, forested rivers at higher elevations. In July‒August males are often seen interacting in a frontal display of territoriality centred around an exposed point such as a stick above the river., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

32. Aeshna williamsoniana Calvert, 1905 Only females were collected. This species was difficult to catch due to high and fast flight, and most collections came from specimens trapped under (blue) tarpaulins in the camps. The species was widespread across CNP in open areas mostly close to or around the large rivers. New record for Honduras., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 462, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

7. Palaemnema sp. 3 (nr. paulirica) Its current status is uncertain pending further studies., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 456, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

8. Palaemnema lorae Jocque & Garrison, n. sp. Figs. 2–4, 6–10 Holotype ♂: HONDURAS: Cortés Dept., CNP, Cantiles, Trail 5, small river close to camp, N15.513457 W88.241681; 1846m, 23 June 2012 collected by Merlijn Jocque, field code: BINCO _HON_12_047 (RBINS). Paratypes: same data but: 20 June 2013, 1♂; same data but: 4 August 2013, 1♂; same data but: 30 July 2015, 1♂; same data but: 9 June 2017, 1♂; Cortecito camp, N15.521825 W88.288277, 1363m, 29 June 2011, collected by Merlijn Jocque, 1♂; field codes: BINCO _HON_11_029, BINCO _HON_13_038-039, BINCO _HON_15_051, BINCO _HON_17_007 (RWG); El Danto camp, Tr 4, CNP, Honduras, N15.53593 W88.2854, 1481m, 28 June 2014, collected by Merlijn Jocque, 2♂♂; field codes: BINCO _HON_14_091-092 (MJ). Etymology: Named lorae (Latinized name) after Lore Geeraert, friend of the senior author who contributed to the study of dragonflies in CNP and in honor of her love for all living things and the rainforest. Description of holotype (colors not well preserved, Fig. 2) Head: labium ivory white with tips of median and lateral lobes and movable hook becoming black; maxilla palp ivory white, maxillary palps black; labrum pale margined apically in black; genae, clypeus and base of mandibles pale; antefrons pale, postfrons pale margined basally with black; remainder of head black with metallic reflections and with an obscure brown spot laterad to lateral ocellus; rear of head entirely black; transverse occipital carina present but poorly developed, its lateral extremity not angular or pronounced but merging with remainder of occipital lobe. Thorax. Prothorax black dorsally, lateral portion of middle lobe pale; propleuron black; most of mesepisterum including dorsal carina black, merging above with black on mesepisternum, pale antehumeral stripe narrow, enlarged basally and narrowing dorsally and ending before antealar sinus; posterior half of mesepimeron and anterior half of metepisternum pale; broad black metepleural stripe present, its posterior margin of varied outline, an obscure pale spot just below antealar carina and anterior to obsolete mesopleural suture (Figs. 2, 6), venter of thorax ivory; coxae and trochanters ivory (possibly blue in life); femora pale but darkened apically, protibia largely black; meso- and metatibiae mostly pale with obscure dark areas along margins; tarsi and armature black. Wings hyaline, venation (Fig. 8, paratype) black; pterostigma elongate, rhomboid, brown, surmounting 1 ¾ cells in all wings; Px Fw: 19/20, Hw 18/17; RP 2 at Fw 8/8, Hw: 7/7; IR 1 at Fw 10/9, Hw 9/9; MP ending at level origin of IR 1 in Fw, 2.5 cells distal to origin of IR 1 in Hw. Abdomen including appendages black except for obscure lateral basal rings on S4–7 (Figs. 2–4). Genital ligula (Fig. 9) of type B of Calvert (1931). Cercus (Fig. 9) semicircular armed above with a small dorsal tooth at 0.5 of appendage length, apex of cercus entire; paraproct semicircular, about ¾ length of cercus, the distal 0.50 laminar, concave medially, its tip ending in a simple medially directed unmodified spine. Dimensions: Hw 33, abdomen 48, total length 57. Variation in paratypes: Extent of black on mesepimeron varies with two males with entire mesepimeron black (Fig. 7). Px Fw: 20–21; Hw: 18–20; RP 2 at Fw 6–8, Hw: 7; IR 1 at Fw 9–11, Hw 8–9; Hw: 32–35; Abdomen:48– 51. Diagnosis: A large species (56–60mm) with pale colors most likely blue in life (Fig. 3) with tip of paraproct ending in a simple acute tip (Fig. 9). Male of P. lorae is larger than any known congener with the exception of P. gigantula Calvert, 1931. However, in the latter species the wings are comparatively shorter with the Hw extending about midway to S5 (Fig. 5) compared to about midway to S 6 in P. lorae (Fig. 2). S8–10 are primarily blue in P. gigantula but entirely black in P. lorae. Male of P. lorae keys in Calvert (1931) to couplet OO (Abdominal segment nine black) then to P. carmelita Ris, 1918 (RR. Superior appendages with apex not excised, superior tooth at 0.48– 0.53 of appendage length; basal tooth of inferiors at a most blunt or triangular tubercle; penis form B; mesepimeron and metepisternum obscure bronze violet) but differs from that species as follows: (contrasting characters for P. carmelita in parentheses): Narrow pale antehumeral stripe present (absent); tip of paraproct ending in a simple medially directed unmodified spine (ending in a spatulate tooth with a shallow apical notch, Fig. 11, enlarged and redrawn from Kennedy, 1938). All examined specimens of P. lorae were preserved in ETOH upon capture and, over time, color pattern became obscured upon drying. Some specimens (including holotype) also suffered thoracic pressure distortions resulting in buckling of the venter of the thorax. Biology: Palaemnema lorae was observed within an elevational range of 1363‒1846m along fast flowing crystal-clear forest streams. Forest was mostly lower montane rain forest. Damselflies were observed moving around during brief moments of sunshine penetrating the often-cloudy environment. Otherwise, a gentle beating of the vegetation close to the river edges could trigger movement of individuals. This species is thus far known only from the type locality where it was rare., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 457, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139, {"references":["Calvert, P. P. (1931) The generic characters and the species of Palaemnema (Odonata: Agrionidae). Transactions American Entomological Society, 57, 1 - 110.","Kennedy, C. H. (1938) Palaemnema picicaudata, P. abbreviata and P. brucelli, new dragonflies from Ecuador, with notes on other Palaemnema. Annals of the Entomological Society America, 31 (2), 249 - 266. https: // doi. org / 10.1093 / aesa / 31.2.249"]}

