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1. The bilobatus valve pattern group The workers within this pattern have basic enteric valves: all six primary cushions (PCs) are similar in their arrangement (Fig. 12A). In the upstream and middle spiny parts, they are armed with relatively strong spines, becoming gradually thinner downstream; moreover, the middle part bears some lateral supporting spindle-like bristles leaning on the funnel membrane. In the downstream bristly part, the spines are rather abruptly replaced with longer and bristle-like setae, first straight, then curved and eventually hooked. The PCs are roughly triangular: their largest width is generally located near their upstream end and their lateral margins converge gradually downstream until the bristly part where they remain parallel (Fig. 11A). The enteric valve shows hexaradial to triradial symmetry, the odd PCs often being somewhat longer than the even PCs. In some cases, PC1 is still longer than PC3 and PC5, tending towards bilateral symmetry. Workers (counts) Code Range (Nb) Fore coxa: number of spines on carina WCx1CN 0–20 Fore coxa: number of spines on fore side WCx1SN 0–4 Valve: number of bristles on PC1 downstream part WVP1DBN 15–160 Valve: number of lateral supporting bristles on PC1 (one side) WVP1SBN 3–37 Workers (raw data) Code Range (mm) Head: length to fore edge of postclypeus WHdL 0.67–1.37 Head: width WHdW 0.84–1.55 Mandible (left): length of apical tooth inner side WMlAL 0.08–0.28 Mandible (left): apical to first marginal WMlAmD 0.09–0.27 Mandible (left): apical tooth width at its base WMlAW 0.07–0.20 Mandible (left): functional length WMlL 0.35–0.75 Mandible (left): length of the molar tooth WMlML 0.13–0.25 Mandible (left): first to third marginal WMlmmD 0.08–0.17 Mandible (left): third marginal to molar WMlmMD 0.05–0.12 Mandible (left): width of the first marginal tooth WMlmW 0.03–0.13 Mandible (left): width WMlW 0.26–0.56 Mandible (right): apical to first marginal distance WMrAmD 0.09–0.26 Mandible (right): first to second marginal distance WMrmmD 0.05–0.12 Valve: length of downstream bristly part of PC1 1 WVP1DL 0.09–0.39 Valve: length of PC1 1 WVP1L 0.45–1.04 Valve: length of middle spiny part of PC1 1 WVP1ML 0.04–0.53 Valve: length of upstream spiny part of PC1 1 WVP1UL 0.07–0.57 Valve: width of PC1 2 WVP1W 0.04–0.15 Valve: average length of PCs2 WVP-AvL 0.43–0.85 Valve: average width of PCs2 WVP-AvW 0.04–0.12 Postclypeus: length WPcL 0.24–0.47 Postclypeus: width WPcW 0.52–0.96 Tibia (fore): length WT1L 0.60–1.18 Tibia (fore): maximal width WT1W 0.11–0.25 Tibia (hind): length WT3L 0.84–1.66 1 Excluding the members of the sulcifrons pattern which bear a spatula on PC1 2 Including all PCs of all species (with and without spatulas) An odd primary cushion (Fig. 11A) is made of (a) a long, upstream, spiny part (38–58% of total length) with relatively strong spines, (b) a short, middle, spiny part (8–15% of total length) with somewhat weaker spines and with few lateral supporting bristles (3–6 on each side), and (c) a long, downstream, bristly part (27–49% of total length) with 15–50 straight or curved short bristles and possibly a few hooked ones (Fig. 11A). The secondary cushions (SCs) are also armed upstream with spines (less robust than on the primary cushions) and downstream with short bristles; the SCs are wide at the upstream end, narrowing noticeably downstream, in most species with a heterogeneous scattering of the spines (the spines are lacking in some irregular spots) (Fig. 12A), more rarely with a homogeneous scattering of the spines. In the soldier’s enteric valve, the primary cushions are barely or not at all outlined (Fig. 12B); the lateral supporting bristles are very short or absent and there are few bristles on the downstream end. The secondary cushions are like those of workers but bear less developed spines. This basic valve pattern is therefore characterised by scarcely developed valves, with very few (± 6) supporting bristles on each side of the PCs and generally heterogeneous spines scattering on the SCs; all of the species are small. Material examined Ten taxa have such enteric valves, Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on pages 30-33, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Cubitermes orthognathus (Emerson, 1928) BURUNDI • 1 ♀ (queen), 1 ♂ (king), 2 soldiers, 2 workers (two samples); Gihofi-Giharo road; 27 Nov. 2013; ULB • 1 soldier, 1 worker; Gisagara; 30 Nov. 2013; ULB • 1 ♀ (queen), 1 ♂ (king), 3 soldiers, 3 workers (three samples); Ruvubu National Park; 29 Nov. 2013; ULB • 4 soldiers, 4 workers (four samples); Ruvubu National Park; 2014; ULB. DEMOCRATIC REPUBLIC OF THE CONGO • 1 soldier, 1 worker, paratypes; Haut-Uélé, Faradje; 1912; RMCA • 1 worker, paratype; Haut-Uélé, Faradje; 1912; PPRI • 2 soldiers, 2 workers (two samples); Haut-Uélé, Toro; 1926; RMCA • 3 ♀♀ (queens), 1 ♀ (alate), 1 ♂ (king), 1 ♂ (alate), 3 soldiers, 3 workers (six samples); Haut-Uélé, Garamba National Park; 1950–1951; RMCA • 1 soldier, 1 worker; Mikembo; 21 Dec. 2011; ULB. KENYA • 1 ♀ (alate), 1 soldier, 1 worker; 7 Aug. 2015; ULB. RWANDA • 1 ♀ (queen), 1 soldier, 1 worker; Kinazi; 8 Jan. 1953; RMCA. UGANDA • 1 ♀ (queen), 1 soldier, 1 worker; Bugiri; 8 Aug. 1957; RMCA., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 50, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Emerson A. E. 1928. Termites of the Belgian Congo and the Cameroon. Bulletin of the American Museum of Natural History 57 (7): 401 - 574."]}

