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Bilton, David T., Gentili, Elio (2014): Laccobius leopardus sp. nov. from the Western Cape of South Africa (Coleoptera: Hydrophilidae). Zootaxa 3835 (3): 397-400, DOI: http://dx.doi.org/10.11646/zootaxa.3835.3.10

Laccobius leopardus sp. n. (Fig. 1) Type locality. South Africa: Western Cape: Cederberg, Matjies River at Matjiesrivier Reserve ca. 3 km N of Cederberg Oasis, 32 �� 31 ��� 10.53 ���S 19 �� 21 ��� 02.63���E, 726 m, D.T. Bilton leg. (Figure 1 f). Type material. Holotype (male): ��� 22���25 /ix/ 2011 South Africa WC, Cederberg���Matjiesrivier Reserve, Matjies River ca. 3 km N of, Cederberg Oasis D. T. Bilton leg.��� (genitalia extracted and mounted in DMHF on same card) and red holotype label (ISAM). Paratypes (9): 3 ♂, 4 ♀ Same data as holotype (CDTB, CGV, NMW); 2 ♀ ��� 21 /ix/ 2010 South Africa WC, Cederberg���Matjiesrivier Reserve, Matjies River ca. 3 km N of, Cederberg Oasis D. T. Bilton leg (CDTB).��� All with red paratype label. Description. Size: Holotype: TL = 3.5 mm, MW = 1.65 mm; Paratypes: TL = 3.4���3.7 mm, MW = 1.65���1.75 mm. Colour: Head, including labrum, dark brown with aeneous purplish-green reflections. Distinct pale preocular spots extending from front margins of eyes to frontal angles of clypeus. Pronotum pale straw yellow-yellowish brown, with approximately parallel-sided dark central patch occupying discal 1 / 3, and broadly reaching front and hind margins. Elytra straw yellow, with diffuse darker patches on disc, and darker rings of pigment around punctures. Venter black, with exception of preocular region of head, pronotal hypomeron and elytral epipleurs and pseudepipleurs, which are straw yellow as upperside, and lateral areas of abdominal ventrites 2���5 which are paler brown. Legs, antennae and palpi straw yellow; trace of infuscation visible on last segment of maxillary palp (see Fig. 1 a���b). Head: Labrum curved at lateral margins, but broadly truncately rounded at front edge. Upper surface of head, including labrum, shining and distinctly shagreened, with sparse, coarse shallow punctures. Frontoclypeal suture distinct, with coarse punctures posterior to suture and inside each eye, each puncture bearing a long, white decumbent hair-like seta. Eyes oval, moderately large, occupying 1 / 3 side margin of head. Inter-ocular distance ca. 3.2 x diameter of eye. Pronotum: Transverse; approximately 3 x broader than long. Broadly rounded at sides, with acutely rounded front angles, and broadly rounded hind angles. Front margin broadly bisinuate around centre; hind margin weakly curved. Upper surface shagreened, this microsculpture being slightly less impressed than on head, and most evident over central dark patch. With sparse, coarse shallow punctures throughout; these punctures being shallower than those on head. Larger punctures bearing long decumbent hair-like setae, visible particularly towards side margins. Elytra: Elongate oval, subparallel in basal half and evenly rounded to apex; widest at middle. Elytral punctures rather irregular, some traces of rows on elytral disc. Each puncture bearing a seta. On disc setae of punctures short; not reaching beyond hind margin of puncture. Setae much longer at sides and in apical ��, where many are 5��� 6 x puncture diameter in length. Elytral margin with stout bristles on shoulder, and similar, slightly longer, stout bristles in apical half. Ventral side: Labrum without speculae, front margin broadly curved, and overhanging oral cavity, especially in the centre. Distinct circular apicomedian emargination visible on front margin of labrum, this occupying the downcurved upper surface and front labral edge. Ventral surface of labrum somewhat excavated either side of apicomedian emargination. Apicomedian emargination with a cluster of short, golden setae. Mentum rectangular, shining, lacking microsculpture, and with fine, scattered punctures. Submentum with similar surface properties. Gula and genae microreticulate, dull, with hydrofuge pubescence. Genae rugosely punctate towards lateral margins of head, with distinct field of long, golden, curved, hair-like erect postocular setae. Prosternum roof-like, with a central longitudinal keel. Surface punctate, with hydrofuge pubescence. Mesoventral keel with a distinct hooked anterior tooth. Tooth surrounded by broad lateral expansions, furnished with long white hairs. Keel narrowed and linear between mesocoxae. Metaventrite pubescent, with exception of a glabrous central section. Hairs thinner towards posterior margin. Glabrous central section with distinct longitudinal furrow occupying posterior �� of metaventrite. Pubescence overlying distinct micoreticulation; imbricate and isodiametric anteriorly, transverse posteriorly. Abdominal ventrites 1���5 shining, with sparse, coarse punctures, some bearing long golden setae arranged in loose transverse rows. Setae 1 / 3 ��� 3 / 4 of segment length. Abdominal ventrites broadly excavated laterally. Ventrite 5 with denser punctation; some punctures situated in short transverse furrows. Ventrite 6 dull, pubescent, with transverse microreticulation. Legs: Moderately long and slender. Segments 2���3 of male protarsi expanded, with smooth sucker plate and modified bristle fields, typical of the genus. Pro and mesotibiae straight; posterior tibiae evenly curved. All tibiae strongly spinose. Procoxae, protrochanters and base of profemora with hydrofuge pubescence. Mesocoxae with hydrofuge pubescence, mesotrochanters and anterobasal portion of mesofemora with scattered long setae. Metacoxae glabrous. Base and posterior margins of metatrochanters and base of metafemora with stout hair-like setae. Aedeagus: Parameres 2.3 x length of phallobase. Paramere apices elongate, finger-like, extending beyond apex of median lobe. Median lobe with broadly rounded apex, and distinct, broad semicircular excavation on ventral face. Minutely dentate internal sac structures visible at base of this excavated area. Female: Concavity of abdominal ventrites weaker than in males. Foretarsi unmodified. Metatrochanters with much fewer, smaller setae than in male. Variability: In addition to size differences, specimens vary somewhat in the extent of the dark central patch on the pronotum (Fig. 1 a���b) and the extent of dark mottling on the elytra, which can be slightly more extensive than seen in the holotype. Differential diagnosis. L. leopardus sp. nov. is morphologically closest to Laccobius caffer, from which it can be distinguished by its larger size (TL 3.4���3.7 mm vs. 2.5���3.1 mm, see Gentili, 1981), darker dorsal pigmentation, with a broader and typically more parallel-sided central dark patch on the pronotum (see Fig 1 a���c). In addition the anterior labral margin is more evenly curved and less truncate dorsally in examined specimens of L. caffer, and the abdominal ventrites have more well developed paler patches in this species. In L. caffer the margins of abdominal ventrites 2���4, the posterior margin of ventrite 4 and much of abdominal ventrite 5 are pale yellow. The two species also differ significantly in male genitalia (Fig. 1 d���e). Distribution. To date only known from the type locality, the sandy gravelly margins of the Matjies River, on the edge of the Cederberg range in the Western Cape Province, South Africa. This locality is situated in a region which is somewhat transitional between the fynbos and succulent karoo biomes (Murcinia & Rutherford, 2006); fynbos occurring at higher altitudes, lower altitude sites such as the type locality being surrounded by succulent karoo. Etymology. Named in reference to the leopards (Panthera pardus pardus (Linnaeus)) which frequent the Cederberg Conservancy in which the type locality is situated, and the spotted appearance of the elytra. The name is a noun in apposition. Ecology. Specimens were puddled by hand from marginal sediment in shallow river backwaters (Fig. 1 f), which were mostly without appreciable surface flow at the time of sampling, but would clearly be connected to the main river at times of higher flow. They occurred together with the water beetles Hydropeplus montanus Omer-Cooper, Herophydrus inquinatus (Boheman), Sharphydrus brincki Bilton, Yola frontalis R��gimbart, Hydroglyphus infirmus (Boheman), H. lineolatus (Boheman), Canthyporus guignoti Omer-Cooper, C. petulans Guignot, Philaccolus lineatoguttatus (R��gimbart), Laccophilus lineatus Aub��, Georyssus spp., Berosus punctulatus Boheman, Laccobius revelieri Perris, L. venustus Gentili, Helochares spp., Crenitis zimmermanni Knisch, Parasthetops nigritus Perkins & J. Balfour-Browne, P. pampinus Perkins, P. rufulus Perkins & J. Balfour-Browne and Prosthetops wolfbergensis Bilton., Published as part of Bilton, David T. & Gentili, Elio, 2014, Laccobius leopardus sp. nov. from the Western Cape of South Africa (Coleoptera: Hydrophilidae), pp. 397-400 in Zootaxa 3835 (3) on pages 397-400, DOI: 10.11646/zootaxa.3835.3.10, http://zenodo.org/record/224747, {"references":["Gentili, E. (1981) Il genere Laccobius nella regione Etiopica (Coleoptera, Hydrophilidae). Annuario Ossevatorio Fisica Terrestre Museo Antonio Stoppani del Seminario Arcivescovile di Milano (N. S.), 3, 31 - 42.","Murcina, L. & Rutherford, M. C. (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19, 1 - 807."]}

Laccobius (s.str.) binotatus Orchymont, 1934 Material examined. Anhui: 1 male (SYSU): Woo-Fu (Wuhu), 29.v. 1932, without collector. Beijing: 4 males, 3 females (SYSU): Shisanling, 10.viii. 1962, Zhelong Pu lgt.; 1 female (SYSU): “Peiping”, no other data. Chongqing: 1 male, 2 females (SYSU): 8.iii. 1940, Zhelong Pu lgt. Gansu: 6 spec. (NMPC): Dagcanglhamo (=Langmusi) env., 34 °05.1ʹN 102 ° 37.9 ʹE, 3540 m a.s.l., stream and pools, 23.– 25.vi. 2005, J. Hájek, D. Král & J. Růžička lgt. Guizhou: 1 male (SYSU): Guizhou, 1939, Zhelong Pu lgt.; 1 male, 1 female (SYSU): Guiyang, Guizhou Normal University Campus, 15.viii. 1982, Zhihe Huang lgt.; 3 males, 4 females (SYSU): Pingba, horse farm, 13.viii. 1982, Zhihe Huang lgt. Heilongjian: 6 males, 5 females (SYSU): Shangzhi County, Maoershan Mt., 3.viii. 1963, Zhelong Pu lgt.; 1 male (SYSU): Wudalianchi, 7.viii. 2008, Fenglong Jia lgt. Hubei: 2 females (SYSU): Hankou, each one with label “A. d’Orchymont det. Laccobius (s. str.) binotatus ”. Hunan: 2 females (SYSU): Nanyue (Hengshan) Mt., 4.ix. 1941, Zhelong Pu lgt. Inner Mongolia: 1 male (SYSU): Huhehaote, viii. 1975, Xianwen Huang lgt.; 1 male (SYSU): Linxi County, Linxi town, 23.vii. 2008, Fenglong Jia lgt.; 1 female (SYSU): suburb of Jining, viii. 1975, without collector. Qinghai: 2 males (NMPC): Huangzhong env., Taer Si lamasery (= Kumbum), 36 ° 28.8–29.5 ʹ 101 °34.0– 34.1 ʹE, 2665–2780 m a.s.l., puddles, 17.vii. 2005, J. Hájek, D. Král & J. Růžička lgt. Shaanxi: 6 males, 6 females (SYSU): Xi’an, Dayu, 12.v. 2011, Fenglong Jia lgt.; 1 female (SYSU): Xi’an, Chaba River, 11.v. 2011, Fenglong Jia lgt.; 2 males, 4 females (SYSU): Huashan Mts., 24.viii. 1984, Zhihe Huang lgt.; 4 males, 1 female (SYSU): Chang’an, Xingguosi, 19.vii. 1984, Zhihe Huang lgt.; 3 males, 3 females (SYSU): Chang’an, Wutai, 23.viii. 1984, without collector; 1 male, 1 female (SYSU): Wugong, 29.iv. 1940, without collector. Sichuan: 2 males, 1 female (SYSU): Xikang, Xichang, 5.x. 1939, Fengying Zheng lgt.; 2 males, 2 females (SYSU): Emei Mt., 6.viii. 1982, Zhihe Huang lgt.; 1 male (SYSU): Qingchengshan Mt., 8.viii. 1982, Zhihe Huang lgt. Yunnan: 2 males (SYSU): Pohui, 2.ix. 1934, Zhelong Pu lgt.; 1 male (SYSU): Lufengcun, 26.iii. 1940, without collector; 5 males, 7 females (SYSU): Chengjiang, 15.v. 1940, Zhelong Pu lgt. Zhejiang: 9 males, 7 females (SYSU): Tianmushan Mt., 27.vii– 10.viii. 2009, Fenglong Jia lgt. Comments. Widely distributed in China, reaching to Russian Far East and Korean Peninsula. New for Chongqing, Gansu, Qinghai and Yunnan. Laccobius (s.str.) cinereus Motschulsky, 1860 Material examined. Inner Mongolia: 3 males, 5 females (SYSU): Hulunbeir, Zhalainur, 27.vii. 2003, Fenglong Jia lgt.; 3 males, 2 females (SYSU): Hailar, 23–26.vii. 2003, Fenglong Jia lgt. Qinghai: 2 males, 1 female (NMPC, MSNV): 7 km NE of Ulan, 3020 m a.s.l., 36 ° 57.6 ʹN 98 ° 30.6 ʹE, 7.vii. 2005, Hájek, Král & Růžička lgt.; 1 male, 1 female (SYSU): Yushu, Batang, vi. 1975, without collector; 1 male (SYSU): Yushu, Shangbatang, 4000 m a.s.l., 20.viii. 1974, Nanying Shen lgt. Shanxi: 1 male (SYSU): W of Datong, 15.vi. 1983, Zhelong Pu lgt. Comments. Distributed from Central Asia and Eastern Siberia to North China (Gentili 1995, 2003). New for Qinghai and Shanxi provinces.

