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Ivanauskiella sutteri sp. nov. (Figs 2–3, 51, 72) Type material. HOLOTYPF: ♀, GREECE: CRETE: Makrigialos,Aspros Potamos, 20 m, LF, 3.ix.2001, R. Sutter leg. (gen. slide 165/20, OB) (SMNK). PARATYPF: 1 J, same data as for holotype but 8.ix.2001 (gen. slide 6630, Sutter) (SMNK). Diagnosis. This new species can be recognized externally by the white head and white forewing strongly mottled with light brown scales. The male genitalia are identifiable from a broad valva (in basal half about as broad as at apex) and uncus strongly widened after 1/4 (after 1/2–3/ 4 in the remaining species). Two short teeth on colliculum and a single four-horned signum are characteristic for the female genitalia. Description. Adult (Figs 2–3). Wingspan 7.6–8.0 mm. Head white; labial palpus white mixed with brown on outer surface; scape of antenna light brown with white apex, flagellum light brown ringed with white; thorax and forewing covered with white, brown-tipped scales, very indistinct markings and diffuse brown spot in fold in one specimen, fringe white; hindwing white. Male genitalia (Fig. 51). Uncus very slender at base, strongly widened after 1/4–1/3, extending to top of valva, apex triangular, posterior margin weakly serrate; tegumen subrectangular, slightly broader than medial portion of uncus; valva broad, weakly narrowed in middle, gradually bent, top and base as broad as top of uncus; sacculus 1/3–1/2 length of valva, as broad as valva in middle; vinculum short; saccus short, triangular, pointed; phallus with 15–17 large needle-shaped and small, short, thorn-shaped spines, distal plate large, of irregular shape. Female genitalia (Fig. 72). Papilla analis subovate, covered with short hair-like setae; apophysis posterioris straight, as long as apophysis anterioris; segment VIII weakly sclerotised, subtrapezoidal, slightly broader than long; apophysis anterioris straight; ductus bursae gradually broadening anteriorly, colliculum with two short thorn-shaped sclerites, situated at 1/3 of ductus bursae; corpus bursae egg-shaped, signum single plate with four horn-shaped processes – posterior ones shorter than anterior ones. Molecular data. No barcode available. Etymology. The species is named in honour of the late German microlepidopterist Reinhard Sutter, who collected the type series of the new species. Biology. Host plant unknown. Adults were collected at light at an altitude of 20 m in early September. Distribution. Greece: South-eastern Crete.

Ivanauskiella psamathias (Meyrick, 1891) (Figs 8–11, 53, 75) Apodia psamathias Meyrick, 1891: 56. TL: Algeria, Biskra Aristotelia psamathias (Meyrick, 1891): CLARKF (1969: 302) Ivanauskiella psamathias (Meyrick, 1891): PlSKUNOv (1990: 309) Type material examined. LFCTOTYPF: J, ALGERIA: ‘ Biskra,Algeria, 21/4/90 | Apodia psamathias Meyr., Holotype, J | Meyrick Coll. B.M. 1938-290 | Type | Lectotype | J genitalia on slide 19.ii.1949, J.F.G.C. 9056 | Slide NHMUK 010316668ʼ (NHMUK). Additional material examined. ALGERIA: 1 J, ‘ Biskra, [18]97, St[au]d[in]g[er].ʼ (gen. slide 297/20J, OB) (MfN). TUNISIA: 6 JJ 5 ♀♀, “Mauritania”, Tunesia-Sud, Oasis Tozeur, 28.iv.–11.v.1981, M. & W. Glaser leg. (gen. slide 323/ 19♀, 5/20J, 24/20J, 97/22J, 98/ 22♀, OB) (SMNK); 12 JJ 28 ♀♀, Nefta area, 1.–4.v.1988, O. Karsholt leg. (gen. slides 4706, 4707 Hendriksen) (NMPC, ZMKU, ZMUC); 9 JJ, 3 specimens without abdomen, 5 km W Douz, 17.–18.iv.1984, K. Mikkola leg. (gen. slides Hk5534, Hk5444, Hk5547, Hk5548, Hk5549; 5425 OK) (MZHF, ZMUC); 1 specimen without abdomen, 1 km W. Douz, 19.iv.1984, K. Mikkola leg. (MZHF). Diagnosis. Ivanauskiella psamathias is distinctive by having light, greyish-brown forewing with three distinct dark spots in cell, one in fold and two below basal 1/3 of costal margin. Ivanauskiella nigripunctata sp. nov. is similar but smaller (6.5–7.7 mm in wingspan). 