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In dieser Arbeit wurden drei Methoden getestet und analysiert, um die Temperatur, den Ladezustand und den Alterungszustand von Lithium-Ionen-Pouchzellen zu messen. Diese Methoden sollen mehr Informationen ��ber den Zustand der Batterie w��hrend des Betriebs liefern, um die Effizienz zu erh��hen, die Lebensdauer zu verl��ngern und die Sicherheit zu steigern. Um die Genauigkeit der Temperaturmessung zu verbessern, wurde ein Sensor in das Elektrodenpaket integriert. Das Experiment hat gezeigt, dass der innere Sensor w��hrend eines ��berladungstests eine Temperatur von ungef��hr 80 C gemessen hat, w��hrend ein Sensor au��erhalb der Zelle meist 60 C nicht ��berschritten hat. F��r die zweite Methode wurde die Laufzeitmessung von akustischen Wellenpaketen gemessen, die durch eine Lithium-Ionen-Pouchzelle geschickt wurden, um den Ladezustand zu ermitteln. Ein Signal, moduliert aus einer Dreieck-Fensterfunktion und einem hochfrequenten Sinussignal, wurde benutzt, dessen Eigenschaften diskutiert und die optimale Zyklenzahl gew��hlt, um genaue und wiederholbare Ergebnisse zu bekommen. Eine lineare Abh��ngigkeit zwischen Laufzeit und dem Ladezustand wurde festgestellt f��r einen Bereich von 100 bis zu 20 % Ladezustand wobei die Laufzeit um 9.4 % k��rzer wurde. F��r die dritte Methode wurde ein force-sensitive-resistor verwendet um die qualitative Volumen��nderung einer Pouchzelle w��hrend des Ladens und Entladens zu messen. Mit einer einfachen und kosteng��nstigen Konstruktion, war es m��glich, an kommerziell erh��ltlichen Pouchzellen Druck��nderungen zu messen, welche ident mit den Resultaten vorangegangener Studien sind. Es war m��glich, die Phasen der Lithium (De-)Interkalation und einen Zusammenhang zwischen Gesamtdruck��nderung und Kapazit��t der Zelle festzustellen. Durch alle drei Methoden war es m��glich, mit einfachen Konstruktionen, wichtige Informationen ��ber den Zustand der Lithium-Ionen-Pouchzelle zu erhalten., In this thesis, three different methods were tested and evaluated to measure the temperature, the state-of-charge and the state-of-health of lithium ion pouch cells. Those methods help to improve the efficiency, extend the service life and increase the safety of lithium ion batteries. To improve the accuracy of the temperature measurement, a sensor was placed inside the electrode pack of the cell to get more accurate data. In the experiments, it was shown that the temperature measurement inside the cell during overcharge tests measured temperatures of about 80 C while outside sensors mostly measured less than 60 C. The second evaluated method measured the time-of-flight of acoustic waves through the pouch cell to determine the state-of-charge. A triangle-windowed sinusoidal wave package was used and its important characteristics were discussed and defined to get accurate and repeatable results. A linear correlation between the change in time-of-flight and state-of-charge was measured for a range from 100 to 20 % charging level with a change of time-of-flight of 9.4 %. The third method used a force-sensitive-resistor to measure the qualitative volume change of a pouch cell for charging and discharging. With a simple and cheap construction, it was possible to measure pressure changes in commercial cells which correspond with the findings of previous studies. It was possible to determine different phases of lithium (de-)intercalation and a relation between total pressure change and cell capacity was measured. All three methods proved to be capable to gather more information of the state of a lithium ion pouch cells with simple and practical constructions.

