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Hyposoter carbonarius (Ratzeburg, 1844) Campoplex carbonaria Ratzeburg, 1844: type ♀, lost (Horstmann, 2013). syn. Anilasta pectinata Thomson, 1887: Horstmann (2013) syn. Campoplex melanarius Holmgren, 1860: Horstmann (2013) Horstmann (2013) records that the type material of this species is lost, but states that the species is easily identifiable from a combination of the original description and host records given for it by Ratzeburg (Orgyia antiqua (Linnaeus) and Dicallomera fascelina, (Linnaeus)). It is a comparatively large and robust species, much more so than any other similar species. Although Horstmann’s draft key separated it on the basis that the maxillary palps are black rather than yellow, which is indeed usually the case, we have seen a number of specimens, mostly males, with yellow palps, so that this criterion is not infallible. Horstmann’s original key has been altered to accomodate this. The hosts for H. carbonarius are recorded by Shaw et al. (2016) as several species of low-feeding heathland Erebidae (Lymantriinae), Dicallomera fascelina (Linnaeus), Orgyia antiqua (Linnaeus) and O. antiquoides (Ḩber). There is a single reared male in NHMUK, from Dicallomera fascelina. In addition, Kukal and Kevan (1987) provide an account of the effect of parasitism by this species (as H. pectinatus) on the lymantriine moth Gynaephora groenlandica (Wocke) at the limits of vegetation in the high Canadian arctic. We have seen specimens from the UK, Germany and the Netherlands. Its reported range includes most northern and central European countries, Greenland and Canada. The few captured specimens we have seen were taken in May and June; however, Shaw et al. (2016) record more than one annual generation in N.W. Europe, with the winter being passed in the hibernating larva of D. fascelina., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 37, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Ratzeburg, J. T. C. (1844) Die Ichneumonen der Forstinsecten in forstlicher und entomologischer Beziehung. Nicolaischen Buchhandlung, Berlin, 224 pp. https: // doi. org / 10.5962 / bhl. title. 11094","Horstmann, K. (2013) Revisionen von Schlupfwespen-Arten XVII (Hymenoptera: Ichneumonidae). Mitteilungen Munchener Entomologischen Gesellschaft, 103, 1 - 14.","Thomson, C. G. (1887) XXXV. F ˆ rs ˆ k till uppstallning och beskrifning af aterna inom slagtet Campoplex (Grav.) Opuscula Entomologica, Lund, XI, 1043 - 1182.","Holmgren, A. E. (1860) F ˆ rs ˆ k till uppstallning och beskrifning af de i Sverige funna Ophionider (Monographia Ophionidum Sueciae). Kongliga Svenska Vetenskapsakademiens Handlingar, 2 (8), 1 - 158.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222.","Kukal, O. & Kevan, P. G. (1987) The influence of parasitism on the life history of a high arctic insect, Gynaephora groenlandica (Wocke) (Lepidoptera: Lymantriidae). Canadian Journal of Zoology, 65 (1), 156 - 163. https: // doi. org / 10.1139 / z 87 - 022"]}

Hyposoter pechipogator Galsworthy & Shaw sp. nov. (Figs. 26–28) Diagnosis. In the group with black metasoma, pale fore and mid coxae, and more or less orange hind tibiae, this species can be distinguished from all but H. monensis by its rugose area petiolaris. It can be distinguished fairly easily from the latter by the presence of strong punctures on the lower mesopleuron, and the emarginate seventh tergite. Description. Female. Fore wing length: 5 mm. Head: face longer than broad, granulate, with rather small punctures spaced more than one diameter apart; clypeus rather broad and only shallowly convex, the punctures a little larger and closer than on face, with apical margin narrowly smooth and outflexed; mandibles yellow, only slightly tapering; frons and vertex sculptured as face, temples abruptly narrowed and almost flat behind eyes; first flagellomere about 5.0x longer than wide, both antennae of unique holotype missing apical flagellomeres. Mesosoma: mesoscutum granulate and rugose at rear and along lines of notauli, which are not however incised; upper part of mesopleuron, including speculum, very evenly and finely granulate throughout, unpunctured except along leading edge; lower mesopleuron granulate with rather close deep punctures throughout; metapleuron and hind coxa similarly sculptured, but with slightly finer punctures; upper face of propodeum finely granulate, but petiolar area, especially lower part, becoming rugose; area basalis a short triangle; area superomedia lozenge-shaped, its front edges clearly defined and continuing into costulae, the edges behind the costulae only faintly indicated, and the rear boundary formed by the apex of the strong triangular carinae defining the petiolar area. Wings narrow, 1 cu-a slightly postfurcal, areolet very small and long-stalked, nervellus slightly curved. Metasoma: petiole of first segment smooth, shiny and unpunctured; postpetiole finely granulate, unpunctured, very rounded and convex, with a marked longitudinal pit at base; other metasomal segments very finely granulate and finely punctured, seventh segment with a V-shaped emargination; ovipositor sheaths slightly shorter than depth of metasoma. Colour: black; antenna dark brown, tips of scape and pedicel with small yellow spots; fore and mid coxae, trochanters and trochantelli yellow, mid coxa orange at base; hind coxa black; rest of legs uniform orange; second metasomal sternite clear yellow, other sternites dark brown, except for a small amount of yellow on distal margins. Holotype: ♀, Wyre Forest, Worcs, SO 744761, UK, ex Pechipogo strigilata, em. 1.8.2002, D.Grundy (per M.Bloxham). (NMS). Male unknown. Biology. The unique holotype was reared in the UK from a larva of Pechipogo strigilata (Linnaeus) (Lepidoptera, Erebidae) by D. Grundy (per M. Bloxham). Remarks. The holotype has the metasoma and the right fore wing mounted separately on a card on the same pin. A second female specimen in the NMS collection, Malaise trapped, agrees very well with the description of the holotype in most respects, including the heavy and obvious punctation on the lower mesopleuron, and the longitudinal pit at the base of the postpetiole, but has a longer area superomedia with the apical edges better defined. It may well belong to the same species. Details are: ♀, Abbots Moss, Cheshire, SJ5868, Malaise trap 2, Quercus/Betula/Pinus, 25.viii.–21.ix.86, R.R.Askew NMSZ 1988.002.

Hyposoter ruficoxator Galsworthy & Shaw sp. nov. (Figs. 37, 38) Diagnosis. One of only two species with an all black metasoma and red hind coxa. It is easily distinguished from the other, Hyposoter coxator, by the colour of the hind tibia, as in the key. Description. Female: fore wing 4 mm. Head: face longer than broad, distinctly narrowed ventrally, with close-set punctures distinctly visible on a granulate surface; clypeus fairly narrow, convex, evenly rounded, sculptured as face; malar space about half basal width of mandibles, latter strongly narrowed towards apex, with a wide ventral flange to about two thirds; frons granulate with close-set punctures, the latter disappearing on vertex, except between the two hind ocelli; temples sharply narrowed behind eyes, very short in dorsal view, granulate with small punctures; hypostomal carina disappearing well before meeting oral carina; antenna slender, gently tapering, with 39 flagellomeres in the unique female. Mesosoma: mesoscutum and scutellum very finely granulate throughout, the lines of the notauli marked by a slight rugosity; pronotum similarly sculptured, without epomia, but with some transverse rugae in lower part; mesopleuron granulate throughout, including speculum, with small rugae in prefoveal depression, and with scattered punctures in ventral part only; metapleuron granulate with punctures; pronotum finely granulate, slightly rougher in petiolar area; carinae very reduced, with only a short basal carina, not reaching the base of the propodeum, the proximal two sides of the area superomedia, and the costulae visible; petiolar area without carinae. Wings narrow, 1 cu-a interstitial; areolet small, rhomboid, short-stalked. Metasoma: first tergite with upper side very finely granulate, petiole narrow and parallel-sided until close to apex, postpetiole longer than broad; second and third tergites elongate, dull with very fine sculpture and minute punctures; fourth to seventh tergites progressively more shiny, with more visible, but still very small, punctures; sixth and seventh tergites entire; ovipositor short, projecting very slightly beyond tip of metasoma. Colour: black; mandibles except for teeth, underside of scape, tegulae, fore and mid coxae and trochanters yellow; apex of clypeus, rest of fore and mid legs, hind coxa, trochanters and femur, and sternites one and two orange; hind femur dull whitish, with very slight subapical dark mark, and apical eighth darkened; hind tarsi dull whitish with extreme apices darkened; tergites two and three with faint reddish wash laterally. Male: very similar to female, but with more extensive carination on the propodeum, including faint borders to the petiolar area; 38 flagellomeres; colour identical, except orange on clypeus more extensive, and red wash at sides of tergites two and three lacking. Holotype: ♀, Mergentheim [Germany], 30.vi.1956, red label ‘1070’, label in Horstmann’s hand ‘cf. coxator Hinz’ (ZSM). Paratypes: 2♁, details as holotype. (ZSM). Remarks. Biology unknown. Identified by Horstmann as a new species.

Hyposoter horstmanni Galsworthy & Shaw sp. nov. (Figs. 11–13) Diagnosis. Immediately recognisable by the unusual sculpture of the propodeum, with a broad area superomedia joined to the area petiolaris, completely lacking costulae, and uniformly granulate throughout. The colour of the metasoma is also distinctive, with the rather elongate second tergite entirely black, and the other tergites dark in the centre with a broad orange wash laterally. This coloration is shared with H. rufovariatus, which however has the seventh tergite entire. Description (female). Fore wing length 5 mm. Head: face slightly wider than long, unpunctured and covered with fine cellular microsculpture; clypeus rather narrow, shallowly convex, sculptured as face, but smooth towards apex and with a sharp, well-defined apical edge; mandibles narrowing only slightly; malar space about equal to basal width of mandibles; frons, vertex and temples sculptured as face, unpunctured; temples narrowing sharply behind eyes, and occiput precipitate behind ocelli; antenna very narrow, first flagellomere about 5.0x longer than wide, 30 flagellomeres. Mesosoma: pronotum granulate, finely striate across depression, without epomia; mesoscutum and lower part of mesopleuron finely and uniformly granulate without punctures; notauli absent; speculum smooth and shining, but small; rest of upper mesopleuron finely striate; metapleuron finely rugose, contrasting with lower mesopleuron; propodeum sloping almost from base, finely granulate throughout; area basalis a very small, narrow triangle; area superomedia 5-sided, broadly elongate, seamlessly joined to egg-shaped petiolar area; costulae absent; legs rather narrow, especially femora; hind coxa uniformly granulate. Wings narrow, 1 cu-a strongly postfurcal, areolet quadrangular and sessile, nervellus reclivous, slightly curved. Metasoma: first segment of metasoma flattened at base and narrowed centrally; base smooth and shining, rest of petiole and postpetiole finely granulate; postpetiole curving outwards from petiole anteriorly, approximately rectangular, with a smooth patch at apex pointing forwards like a tooth; 2nd tergite about 2.0x as long as broad, finely granulate, as are remaining tergites; 7th tergite deeply incised; ovipositor length 1.3x depth of metasoma. Colour: black; mandibles, palps and tegulae yellow; all coxae and hind trochanter dark brown; rest of legs orange, hind tibia with only faint darkening at extreme apex; tergites from 3 onwards with orange wash laterally; antenna uniformly dark brown. Male: unknown. Holotype: ♀, Ober-Harz, Torfhaus [Germany], ca 800m, 11.ix.1955, E. Bauer, Omorgus? nov. spec. ♀, sp. nov. 2 Horstm. (ZSM). Paratypes: 1♀, same data as holotype (ZSM). UK: 2♀, Easter Ross NH 4689, Mal. Tr. Native pinewood, 8.89, I. MacGowan, NMSZ 1992.144 ; 1♀, Rannoch, Perths, Native Pinewood, Mal. Trap, NN 5655, I. McGowan, 8.90, NMSZ 1992.002 ; 1♀, Loch Garten, Inv., Malaise Trap, Pinus and Betula, J.A. Owen, July 84 (all in NMS). Biology: unknown; there is a probable association with old pine forest. Remarks: identified as a new species by Horstmann.