Laccobius (Dimorpholaccobius) simulans Orchymont, 1923 Material examined. Sichuan: 1 male, 1 female (SYSU): Batang, Yidun, 3370 m a.s.l., 17.viii. 1982, Shuyong Wang lgt. Yunnan: 4 males, 3 females (NMPC, MSNV): 3 km W Daju, 27 �� 19.8 ��N 100 �� 14.4 ��E, 1685 m a.s.l., individually collected on gravel bank of a stream, 17���20.vi. 2007, J. H��jek & J.Růžička lgt.; 1 female (NMPC): Rehai hot springs, 6 km SW of Tengchong, 24 �� 57.1 ��N 098�� 26.2 ��E, 1400 m, individually collected in small pool with clay bottom, 5.vi. 2007, J. H��jek & J. Růžička lgt.; 21 spec. (SYSU): Chengjiang, 15���19.v. 1940, Zhelong Pu lgt.; 31 spec. (SYSU): Lufengcun, 24���26.iii. 1949, Zhelong Pu lgt.; 1 male, 2 females (SYSU): Kunming, 28.x. 1939, Zhelong Pu lgt.; 1 male (SYSU): Xinping, Mamushu cun, 24.021 ��N 102.128 ��E, 1127 m a.s.l., 21.v. 2011, Keqing Song lgt. Comments. Distributed from northwestern India (Punjab) through Himalaya and Yunnan to central Sichuan. New for Sichuan Province., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on page 411, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Dimorpholaccobius) bipunctatus (Fabricius, 1775) Material examined. Xinjiang: 12 spec. (SYSU, MSNV): Wulumuqi, Liudaowan, 18.vii. 1984, lgt. Zhihe Huang; 2 spec. (SYSU): Wulumuqi City, 18.vii. 1984, lgt. Zhihe Huang; 1 spec. (SYSU): Yishadong, 22.vii. 1984, lgt. Zhihe Huang; 1 spec. (SYSU): Chabuchar town, 4.viii. 1984, lgt. Zhihe Huang. Distribution. Widely distributed from North Africa (Tunisia) through Europe and Central Asia to Xinjiang, China. First record for China., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on pages 408-409, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Cyclolaccobius) hingstoni Orchymont, 1926 (Fig. 17) Material examined. Xizang: 2 males, 3 females (CASB): Yangbajing, ca. 4220 m a.s.l., 22.v. 1960, Wang Chunguang lgt.; 1 female (SYSU): suburb of Rikaze, 3826 m a.s.l., 20���23.vii. 1986, Liang Geqiu lgt. Comments. Known from Tibet Plateau and Nepal. Here (at SYSU) we are providing additional records from eastern Xizang. Laccobius (Cyclolaccobius) nitidus Gentili, 1984 Material examined. Hunan: 1 male (NHMW): ca. 25 km N of Pingjiang City, 28 �� 50 �� 10 ��N 113 �� 38 ��0 5 ��E, ca. 200 m a.s.l., 20.iii. 2003, Sch��nmann, Komarek & Wang lgt. (CWBS 500). Jiangxi: 1 male (NHMW): Luang Shi Peng Mt., 29 �� 26 �� 58 ��N 115 �� 58 �� 48 ��E, ca. 250 m, 26.iii. 2003, Sch��nmann, Komarek & Wang lgt. (CWBS 512); 4 males, 7 females (SYSU): Shangrao Sanqingshan Mt., 15���20.iv. 2007, Fenglong Jia lgt. Shaanxi: 4 males, 8 females (NMPC, MSNV): Hua Shan Mts., 110 km NEE Xian, Huayin village, 34 �� 31.2 ��N 110 ��04.6��E, ca. 530 m a.s.l., river valley, 10.v. 2011, M. Balke & J. H��jek lgt.; 1 female (NMPC): Qinlingshan Mts., ca. 30 km SSW of Xian, Fengyukou, 34 ��01.4��N 108 �� 49.2 ��E, 580 m a.s.l., wet cliff and waterfall, 12.v. 2011, M. Balke & J. H��jek lgt.; 4 males, 5 females (NMPC): Hua Shan Mts., 110 km NEE Xian, Huayin village, 34 �� 29.5 ��N 110 ��05.1��E, 1275 m a.s.l., granite cliff, 8���9.v. 2011, M. Balke & Jiř�� H��jek lgt.; 2 males, 2 females (SYSU): Hua Shan Mts., 24.viii. 1984, Zh.H. Huang lgt.; 2 males, 2 females (SYSU): Hua Shan Mts., the old road, 9.v. 2011, Fenglong Jia lgt.; 1 male, 3 females (SYSU): Qinlingshan Mts., Fengyukou, wet cliff and waterfall, 12.v. 2011, Fenglong Jia lgt. Zhejiang: 12 males, 10 females (SYSU): Tianmushan Mt., 27���28.vii. 2009, Fenglong Jia lgt. Comments. So far known from Anhui, Fujian, Hunan, Shaanxi, Sichuan and Zhejiang (Gentili 1995, 2003)., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on pages 409-410, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Microlaccobius) florens Gentili, 1979 Material examined. Guangdong: 1 male, 2 females (SYSU): Xinyi, 7.vi. 1960, without collector. Guangxi: 2 males, 2 females (SYSU): Yangshuo, 1985, Shoujian Chen lgt. Comments. Distributed in the maritime areas of East and South China. New for the Guangxi province., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on page 413, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Dimorpholaccobius) striatulus (Fabricius, 1801) Material examined. Xinjiang: 3 spec. (SYSU, MSNV): Kashi, 7.viii. 2006, lgt. Ling Zhao; 1 ex. (SYSU): Huocheng, marshland, 27.vii. 2005, lgt. Ling Zhao. Distribution. Widely distributed from Europe through Central Asia to Xinjiang, China. First record for China., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on page 409, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Microlaccobius) elegans Gentili, 1979 Material examined. Sichuan: 10 males 7 females (SYSU): Xikang, Kangding, 2.viii. 1939, Fengying Zheng lgt. Yunnan: 10 males, 14 females (SYSU): Chengjiang, 15.v. 1940, Zhelong Pu lgt.; 1 male (SYSU): Lufengcun, 25.vii. 1939, Zhelong Pu lgt.; 1 male, 2 females (SYSU): Yingjiang, 850 m a.s.l., 25.v. 1983, Lizhong Hua lgt. Comments. Distributed from SE Asia (Laos, Thailand, Vietnam) to South and Central China (Fujian, Henan, Shaanxi, Shandong, Sichuan,Yunnan)., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on page 413, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Jia, Fenglong, Gentili, Elio, Fikáček, Martin (2013): The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae). Zootaxa 3635 (4): 402-418, DOI: http://dx.doi.org/10.11646/zootaxa.3635.4.4

Laccobius (Cyclolaccobius) politus Gentili, 1979 (Figs. 14���16) Material examined. Yunnan: 1 male, 1 female (NMPC): Gudong env., Yunfeng Shan Mt., 25 �� 22.7 ��N 98 �� 25.4 ��E, 1825 m a.s.l., collected individually in rapid stream at margin of mixed forest with Pinus, Quercus and Rhododendron, 8.vi. 2007, J. H��jek & J. Růžička lgt. Comments. Gentili (2012) synonymized Oocyclus flaveolus Hebauer & Wang, 1998, with L. politus. So far known from Taiwan only. New for Yunnan Province. Laccobius (Cyclolaccobius) yunnanensis Gentili, 2003 Material examined. Yunnan: 1 male (NMPC): Rehai hot springs, 6 km SW of Tengchong, 24 �� 57.1 ��N 098�� 26.2 ��E, 1400 m, individually collected in small pool with clay bottom, 5.vi. 2007, J. H��jek & J. Růžička lgt. Comments. Described from Myanmar and southern Yunnan. The above specimen is the second known locality of L. yunnanensis in China., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on pages 410-411, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Cyclolaccobius) hainanensis sp. nov. (Figs. 8���13) Type locality. China, Hainan Island, Limushan Mts., 19 �� 9.1���9.2 ��N 109 �� 45���46 ��E, 550���750 m a.s.l. Type material. Holotype: ♂ (SYSU): China: Hainan Isl. [MF 18] / Limushan Mts., 5.v. 2011 / 19 �� 9.1���9.2 ��N 109 �� 45���46 ��E / 550���750 m; along the road / Fik��ček & Zhao lgt. // seepages with thin algae / layer + nearly bare wet rock / without distinct algae along / most remote end of road. Paratypes (32): 31 spec. (SYSU, NMPC, MSNV): same data as holotype; 1 male (MSNV): China, Hainan Prov. / Limushan, Limu Temple / 5.v. 2011, Shuan Zhao leg. Description. Total length 1.8���2.1 mm (holotype 2.0 mm); maximum width 1.2���1.4 mm (holotype 1.3 mm). Total length / total width ratio = 1.5. Body oval, moderately convex, with greatest width in anterior third of elytra, continually narrowing more posteriad. Head. Labrum black, without specula in both sexes, slightly convex on anterior margin, closely adhering to clypeus; surface densely punctured, anteriorly with stiff setae. Clypeus and frons shiny black, preocular pale spots extremely small and vague; surface with irregular, scarcely arranged fine punctures, punctures slightly impressed but larger than labral punctures, each puncture with decumbent seta; microsculpture absent. Clypeus nearly 1.5 �� as wide as long, deeply emarginate anteriorly; frontoclypeal suture scarcely distinct. Eyes transversely oblique, nearly 2.5 �� as large as long, reniform posteriorly, closest to each other posteriorly than anteriorly; minimum interocular distance in frontal view 2.2 �� of width of one eye. Ventral surface black, except the brown maxillae. Mentum flat, uniformly microreticulated, with scarce punctures. Postmentum smooth, gula granulate. Maxillary palpi yellowish; palpomere 3 inflated, palpomere 4 asymmetrical, straight medially and convex laterally, scarcely infuscate at extreme apex. Antennae with 8 antennomeres, yellowish with darker club; scape (antennomere 1) longer than antennomeres 2���4 combined, pedicel (antennomere 2) cone-shaped, two intermediate antennomeres (antennomeres 3���4) very short, cupule (antennomere 5) asymmetrical, oval ventrad, crescent-shaped and emarginated dorsad, antennomere 7 expanded, larger than remaining club antennomeres, antennomere 8 constricted near apex. Thorax. Pronotum black, laterally with a yellowish irregular band widening posteriorly, external borders slightly explanate; microsculpture absent, punctation consisting of fine sparsely arranged setiferous punctures, only with irregular serial arrangement of punctures along margins. Prosternum with narrow longitudinal carina. Scutellar shield equilaterally triangular, black, impunctate. Mesoventrite with longitudinal keel, anteriorly forming a beak. Elytra combined ca. as long as wide (i.e., elytral index = 1.0), black or dark on disc, yellowish along lateral margins, lateral pale stripe slightly widening anteriorly, more widening posteriorly, very narrow at midlength. Elytral punctation consisting of fine setiferous punctures arranged in ca. twenty longitudinal rows, with alternating series of larger and denser punctures (primary rows) and more irregularly distributed finer punctures (secondary rows). Parasutural stria absent, fifth elytral row slightly sulciform (Fig. 8). Microsculpture absent. Lateral margins of elytra slightly explanate with a narrow bead. Epipleuron ending at level of metafemora. Metaventrite with decumbent pubescence, which is absent from a small medioposterior area. Legs. Trochanters dark brown, femora dark brown except apical part, tibiae and tarsi yellowish brown. Femora pubescent basally, metafemur with coarser setiferous punctures. Tibiae with ranges of spines more developed on outer margin and on mesotibiae, metatibiae curved. Tarsal natatory setae nearly absent. Legs with five tarsomeres; protarsomere 5 nearly 3 �� as long as other protarsomeres; mesotarsomeres 2 and 5 each nearly 2 �� as long as other mesotarsomeres; metatarsomeres 1 and 2 each longer than other metatarsomeres. Claws shorter than tarsomere 5, arcuate. Abdomen. Ventrites nearly smooth, sparsely pubescent; ventrite 5 and 6 wrinkled, pubescence denser on ventrite 6 than on ventrites 1���5. Aedeagus. Total length 0.45 mm. Parameres nearly 1.5 x as long as phallobase. Phallobase 1.3 x as long as wide (Figs. 10���12). Parameres slightly narrowing from base to apex, lateral and medial margins very slightly arcuate; distance between lateral margins of phallobase slightly wider than distance between lateral margins of parameres; apex of parameres asymmetrical, obtusely pointed. Median lobe as long as parameres, conically narrowing from base to midlength, then widened, conspicuously narrowing to wide subdivided apex (hence the apex lanceolate in shape). Differential diagnosis. Laccobius hainanensis sp. nov. is very close to L. politus Gentili, 1979, but may be distinguished from it by the wide lateral pale stripe on elytra suddenly widening near the elytral apex (more or less uniformly wide in L. politus), the elytral puncture rows more conspicuous (scarcely evident in L. politus), fifth elytral row slightly sulciform (not sulciform in L. politus), the primary elytral puncture rows conspicuously setiferous (setae inconspicuous or barely conspicuous in L. politus), the median lobe constricted subapically, weakly narrowing apicad (indistinctly constricted subapically and widely rounded at apex in L. politus), and parameres wider, flat (not rod-like) and with pointed apex (parameres narrower and widely rounded apically in L. politus). The Chinese species of the subgenus Cyclolaccobius may be identified according to the following key (modified from Gentili 1995, 2003): 1. Elytral punctures irregularly arranged, scattered; body length over 3 mm.......................................... 2 - Elytra with longitudinal rows of punctures; body length under 3 mm............................................. 3 2. Body blackish, convex in lateral view; head and pronotum smooth; male with specula; parameres longer than median lobe. 3.4���4.3 mm..................................................................... L. zugmayeri Knisch, 1910 - Body testaceous, depressed in lateral view; head and pronotum shagreened; male without specula; parameres shorter than median lobe (Fig. 26). 3.0��� 3.2 mm................................................ L. hingstoni Orchymont, 1926 3. Elytral base with a small yellow spot; median lobe longer than parameres, swollen in the basal half, narrow in apical half. 1.8 ���2.0 mm................................................................... L. yunnanensis Gentili, 2003 - Elytral base entirely dark................................................................................ 4 4. Body length 2.0��� 2.8 mm. Elytral rows with punctures larger, very conspicuous at 40 �� magnification. Median lobe narrow throughout, narrower than parameres in its whole length..................................... L. nitidus Gentili, 1984 - Body length 1.8���2.1 mm. Elytral rows of punctures hardly conspicuous at 40 �� magnification. Median lobe widened at base and near apex......................................................................................... 5 5. Elytral primary puncture rows conspicuously setiferous at 100 ��; fifth row slightly sulciform. Elytral pale lateral strip constricted at midlength, then abruptly widening near elytral apex. Median lobe constricted subapically, weakly narrowing apicad................................................................................. L. hainanensis sp. nov. - Elytral primary puncture rows not or scarcely setiferous at 100 ��; fifth row normal, not sulciform. Elytral pale lateral strip uniformly wide, only widening near elytral apex. Median lobe indistinctly constricted subapically, widely rounded at apex........................................................................................ L. politus Gentili, 1979 Biology. The type specimens were collected on a seepage and wet rocks along a road through secondary tropical forest. The habitat contained patches with a thin algae layer as well as areas of wet rock without any apparent algae. Etymology. The species name refers to Hainan Island and Province, where the species was collected. Distribution. Hainan Island. Known only from the type locality in Limushan Mts. in central Hainan., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on pages 406-408, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Glyptolaccobius) qinlingensis sp. nov. (Figs. 1–7) Type locality. China, Shaanxi Province, Qinlingshan Mts., Fengyukou, 34 °01.4ʹN 108 ° 49.2 ʹE, 580 m a.s.l. (the series including the holotype, collected by the first author, was taken simultaneously and in the same spot that was collected by Balke & Hájek, hence we provide the complete data on the type locality combining the label data of both these series). Type material. Holotype: ♂ (SYSU): CHINA, Shaanxi, Qinling, Qinlingliang, Fengyukou, 12.v. 2011, lgt. Fenglong Jia [transcribed from Chinese]. Paratypes (58): 32 spec. (SYSU): same data as the holotype; 2 spec. (MSNV): China, Shaanxi, Prov. / Huashan, rock of old road / 12.v. 2011 / Fenglong Jia leg.; 9 spec. (NMPC, MSNV): CHINA: Shaanxi prov. 12.v. 2011 / ca. 30 km SSW of Xian / Qinlingshan Mts., Fengyukou / (wet cliff, waterfall), 34 °01.4ʹN / 108 ° 49.2 ʹE, 580 m / M. Balke & J. Hájek lgt.; 12 spec. (NMPC, MSNV): CHINA: Shaanxi Prov. 10.v. 2011 / 110 km NNE Xian, Huayin vill. / Hua Shan Mt. (river valley), / ca. 530 m, 34 ° 31.2 ʹN 10 °04.6ʹE / M. Balke & J. Hájek lgt.; 3 spec. (NMPC): CHINA: Shaanxi prov., 8–9.v. 2011 / 110 km NEE Xian, Huyain vill. / Hua Shan Mt. (granite cliff) / 1275 m, 34 ° 29.5 ʹN 110 °05.1ʹE / M. Balke & J. Hájek lgt. Description. Total length 2.7–2.9 mm (holotype: 2.8 mm), total width 1.4–1.5 mm (holotype: 1.5 mm). Body oval, moderately convex, maximum width in anterior elytral third. Total length / total width ratio = 1.9. Dorsal surface finely pubescent. Head. Coloration black, without pale preocular spots, surface with dense and coarse punctures, without microsculpture; ‘systematic punctures’ indistinct; periocular sulci vaguely defined, reaching anterolateral branches of frontoclypeal suture. Labrum blackish, about 2.3 × as wide as long, without specula in both sexes; anterior margin very feebly emarginated medially; lateral portions sharply angulated. Anterior margin of clypeus deeply excised. Eyes oblong, oblique, closest to each other posteriorly, slightly protruding laterad, separated by distance equal to 2.35 × of width of one eye in dorsal view; posterior margin of eye nearly straight in dorsal view, distinctly reniform in lateral view; postocular portion of tempora short. Mentum strongly rugose, bearing few coarse punctures, not emarginate anteriorly. Submentum not microgranulate. Gula microgranulate, gular sutures widely separated, tentorial pits moderately large, very distinct. Maxillary palpi yellowish brown, dark at apex; palpomere 1 thin and short; palpomeres 2 and 3 nearly equal in length, palpomere 3 dilated at apex; palpomere 4 elongate, nearly 1.4 × as long as palpomere 3, asymmetrical, inner margin straight and outer margin convex, apex truncate. Antenna with eight antennomeres; scape (antennomere 1) longer than antennomeres 2–4 combined; pedicel (antennomere 2) cone-shaped; two intermediate antennomeres (antennomeres 3–4) very short; cupule (antennomere 5) asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, bearing densely arranged setae, antennomeres 6 and 8 nearly equal in length, antennomere 6 widest, antennomere 7 shortest and narrowest. Thorax. Pronotum transverse, slightly impressed sublaterally, 0.4 × as long as wide; black with yellowish lateral margins; yellow area about 0.5 × as wide as posterior margin of eye along base, sometimes yellow edge vaguely defined from black disc; surface smooth as that on head, without microsculpture; punctures coarse and dense, bearing decumbent whitish setae. Prosternum black, tectiform, with longitudinal keel. Scutellar shield equilaterally triangular, black, with few punctures. Mesoventrite with longitudinal, anteriorly arrow-shaped keel. Elytra combined slightly elongate, ca. 1.2 × as long as wide, blackish with yellowish lateral margins and apices; without pale yellow basal spot; 10 longitudinal rows of regularly impressed primary punctures alternating with 10 rows (interstriae) of sparser and more or less irregular secondary punctures; secondary punctures of same size as primary punctures; all punctures bearing long whitish semierect setae. Parasutural furrow reaching anterior third of elytral length, very conspicuous on posterior elytral third and elytral apex. Epipleura and pseudoepipleura oblique, nearly vertical. Metaventrite setiferous with an oval median glabrous area and a longitudinal sulcus posteromedially. Legs. Procoxae and trochanters pubescent; profemora setiferous on basal half, smooth distally, with tibial grooves; protibiae smooth, each bearing stiff setae and two apical spurs. Mesofemora smooth, with tibial grooves; ventral side of mesotibiae with three longitudinal rows of stiff setae and sulcus between two longitudinal ridges. Metatrochanters smooth and shining, only very finely pubescent mesally; metafemora smooth, each with scarce punctures and tibial grooves; metatibiae curved, ventral side of each tibia with three longitudinal rows of stiff setae and longitudinal sulcus between two ridges. Natatory setae nearly absent on dorsal surface of tarsi. Metatarsi with second tarsomere as long as third and fourth combined; fourth metatarsomere about 0.65 × as long as third tarsomere. Abdomen. Ventrites 1–4 smooth, without microsculpture, with transverse series of setae; last two ventrites densely pubescent. Aedeagus (Figs. 3–5). Total length 0.6 mm. Parameres nearly 1.65 × as long as phallobase. Median lobe almost as long as parameres, with a series of stout spines laterally, its apex rounded, subdivided medially. Differential diagnosis. Laccobius qinlingensis sp. nov. is easily distinguished from other species of Glyptolaccobius by the combination of following characters: body large (2.7–2.9 mm long); head without preocular spots; pronotum with pale lateral margins; mesoventrite with longitudinal keel, arrow-shaped anteriorly; parasutural stria of elytra distinct, reaching anterior third of elytral length; elytra with 10 rows of distinct primary punctures and somewhat irregular secondary rows of punctures that are of the same size as primary rows; lateral margins of elytra and elytral apex pale; pale spots near scutellum absent; elytral rows of punctures not sulciform; mentum rugose. Besides the external characters, Laccobius qinlingensis sp. nov. differs from all Glyptolaccobius by the median lobe bearing lateral subapical rows of stout spines (Fig. 3, 5). These spines are unique for L. quinlingensis; no similar structure is known in any other Laccobius species except L. (Yateberosus) elevatus Fauvel, 1883 from New Caledonia, although the spines are much thinner in that species (for details, see Gentili 1988). The key to the species of the subgenus Glyptolaccobius published by Gentili & Fikáček (2009) should be modified as follows: 4. Entire elytra shining black, without yellow lateral margins. Apex of median lobe widely expanded (Gentili & Fikáček 2009: Fig. 4). Body length 2.3–2.5 mm........................................ L. moriyai Kamite, Ogata & Hikida, 2007 – At least lateral margins and apex of elytra yellowish. Apex of median lobe widened or not. Body length less than 2.3 mm or more than 2.7 mm.................................................................................... 4 a 4 a. Body length 2.7–2.9 mm. Posterior pronotal margin yellowish laterally, median lobe somewhat widened before apex, with a series or large spine-like projections laterally (Figs. 3, 5).................................... L. qinlingensis sp. nov. – Body length smaller than 2.3 mm. Posterior pronotal margin yellowish or entirely black, median lobe without a series of spinelike projections laterally................................................................................ 5 Etymology. The species name refers to the Qinlingshan Mts. in which all known localities of this species are situated. Biology. Part of the type series was collected on wet granite rocks at the bottom of the waterfall in Fengyukou (Fig. 6) or in small hygropetric “streamlets” flowing over the granite cliffs of the Huashan Mts. near Huayin village (Fig. 7) (in the latter case together with Aspidytes wrasei Balke, Ribera & Beutel, 2003 (Adephaga: Aspidytidae)). Second part of the type series (in Fengyukou and in river valley near Huayin village) was collected on wet rocks at sides of the roads (the rocks were exposed during the construction of the road). Distribution. Only known from three localities in the granite massif of the Qinlingshan Mts.