4. Cionus osmanlis sp. n. Figs 4 a���f. http://zoobank.org/ urn:lsid:zoobank.org:act: 18BF4A18-C98A-4036-A3AE-82C9866E747F Type locality. Artvin (Turkey). Type series. Holotype: 3.50 mm long, well-preserved male labeled ��� 15.06.2002 NE Turkey Macahel Gecidi N Artvin leg. P. Bialooki / coll. Białooki / HOLOTYPUS Cionus osmanlis sp. n. M.Ko��ť��l et R.Caldara des. 2016 [printed red label]��� (UWCP). Paratype: ���J��gaz Dagh Paphlag Turc Staněk lgt. / PARATYPUS Cionus osmanlis sp. n. M.Ko��ť��l et R.Caldara des. 2016��� (1 ♂ NMPC). Description. Male (holotype). Body subrotund. Head: rostrum stout, medium long (l/ w 4.4, Rl/Pl 1.41), black; in lateral view clearly evenly curved, same width from base to antennal insertion, shortly beyond antennal insertion narrower, apically bluntly tapered; in dorsal view narrowest at base, slightly broadened to apex, basal part not laterally constricted, apical part shortly beyond antennal insertion moderately dorsoventrally flattened; from base to antennal insertion with very dense to confluent rugulose punctures, apical part, especially on sides, semi-densely covered with round punctures, at antennal insertion finely longitudinally ribbed; basal part with mostly backwardly oriented, recumbent whitish small thin seta-like scales, apical part with long, forwardly oriented, suberect yellowish seta-like scales. Head between eyes narrow, about 0.3 rostrum width at base. Eyes large, very slightly rounded to flat, not protruding from head outline. Antennae reddish-brown, with very slightly darkened club, inserted at 0.7 of rostrum length; funicle of 0.7 scape length, segment 1 wider than segment 2, as long as segment 2, segment 1 more than twice as long as wide, segment 2 3 �� as long as wide, segment 3 subquadrate, segments 4���5 transverse; club oblong suboval, 2.2 �� as long as wide, of approximately 0.9 funicle length, completely covered with recumbent, densely arranged, brown tiny hairs and sparsely distributed, erect, dark long sensilla. Pronotum: black, markedly wider than long (Pl/Pw 0.60), densely finely, evenly covered with small round punctures; on sides densely covered with subrecumbent, forwardly and medially oriented reddish-brown, elongate (l/w 4���6) scales, at midline with very sparse broad scales, longitudinal area in basal part narrowed forwardly, then slightly broadened to anterior margin, anteriorly with small recumbent, forwardly oriented gray to yellowish scales, lateral scaled pronotal area with small bare ���window���; widest at base, subconically narrowed to 1/2 of pronotal length, then very moderately constricted and narrowed to anterior margin, in lateral view in basal half flat on disc, then slightly falling to anterior margin. Prosternum: anterior margin with deep, nearly rectangularly incised emargination, bounded by sharp elevated margins, not reaching coxae. Scutellum: broadly subtriangular, blunt at apex, at posterior margin with comb of densely arranged, backwardly oriented yellowish elongate scales. Elytra: black, in basal 2/3 evenly, in apical third broadly rounded, short (El/Ew 1.09), widest shortly before mid-length, at base markedly wider than pronotum (Ew/Pw 1.77), humeri clearly but not strikingly prominent, with shallow posthumeral impression; very slightly convex on disc; interstria 1 beyond 1/4 of medial length distinctly, before apex slightly broadened, interstria 2 at same lengths narrowed and markedly constricted laterally, interstria 3 slightly constricted laterally encompassing large dorsal elongate black tomentous and smaller preapical elongate maculae, dorsal macula at posterior, preapical macula at anterior margin with relatively large patch formed by densely arranged overlapping whitish-yellow scales, odd interstriae markedly convex and clearly broader than even ones; odd interstriae with unevenly large, alternating light brownish and black velvet-like patches; striae shallow, formed by irregular single rows of round punctures; entire surface excepting described pattern covered with recumbent gray, light brown and blackish elongate (l/w 3���5) scales, sparsely distributed and leaving integument partially visible. Venter: relatively densely covered with subrecumbent, gray thin long hairs, lateral parts of meso- and metasternum, metepisternum and both epimera covered with densely arranged whitish and reddish elongate scales, most lateral parts of ventrites 3 and 4 with small patches of whitish scales; mesosternal process very short, broad, blunt at apex; metasternum concave, punctured; ventrite 1 with relatively deep, ventrite 2 with shallow medial impression, punctured, with suberect, backwardly oriented, long gray hairs; ventrite 1 1.7 �� as long as ventrite 2, ventrites 1���2 combined 5.1 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.7 length of ventrite 5. Legs: femora and tibiae brown, tarsi light brown; profemora with very small teeth emphasized by several erect scales, meso- and metafemora with large triangular sharp teeth; femora and tibiae covered with unevenly distributed, subrecumbent, whitish, reddish and blackish scales, clustered especially on profemora into feebly distinct transverse bands, tarsi including onychia with whitish suberect hairs; onychia of anterior legs of normal length, maximally of 0.8 length of tarsomeres 1���3 combined; lateral protarsal claws of half length of medial ones, medial claws of meso- and metatarsi only slightly shorter than lateral ones. Penis: Figs 4 d���f, its body broadly pointed at apex. Female. Unknown. Variability. The paratype (3.47 mm long) differs from the holotype by lighter antennae and tarsi, and more pointed apex of the body of penis. Diagnosis. This species is recognizable by smaller size, stout and curved rostrum, clearly vaulted odd interstriae, especially at elytral base, not elongate onychia in males, and penis shape. Comparative notes. Cionus osmanlis is most closely related to C. tuberculosus, from which it differs by not elongate antennal club, more curved rostrum, brown femora and tibiae, shorter protarsal onychia in male, and broader and pointed body of penis. Biological notes. Biology unknown. Distribution. North-eastern and northern (Paphlagonia) Turkey. Etymology. Derived from the genitive of a Latin name of a Turkish inhabitant of Ottoman empire. Non-type specimens examined. None., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 18-19, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117