Cubitermes comstocki (Emerson, 1928) CAMEROON • 1 ♀ (queen), 1 worker, paratypes; Bipindi; 1920; AMNH • 1 soldier, 1 worker, paratypes; Bipindi; 1920; RMCA., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 40, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Emerson A. E. 1928. Termites of the Belgian Congo and the Cameroon. Bulletin of the American Museum of Natural History 57 (7): 401 - 574."]}

Cubitermes schmidti (Emerson, 1928) CAMEROON • 1 soldier, 1 worker, paratypes; Bipindi; 1920; RMCA • 1 soldier, 1 worker, paratypes; Bipindi; 1920; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 41, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Emerson A. E. 1928. Termites of the Belgian Congo and the Cameroon. Bulletin of the American Museum of Natural History 57 (7): 401 - 574."]}

Cubitermes oblectatus Harris, 1958 DEMOCRATIC REPUBLIC OF THE CONGO • 1 ♀ (queen), 1 soldier, 1 worker, paratypes (two samples); Katanga, Upemba National Park; 14 Jul. 1947; RMCA • 1 ♀ (queen), 1 soldier, 1 worker (two samples); same data as preceding; RMCA. This species has only been collected in East Africa (Fig. 19D)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 54, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Harris W. V. 1958. Isoptera. In: De Witte G. F. (ed.) E xploration du Parc National de l'Upemba, I. Mission G. F. De Witte 52: 3 - 26. Institut des Parcs nationaux du Congo belge, Bruxelles."]}

Cubitermes hamatus Sjöstedt, 1926 CONGO • 1 soldier, 1 worker, syntypes; Brazzaville; 1907; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 55, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Sjostedt Y. 1926. Revision der Termiten Afrikas. 3. Monographie. Kungliga Svenska Vetenskapsakademiens Handlingar (Ser. 3) 3 (1): 1 - 419."]}

