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5. Gordonoryssomus everardoi Almeida & Santos, sp. nov. Type Material: Holotype, male (DZUP). Type locality: “Nova Friburgo, Brazil ”. Geographical distribution: Brazil (Fig. 17).

Gordonoryssomus everardoi sp. nov. (Fig. 12) Description of holotype (male). Length 3.44, width 2.26 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 12 C). General color yellowish red, middle of thorax and abdomen dark brown. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal triangular black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 12 D). Elytra yellowish red with a large black spot, extending from the base almost to the apex, reaching the middle of humeral callus (Fig. 12 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs yellowish brown. Metaventrite finely, sparsely punctured medially and anteriorly, coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally (Fig. 12 B). First abdominal ventrite with postcoxal line extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 12 E). Genitalia simple; basal lobe as longer than the parameres, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere straight in lateral view, nearly parallel-sided (Fig. 12 G, H); sipho flattened and rounded apex (Fig. 12 F). Female. Unknown. Etymology. This species is named in honor of Dr. Everardo José Grossi, a doctor with a passion for beetles. Discussion. The color pattern of this species is very similar with G. deyrollii, but it differs in body length, and also male genitalia, with basal lobe almost the same length as the parameres, and J-shaped sipho. It also differs from all other species of the genus by the color pattern and male genitalia. Type material. “ BRAZIL, R.J., Nova/Friburgo, Sans Souci/ 21 -I- 2007 1050 m /P. Grossi col.” “♂” [white label]. HOLOTYPE / Gordonoryssomus / everardoi /Almeida & Santos, 2014 ” [red label]. 1 ex. “ DZUP / 131938 ” [DZUP]. Geographical distribution. Nova Friburgo, Sans Souci, Rio de Janeiro, Brazil (Fig. 17).

Gordonoryssomus limeirai sp. nov. (Fig. 13) Description of holotype (male). Length 3.71, width 2.78 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 13 C). General color and middle of thorax and abdomen brownish yellow. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal triangular black spot, translucent at apex; finely, densely punctured; pubescence of semierect yellowish white hairs (Fig. 13 D). Elytra yellowish red with a large black spot, extending from the base almost to the apex, reaching the middle of humeral callus. With a brownish yellow spot in sutural margin along almost 2 / 3 of the suture (Fig. 13 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent (Fig. 13 B). Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally. First abdominal ventrite with postcoxal line almost touching the posterior margin of the first abdominal ventrite; precoxal area coarsely punctured on base of cavity (Fig. 13 E). Genitalia simple; basal lobe equal in length to the paramere, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere straight in lateral view, nearly parallel-sided (Fig. 13 G, H); sipho flattened, sinuous at apex (Fig. 13 F). FIGURE 13. Gordonoryssomus limeirai sp. nov. (A���E) Habitus: (A) dorsal; (B) ventral; (C) lateral; (D) frontal view. (E) Abdomen; (F���H) male genitalia: (F) sipho; (G) Tegmen, ventral; (H) lateral view. (I���J) Female terminalia: (I) spermatheca; (J) coxites. Female. Similar to male in all aspects externally. Genitalia with genital plates triangular, not elongated, styli with setae; spermatheca elongate, not uniform, with narrow apex (Fig. 13 I, J). Etymology. This species is named in honor of the dedicated entomologist, Dr. Francisco Limeira de Oliveira. Discussion. The color pattern is similar to that of G. delicatus, but it differs in length and the size of the sutural spot. Female genitalia with genital plate not elongate, somewhat triangular as in G. deyrollii, G. delicatus and G. mirnae. G. limeirai differs from all other species of the genus by the postcoxal line which almost touches the posterior margin of the first abdominal ventrite and male genitalia with wider basal lobe. Type material. ��� BRAZIL (PI), Caracol/Parque Nacional Serra das/Confus��es, Andorinha, 515 m,/ 09��08��� 27.8 ���S / 43 �� 33 ��� 42.1 ���W / CZMA ��� ���Suspensa dupla (20 metros)/ 20-30.IX. 2013, J.A. Rafael,/F. Limeira de Oliveira &/T.T.A. Silva cols./ CZMA ��� ���♂��� [white label]. ��� HOLOTYPE / Gordonoryssomus / limeirai /Almeida & Santos, 2014 ��� [red label]. 1 ex. [CZMA]; ��� BRAZIL (PI), Caracol/Parque Nacional Serra das/Confus��es, Andorinha, 515 m,/ 09��08��� 27.8 ���S / 43 �� 33 ��� 42.1 ���W / CZMA ��� ���Suspensa dupla (20 metros)/ 20-30.IX. 2013, J.A. Rafael, / F. Limeira de Oliveira & /T.T.A. Silva cols./ CZMA ��� ���♀��� [White label].��� PARATYPE / Gordonoryssomus / limeirai / Almeida & Santos, 2014 ��� [yellow label]. 1 ex. ��� DZUP / 245996 ��� [DZUP]. Geographical distribution. Brazil (PI) (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