Laccobius (Microlaccobius) nepalensis Gentili, 1982 Material examined. Hainan: 2 males, 2 females (SYSU): Tongshi, 19.xii. 1957, Cuiying Li lgt.; 1 female (SYSU): Xinglong, 24.xii. 1957, Cuiying Li lgt. Comments. Reaching from Himalaya (Nepal, West Bengal) to SE Asia (Laos, Thailand) and Hainan island., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on page 416, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Cyclolaccobius) martini sp. nov. (Figs. 1���9) Type locality. China, Guangxi, Shiwandashan Mts., 2189900 ˚N, 107.90570 ˚E. Type material. Holotype: ♂ (SYSU): China: Guangxi, Shiwandashan Mt., in the upper part of Shitouhe River (=Stone River), Tri-channel river, in sands of water margin among large stones, 21.89900˚N, 107.90570 ˚E, 339 m a.s.l., 9.vii. 2011, Keqing Song leg. (with Chinese and English labels). Paratypes: 12 ♂♂, 9 ♀♀ (SYSU, MSNV, IZCAS): same data as holotype. Differential diagnosis. Laccobius martini sp. nov. is very close to L. hainanensis Jia, Gentili & Fik��ček, 2013, but can be distinguished from it by the whole dorsal surface with white setae (yellowish brown setae in L. hainanensis), the setae forming 10 white lines along primary punctural rows (specify the state of this character in L. hainanensis here), secondary rows of punctures somewhat sparser, and the median lobe of aedeagus not constricted subapically (constricted subapically in L. hainanensis). The new species can be distinguished from L. politus Gentili, 1979, another similar species, by the wider lateral pale stripe on elytra suddenly widening near the elytral apex (more or less uniformly wide in L. politus), the elytral puncture rows more conspicuous (scarcely evident in L. politus), the primary rows with dense and conspicuous white setae that form 10 lines along primary punctural rows (setae inconspicuous or barely conspicuous in L. politus), and parameres wider than median lobe (parameres narrower in L. politus). Description. Total length 1.8���2.1 mm (holotype 2.0 mm); maximum width 1.3���1.4 mm (holotype 1.3 mm). Total length / total width ratio = 1.5. Body oval, moderately convex, with greatest width in anterior third of elytra, evenly narrowed posteriad (Fig. 1). Head. Labrum black, without specula in both sexes, slightly convex on anterior margin, closely adhering to clypeus; surface densely punctured, anteriorly with stiff white setae. Clypeus and frons shiny black, preocular pale spots extremely small and vague or completely absent (Fig. 3); surface with irregular, sparsely arranged fine punctures, punctures slightly impressed but larger than labral punctures, each puncture with decumbent white seta; microsculpture absent. Clypeus nearly 1.5 �� as wide as long, deeply emarginate anteriorly; frontoclypeal suture scarcely apparent. Eyes transversely oblique, nearly 2.5 �� as large as long, reniform posteriorly (Fig. 3), closer to each other posteriorly than anteriorly; minimum interocular distance in frontal view 2.3 �� of width of one eye. Ventral surface black, except the brown maxillae. Mentum flat, uniformly microreticulated, with sparse punctures. Postmentum smooth, gula granulate. Maxillary palpi yellowish; palpomere 3 inflated, palpomere 4 asymmetrical, straight medially and convex laterally, scarcely infuscate at extreme apex. Labial palpi same colour as maxillary palpi. Antennae with 8 antennomeres, yellowish with darker club; scape (antennomere 1) longer than antennomeres 2���4 combined, pedicel (antennomere 2) cone-shaped, two intermediate antennomeres (antennomeres 3���4) very short, cupule (antennomere 5) asymmetrical, oval ventrad, crescent-shaped and emarginated dorsad, antennomere 7 expanded, larger than remaining club antennomeres, antennomere 8 constricted near apex. Thorax. Pronotum black with green reflection, laterally with a sinuate yellowish irregular band widening posteriorly (Fig. 3), external borders slightly explanate; microsculpture absent, punctation consisting of fine sparsely arranged white-setiferous punctures, only with irregular serial arrangement of punctures along margins. Prosternum with narrow longitudinal carina medially. Scutellar shield equilaterally triangular, black with strong green reflection, with a few punctures. Mesoventrite with longitudinal keel, anteriorly forming a ���beak���. Elytra combined ca. as long as wide (i.e., elytral index = 1.0), black on disc, yellowish along lateral margins, lateral pale stripe slightly widening anteriorly, more widening posteriorly, very narrow at midlength. Elytral punctation consisting of fine setiferous punctures (each bearing a white seta) arranged in ca. twenty longitudinal rows, with alternating series of larger and denser punctures (primary rows) and more irregularly and sparsely distributed finer punctures (secondary rows), the setae forming 10 white lines along primary punctural rows (Figs. 1���2). Parasutural furrow absent. Microsculpture absent. Lateral margins of elytra slightly explanate with a narrow bead. Epipleura ending at level of metafemora. Metaventrite with decumbent pubescence (Fig. 4), which is absent from a small medioposterior area. Legs. Coxae dark brown, trochanters and other segments of legs yellowish brown. Trochanters with sparse pubescence, apical part of metatrochanter not abutted to femur. Femora pubescent basally, metafemur with coarser setiferous punctures. Tibiae with rows of spines more developed on outer margin and on mesotibiae, metatibiae curved. Tarsal natatory setae nearly absent. Legs with five tarsomeres; protarsomere 5 nearly 3 �� as long as other protarsomeres; mesotarsomeres 2 and 5 each nearly 2 �� as long as other mesotarsomeres; metatarsomeres 1 and 2 each longer than other metatarsomeres. Claws shorter than tarsomere 5, arcuate. Male with basal 2 protarsomeres dilated. Abdomen. Ventrites nearly smooth, sparsely pubescent, covered with broken-glass-like material; ventrite 5 and 6 wrinkled, pubescence denser on ventrite 6 than on ventrites 1���5. Aedeagus. Total length 0.42 mm. Parameres nearly 1.9 x as long as phallobase. Phallobase 1.2 x as long as wide (Figs. 5���6). Parameres slightly narrowing from base to apex, lateral and medial margins very slightly arcuate; distance between lateral margins of phallobase almost as wide as distance between lateral margins of parameres; apex of parameres asymmetrical, obtusely pointed. Median lobe as long as parameres, conically narrowing from base to midlength, then almost parallel, conspicuously narrowing apically. Sternite 9 as in Fig. 7. Etymology. Named in honour of Dr. Martin Fik��ček, Department of Entomology, National Museum, Czech Republic, and Department of Zoology, Faculty of Science, Charles University in Prague, a hydrophilid specialist who has collaborated with the first and third authors many times. Biology. All known specimens were collected in an aquatic situation, living underwater in the sand at a river edge (Figs. 8���9). When the sand underwater is stirred, the beetles float up to water���s surface and swim on the surface or swim down quickly. When they reach the substrate, they immediately walk downwards and hide beneath it. Remarks. The key to Chinese species of the subgenus Cyclolaccobius (Jia et al. 2013) may be modified as following: 5 a. Dorsum with white setae, the setae forming 10 white lines along primary punctural rows, secondary rows of punctures somewhat sparser............................................................................ L. martini sp. nov. - Dorsum with yellowish brown setae or without conspicuous setae, the setae not forming lines along primary punctural rows, secondary rows of punctures somewhat denser............................................................. 5 b 5 b. Elytral primary puncture rows conspicuously setiferous at 100 ��; fifth row slightly sulciform. Elytral pale lateral strip constricted at midlength, then suddenly widening near elytral apex. Median lobe constricted subapically, weakly narrowing apicad.............................................................. L. hainanensis Jia, Gentili & Fik��ček, 2013 - Elytral primary puncture rows not or scarcely setiferous at 100 ��; fifth row normal, not sulciform. Elytral pale lateral strip uniformly wide, only widening near elytral apex. Median lobe indistinctly constricted subapically, widely rounded at apex........................................................................................ L. politus Gentili, 1979, Published as part of Jia, Fenglong, Song, Keqing & Gentili, Elio, 2013, A new species of Laccobius Erichson, 1837 from China (Coleoptera: Hydrophilidae), pp. 91-95 in Zootaxa 3734 (1) on pages 92-95, DOI: 10.11646/zootaxa.3734.1.11, http://zenodo.org/record/285187

Jia, Fenglong, Song, Keqing, Gentili, Elio (2013): A new species of Laccobius Erichson, 1837 from China (Coleoptera: Hydrophilidae). Zootaxa 3734 (1): 91-95, DOI: http://dx.doi.org/10.11646/zootaxa.3734.1.11

Laccobius (Cyclolaccobius) Gentili, 1991 Laccobius (Cyclolaccobius) Gentili, 1991: 381. Type species: Laccobius rectus Sharp, 1890 (by original designation). Diagnosis. The following character combination can be used to separate Cyclolaccobius from other subgenera of Laccobius Erichson: 1) eyes reniform and oblique in dorsal view (see Gentili 1995: fig. 1); 2) pronotum broadest at base; 3) hind tibiae clearly curved; 4) elytra without parasutural furrow; 5) tarsal natatory setae nearly absent., Published as part of Jia, Fenglong, Song, Keqing & Gentili, Elio, 2013, A new species of Laccobius Erichson, 1837 from China (Coleoptera: Hydrophilidae), pp. 91-95 in Zootaxa 3734 (1) on page 92, DOI: 10.11646/zootaxa.3734.1.11, http://zenodo.org/record/285187