Cubitermes breviceps (Sjöstedt, 1913) TANZANIA • 1 soldier; Dodoma; 8 Jan. 1935; note that the inclusion of C. breviceps in this species group is based on a single non-type soldier; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 38, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Sjostedt Y. 1913. Uber Termiten aus dem inneren Kongo, Rhodesia und Deutsch-Ostafrica. Revue de Zoologique Africaine 2 (3): 354 - 391."]}

Cubitermes bulbifrons Sjöstedt, 1924 DEMOCRATIC REPUBLIC OF THE CONGO • 1soldier, 1 worker, syntypes; Bas-Congo, Mukimbungu; 1904?; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 43, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Cubitermes testaceus Williams, 1966 BURUNDI • 1 ♀ (alate), 1 ♂ (alate), 1 soldier, 1 worker; Mahanga; 26 Nov. 2013; ULB. RWANDA • 1 ♀ (queen), 2 soldiers, 2 workers (two samples); Rukara; 5 Feb. 1953; RMCA. UGANDA • 1 ♀ (queen), 1 soldier, 1 worker, paratypes; Hoima-Fort Portal road; 17 Feb. 1955; NHMUK • 1 soldier, 1 worker, paratypes; Kawanda; 28 Dec. 1949; PPRI., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 51, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Williams R. M. C. 1966. The East African termites of the genus Cubitermes (Isoptera: Termitidae). Transactions of the Royal Entomological Society of London 118 (4): 73 - 118. https: // doi. org / 10.1111 / j. 1365 - 2311.1966. tb 00835. x"]}

Cubitermes sulcifrons Wasmann, 1911 CAMEROON • 1 ♀ (queen), 1 soldier, 1 worker, paratypes; Lolodorf; Nov. 1895; NHMM. DEMOCRATIC REPUBLIC OF THE CONGO • 1 soldier, 1 worker; Yangambi; 15 Jul. 2013; ULB. GABON • 1 soldier, 1 worker; Nkobissimo; 15 Apr. 2017; ULB., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 55, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Wasmann E. 1911. Zur Kenntnis der Termiten und Termitengaste vom Belgischen Congo. Revue de Zoologique Africaine 1 (1 - 2): 91 - 117 & 145 - 176."]}

Cubitermes niokoloensis Roy-Noël, 1969 SENEGAL • 1 ♀ (alate), 1 soldier, paratypes; Bangare-Niokolo-Koba road; Jan. 1965; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 47, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Roy-Noel J. 1969. Isoptera. In: Le Parc national du Niokolo-Koba (Senegal). Memoires de l'Institut fondamental d'Afrique noire 84: 113 - 178."]}

Cubitermes bugeserae Bouillon & Vincke, 1971 BURUNDI • 1 soldier, 1 worker; Mahanga; 26 Nov. 2013; ULB • 1 ♀ (alate), 1 ♂ (alate), 4 soldiers, 4 workers (four samples); Ruvubu National Park; 2014; ULB. RWANDA • 1 ♀ (queen), holotype; Bugesera; 16 Nov. 1963; RMCA • 1 soldier, paratype; same data as preceding; RMCA., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 55, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Bouillon A. & Vincke P. P. 1971. Valvule enterique et revision du genre Cubitermes Wasmann. Cubitermes bugeserae sp. nov. (Isoptera, Termitidae). Revue de Zoologie et de Botanique Africaines 84 (3 - 4): 269 - 280."]}

Cubitermes zenkeri (Desneux, 1904) CAMEROON • 1 ♀ (alate), 1 soldier, 1 worker, syntypes; 1904?; NHMM • 1 ♀ (queen), 1 ♀ (alate), 1 soldier, 1 worker; Bipindi?; 1904?; AMNH. The above were collected in West, Central and East Africa (Fig. 15D)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 42, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Desneux J. 1904. Notes termitologiques. Annales de la Societe entomologique de Belgique 48 (3): 146 - 151."]}