9. Amphipteryx meridionalis González-Soriano, 2010 A common forest damselfly at mid-elevation (1500masl) in CNP. Associated with large rivers and omnipresent in sunspots along the forested rivers. Largely absent along small, shaded streams at high elevation or open rivers at low elevation. The female was first described from CNP (Jocque & Argueta 2014)., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 459, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139, {"references":["Gonzalez-Soriano, E. (2010) A synopsis of the genus Amphipteryx Selys 1853 (Odonata Amphipterygidae). Zootaxa, 2531 (1), 15 - 28. https: // doi. org / 10.11646 / zootaxa. 2531.1.2","Jocque, M. & Argueta, I. (2014) A new species in the genus Amphipteryx Selys, 1853 (Odonata, Amphipterygidae) from Pico Bonito National Park, Honduras. ZooKeys, 408, 71 - 80. https: // doi. org / 10.3897 / zookeys. 408.7174"]}

30. Paraphlebia itzamna Ortega-Salas, Jocque & González-Soriano in Ortega-Salas, González-Soriano & Jocque, 2022 A lowland species that in CNP is most reliably observed in the shaded parts of where smaller streams enter the large river close to Guanales camp., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 461, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139, {"references":["Ortega-Salas, H., Gonzalez-Soriano, E. & Jocque, M. (2022) Untangling the waterfall damsels: a review of the Mesoamerican genus Paraphlebia Selys in Hagen, 1861 (Odonata: Thaumatoneuridae) with descriptions of 11 new species. Zootaxa, 5089 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 5089.1.1"]}

20. Argia funebris ( Hagen, 1861) A rare species in CNP, currently only recorded from a single spot at higher elevation close to a waterfall with a lot of bare exposed rock along Río Cusuco, a large rapid flowing river. New record for Honduras., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 461, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

33. Oplonaeschna armata (Hagen, 1861) New record for Honduras., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 462, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

2. Palaemnema angelina Selys, 1860 A lowland species., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 456, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

10. Hetaerina capitalis Selys, 1873 This species is rarely collected within the CNP boundaries, but with occasional observations as high as Guanales camp (1331 masl.)., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

6. Palaemnema sp. 2 Its current status is uncertain pending further studies., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 456, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

29. Paraphlebia kukulkan Jocque & Ortega-Salas in Ortega-Salas, González-Soriano & Jocque, 2022 Common along small, shaded forest streams and creeks. This species was only found at higher elevations around 1500 to over 1900masl., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 461, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139, {"references":["Ortega-Salas, H., Gonzalez-Soriano, E. & Jocque, M. (2022) Untangling the waterfall damsels: a review of the Mesoamerican genus Paraphlebia Selys in Hagen, 1861 (Odonata: Thaumatoneuridae) with descriptions of 11 new species. Zootaxa, 5089 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 5089.1.1"]}

18. Argia cuprea (Hagen, 1861) Only observed at low elevation., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

36. Epigomphus subobtusus Selys, 1878 A relatively common gomphid in CNP that was encountered from low to mid elevation. Sometimes occurring with E. schausi. Adults were regularly seen basking in the sun on higher vegetation, not necessarily close to the water., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 462, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

14. Hetaerina rudis Calvert, 1901 This is a rare species in CNP, only observed at higher elevations. The female is as yet known from the original description only and has not been observed in CNP yet. New for Honduras., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

19. Argia elongata Garrison & von Ellenrieder, 2019 Only observed at low elevation. The species was described based on material from Cusuco NP (Garrison & von Ellenrieder 2017)., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139, {"references":["Garrison, R. W. & von Ellenrieder, N. (2017) New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae). Zootaxa, 4035 (1), 1 - 93. https: // doi. org / 10.11646 / zootaxa. 4235.1.1"]}

12. Hetaerina infecta Calvert, 1901 Together with H. cruentata this is the most encountered Hetaerina species in CNP., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

4. Palaemnema nathalia Selys, 1886 Individuals from this species are amongst the largest examples seen. This is a widespread species occurring from southern Mexico south through Venezuela., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 456, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

44. Sympetrum illotum illotum (Hagen, 1861) Few specimens were collected of this species; mostly from open and sunlit areas around basecamp., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 463, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