Laccobius (Compsolaccobius) pallidissimus Reitter, 1899 Material examined. Xinjiang: 41 exs. (32 exs. SYSU, 8 exs. MSNV, 1 ex. NMPC): Chabuchaer, 4.viii. 1984, lgt. Zhihe Huang; 1 ex. (SYSU): Yili Farmland, 2.viii. 1984, lgt. Zhihe Huang; 3 exs. (SYSU): Kuerle city, 20.vii. 1984, lgt. Zhihe Huang; 11 exs. (SYSU): Kuerle, Yishadong, 22.vii. 1984, lgt. Zhihe Huang; 1 ex. (SYSU): Bazhou, Yishadong, 22.vii. 1984, lgt. Zhihe Huang; 2 exs. (SYSU): Jinghe, Gaoquan, 31.vi. 1984, lgt. Zhihe Huang; 26 exs. (SYSU): Yili Agricultural School, 3.viii. 1984, lgt. Zhihe Huang. Distribution. Known from Afghanistan, Tajikistan, Turkmenistan, Uzbekistan. First record for China. Laccobius (Microlaccobius) tonkinensis Gentili, 1979 Material examined. Shaanxi: 2 females (NHMW): Qinling Shan Mts., river bank at road Zhouzhi-Foping, 96 km WSW of Xi���an, 33 �� 53 ��N 108 ��02��E, 950 m a.s.l., in fine to coarse gravel at river bank, 4.vii. 2001, Wrase lgt. Distribution. Known from Laos, Thailand and Vietnam. First record for China., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on page 409, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Laccobius (Compsolaccobius) decorus (Gyllenhal, 1827) Material examined. Qinghai: 1 male (NMPC): 13.4 km NW of Golmud, 36 �� 25.7 ��N 94 �� 45.9 ��E, 2800 m a.s.l., saline, 8.vii. 2005, H��jek, Kr��l & Růžička lgt. Distribution. Widely distributed from North Europe (Baltic area) through Ukraine and Russia (West Siberia) and Central Asia (Afghanistan, Mongolia) to northwestern China (Qingshai). First record for China., Published as part of Jia, Fenglong, Gentili, Elio & Fik����ek, Martin, 2013, The genus Laccobius in China: new species and new records (Coleoptera: Hydrophilidae), pp. 402-418 in Zootaxa 3635 (4) on page 409, DOI: 10.11646/zootaxa.3635.4.4, http://zenodo.org/record/219563

Georissus (Neogeorissus) nemo sp. nov. (Figs. 8, 9–14, 17–18) Type locality. Yemen, Socotra Island, Hallah Arhar, 12°33.0′N 54°27.6′E, 15 m a.s.l. Type material. HOLOTYPE: J (NMPC): ‘ YEMEN: Socotra Isl. / Hallah Arhar (spring) / 12°33.0′N 54°27.6′E, 15 m / 11.xi.2010, leg. J. Hájek’. PARATYPES: 1 ♀, 6 spec. (NMPC, KSEM, NHMW, IRSNB): same label data as the holotype; 62 spec. (BMNH; NMPC, KSEM, 10 spec. in DNA grade in coll. NMPC): ‘ YEMEN: Socotra island / Halla area, Arher; freshwater / spring in sand dune / 9.-10.+ 15.vi.2012 / 12°33.0′N 54°27.6′E, 5 m // Socotra Expedition 2012 / J. Bezděk, J. Hájek, V. Hula / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart’; 1 spec. (NMPC): ‘ YEMEN: Socotra Island / ca. 3 km NE of Shuab / Avicennia marina mangrove; / sand dunes, 20.-21.vi.2012 / 12°34.1′N 53°23.9′E, 3 m // SOCOTRA expedition 2012 / J. Bezděk, J. Hájek, V. Hula / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart leg. ’. Description. Body weakly elongate, lowly convex in lateral view. Body length 1.3–1.5 mm (holotype 1.4 mm), width of head 0.35–0.45 mm (holotype 0.37 mm), maximum width of pronotum 0.5–0.6 mm (holotype 0.5 mm), maximum width of elytra 0.75–0.95 mm (holotype 0.80 mm). Coloration elytra and pronotum reddish brown to brown with olive reflections, head dark brown. Head (Fig. 10). Clypeus weakly convex, with scattered sparsely arranged granules, anterior margin with marginal row of densely arranged granules; each posterolateral portion of clypeus anteriorly to eye declined, divided from mesal portion by ridge arising from frons, ridge high and sharp posteriorly, but more weakly defined anteriorly before joining with marginal clypeal row of granules. Frons with central elongate depression surrounded by elevated ridges each bearing several weakly pronounced granules; anterior portion of frons with sharp sublateral ridge without granules at midlength between central depression and inner margin of each eye (this ridges continues to clypeus more anteriorly); submesal portion with transverse blunt ridge connecting midlength of central ridge with posterior portion of sublateral ridge. Eyes large, oval in lateral view. Prothorax (Figs. 11–12). Pronotum 1.1× as wide as long, with maximum width at posterior 0.4; lateral portions very small, slightly projecting laterad as blunt elongate lobes only. Anterior portion with two closely situated submedian ridges delimiting shallow median groove, each side laterally of ridges weakly convex, with scattered granules. Central portion of pronotum with large but rather shallow rhomboid depression delimited by wide granuliferous ridges anteriorly and narrow ones posteriorly, posterior ridges not joining but converging to posterior margin; posterolaterally of central depression with pair of large but shallow impressions delimited laterally by small granuliferous bulge; posterolateral portion of pronotum each with high granuliferous protuberance. Posterolateral pits absent. Ventral morphology of prothorax corresponding with G. crenulatus. Mesothorax. Elytra (Figs. 9, 13) combined 1.2× as long as wide, 2.2× as long as pronotum; base of elytra ca. as wide as maximum width of pronotum, maximum width of elytra between anterior 0.1–0.5, elytra gradually narrowing in posterior 0.5–0.2, apex strongly narrowed. Elytral suture and intervals 2, 4 and 6 elevated into high narrow ridges, ridge on interval 6 arising from large humeral protuberance; lateral portion with Y-shaped structure formed by highly elevate interval 8 and anterior portion of interval 9; odd intervals (1, 3, 5) flat, not elevate, interval 7 flat anteriorly and becoming slightly elevate posteriorly. All ridges and humeral protuberance with very weak and low granules only, hence the elytral ridges nearly smooth; odd elytral intervals only with scattered and extremely low granules, hence appearing nearly totally flat. Elytral series regular, serial punctures small but sharply impressed. Lateral-most portion of elytron declined, hence elytral laterally without clear projections. Median pentagonal protuberance of mesoventrite flat, without distinct pits. Metathorax. Metaventrite ca. 2× as long as mesoventrite, flat, only with few scattered indistinct granules, without distinct median discrimen. Metathoracic wings present in specimens examined for this character (n=2). Abdomen (Fig. 14) gradually narrowing posteriad. Ventrites 1 flat, only with sparsely arranged indistinct granules especially in posterior third (anterior portion of some specimens totally bare), ventrites 2–4 without granules, ventrite 5 flat with few indistinct granules along posterior margin. Male genitalia (Fig. 8). Aedeagus 0.4 mm long (measured in holotype only). Parameres 0.85× as long as phallobase, their combined width slightly narrower than maximum width of phallobase; lateral margins of parameres very slightly convergent in basal 0.6, arcuately bent inward apically; apex widely obtusely pointed; inner margins straight, well sclerotized. Median lobe 0.75× length of parameres, narrowly triangular apically, apex sharp, gonopore situated subapical, struts ca. 0.4× as long as apical portion of median lobe. Phallobase slightly widening posteriad, with wide but indistinct latero-posterior band along margins, without posterior opening. Differential diagnosis. Based on the general sculpture of the pronotum (i.e., rhomboid central depression and low submedian and higher lateral tubercles, Fig. 11, 17) and elytra (i.e., odd intervals more elevated than even ones), Georissus nemo sp. nov. belongs to the G. costatus species group sensu DELÈVE (1967a,b). It may be distinguished from other species of the group by the combination of the following characters: elytral ridges highly elevated, very weakly denticulate in lateral view; intervals 2, 4 and 6 nearly completely flat with very few indistinct granules; lateral portions of pronotum not projecting into acute lobes; abdomen with very indistinct granules on ventrites 1 and 5 only; phallobase slightly longer than parameres, only indistinctly widened posteriad; parameres widely arcuate apically. By elytral morphology, G. nemo sp. nov. especially resembles the African species G. alticosta Grouvelle, 1909 and G. decorsei Paulian & Legros, 1943, and G. decoratus Delève, 1972 from Sri Lanka, all of which may be easily distinguished by the morphology of the aedeagus which is much narrower in both African species (DELÈVE 1967a, Fig. 8) and much wider in G. decoratus (DELÈVE 1972, Fig. 16). Georisus nemo sp. nov. also differs from both African species by the indistinctly denticulate elytral costae (costae are totally smooth in G. alticosta and G. decorsei) and from G. decoratus by flat elytral intervals 2 and 4 (slightly convex in the latter species). Georissus nemo sp. nov. may be easily distinguished from the following unidentified species from Socotra Island by its smaller body size (the other species is 1.8–2.0 mm long), elytral costae with very indistinct denticulation (strongly denticulate in the other species), elytral intervals 2 and 4 flat (convex and bearing many distinct granules in the other species), abdominal ventrites with extremely indistinct granulation (with very distict granulation on whole ventrites 1 and 5 in the other species), and by the weakly metallic coloration (head and pronotum strongly and elytra moderately metallic in the other species). From the third Socotran species of Georissus, G. maritimus sp. nov., it differs by its costate odd elytral intervals. Etymology. The species name refers to Captain Nemo, a fictional character of two novels by Jules Verne, who lived underseas (in a submarine Nautilus), hence in an environment unusual for a human. This resembles specimens of Georissus nemo sp. nov. collected in 2010 which were found underwater, in an environment unusual for this genus. Collection circumstances. The vast majority of specimens in the type series were collected in Arhar along the permanent stream rising below the sand dunes on the base of rock cliffs of the Socotra Plateau falling to the sea coast. The specimens inhabited the sandy waterlogged surroundings of the stream partly overgrown by short-grazed lawn of few undetermined Poaceae and Cyperaceae and surrounded by shrubs of Tamarix nilotica (Ehrenb.) Bunge (Tamaricaceae). The majority of the specimens were collected at night, creeping on the bare wet sand, while a portion of the specimens were attracted at light trap installed close to the stream. In 2010, when the locality was only shortly visited in the daytime, few specimens were found on the submerged underside of the stones directly in the stream. One specimen was also found among a large number of G. maritimus sp. nov. from the type locality of the latter species (see above). Distribution. Known from two distant localities on Socotra Island, indicating that the species may be widely distributed on suitable habitats of the island. Notes. Although the relationships between the species of Georissus Latreille, 1809 are poorly known and the species groups defined by the pronotal and elytral sculpture may be easily polyphyletic, the strong resemblance of G. nemo sp. nov. to some African and Sri Lankan species (see Differential diagnosis) is rather striking. Moreover, there is a long series of an undescribed Georissus species in NMPC which was collected in southern India (Tamil Nadu state, S of Tuticorin) in salt marshes at the Támbrapathi river estuary, hence in a habitat somewhat resembling that of G. nemo sp. nov. The Indian species is very similar to G. nemo sp. nov. by dorsal sculpture, male genitalia and highly reduced abdominal granulation, although it shows some weak differences from G. nemo sp. nov. (elytral intervals 2 and 4 slightly convex, body coloration strongly metallic) and therefore seems to represent a separate species. Even though further studies of the Indian specimens are needed to completely understand their identity, the strong resemblance between both taxa suggests that G. nemo may possibly represent a South Indian element of Socotran fauna.

Sternolophus (Sternolophus) unicolor Laporte de Castelnau, 1840 Material examined. 1 ♀ (CULS): wadi Far, 1.iv.2001, lgt. J. Farkač; 1 J (NMPC): Firmin, x.2000, lgt. V. Bejček & K. Šťastný; 2 spec. (CULS, NMPC): Calanthia, 29.–30.iii.2001, lgt. J. Farkač; 2 JJ, 1 spec. (CULS): Ayhaft, 15.iii.2000, lgt. J. Farkač; 1 spec. (NMPC): wadi Ayhaft, 12°36.5′N 53°58.9′E, 200 m a.s.l., 7.–8.xi.2010, lgt. J. Hájek; 1 spec. (NMPC): same locality and date, lgt. P. Hlaváč; 3 spec. (NMPC): Firmihin plateau, 12°28′46″N 54°01′E, 400–500 m a.s.l., 18.–19.vi.2010, lgt. V. Hula & J. Niedobová; 1 spec. (IRSNB): hills near Hadibu, 29.ii.2008, G. 31.496, lgt. A. Saldaitis; 8 spec. (CULS, NMPC): wadi Faar, 12.433°N 54.195°E, 69 m a.s.l., 3.xii.2000, lgt. V. Bejček & K. Šťastný. Distribution. African species widely distributed in Madagascar and east Africa. First record from Socotra Island. Note. WRANIK (2003) lists the Near East/Arabian species S. decens Zaitzev, 1909 for the Socotran fauna, but this record seems to be based on the misidentification. The taxonomy of the genus Sternolophus Solier, 1834 is still not properly resolved and no modern revision exists for the Old World species of the genus. The Socotran specimens clearly differ from S. decens by the diagnostic characters given in the identification key by ZAITZEV (1909): they are generally wider than the Arabian specimens of S. decens and bear a longer and stouter metaventral spine. They agree well with the specimens of S. unicolor from Madagascar present in the collection of NMPC. Few Sternolophus specimens with a wider general body form from southern Yemen are present in the collection of NMPC, but they clearly differ from the Socotran specimens by a shortened and apically blunt metaventral spine and may be conspecific with the Yemeni specimens which BALFOUR- BROWNE (1951) identified as possibly belonging to S. solieri Laporte de Castelnau, 1840., Published as part of Fikáček, Martin, Delgado, Juan A. & Gentili, Elio, 2012, The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae), pp. 107-130 in Acta Entomologica Musei Nationalis Pragae 52 on page 125, DOI: 10.5281/zenodo.5334580, {"references":["WRANIK W. 2003: Fauna of the Socotra Archipelago. Field guide. Universitat Rostock, Rostock, 542 pp.","ZAITZEV F. A. 1909: Analytische Uebersicht der mir bekannten Arten der Gattung Sternolophus Solier nebst Bemerkungen uber die anderen Arten dieser Gattung (Coleoptera, Hydrophilidae). Russkoe Entomologicheskoe Obozrenie 8 (1908): 228 - 233."]}

Laccobius (Microlaccobius) minor (Wollaston, 1867) Material examined. 1 J, 1 ♀ (NMPC): Qadub, 12°38.3′N 53°57.3′E, saline, 14.vi.2012, Socotra Expedition 2012 lgt. Distribution. Laccobius minor is a species widespread in savannah and semidesert areas of Africa, reaching northwards to Yemen, southern Saudi Arabia, Israel and Lebanon (GENTILI 1981, 1988, 1989; HEBAUER 1994). First record from Socotra Island. Note. The Socotran specimens were collected in shallow exposed pools in the salt marsh ca. 20 m of the seacoast., Published as part of Fikáček, Martin, Delgado, Juan A. & Gentili, Elio, 2012, The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae), pp. 107-130 in Acta Entomologica Musei Nationalis Pragae 52 on page 124, DOI: 10.5281/zenodo.5334580, {"references":["GENTILI E. 1981: Il genere Laccobius nella regione etiopica (Coleoptera: Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (N. S.) 3 (1980): 31 - 42.","GENTILI E. 1988: Verso una revisione del genere Laccobius (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (N. S.) 9 (1986): 31 - 47.","GENTILI E. 1989: The Laccobius (Coleoptera, Hydrophilidae) of the Arabian Peninsula and Sinai. Fauna of Saudi Arabia 10: 95 - 102.","HEBAUER F. 1994: The Hydrophiloidea of Israel and the Sinai (Coleoptera, Hydrophiloidea). Zoology in the Middle East 10: 73 - 137."]}