4. Gordonoryssomus delicatus Almeida & Lima, 1995 Gordonoryssomus delicatus Almeida & Lima, 1995: 715���717 (original description, morphological details and male genitalia figured). Type material: Holotype, male (DZUP); paratypes (4 total), 1 Rio de Janeiro and 1 Minas Gerais (USNM), 1 (BMNH), 1 Santa Catarina (DZUP). Type locality: ���Santa Catarina, Brazil ���. Geographical distribution: Brazil (RJ, MG, SC, RS) (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024"]}

Gordonoryssomus mirnae sp. nov. (Fig. 14) Description of holotype (male). Length 5.64, width 3.82 mm. Form elongate oval, widest at middle of elytra, not flattened dorsoventrally (Fig. 14 C). General color reddish yellow (Fig. 14 A, B). Outline of pronotum and elytron discontinuous. Pronotum and elytra without spots. Pronotum translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 14 D). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs reddish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites finely punctured, punctures separated by about a diameter or more, punctures becoming thicker laterally. First abdominal ventrite with postcoxal line slightly angulate, extending beyond middle of first ventrite (Fig. 14 E). Genitalia simple; basal lobe longer than paramere, lateral margin thickened and abruptly widened in apical half, apex rounded; paramere slightly curved in lateral view, nearly parallel-sided (Fig. 14 G, H); sipho curved and apex acuminate (Fig. 14 F). Female. Similar to male in all aspects externally. Genitalia with genital plates triangular, not elongate, styli with setae; spermatheca elongate, uniform (Fig. 14 I). Etymology. This species is named in honor of a great lepidopterist, Dr. Mirna Martins Casagrande. Discussion. G. m i r n a e differs from the other species of the genus because it is the only species with uniform color pattern reddish yellow. FIGURE 14. Gordonoryssomus mirnae sp. nov. (A���E) Habitus: (A) dorsal; (B) ventral; (C) lateral; (D) frontal view. (E) Abdomen; (F���H) male genitalia: (F) sipho; (G) tegmen, ventral; (H) lateral view. Female terminalia: (I) coxites. Type material. ��� PERU, M. de Dios, Par-/que Manu, Pakitza 340m / 11 �� 55 ¨ 48 ���S 71 �� 15 ¨ 18 ���W/ 27 Sep 1991 / Leg. M. Casagrande��� ���♂��� [white label]. ��� HOLOTYPE / Gordonoryssomus / mirnae /Almeida & Santos, 2014 ��� [red label]. 1 ex. ��� DZUP / 188196 ��� [DZUP]; ���Chuani, Dept./La Paz/ Bol��via /May, 1925,/G. L. Harrington��� ���♀��� [white label]. ��� PARATYPE / Gordonoryssomus / mirnae /Almeida & Santos, 2014 ��� [yellow label]. 1 ex. [USNM]. Geographical distribution. Peru and Bolivia (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

6. Gordonoryssomus limeirai Almeida & Santos, sp. nov. Type Material: Holotype, male (CZMA); paratype (1), same data of holotype (DZUP). Type locality: “Caracol/ Parque Nacional Serra das/Confusões, Andorinha, Piauí, Brazil ”. Geographical distribution: Brazil (Fig. 17).

5. Gordonoryssomus everardoi Almeida & Santos, sp. nov. Type Material: Holotype, male (DZUP). Type locality: ���Nova Friburgo, Brazil ���. Geographical distribution: Brazil (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

7. Gordonoryssomus mirnae Almeida & Santos, sp. nov. Type Material: Holotype, male (DZUP); paratype (1), La Paz, Bolivia (USNM). Type locality: ���M. de Dios, Parque Manu, Peru ���. Geographical distribution: Peru and Bolivia (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

6. Gordonoryssomus limeirai Almeida & Santos, sp. nov. Type Material: Holotype, male (CZMA); paratype (1), same data of holotype (DZUP). Type locality: ���Caracol/ Parque Nacional Serra das/Confus��es, Andorinha, Piau��, Brazil ���. Geographical distribution: Brazil (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