35. Rhionaeschna jalapensis (Williamson, 1908) A single specimen was collected at mid-elevation in an open sunny spot in mixed pine dominated forest close to Río Cusuco (at basecamp). This species is possibly more common but in flight can easily be confused with R. cornigera., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 462, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

5. Palaemnema sp. 1 (nr. paulitaba) The most commonly encountered Palaemnema species in CNP. Its current status is uncertain pending further studies., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 456, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

17. Argia chelata Calvert, 1902 This large Argia Rambur, 1842 is common in CNP and often abundant along mid-sized to large rivers., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

24. Megaloprepus caerulatus (Drury, 1782) The largest dragonfly of the world is occasionally observed at low- to mid (1600masl.) elevations in CNP. Observations were mostly opportunistic., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 461, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

11. Hetaerina cruentata (Rambur, 1842) Together with H. infecta this is the most encountered Hetaerina Hagen in Selys, 1853 species in CNP. Hetaerina cruentata is found along the whole elevational range from just 500m in the lowlands to 1900m. Two morphological variations are observed separated by elevation; a small slender and paler variant at low elevation, and a larger, darker variant with more intense red colouring on the wing from around 1500m., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 460, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

1. Archilestes grandis (Rambur, 1842) Only recorded at low elevation. New record for Honduras., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 456, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

42. Libellula herculea Karsch, 1889 A widely distributed species that was often seen around muddy pools after rains on the dirt road between Buenos Aires and the visitor center in CNP., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 463, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

31. Cora marina Selys, 1868 A single specimen was recorded. Localized, currently only known from a population found along a mid-sized forested river at 1363masl. Honduras, Cortés, CNP, on the West side of CNP. Apparently endemic to Honduras., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on pages 461-462, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

38. Cordulegaster diadema Selys, 1868 This large dragonfly was sometimes caught in mist nets for birds mounted over large rivers in the forest. Some collections came from a central open spot at mid elevation in CNP. New record for Honduras. NOTE: Cordulegaster godmani McLachlan, 1876, was recently considered a junior synonym of C. diadema (Novelo-Gutiérrez 2018)., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 462, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139, {"references":["Novelo-Gutierrez, R. (2018) Cordulegaster virginiae sp. nov. from Mexico, including a comparison with C. diadema Selys, 1868, and a redescription of its larva (Odonata: Cordulegastridae). Zootaxa, 4394 (3), 371 - 382. https: // doi. org / 10.11646 / zootaxa. 4394.3.3"]}

40. Brechmorhoga rapax rapax Calvert, 1898 This species shared the same river habitat as B. pertinax but preferred sunlit areas., Published as part of Jocque, Merlijn & Garrison, Rosser, 2022, Dragonflies of Cusuco National Park, Honduras; checklist, new country records and the description of a new species of Palaemnema Selys, 1860 (Odonata: Platystictidae), pp. 453-476 in Zootaxa 5188 (5) on page 463, DOI: 10.11646/zootaxa.5188.5.3, http://zenodo.org/record/7099139

A ten­day collecting trip to Napo province was conducted between January 13 and 23, 2020, visiting localities where W. C. Macintyre originally collected Argia schneideri Garrison & von Ellenrieder, 2017 between 1935 and 1942, with the intention of documenting its life habits and obtaining photographs in life. A total of 65 odonate species in 36 genera were collected, including four new records for Napo province, but the target species was not found. A list of species recorded and color scans of species that have so far not been photographed are included. Color photographs and notes on the habitat of Argia schneideri are made available through the courtesy of colleagues who found it elsewhere, and its current known distribution is discussed.