Hyposoter rufovariatus (Schmiedeknecht, 1909) Anilastus rufovariatus Schmiedeknecht, 1909: lectotype ♀ in TLHR, designated by Oehlke & Townes (1969) and examined by Horstmann. We have seen two specimens in Horstmann’s collection identified by him as this species. The species is very similar in coloration to H. horstmanni sp. nov., with the tergites beyond the second darkened in the centre and orange laterally, but the seventh tergite is entire, and the second tergite shorter and broader. The propodeum has costulae present, and the sculpture is much coarser than in horstmanni, and rugose in the area petiolaris. The biology is unknown. We have seen specimens from Germany, and one from Georgia. There are records of uncertain reliability from France, and Tunisia. One specimen from the third author’s collection was barcoded (BOLD Sample ID CollHH3496)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 67, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Schmiedeknecht, O. (1909) Opuscula Ichneumonologica. IV. Band. Fasc. XXI - XXIII. Ophioninae. s. n., Blankenburg in Th ¸ ringen. [unknown pagination]","Oehlke, J. & Townes, H. K. (1969) Schmideknechts Ichneumonidentypen aus der Kollektion des Museums Rudolstadt. Beitrage sur Entomologie, 19, 395 - 412."]}

Hyposoter ebenitor Aubert, 1972 Hyposoter ebenitor Aubert, 1972: holotype ♀ in MZLS (Klopfstein et al., 2011). This species has an ovipositor which projects well beyond the metasoma, and is therefore superficially similar to H. ebeninus and H. rhodocerae. It is, however, easily distinguished from both (and from virtually any other European Hyposoter) by the smooth surface of most of the mesopleuron, covered with fairly close punctation. Both the other species have the same area finely granulate, with any punctures almost invisible. The species has a more southerly distribution than Hyposoter ebeninus, and has been reared from various species of Pieridae in the genera Euchloe, Pontia and Zegris (Lepidoptera, Pieridae) (Shaw et al. 2016). 1 specimen in NHMUK was reared from Pontia daplidice (Linnaeus) in Israel. We have seen specimens from France, Spain, Cyprus, Turkey, Israel and Iran. Specimens in NHMUK were all taken in June., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 43, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Aubert, J. - F. (1972) Etude commentee de nouveaux lectotypes choisis dans les collections HOLMGREN et THOMSON (Hym. Ichneumonidae). Entomologica Scandinavica, 3, 145 - 152. https: // doi. org / 10.1163 / 187631272 X 00229","Klopfstein, S. & Baur, H. (2011) Catalogue of the type specimens of Ichneumonidae (Hymenoptera) in the Jacques F. Aubert collection at the Musee de Zoologie, Lausanne, Switzerland. Zootaxa, 3081 (1), 1 - 90. https: // doi. org / 10.11646 / zootaxa. 3081.1.1","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222."]}

Hyposoter orbator (Gravenhorst, 1829) Campoplex orbator Gravenhorst, 1829: lectotype ♀ in UWCP, designated by Horstmann (2000), examined by Horstmann. syn. Limneria rufa Bridgman, 1882 (Horstmann, 1972) The species is notable for its entirely red metasoma, without any black markings. The whole postpetiole is usually red, with only the petiole black. There is always a clear glymma. There is one modern rearing record, from Minucia lunaris ([Denis & Schifferm̧ller]) (Lepidoptera, Erebidae) in France (Shaw et al. 2016). A specimen from Belgium in Horstmann’s collection is mounted with a cocoon and marked simply ‘ex Lep’, but the larval head capsule attached to the cocoon has been identified by Dr. A. Giusti at the NHM as that of Malacosoma neustria Linnaeus (Lepidoptera, Lasiocampidae). There are nineteenth century records in the UK of rearing (as Limneria rufa) from Lasiocampa quercus, which are credible (eg. Bignell 1883). The lectotype is unlabelled, but probably from northern Italy (Horstmann 2000). We have seen specimens from the UK, France, and Belgium: the species has also been reported from Austria, Czechoslovakia, Germany, and Israel., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 57, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Gravenhorst, J. L. C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https: // doi. org / 10.5962 / bhl. title. 65750","Horstmann, K. (2000) Typenrevisionen der von Gravenhorst beschriebenen oder gedeuteten Campoplex-Arten (Hymenoptera, Ichneumonidae). Linzer biologische Beitrage, 32 (2), 1203 - 14.","Bridgman, J. B. (1882) Further additions to the Rev. T. A. Marshall's catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1882, 141 - 164. https: // doi. org / 10.1111 / j. 1365 - 2311.1882. tb 01574. x","Horstmann, K. (1972) Type revision of the species of Cryptinae and Campopleginae described by J. B. Bridgman (Hymenoptera: Ichneumonidae). Entomologist, 105, 217 - 228.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222.","Bignell, G. C. (1883) Note on Limneria rufa, Brdg., L. brischkei, Brdg. and Rhogas reticulator Nees. Entomologist, 16, 69."]}

Limneria aliena Brischke, 1880 Horstmann’s notes indicate that the type material of this species has been destroyed, as is the case with much of Brischke’s collection. It was described in Limneria from a single male, and Schmiedeknecht (1909) confirms that it was at that point still known only from the male. The original description is insufficient to show that it belongs to Hyposoter at all, but if it did, it could apply to more than one species. Aubert (1968a) transferred the taxon to Hyposoter, but this was on the basis of a female in his collection which he described as ‘femelle nouvelle’; although he stated that it conformed in all respects to Brischke’s description, he went on to state that as it lacked almost all red coloration on the metasoma (in contrast to Brischke’s description) it might be described as a new form ‘ nigrior ’. This is hardly a convincing attribution, and we regard alienus as a nomen dubium. There are five specimens in the Horstmann collection marked ‘ Hyposoter alienus’ by R.Bauer. One, which has been marked ‘? Olesicampe sp. ’ by Horstmann, belongs to the species similar to H. caedator and H. dubitatus referred to under H. caedator (q.v.) as a probable Olesicampe. The other four are specimens of H. castaneus sp. nov. described below. They do not match Brischke’s description of alienus. Hyposoter alienus is on the British checklist (Broad, 2016) by virtue of a record in Ireland by Johnson (1929), who lists a single male in a faunistic list with no indication of the basis for the identification, and we suggest that the name should be removed from the checklist., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 7, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Brischke, C. G. A. (1880) Die Ichneumoniden der Provinzen West- und Ost-Preussen. Schriften der Naturforschenden Gesellschaft in Danzig, 4 (4), 108 - 210.","Schmiedeknecht, O. (1909) Opuscula Ichneumonologica. IV. Band. Fasc. XXI - XXIII. Ophioninae. s. n., Blankenburg in Th ¸ ringen. [unknown pagination]","Aubert, J. - F. (1968 a) Revision des travaux concernant les Ichneumonides de France et 6 e supplement au catalogue de Gaulle. Bulletin de la Societe Linneenne de Lyon, 37, 133 - 4. https: // doi. org / 10.3406 / linly. 1968.6002","Broad, G. R. (2016) Checklist of British and Irish Hymenoptera - Ichneumonidae. Biodiversity Data Journal, 4, e 9042. https: // doi. org / 10.3897 / BDJ. 4. e 9042","Johnson, W. F. (1929) Irish Ichneumonidae. Entomologist's Monthly Magazine, 65, 134 - 137."]}

Rhythmonotus singularis Schmiedeknecht of Campoplex tricolor Ratzeburg syn. n. He records the type material of R. singularis as follows “ Holotype (♁) of R. singularis: square uninscribed pink label, Rhytmonotus sp. an Fenster 25. Juli 1889, „Rhytmonotus singularis # Schmiedek. (Rudolstein).”, Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 7, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256

Hyposoter dolosus (Gravenhorst, 1829) Campoplex dolosus Gravenhorst, 1829: lectotype ♁ designated by Aubert (1968b) in UWCP, examined by Horstmann. syn. Campoplex rufimanus Gravenhorst, 1829: Horstmann (2000) syn. Limneria oculata Tschek, 1871: Hinz (1957) A large species, in comparison with other species running to the same section of the key. The tegula is usually blackish in the female, and may be yellow or blackish in the male, or almost any shade in-between. The centre of the metasoma is a rich red colour, often with a varying amount of irregular blackish wash darkening parts of it. The sixth and seventh tergites are always black. The areae externae and the proximal part of the area superomedia of the propodeum are finely granulate, contrasting with the strongly rugose sculpture of the rest of the propodeum. A parasitoid of moths of the erebid subfamily Arctiinae, including Arctia caja (Linnaeus), Coscinia striata (Linnaeus), Cymbalophora pudica (Esper) and Spilosoma lutea (Hufnagel) (Shaw et al. 2016). We have seen specimens from the UK, Spain and Germany. It has been reported from most European countries as far east as Ukraine, and also from Tunisia. Phenological data shows specimens taken from March to September, in every month except July; rearings suggest that it is plurivoltine, overwintering in the host larva (Shaw et al. 2016)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 41, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Gravenhorst, J. L. C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https: // doi. org / 10.5962 / bhl. title. 65750","Aubert, J. - F. (1968 b) Fixation des types, lectotypes et paratypes dans les collections d'ichneumonides et premiѐre liste de types perdus ou conserves. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 41, 175 - 201.","Horstmann, K. (2000) Typenrevisionen der von Gravenhorst beschriebenen oder gedeuteten Campoplex-Arten (Hymenoptera, Ichneumonidae). Linzer biologische Beitrage, 32 (2), 1203 - 14.","Tschek, C. (1871) Ichneumonologische Fragmente. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 21, 37 - 68.","Hinz, R. (1957) Zur systematik und ˆ kologie der Ichneumoniden 1 (Hym). Deutsche Entomologische Zeitschrift, 4, 86 - 90. https: // doi. org / 10.1002 / mmnd. 19570040112","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222."]}

Hyposoter maculatus (Hedwig, 1938) Anilasta maculata Hedwig, 1938: lectotype ♀ in SDEI, designated by Oehlke (1963), examined by Horstmann. We have seen a specimen from Germany in Horstmann’s collection, which he had marked as identical to the lectotype. The species is distinctive, with a broad dark central streak along the metasoma, which is otherwise orange from the third tergite on. It is most similar to Hyposoter prolixa but, apart from the colour differences, it has a very different sculpture on the propodeum, as set out in the key. Nothing is known of the biology. The species is recorded from Poland and Germany., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 52, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Hedwig, K. (1938) Neue schlesische Ichneumoniden (Hymenoptera). Arbeiten uber Morphologische und Taxonomische Entomologie, 5, 221 - 227.","Oehlke, J. (1963) Revision der im Deutschen Entomologischen Institut aufbewahrten Typen palaarktischer Ichneumoniden (Hymenoptera: Ichneumonidae). Beitrage zur Entomologie, 13, 403 - 410."]}

Hyposoter inquinatus (Holmgren, 1860) Limneria inquinata Holmgren, 1860: lectotype ♀ in NHRS, Sweden, designated by Aubert (1980), not examined. Horstmann’s notes suggest that he had not seen the lectotype, but relied on a specimen from Hannover, Germany, in ZSM, which had been compared with the lectotype by Hinz. Usually fairly easy to identify by combination of pale fore and mid coxae, darkish hind femur, and rather dark reddish hind tibia with strong basal and apical infuscations. A fair number of specimens have the hind femur more reddish, especially on the inside, but it is always darker than in other similar species. The bright yellow underside to the scape is a helpful pointer. Specimens with more reddish-brown hind femora might key to Hyposoter rivulator sp. nov., but the latter has a much paler colour in the middle of the hind tibia, and normally a brown underside to the scape. We have seen specimens from the UK and France. The lectotype was from Sweden. The species has been recorded with uncertain reliability from most countries in Europe. There is an early rearing record of doubtful credibility (very possibly a misidentification of Hyposoter tricolor) from Abraxas grossulariata (Linnaeus) (Lepidoptera, Geometridae, Ennominae) (Plaas, 1932). We have seen specimens taken from July to September, but with the overwhelming majority in August., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 48, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Holmgren, A. E. (1860) F ˆ rs ˆ k till uppstallning och beskrifning af de i Sverige funna Ophionider (Monographia Ophionidum Sueciae). Kongliga Svenska Vetenskapsakademiens Handlingar, 2 (8), 1 - 158.","Aubert, J. - F. (1980) Notes sur diverse Ichneumonides mal connues ou inedites. Bulletin de la Societe Entomologique de Mulhouse, 1980 (Janvier - Mars), 1 - 6.","Plaas, G. (1932) Der Stachelbeerspanner Abraxas grossulariata L. in Schleswig-Holstein. Zeitschrift fur Wissenschaftliche Insektenbiologie, 26 & 27, 183 - 191 & 12 - 17."]}

Hyposoter tibialis (Hedwig, 1938) Anilasta tibialis Hedwig, 1938: lectotype ♁ in SDEI, designated by Oehlke (1963) and examined by Horstmann. syn. Hyposoter nigrior Aubert, 1993 syn. nov.: synonymy proposed in Horstmann’s notes after examination of type material. Horstmann made full notes on the lectotype, unfortunately in near illegible handwriting. We have seen a male and a female determined by Horstmann in his collection. The dark hind tibia is very striking in this species, and should make determination straightforward. Of Horstmann’s two specimens, only the male has the reddish underside to the hind tibia, while both have a reddish flush on the underside of the scape, and very regular granulation over the whole of the propodeum. The type is from southern Poland, while Horstmann’s specimens are from Malsch, near Karlsruhe in southern Germany. H. nigrior was described from France. Nothing is known of its biology., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 68, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Hedwig, K. (1938) Neue schlesische Ichneumoniden (Hymenoptera). Arbeiten uber Morphologische und Taxonomische Entomologie, 5, 221 - 227.","Oehlke, J. (1963) Revision der im Deutschen Entomologischen Institut aufbewahrten Typen palaarktischer Ichneumoniden (Hymenoptera: Ichneumonidae). Beitrage zur Entomologie, 13, 403 - 410."]}