3–5 large thorn-shaped and a few small spine-like cornuti in vesica in combination with narrow saccus are characteristic for the male genitalia. Other species of the genus have more, but smaller cornuti. In the female genitalia both signa are rounded and densely covered with short thorns. Redescription. Adult (Figs 8–11). Wingspan 8.7–10.0 mm. Head, thorax and tegulae covered with white brown-tipped scales, frons white; labial palpus upcurved, segment 2 brown with white basal and apical ring, segment 3 white with brown subapical ring; scape of antenna light brown with base and apex white, flagellum light brown ringed with white; forewing covered with white brown-tipped scales, two distinct brown spots in cell and two spots in fold surrounded with white, indistinct brown spots under 1/3 of costal margin and diffuse white tornal and costal spots at 3/ 4 in some specimens, fringe white, brown tipped; hindwing white. Male genitalia (Fig. 53). Uncus slender from base to 2/3, then strongly widened towards triangular apex, extending to top of valva, posterior margin weakly serrate; tegumen subrectangular; valva comparatively broad, in middle twice as broad as medial portion of uncus, dorsal margin straight, ventral margin bent, apical 1/3 as broad as top of uncus; sacculus about half length and as broad as valva in middle; vinculum short; saccus very slender, pointed; phallus with 3–5 elongate, needle-shaped and 5–7 very small thorn-shaped spines, distal plate large, of irregular shape. Female genitalia (Fig. 75). Papilla analis subovate, covered with short hair-like setae and strong basal setae; apophysis posterioris slightly longer than apophysis anterioris; segment VIII weakly sclerotised, subtrapezoidal, slightly broader than long; apophysis anterioris straight; ductus bursae of even width except for gradually broadened anterior portion, colliculum at 1/2 of ductus bursae with one large thorn-shaped sclerite; corpus bursae ovate, both signa rounded serrate plates: anterior one larger than posterior one. Molecular data. No barcode available. Biology. Early stages and host-plant unknown. The specimens from Tunisia were collected at light in a semi-desert habitat. Distribution. The species is restricted to North-West Africa: Algeria and Tunisia (MFYRlCK 1891, CARADIA 1920). Records from Europe and Asia should be referred either to I. turkmenica or I. limoniella sp. nov. or I. occitanica (details under those species). ‘Mauretaniaʼ on the labels on some specimens refers to an old concept of Mauretania, covering all of North-West Africa. Remarks. Apodia psamathias was described based on two males collected in April 1890 at the oasis of Biskra, Algeria. CLARKF (1969: 302, figs 1–1b) designated one specimen as lectotype and provided a monochrome photographs of the left wings, phallus and ventral view of the male genitalia of this specimen. Photographs of the lectotype in high resolution and a slide of its genitalia have been examined by authors and compared with specimens from Tunisia. Despite the lectotype looking more unicoloured with less distinct spots in the fold, its wing pattern with characteristic markings in the cell corner and apex spotted with white match well ‘fresher ʼ specimens collected in Tunisia. Four large thorn-shaped cornuti in the vesica of the lectotype of A. psamathias also justify its conspecificity with specimens from Tunisia., Published as part of Bidzilya, Oleksiy, Karsholt, Ole & Šumpich, Jan, 2023, A revision of the genus Ivanauskiella (Lepidoptera: Gelechiidae), pp. 135-164 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 63 (1) on pages 139-145, DOI: 10.37520/aemnp.2023.007, http://zenodo.org/record/8005865, {"references":["MEYRICK E. 1891: A fortnight in Algeria, with descriptions of new Lepidoptera. Entomologist ʼ s Monthly Magazine 27: 9 - 13, 55 - 62.","CHRETIEN P. 1908: Description de deux Gelechides nouvelles dʼAlgerie. Bulletin de la Societe Entomologique de France 13: 91 - 93.","NEL