Indolestes anomalus Fraser, 1946 Figs. 1–7 Indolestes anomala Fraser 1946: 43 –44, figs. 1c, 2e—original description of “a single rather teneral female” from “ Siam: Doun-moi”, drawings of the pterostigma and thorax in lateral view; Hämäläinen & Pinratana 1999: 32 —report for Tak (June) [Mae Sot, 1 vi 1986, 1 ♂, 1 ♀, A. Pinratana’s collectors leg.—M. Hämäläinen, pers. comm.], Lampang (October) [Huay Tak Teak Plantation, 23 x 1986, 1 ♂, M. Hämäläinen leg.—M. Hämäläinen, pers. comm.] and Chaiyaphum (July) [Phu Khieo, 20 vii 1996, 1 ♀, A. Pinratana leg.—M. Hämäläinen, pers. comm.] Provinces of Thailand; Ng et al. 2011: 12, fig. 1—report for the Peninsular Malaysia, Cameron Highlands of 2 ♂♂ collected on 20 ix 2008 at Sg. Pos Terisu area and of 1 ♂ collected on 24 ix 2008 at Sg. Telom area; drawings of the variable pattern of synthorax of males from Malaysia, Cameron Highlands, and Thailand, Lampang and Tak Provinces Lestes (Indolestes) anomala (Fraser): Kimmins 1966: 178 —information on the holotype in the collection of British Natural History Museum: “ Holotype ♀. Indolestes anomala Fraser, Dalat, Siam, 17.x. 20, Type [label F.C.F.]. The locality given in the original description is Doun-moi, without date, but the specimen otherwise agrees.” Indolestes anomalus: Day et al. 2012: 18 —report of 2 ♂♂, 1 ♀ collected on 09– 10.07.2011 in Thailand, Petchabun Province, Nam Nao National Park. Specimens examined. Vietnam. 1 ♀ (Fig. 3), Đỗng Nai Province, Cát Tiên National Park, 11.440 N, 107.422 E, 6 ii 2016, T. Kompier leg. Thailand: 2 ♂, 5 ♀♀ (1 ♂ (Fig. 6), 1 ♀ (Fig. 4) in tandem), Chaiyaphum Province, small muddy pond at Tung Lui Lai, 16.6089 N, 101.8906 E, 19 vii 1996, R.W. & J.A. Garrison leg. The type locality. The type locality of I. anomalus was given in the original description as “ Siam: Doun-moi”. Siam is the historical name of Thailand, but no site with this or a similar name could be located in that country, so Hämäläinen & Pinratana (1999: page 32) noted that “The whereabouts of the type locality is unknown”, and Dow (2009) noted that “The type locality, " Siam, Doun Moi" (Fraser 1946) cannot be relocated.” “Doun Moi” might be variant spelling of “Than Moi” or “Than Muoi”, a place in northern Vietnam (or in Tonkin, as that was referred to at that time), well known for numerous insect specimens collected there by Hans Fruhstorfer in June–July 1901. Presently this location is called Đỗng Mỏ town, the capital of Chi Lăng District, Lạng SƠn Province (21.66° N, 106.58° E) (Hämäläinen 2015). Thus “ Siam ” could result from a confusion with Tonkin or used in a sense to include all of Indochina. (For instance, the holotype of Macrogomphus borikhanensis Fraser, 1933 is labelled “Borikhane/ Laos, Siam ”, see Kimmins 1966). But the holotype label has no indication of the collector, which as a rule is present on labels of Fruhstorfer’s specimens. However, Kimmins (1966) transcribed the holotype label (Fig. 2d) as follows: " Indolestes anomala Fraser, Dalat, Siam, 17.x.20, Type" and added: "Type locality given in the original description is Doun-moi, without date, but the specimen otherwise agrees". Kimmins (1966) noted a number of cases where actual labels of the type specimens did not coincide with information given by Fraser in his original descriptions. There are a Dalet or Dalat Town in Myanmar and Đà Lạt Town in Vietnam, the capital of Lâm Đỗng Province. The Cát Tiên District of the latter province is the location of the headwaters of the Đỗng Nai River. This river gave rise to the name of Đỗng Nai Thưọng commune in Cát Tiên District and Đỗng Nai Province (with the capital Biên Hòa), which borders Lâm Đỗng Province to the southwest. It is not clear which were the local toponyms and administrative divisions during Fraser’s time but it is very plausible that the holotype originated from the Đỗng Nai River basin in South Vietnam and that the river (or town) name was misspelled as “Doun-moi”. The original label of the specimen (or otherwise communicated information on the locality) could include both ‘Dalat’ and ‘Doun-moi’, two different toponyms from the same area, while Fraser included only the former on the label under the holotype and mentioned only the latter in the original description. “ Siam ” still was also an error or was used in too broad a sense. So any topotypes should be sought within the Đỗng Nai River basin in Lâm Đỗng Province or Đỗng Nai Province. Cát Tiên National Park is located along the Dong Nai River on the border between these provinces and so would be a good place to search for this species. The holotype. The holotype is kept in Natural History Museum, United Kingdom (BMNH) under reference number NHMUK 0 13324318. Photographs of its details were kindly provided by Benjamin Price (Fig. 2). The holotype is a somewhat teneral female in poor condition. The left hind wing and abdominal segments [hereafter S] 8–10 are missing (but were described in the original description). There is a pair of pale spots on the occiput lateroposteriorly of the lateral