Cubitermes pretorianus Silvestri, 1914 REPUBLIC OF SOUTH AFRICA • 1 soldier, 1 worker, syntypes; Transvaal, Pretoria; 14 Mar. 1913; IEAP., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 34, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Silvestri F. 1914. Contribuzione alla conoscenza dei Termitidi e Termitofili dell'Africa occidentale. I. Termitidi. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura, Portici 9: 1 - 146."]}

Cubitermes exiguus Mathot, 1964 DEMOCRATIC REPUBLIC OF THE CONGO • 1 ♀ (alate), 1 worker; Kinshasa, University campus; 20 Aug. 1963; PPRI • 1 ♂ (king), 1 ♀ (queen), 1 soldier, 1 worker; same data as preceding; 6 Jun. 1964; RMCA • 1 soldier, 1 worker; Kinshasa, Njili–Sanda road; 19 May 1964; RMCA • 1 ♀ (queen), 1 soldier, 1 worker; Kinshasa, Kimwenza; 11 Mar. 1965; RMCA • 1 ♀ (queen), 1 soldier, 1 worker; Kinshasa, Funa valley; 18 Oct. 2014; ULB • 1 soldier, 1 worker; Kinshasa, Funa valley; 24 Apr. 2017; ULB • 1 ♀ (queen), 1 soldier, 1 worker; Kinshasa, university campus; 24 Apr. 2017; ULB., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 34, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Mathot G. 1964. Description d'une nouvelle espece de Cubitermes du Congo (Isoptera, Termitinae). Cubitermes exiguus n. sp. In: Bouillon, A. (ed.) Etudes sur les termites africains, Un Colloque international, Universite Lovanium: 15 - 21. Edition de l'Universite Leopoldville [Kinshasa]."]}

Cubitermes sankurensis var. elongata Sjöstedt, 1926 DEMOCRATIC REPUBLIC OF THE CONGO • 1 soldier, syntype; no date; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 51, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Sjostedt Y. 1926. Revision der Termiten Afrikas. 3. Monographie. Kungliga Svenska Vetenskapsakademiens Handlingar (Ser. 3) 3 (1): 1 - 419."]}

Cubitermes duplex forma nduma Fuller, 1925 REPUBLIC OF SOUTH AFRICA • 1 ♀ (alate), lectotype (designated by Ruelle 1971); Eastern Cape, Nduma; 20 Sep. 1922; PPRI, registration number: TYPH 0 2127 • 1 ♂ (alate), paralectotype; Eastern Cape, Nduma; 20 Sep. 1922; PPRI., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 49, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Fuller C. 1925. The termites of South Africa. South African Journal of Natural History 5: 167 - 246."]}

Cubitermes weissi Silvestri, 1912 CONGO • 1 ♀ (alate), 1 soldier, 1 worker, syntypes; Brazzaville; 1907?; IEAP • 1 soldier, 1 worker; Brazzaville; 8 Jun. 1946; RMCA. DEMOCRATIC REPUBLIC OF THE CONGO • 2 ♀♀ (queens), 2 soldiers, 2 workers (two samples); Kinshasa, Bateke Plateau near Menkao; 29 Jan. 1972; RMCA • 1 soldier, 1 worker; Kinshasa, Bateke Plateau, Bombo-Lumene reserve; 15 Dec. 2012; ULB. The above were collected in West, Central and Southern Africa (Fig. 13C)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 37, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Silvestri F. 1912. Termiti raccolte da L. Fea alla Guinea Portoghesa e alla isole S. Thome, Annobon, Principe e Fernando Poo. Annali del Museo Civico di Storia Naturale (Ser 3) 5: 211 - 255."]}