2. Gordonoryssomus nigripilosus Almeida & Lima, 1995 Gordonoryssomus nigripilosus Almeida & Lima, 1995: 713���714 (original description, morphological details and male genitalia figured). Type material: Holotype, male (MNHN). Type locality: ���Esp��rito Santo, Brazil ���. Geographical distribution: Brazil (ES) (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024"]}

Gordonoryssomus everardoi sp. nov. (Fig. 12) Description of holotype (male). Length 3.44, width 2.26 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 12 C). General color yellowish red, middle of thorax and abdomen dark brown. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal triangular black spot, translucent at apex; finely, densely punctured; pubescence of semidecumbent yellowish hairs (Fig. 12 D). Elytra yellowish red with a large black spot, extending from the base almost to the apex, reaching the middle of humeral callus (Fig. 12 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs yellowish brown. Metaventrite finely, sparsely punctured medially and anteriorly, coarsely, densely punctured laterally and posteriorly with punctures large, often confluent. Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally (Fig. 12 B). First abdominal ventrite with postcoxal line extending beyond middle of first ventrite; precoxal area coarsely punctured on base of cavity (Fig. 12 E). Genitalia simple; basal lobe as longer than the parameres, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere straight in lateral view, nearly parallel-sided (Fig. 12 G, H); sipho flattened and rounded apex (Fig. 12 F). Female. Unknown. Etymology. This species is named in honor of Dr. Everardo Jos�� Grossi, a doctor with a passion for beetles. Discussion. The color pattern of this species is very similar with G. deyrollii, but it differs in body length, and also male genitalia, with basal lobe almost the same length as the parameres, and J-shaped sipho. It also differs from all other species of the genus by the color pattern and male genitalia. Type material. ��� BRAZIL, R.J., Nova/Friburgo, Sans Souci/ 21 -I- 2007 1050 m /P. Grossi col.��� ���♂��� [white label]. HOLOTYPE / Gordonoryssomus / everardoi /Almeida & Santos, 2014 ��� [red label]. 1 ex. ��� DZUP / 131938 ��� [DZUP]. Geographical distribution. Nova Friburgo, Sans Souci, Rio de Janeiro, Brazil (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860

Gordonoryssomus limeirai sp. nov. (Fig. 13) Description of holotype (male). Length 3.71, width 2.78 mm. Form elongate oval, widest at middle of elytra, somewhat flattened dorsoventrally (Fig. 13 C). General color and middle of thorax and abdomen brownish yellow. Outline of pronotum and elytron discontinuous. Pronotum yellowish red, with a basal triangular black spot, translucent at apex; finely, densely punctured; pubescence of semierect yellowish white hairs (Fig. 13 D). Elytra yellowish red with a large black spot, extending from the base almost to the apex, reaching the middle of humeral callus. With a brownish yellow spot in sutural margin along almost 2 / 3 of the suture (Fig. 13 A). Elytral punctures intermixed coarse and fine, dense, slightly larger than on pronotum; sparse in the discal area. Antenna, mouthparts, hypomera, epipleura, and legs brownish yellow. Metaventrite finely, sparsely punctured medially and anteriorly; coarsely, densely punctured laterally and posteriorly with punctures large, often confluent (Fig. 13 B). Abdominal ventrites coarsely punctured, punctures separated by about a diameter or less, punctures becoming coarser laterally. First abdominal ventrite with postcoxal line almost touching the posterior margin of the first abdominal ventrite; precoxal area coarsely punctured on base of cavity (Fig. 13 E). Genitalia simple; basal lobe equal in length to the paramere, lateral margin thickened and abruptly widened in apical half, apex bluntly pointed; paramere straight in lateral view, nearly parallel-sided (Fig. 13 G, H); sipho flattened, sinuous at apex (Fig. 13 F). FIGURE 13. Gordonoryssomus limeirai sp. nov. (A–E) Habitus: (A) dorsal; (B) ventral; (C) lateral; (D) frontal view. (E) Abdomen; (F–H) male genitalia: (F) sipho; (G) Tegmen, ventral; (H) lateral view. (I–J) Female terminalia: (I) spermatheca; (J) coxites. Female. Similar to male in all aspects externally. Genitalia with genital plates triangular, not elongated, styli with setae; spermatheca elongate, not uniform, with narrow apex (Fig. 13 I, J). Etymology. This species is named in honor of the dedicated entomologist, Dr. Francisco Limeira de Oliveira. Discussion. The color pattern is similar to that of G. delicatus, but it differs in length and the size of the sutural spot. Female genitalia with genital plate not elongate, somewhat triangular as in G. deyrollii, G. delicatus and G. mirnae. G. limeirai differs from all other species of the genus by the postcoxal line which almost touches the posterior margin of the first abdominal ventrite and male genitalia with wider basal lobe. Type material. “ BRAZIL (PI), Caracol/Parque Nacional Serra das/Confusões, Andorinha, 515 m,/ 09º08’ 27.8 ”S / 43 º 33 ’ 42.1 ”W / CZMA ” “Suspensa dupla (20 metros)/ 20-30.IX. 2013, J.A. Rafael,/F. Limeira de Oliveira &/T.T.A. Silva cols./ CZMA ” “♂” [white label]. “ HOLOTYPE / Gordonoryssomus / limeirai /Almeida & Santos, 2014 ” [red label]. 1 ex. [CZMA]; “ BRAZIL (PI), Caracol/Parque Nacional Serra das/Confusões, Andorinha, 515 m,/ 09º08’ 27.8 ”S / 43 º 33 ’ 42.1 ”W / CZMA ” “Suspensa dupla (20 metros)/ 20-30.IX. 2013, J.A. Rafael, / F. Limeira de Oliveira & /T.T.A. Silva cols./ CZMA ” “♀” [White label].“ PARATYPE / Gordonoryssomus / limeirai / Almeida & Santos, 2014 ” [yellow label]. 1 ex. “ DZUP / 245996 ” [DZUP]. Geographical distribution. Brazil (PI) (Fig. 17).