ocelli, surrounding small tubercles, mentioned in the original description (Fraser 1946). The synthoracic pattern (Fig. 2c) fits the drawing in the original description (Fraser 1946: Fig. 1b) where, however, the black spots at the posteroventral corner of the mesepisternum and anteroventral corner of the metepimeron were shown as smaller, while the dark stripe from the latter to the spiracle was shown as distinct although this is barely traceable in the holotype. In describing the most conspicuous diagnostic character for the species, the pterostigma unusually short for Lestidae, Fraser (1946: 45) made somewhat confusing statements: “pterostigma... of fore wing about one-third longer than deep, that of the hind-wing about half as long again as deep” meaning that the pterostigmata length in the fore and hind wings was, respectively, ca 1.33 and 1.5 times its height. Fraser did not mention which pterostigma he figures (Fig. 2e), showing the maximum length comprising 1.68 of the maximum width (Fig. 1a). The actual pterostigmata of the holotype are substantially longer: 1.90 times as long as high along the dorsal side and 2.25 times as long as high along the ventral side in the fore wing (Fig. 2e) and, respectively, 1.75 and 2.20 times as long as high in the hind wing (Fig. 2f). In both wings, the pterostigmata covers a bit less than two underlying cells (Fig. 2e, f), but one and half cells in Fraser’s figure. Fraser clearly stated that he had only one specimen and the BMNH specimen labeled as the holotype of “ Indolestes anomala ” fits very well in other respects, e.g. the synthoracic pattern, thus excluding the possibility of other specimens. The left hind wing is missing in the holotype (Fig. 2a); perhaps it had a shorter pterostigma, which Fraser may have figured and he may have detached that wing for convenience. Some of the discrepancies mentioned above between Fraser’s own figure and his textual description of the pterostigmata proportions indicates that he likely did not aim for high degree of accuracy of his measurements and/or drawings. Other incorrect proportions in Fraser’s drawings are well known (e.g. the wrong shape of the paraprocts shown for the lectotype of Lestes nigriceps Fraser, 1924, see Kosterin 2018). A nearly topotypic female. On 6 ii 2016 one of us (T.K.) collected a female in Cát Tiên National Park (Fig. 3). It fits, with minor differences, the original description well. It displays the short pterostigmata (Fig. 3b), and has a pair of pale-coloured tubercles on the occiput, behind the lateral ocelli, a pair of dark spots on the anteclypeus, and a synthoracic pattern consisting of a posthumeral bronze stripe split into three irregular spots and a blunt projection of the dorsal bronze stripe at ca. three-fourths of its length (Fig. 3a). The central and posterior bronze spots on the mesepimeron are somewhat shorter (Fig. 3a) than in the holotype (Fig. 2c). The pale occipital spots extend lateroposteriorly of the pair of tubercles (Fig. 3a). The length of the hindwing is 20 mm (as in the original description); of the abdomen 28.5 mm (vs 27 mm in the original description). The hind wing pterostigma of the female from Cát Tiên is 1.63 and 1.78 times as long as high along the dorsal and ventral margins, respectively (Fig. 3b), which is even shorter than in the holotype (Fig. 2f). The characters of the end of abdomen differ considerably from the original description, which states “segments 8 to 10 dark brown with obscure indications of large yellow lateral spots. Anal appendages brownish, shortly conical”. This statement cannot be verified since the holotype lacks S8–10. In the Cát Tiên female, S8 is dark dorsally and pale ventrally and posteriorly, with a pair of smaller dark lateral spots, all colour transitions of which are diffuse; S9 dark becoming paler laterally, while S10 is entirely pale (Fig. 3c). Cerci are also pale with black tips and are conical but are as long as S10 (Fig. 3c). The colorational differences could result from the post-mortem changes or from the teneral state of the holotype, while the Cát Tiên female was photographed upon capture (Fig. 3c). This was the only specimen collected during many visits to the park, nor has it been found during the extensive explorations of the area by James Holden (pers. comm.), so it may be rare. However, large parts of the park have not been surveyed. Thai females. We have five females from a small muddy pond at Tung Lui Lai, Chaiyaphum Province, central Thailand collected by R. W. and J. A. Garrison. One female was collected in tandem with a male and is illustrated here (Fig. 4). It has somewhat longer pterostigmata: 2.21 and 2.14 as long as high along the ventral margin in the fore and hind wings, respectively (Fig. 4g, h), which fits the holotype. The synthoracic pattern and such structural characters as a pair of pale-coloured occipital (postocellar) tubercles, although very slight (Fig. 4b, c), also correspond to I. anomalus, and the postocellar light spots are confined to the immediate