Cubitermes pretorianus var. heidelbergi Fuller, 1925 REPUBLIC OF SOUTH AFRICA • 1 soldier, lectotype (designated by Ruelle 1971); Gauteng; 1917; PPRI, registration number: TYPH 0 2122., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 34, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Fuller C. 1925. The termites of South Africa. South African Journal of Natural History 5: 167 - 246."]}

Cubitermes pallidiceps (Sjöstedt, 1913) DEMOCRATIC REPUBLIC OF THE CONGO • 1 soldier, 1 worker, syntypes; Katanga, Lubumbashi; 19 Mar. 1912; RMCA • 2 ♀♀ (queens), 1 ♂ (king), 5 soldiers, 5 workers (five samples); Katanga, Lubumbashi; 20 Feb.–19 Apr. 1912; RMCA • 1 ♀ (queen), 1 ♂ (king), 1 soldier, 1 worker; Katanga, Kasenga; 2 Feb. 1912; RMCA • 1 soldier, 1 worker; Katanga, Lubumbashi-Kasenga road; Jul. 2014; ULB • 3 soldiers, 3 workers (three samples); Katanga, Mikembo; Jun. 2016; ULB., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 38, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Sjostedt Y. 1913. Uber Termiten aus dem inneren Kongo, Rhodesia und Deutsch-Ostafrica. Revue de Zoologique Africaine 2 (3): 354 - 391."]}

Cubitermes umbratus Williams, 1954 TANZANIA • 1 ♂ (alate), 1 soldier, 1 worker, paratypes; Usambara; 1 Apr. 1951; NHMUK • 1 ♀ (queen), 1 soldier, 1 worker, paratypes; Amani; 3 Jan. 1952; NHMUK. The above were collected in Central and East Africa (Fig. 20D)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 55, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Williams R. M. C. 1954. New East African Termitinae (Isoptera: Termitidae). Proceedings of the Royal Entomological Society of London B 23 (11 - 12): 215 - 227."]}

Cubitermes glebae (Sjöstedt, 1913) KENYA • 1 ♀ (queen), 1 ♂ (king), 1 soldier, 1 worker; near Nairobi; 20 Nov. 1934; AMNH. TANZANIA • 1 ♀ (alate), 1 soldier, 1 worker; Kilimanjaro, Kibongoto; 2 Mar. 1954; NHMUK., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 38, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Sjostedt Y. 1913. Uber Termiten aus dem inneren Kongo, Rhodesia und Deutsch-Ostafrica. Revue de Zoologique Africaine 2 (3): 354 - 391."]}

Cubitermes muneris (Sjöstedt, 1913) DEMOCRATIC REPUBLIC OF THE CONGO • 1 soldier, syntype; Katanga, Lumata; 6 Jun. 1912; RMCA • 6 ♀♀ (queens), 3 ♂♂ (kings), 3 soldiers, 3 workers (six samples); Katanga, Upemba National Park; 28 Mar.–29 Nov. 1947; RMCA. KENYA • 2 soldiers, 2 workers (two samples); 6–7 Aug. 2015; ULB., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 38, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Sjostedt Y. 1913. Uber Termiten aus dem inneren Kongo, Rhodesia und Deutsch-Ostafrica. Revue de Zoologique Africaine 2 (3): 354 - 391."]}