3. Gordonoryssomus nigrus Almeida & Lima, 1995 Gordonoryssomus nigrus Almeida & Lima, 1995: 713���715 (original description, morphological details and male genitalia figured). Type material: Holotype, male (USNM). Type locality: ���Santa Cruz, Bolivia ���. Geographical distribution: Bolivia (Fig. 17)., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024"]}

1. Gordonoryssomus deyrollii (Crotch, 1874) Oryssomus deyrollii Crotch, 1874: 292 (original description); Gordon, 1987: 35 (catalogue). Oryssomus deyrollei: Korschefsky, 1931: 176 (catalogue); Gordon, 1974: 147 (systematics). Gordonoryssomus deyrollei: Almeida & Lima, 1995: 711���712 (new combination, description, morphological details and male genitalia figured). Type material: Holotype, male (UMZC, ��� Deyrollii Braz��� [genitalia slide-mounted]). Type locality: Brazil. Geographical distribution: Brazil (RJ, MG, SC), Chile (?) (Fig. 17). Remarks: All the material studied has distribution in Brazil, although Almeida & Lima (1995) listed 2 specimens from Chile, belonging to Germain material, from Sicard Collection (MNHN). Gonz��lez (2008) remarked that this material is possibly mislabeled, since there are other mistakes in this collection., Published as part of Almeida, L��cia M. & Santos, Paula B., 2014, Synopsis of Oryssomini Gordon (Coleoptera: Coccinellidae) from the Neotropical region with new species of Oryssomus Mulsant, Pseudoryssomus Gordon and Gordonoryssomus Almeida & Lima in Zootaxa 3846 (1), DOI: 10.11646/zootaxa.3846.1.2, http://zenodo.org/record/250860, {"references":["Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, 311 pp.","Gordon, R. D. (1987) A catalogue of the Crotch collection of Coccinellidae (Coleoptera). Occasional Papers on Systematic Entomology, 3, 1 - 46.","Korschefsky, R. (1931) Coccinellidae. I. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Pars 118. W. Junk, Berlin, pp. 1 - 224.","Gordon, R. D. (1974) A review of the Oryssomini, a new tribe of Neotropical Coccinellidae (Coleoptera). The Coleopterists Bulletin, 28 (3), 145 - 154.","Almeida, L. M. & Lima, I. M. M. (1995) Revisao do genero Oryssomus Mulsant (Coleoptera, Coccinellidae, Exoplectrinae, Oryssomini) e descricao de Gordonoryssomus, gen. n. Revista Brasileira de Zoologia, 12, 701 - 718. http: // dx. doi. org / 10.1590 / s 0101 - 81751995000300024","Gonzalez, G. (2008) Lista y distribucion geografica de especies de Coccinellidae (Insecta: Coleoptera) presentes en Chile. Boletin del Museo Nacional de Historia Natural, 57, 77 - 107."]}