Hyposoter placidus (Desvignes, 1856) Campoplex placidus Desvignes, 1856: lectotype ♀ in NHMUK, designated by Fitton (1976), examined. In the female the length of the ovipositor distinguishes this species readily from ebeninus and rhodocerae. The yellow spot at the base of the hind tibia is faint, but usually present. For distinction from clausus, see notes under the latter species. Males may not always be distinguishable from those of ebeninus, but see also under the latter. This is a well-known parasitoid of species of Lycaena (Lycaenidae), including Lycaena dispar (Haworth), L. hippothoe (Linnaeus), L. helle ([Denis & Schifferm̧ller]), and L. phlaeas (Linnaeus) (Shaw et al. 2009, 2016). There are numerous specimens in Horstmann’s collection reared from L. dispar. There are three specimens reared from the same host in NHMUK. We have seen specimens from the UK, Germany, France, the Netherlands, Belgium, Spain and Andorra. We are not aware of records from elsewhere. Limited phenological data shows specimens collected from June to September with a fairly even distribution over the period. One specimen from the third author’s collection was barcoded (BOLD Sample ID CollHH1214)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 59, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Desvignes, T. (1856) Catalogue of British Ichneumonidae in the collection of the British Museum. British Museum, London, 120 pp.","Fitton, M. G. (1976) The western Palaearctic Ichneumonidae (Hymenoptera) of British Authors. Bulletin of the British Museum (Natural History), Entomology Series, 32, 301 - 373.","Shaw, M. R., Stefanescu, C. & van Nouhuys, S. (2009) Parasitoids of European butterflies. In: Settele, J., Shreeve, T., Konvicka, M. & van Dyck, H. (Eds.), Ecology of Butterflies in Europe. Cambridge University Press, Cambridge, pp. 130 - 156.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222."]}

Hyposoter aglyphus Galsworthy & Shaw sp. nov. (Figs. 1, 2) Diagnosis. A combination of red-marked metasoma, pale tegulae, dark fore trochanter and trochantellus, and lack of a glymma separate this species from all but H. morairae. It is easily separated from the latter by the longer ovipositor and longer face. Some specimens have traces of a glymma in the form of an outline, but it is never recessed. Description (female). Wing length ca 5 mm. Head: face slightly longer than broad, longitudinally rugose, with punctures well hidden between the rugae; clypeus distally curved, only slightly convex, apex sharp, similarly but less heavily sculptured than face; mandibles strongly tapering, with broad flange in basal two thirds; malar space about equal to base of mandible; frons granulose in centre, granulate laterally; vertex and temples finely granulate; antenna slender and tapering, first flagellomere about 3x as long as wide; 27–28 flagellomeres. Mesosoma: mesoscutum with fairly large punctures about one diameter apart spread over its whole surface on a strongly granulate background, and becoming centrally slightly rugose; anterior and lower part of mesopleuron with small punctures about 2 diameters apart on a lightly granulate background; speculum shining, faintly striate; area in front of speculum striate, striations continuing towards upper edge of mesopleuron where they are strong; metapleuron with larger punctures on a granulate background; propodeum punctate on anterior lateral areas, otherwise rather coarsely rugose; area basalis reduced to a broad keel; area superomedia about as long as wide, 5-sided, its posterior end marked only with rugae; carinae of lateral areas, including costulae, strong. Wings narrow, 1 cu-a slightly postfurcal, areolet short-stalked, nervellus reclivous. Metasoma: petiole of first tergite smooth and polished, setose from about one third, and with faint very fine cellular microsculpture, rather square in cross section, and without glymma; dorsolateral carina absent; postpetiole broad and rounded; 2nd tergite about as long as wide; 2nd tergite onwards with unpunctured, almost smooth surface, but somewhat dull as a result of very fine transverse microsculpture; 6th and 7th tergites entire; ovipositor about 1.5x apical depth of metasoma, minimally curved upwards, with fine subapical notch. Colour: black; mandibles dark brown at base, yellowish towards apex, with teeth deep red-brown; tegula yellow; all coxae, trochanters and trochantelli black; all femora orange; fore tibia orange inside, white outside; mid tibia orange, but proximal half white on outside; hind tibia orange, with broad pale proximal ring and faint dark subproximal and distal rings; fore tarsus orange, middle and hind tarsi orange-brown with basitarsus broadly white proximally; tergite one of metasoma black with a variable amount of orange ranging from centre of postpetiole to almost whole of postpetiole; tergites 2 and 3 orange; tergite 4 orange proximally; rest of metasoma dark brown to black. Holotype: ♀, [UK] Llangristiolus, Anglesey, SH434736, Malaise Trap by hayfield, 7–27.viii.82; S.A. & D.C.Wilkinson, RSMNH1982.066 (NMS). Paratypes: 1♀, details as holotype ; 1♀, Praa Sands, Cornwall 29.7.76, M.R.Shaw, Hyposoter sp. det R. Hinz ; 1♀, Ainsdale NNR, Lancs, 13.6.75, M.R.Shaw ; 1♀, France, Bédoin, Vaucluse, 7–16.vii.91, M.R.Shaw (all in NMS). Biology: unknown. A male in the NMS collection which is generally similar to this species, and would run to it in the key, was reared from Aspitates ochrearia (Rossi) (Lepidoptera, Geometridae) found at Dungeness in Kent. The specimen lacks the pale proximal ring on the hind tibia which is present in the type series of females, and has a much darker hind tarsus, and therefore may not belong to the same species, but this host would fit the generally seashore or xerophilic localities of most of the type series., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on pages 31-32, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256

Hyposoter morairae Galsworthy & Shaw sp. nov. (Figs. 21, 22) Diagnosis. Separated from Hyposoter orbator, which it otherwise resembles, by its complete lack of a glymma. Other similar species all have the apex of the metasoma darkened or black. Description. Female. Wing length 5 mm. Head: face quite strongly transverse, longitudinally rugose, with punctures well hidden between the rugae; clypeus distally curved, only slightly convex, apex sharp, similarly but less heavily sculptured than face; mandibles strongly tapering, with broad flange in basal two thirds; malar space small, about one third width of base of mandibles; genal carina meeting hypostomal carina at acute angle short of base of mandible; frons rugose in centre, granulate laterally; vertex and temples finely granulate; antenna slender and tapering, first flagellomere about 3x as long as wide. The unique female is missing one antenna and the tip of the other. Mesosoma: mesoscutum with fairly large punctures about one diameter apart spread over its whole surface, on a strongly granulate background; anterior and lower part of mesopleuron with small punctures about 2 diameters apart on a lightly granulate background; speculum shining and partly smooth; area in front of speculum striate, the striations continuing towards the upper edge of the mesopleuron where they are strong; metapleuron with larger punctures on a granulate background; propodeum unevenly punctate on a coarse granulate background; area basalis V-shaped; area superomedia longer than wide, 5-sided, its posterior end marked only with rugae; carinae of lateral areas, including costulae, strong. Wings narrow, 1 cu-a slightly postfurcal, areolet short-stalked. Metasoma: petiole of first tergite smooth and polished, rather square in cross section, and without glymma; dorsolateral carina present, running faintly on as far as spiracle; postpetiole narrow, an elongate oval, convex, finely granulate; 2nd tergite 1.5x as long as wide; 2nd tergite onwards with unpunctured almost smooth surface, but somewhat dull as a result of very fine transverse microsculpture; 6th and 7th tergites entire; ovipositor shorter than apical depth of metasoma, minimally curved upwards, with strong subapical notch. Colour: black; mandibles dark brown at base, yellowish towards apex, with teeth deep red-brown; tegulae yellow; all coxae, trochanters and trochantelli black; all femora orange; fore tibia orange inside, white outside; mid tibia brown, but basal half white on outside; hind tibia orange, with broad white basal ring and dark subbasal and apical rings; fore tarsi orange, middle and hind tarsi orange-brown with basitarsus broadly white at base; petiole of first tergite black, postpetiole entirely red-orange; 2nd tergite orange with broad basal and narrow apical black bands; tergites 3-5 entirely orange; tergite 6 orange with strong brown suffusion; tergite 7 brown. Holotype: ♀, Esp [Spain], Alicante, Moraira, 90m ; 14–22.v.1989 ; garrigue; rec. R. Wahis ; Mal. trap; 2 labels in Horstmann’s handwriting “ Hyposoter sp 3 ” and “ cf orbator ♀ ”. In ZSM. Biology. unknown. Remarks. Identified by Horstmann as an undescribed species., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 54, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256

Hyposoter castaneus Galsworthy & Shaw sp. nov. (Figs. 3–5) Diagnosis. Most specimens have the metasoma darkish brown, which is unusual in the genus and an immediately obvious distinguishing feature; but the metasoma can be more orange, and the species is therefore included twice in the key. Apart from the key characters, the species is distinguished by an unusually broad clypeus and face, which separate it readily from the otherwise quite similar (in some colour forms) H. caedator. The sculpture of the mesopleuron, with clear transverse rugae in front of a smooth speculum, is also distinctive. Description. Fore wing 3.5–4 mm. Head: face twice as broad as long, rugosely sculptured throughout, without evident punctures; clypeus short, curved at sides, similarly sculptured, with narrow, sharp, unsculptured apex; malar space equal to basal width of mandible; mandible tapering, with ventral flange narrow; genal carina complete, joining oral carina just short of base of mandible; frons and vertex rugosely sculptured, but less strongly than face; temples sharply narrowed behind eyes, very short, similarly sculptured; antenna slender, flagellum with 24–25 flagellomeres, first flagellomere 2.5x as long as wide. Mesosoma: epomia present; mesoscutum centrally with close-set deep punctures on a granulate background, lateral and posterior parts becoming strongly rugose; mesopleuron shining, but with faint linear sculpture throughout; groove in front of fovea strongly striate; anterior and ventral areas, and metapleuron, with strong, evenly spaced punctures on a granulate background; propodeum rather short and rounded; area basalis narrow, V-shaped; area superomedia closed, broader than long, 5-sided, with front lateral edges much longer than rear ones; costulae complete; area petiolaris broadly triangular; whole propodeum rugosely sculptured, most strongly on petiolar area; propodeal spiracle slightly oval. Wings narrow, 1 cu-a interstitial, areolet almost sessile to short-stalked with vein 2 rs -m entering well beyond middle, nervellus gently curved, almost vertical. Metasoma: first tergite with petiole narrowed before postpetiole, and ventral edge sinuate in lateral view; glymma obvious, but shallow; dorsolateral carinae present, but weak; postpetiole triangular, faintly granulate, with symmetrical depressions above and behind spiracles; tergite 2 strongly granulate to two thirds, its posterior part and other tergites only faintly sculptured, and with scattered fine punctures; tergite 6 with posterior border entire; ovipositor slightly longer than apical depth of metasoma, almost straight, with strong subapical notch. Colour: black; apex of underside of scape, mandibles other than extreme base and teeth (which are brown), tegulae, 2nd metasomal sternite yellow; femora, tibiae and fore and mid tarsi orange; hind tarsus brown, basitarsus pale at base; tergite 1 black; tergite 2 black, with posterior edge sometimes orange, tergite 3 orange to brown with orange wash; tergite 4 orange to pale brown; tergites 5 onwards dark brown. Holotype: ♀, [UK] Santon Downham, Norfolk, TL818883, Malaise trap: heath with birch and pine, 25.7– 5.8.83, J.Field, RMSNH 1986.021 (NMS); paratypes: 1♀, Chippenham Fen, Cambs, TL 650693, Malaise trap: carr at reedbed edge, 16–24.6.83, J.Field, RMSNH 1986.021; 1♀, Catfield, Norfolk, TG379201, Malaise trap: abandoned wet meadow, R.T.J.Jarvis, 27.7.–10.8.84 RSMNH 1983.102 (all in NMS); 2♀, Merton Park, London SW 19, TQ2469, Malaise trap in garden, 12–19.vii.14, A.C.Galsworthy (NHMUK); 1♀, [Germany] Augsburg, 4.vii.1959, Hyposoter carbonarius ♀ det R.Bauer, [in Horstmann’s hand] Hyposoter? insulator Aub. Horstm. 2013 (ZSM); 4♀, Nürnberg [Germany], 7 & 11.7.76, 23.7.88, and 27.5.89, all labelled Hyposoter alienus Brke, R.Bauer (ZSM); 1♀, Messkirch, Baden [Germany], 1.9.1969, I.Wall, Hyposoter? meridionellator Aubert det. Riedel, 2014 (private collection Matthias Riedel). Male and biology unknown.