Laccobius (Microlaccobius) eximius Kuwert, 1890 Material examined. 6 JJ, 8 ♀♀ (NMPC, MSNV): Haghier Mts., Wadi Madar, 12°33.2′ N, 54°00.4′E, 1180–1230 m, 12–14.xi.2010, lgt. Jiří Hájek. Distribution. The distribution of the species is unclear due to changes of its taxonomic position by various authors until it was confirmed as a separate species by FIKÁČEK et al. (2010) (see p. 147 in the latter paper for history of the taxonomic concepts of the species). The species was described from ‘Hejaz’ and since then recorded from various parts of the Arabian Peninsula (HEBAUER 1997, as L. praecipuus) and Egypt (KUWERT 1890), but most these records require confirmation as they may concern the widespread L. praecipuus. So far, the only specimens examined by us reliably belonging to L. eximius were all collected in western Saudi Arabia at high altitudes (ca. 2000 m a.s.l.) of the mountain range along the Red Sea. First record from Socotra Island. Note. Laccobius eximius was found at a single locality at high altitude in Socotra Island. This corresponds to its distribution on the Arabian Peninsula, where the species inhabits the mountains in the western part of Saudi Arabia where it replaces the common lowland species L. praecipuus. The dorsal coloration of the Socotran specimens is paler than in the majority of the Saudi Arabian specimens which are nearly completely dark (see FIKÁČEK et al. 2010, Fig. 11). Pale specimens of L. eximius easily differ from L. praecipuus by darker pronotum (uniformly black on a larger surface) and elytral longitudinal puncture rows uniformly darkened (sometimes these rows join one another forming a nearly black elytral surface). The morphology of the aedeagus of the Socotran specimens agrees in all details with those from the Arabian Peninsula (see FIKÁČEK et al. 2010, Figs. 7–9) and L. eximius is therefore easily distinguished from L. praecipuus by genital morphology. Socotran specimens of L. eximius are generally smaller than those from the Arabian Peninsula., Published as part of Fikáček, Martin, Delgado, Juan A. & Gentili, Elio, 2012, The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae), pp. 107-130 in Acta Entomologica Musei Nationalis Pragae 52 on page 124, DOI: 10.5281/zenodo.5334580, {"references":["KUWERT A. 1890: Bestimmungs-Tabellen der europaischen Coleopteren. XIX. Heft. Hydrophilidae. I. Abteilung: Hydrophilini. Verhandlungen des naturforschenden Vereins in Brunn 28 (1889): 1 - 121.","FIKACEK M., GENTILI E. & SHORT A. E. Z. 2010: Order Coleoptera, family Hydrophilidae. Pp. 135 - 165. In: HARTEN A. VAN (ed.): Arthropod fauna of the UAE, volume 3. Multiply Marketing Consultancy Services, Abu Dhabi, 700 pp.","HEBAUER F. 1997: Annotated checklist of the Hydrophilidae and Helophoridae (Insecta: Coleoptera) of the Arabian Peninsula with a description of a new genus and species. Fauna of Saudi Arabia 16: 255 - 276."]}

Georissus (Neogeorissus) maritimus sp. nov. (Figs. 1–7, 15–16, 19–20) Type locality. Yemen, Socotra Island, ca. 3 km NE of Shuab, 12°34.1′N 53°23.9′E, 3 m a.s.l. Type material. HOLOTYPE: J (NMPC): ‘ YEMEN: Socotra island / ca. 3 km NE of Shuab / mangrove, Avicennia marina / 12°34.1′N 53°23.9′E; 3 m a.s.l. / saline, 20-21.vi.2012 // Socotra Expedition 2012 / J. Bezděk, J. Hájek, V. Hula, / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart leg. ’. PARATYPES: 4 JJ, 1 ♀, 41 spec. (BMNH, IRSNB, NHMW, NMPC, KSEM, 5 spec. in DNA grade kept in NMPC): same label data as the holotype. Description. Body narrowly elongate, weakly convex in lateral view. Body length 1.20– 1.60 mm (holotype 1.40 mm), width of head 0.30–0.40 mm (holotype 0.35 mm), maximum width of pronotum 0.45–0.60 mm (holotype 0.55 mm), maximum width of elytra 0.65–0.85 mm (holotype 0.75 mm). Coloration of elytra brown to piceous, pronotum dark brown with paler anterior portion, head piceous to black. Head (Fig. 2). Clypeus weakly convex, with few weakly developed granules anteromedially, anterior margin with elevated bead without distinct granules; each side of clypeus with deep pit anterolaterally; lateral portion of clypeus posteriorly divided from mesal portion by high ridge without granules, ridges bent laterad and bearing more or less distinct granules more anteriorly. Frons elevated above clypeus, bearing large elongate rhomboid plate mesally reaching to posterior part of frons; laterally of mesal plate with high and wide ridge on each side; posterior part of median rhomboid and longitudinal ridges connected by group or large and low granules posteriorly; frons with two pairs of deep pits, one situated anteromesally, second laterally. Clypeus and median portion of frons with weak mesh-like microsculpture. Antennal club with three antennomeres. Prothorax (Figs. 3–4). Pronotum 1.15× as wide as long, with maximum width at midlength; lateral portions rounded, without distinct projections. Median portion with two closely situated submedian ridges throughout, delimiting shallow median groove; each lateral lobe divided from mesal portion by longitudinal ridge falling into separate granules posteriorly. Anterior half of pronotum without granules, only bearing small pits intermixed with sparsely arranged punctures. Posterior half of pronotum without distinct bulges, with sparsely arranged large but low granules throughout the surface. Pronotum with two pairs of deep pits, one anteriorly and the other posteriorly of lateral lobe. Whole surface between granules with weak meashlike microsculpture. Ventral portion of prothorax with extremelly large antennal grooves, otherwise corresponding with G. crenulatus (Rossi, 1794). Mesothorax. Elytra (Figs. 1, 5) combined 1.25× as long as wide, 1.75× as long as pronotum; base of elytra ca. as wide as maximum width of pronotum, maximum width of elytra in anterior third, then weakly narrowing posteriad to posterior fifth, apex strongly narrowed. Elytra completely devoid of granules, interstices without microsculpture. Elytral suture and intervals 1–7 evently convex, interval 8 reduced, interval 9 slightly elevated; lateral portion with suboval depression at anterior third across intervals 8–9; all intervals more convex posteriorly than anteriorly. Elytral series 1–11 regular, serial punctures small but sharply impressed. Lateral-most portion of elytron declined, hence elytra laterally without clear projections. Median pentagonal protuberance of mesoventrite flat, with distinct pits. Metathorax. Metaventrite ca. 1.2× as long as mesoventrite, flat, with few scattered indistinct granules along posterior margin and four deep pits along anterior margin; lateral portions divided from mesal part by longitudinal ridges. Metathoracic wings absent in specimens examined for this character (n=5). Abdomen (Fig. 6) gradually narrowing posteriad. Ventrite 1 flat mesally, with lateral portion declines and divided by ridge; median portion with sparsely arranged punctures, without granules; posterior margin of ventrite 1 with pair of large tubercles facing enlarged granules of ventrite 2. Ventrites 3–4 with weakly developed granules along anterior margin. Male genitalia (Fig. 7). Aedeagus 0.35–0.45 mm long. Parameres 1.6× as long as phallobase, their combined width narrower than maximum width of phallobase; parameres nearly parallel-sided throughout except subbasally, subbasally slightly constricted; apex rounded at outer margin, bluntly pointed mesally; apex with numerous large pores, ventral portion of paramere with numerous sparsely arranged micropores. Median lobe 0.70× length of parameres, narrowly triangular apically, apex sharp, gonopore situated subapically, struts ca. 0.25× as long as apical portion of median lobe. Phallobase widening posteriad, without large posterior opening. Differential diagnosis. Based on the evenly convex elytral intervals, Georissus maritimus sp. nov. may be easily diagnosed from the species of the G. costatus and G. caelatus species groups sensu DELÈVE (1967a,b). It may be easily distinguished from the majority of the species with equally elevated elytral intervals by the absence of a median rhomboid impression on the pronotum, which it shares with the African species G. sordidus Grouvelle, 1915 and G. bicolor Grouvelle, 1909. The latter two species may be however immediately distinguished from G. maritimus sp. nov. by much larger and wider pale-coloured body, morphology of male genitalia and the sculpture of the abdomen (see DELÈVE 1967a), as well as by the sculpture of the pronotum which lacks any bulges in the posterior half in G. maritimus sp. nov. In fact, Georissus maritimus sp. nov. seems to be unique among all described species of the genus by its pronotal sculpture, i.e. the combination of the absence of a median rhomboid depression, absence of granules in anterior half of the pronotum and absence of elevated granulate bulges posteriorly. By the extremely long and narrow parameres and posteriorly widening phallobase, The new species resembles G. alticosta Grouvelle, 1909 (which has costate even elytral intervals), G. alluaudi Delève, 1967 (which has the pronotum with a rhomboid central depression), G. acutecostatus Fairmaire, 1898 and G. biroi Delève, 1969 (both with costate even elytral intervals). Etymology. The species name refers to the type locality of this species, which is on the bank of a brackish lagoon situated at the estuary of a temporary stream to the Arabian Sea. Collection circumstances. The type series was collected at night on a sandbar between the sea and a brackish lake in an estuary of a temporary stream (Figs. 19–20). The specimens were collected on places without vegetation which were wet due to the seepage of the subsurface water. The surrounding vegetation (which usually began within ca. 2 meters of the sites with Georissus maritimus sp. nov.) consisted of the low succulent shrubs of Arthrocnemum macrostachyum (Moric.) Moris (Amaranthaceae), Limonium socotranum (Vierh.) Radcl. -Sm. (Plumbaginaceae) and isolated groups of mangrove trees Avicennia marina (Forssk.) Vierh. (Avicenniaceae). No specimen of Georissus maritimus sp. nov. bore the substrate layer on the body dorsum. Distribution. Known only from the type locality. Note. The unique morphology of G. maritimus sp. nov. makes it impossible to assign this species to any of the species groups proposed by DELÈVE (1967a,b) and it seems to be rather isolated from the known species of the genus. Further studies are thus necessary to understand the geographic origin of the species., Published as part of Fikáček, Martin, Delgado, Juan A. & Gentili, Elio, 2012, The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae), pp. 107-130 in Acta Entomologica Musei Nationalis Pragae 52 on pages 109-112, DOI: 10.5281/zenodo.5334580, {"references":["DELEVE J. 1967 a: Notes sur les Georissus d'Afrique et descriptions d'especes nouvelles (Coleoptera Georissidae). Bulletin de l ' Institute Royal des Sciences Naturelles de Belgique 43 (26): 1 - 23."]}

Cercyon (Cercyon) nigriceps (Marsham, 1802) Material examined. 1 J, 1 ♀, 5 spec. (NMPC): Al Haghier Mts., wadi Madar, 12°33.2′N 54°00.4′, 1180–1230 m a.s.l., 12.–14.xi.2010, J. Hájek lgt.; 2 spec. (NMPC): Al Hagier Mts., W slopes, Skant area, 12°35′52″N 54°00′01″, 1240 m a.s.l., 2.xii.2003, D. Král lgt.; 1 spec. (NMPC): Hadiboh env., 12°65′02″N 54°02′04″E, ca. 10–100 m a.s.l., 21.xi.–12.xii.2003, D. Král lgt.; 1 spec. (NMPC): Zemhon area, 12°30′58″N 54°06′39″E, 270–350 m a.s.l., 3.–4.ii.2010, Purchart & Vybíral lgt.; 2 spec. (CULS): Dixam plateau, Sirhin area, 12°31′08″N 53°59′09″E, 812 m a.s.l., 1.–2.xii.2003, J. Farkač lgt.; 1 spec. (NMPC): same label data, but P. Kabátek lgt.; 4 spec. (NMPC): Dixam plateau, Firmihin (Dracaena forest), 12°28.6′N 54°01.1′E, 490 m a.s.l., 15.–16.xi.2010, J. Bezděk lgt; 1 spec. (PLCL): wadi Ayhaft, 28.ii.–1.iii.2009, P. Lo Cascio & F. Grita lgt.; 6 spec. (NMPC): Dixiam plateau, wadi Esgego, 12°28′09″N 54°00′36″E, 300 m a.s.l., 2.–3.xii.2003, lgt. D. Král. Distribution. Originally most probably Oriental (S. Ryndevich, pers. comm.), this species was introduced during or prior to the 19th century to tropical and subtropical areas all over the world, and occurs rarely in adjacent temperate regions. Previously recorded from Socotra Island by WRANIK (2003)., Published as part of Fikáček, Martin, Delgado, Juan A. & Gentili, Elio, 2012, The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae), pp. 107-130 in Acta Entomologica Musei Nationalis Pragae 52 on page 127, DOI: 10.5281/zenodo.5334580, {"references":["WRANIK W. 2003: Fauna of the Socotra Archipelago. Field guide. Universitat Rostock, Rostock, 542 pp."]}