Cubitermes severus Silvestri, 1914 BENIN • 1 ♀ (queen), 5 soldiers, 5 workers (five samples); Birni forest; Nov. 2016; ULB. BURKINA FASO • 1 soldier, 1 worker; Bama; 2012; ULB. CAMEROON • 1 ♀ (alate), 1 ♂ (alate), 1 soldier, 1 worker; Batié; ULB • 1 ♀ (alate), 1 ♂ (alate), 1 soldier, 1 worker; Boulourou; ULB • 1 ♀ (alate), 1 ♂ (alate), 1 soldier, 1 worker; Ekoua; ULB • 1 ♀ (alate), 1 ♂ (alate), 2 soldiers, 2 workers (two samples); Koutaba; ULB. CÔTE D’IVOIRE • 1 soldier, 1 worker; Youhouli; 7 Sep. 1953; AMNH • 2 soldiers, 2 workers (two samples); Dabou; 18 Feb. 2015; ULB • 1 ♀ (alate), 1 ♂ (alate), 2 soldiers, 1 worker (two samples); Korhogo; 2011-2013; ULB • 1 soldier, 1 worker; Katiola; Feb. 2015 • 1 ♀ (queen), 3 soldiers, 3 workers (three samples); Man; Feb. 2015; ULB • 1 ♀ (queen), 1 ♂ (king), 2 soldiers, 2 workers (two samples); Zéalé; 18 Feb. 2015; ULB • 1 ♂ (alate), 3 soldiers, 3 workers (three samples); Lamto; 2010; ULB • 1 soldier, 1 worker; Lamto; 21 Feb. 2015; ULB • 1 soldier, 1 worker; Taabo; 21 Feb. 2015; ULB. GAMBIA • 1 soldier, 1 worker; Brikama; 7 Sep. 1953; ULB. GUINEA • 2 soldiers, 2 workers, syntypes (two samples); Camayenne and Kakoulima; Aug.–Oct. 1912; AMNH • 1 ♀ (queen), 2 soldiers, 1 worker (two samples); Camayenne; 1934; MNHN • 1 soldier; Forecariah; 21 Jun. 1996; RMCA • 1 soldier, 1 worker; Haut-Niger National Park; 22 Dec. 1996; MURS., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 41, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Silvestri F. 1914. Contribuzione alla conoscenza dei Termitidi e Termitofili dell'Africa occidentale. I. Termitidi. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura, Portici 9: 1 - 146."]}

Cubitermes modestior Silvestri, 1914 CÔTE D’IVOIRE • 1 soldier, 1 worker; Banco forest; 21 Aug. 1968; RMCA. GUINEA • 1 ♀ (queen), 1 soldier, 1 worker, syntypes; Kindia; 20 Aug. 1912; IEAP • 1 ♂ (king), 1 soldier, 1 worker, syntypes; Kindia; 20 Aug. 1912; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 41, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Silvestri F. 1914. Contribuzione alla conoscenza dei Termitidi e Termitofili dell'Africa occidentale. I. Termitidi. Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura, Portici 9: 1 - 146."]}

Cubitermes antennalis Sjöstedt, 1924 DEMOCRATIC REPUBLIC OF THE CONGO • 1 ♀ (queen), 1 ♀ (alate), 1 ♂ (alate), 1 soldier, 1 worker, syntypes; Haut-Uele, Moto; 1923; RMCA., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 40, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Cubitermes zulucola Sjöstedt, 1924 REPUBLIC OF SOUTH AFRICA • 1 soldier, lectotype (designated by Ruelle 1971); Kwazulu-Natal, Hlabisa; 29 Sep. 1922; PPRI, registration number: 0 2129 • 1 ♀ (alate), 1 soldier, paralectotypes; same data as preceding; PPRI • 1 soldier, 1 worker; Kwazulu-Natal, Hlabisa; 2 Aug. 1921; AMNH • 1 soldier, 1 worker; Kwazulu-Natal, Somkele; 25 Apr. 1935; AMNH. The above were collected in Central and Southern Africa (Fig. 12 C)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 35, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Unknown, new species DEMOCRATIC REPUBLIC OF THE CONGO •, 1 ♀ (queen), 1 soldier, 1 worker; Kasaï, Kondué; 1905; RMCA • 1 soldier, 1 worker; Kinshasa, Bateke Plateau, Duale; Jan. 2011; ULB • 1 soldier, 1 worker; Kinshasa, Bateke Plateau, Bombo-Lumene reserve; 22 Apr. 2017; ULB. The above were all collected only in Central Africa (Fig. 16C)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 44, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Cubitermes gibbifrons Sjöstedt, 1924 DEMOCRATIC REPUBLIC OF THE CONGO • 1soldier, 1 worker, syntypes; Bas-Congo, Mukimbungu; 1904?, NHMM • 1 soldier, 1 worker; Bas-Congo, Luali; 26 Aug. 1912; RMCA • 3 ♀♀ (alates), 3 ♂♂ (alates), 3 soldiers, 4 workers (four samples); Kasaï, Kondué; 1924; RMCA., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 44, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Cubitermes zavattarii Ghidini, 1937 ETHIOPIA • 1 worker, paratype; Moiale; 18 May 1937; MCGD • 1 worker; Moiale; 13 Apr. 1937; MCGD. KENYA • 1 soldier, 1 worker; 14-falls; 15 May 1952; AMNH. The above were collected in East and Southern Africa (Fig. 14C)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 39, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Ghidini G. M. 1937. Missione del Prof. Edoardo Zavattari nel paese del Borana. Nuove specie di termitidi (diagnosi preventive). Bollettino della Societa Entomologica Italiana 69 (9 - 10): 141 - 143."]}