Hyposoter flavicoxa Galsworthy & Shaw sp. nov. (Figs. 7, 8) Diagnosis. Most similar to pale specimens of H. caedator, but easily distinguished by the sculpture of the propodeum, which is granulate throughout, with a broad, ill-defined area superomedia, and by the yellow underside to the scape. The reduced sculpture of the propodeum also distinguishes this species from H. alpicola. Description. Wing length: 4.5 mm. Head: face longer than broad, face and clypeus rather coarsely granulate, making punctures almost invisible; clypeus relatively narrow and strongly convex, with narrow and smooth apical margin; malar space shorter than basal width of mandibles; mandibles strongly narrowed towards apex; frons sculptured much as face, but vertex and temples much more finely granulate; occiput descending immediately behind ocelli, and temples very strongly narrowed behind eyes; antenna somewhat broadened in distal half, but distally tapering: 32 flagellomeres. Mesosoma: pronotum without epomia; mesoscutum finely and evenly granulate throughout, position of notauli marked only by bands of slightly outstanding short ridges; mesopleuron, including speculum, finely granulate virtually throughout without visible punctures, apart from some short longitudinal striae in front of mesopleural fovea and a few longer striae beneath tegular ridge; metapleuron slightly more coarsely rugose, especially in a band along dorsal edge; scutellum finely granulate, keeled only at extreme base; propodeum without clear keels, edges of area superomedia and area petiolaris outlined by irregular lines of rugosity; first lateral areas and front part of area superomedia finely granulate, rest of propodeum strongly transversely rugose; costulae absent, but their position indicated by abrupt change in surface sculpture. Wings narrow; fore wing 1 cu-a postfurcal; areolet sessile; nervellus vertical, slightly curved but not intercepted. Metasoma: first metasomal tergite with almost parallel sides to petiole, widening distally to a rectangular postpetiole, all very finely granulate almost to base of petiole; glymma large and obvious; 2nd tergite 1.2x as long as wide, thyridia small and close to base; surface of 2nd tergite onwards smooth but not polished, apart from a slight rugosity in basal half of 2nd tergite; 6th and 7th tergites entire; ovipositor shorter than apical depth of metasoma, very slightly upcurved, with a strong subapical notch. Colour: black; mandibles, underside of scape, fore and mid coxae, trochanters and trochantelli yellow, sometimes shaded with orange brown on mid coxa; first tergite black with reddish tinge at apex of postpetiole, 2nd tergite orange, darkened at least in basal half, often up to five-sixths, with a narrow black line on apex of tergite; tergites 3–5 orange, 6–7 dark brown to black; all femora, tibiae and tarsi orange, hind tibia usually only faintly darkened at extreme base and apex, sometimes more extensively at apex; visible sternites orange. Male (based on a single specimen): very similar to female, but with fore and mid legs paler, almost clear yellow, and more extensive darkening on the hind part of the metasoma. Remarks: biology unknown. We have seen specimens only from the UK and France. Holotype: ♀, [UK] Tarrington, Herefordshire, SO6140, Malaise trap, 28.vii–6.viii.18, A.C.Galsworthy (NHMUK). Paratypes: 1♀, Chippenham Fen, Cambs, TL 650293, Malaise trap, carr at reedbed edge B, 30.7–11.8.85, J. Field, RMSNH 1986.021 ; 1♀, Abbots Moss, Cheshire, SJ 5868, Malaise Trap 2, Quercus / Betula / Pinus, 5–25.viii.86; R.R.Askew, NMSZ 1988.002 ; 1♀, Lot-et-Garonne, Bernac, Mal. TR. 26.6–3.7.91, R.R.Askew (all NMS). 2♀, Tarrington, Herefordshire, SO6140, light trap, 18–23.viii.16 and 15–22.viii. 2017 ; 1 ♁, 5♀, same location, Malaise trap, 28.vii–6.viii.18, A.C.Galsworthy (all NHMUK)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on pages 43-45, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256

Hyposoter alpicola (Smits van Burgst, 1914) comb.nov. Angitia alpicola Smits van Burgst, 1914 ; holotype ♁ in LEW, examined by Horstmann. syn. Hyposoter caedator postcaedator Aubert, 1964 syn. nov.: lectotype, here designated. Label data: TYPE; HYPOSOTER CAEDATOR Grav. POSTCAEDATOR Aub; J F Aubert 1962 BdR VAUVERT; Si ce n’est pas une forme de CAEDATOR ne correspond aucune autre espèce decrite [in Aubert’s handwriting]; syntypes 1 & 2/?22 Hyposoter caedator postcaedator Aubert 1964 c, labelled by S. Klopfstein 2009. GBIFCH 00832761 [with barcode]; Note: there are two specimens on the pin; the top one is chosen as the lectotype. (Fig. 43) Horstmann (1969), after examining the holotype of alpicola, speculated that this species belonged in Hyposoter. His notes, and his draft key, confirm that he had intended to transfer it to Hyposoter, and that he regarded Hyposoter postcaedator Aubert, 1964, the type material of which he had also examined, as a junior synonym of it. These intentions are now formalised here. We have not ourselves seen the holotype of alpicola, and rely on Horstmann’s notes. We have however seen the lectotype of postcaedator designated above. This is distinct from H. praecaedator in having the last 2 tergites of the metasoma darkened, and a narrow dark apex to tergite 2. The mesopleuron is fully granulate. The species resembles H. caedator more than praecaedator but, unlike caedator, it has a yellow underside to the scape. The holotype of H. alpicola was from near Bolzano in the Italian Alps. Hyposoter postcaedator was described from the south of France. We have seen two specimens of H. alpicola from Norway, both taken in July. The biology is unknown., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 34, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Smits van Burgst, C. A. L. (1914) Ichneumonidae captured in the environs of Bozen (Tyrol) in June 1913. Deutsche Entomologische Zeitschrift, 1914, 325 - 333.","Aubert, J. - F. (1964) Les Ichneumonides du rivage mediterraneen francais (Hym.). 7 e serie: Ichneumoninae, Cryptinae, Ophioninae et Mesochorinae de l'Herault et des Bouches-du-Rhone. Bulletin de la Societe Entomologique de France, 69, 144 - 164. https: // doi. org / 10.3406 / bsef. 1964.20764","Horstmann, K. (1969) Typenrevision der europaischen Arten der Gattung Diadegma F ˆ rster (syn. Angitia Holmgren). Beitrage zur Entomologie, 19, 413 - 472."]}

Hyposoter tricoloripes Viereck, 1911 Anilastus tricoloripes Viereck, 1911: holotype ♀ in USNM, not examined. Similar to H. tricolor, but easily distinguished by the colour of the hind tibia, as in the key. A well-known parasitoid of the Gypsy moth, Lymantria dispar (Linnaeus) (Erebidae; Lymantriinae), for the control of which it was introduced to the USA. It has been recorded from much of central Europe westward to France. It had seemed possible that with the recent spread of the Gypsy moth back to the UK, this parasitoid might follow it. The first author has now taken two specimens in a Malaise trap in northwest Surrey in the UK, and the opportunity is here taken to record it for the first time from the UK: 2 females, Malaise trap, Queen Mary Reservoir, Surrey, TQ0569, 17.vi–9.vii.22 and 16–27.vii.22. We have otherwise seen specimens only from Germany., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 68, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Viereck, H. L. (1911) Descriptions of one new genus and eight new species of Ichneumon flies. Proceedings of the United States National Museum, 40 (1832), 475 - 480. https: // doi. org / 10.5479 / si. 00963801.1832.475"]}

Hyposoter insulator Aubert, 1960 stat. nov. (Fig. 41) Hyposoter carbonarius insulator Aubert, 1960: lectotype ♀, here designated, in MZLS. Label details: TYPE; ♀ ; HYPOSOTER CARBONARIUS Ratz. INSULATOR Aub. J.F.AUBERT 15.8.1959 AJACCIO (CORSE). Syntype 1/3 (3) Hyposoter carbonarius insulator Aubert 1960 g, labelled by S. Klopfstein 2009. GBIFCH 00832765. This was described by Aubert (1960) as a subspecies of Hyposoter carbonarius Ratzeburg, though in Aubert (1961) he makes clear that he was uncertain whether to describe the specimens as a new species or a subspecies of carbonarius. Horstmann examined the syntype series and it is clear from his notes and his draft key that he intended to elevate the taxon to full species status. We have examined one of the syntypes (that designated as lectotype above). The type series is well distinguished from carbonarius, inter alia by the yellow tegulae and palps, the quadrate to elongate area superomedia and the partly smooth speculum and rather finely granulate mesopleuron, with no visible punctures. We therefore here raise it to species status. The partly smooth speculum should also distinguish insulator from other species in this part of the key. The seventh tergite is not emarginate in the female, as it is in ebeninus and rhodocerae. The biology is unknown, and the species is known only from Corsica., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on pages 48-49, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Aubert, J. - F. (1960) Description preliminaire de quelques espѐces et sous-espѐces mediteraneennes de la famille des Ichneumonides. Bulletin de la Societe Entomologique de Mulhouse, 1960 (Septembre - Octobre), 62 - 65. https: // doi. org / 10.3406 / bsef. 1960.20555","Aubert, J. - F. (1961) Les Ichneumonides de Corse (Hym). Annales de la Societe Entomologique de France, 130, 159 - 188."]}

Hyposoter sicarius (Gravenhorst, 1829) Campoplex sicarius Gravenhorst, 1829: holotype ♀ in ZMHB, examined by Horstmann. We have seen a specimen from Germany identified by Horstmann as sicarius by comparison with the type. The species is readily identifiable by the unusual structure of the genal carina, which disappears completely at a level well above the bottom of the eye, and a long way before it would join the hypostomal carina. Sternites 2–4 are uniform bright orange, similar to the tergites. There is an early record (Rudow, 1917) of rearing from Panolis flammea (Denis & Schifferm̧ller) (Lepidoptera, Noctuidae), but this is of doubtful reliability. The type is from Germany and we have seen specimens only from that country. There are published records of the species from many European countries from France to Russia, but possible confusion with H. dubitatus (q.v.) makes these particularly unreliable.

Hyposoter dubitatus (Holmgren, 1860) stat. rev. Limneria dubitata Holmgren, 1860: lectotype ♀ in NHRS, designated by Hinz (1962), examined. Limneria dubitata Holmgren was synonymized with Campoplex sicarius Gravenhorst, 1829 by Schmiedeknecht (1909), though without explanation. Aubert (1972), after examining the lectotype, stated that the synonymy with sicarius was ‘doubtful’. Horstmann’s draft key makes it clear that he had intended to separate dubitatus from sicarius, but we have not found evidence in his notes that he had seen the lectotype of dubitatus. However, we have seen the lectotype, and confirm that it is quite different from the specimen of sicarius in Horstmann’s collection which he had compared with the type of the latter. We therefore formalise here Horstmann’s intention to restore dubitatus to valid species status stat. rev. Three specimens (one male and two females) in Horstmann’s collection from northern Germany had originally been identified by Horstmann as H. thuringiacus Schmiedeknecht, but bear Horstmann’s labels correcting this to H. dubitatus. They match the type of dubitatus very well (see above under H. thuringiacus in the section on species excluded from the key). The species is very similar to H. caedator: see notes under that species for separation of the two. The type material is from Sweden. The two female specimens in Horstmann’s collection mentioned above bear labels ‘ex Epione parallelaria’, i.e. Epione vespertaria (Linnaeus) (Lepidoptera, Geometridae), though there are no cocoons or larval remains. One specimen in NMS was reared in Italy from an unknown geometrid and a further specimen in the French Alps also from an indeterminate geometrid, possibly Eulithis. There are also further specimens in NMS from France and from the UK.

Hyposoter leucomerus (Thomson, 1887) Anilasta leucomera Thomson, 1887: lectotype ♀ in MZLU, designated by Aubert (1966b), examined by Horstmann. syn: Limneria tricincta Holmgren, 1858: Horstmann, 1999 Larger than other similar species other than Hyposoter flavicoxa sp. nov. The hind tibia is striking, with the central colour pale straw, contrasting strongly with the dark basal and apical rings. There is normally a clear black transverse line at the extreme apex of the second metasomal tergite. This species has been reared from Hellinsia tephradactylus (Lepidoptera, Pterophoridae) (Shaw et al. 2016). We have seen specimens from Norway, Germany and the UK. The lectotype is from Sweden. The species has been reported from countries across Europe from Ireland to Turkey, but many of these are older records whose accuracy is doubtful. Very limited phenological data shows specimens taken in May and August. One specimen from the third author’s collection was barcoded (BOLD Sample ID CollHH1256)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 52, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Thomson, C. G. (1887) XXXV. F ˆ rs ˆ k till uppstallning och beskrifning af aterna inom slagtet Campoplex (Grav.) Opuscula Entomologica, Lund, XI, 1043 - 1182.","Aubert, J. - F. (1966 b) Fixations d'Ichneumonides lectotypes dans la collection C. G. Thomson conservee Lund. Opuscula Entomologica, 31 (1 / 2), 125 - 132.","Horstmann, K. (1999) Revisionen von Schlupfwespen-Arten III (Hymenoptera: Ichneumonidae). Mitteilungen Munchener Entomologischen Gesellschaft, 89, 47 - 57.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222."]}

Hyposoter validus (Pfankuch, 1921) Anilastus validus Pfankuch, 1921: holotype ♁ in ZMHB, examined by Horstmann. Horstmann (1986) gave details of the holotype, which he had examined. The partially red metasoma, coupled with dark tegulae and hind femora, and the presence of a glymma, should distinguish the species from all others, except the occasional darker specimen of H. dolosus, from which it should be distinguishable by the key characters. For distinction from pseudovalidus sp. nov., see under the latter. This species has been reported by Kolomiets (1962) as a parasitoid in Russia of the defoliating moth Dendrolimus sibiricus Chetverikov (Lepidoptera, Lasiocampidae), but the accuracy of this determination needs to be checked. We have seen a specimen from Norway in Dr M.Riedel’s collection. Apart from this, its description from Germany and the report from Russia, it has also been reported from Poland., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 69, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Pfankuch, K. (1921) Neue Ichneumoniden. Entomologisches Jahrbuch, 30, 149 - 159.","Horstmann, K. (1986) Typenrevision der von Karl Pfankuch beschriebenen Arten und Formen der Familie Ichneumonidae (Hymenoptera). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 8, 25 - 264.","Kolomiets, N. G. (1962) [Parasites and predators of Dendrolimus sibiricus Tschet]. Izdatel'stvo Sibirskogo Otdeleniya An SSSR, Novosibirsk, 173 pp. [in Russian]"]}