Laccobius (Glyptolaccobius) sipeki sp. nov. (Figs. 6, 10) Type locality. India, Meghalaya, Khasi Hills, Laitkynsew, 11 km SW of Cherrapunjee. Type material. HOLOTYPE: J (NMPC): ���INDIA, Meghalaya State (6) / E Khasi Hills, 11 km SW Cherra- / punjee, Laitkynsew, 21���24.IV. / 2008, 25��12���N, 91��40���E, 460 m / Fik��ček, Podskalsk��, ����pek lgt. // wet rock with algae/ blue algae / and fallen leaves at side of / waterfall on small river surround. / by tropical forest, ca. 200m / upstream from living bridge���. PARATYPE: INDIA: MEGHALAYA: 1 J (NHMW): ���INDIA: Meghalaya / SW of Cherrapunjee / 25��13������14���N 91��40���E, 900m / P. Pachol��tko leg. / 5.���24.v.2005 ���. Description. Small-sized species, body length 1.85 mm, width 1.09 mm. Maximum length / maximum width ratio 1.7. Body oval, moderately convex. Head. Labrum of males without specula; anteriorly and posteriorly crescent-shaped in dorsal view, shiny black. Clypeus moderately convex, shiny black, with sparse and faint punctures, without distinct ���systematic punctures���. Only terminal branches of frontoclypeal suture conspicuous. Frons shiny black with sparse and faint punctures as those on clypeus. Eyes oblique, weakly convex, not protruding laterad, separated by distance equal to 2.5�� of width of one eye, nearly reniform in lateral view, emarginated posteriorly. Maxillary palpi less than 0.5�� as long as head width; palpomere 2 slightly swollen, nearly as long as palpomere 3; palpomere 4 1.5�� as long as palpomere 3, outer and inner margin rounded, with ogival apex. Mentum ca 0.5�� as wide as long, almost flat, at least medially, covered with faint punctures. Eight antennomeres; scape longer than antennomeres 2���4 combined; pedicel cone-shaped; two intermediate antennomeres (antennomeres 3���4) very short; cupule asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, with antennomeres 6 and 8 nearly equal in length, antennomere 6 wider, antennomere 7 shorter and narrower. Thorax. Pronotum without distinct ���systematic punctures���, covered with sparse and faint punctation, interstices smooth and shining; black in centre and along posterior margin, yellow coloration of lateral margins widening posteriorly. Prosternum well developed, tectiform medially, bearing fine but distinct median carina. Scutellum black, equilateral. Mesoventrite reaching anterior mesothoracic margin at single point, rather flat with longitudinal carina between mesocoxae raised to small acute tooth slightly anterior to mesocoxae. Metaventrite with weakly raised middle portion slightly projecting anteriorly between mesocoxae and posteriorly between metacoxae, bearing hydrofuge pubescence except for posteromedian glabrous area on raised middle portion. Anepisternum 3 ca. 4.5�� as long as wide, subparallel. Elytra black, with yellowish lateral margins and apices, each elytron with wide parasutural furrow engraved from base to nearly anterior two thirds of elytral length and with ca. 20 longitudinal series of punctures: 10 primary rows consisting of well-discernible setiferous punctures, 10 alternate, scarcely visible rows, consisting of a small number of punctures; lateral margins neither serrate nor denticulate; epipleura oblique, pseudoepipleura nearly vertical and separated by distinct ridge; their anterior dilated portion ending anterior to metacoxae. Legs. Fore coxae almost contiguous, pubescent; fore trochanters pubescent; middle coxae separated by median carina of mesoventrite; tip of hind trochanters free, not abutted to hind femora. Femora with distinct tibial grooves distally on inner faces; basal third of ventral side of fore femora covered with hydrofuge pubescence; middle and hind femora nearly glabrous. Tibiae relatively short and stout, progressively wider towards apices, spiny, lacking swimming hairs; hind tibiae curved inwards. Middle and hind tarsi nearly without fine and sparse swimming hairs on dorsal face. Abdomen. Six distinct ventrites, ventrites 1���5 rather shiny and sparsely pubescent, ventrite 6 more rugose, pubescent and somewhat retractable; ventrite 1 not carinate; posterior margin of ventrite 5 subtruncate. Aedeagus. Total length 0.44 mm. Parameres shorter than median lobe, 1.3�� as long as phallobase, outer side nearly straight, inner side strongly diverging in terminal 0.4. Median lobe slender, slightly narrowing apicad, rounded at apex; margins of longitudinal excision bearing long hairs (Fig. 6). Differential diagnosis. Laccobius sipeki sp. nov. belongs to a group of Glyptolaccobius species with curved hind tibiae and elytra bearing longitudinal rows of punctures (together with L. celsus, L. silvester, L. shorti, L. munus, and L. sharmai). It differs from the other species by the wide parasutural furrow, not strictly corresponding to a single row of punctures, and by the lack of yellowish dots and stripes near the elytral base and suture; the shape of parameres of this species resembles L. munus, but the divergent interior part is longer. Etymology. The species is dedicated to Petr ����pek, who collected some of the Glyptolaccobius species described in this paper. Bionomics. The holotype was collected on a wet rock with algae, blue algae and fallen leaves at the side of a waterfall on a small river surrounded by dense tropical forest (Fig. 11) together with the following beetles: Laccobius pluvialis, L. eliogentilii, L. (Cyclolaccobius) sp., Oocyclus sp., Dactylosternum sp. (Hydrophilidae), Hydraena sp. (Hydraenidae), Ceradryops sp. (Dryopidae) and two species of the genus Hydroscapha (Myxophaga: Hydroscaphidae) (FIK��ČEK & ����PKOV�� 2009). Distribution. India, Meghalaya., Published as part of Gentili, Elio & Fika��ek, Martin, 2009, Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae), pp. 607-623 in Acta Entomologica Musei Nationalis Pragae 49 (2) on pages 618-622, DOI: 10.5281/zenodo.5321314, {"references":["KAMITE Y., OGATA T. & HIKIDA N. 2007: Two new species of the genus Laccobius (Coleoptera, Hydrophilidae) from Japan. Elytra (Tokyo) 35: 34 - 41.","FIKACEK M. & SIPKOVA H. 2009: New Asian Hydroscapha, with comments on male-female association of co-occuring [sic!] species (Coleoptera: Myxophaga: Hydroscaphidae). Zootaxa 2286: 31 - 48."]}

Laccobius (Glyptolaccobius) hanka sp. nov. (Figs. 2) Type locality. India, Arunachal Pradesh, 1 km N of Bhalukpong, 27��01���21���N 92��38���06���E, 240 m a.s.l. Type material. HOLOTYPE: J (NMPC): ���INDIA, Arunachal Pradesh (16a) / 1 km N of Bhalukpong / 7���8.V. 2008, 240 m / 27��01���21���N 92��38���06���E / Fik��ček, Podskalsk��, ����pek lgt. // seepage: wet rock with moos [= moss] / and Nostoc below steep slope / with tropical evergreen forest���. PARATYPES: INDIA: ARUNACHAL PRADESH: same data as holotype, 26 spec. (KSEM, MSNV, NMPC). Description. Total length 1.45���1.85 mm (holotype: 1.60 mm), total width 0.90���1.10 mm (holotype: 1.00 mm). Body widely oval, convex, maximum length / maximum width ratio 1.6; maximum width between anterior elytral third and elytral midlength. Head. Coloration shining black, with rare and faint punctures, surface smooth without microsculpture; ���systematic punctures��� indistinct; anterolateral branches of frontoclypeal suture conspicuous at 100��. Labrum black, trapezoid, without specula; anterior margin nearly straight, slightly emarginated medially; lateral margins oblique, posterior margin straight. Eyes oblong, oblique, closest to each other posteriorly, not protruding laterad, separated by distance equal to 2.5�� of one eye; postocular portion of tempora short. Mentum flat or slightly convex, smooth without punctures, emarginated anteriorly. Submentum and gula smooth. Maxillary palpi yellowish; palpomere 1 thin and short; palpomeres 2 and 3 nearly equal in length, both dilated at apex; palpomere 4 elongate, nearly twice as long as palpomere 3, symmetrical, inner and outer margin convex, apex truncate. Eight antennomeres; scape longer than antennomeres 2���4 combined; pedicel cone-shaped; intermediate antennomeres (antennomeres 3���4) very short; cupule asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, with densely arranged setae, antennomeres 6 and 8 nearly equal in length, antennomere 6 wider, antennomere 7 shorter and narrower. Thorax. Pronotum transverse, 0.38�� as long as wide, widest at posterior margin; blackish with yellowish, posteriorly widened lateral margins, yellow space reaching width of eye; surface smooth, without microsculpture. Prosternum black, with longitudinal keel. Scutellar shield equilateral, dark, impunctate. Mesoventrite with longitudinal keel, tuberculate anteriorly. Elytra combined ca 1.15�� as long as wide; brown to black with yellowish lateral margins and apices; each elytron with indistinct pale yellow dot at base; 10 longitudinal rows of larger setiferous punctures alternating with 10 rows (interstriae) of sparser and fainter punctures; all rows less conspicuous in darker specimens. Black parasutural space forming a flat and shining stripe extended over parasutural furrow. Epipleura and pseudoepipleura oblique, nearly vertical. Metaventrite setiferous with three postero-median, posteriorly converging sulci. Legs. Fore coxae and trochanters granulate, pubescent; fore femora setiferous on basal third, smooth distally, with tibial groove; fore tibiae smooth, each with stiff setae and two apical spurs. Middle femora with tibial grooves; ventral side of each middle tibia with three longitudinal rows of stiff setae and sulcus between two longitudinal ridges. Hind trochanters smooth and shining; hind femora smooth, with setiferous punctures and tibial grooves; hind tibiae curved, ventral side of each tibia with three longitudinal rows of stiff setae and longitudinal sulcus between two ridges. Abdomen. Ventrite 1 granulated, ventrites 2���4 smooth, ventrites 5���6 granulated with short setae. Aedeagus (Fig. 3): Total length 0.37���0.49 mm. Parameres nearly as long as phallobase; medial margin of parameres subparallel, apices slightly diverging in dorsal view. Median lobe wide, tongue-shaped, slightly constricted at midlength, barely rounded apically, nearly as long as parameres. Differential diagnosis. Laccobius hanka sp. nov. belongs to a group of Glyptolaccobius species with curved hind tibiae and pubescent elytra with longitudinal rows of punctures (together with L. silvester, L. shorti and L. sipeki sp. nov.). It differs from all three mentioned species by the pale spot on the elytral base and the wide median lobe. Etymology. The new species is named after Hanka, a diminutive of the first name of Hana ����pkov�� (her maiden name was H. Podskalsk��), who collected part of the type series of this species. To be treated as a noun in apposition. Bionomics. All specimens were collected at a seepage site at a roadside below a steep slope covered with moss and a mat of Nostoc (Fig. 13). The locality is situated at the margin of the Assam Valley where the first highlands of the Himalaya Range begin to rise. Distribution. India, lowland border regions of Arunachal Pradesh. So far known only from the type locality., Published as part of Gentili, Elio & Fika��ek, Martin, 2009, Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae), pp. 607-623 in Acta Entomologica Musei Nationalis Pragae 49 (2) on pages 612-614, DOI: 10.5281/zenodo.5321314

Laccobius (Glyptolaccobius) josefi sp. nov. (Fig. 3, 9) Type locality. India, Arunachal Pradesh, 1 km N of Bhalukpong, 27��01���21���N 92��38���06���E, 240 m a.s.l. Type material. HOLOTYPE: J (NMPC): ��� INDIA, Arunachal Pradesh (16a) / 1 km N of Bhalukpong / 7���8.V. 2008, 240 m / 27��01���21���N 92��38���06���E / Fik��ček, Podskalsk��, ����pek lgt.��� // seepage: wet rock with moos [= moss] / and Nostoc below steep slope / with tropical evergreen forest���. PARATYPES: 1 J 1 ♀ 2 spec. (NMPC, MSNV): same data as holotype. Description. Total length 2.20 mm, total width 1.15 mm. Body widely oval, convex, maximum width in anterior elytral third. Ratio total length / total width = 1.9. Head. Coloration blackish with two pale preocular spots, surface with rare but distinct punctures, without microsculpture; ���systematic punctures��� indistinct; periocular sulci distinct and punctate at 100��, reaching anterolateral branches of frontoclypeal suture; metopico-sagittal suture scarcely detectable. Labrum blackish, without specula; anterior margin nearly straight, slightly emarginated medially; posterior margin arched, strongly bent. Eyes oblong, oblique, closest to each other posteriorly, not protruding laterad, separated by distance equal to 2.35�� of width of one eye, posterior margin of eye, in dorsal view nearly straight, not reniform; postocular portion of tempora short. Mentum flat, microgranulated and punctured, emarginated anteriorly. Submentum and gula microgranulated. Maxillary palpi yellow-brown; palpomere 1 thin and short; palpomeres 2 and 3 nearly equal in length, palpomere 3 dilated at apex; palpomere 4 elongate, nearly twice as long as palpomere 3, asymmetrical, inner margin straight and outer margin convex, apex truncate. Eight antennomeres; scape (antennomere 1) longer than antennomeres 2���4 combined; pedicel (antennomere 2) cone-shaped; two intermediate antennomeres (antennomeres 3���4) very short; cupule (antennomere 5) asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, with dense setae, antennomeres 6 and 8 nearly equal in length, antennomere 6 wider, antennomere 7 shorter and narrower. Thorax. Pronotum transverse, 0.44�� as long as wide; black with yellowish lateral margins; yellow area as wide as posterior margin of eye along the base; surface smooth as that on head, without microsculpture except some sparse and faint punctures. Prosternum black, tectiform, with longitudinal keel. Scutellar shield equilateral, black, with some punctures. Mesoventrite granulate, with longitudinal, anteriorly tuberculate keel. Elytra combined slightly elongate, ca. 1.20�� as long as wide, blackish with yellowish lateral margins and apices; each elytron with pale yellow dot near middle of base; 10 longitudinal rows of regularly impressed punctures alternating with 10 rows (interstriae) of sparser and fainter punctures. Parasutural furrow scarcely conspicuous on posterior elytral third, disappearing before elytral apex. Epipleura and pseudoepipleura oblique, nearly vertical. Metaventrite setiferous with median longitudinal glabrous area and three longitudinal sulci converging posteriorly. Legs. Fore coxae and trochanters granulate, pubescent; fore femora setiferous on basal third, smooth distally, with tibial grooves; fore tibiae smooth, each bearing stiff setae and two apical spurs. Middle femora smooth, with tibial grooves; ventral side of middle tibiae with three longitudinal rows of stiff setae and sulcus between two longitudinal ridges. Hind trochanters smooth and shining; hind femora smooth, each with scarce punctures and tibial grooves; hind tibiae curved, ventral side of each tibia with three longitudinal rows of stiff setae and longitudinal sulcus between two ridges. Natatory setae nearly absent on dorsal surface of tarsi. Abdomen. All six ventrites smooth, without microsculpture. Aedeagus (Fig. 3). Total length 0.49 mm. Parameres nearly 1.5�� as long as phallobase. Median lobe slightly longer than parameres, its apex widened and separated into two lobes. Differential diagnosis. Laccobius josefi sp. nov. belongs to the group of species with curved hind tibiae, longitudinal rows of elytral punctures and black elytral surface with yellowish basal dots (together with L. guttalis sp. nov., L. munus and L. sharmai). It differs from the other three species by the clear preocular spots and by the shape of the aedeagus with the widely swollen apical portion of the median lobe; from L. guttalis sp. nov. it also differs by the lack of transverse lines on the parasutural stripe in anterior elytral third and from L. munus and L. sharmai by the less convex body. Etymology. We dedicate this species to Josef Jel��nek on the occasion of his 70 th birthday. Bionomics. See L. hanka sp. nov. Distribution. India, lowland border regions of Arunachal Pradesh., Published as part of Gentili, Elio & Fika��ek, Martin, 2009, Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae), pp. 607-623 in Acta Entomologica Musei Nationalis Pragae 49 (2) on pages 615-616, DOI: 10.5281/zenodo.5321314

Laccobius (Glyptolaccobius) pluvialis Gentili, 2006 (Fig. 5) Type locality. INDIA NE, Meghalaya State, SW of Cherrapunjee, 25��13������14���N 91��40���E, 900 m. Additional material examined. INDIA: MEGHALAYA: E Khasi Hills, 11 km SW of Cherrapunjee, Laitkynsew, 25��13���N, 91��39���E, 810 m, Fik��ček, Podskalsk��, ����pek leg., 21.���24.iv.2008, seepage: wet rock with algae, blue algae, moss, ca. 1.5���2 km via rd. from ���Cherrapunjee Holid. Resort��� in direct. Cherrapunjee, exposed���, 109 spec. (KSEM, NMPC, MSNV); E Khasi Hills, 11 km SW of Cherrapunjee, Laitkinsew, 25��12���N, 91��40���E, 460 m, Fik��ček, Podskalsk��, ����pek leg. 21���24.iv.2008, wet rock with algae, blue algae and fallen leaves at side of waterfall on small river surrounded by tropical forest, ca. 200 m upstream from living bridge [= bridge made of living roots of large Ficus trees], 1 J (NMPC); SW of Cherrapunjee, 25��13������14���N 91��40���E, 900 m, P. Pachol��tko leg., 5.���24.v.2005 (same data as holotype), 80 spec. (NHMW, MSNV). Note. The aedeagus was illustrated by GENTILI (2006); the illustrations in this paper are based on new slides. Bionomics. Laccobius pluvialis has been repeatedly found in large numbers on loamy seepages as well as wet rocks below these seepages and at sides of small temporary streams on the southern slopes of the Meghalaya Plateau. Most of the specimens collected in 2008 were found at sides of roads in exposed microhabitats sparsely covered with moss, algae and bluegreen algae (Fig. 12, see also FIK��ČEK & ����PKOV�� 2009, SHORT 2009). Only one specimen of L. pluvialis was found on partly shaded wet rocks at the side of a larger waterfall surrounded by a dense secondary rainforest (see Bionomics of L. sipeki sp. nov. and Fig. 11 for details)., Published as part of Gentili, Elio & Fika��ek, Martin, 2009, Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae), pp. 607-623 in Acta Entomologica Musei Nationalis Pragae 49 (2) on page 616, DOI: 10.5281/zenodo.5321314, {"references":["GENTILI E. 2006: Revisional notes on the genus Laccobius. I. Subgenus Glyptolaccobius (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 46: 57 - 76.","FIKACEK M. & SIPKOVA H. 2009: New Asian Hydroscapha, with comments on male-female association of co-occuring [sic!] species (Coleoptera: Myxophaga: Hydroscaphidae). Zootaxa 2286: 31 - 48.","SHORT A. E. Z. 2009: Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49: 625 - 630."]}