circumference of those (Fig. 4b, c). The coloration of the end of the abdomen is similar to the Cát Tiên female, with S8 somewhat darker; and the cerci too are as long as S10 (Fig. 4e). The posthumeral dark spots are reduced in comparison with the holotype and Cát Tiên female: the posterior one narrow but parallel-sided, without a ‘neck’, the middle one small, the anterior one missing (Fig. 4a). Measurements (N=5, in mm): Hw: 20–22.5, Abd. 27–31, total length: 35–38. The photos from Nam Nao National Park in neighbouring Phetchabun Province by Dennis Farrell show mature females (Fig. 5c, d) in which the synthorax has a saturated blue ground colour, while that of the prothorax and mesepisternum is brownish. The pale colour of S1–3 and S10 is also blue, while S4–9 is brownish with indistinct darker dorsal spots expanding at the segment joints, with blue present only on the anterior margins of S4–7; the eyes are brownish-grey. Teneral females on his photos (not reproduced) have a brownish-grey ground colour. Male. Since the male has not been described and its diagnostic characters remain unknown, we provide below a description of a male (Figs. 6, 7) collected in tandem with a female at Chayaphum Province of Thailand. Habitus. Resembles the female, ground colour pale-bluish (further referred to as ‘pale’), dark pattern bronzeblack (further ‘dark’). Head. Labrum, base of mandibles, genae and anteclypeus pale; postclypeus pale anteriorly, dark posteriorly, with a pair of large semicircular spots touching each other at base; epicranium and rear of head dark (Fig. 6b). Thorax. Prothorax pale dorsally, with following areas dark: a pair of dorsolateral spots on middle lobe, wash of brown along dorsal margin of pleuron and a pair of ill-defined dorsolateral brown spots on rounded posterior lobe (Fig. 6a,c). Synthorax (Fig. 6c, d). Mesepisternum with dark middorsal stripe occupying over half of mesepisternum, its sides parallel except for a small rounded lobe almost touching mesopleural suture at upper fifth and again at upper margin and with an offshoot extending posteriorly over mesopleural fossa; mesepimeron pale with following areas dark: irregular elongate spot extending posteriorly from mesopleural fossa and curving ventrally to level of semicircular spot of middorsal stripe, isolated irregular spot at middle of mesopleural suture and a small subtriangular spot at juncture of mesinfraepisternum, dorsal carina and upper fifth of interpleural and metapleural sutures dark; a narrow elongate dark spot bordering posterior margin of pale metinfraepisternum (Fig. 6c); venter of thorax as in Fig. 6d. Coxae pale but diffusely brownish anteriorly. Femora black, becoming brown externally (Fig. 6a, c). Wings hyaline; venation brownish. Postnodals: FW 10/11, Hw 10/10; RP 2 at Fw starts at postnodal 4/4, at Hw at postnodal 3/3. Pterostigmata: FW: 1.6 2–1.69/1.87–1.95 times as long as high (along dorsal/ventral side); HW 1.72–1.94/1.84–1.98 times as long as high, surmounting surmounting 1.5/cells in both wings. In one male specimen, costal vein at pterostigma is noticeably convex at FW and distinctly convex at HW; in the other male specimen it is only very slightly convex at both wings. Abdomen with the following dark markings (Fig. 6a): S1 with a small dorsal quadrate spot at basal 0.30 and with narrow apical ring; S3–7 with a long dorsal stripe beginning at basal tenth, its base moderately expanded proximally and with a quadrate expansion laterally at distal end (that of S2 shaped as an inverted key-hole), but almost interrupted dorsally by an incomplete apical ring; basal 0.60 of S8 black, distal 0.30 pale, narrowly divided above by thin inverted triangular dark spot; S9 with irregular dark narrow basal ring, otherwise pale; S10 entirely pale, with a deep triangular dorsal incision. Genital ligula (Fig. 7) of usual lestine form, basal segment ovate, ectal arm with small semihyaline coiled tip that is grooved entally and ectally. Cercus (Fig. 6 e–g) with proximal half pale, becoming blackish-brown distally, mesal margin with a small semicircular lobe followed distally by a strong posteromedially directed tooth, apical third of cercus slightly swollen and attenuate distally; paraproct semicircular, black, about one-fifth the length of cercus. Measurements (in mm): HW 20, Abdomen (including cerci) 32, total length 38 The other male specimen from the same series shows no significant differences. Mature males photographed by Dennis Farrell (Fig. 5a, b, d) have a saturated blue ground colour and dark-blue eyes with a green hue below, while these are brownish-grey in teneral ones (not shown); the expansion of the dorsal synthoracic dark stripe sometimes reaches the mesopleural suture and merges with the posterior posthumeral spot. Habitat. In Cát Tiên National Park in Vietnam the female was found in January, which is during the dry season, in seasonally humid tropical forest with dense and thorny undergrowth. In Chayaphum Province of Thailand the species was found by R.G. at a small muddy