Hyposoter anglicanus (Habermehl, 1923) Anilasta anglicana Habermehl, 1923: holotype ♀ in NHMN, examined. We have examined the female holotype, as had Horstmann. The species is something of a mystery, as it appears to be reliably known only from the holotype specimen. Although Habermehl stated that this specimen was from the environs of Lichfield UK (L. Carr collection), the specimen itself bears no locality label, and the species has now been excluded from the current British checklist because of doubts about the origin of Carr’s specimens (see Perkins 1953; Shaw 2003). The holotype is very similar to some specimens of H. longulus, and may well belong to that species. The biology is unknown., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 34, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Habermehl, H. (1923) Neue englische Ichneumoniden nebst Bemerkungen ¸ ber Ichneumon macrocerus C. G. Thoms. ♀, Cratichneumon foersteri Wesm. ♀, und Grypocntrus cinctellus Ruthe ♁ (Hym.). Konowia, 2, 34 - 43.","Perkins, J. F. (1953) Notes on the British Ichneumoninae with descriptions of new species (Hym., Ichneumonidae). Bulletin of the British Museum (Natural History), Entomology, 3 (4), 9 - 176. https: // doi. org / 10.5962 / bhl. part. 1050","Shaw, M. R. (2003) Adverse comment on the supposed British status of the web-spinning sawfly Cephalcia arvensis Panzer (Hymenoptera: Pamphiliidae). British Journal of Entomology and Natural History, 16 (1), 2."]}

Hyposoter ebeninus (Gravenhorst, 1829) Campoplex ebeninus Gravenhorst, 1829: lectotype ♀ in UWCP, designated by Townes et al. (1965), examined by Horstmann. syn. Limneria vulgaris Tschek, 1871: Hinz (1957) syn. Limnerium brevicaudis Szépigleti, 1916: Horstmann (1978) syn. Angitia rapae Meyer, 1926: Horstmann (1969) Females are easily distinguished from all other species with all dark tergites and sternites in the metasoma, except Hyposoter rhodocerae, by the length of the ovipositor, and can be separated from the latter by the key characters. Males can be more difficult to separate, especially from placidus, but the dorsal face of the propodeum is longer in placidus, especially viewed laterally. The faint yellow spot at the base of the hind tibia is usually visible in placidus, but lacking in ebeninus. Shaw et al. (2016) gave details of rearings of Hyposoter ebeninus from three species of Carcharodus (one since transferred to Muschampia) (Hesperidae) and several Pieridae, notably species of Pieris and Euchloe. Reared specimens in NHMUK are as follows: from Pieris brassicae (Linnaeus): 2♁ 5♀ (Algeria), 1♁ 1♀ (India), 2♁ 2♀ (Turkey), 6♁ 4♀ (Egypt); from Pieris rapae on cabbage (Linnaeus): 3♁ 1♀ (Cyprus); from Euchloe ausonia (Ḩbner): 1♀ (Israel). Also in NHM 1♁ 1♀ are labelled as having been reared from Colias (Pieridae) larvae on cabbage in Cyprus; but no Colias species feed on cabbage and this is doubtless a misidentification of P. rapae larvae, which resemble those of some Colias species. There are many older records of rearings from Lepidoptera in other families in the literature, but their reliability is highly questionable. We have seen specimens from Germany, Austria, France, Portugal, Spain, Italy, Sardinia, former Yugoslavia, Greece, Cyprus, Israel, Iran, Algeria and India. There are also reports from right across Asia to China. The fact that it does not occur in Britain, despite the abundance of Pieris spp, may be a result of its possible overwintering hosts (Carcharodus and perhaps Muschampia spp) not occurring in Britain (cf. Shaw et al. 2016). One specimen from the third author’s collection was barcoded (BOLD Sample ID CollHH3480)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 42, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Gravenhorst, J. L. C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https: // doi. org / 10.5962 / bhl. title. 65750","Townes, H. K., Momoi, S. & Townes, M. (1965) A catalogue and reclassification of the eastern Palearctic Ichneumonidae. Memoirs of the American Entomological Institute, 5, 1 - 661.","Tschek, C. (1871) Ichneumonologische Fragmente. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 21, 37 - 68.","Hinz, R. (1957) Zur systematik und ˆ kologie der Ichneumoniden 1 (Hym). Deutsche Entomologische Zeitschrift, 4, 86 - 90. https: // doi. org / 10.1002 / mmnd. 19570040112","Horstmann, K. (1978) Typenrevision der von G. Szepigleti beschriebenen palaarktischen Arten der Unterfamilie Campopleginae (Hymenoptera: Ichneumonidae). Folia Entomologica Hungarica, 31, 37 - 45.","Meyer, N. F. (1926) Einige neue Ichneumoniden und Cynipiden. Revue Russe d'Entomologie, 20, 260 - 264.","Horstmann, K. (1969) Typenrevision der europaischen Arten der Gattung Diadegma F ˆ rster (syn. Angitia Holmgren). Beitrage zur Entomologie, 19, 413 - 472.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222."]}

Hyposoter karmensis Galsworthy & Haraldseide sp. nov. (Figs. 18, 19) Diagnosis. Among red-bodied species with mainly dark trochanters, generally similar to H. notatus and H. caudator, but well diffentiated by the key characters, in particular the narrower and darker 2nd abdominal tergite, which more closely resembles that of H. ruficrus, the latter however having dark tegulae. Description (female). Fore wing length: 4.5 mm. Head: face broader than long, quite coarsely granulate; clypeus narrow and rounded, sculpture as face, with a narrow and sharp shining apical margin; mandibles strongly narrowed, with a broad ventral flange; malar space about equal to proximal width of mandibles; teeth equal in size; genal carina joining hypostomal carina at some distance from base of mandible; frons, vertex and temples evenly granulate; temples in dorsal view very short and very strongly narrowed behind eyes, in lateral view about one third width of eyes; antenna slender and slightly tapering, with 28–29 flagellomeres, first flagellomere 2.4x as long as wide, distal flagellomeres quadrate to slightly transverse. Mesosoma: pronotum with strong epomia, granulate with transverse striae; mesoscutum granulate with many fine rugae, and scattered punctures visible under high magnification; mesopleuron including speculum matt with very fine irregularly reticulate sculpture, forming fine striae in hollow in front of speculum, and with many fine punctures confined to lower mesopleuron; epicnemial carina complete laterally and ventrally; metapleuron coarsely granulate; propodeum with anterior fields and area superomedia finely granulate, rest of surface rugose; carinae well defined, and complete except for costulae, which terminate half way across their fields; area basalis narrowly triangular; area superomedia longer than broad, not closed distally. Wings narrow, 1 cu-a postfurcal; areolet stalked; nervellus vertical. Metasoma: petiole smooth and shining, without carinae; postpetiole broadly rounded, with minute reticulate sculpture; small glymma present close to postpetiole; 2nd tergite elongate, 1.4x longer than wide, third tergite quadrate, rest transverse; 2nd tergite onwards apparently smooth, but rather matt with irregular minute microsculpture; 6th and seventh tergites entire; ovipositor short, only slightly curved, with a small dorsal notch slightly beyond half its length, sheaths less than distal height of metasoma. Colour: black; yellow: tips of mandibles, tegulae, wing bases, fore and mid trochantelli; orange: fore and mid femora, tibiae and tarsi, hind trochantellus and femur, distal one third of second tergite, third and fourth tergites, and fifth tergite laterally; hind tibia orange, darkened proximally and distally, in some specimens with a small yellow fleck dorsoproximally; hind tibial spurs white, tipped dark brown; hind tarsi dark brown, basitarsus with only a tiny white fleck proximally. Male and biology unknown. Holotype: ♀, Norway, EIS 13 RY, KARMØY, Skår August 2016 (Malaise) leg. Håkon Haraldseide (in NHMUK). Paratypes: 4♀, data as holotype, one June 2017 (BOLD Sample ID CollHH1208) (in ZSM), two July 2018 (one in NMS, one in coll. H.H. Haraldseide), and one August 2018, (in coll. A C Galsworthy) leg. Håkon Haraldseide; 2 ♀, Germany: Bayern, Allgäu Oberstdorf, Engenkopf, Malaise trap, 1210m and 1230m, 29.05– 10.6.2015 (BOLD Sample ID CollHH3327), and 21.08– 4.9.2015; leg. D. Doczkal & V. Voith (in coll. Haraldseide). Remarks. The barcode data for the German specimen agrees closely with that of the Norwegian specimen.

Hyposoter brischkei (Bridgman, 1882) Limneria brischkei Bridgman, 1882: holotype ♀ in NHMUK, examined. There has been, and perhaps still is, confusion between this species and H. boops (Thomson), which were first synonymized by Aubert (1975), and then again separated in Aubert (1997). Horstmann (2013) dealt with this problem and concluded that there was a constant difference between the two taxa in the sculpture of the propodeum, with the carinae defining the distal part of the area superomedia blurred or absent in brischkei but clearly defined in boops. In the very long series of brischkei in NMS there is a good deal of variation in this feature. In most cases the carinae to the rear of the costulae are faint or absent, and the area petiolaris is evenly and finely granulate. However, the lateral carinae of the distal half of the area superomedia are certainly visible in some specimens. We have found only one specimen marked by Horstmann as ‘boops’ in Horstmann’s collection: this is from the Harz mountains and might be the one mentioned in his 2013 paper: it has a strongly defined area superomedia and a much more rugose area petiolaris, but it also has very dark second and third sternites. Aubert described the area superomedia of the lectotype of H. boops, in translation, as ‘narrow, weakly defined, but with a trace of longitudinal carinae’. We have not ourselves seen the lectotype of H. boops, as Horstmann had, and have therefore left Horstmann’s draft key unchanged in this respect, continuing to treat the two species as distinct. But this is a stance that will certainly need to be revisited. H. brischkei has been reared from a wide variety of noctuid and geometrid larvae, but also from two tortricids (details in Shaw et al. 2016). The holotype was reared from Xestia triangulum (Hufnagel) (Lepidoptera, Noctuidae). In NHMUK there are also 16 specimens reared in Scotland from Thera juniperata (Linnaeus) and 9 labelled as reared in southern England from “ T. variata ”, almost certainly T. obeliscata (Ḩbner) (Lepidoptera, Geometridae). This appears to be one of the commonest species of Hyposoter in the UK, and there are literature records from most countries in central and northern Europe. There is a very long series from Norway in the third author’s collection and we have seen specimens from Germany. We have seen specimens taken in all months from May to October, but most from June to September, with a strong peak in July and August. Eight specimens from the third author’s collection were barcoded (BOLD Sample IDs CollHH1213, 1215, 3471, 3472, 3473, 3474, 3475, and 3487)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on pages 35-36, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Bridgman, J. B. (1882) Further additions to the Rev. T. A. Marshall's catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1882, 141 - 164. https: // doi. org / 10.1111 / j. 1365 - 2311.1882. tb 01574. x","Aubert, J. - F. (1975) Les Ichneumonides petiolees ouest-palearctiques de MORLEY. Bulletin de la Societe Entomologique de Mulhouse, 1975, 13 - 17.","Aubert, J. - F. (1997) 10 eme supplement au catalogue de GAULLE: 129 especes d'Ichneumonides nouvelles pour la faune francaise (Hymenoptera, Ichneumonidae). Nouvelle Revue d'Entomologie, 14, 99 - 114.","Horstmann, K. (2013) Revisionen von Schlupfwespen-Arten XVII (Hymenoptera: Ichneumonidae). Mitteilungen Munchener Entomologischen Gesellschaft, 103, 1 - 14.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222."]}

Hyposoter nigromaculatus (Strobl, 1904) Anilasta nigromaculata Strobl, 1904: lectotype ♀ in Naturhistorisches Museum, Admont, Austria, designated by Horstmann (1995), examined by Horstmann. syn Holocremna nasuta Hedwig, 1932 (Horstmann (1995) There is a full description in Horstmann’s notes. We have seen a single specimen in Horstmann’s collection from 2550m in the Austrian Alps. The species has sculpture not dissimilar to H. carbonarius, but it is much smaller and is easily distinguished by the black colour at the base of the hind femur. Recorded from Austria and Germany; there are no host records., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 57, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Strobl, G. (1904) Ichneumoniden Steiermarks (und der Nachbarlander), V. Fam. Ophionidae. Mitteilungen Naturwissenschaftlichen Vereines fur Steiermark, Graz, 40, 43 - 160. [1903]","Horstmann, K. (1995) Typenrevision einiger Ichneumonidae aus der Sammlung Strobl (Hymenoptera). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 47 (1 / 2), 31 - 35.","Hedwig, K. (1932) Alte und neue Ichneumoniden. (Hym.). Deutsche Entomologische Zeitschrift, 1932, 24 - 35"]}

Hyposoter caudator Horstmann, 2008 Hyposoter caudator Horstmann, 2008: holotype ♀ in NMBE. We have seen both the holotype and a female paratype in NMS. Very similar to H. notatus, which is also a parasitoid of polyommatine Lycaenidae and much more frequently found. The females can be separated by the length of the ovipositor: in the present species the ovipositor sheaths are about the same length as the first tergite, whereas in notatus they are only about half the length. Horstmann in his draft key commented that males could not always be separated from those of H. notatus. In the present species there is usually less darkening of the orange hind tibia, and the coloration of the median outer part is yellowish rather than red, but this is not a very reliable guide. This species has been reared from the larvae of the lycaenid butterflies Agriades pyrenaicus (Boisduval), and Plebicula dorylas (Denis & Schifferm̧ller). Specimens reared from the latter host are in Bern, 3 specimens, including a female paratype, reared from the former host are in NMS and a further female paratype from the same host is in ZSM. We have seen specimens from Switzerland, France and Spain., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 38, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Horstmann, K. (2008) Neue westpalaarktische arten der campopleginae (Hymenoptera: Ichneumonidae). Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen, 60 (1 - 2), 3 - 27."]}