J. & VARENNE T. 2013: Spatuncusella occitanica gen. nov., sp. nov., decouvert dans le Midi de la France (Lepidoptera, Gelechiidae, Gelechiinae, Anomologini). Revue de l'Association Roussillonaise d'Entomologie 22 (1): 35 - 40.","IVINSKIS P. & PISKUNOV V. 1980: Opisanie roda semeistva vyemchatokrylykh molei Ivanauskiella Ivinskis et Piskunov, gen. n. i vida Ivanauskiella turkmenica Ivinskis et Piskunov, sp. n. iz Turkmenskoi SSR. [Description of Gelechiid moth genus Ivanauskiella Ivinskis et Piskunov, gen. n. and species Ivanauskiella turkmenica Ivinskis et Piskunov, sp. n. from the Turkmenian SSR]. Lietuvos TSR Moksl u Akademijos Darbai, C Serija 2 (90): 23 - 27 (in Russian)."]}

Ivanauskiella ainella (Chrétien, 1908) (Figs 12–15, 45, 54, 76) Apodia ainella Chrétien, 1908: 91. TL: Algeria, Biskra Ivanauskiella ainella (Chrétien, 1908): PlSKUNOv (1990: 309) Type material examined. PARATYPFS: J, ‘ Apodia ainella Chret. | 11.4 | Paratype ʼ (gen.slide 193/20, OB) (SMNK); 2 ♀♀, ‘ Biskra, 30.4.[19]07 ʼ (MNHN). Additional material examined. ALGERIA: 1 J, Biskra, i.–v.1901, Steinbach leg. (genitalia in vial, 139/20, OB) (MfN). MOROCCO: 12 JJ 10 ♀♀, Erfoud/Rissani area, 13.–14.iv. 1989, 900 m, O. Karsholt leg. (gen.slide 177/22J; 178/ 22♀; 180/22J, OB) (NMPC, ZMKU, ZMUC). Diagnosis. Ivanauskiella ainella is a comparatively large light brown species with dark brown markings (spots and streaks). The male genitalia are characterised by 10–11 slender spine-like cornuti in vesica in combination with equilateral triangular saccus. The female genitalia resemble those of I. nigripunctata sp. nov. but differ in upcurved rather than transverse processes of posterior signum. Redescription. Adult (Figs 12–15). Wingspan 10.0–13.0 mm. Head and tegulae uniformly greyish brown, frons light grey, thorax dark brown; labial palpus slender, upcurved, light brown, darker on outer surface, segment 2 with light apical ring; scape of antenna brown, flagellum brown, indistinctly ringed with grey; forewing light greyish-brown with dark brown suffusion along veins, in fold, below costal margin and in subapical 1/4, diffuse blackish spots in cell and below base of costa, blackish elongate spot and slender streak in fold; fringe grey, brown tipped; hindwing light grey. The examined specimens show variation in the colour of the forewing, from light brown to blackish brown. In specimens with dark forewings the blackish spots become obsolete. Male genitalia (Figs 45, 54). Uncus slender from base to 2/3, then strongly widened towards rounded apex, extending to top of valva, posterior margin serrate; tegumen subrectangular; valva gradually bent, in middle twice as broad as medial portion of uncus, apical 1/3 as broad as top of uncus; sacculus about half length and as broad as valva in middle; vinculum short; saccus broad, triangular, weakly acute; phallus with 12–15 elongated, needle-shaped spines and a few small thorns, distal plate large, of irregular shape. Female genitalia (Fig. 76). Papilla analis subovate, covered with short hair-like setae and strong basal setae; apophysis posterioris straight, slightly longer than apophysis anterioris; segment VIII weakly sclerotised, subtrapezoidal, as broad as long; apophysis anterioris straight; ductus bursae weakly broadened anteriorly, colliculum with one large thorn-shaped sclerite, situated at 2/3 of ductus bursae; corpus bursae egg-shaped, anterior signum rounded plate, posterior signum irregular plate with two upcurved horns. Molecular data. No barcode available. Biology. Host plant unknown. Adults were recorded in April–May. Distribution. Algeria, Morocco (new record). Remarks. Apodia ainella was described based on an unspecified number of specimens collected in April and May at the oasis of Biskra, Algeria. It was compared with I. psamathias, and stated to differ in larger size, darker wings and elongate (instead of rounded) black spots in the forewing (CHRḖTlFN 1908: 91).A