Unknown, new species 2 CAMEROON • 1 soldier, 1 worker; Ekoua; 28 Mar. 2015; ULB • 1 ♀ (alate), 1 soldier, 1 worker; Akolo; 1 Apr. 2015; ULB • 1 soldier, 1 worker; Boulourou; 1 Apr. 2015; ULB • 1 soldier, 1 worker; Foumban; 5 Apr. 2015; ULB • 1 soldier, 1 worker; Batié; 7 Apr. 2015; ULB • 1 soldier, 1 worker; Koutaba; 8 Apr. 2015; ULB • 1 soldier, 1 worker; Néfanté; 9 Apr. 2015; ULB. The above were collected in West, Central, East and Southern Africa (Fig. 17G)., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 52, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

4. The fungifaber valve pattern group In the worker’s enteric valve, the primary cushions are of two types: the even PCs are similar to the PCs of the basic enteric valves but the odd PCs, in their downstream part, are crested and protrude within the lumen of the valve (as in Fig. 4A). The primary cushions are either fusiform (their largest width is located near the middle and their lateral margins converge gradually towards both ends) or triangular (their largest width is located near their upstream end and their lateral margins converge gradually downstream). The odd PCs bear crests that are as high as or higher than they are wide, with longer and stronger bristles (Fig. 11E). On a microscope slide, it is quite frequent that at least one of these PCs is twisted, showing the crest in profile (Fig. 11 E’). The enteric valve thus shows triradial symmetry. In some samples, however, the crests are weakly developed; their valves are then sometimes difficult to distinguish from the basic enteric valves of the bilobatodes or the muneris patterns. An odd PC is made of (a) an upstream spiny part (18–42% of total length) with relatively strong spines, (b) a middle spiny part (34–55% of total length) with somewhat weaker spines and with 13–30 lateral supporting bristles on each side, and (c) a bristly part (21–36% of total length) with 35–85 long, straight or curved bristles on a crest generally higher than wide; behind the crest, the bristly part bears some curved or hooked bristles (Fig. 15A). The secondary cushions are wide at the upstream end, narrowing noticeably downstream with a homogeneous spine scattering (Fig. 15A). In the soldier’s enteric valve (Fig. 15B), the crests (on the odd PCs) are generally absent or weakly developed, and often their valves can hardly be distinguished from the basic enteric valves of the bilobatodes or muneris patterns. The secondary cushions are like those of workers but bear less developed spines. The imago’s enteric valve is much smaller than those of the soldiers and workers; the primary cushions are simple, without any crest, the secondary cushions are wide and homogeneous (Fig. 15C). This basic valve pattern is therefore characterised by high and narrow hairy crests on the downstream end of the odd PCs; most species are large. Material examined Eleven species have such enteric valves