Galsworthy, Anthony, Shaw, Mark R., Haraldseide, Håkon (2023): A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species. Zootaxa 5290 (1): 1-73, DOI: https://doi.org/10.11646/zootaxa.5290.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5290.1.1

Hyposoter praecaedator Aubert, 1963 (Fig. 42) Hyposoter praecaedator Aubert, 1963: lectotype ♀ in MZLS, here designated, labelled “TYPE // HYPOSOTER ♀ ; (=ANILASTUS) Aub; PRAECAEDATOR // J.F.AUBERT; 28.8.1961 VAR; LE PRADET. Syntype 1/6 Hyposoter praecaedator Aubert, labelled by S. Klopfstein 2009 “ The species is based on a series of six syntypes (Klopfstein et al. 2011), which had been examined by Horstmann. We have seen the single syntype designated above as lectotype. This is a small species, very similar to Hyposoter caedator in build, but differs from examples of the latter with pale fore and mid coxae mainly in the more shiny speculum, the extensive reddish coloration of the apex of the hind coxa, the stronger punctation of the lower mesopleuron, and the paler colour of the hind tibia. Hyposoter leucomerus is larger, and has a similarly coloured hind tibia (though the medial colour is usually paler), but always has a dark rear edge to the second tergite, lacking in praecaedator. The biology is unknown, and the species is known only from the south of France., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 59, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Klopfstein, S. & Baur, H. (2011) Catalogue of the type specimens of Ichneumonidae (Hymenoptera) in the Jacques F. Aubert collection at the Musee de Zoologie, Lausanne, Switzerland. Zootaxa, 3081 (1), 1 - 90. https: // doi. org / 10.11646 / zootaxa. 3081.1.1"]}

Anilastus thuringiacus Schmiedeknecht, 1909 This species is absent from Horstmann’s key and there is only a short MS note, which has not been completed, suggesting that he failed to find the holotype. It is not mentioned in Oehlke and Townes’ (1969) review of the type material from Schmiedeknecht’s collection in Rudolstadt, and we have not been able to find any published record of its location. The species was described from a single female from Blankenburg in Germany. There is a series in Horstmann’s collection (2♀, 1♁) which had originally been marked with this species name by Horstmann, but with labels subsequently added that correct the determination to Hyposoter dubitatus Holmgren. They are identical to the holotype of H. dubitatus, which we have examined, but there is no evidence as to whether Horstmann intended to synonymize the two names. In the absence of further information on the holotype of thuringiacus, we regard it as a nomen dubium. It is on the British checklist (Broad, 2016) by virtue of some dubious references in Johnson (1929) and Carr (1924), and we suggest that it should be removed., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 8, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Schmiedeknecht, O. (1909) Opuscula Ichneumonologica. IV. Band. Fasc. XXI - XXIII. Ophioninae. s. n., Blankenburg in Th ¸ ringen. [unknown pagination]","Broad, G. R. (2016) Checklist of British and Irish Hymenoptera - Ichneumonidae. Biodiversity Data Journal, 4, e 9042. https: // doi. org / 10.3897 / BDJ. 4. e 9042","Johnson, W. F. (1929) Irish Ichneumonidae. Entomologist's Monthly Magazine, 65, 134 - 137.","Carr, L. A. (1924) The Ichneumonidae of the Lichfield District, Staffordshire. Transactions of the North Staffordshire Field Club, 58 (Appendix), 1 - 70."]}

Hyposoter nigrior Aubert of Anilasta tibialis Hedwig syn. n. On the type material he wrote (in translation) „ One syntype (♁) of H. nigrior is present in Coll. Aubert (Lausanne) (Klopfstein & Baur 2011: 58). Further syntypes, which should be preserved in Lund, are not traceable there (Hansson, in lit.).“, Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 7, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Klopfstein, S. & Baur, H. (2011) Catalogue of the type specimens of Ichneumonidae (Hymenoptera) in the Jacques F. Aubert collection at the Musee de Zoologie, Lausanne, Switzerland. Zootaxa, 3081 (1), 1 - 90. https: // doi. org / 10.11646 / zootaxa. 3081.1.1"]}

Hyposoter horticola (Gravenhorst, 1829) Campoplex horticola Gravenhorst, 1829: lectotype ♁ in UWCP, designated by Horstmann (2000) and examined by him. The female is easily distinguished from all other Hyposoter by the unique structure of the eighth tergite, whereby the lateral edges are extended backwards to form a covering for the ovipositor and its sheaths. The male has a deeply incised seventh tergite. The species is a parasitoid of various species of melitaeine Nymphalidae (Lepidoptera): Melitaea athalia (Rottemburg), M. cinxia (Linnaeus), and Mellicta aurelia (Nickerl) (Shaw et al. 2016). There is a detailed account of its life history in Finland by Nouhuys & Ehrnsten (2004) and Nouhuys & Kaartinen (2008) as a parasitoid of M. cinxia, which is parasitized just before it leaves its egg by females that have located and monitored the egg batch over time. Oviposition through the host’s eggshell may account for the unusual structure of the female’s 8th tergite. We have seen specimens from Finland, Poland, Germany, France and Spain. The species has also been reported from Sweden, Hungary and Romania. Two specimens from the third author’s collection were barcoded (BOLD Sample IDs CollHH3481 and 3482)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 48, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Gravenhorst, J. L. C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https: // doi. org / 10.5962 / bhl. title. 65750","Horstmann, K. (2000) Typenrevisionen der von Gravenhorst beschriebenen oder gedeuteten Campoplex-Arten (Hymenoptera, Ichneumonidae). Linzer biologische Beitrage, 32 (2), 1203 - 14.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222.","Nouhuys, S. van & Ehrnsten, J. (2004) Wasp behaviour leads to uniform parasitism of a host available only a few hours per year. Behavioural Ecology, 15, 661 - 665. https: // doi. org / 10.1093 / beheco / arh 059","Nouhuys, S. van & Kaartinen, R. (2008) A parasitoid wasp uses landmarks while monitoring potential resources. Proceedings of the Royal Society (B), 275, 377 - 385. https: // doi. org / 10.1098 / rspb. 2007.1446"]}

Hyposoter naso Galsworthy & Shaw sp. nov. (Figs. 23–25) Diagnosis. Superficially similar to some paler specimens of H. caedator, though distinguished from them by the dark trochanters and trochantelli on all legs. The unusual shape of the clypeus, which is unique in Hyposoter, is the most distinctive feature. From some angles this can appear to be similar to the clypeal tooth of Campoletis, but seen in profile it is clearly a nose-like bulge, not a tooth. Description. Female. Fore wing: 5 mm. Head: face broader than long, clypeus narrow, the central part strongly and narrowly prominent subdistally, surface of both rugosely reticulate; malar space slightly broader than width of mandibles at base, the latter strongly narrowed at about two thirds; frons and vertex strongly granulate, temples lightly so, the latter strongly narrowed behind the eyes, making the temples very short in dorsal view; antenna slender, slightly tapered in apical half. 31–32 flagellomeres. Mesosoma: mesoscutum closely and rugosely punctate throughout; speculum smooth and shining; area above and in front of speculum with strong transverse striation; front and ventral area of mesopleuron, and metapleuron, with close strong punctation; all areas of propodeum rugose, petiolar area with strong transverse rugosity; propodeal carinae complete except for hind carina of area superomedia; basal area narrowly trapezoid; area superomedia about as broad as long. Wings narrow; 1 cu-a very slightly postfurcal; areolet small and squarish, long-stalked; nervellus slightly curved. Metasoma: basal part of first tergite parallel-sided, smooth and shining, sharply narrowing before postpetiole; latter a broad oval, with a linear depression in the basal half, lightly granulate; glymma a very shallow linear impression at base of postpetiole. 2nd tergite longer than broad, widening to apex; third tergite quadrate, the rest transverse; 2nd tergite with light granulation, the rest almost smooth, but dull with very fine microsculpture; sixth tergite entire, seventh with a semicircular emargination at apex; ovipositor shorter than height of metasoma. Colour: black; tegulae yellow; fore and mid legs orange except for black trochanters and trochantelli; hind femur orange; hind tibia with a pale yellow spot dorsally at the base, rather uniform brownish dorsally, orange ventrally, with faint darkening subproximally and distally; hind tarsi brown dorsally, orange ventrally; tergite 1 black; tergite 2 black in basal half, orange distally, apart from a narrow black band at extreme apex, which extends to base of tergite 3; rest of tergite 3 orange, as are tergites 4–6; tergite 7 brown; sternites 2–3 yellow with broad pale brown bands. Male. Very similar to female, but sides of postpetiole orange. Holotype: ♀, Spain: Madrid, 600m, Ciempozuelos, G.E.King, ex Perigune convergata (de Villers) (Lepidoptera, Geometridae), Thymus, 16.4.2005, coc 23.4.2005, em 2.5.2005 (NMS). Paratypes: 1♀, Spain, Madrid, 600m, Ciempozuelos, G.E.King, ex Perigune narbonea (Linnaeus) (Lepidoptera, Geometridae), Teucrium, 28.11.2004, coc 3.12.04, em 31.12.04; 1♀, Ciempozuelos, ex Petrophora convergata, (de Villers) (Lepidoptera, Geometridae), Thymus, 8.4.2004, em by 20.5.2004, G.E.King ; 1♀, Spain, Madrid, 600m, Ciempozuelos, G.E.King, Perigune convergata (de Villers), Thymus, 8.4.2004, coc 27.4.04, em 16/ 20.5.04 ; 1♀, Spain, Madrid, 600m, Ciempozuelos, “ Scopula ” Glossotrophia asellaria Herrich-Schäffer (Lepidoptera, Geometridae), Antirrhinum, 13.5.05 [sic: presumably 13.3.05], pupa 3.4.05, em 1.5.05 ; 1♀, Spain: Madrid, 600m, Ciempozuelos,? Chemerina caliginearia (Rambur) (Lepidoptera, Geometridae), 4.5.08, em 21.5.08, G.E.King ; 1♁, Spain, Madrid, 600m, Ciempozuelos, G.E.King, ex Antilurga alhambrata (Staudinger) on Helianthemum hirtum, 6.3.2005 em. 29.3.2005 ; 1♁, Spain, Madrid, 600m, Ciempozuelos, G.E.King, uncertain host, coc. 12.4.08, em 18.4.2008 ; 1♁, Spain, Madrid, 600m, Ciempozuelos, G.E.King, ex Petrophora convergata (de Villers) (all in NMS). All the above are accompanied by cocoons and host remains, except for the single female reared from the sterrhine G. asellaria which is mounted with an intact lepidopteran pupa. This raises a doubt as to the identity of the actual host in that case. The rearing data make clear that H. naso is a plurivoltine species. Remarks. First identified by Horstmann as an undescribed species.

Hyposoter kontzeii (Kiss, 1924) Omorgus kontzeii Kiss, 1924: holotype ♀ in HNHM, examined by Horstmann. We have not seen this species. Horstmann (2009) transferred it to Hyposoter and included it in his draft key. So far as we know, the species is known only from the holotype. Horstmann’s notes indicate that the label of the holotype states that it was obtained ‘ex larva’, but there is no indication of the kind of larva involved, nor was there any mention of this in the original description. The holotype was collected in Romania., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 50, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Kiss, von Zilah A. (1924) Beitrag zur Kenntnis der ungarischen und siebenb ¸ rgischen Ichneumoniden- (Schlupfwespen-) Fauna. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften in Hermannstadt, 72 / 74, 32 - 146.","Horstmann, K. (2009) Typenrevisionen der von Kiss beschriebenen Taxa der Ichneumonidae III. Verschiedene Unterfamilien (Hymenoptera, Ichneumonidae). Linzer Biologische Beitrage, 41 (1), 673 - 689."]}