Laccobius (Glyptolaccobius) guttalis sp. nov. (Figs. 1, 7) Type locality. Central Nepal, Bagmati, Sindhupalchok, Sarmatang, 2500 m a.s.l.. Type material. HOLOTYPE: J (NKME): ‘C. NEPAL, Bagmati / Sindhupalchok / 2500 m, Sarmatang / 03–08.VI.1989 / leg. C. Holzschuh’. Description. Total length 2.15 mm, total width 1.25 mm. Maximum length / maximum width ratio 1.7. Body wide oval, convex, maximum width in anterior elytral third. Head. Coloration entirely black, with rare and inconspicuous punctures; shining, smooth without microsculpture; ‘systematic punctures’ indistinct; anterolateral branches of frontoclypeal suture conspicuous and punctate at 100×. Labrum black, trapezoid, without specula; anterior margin nearly straight, slightly emarginated medially; lateral margins oblique, posterior margin crescent-shaped. Eyes oblong, oblique, closest to each other posteriorly, not protruding laterad, interocular distance equal to 2.5× of width of one eye; postocular portion of tempora short. Mentum flat, smooth without punctures, emarginated anteriorly. Submentum and gula microgranulate. Maxillary palpi yellow-brown; palpomere 1 thin and short; palpomeres 2 and 3 nearly equal in length; palpomere 3 dilated at apex; palpomere 4 elongate, nearly twice as long as palpomere 3, asymmetrical, inner margin straight and outer margin convex, apex truncate. Eight antennomeres; scape longer than antennomeres 2–4 combined; pedicel cone-shaped; intermediate antennomeres (antennomeres 3–4) very short; cupule asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; antennal club loose, bearing densely arranged setae, antennomeres 6 and 8 nearly equal in length, antennomere 6 wider, antennomere 7 shorter and narrower. Thorax. Pronotum transverse, 0.35× as long as wide; black with yellowish lateral margins, yellow margin widened posteriorly and continuing as fine yellow strip along base; surface smooth, without microsculpture, with some sparse and faint punctures. Prosternum black, with longitudinal keel. Scutellar shield equilateral, black, bearing very small punctures. Mesoventrite with longitudinal keel simple, not tuberculate anteriorly. Elytra combined slightly elongate, ca 1.20× as long as wide; shining black with yellowish-brown lateral margins and apices, and pale yellow dots and stripes; 10 longitudinal rows of setiferous, more impressed punctures alternating with 10 rows (interstriae) of sparser and fainter punctures. Parasutural space covered by oblique lines reaching first punctural row in anterior fourth and then forming a flat and shining stripe separating suture from parasutural furrow, stripe and furrow disappearing just before elytral apex. Epipleura and pseudoepipleura oblique, nearly vertical. Metaventrite setiferous, with median longitudinal glabrous area posteriorly. Legs. Fore coxae and trochanters granulate, pubescent; fore femora setiferous on basal third, smooth distally, with tibial groove; fore tibiae smooth, with stiff setae and two apical spurs. Middle femora with tibial grooves; ventral face of middle tibiae with three longitudinal rows of stiff setae and sulcus between two longitudinal ridges. Hind trochanters smooth and shining; hind femora smooth, with scarce punctures and tibial grooves; hind tibiae curved, ventral face with three longitudinal rows of stiff setae and longitudinal sulcus between two ridges. Abdomen. Ventrites 1–4 smooth, with lateral setae; ventrites 5–6 wrinkled. Aedeagus (Fig. 1). Total length 0.68 mm. Parameres slightly longer than phallobase, slightly overlapping apex of median lobe. Median lobe narrow, slightly widened in apical 0.4, with longitudinal excision ventrally; apical portion broadly rounded. Differential diagnosis. Laccobius guttalis sp. nov. belongs to a group of species with curved hind tibiae, longitudinal rows of elytral punctures and black elytral surface with yellowish dots and stripes (together with L. josefi sp. nov., L. munus and L. sharmai). It differs from the other three species by the presence of transverse lines in the anterior part of the parasutural stripe and the shape of the aedeagus, with the apical portion of the median lobe swollen and the median lobe bearing a longitudinal ventral excavation (Fig. 1). Etymology. From a drop (Latin: gutta), alluding to the preferred seepage habitat of Glyptolaccobius. Bionomics. Unknown. Distribution. Known only from the type locality in Nepal.

Laccobius (Glyptolaccobius) sharmai Gentili, 1995 Type locality. Nepal E, Solukhumbu, Surka La Pass, Karka Khola, 2875 m a.s.l. Additional material examined. NEPAL: Manasiu Mts., E slope of Ngadi Khola Valley, 2000���2300 m, 28��22���N 84��29���E, 14���16.v.2005, J. Schmidt leg., 17 spec. (NHME; MSNP; MSNV); Manasiu Mts., SE slope W Gupchi Danda, 2200���2300 m, 28��08���37N 84��44���42E, 18.v.2006, J. Schmidt leg., 1 J 1 ♀ (CSHS); Ganesh Himal, NNW Trisuli Bazar, Singhen Khola, 2400���2500 m, 19.iv.1999, Ghal�� & Gurung leg., 1 J 1 (NHME); Baglung Lekh, ca. 30 km W Baglung, N Tara Khola, 2500���2700 m, 28��22���N 83��20���E, 18.v.2004, J. Schmidt leg., 3 spec. (NHME, MSNV); Baglung Lekh, ca. 30 km W Baglung, N Tara Khola, 2700���2800 m, 28��22���N 83��20���E, 19.���21.v.2004, J. Schmidt leg., 2 spec. (NHME); Annapurna Mts., Banthanti S Gorapani, 2400 m, 26.v.2004, J. Schmidt leg., 1 spec. (NHME). Note. This species is known from the foothills of high mountains in Nepal: Manasiu, Annapurna and Everest (Sagarmatha). It is recorded here also from the Dhaulagiri mountains and Langtang National Park (Ganesh Himal)., Published as part of Gentili, Elio & Fika��ek, Martin, 2009, Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae), pp. 607-623 in Acta Entomologica Musei Nationalis Pragae 49 (2) on page 618, DOI: 10.5281/zenodo.5321314, {"references":["GENTILI E. 1995: Hydrophilidae: 3. The genus Laccobius Erichson in China and neighbouring areas (Coleoptera). Pp. 411 - 429. In: JACH M. A. & JI L. (eds.): Water Beetles of China, Vol. 1. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, vi + 572 pp."]}

Laccobius (Glyptolaccobius) moriyai Kamite, Ogata & Hikida, 2007 (Figs. 4, 8) Type locality. Japan, Honshu Island, Yamagata Prefecture, Asahi-Mura, Arasawa. Type material (not examined). HOLOTYPE: J (ELEU): ���Arasawa, Asahi-Mura, Yamagata Pref., H. Moriya leg. 23-IX-2004 ���. PARATYPES: 4 spec., same data as holotype. Length 2.30���2.50 mm; width 1.40���1.50 mm. Additional material examined: JAPAN: HONSHU ISLAND: Yamagata Pref., Arasawa-dam, Tsuruoka-shi, T. Ikeda leg. 16.viii.2008, 1 J 1 ♀ (MSNV, NMPC). Differential diagnosis. Laccobius moriyai belongs to a group of species with curved hind tibiae, shining black elytra with longitudinal rows of punctures of which only the first one is sulciform. The combination of uniformly black dorsal colouration and apically strongly widened apex of the median lobe is not found in any other species of Glyptolaccobius. Distribution. Laccobius moriyai is the first representative of the subgenus Glyptolaccobius occurring in the Palaearctic Region. So far the species is known only from the environs of the Arasawa Dam in the Yamagata Prefecture in Japan (Honshu island)., Published as part of Gentili, Elio & Fika��ek, Martin, 2009, Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae), pp. 607-623 in Acta Entomologica Musei Nationalis Pragae 49 (2) on page 616, DOI: 10.5281/zenodo.5321314, {"references":["KAMITE Y., OGATA T. & HIKIDA N. 2007: Two new species of the genus Laccobius (Coleoptera, Hydrophilidae) from Japan. Elytra (Tokyo) 35: 34 - 41."]}

Laccobius (Glyptolaccobius) affinis Knisch, 1927 (Figs. 5-7) Laccobius (s. str.) affinis Knisch, 1927: 132. Laccobius (s. str.) affinis: D’ ORCHYMONT (1928): 140. Laccobius (Platylaccobius) affinis: GENTILI (1979): 31. Laccobius (Glyptolaccobius) affinis: GENTILI (1989): 36; GENTILI (1995): 253; HANSEN (1999): 130; HEBAUER (2002): 12. Type locality. India, Uttar Pradesh, Kumaon, Gori Valley, 2300 m a.s.l. Type material examined. LECTOTYPE: J, 2.0 × 1.1 mm (NHML): ‘Gori Valley, Kumaon, India, 7000 ft H.G.C. / Knisch det. 1925 Laccobius s.str. affinis Kn. [handwritten] / Coll. A.Knisch COTYPUS’. PARALECTOTYPES: 6 spec. (3 NHML, 3 ISNB). The three paralectotypes from NHML are mounted on separate cards, with labels as for the lectotype. Two of the three paralectotypes from ISNB (male and female) are mounted on one card, with labels: ‘Coll. R.I.Sc.N.B., Inde, Gori Valley, Kumaon, India, 7000 ft H.G.C. / Knisch det. 1925 Laccobius s.str. affinis Kn. / Coll. A.Knisch COTYPUS’. The last specimen is a male of L. egregius (see under that species). Following the text of KNISCH (1927): ‘Typus im British Museum, London’ I designated one of the specimens in NHML as a lectotype (GENTILI 1978). The three specimens from ISNB come from the Knisch collection: ‘Sieben Exemplare liegen nur durch Güte des Herrn G. C. Champion vor, von welchen drei freundl. meiner Sammlung überlassen wurden’ [‘There are seven specimens available due to the kindness of Mr. G. C. Champion, three of which were kindly donated for my collection.’] (KNISCH 1927). Additional material examined. INDIA: UTTAR PRADESH, Gori Valley, Kumaon 7000′, H. G. Champion leg., 1 J 1 ♀ (MSNV). NEPAL: Annapurna Mts., Baglung Lekh, ca 18 km W Baglung, upp. Okhle village, 2300 m a.s.l., 13.v.2004, J. Schmidt leg., 1 J 2 ♀♀ (NHMW); Sankua Sabha Distr., above Pahakhola, 2600-2800 m a.s.l., Quercus semecarpifolia, Rhododendron, 31.v-3.vi.1988, Martens & Schawaller leg., 1 J (CFHG); Solukhumbu Distr., above Nunthala, 2300-2500 m a.s.l., 13.v.1997, W. Schawaller leg., 1 J (CFHG). Material not examined. NEPAL: ‘Prov. GANDAKI, Distr. Kaski, Ghorepani nr. Tatapani, 2800- 2500 m, leg. Wolf 23.4.2000 (NMEG)’ (HEBAUER 2002). Differential diagnosis. Laccobius affinis belongs to species with unordered elytral punctures and curved hind tibiae. It resembles L. egregius and L. incisus by the dark elytral colour without clear spots and stripes. From these species it is chiefly distinguished by its aedeagus (Figs. 5-7) with simple parameres, i.e. not excised or hooked at apices. Externally, L. affinis differs from the latter two species by the lack of shagreen and black hairs on the head and pronotum, arrowhead-shaped anterior margin of the mesosternal keel, and the parasutural furrow reaching elytral apex. Distribution. India (Uttar Pradesh), Nepal., Published as part of Gentili, Elio, 2006, Revisional notes on the genus Laccobius. I. Subgenus Glyptolaccobius (Coleoptera: Hydrophilidae), pp. 57-76 in Acta Entomologica Musei Nationalis Pragae 46 on page 61, DOI: 10.5281/zenodo.5176694, {"references":["KNISCH A. 1927: Neue Hydrophiliden der Orientalfauna. Spolia Zeylanica 14: 129 - 133.","D' ORCHYMONT A. 1928: Catalogue of Indian Insects. Part 14 - Palpicornia. Government of India Central Publication Branch, Calcutta, 2 + 146 pp.","GENTILI E. 1979: I Laccobius della Regione Orientale (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 1 (1978): 27 - 50.","GENTILI E. 1989: Alcune novita sul genere Laccobius (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 10 (1987): 31 - 39.","GENTILI E. 1995: Hydrophilidae: 3. The genus Laccobius Erichson in China and neighbouring areas (Coleoptera). Pp. 245 - 286. In: JACH M. A. & JI L. (eds.): Water Beetles of China. Vol. I. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, 410 pp.","HANSEN M. 1999: Hydrophiloidea (Coleoptera). In HANSEN M. (ed.): World Catalogue of Insects, Vol. 2. Apollo Books, Stenstrup, 416 pp.","HEBAUER F. 2002: Hydrophilidae of Northern India and Southern Himalaya (Coleoptera: Hydrophilidae). Acta Coleopterologica 18: 3 - 72."]}

Laccobius (Glyptolaccobius) pluvialis sp. nov. (Figs. 1-3, 27-29) Type locality. India NE, Meghalaya, SW of Cherrapunjee, 900 m a.s.l. Type material. HOLOTYPE: J, 1. 9 × 1.1 mm (NHMW): ‘NE INDIA, MEGHALAYA, / SW of CHERRAPUNJEE, / 25°13-14′ N, 91°40′E, / 5.-24.V. 2005, 900 m, / P. Pacholatko leg. ’. PARATYPES: 180 spec. (NHMW, MSNV, CFHG), same data as holotype. Description. Total length 1. 7-2.2 mm, total width 1. 0-1. 2 mm. Body (Fig. 1) oval, convex, its outline not interrupted between pronotum and elytra; elytra not explanate. Head. Labrum of males without specula; trapezoidal in dorsal view, with straight anterior and posterior margins and faintly oblique sides. Clypeus strongly convex, not deflexed towards margins, black and shining, without distinct ‘systematic punctures’. Eyes oblique, slightly reniform in lateral view, straight anteriorly and emarginate posteriorly, weakly convex, not protruding, separated by a little more than 2.5 times the width of one eye. Maxillary palpi less than 0.5x as long as head wide; palpomere 2 not swollen, as wide as palpomere 3; apical segment nearly 0.3 times as long as whole palpus, asymmetrical, outer margin nearly straight, inner margin rounded, with truncate apex. Mentum ca 1. 5 times as wide as long, surface punctate, almost flat. Eight antennomeres (Fig. 3); scape (antennomere 1) longer than antennomeres 2-4 together; pedicel (antennomere 2) cone shaped; two intermediate antennomeres (3-4) very short; cupule (antennomere 5) slightly asymmetrical, crescent-shaped in ventral view, nearly flat in dorsal view; segment 3 of club nearly twice as long as segment 1, the latter more stout but nearly as long as segment 2 of club. Thorax. Pronotum smooth and shining, without distinct ‘systematic punctures’, only with sparse and faint punctures; disc black, yellow lateral margins wider posteriorly. Prosternum well developed, disc slightly bulging; median carina fine and in part obsolete. Mesoventrite only reaching anterior margin of mesothorax in a single point, rather flat except for median carina, the latter more raised anteriorly and forming a small acute tooth anterior of mesocoxae. Metaventrite with middle portion rather weakly raised and slightly projecting anteriorly between mesocoxae, covered with hydrofuge pubescence except for posteromedian glabrous area on raised middle portion. Anepisternum 3 ca 3.5 times as long as wide, subparallel. Elytra with a hardly visible parasutural stria on posterior third, without trace of serial punctures; dark on disc, near suture and along anterior margin, chestnut-yellow near posterior and lateral margins, with yellow and darker colours mixed in contact zone; lateral margins not serrate nor denticulate; epipleura oblique, pseudepipleura nearly vertical, separated by distinct ridge consisting of small arcs; their anterior widened portion ending before metacoxae. Hind wing (Fig. 2) with r-m crossvein rising from distal half of the pigmented area at anterior wing margin; wedge cell only slightly more than half as long as basal cell; jugal lobe distinctly demarcated from remainder of wing by sharp and rather deep excision at posterior wing margin. Legs. Fore coxae pubescent, almost contiguous; middle coxae separated by median carina of mesoventrite; trochanters pubescent, especially middle ones; tips of hind trochanters free, not abutted to hind femora. Femora with distinct tibial grooves distally on inner face; base of fore femora pubescent, middle and posterior femora nearly glabrous, middle femora with a subbasal patch of dense stiff setae in both sexes. Tibiae relatively short and stout, progressively wider towards apices, spiny, with three longitudinal series of short spines and two large apical thorns; without swimming hairs; hind tibiae curved inward. Fore tarsomeres 2 and 3 expanded in males; middle and hind tarsi with fine and sparse swimming hairs on dorsal face; hind tarsomere 1 much shorter than tarsomere 2. Claws of moderate size, robust, weakly curved. Abdomen. Six visible ventrites, ventrites 1-5 rather shiny and sparsely pubescent, ventrite 6 more dull, densely pubescent and somewhat retractable; ventrite 1 not carinate; posterior margin of ventrite 5 subtruncate. Male genitalia.Aedeagus as in Figs. 27-29; phallobase nearly 0.5 times as long as parameres, median lobe with swelling before apex. Differential diagnosis. Laccobius pluvialis sp. nov. is extremely close to L. jaechi and L. eliogentilii in body shape and colour, including the pubescent surface with irregular punctation and the parasutural furrow recognizable barely in the apical fifth. The elytra lack basal spots and parasutural stripes but are widely chestnut-yellowish near the lateral margins and apex. The phallobase is distinctly shorter than in L. jaechi; finally, the subapical swelling of the median lobe differentiates this species from L. eliogentilii. Etymology. This species is named in reference to the extreme rainfall records reported from the area close to the type locality. Distribution. India (Meghalaya). So far known only from the type locality., Published as part of Gentili, Elio, 2006, Revisional notes on the genus Laccobius. I. Subgenus Glyptolaccobius (Coleoptera: Hydrophilidae), pp. 57-76 in Acta Entomologica Musei Nationalis Pragae 46 on pages 68-70, DOI: 10.5281/zenodo.5176694, {"references":["GENTILI E. 1995: Hydrophilidae: 3. The genus Laccobius Erichson in China and neighbouring areas (Coleoptera). Pp. 245 - 286. In: JACH M. A. & JI L. (eds.): Water Beetles of China. Vol. I. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, 410 pp."]}