pond. In Phetchabun Province, in Nam Nao National park, the species was regularly found in forest ponds, including a large one with reeds, from January to August, sometimes as extremely abundant (Farrell 2018). Distribution. The species is presently known from three separated areas: northern Thailand (Tak, Lampang, Phetchabun and Chaiyaphum Provinces), southern Vietnam (Đỗng Nai and possibly Lâm Đỗng Provinces) and Cameron Highlands in Peninsular Malaysia (Fraser 1946; Day et al. 2012; Ng et al. 2011; Farrell 2018; this paper), as shown in Fig. 8. It was observed as abundant only in Phetchabun Province of Thailand (Farrell 2018). It can be expected in Cambodia, which lies between these regions. Diagnosis. Indolestes anomalus is characterized by the synthoracic pattern with (i) the middorsal black stripe having a small extension at ca. three-fourths of its length and posteriorly expanding to the mesopleural suture and (ii) three irregular and variable posthumeral dark spots: an elongate dorsal one followed by a smaller quadrate one and finally by a smaller triangular spot at the margin of mesinfraepisternum (this small spot may be missing) (Figs. 1a, 3a, 4a, 5, 6a, c). These spots rarely become confluent thus forming a broad irregular stripe confluent with the middorsal stripe (e. g. Ng et al. 2011: Fig. 1b). The synthoracic maculation of I. anomalus differs from that in any other Asian Indolestes spp. Indolestes indicus Fraser, 1922, I. gracilis gracilis (Hagen in Selys, 1862) and I. peregrinus (Ris, 1916) also have or may have a number of separate posthumeral spots, but in I. gracilis the dorsal dark stripe has straight margins whereas in I. indicus and I. peregrinus this stripe forms long trapezoid extensions medially rather than a small extension as in I. anomalus, but also expanding at the posterior end, like in I. anomalus (Fraser 1933; Asahina 1976). The male cerci of I. anomalus (Fig. 6 e–g) have attenuated apices similar to those of Indolestes cyaneus (Selys, 1862), I. gracilis, I. guizhouensis Zhou & Zhou, 2005 (if this is a bona fide species) and I. peregrinus among Asian species (Kosterin 2015), but differ clearly from those species by the much larger and prominent, rounded inner swellings. Females have a characteristic pair of pale tubercles at the occiput lateroposterior of the lateral ocelli (Figs. 3a, 4b, c). Females resemble those of I. peregrinus by pale S10 and cerci contrasting with dark S9 (Figs. 3c, 4a,d). Pterostigmata at its ventral margin 1.8–2.2 as long as high (Figs. 3b, 4f, g); that is shorter than in the related species (e.g. 3.0– 3.2 in I. peregrinus). In his description, Fraser (1946) especially stressed the short pterostigmata of the holotype but it is longer than he depicted and becomes a less conspicuous diagnostic character. Nevertheless, it is still relatively shorter than in all other Indolestes species. Indolestes anomalus most resembles I. peregrinus, with which it shares separated posthumeral spots in both sexes, but differs by its smaller size and a more posterior position of the prominence of the dorsal synthoracic, Published as part of Kosterin, Oleg E., Garrison, Rosser, Kompier, Tom & Farrell, Dennis, 2019, Taxonomic notes on Indolestes Fraser, 1922 (Lestidae, Zygoptera). 3. Male and clarified type locality of Indolestes anomalus Fraser, 1946, pp. 67-78 in Zootaxa 4555 (1) on pages 68-78, DOI: 10.11646/zootaxa.4555.1.5, http://zenodo.org/record/2624152, {"references":["Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x","Hamalainen, M. & Pinratana, A. (1999) Atlas of the Dragonflies of Thailand. Distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp.","Ng, J. F., Dow, R. W. & Choong, C. Y. (2011) New records of Odonata (Insecta) from the Cameron Highlands, with first records of two species for Malaysia. Journal of Science and Technology in Tropics, 7, 9 - 16.","Kimmins, D. E. (1966) A list of the Odonata types described by F. C. Fraser, now in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 18, 173 - 227.","Day, L., Farrell, D., Guenther, A., Hamalainen, M., Klimsa, E., Korshunov, A., Kosterin, O., Makbun N., Pelegrin, A., Roeder, U., Ruangrong R. & Vikhrev, N. (2012) New provincial records of Odonata from Thailand mostly based on photographs. Agrion, 16 (1), 16 - 24.","Dow, R. A. (2009) Indolestes anomalus. The IUCN Red List of Threatened Species. Version 2014.3. Available from: www. iucnredlist. org (accessed February 2018)","Hamalainen, M. (2015) From Echo maxima to Archineura maxima - a slow taxonomic process (Odonata: Calopterygidae). Notulae odonatologicae, 8 (6), 156 - 201.","Fraser, F. C. (1933) The fauna of British India, including Ceylon and Burma. Odonata. Uol. I. Taylor and Francis, London, 423 pp.","Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4526.4.8","Farrell, D. (2018) Dragonflies & Damselflies of Thailand. Available from: http: // thaiodonata. blogspot. ru / (accessed 19 February 2018)","Asahina, S. (1976) Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces. Kontyu, 44, 1 - 12."]}