Hyposoter jubator Galsworthy & Shaw sp. nov. (Figs. 14–17) Diagnosis. One of only two known European Hyposoter species with an open areolet in the fore wing. Easily distinguished from the other, Hyposoter fitchii, by the dark brown hind femur and the pale proximal third of the hind tibia contrasting with the dark distal third. In H. fitchii the hind femur and hind tibia are both bright orange. Description. Female. Fore wing length 4.5 mm. Head. Temples, frons, face, clypeus and genae uniformly finely granulate, with silvery adpressed hairs which are longest on lower part of face, clypeus and genae; clypeus flat, not separated from face, its apex evenly curved and sharp; malar space approximately equal to basal width of mandible; mandibles evenly tapered, with narrow ventral flange basally to one half; temples short, very strongly narrowed behind eyes; antenna with 31 flagellomeres, strongly tapered distally. Mesosoma. Mesoscutum evenly granulate, slightly more coarsely so than head; scutellum more finely granulate; floor of scuto-scutellar groove polished, shining and punctate; mesopleuron sculptured as mesoscutum, apart from some small vertical striations at posterodorsal corner and a small almost smooth area in the speculum, and punctate in lower third; metapleuron coarsely granulate; propodeum finely granulate, with transverse striations in area petiolaris; area basalis triangular; area superomedia pentagonal, approximately as broad as long, the front two edges formed of strong carinae which extend to form the costulae, the side two edges of weaker carinae, and the posterior side continuous with the area petiolaris; area petiolaris diamond-shaped with strong carinae. Wings, Fore wing 1 cu-a postfurcal. Areolet open. Metasoma. Petiole of first tergite straight-sided, with strong dorsolateral carinae, its surface smooth; postpetiole about as broad as long, slightly trapezoid, with granulate sculpture; a small glymma, located just anterior to postpetiole; tergite 2 markedly elongate, tergite 3 slightly so, both finely granulate; rest of tergites transverse, more or less smooth with fine punctures; sixth and seventh tergites entire; ovipositor sheaths very slightly protruding beyond abdomen; ovipositor not visible in type material. Colour. Black. Yellow: apex of underside of antennal scape (in holotype only), mandibles other than teeth, tegulae, fore coxa and trochanters, distal half of mid coxa and whole of mid trochanters, second and third sternites. Brown: antenna, underside of fore femur, hind trochanters and femur, distal third of hind tibia, mid and hind tarsi. Dull orange: rest of fore and mid legs, proximal two thirds of hind tibia. Holotype. ♀, England SS, [Somerset] Withypool, 1948, F.H.Lyon, B.M.1948-355, Ex C. glabraria Schiff., Anilastus (Eriborus) sp., [in Horstmann’s writing] Hyposoter sp. nov. ♀ (NHMUK). Paratype 1♀, same data as holotype, with addition of label reading “No 6” [head mounted separately on card] (NHMUK). Biology. The cocoons from which the two type specimens were reared are mounted on a card on a separate pin. Both these and the specimens bear labels reading “Ex C. glabraria Schiff ”. This is presumably Cleora glabraria (Ḩbner), which is now regarded as a junior synonym of Alcis jubata (Thunberg). The larval remains attached to the cocoons are consistent with this, showing the double row of dark spots along the dorsal side which are characteristic of the larva of this species. The cocoons are fairly typical Hyposoter cocoons, whitish with discrete dark patches. Remarks. A further female in NMS, reared from Chloroclysta truncata (Hufnagel) in Aberdeenshire, Scotland, is very similar, and probably belongs to this species. It differs slightly in having shorter antennae (29 flagellomeres), stronger carination on the propodeum, and a shorter ovipositor, and is therefore not included in the type series. Although he had labelled these specimens some years earlier as a new species, and they are recorded in his notes, Horstmann did not include them in his draft key. We do not know why, but it may have been simply an oversight., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 49, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256

Hyposoter seniculus (Gravenhorst, 1829) Campoplex seniculus Gravenhorst, 1829: holotype ♀ in UWCP, examined by Horstmann syn. Anilastus melaleucus Schmiedeknecht,1909: Hedwig, 1940 syn. Anilasta berberatae Habermehl, 1922 syn. n.: synonymy proposed in Horstmann’s notes after examination of type material. Horstmann left full notes on the holotype. According to these, the central section of the hind tibia is bright yellow, with the base and apex abruptly dark. The species most likely to be confused with it would be H. albonotatus (with which Horstmann had earlier confused it), but the latter has the fore and mid coxae yellow, whereas they are dark in seniculus. The holotype is labelled ‘ Silesia’, but it is not clear whether the locality is in modern Germany or Poland. There are records in the literature of uncertain reliability from Germany, Austria, Romania, the Netherlands and France., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 67, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Gravenhorst, J. L. C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https: // doi. org / 10.5962 / bhl. title. 65750","Schmiedeknecht, O. (1909) Opuscula Ichneumonologica. IV. Band. Fasc. XXI - XXIII. Ophioninae. s. n., Blankenburg in Th ¸ ringen. [unknown pagination]","Hedwig, K. (1940) Alte und neue Ichneumoniden. Deutsche Entomologische Zeitschrift, 1940, 190 - 203.","Habermehl, H. (1922) Neue und wenig bekannte palaarktische Ichneumoniden (Hym.) Deutsche Entomologische Zeitschrift, 1922, 348 - 359."]}

Hyposoter rapacitor Aubert, 1971 Hyposoter rapacitor Aubert, 1971: lectotype ♀, here designated (in MZLS). Label data: TYPE ♀ HYPOSOTER RAPACITOR Aubert (Anilastus rapx Auctt); J F AUBERT 26.7.1960 LECLERQ LEG. PONT MOUSSON; syntype 1/10 (5) Hyposoter rapacitor Aubert 1971 labelled by S. Klopfstein 2009. GBIFCH 00832765. Based on ten syntypes (Klopfstein et al. 2011), some of which Horstmann had examined. We have examined the syntype designated as lectotype above. There are three specimens in Horstmann’s collection. The species is most similar in build to H. rufonigrator sp. nov., but has a longer ovipositor (slightly longer than first tergite in rapacitor, slightly shorter in rufonigrator). In addition rapacitor has more orange coloration on the central tergites: in rufonigrator the orange coloration is reduced to a few patches, and is often absent altogether. Brighter specimens of rapacitor resemble H. caudator superficially, but can be distinguished by the weaker carination on the propodeum. The biology is unknown. The type series is from France, Spain and Switzerland (Klopfstein et al. 2011). Horstmann’s specimens are from Germany., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 62, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Aubert, J. - F. (1971) Supplement aux Ichneumonides petiolees avec neuf especes nouvelles. Bulletin de la Societe Entomologique de Mulhouse, 1971, 35 - 43.","Klopfstein, S. & Baur, H. (2011) Catalogue of the type specimens of Ichneumonidae (Hymenoptera) in the Jacques F. Aubert collection at the Musee de Zoologie, Lausanne, Switzerland. Zootaxa, 3081 (1), 1 - 90. https: // doi. org / 10.11646 / zootaxa. 3081.1.1"]}

Hyposoter sanguinator Aubert, 1960 Hyposoter sanguinator Aubert, 1960: holotype ♀ in MZLS, examined. The combination of red hind coxae and entirely red metasoma, including the whole of the first tergite, make the species easy to identify. Nothing is known of the biology. The species is known only from the south of France and from Corsica., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 67, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Aubert, J. - F. (1960) Description preliminaire de quelques espѐces et sous-espѐces mediteraneennes de la famille des Ichneumonides. Bulletin de la Societe Entomologique de Mulhouse, 1960 (Septembre - Octobre), 62 - 65. https: // doi. org / 10.3406 / bsef. 1960.20555"]}

Anilasta berberatae Habermehl of Campoplex seniculus Gravenhorst syn. n. The type material of berberatae is recorded as “ Holotype (♀) of A. berberatae: „Frankfurt, „Saalm., „from Cid[aria]. berberata a. 20.4.85 [= Pareulype berberata (Denis & Schifferm̧ller) (Geometridae)] (Frankfurt)”., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 7, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256

Hyposoter fitchii (Bridgman, 1881) Limneria fitchii Bridgman, 1881: lectotype ♀ in NCMK, designated by Horstmann (1972), and examined by him. We have examined 2 paralectotypes in NHMUK designated by Horstmann. The species is one of only two among European Hyposoter that lack vein 3 rs-m, and thus an areolet, which makes its identification straightforward. The colours of the hind femur and tibia distinguish it from the other, H. jubator sp. nov. with dark brown hind femur. It appears to be rare in collections. Bridgman’s four original specimens were reared in the UK by G.C.Bignell. Horstmann’s collection contains three specimens, two from the Frisian Islands (Germany) and one from northern Spain. There are also records from Finland and France. The type series was reared in England from Meganola albula (Denis & Schifferm̧ller) (Lepidoptera, Nolidae). It has also been reared more recently from Meganola togatulalis (Ḩbner) in France (Shaw et al. 2016)., Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on page 43, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Bridgman, J. B. (1881) Some additions to Mr. Marshall's catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1881, 143 - 168. https: // doi. org / 10.1111 / j. 1365 - 2311.1873. tb 01517. x","Horstmann, K. (1972) Type revision of the species of Cryptinae and Campopleginae described by J. B. Bridgman (Hymenoptera: Ichneumonidae). Entomologist, 105, 217 - 228.","Shaw, M. R., Horstmann, K. & Whiffin, A. L. (2016) Two hundred and twenty-five species of reared western Palaearctic Campopleginae (Hymenoptera: Ichneumonidae) in the National Museums of Scotland, with descriptions of new species of Campoplex and Diadegma, and records of fifty-five species new to Britain. Entomologist's Gazette, 67, 177 - 222."]}