Unknown, new species 1 BURKINA FASO • 1 soldier, 1 worker; Bala; Aug. 2016; ULB • 2 soldiers, 2 workers (two samples); Bama; Aug. 2016; ULB • 1 ♂ (king), 1 soldier, 1 worker; Bana; Aug. 2016; ULB • 2 soldiers, 2 workers (two samples:); Bekuy; Aug. 2016; ULB • 1 soldier, 1 worker; Bobo Dioulasso; Aug. 2016; ULB • 1 soldier, 1 worker; Poundoun; Aug. 2016; ULB • 1 ♀ (queen), 4 soldiers, 4 workers (four samples); Tiogo; Aug. 2016; ULB. CÔTE D’IVOIRE • 1 soldier, 1 worker; Lamto; no date; ULB • 3 ♀♀ (alates), 3 ♂♂ (alates), 4 soldiers, 4 workers (four samples); Toumodi; Feb.–Apr. 2015; ULB. GUINEA • 1 soldier; Kindia; Nov. 1995; RMCA. SENEGAL • 1 soldier, 1 worker; Casamance, Balingore; 13 Sept. 2011; ULB • 1 soldier, 1 worker; Indiga; 22 Nov. 2016; ULB • 1 soldier, 1 worker; Tambacounda; 22 Nov. 2016; ULB., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 52, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Cubitermes fungifaber var. elongata Sjöstedt, 1924 (invalid name) DEMOCRATIC REPUBLIC OF THE CONGO • 1soldier, 1 worker, syntypes; Bas-Congo, Mukimbungu; 1904; AMNH • 1 ♀ (alate), 2 ♂♂ (alates), 2 workers (two samples); Kasaï, Kondué; no date; RMCA., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 44, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175

Cubitermes anatruncatus (Fuller, 1925) REPUBLIC OF SOUTH AFRICA • 1 soldier, lectotype (designated by Ruelle 1971); Kwazulu-Natal, Conjeni; 29 Jul. 1921; PPRI, registration number: TYPH 0 2119 • 1 ♀ (alate), 1 ♂ (alate), 1 soldier, 1 worker, paralectotypes; Kwazulu-Natal, Conjeni; 29 Jul. 1921; PPRI., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 49, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Fuller C. 1925. The termites of South Africa. South African Journal of Natural History 5: 167 - 246."]}

Cubitermes undulatus (Fuller, 1925) REPUBLIC OF SOUTH AFRICA • 1 soldier, lectotype (designated by Ruelle 1971); Gauteng, Pretoria; 1980; PPRI, registration number: TYPH 0 2128 • 1 soldier, 1 worker, paralectotypes; Gauteng, Pretoria; 1980; PPRI., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 39, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Fuller C. 1925. The termites of South Africa. South African Journal of Natural History 5: 167 - 246."]}

Cubitermes tenuiceps (Sjöstedt, 1913) DEMOCRATIC REPUBLIC OF THE CONGO • 1 soldier, 1 worker, syntypes; Katanga, Lubumbashi; 20 Mar. 1912; RMCA • 2 soldiers, 2 workers (two samples); Katanga, Lubemba; Jul. 2014; ULB • 2 soldiers, 2 workers (two samples); Katanga, Mikembo; Jun. 2016; ULB., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 35, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Sjostedt Y. 1913. Uber Termiten aus dem inneren Kongo, Rhodesia und Deutsch-Ostafrica. Revue de Zoologique Africaine 2 (3): 354 - 391."]}

Cubitermes kemneri (Emerson, 1928) CAMEROON • 1 soldier, 1 worker, paratypes; Bipindi; 1920; AMNH., Published as part of Josens, Guy & Deligne, Jean, 2019, Species groups in the genus Cubitermes (Isoptera: Termitidae) defined on the basis of enteric valve morphology, pp. 1-72 in European Journal of Taxonomy 515 on page 44, DOI: 10.5852/ejt.2019.515, http://zenodo.org/record/2638175, {"references":["Emerson A. E. 1928. Termites of the Belgian Congo and the Cameroon. Bulletin of the American Museum of Natural History 57 (7): 401 - 574."]}