Gentili, Elio (2006): Revisional notes on the genus Laccobius. I. Subgenus Glyptolaccobius (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 46: 57-76, DOI: http://doi.org/10.5281/zenodo.5176694, {"references":["BERGE HENEGOUWEN A. L. VAN 1982: De nederlandse soorten van het genus Laccobius Erichson (Coleoptera, Hydrophilidae), een systematische en faunistische studie. [On the Dutch species of the genus Laccobius Erichson (Coleoptera, Hydrophilidae), a systematic and faunistic study]. Zoologische Bijdragen 28: 59-84 (in Dutch, English abstract without title).","BLACKBURN T. 1895: Further notes on Australian Coleoptera, with descriptions of new genera and species. XVII. Transactions of the Royal Society of South Australia 19 (1894-95): 27-60.","CHEARY B. S. 1971: The Biology, Ecology and Systematics of the genus Laccobius (Laccobius) (Coleoptera: Hydrophilidae) of the New World. University of California, Riverside, 178 pp.","GANGLBAUER L. 1904: Die Kafer von Mitteleuropa. Vol. 4, part 1. Karl Gerold's Sohn, Wien, 286 pp.","GENTILI E. 1974: Descrizione di nuove entita appartenenti al genere Laccobius Erichson, 1837 e proposta per un nuovo inquadramento sottogenerico (Coleoptera Palpicornia). Memorie del Museo Civico di Storia Naturale Verona 20 (1972): 549-565.","GENTILI E. 1979: I Laccobius della Regione Orientale (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 1 (1978): 27-50.","GENTILI E. 1981: Il genere Laccobius nella Regione Etiopica (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 3 (1980): 31-42.","GENTILI E.1985:I Laccobius americani.I. I Laccobius del Canada (Coleoptera,Hydrophilidae).Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 6 (1983): 31-45.","GENTILI E. 1986a: I Laccobius americani. II. Il genere Laccobius a Sud del Canada (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 7 (1984): 31-40.","GENTILI E. 1986b: I Laccobius americani. III. Il genere Laccobius a Sud del Canada (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 8 (1985): 31-52.","GENTILI E. 1988: Verso una revisione del genere Laccobius (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 9 (1986): 31-47.","GENTILI E. 1989: Alcune novita sul genere Laccobius (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 10 (1987): 31-39.","GENTILI E. 1991: Elementi per una revisione del genere Laccobius (Coleoptera, Hydrophilidae). Giornale Italiano di Entomologia 5: 381-389.","GENTILI E. 1995: Hydrophilidae: 3. The genus Laccobius Erichson in China and neighbouring areas (Coleoptera). Pp. 245-286. In: JACH M. A. & JI L. (eds.): Water Beetles of China. Vol. I. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, 410 pp.","GENTILI E. 2003: Hydrophilidae: III. Additional notes on the genus Laccobius Erichson in China and neighbouring areas (Coleoptera). Pp. 411-429. In: JACH M. A. & JI L. (eds.): Water Beetles of China, Vol. 3. Zoologisch- Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, vi+572 pp.","GENTILI E. 2005: The genus Laccobius Erichson, 1837 in the Australian Region (Coleoptera, Hydrophilidae). Pp. 317-370. In: DACCORDI M. & GIACHINO P. M. (eds.): Results of the Zoological Missions to Australia of the Regional Museum of Natural Sciences of Turin, Italy. Monografie del Museo Regionale di Scienze Naturali Torino 42: 1-643.","GENTILI E. & CHIESA A. 1975: Revisione dei Laccobius paleartici. Memorie della Societa Entomologica Italiana 54: 1-187.","HANSEN M. 1991: The Hydrophiloid Beetles. Phylogeny, classification and a revision of the genera (Coleoptera, Hydrophiloidea). Biologiske Skrifter. Vol. 40. The Royal Danish Academy of Sciences and Letters, Munksgaard - Copenhagen, 368 pp.","HANSEN M. 1999: Hydrophiloidea (Coleoptera). In HANSEN M. (ed.): World Catalogue of Insects, Vol. 2. Apollo Books, Stenstrup, 416 pp.","HEBAUER F. 2002: Hydrophilidae of Northern India and Southern Himalaya (Coleoptera: Hydrophilidae). Acta Coleopterologica 18: 3-72.","KNISCH A. 1924: Neue Palpicornier aus dem sudlichen Himalaya. Wiener Entomologische Zeitung 41: 29-41.","KNISCH A. 1927: Neue Hydrophiliden der Orientalfauna. Spolia Zeylanica 14: 129-133.","D' ORCHYMONT A.1925:Contributions a l'etude des Hydrophilides I. Bulletin et Annales de la Societe Entomologique de Belgique 65: 63-77.","D' ORCHYMONT A.1928:Catalogue of Indian Insects.Part 14 - Palpicornia. Government of India Central Publication Branch, Calcutta, 2 + 146 pp.","SHATROVSKIY A.G. 1984.Obzor vodolyubov roda Laccobius Er.(Coleoptera,Hydrophilidae) fauny SSSR.(Revision of the genus Laccobius Er. of the Soviet Union (Coleoptera, Hydrophilidae)). Entomologicheskoe Obozrenie 63: 301-325 (in Russian, English abstract).","WOLLASTON T. V. 1867: Coleoptera Hesperidum, being an enumeration of the Coleopterous insects of the Cape Verde Archipelago. J. von Voorst, London, 39 + 285 pp.","ZAITZEV F.A. 1938: Vidy r. Laccobius Er. v faune SSSR i sopredel'nykh stran (Coleoptera, Hydrophilidae).[Species of Laccobius in the fauna of the USSR and adjacent lands (Coleoptera, Hydrophilidae)]. Trudy Zoologicheskogo Sektora. Akademiya Nauk SSSR, Gruzinskoe Okdelenie, Zakavkaskii Filial (Tiflis [= Tbilisi]) 2: 109-124 (in Russian)."]}

Laccobius (Glyptolaccobius) silvester sp. nov. (Figs. 34-35) Type locality. India, Uttar Anchal, W Loharket Village, 30 km N Bageshwar, 1800-1900 m a.s.l. Type material. HOLOTYPE: 1 J, 2.2 × 1. 2 mm (NHMW): ‘ INDIA: Uttaranchal / 30 km N Bageshwar / W Loharket [village] // 1800-1900 m. / 24.VI.2003; leg. Z. / Kejval & M. Tryzna’. Description. Small-sized species (holotype 2.2 mm long and 1. 2 mm wide). Body oval, moderately convex, its outline not interrupted between pronotum and elytra; elytra not explanate. Head. Labrum of males without specula; trapezoidal in dorsal view, with straight anterior and posterior margins and feebly oblique sides, shiny black. Clypeus moderately convex, not deflexed towards margins, shiny black, with sparse and faint punctures, without distinct ‘systematic punctures’. Eyes oblique, weakly convex, not protruding, separated by slightly more than two times width of one eye, nearly reniform in lateral view, emarginate posteriorly. Maxillary palpi less than 0.5 times as long as head wide; palpomere 2 not swollen, narrower and nearly as long as palpomere 3; palpomere 4 1. 5 times as long as palpomere 3, outer margin nearly straight, inner margin rounded, with truncate apex. Mentum ca 0.5 times as wide as long, almost flat, at least in centre, covered with faint punctures. Eight antennomeres; scape (antennomere 1) longer than three following antennomeres together; pedicel (antennomere 2) cone-shaped; two intermediate antennomeres (3-4) very short; cupule (antennomere 5) asymmetrical, oval in ventral view, crescent-shaped and emarginate in dorsal view; club with segments 1 and 3 nearly equal in length, segment 1 more stout, segment 2 shorter and narrower. Thorax. Pronotum without distinct ‘systematic punctures’ but covered with sparse and faint punctation; smooth and shining; black in centre and along anterior and posterior margins, yellow coloration of lateral margins widening posteriorly. Prosternum well developed, tectiform medially and with fine, clearly traced median carina. Scutellum black, equilateral. Mesoventrite reaching anterior mesothoracic margin at single point, rather flat except for median carina, the latter more raised anteriorly and forming small acute tooth slightly anterior of middle coxae. Metaventrite with rather weakly raised middle portion, the latter slightly projecting anteriorly between middle coxae, with hydrofuge pubescence except for posteromedian glabrous area on raised middle portion. Anepisternum 3 ca 4.5 times as long as wide, subparallel. Elytra with complete parasutural furrow and ca 20 longitudinal series of punctures: ten primary rows of well discernible setigerous punctures, and ten alternate, scarcely visible rows; lateral margins neither serrate nor denticulate; epipleura oblique, pseudepipleura nearly vertical and separated by distinct ridge consisting of small arcs; their anterior dilated portion ending before hind coxae. Legs. Fore coxae almost contiguous, fore trochanters pubescent; middle coxae separated by median carina of mesoventrite; tip of hind trochanters free, not abutted to hind femora. Femora with distinct tibial grooves distally on inner faces; basal third of ventral side of fore femora covered with hydrofuge pubescence; middle and hind femora nearly glabrous. Tibiae relatively short and stout, progressively wider towards apices, spiny (ventral face with three longitudinal rows of stiff setae, apices with two long spurs), without swimming hairs; hind tibiae curved inwards. Fore tarsomeres 2 and 3 expanded in males. Middle and hind tarsi with fine and sparse swimming hairs on dorsal face; hind tarsomere 1 much shorter than tarsomere 2. Claws of moderate size, robust, weakly curved. Abdomen. Six distinct ventrites, ventrites 1-5 rather shiny and sparsely pubescent, ventrite 6 more dull, densely pubescent and somewhat retractable; ventrite 1 not carinate; posterior margin of ventrite 5 subtruncate. Male genitalia. Median lobe slender, straight, excised at apex; parameres nearly as long as median lobe and slightly longer than phallobase (Figs. 34-35). Differential diagnosis. Laccobius silvester sp. nov. pertains to the group of Glyptolaccobius species with curved hind tibiae and elytra with longitudinal rows of punctures. Within the group it differs from L. celsus and L. munus by the lack of sulciform punctural rows, from L. sharmai by the lack of yellowish dots and stripes near the elytral base and suture, and from L. shorti sp. nov. by the parameres nearly as long as phallobase. Etymology. The mountain zones of Uttar Anchal are rich of luxuriant forests. Distribution. India (Uttar Anchal)., Published as part of Gentili, Elio, 2006, Revisional notes on the genus Laccobius. I. Subgenus Glyptolaccobius (Coleoptera: Hydrophilidae), pp. 57-76 in Acta Entomologica Musei Nationalis Pragae 46 on pages 72-74, DOI: 10.5281/zenodo.5176694, {"references":["GENTILI E. 1979: I Laccobius della Regione Orientale (Coleoptera, Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (n. s.) 1 (1978): 27 - 50.","GENTILI E. 1995: Hydrophilidae: 3. The genus Laccobius Erichson in China and neighbouring areas (Coleoptera). Pp. 245 - 286. In: JACH M. A. & JI L. (eds.): Water Beetles of China. Vol. I. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, 410 pp."]}

Gentili, Elio, Fikaček, Martin (2009): Taxonomic notes on Laccobius, subgenus Glyptolaccobius, with new records and description of four new species (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49 (2): 607-623, DOI: http://doi.org/10.5281/zenodo.5321314, {"references":["ERICHSON W. F. 1837: Die Kafer Mark Brandenburg. Vol. 1. F. H. Morin, Berlin, viii + 740 pp.","FIKACEK M. & SIPKOVA H. 2009: New Asian Hydroscapha, with comments on male-female association of co-occuring [sic!] species (Coleoptera: Myxophaga: Hydroscaphidae). Zootaxa 2286: 31-48.","GENTILI E. 1979:I Laccobius della regione orientale (Coleoptera, Hydrophilidae).Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (N. S.) 1 (1978): 27-50.","GENTILI E. 1988: Verso una revisione del genere Laccobius (Coleoptera: Hydrophilidae). Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (N. S.) 9 (1986): 31-47.","GENTILI E.1989:Alcune novita sul genere Laccobius (Coleoptera:Hydrophilidae).Annuario Osservatorio di Fisica Terrestre e Museo Antonio Stoppani del Seminario Arcivescovile di Milano (N. S.) 10 (1987): 31-39.","GENTILI E. 1995: Hydrophilidae: 3. The genus Laccobius Erichson in China and neighbouring areas (Coleoptera). Pp. 411-429. In: JACH M.A. & JI L. (eds.): Water Beetles of China, Vol. 1. Zoologisch-Botanische Gesellschaft in Osterreich and Wiener Coleopterologenverein, Wien, vi + 572 pp.","GENTILI E. 2006: Revisional notes on the genus Laccobius. I. Subgenus Glyptolaccobius (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 46: 57-76.","HEBAUER F. 2002: Hydrophilidae of Northern India and Southern Himalaya (Coleoptera: Hydrophilidae). Acta Coleopterologica 18: 3-72.","KAMITE Y., OGATA T. & HIKIDA N. 2007:Two new species of the genus Laccobius (Coleoptera, Hydrophilidae) from Japan. Elytra (Tokyo) 35: 34-41.","KNISCH A. 1927: Neue Hydrophiliden der Orientalfauna. Spolia Zeylanica 14: 129-133.","SHORT A. E. Z. 2009: Two new species of the hygropetric genus Oocyclus from eastern India (Coleoptera: Hydrophilidae). Acta Entomologica Musei Nationalis Pragae 49: 625-630."]}