Argia mauffrayi Garrison & Ram��n-Cabrera, n. sp. Figs. 1 (head, thorax, S1���4), 2 (S7���10 lateral), 3 (S5���10 dorsal); 4 (appendages mediodorsal); 5 (appendages lateral), 6 (cerci dorsal), 7 (genital ligula lateral), 8 (genital ligula entolateral); 9 (genital ligula ectal); 10 (thorax), 18 (map). Etymology. Named mauffrayi (Latinized name) in honor of the collector William F. Mauffray in recognition to his valuable contributions to the knowledge of Neotropical Odonata. Specimens examined. 2 ♂♂. Types. Holotype ♂: ECUADOR, Pichincha Province, San Miguel de Los Bancos Cant��n, Recinto Milpe, Milpe Bird Sanctuary, Mirador Uno Trail, 0.0333�� N, 78.8661 W, 1101m., 4 ix 2018, WFM leg. (ZSFQ); Paratype ♂: Esmeraldas Province, Reserva Canande, 0.5203�� N, 79.0866�� W, 470m, 15 v 2012, Jessica Ware & Melissa Sanchez leg. (RWG). A largely orange species with forcipate cercus armed terminally with a decumbent tooth and with a small unilobate paraproct. Description of holotype. Head (Fig. 1). Labrum, genae, clypeus and antefrons orange, epicranium black with small orange spot anterolateral to lateral ocellus, large orange postocular spots confluent laterally with margin of compound eye; scape orange, pedicel black; rear of head black above, pale orange below with an ill-defined isolated black spot above gena. Thorax. Prothorax orange except for anterior margin of anterior lobe black and dorsal portion of middle and hind lobes black with narrow middorsal stripe of orange on middle and posterior lobes. Mesothorax (Fig. 1) with mesepisternum orange, paler orange laterally, black parallel middorsal stripe as wide as pale antehumeral stripe; black humeral stripe divided at upper 0.60, the anterior branch a narrow line along mesopleural suture with mesopleural fossa black, the posterior branch gradually narrowing and ending before subalar carina; metapleural stripe black, slightly narrower than humeral stripe, metapleural carina edged with black; anterior half of mesepisternum and most of metepimeron black; venter of thorax pale orange. Wings. Slightly flavescent with venation black; pterostigma trapezoidal, dark brown, surmounting one cell in all wings; postnodals Fw 13/13, Hw 12/11; postquadrangular cells Fw 3/3, Hw 3/3; RP 2 at Fw 6/7, Hw 5/5. Coxae and trochanters orange, femora orange with tip becoming black, tarsi and armature black, tibiae orange, black basally. Abdomen (Figs. 1���3). Mostly orange; S1 orange with a wash of black at basal 0.30; S2 orange laterally with an ill-defined black stripe constricted medially, apical annulus black; S3 orange with a poorly defined narrow dark lateral stripe interrupted medially, annulus black; S4���6 similar to S3 but posterior darker portion of succeeding segments becoming more extensive and defined; S7 with orange basal annulus followed by dorsolateral black interrupted dorsally with orange terminating with black at apical fifth; S8 black dorsally, orange laterally (Fig. 2); S9 black with a small longitudinal tear-shaped, middorsal spot (Fig. 3); 10 entirely black; torus orange, appendages black. Genital ligula (as in Figs. 7 ��� 9). With a long stiff flagellum, and a shorter medial trilobed process, the two lateral lobes of which are triangular in ventral (ectal) view (as in Fig. 9), and the decumbent median lobe is quadrate apically (as in Fig. 8); the latter process with an evenly curved arcuate ridge; more basally at base of flagellum a pair of decumbent rounded lobes. Caudal appendages. Torifer orange, torus large, planar, roundly triangular (as in Fig. 4), occupying entire ventral margin of torifer and not overlapping bilobed epiproct; epiproct black, small, slightly bilobed (as in Fig. 6); cercus (as in Figs. 4���6) slightly longer than paraproct, about three times longer than wide, slightly swollen basally with medial basal margin slightly convex and slightly narrowing distally with a strongly acuminate mediolaterally directed tooth, dorsal surface of cercus with long cluster of thick hairs; mediodorsal surface of cercus slightly depressed along medial margin; paraproct unilobate, almost quadrate in lateral view with a narrow tuberculate tip (as in Fig. 5). Female. unknown. Measurements. Hw 19.0, abdomen 26.5, total length 33.0. Variation in paratype. Similar to holotype but with black humeral stripe divided at upper 0.70 (e. g. Fig. 10), wings slightly more flavescent and with small middorsal orange stripe confined to hind lobe of prothorax and S9 entirely black. Pterostigma surmounting one cell in all wings; postnodals: Fw 12���12, Hw 11���10; postquadrangular cells Fw 3���3, Hw 3���3; RP 2 at Fw 6���5, Hw 4���4. Dimensions. Hw 18.0, abdomen 24.5, total length 31.0. Diagnosis. This species is unique by overall orange coloration, mostly black S9 and 10, and genital ligula and caudal appendage morphology. The only other South American species of Argia with a largely orange body is A. infumata Selys but it differs greatly in genital and appendage morphology (Garrison & von Ellenrieder 2015). Genital ligula and appendage morphology of A. mauffrayi resemble those of A. cuspidata Garrison & von Ellenrieder, 2018 and A. acridens, although appendage morphology is most similar to that of A. acridens (Fig. 11���17). However, they differ by: 1) dorsal surface of cercus covered with thick hairs in A. mauffrayi (Figs. 4, 5) compared to being bare in A. acridens (Figs. 13, 14), and 2) the cercus in dorsal view being slightly forcipate, with basomedial margin only slightly convex in A. mauffrayi (Fig. 6) versus greatly expanded into a basomesal round lobe as in A. acridens (Fig. 15). Additionally, the torus in A. mauffrayi (Fig. 4) is roughly triangular but is smaller and more oval in A. acridens (Fig. 13). The apical process of the paraproct of A. mauffrayi in lateral view is small and minutely bilobed (Fig. 5) whereas the apical process in A. acridens is broader and roughly quadrate (Fig. 14). The basal trilobate process and paired lateral lobes in the genital ligula in A. mauffrayi (Figs. 7���9) are lacking in A. acridens (Figs. 16, 17). Remarks: Argia mauffrayi is unusual in its overall orange coloration and lack of any pale color on the terminal abdominal segments. Specimens are noticeable in the field as Sanchez, one of the collectors of the paratype, noted its orange color when reporting its capture to Garrison. Mauffray and Tennessen noted the following: "[The holotype] was collected at approx 9:30am along forested trail on a very steep hill with a few small sun-lit spots. It was collected as it flew upward and perched on a leaf in one of these sunny spots along with Argia variegata [F��rster], Argia oculata [Hagen in Selys] and Polythore gigantea [(Selys)]. There was no seepage nor stream at that location; however, there was a small seepage stream about 50 meters from the location. Ken and I continued to look for other specimens and even returned to the site about an hour later, but the weather had turned cloudy" and "..when I first saw the specimen flying���it appeared mostly orange���[and thought] it's a small Heteragrion !" Although the female of A. mauffrayi is unknown, we suspect that the mesostigmal plates will likely be poorly developed similar to the female sex of A. acridens, A. cuspidata (Garrison & von Ellenrieder 2018, Figs. 69 & 72 respectively) and A. mishuyaca Fraser, 1946 (Garrison & von Ellenrieder 2015, Fig. 7c). The curvilinear cercus in the males of these three species are longer than the paraproct and are armed terminally with a apical decumbent tooth���a condition similar to that illustrated here for A. mauffrayi. Argia mauffrayi, which is sympatric with A. acridens and A. cuspidata, is thus far known only from Esmeraldas and Pichincha Provinces in western Ecuador (Fig. 18)., Published as part of Garrison, Rosser W. & Ram��n Cabrera, Giovanni M., 2019, Argia mauffrayi n. sp. from Ecuador (Odonata: Coenagrionidae), pp. 286-292 in Zootaxa 4545 (2) on pages 287-291, DOI: 10.11646/zootaxa.4545.2.8, http://zenodo.org/record/2618856, {"references":["Garrison, R. W. & von Ellenrieder, N. (2015) Damselflies of the genus Argia of the Guiana Shield (Odonata: Coenagrionidae). Zootaxa, 4042 (1), 1 - 134. https: // doi. org / 10.11646 / zootaxa. 4042.1.1","Garrison, R. W. & von Ellenrieder, N. (2018) Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species. Zootaxa, 4470 (1), 1 - 69. https: // doi. org / 10.11646 / zootaxa. 4470.1.1"]}