male and a female from the type series deposited in MNHN were examined by D. Povolný, who designated the male as lectotype (POvOLNỶ 1983: 184–185) and provided drawings of its left wings (POvOLNỶ 1983: fig. 29) and genitalia (POvOLNỶ 1983: pl. 6, fig. 31)., Published as part of Bidzilya, Oleksiy, Karsholt, Ole & Šumpich, Jan, 2023, A revision of the genus Ivanauskiella (Lepidoptera: Gelechiidae), pp. 135-164 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 63 (1) on page 145, DOI: 10.37520/aemnp.2023.007, http://zenodo.org/record/8005865, {"references":["CHRETIEN P. 1908: Description de deux Gelechides nouvelles dʼAlgerie. Bulletin de la Societe Entomologique de France 13: 91 - 93."]}

Ivanauskiella annekristinae sp. nov. (Figs 23–24, 59) Type material. HOLOTYPF: J, FRANCE: CORSICA: Calvi, 10 m, 18.–19. viii.1998,O.Karsholt leg.(ZMUC).PARATYPFS: 9 JJ, same data as for holotype (gen.slide 137/20,OB;3099J, Hendriksen) (NMPC, ZMKU, ZMUC). Diagnosis. Ivanauskiella annekristinae sp. nov. is characterized by the pure white head, thorax, tegulae and labial palpus, and the white forewings with rather large, dark brown markings. Ivanauskiella nigripunctata sp. nov. is similar but differs in the colour of segment 2 of the labial palpus being brown with white apex, and light grey forewings with smaller brown markings. The male genitalia are similar to those of I. nigripunctata sp. nov. except for the broader uncus and the U-shaped, rather than triangular saccus in I. nigripunctata sp. nov. Description. Adult (Figs 23–24). Wingspan 7.0 mm. Head, thorax and tegulae white; labial palpus white, segment 3 about 2/3 length of segment 2 and at its base almost as wide as segment 2, acute; scape of antenna light brown, flagellum light brown, ringed with white; forewing white with gentle light brown suffusion at base, along costal margin and in fold, apex covered with darker, light brown scales with brown tips, two small blackish brown spots in fold, large blackish brown spot at 2/3 and another small spot at 3/4; fringe white, tipped with brown; hindwing and fringe white. Male genitalia (Fig. 59). Uncus moderately broad, apex slightly widened, rounded, extending to top of valva, posterior margin weakly serrate; tegumen subrectangular, slightly broader than medial portion of uncus; valva moderately broad, in middle twice as broad as uncus, gradually bent, top and base as broad as top of uncus; sacculus about half length of valva, as broad as valva in middle; vinculum short; saccus short, U-shaped; phallus with 25–30 dense needle-shaped spines, distal plate ovate. Female genitalia. Unknown. Molecular data. No barcode available. Etymology. The new species is named after Anne Kristine Karsholt for supporting OK by collecting the type series during their honeymoon to Corsica 25 years ago. Biology. Host plant unknown, but possibly Limonium sp., which grew at the type locality. The adults were collected in a light trap with an 8 Watt actinic tube during mid-August. The trap was placed on a cliff ledge facing the Mediterranean Sea. Distribution. France: Corsica. Remarks. Ivanauskiella annekristinae is superficially similar to Phragmatodes cyrneogonella Nel & Varenne, 2012, which is endemic to Corsica and Sardinia. The latter has larger blackish spots and more blackish brown suffusion in the forewing, and the outside of segment 2 of the labial palpus is blackish brown., Published as part of Bidzilya, Oleksiy, Karsholt, Ole & Šumpich, Jan, 2023, A revision of the genus Ivanauskiella (Lepidoptera: Gelechiidae), pp. 135-164 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 63 (1) on pages 152-153, DOI: 10.37520/aemnp.2023.007, http://zenodo.org/record/8005865