Key to European species of Hyposoter (males and females) 1. Metasoma black, sometimes with the posterior borders of the tergites narrowly lightened or the central tergites weakly washed with brown..............................................................................................2 - Metasoma marked with red-brown, at least the third and/or fourth tergites laterally marked with red-brown (a few intermediate species can be reached via either alternative).................................................................43 2. Tegula brown or blackish, sometimes paler on the edge........................................................... 3 - Tegula completely yellowish...............................................................................8 3. Glymma lacking, third metasomal tergite washed with red-brown laterally.................... calcaneus (Strobl, 1904) ♀ - Glymma obvious and deep; metasoma various.................................................................4 4. Hind tibia almost completely blackish, somewhat washed with dark brown, with a longitudinal red-brown stripe ventrally (a)..................................................................................... tibialis (Hedwig, 1938) ♁ ♀ - Hind tibia red-brown, sometimes a little darkened at base and apex (aa)............................................. 5 5. Hind femora strongly marked with black proximoventrally (a); second metasomal sternite yellow-brown, without dark bands; speculum dorsally strongly and thickly punctured and rugose, smooth in a small area ventrally (rest of mesopleuron, mesoscutum and scutellum strongly, closely and evenly punctate)........................ nigromaculatus (Strobl, 1904) (part) ♀ - Hind femora red-brown (aa), at most a little darkened proximally; second metasomal sternite marked with brown or black; speculum finely granulate or predominantly smooth.............................................................6 6. Large robust species (fore wing 7–8 mm); palps usually dark brown or blackish (sometimes yellow, especially in males) (a); hind claws clearly longer than arolium; area superomedia broader than long (b); lower half of mesopleuron strongly punctate (c)...................................................................... carbonarius (Ratzeburg, 1844) ♁ ♀ - Smaller, more delicate species (fore wing length 4–5mm); palps yellowish or yellow-brown (aa); hind claws as long as arolium; area superomedia slightly longer than broad (bb); lower half of mesopleuron without visible punctures (cc)............... 7 7. Hind tibia proximally narrowly marked with brown; speculum relatively smooth; hind basitarsus almost completely dark brown; metasoma shorter, tergite 3 quadrate to transverse; propodeum with areas clearly defined with strong carinae, area petiolaris clearly bordered laterally.................................................... culminator Aubert, 1974 ♁ ♀ - Hind tibia proximally yellow-red; speculum strongly granulate; hind basitarsus almost completely yellow-red; metasoma longer, tergite 3 clearly elongate; propodeum with more weakly defined areas, area petiolaris not clearly bordered laterally....................................................................................... nigritus (Holmgren, 1860) ♀ 8. Hind femur dark brown or black, sometimes red-brown on the inner or outer side (a)..................................9 - Hind femur bright reddish, sometimes a little darkened proximally or distally (aa)................................... 15 9. Clypeus and genae yellow (a)................................................ pallidirostris (Schmiedeknecht, 1909) ♀ - Clypeus and genae black.................................................................................... 10 10. Second metasomal sternite white-yellow, without dark bands (a)................................................... 11 - Second metasomal sternite banded with brown or black, or completely dark (aa)..................................... 13 11. Fore wing areolet open (a)....................................................... jubator sp.n. Galsworthy & Shaw ♀ - Fore wing areolet closed (aa)..................................................................................... 12 12. Scape ventrally yellow (a); fore and mid coxae bright yellow (b).................. inquinatus (Holmgren, 1860) (part) ♀ ♁ - Scape ventrally black (aa); fore and mid coxae predominantly or completely black (bb)..... neglectus (Holmgren, 1860) ♀ ♁ 13. Hind tibia clearly marked with whitish yellow proximally, otherwise brownish with a darkened apex (a) (Area superomedia more than twice as broad as long, rather kidney-shaped, posteriorly closed (b); speculum granulate, matt)........................................................................................... tricoloripes (Viereck, 1911) ♀ ♁ - Hind tibia proximally and distally black, medially whitish to yellowish red, with strong contrast (aa).................... 14 14. Hind basitarsus pale orange (a)........................................................ tricolor (Ratzeburg,1844) ♁ ♀ - Hind basitarsus white with distal quarter black (aa)............................... albonotatus (Bridgman, 1889) (part) ♀ 15. Hind coxa yellowish red or bright red-brown, sometimes marked with a little light brown at base (a).................... 16 - Hind coxa black, sometimes marked with a little white or yellow at apex (aa)....................................... 17 16. Hind tibia proximally and distally broadly blackish, the proximal ring at least twice as long as the breadth of the tibia, medially pale to reddish yellow; hind tarsus blackish, the basitarsus only narrowly lightened proximally (a); metasoma completely black........................................................................ coxator (Thomson, 1887) ♀ ♁ - Hind tibia predominantly whitish yellow, with a small brown spot subproximally, and narrowly blackish at apex; first to third segments of the hind tarsus predominantly whitish yellow, each only spotted with brown at the apex (aa); third metasomal tergite narrowly washed with reddish brown............................... ruficoxator sp. n. Galsworthy & Shaw ♀ ♁ 17. Fore and mid coxae whitish yellow, sometimes tinged with brownish or with brown areas, but not black................ 18 - Fore and mid coxae entirely or largely black, mid coxa at least broadly black at base................................. 25 18. Areolet open.......................................................................... fitchii (Bridgman,1881) ♀ - Areolet obviously closed by vein 3 rs-m................................................................................19 19. Hind tibia broadly black proximally and distally, medially white or pale yellow to orange, with more or less strong contrast (a)................................................................................................ 20 - Hind tibia predominantly yellowish red, in the female hardly marked with darker (aa), in the male proximally and distally blackish (area superomedia about as long as broad, but carinae sometimes reduced).................................... 22 20. Hind tibia medially bright white (a)....................................................................................21 - Hind tibia medially pale yellow to orange (aa)................................... inquinatus (Holmgren, 1860) (part) ♀ ♁ 21. Ovipositor sheaths projecting well beyond apex of metasoma (a); fore and especially mid coxae with large brown patches alternating with clear yellow (b); hind basitarsus dorsally orange brown (c).......... albosignatus sp. n. Galsworthy & Shaw ♀ - Ovipositor sheaths shorter than apical height of metasoma (aa); fore and mid coxae usually uniform pale yellow (bb); hind basitarsus white with short black apex (cc).................................. albonotatus (Bridgman, 1889) (part) ♀ ♁ 22. Area petiolaris in female clearly rugose (a)...................................................................... 23 - Area petiolaris in female granulate and with a few very fine wrinkles (aa)........................................... 24 23. Mesopleuron granulate without obvious punctures (a); seventh tergite entire (b)... monensis sp. n. Galsworthy & Shaw ♀ ♁ - Mesopleuron with obvious punctures visible over the granulation (aa); seventh tergite emarginate (bb)........................................................................................ pechipogator sp. n. Galsworthy & Shaw ♀ 24. Area basalis as long as broad, trapezoid; second metasomal sternite yellowish with brown transverse band................................................................................................. dumeticola (Holmgren, 1860) ♀ - Area basalis narrowly triangular, posteriorly forming a short keel (a); second metasomal sternite yellowish, with no brown marking (b)........................................................... rivulator sp. n. Galsworthy & Shaw ♀ ♁ 25. Second metasomal sternite whitish yellow or bright yellow brown (a)............................................... 26 - Second metasomal sternite mid-brown to black, or banded with this colour (aa) (H. vividus is intermediate and can be reached via either alternative).....................................................................................30 26. Hind tibia proximally and medially whitish, subproximally and distally blackish, with strong contrast (a); (second metasomal sternite distally sometimes washed with red-brown; lower mesopleuron with clear punctures; fore and mid coxae often with some yellow, but mid coxa with black at least at base)................................. vividus (Holmgren, 1860) ♀ ♁ - Hind tibia medially yellow or yellowish red, proximally and distally only narrowly or not marked with brown (aa)........27 27. Speculum very finely granulate, strongly shining, smooth in places (a).............................................. 28 - Speculum completely granulate, hardly shining (aa)............................................................. 29 28. Area in front of speculum with transverse rugae (a); propodeal carinae complete and petiolar area rugose (b).......................................................................................... castaneus sp. n. Galsworthy & Shaw ♀ - Area in front of speculum granulate, without rugae (aa); propodeal carinae reduced and propodeum entirely granulate, including petiolar area (bb)................................................. flaviventer sp. n. Galsworthy & Shaw ♀ ♁ 29. Propodeum with finely delimited areas, sides of area superomedia and/or area basalis blurred with fine wrinkles......................................................................................... brischkei (Bridgman, 1882) ♀ ♁ - Propodeum with clear areas, sides of area superomedia clearly developed...................... boops (Thomson, 1887) ♀ 30. Trochanters and trochantelli black, only fore trochantellus distally reddish-brown (a); speculum almost smooth, strongly shining; centre of mesopleuron with regular punctures on a finely sculptured and rather shiny background (b) (ovipositor sheaths about as long as the petiole, ovipositor clearly upcurved; third metasomal tergite often marked with red-brown laterally)..................................................................... rufonigrator sp. n. Galsworthy & Shaw ♀ ♁ - Trochanters and trochantelli extensively marked with yellow or bright reddish-brown (aa); centre of mesopleuron frequently unpunctured or finely punctured on a granulate background and matt (bb)........................................ 31 31. Hind tibia proximally and distally black or clearly infuscate, medially whitish yellow to red-brown with more or less strong contrast, and without pale spot at base (a)................................................................. 32 - Hind tibia with pale spot at base and/or lacking strong darkening subproximally and distally (aa)....................... 34 32. Speculum fully granulate, with some reflection; hind tibia dorsomedially whitish yellow, proximally and distally broadly blackish; hind claws long pectinate, with pectinations almost as long as length of claw beyond pectinations............................................................................................. seniculus (Gravenhorst, 1829) ♀ - Speculum smooth or almost smooth in a small area, strongly shining; hind tibia dorsomedially yellowish-red; hind claws very short pectinate, pectinations much shorter than length of claw beyond pectinations................................ 33 33. Area superomedia generally 1.2–1.4 as long as broad (a); hind tibia proximally brownish, distally brownish or blackish, spurs pale (b)................................................................ caedator corsicator Aubert, 1960 ♀ ♁ - Area superomedia a little broader than long (aa); hind tibia proximally and distally blackish, spurs dark (bb)........................................................................................ anglicanus (Habermehl, 1923) ♀ 34. Hind tibia proximally marked with whitish or yellowish (this spot sometimes unclear), subproximally and distally brownish or blackish, dorsomedially whitish to yellowish red (a).............................................................. 35 - Hind tibia bright red-brown, sometimes with a proximal yellow spot or narrow apical darkening, but not with a dark subproximal ring (aa).............................................................................................. 38 35. Area superomedia clearly broader than long (a); seventh metasomal tergite dorsally deeply triangularly emarginate (b); ovipositor clearly protruding beyond the apex of the metasoma, ovipositor sheaths longer than the petiole (c) (metasoma black or slightly washed with red)................................................. virginalis (Gravenhorst, 1829) (part) ♀ - Area superomedia at least as long as broad (aa); seventh metasomal tergite dorsally not emarginate (bb); ovipositor protruding little beyond the apex of the metasoma, ovipositor sheaths shorter than the petiole (cc)............................... 36 36. Hind tibia yellow proximally (sometimes not clear), subproximally bright brown to mid-brown, dorsomedially pale to darker yellowish or yellowish red, dorsodistally black (a).................................... longulus (Thomson, 1887) ♀ ♁ - Hind tibia strongly whitish yellow proximally and medially, subproximally and dorsodistally strongly black, with strong and abrupt contrast (aa)........................................................................................ 37 37. Fore coxa black (a); metasomal tergites usually at least marked with red-brown laterally; ventral edge of hind tibia usually orange (b) (in specimens with extended red-brown markings on metasoma the dark rings of the hind tibia are brown instead of black; a species which is very variable in colouring)............................. didymator (Thunberg,1822) (part) ♀ ♁ - Fore coxa broadly whitish yellow at apex (aa), and sometimes mid coxa with some yellow at apex; central metasomal tergites black, sometimes a little washed with red-brown; hind tibiae always with very strong contrast, and with ventral edge narrowly dark brown (bb).................................................................... vividus (Holmgren,1860) ♁ ♀ 38. All or almost all of centre of mesopleuron punctured on a smooth background (a) (seventh metasomal tergite dorsally clearly emarginate; ovipositor clearly extending beyond the apex of the metasoma)................... ebenitor Aubert, 1972 ♀ ♁ - Speculum frequently granulate; centre of mesopleuron with granulate background (aa)............................... 39 39. Genal carina disappearing subventrally (at level of bottom of eye) and replaced by a bulge covered with long grey setae, the latter turning forwards and meeting the oral carina very close to the base of the mandible (a); in the male, and sometimes also the female, scape ventrally broadly marked with yellow (b); in the female ovipositor short, the sheaths usually about half the apical height of the metasoma (c); seventh tergite dorsally not emarginate (in both sexes lower mesopleuron finely granulate with scattered punctation)........................................................... clausus (Brischke, 1880) ♀ ♁ - Genal carina subventrally low but distinct, not turning forwards, and meeting the oral carina at a distance from the base of the mandible (aa); in male scape ventrally at most washed with a little red brown (bb); in female ovipositor longer (cc) and seventh tergite dorsally in some cases clearly emarginate........................................................... 40 40. Speculum smooth in a small area, clearly shining; area superomedia about 1.1x as long as broad (hind tibia bright red-brown, without yellow proximal fleck, with a brown stripe dorsodistally)............................ insulator Aubert, 1960 ♀ - Speculum granulate, or with granules in rows, matt or with silky reflection; area superomedia generally at most as long as broad.............................................................................................. 41 41. Hind tibia with a yellow spot proximally (this sometimes disappearing) (a); in male fore coxa distally often about half yellow; in female ovipositor sheaths about as long as the postpetiole, ovipositor reaching a little beyond apex of metasoma (b); seventh metasomal tergite dorsally not clearly emarginate.................................... placidus (Desvignes, 1856) ♀ ♁ - Hind tibia without yellow spot proximally (aa); in male fore coxa at most narrowly yellowish distally; in female ovipositor sheaths as long as petiole, ovipositor reaching well beyond apex of metasoma (bb); seventh metasomal tergite clearly emarginate dorsally......................................................................................... 42 42. Base and middle of mandible yellow (a); hind tibia not or hardly darkened distally (b).... ebeninus (Gravenhorst, 1829) ♀ ♁ - Mandible clearly marked with black proximally, yellow medially (aa); hind tibia narrowly marked with dark brown distally (bb)........................................................................... rhodocerae (Rondani, 1877) ♀ ♁ 43. Hind coxa bright red-brown (a)................................................................................ 44 - Hind coxa mostly or completely black (aa)................................................................, Published as part of Galsworthy, Anthony, Shaw, Mark R. & Haraldseide, Håkon, 2023, A key to European species of Hyposoter Förster, 1869 (Ichneumonidae: Campopleginae) with descriptions of 18 new species, and notes on all included species, pp. 1-73 in Zootaxa 5290 (1) on pages 8-31, DOI: 10.11646/zootaxa.5290.1.1, http://zenodo.org/record/7959256, {"references":["Strobl, G. 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[unknown pagination]","Viereck, H. L. (1911) Descriptions of one new genus and eight new species of Ichneumon flies. Proceedings of the United States National Museum, 40 (1832), 475 - 480. https: // doi. org / 10.5479 / si. 00963801.1832.475","Bridgman, J. B. (1889) Further additions to the Rev. T. A. Marshall's catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1889, 409 - 439. https: // doi. org / 10.1111 / j. 1365 - 2311.1889. tb 02334. x","Thomson, C. G. (1887) XXXV. F ˆ rs ˆ k till uppstallning och beskrifning af aterna inom slagtet Campoplex (Grav.) Opuscula Entomologica, Lund, XI, 1043 - 1182.","Bridgman, J. B. (1881) Some additions to Mr. Marshall's catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1881, 143 - 168. https: // doi. org / 10.1111 / j. 1365 - 2311.1873. tb 01517. x","Bridgman, J. B. (1882) Further additions to the Rev. T. A. Marshall's catalogue of British Ichneumonidae. Transactions of the Entomological Society of London, 1882, 141 - 164. https: // doi. org / 10.1111 / j. 1365 - 2311.1882. tb 01574. x","Gravenhorst, J. L. C. (1829) Ichneumonologia Europaea. Pars III. Sumtibus auctoris, Vratislaviae, 1097 pp. https: // doi. org / 10.5962 / bhl. title. 65750","Aubert, J. - F. (1960) Description preliminaire de quelques espѐces et sous-espѐces mediteraneennes de la famille des Ichneumonides. Bulletin de la Societe Entomologique de Mulhouse, 1960 (Septembre - Octobre), 62 - 65. https: // doi. org / 10.3406 / bsef. 1960.20555","Habermehl, H. (1923) Neue englische Ichneumoniden nebst Bemerkungen ¸ ber Ichneumon macrocerus C. G. Thoms. ♀, Cratichneumon foersteri Wesm. ♀, und Grypocntrus cinctellus Ruthe ♁ (Hym.). Konowia, 2, 34 - 43.","Aubert, J. - F. (1972) Etude commentee de nouveaux lectotypes choisis dans les collections HOLMGREN et THOMSON (Hym. Ichneumonidae). 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(1971) Supplement aux Ichneumonides petiolees avec neuf especes nouvelles. Bulletin de la Societe Entomologique de Mulhouse, 1971, 35 - 43.","Aubert, J. - F. (1965) Les Ichneumonides du rivage mediterraneen francais (8 e serie). Vie et Milieu, 16, 549 - 573.","Kiss, von Zilah A. (1924) Beitrag zur Kenntnis der ungarischen und siebenb ¸ rgischen Ichneumoniden- (Schlupfwespen-) Fauna. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften in Hermannstadt, 72 / 74, 32 - 146."]}