Ivanauskiella bovis sp. nov. (Figs 4–7, 46, 52, 73–74) Type material. HOLOTYPF: ♀, MOROCCO: HIGH ATLAS: 7 km S of Ouirgane, 950 m, 31°8′19″N 8°5′51″W, 1.–5.v.2016, C. Hviid, K. Larsen & D. Nilsson leg. (gen. slide 164/20, OB) (ZMUC). PARATYPFS: MOROCCO: HIGH ATLAS: 3 JJ, 27 km NW Taroudant, 670 m, N31.2519, W87.76916, 29.iv.2013, J. Tabell leg. (ZMUC); 3 JJ 6 ♀♀, 6 km NW Ouirgane, road to Amizmiz, 1050 m, 31°12′29″N, 8°4′23″W, 1.–2.v.2016, C. Hviid,K.Larsen & D.Nilsson leg.(gen.slide 132/20, OB; 5302 OK) (ZMUC); 19 JJ 13 ♀♀, 7 km S Ouirgane, 950 m, 31°8′19″N, 8°5′51″W, 1.–5.v.2016, C. Hviid, K. Larsen & D. Nilsson leg. (gen.slide 164/20, OB) (NMPC, ZMKU, ZMUC); 1 J 1 ♀, 3 km S Ouirgane, 950 m, 31°11′31″N, 8°3′16″W, 1.–5.v.2016, C. Hviid, K.Larsen & D. Nilsson leg. (gen. slide 132/20, OB) (ZMUC); 5 JJ, Guelmim-Qued Noun, Sidi Ifni, 29°23′00″N, 10°10′21″W, 5.–7.iii.2017, sea level, C. Hviid, O. Karsholt, K. Larsen & D. Nilsson leg. (gen. slide 131/20, OB) (ZMUC). ANTI ATLAS: 1 J, 12 km E Tafraout, 1690 m, N29.72454, W8.84551, 27.iv.2013, J. Tabell leg. (ZMUC). Diagnosis. This new species is identifiable externally by the uniformly coloured ochreous brown forewing, which has not been observed in other species of Ivanauskiella. The male genitalia of I. bovis sp. nov. are rather similar to those of I. ainella and I. limoniella sp. nov., but can be separated by the shape of the cornuti in the vesica, which are comparatively short and broad in I. bovis sp. nov. contrary to the slender needle-shaped cornuti in related species. In the female genitalia colliculum with one large tooth situated at 2/3 of ductus bursae and the connected signa with bull skull shaped posterior ‘hornsʼ are diagnostic. Description. Adult (Figs 4–7). Wingspan 10.0– 12.5 mm. Head, thorax and tegulae ochreous brown with slight sheen, frons whitish; labial palpus uniformly brown, with inner surface light brown, scape of antenna and flagellum as head, with indistinct light rings; forewing ochreous brown, fold and some veins look darker in some specimens; hindwing light grey. Variability. The specimens of the type series vary slightly in the colour of the forewing, from lighter to darker ochreous brown. Some specimens have the fold slightly darker and a tendency to a dark spot at the end of the cell. In other specimens there is a diffuse whitish subapical fascia. Male genitalia (Figs 46, 52). Uncus very slender from 1/3 to 3/4, apex strongly widened, rounded, extending to top of valva, posterior margin weakly serrate; tegumen sub-rhomboid; valva moderately broad, in middle twice as broad as uncus, gradually bent, apical 1/3 and base as broad as top of uncus; sacculus about 1/2 length of valva, as broad as valva in middle; vinculum short; saccus short, triangular; phallus with 10–12 prominent comparatively short and broad cornuti and 8–10 small thorn-shaped spines, distal plate large, of irregular shape. Female genitalia (Figs 73–74). Papilla analis subovate, covered with short hair-like setae and strong basal setae; apophysis posterioris straight, as long as apophysis anterioris; segment VIII weakly sclerotised, subtrapezoidal, slightly broader than long; apophysis anterioris straight; ductus bursae of even width, colliculum with one large thorn-shaped sclerite, situated at 2/3 of ductus bursae; corpus bursae egg-shaped, signa connected: anterior one rounded to weakly elongate plate, posterior one shaped as bull skull with more or less strongly curved horns. Molecular data. No barcode available. Etymology. The species name is genitive of the Latin word bos, bull, and refers to the bull skull shaped signum that is characteristic for the new species. Biology. Host plant unknown. Adults were collected at light at stony localities from March to May – at altitudes from near sea level near the coast of the Atlantic Ocean up to 1690 m in the Anti-Atlas Mts. Distribution. Morocco., Published as part of Bidzilya, Oleksiy, Karsholt, Ole & Šumpich, Jan, 2023, A revision of the genus Ivanauskiella (Lepidoptera: Gelechiidae), pp. 135-164 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 63 (1) on page 139, DOI: 10.37520/aemnp.2023.007, http://zenodo.org/record/8005865