45. Cionus griseopubens Wingelm��ller, 1914 Figs 45 a���f. Cionus griseopubens Wingelm��ller, 1914: 200; 1921: 117; 1937: 181. Caldara, 2013: 123. Alonso-Zarazaga et al., 2017: 185. Type locality. Paleopanagia, Peloponnesos S Taygetos Mts., 400-500 m (Greece). Type series. The species was described based on a single male specimen. Despite our considerable efforts, we could not find the type in coll. Wingelm��ller (NHMW) or elsewhere. Therefore, in order to fix the taxon with regard to its very closely related species we designated the neotype of Cionus griseopubens Wingelm��ller as currently understood. Given that Wingelm��ller described the species from a Greek specimen, we designated a 4.48 mm long, completely preserved male labeled ���GRAECIA, Peloponnesos S Taygetos Mts., 400-500 m Paleopanagia env. 30.iv.2008. machia S. Benedikt leg.��� by adding the label ��� NEOTYPUS Cionus griseopubens Wingelm��ller R.Caldara et M.Ko��ť��l des. 2017 [printed red label]���. The neotype is deposited in NHMW. Synonyms. None. Redescription. Male. Body stout, suboval to subrotund. Head: rostrum moderately stout, medium long (l/ w 5.0, Rl/Pl 1.29), black; in lateral view moderately evenly curved, of same width from base to antennal insertion, then moderately tapered to apex; in dorsal view near base slightly constricted laterally, in apical part slightly flattened; texture and vestiture as in C. ganglbaueri. Head between eyes and eyes as in C. ganglbaueri except slightly more rounded eyes. Antennae reddish-brown, with darkened club and last funicular segments, inserted before 0.7 of rostrum length; funicle of more than 0.6 scape length, segment 1 slightly wider than segment 2, segment 1 about twice, segment 2 more than three times as long as wide, segments 3���5 as long as wide; club elongate, about 2.8 �� as long as wide, of almost 0.9 funicle length, otherwise as in C. ganglbaueri. Pronotum: dark, moderately wider than long (Pl/Pw 0.71), texture and vestiture as in C. ganglbaueri; widest at base, in basal half moderately subconically narrowed, then abruptly strongly conically narrowed to anterior margin, without constriction, in lateral view in basal part flat, then almost abruptly falling to anterior margin. Prosternum: anterior margin with sharply incised, deep emargination bounded by two shiny uneven tubercles, separated from coxae by narrow prosternal area. Scutellum: as in C. ganglbaueri. Elytra: black, in basal 2/3 moderately rounded, in apical third very broadly, evenly rounded, short (El/Ew 1.11), widest in mid-length, at base markedly broader than pronotum (Ew/Pw 1.76), humeri rounded, prominent, with shallow posthumeral impression; slightly convex on disc; interstria 1 at about 1/3 of medial length and in preapical area moderately broadened, interstria 2 at same lengths narrowed and constricted laterally encompassing medium-sized black tomentous maculae without differently colored scale border; interstriae except perimacular areas of equal width; odd interstriae as in C. ganglbaueri except for black patches being slightly more sparsely distributed; striae and entire surface as in C. ganglbaueri, but scales slightly thinner, more elongate, almost completely concealing integument. Venter: almost evenly densely covered with recumbent, strongly elongate scales and hair-like scales, without distinct clusters of scales; mesosternal process flat, broad, blunt at posterior margin, scaled, densely punctured; metasternum concave, densely transversally punctured, with a few indistinct transverse ribs; ventrites 1 and 2 with deep punctured impression; ventrite 1 1.6 �� as long as ventrite 2, ventrites 1���2 combined 3.6 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of approximately same length as ventrite 5. Legs: as in C. ganglbaueri except smaller profemoral teeth and femora covered with more evenly dense scales. Penis: Figs 45 d���f, its body medium long, parallel-sided, rounded at apex. Female. Rostrum longer (Rl/Pl 1.59), antennal insertion closer to rostrum mid-length, at 0.6 of rostrum length. Ventrites 1 and 2 without impression. Protarsal onychia slightly shorter, claws equally long. Variability. Length ♂♂ 3.86���4.67 mm, ♀♀ 4.19���5.24 mm. The vestiture can be seldom yellowish instead of grayish. The rostrum, especially in females, slightly varies in its length. Diagnosis. This species is recognizable by concealed elytral integument, alternating black patch pattern on odd interstriae, mostly grayish integument, elongate antennal club, short protarsal onychia in both sexes, and penis shape. Comparative notes. Cionus griseopubens is most closely related to C. ganglbaueri and C. neglectus. It differs from the former by bigger body size, antennal insertion closer to the rostrum mid-length, smaller dorsal macula, and penis shape, from the latter by subrotund elytra, and penis shape. Some specimens, especially yellowish colored, are reminiscent of C. olivieri, from which they differ by more distally inserted antennae and shorter protarsal onychia in both sexes, and by penis shape. Biological notes. Biology unknown. As closely related species C. ganglbaueri and C. colonnellii live on various Verbascum species, the same host plant genus is to be expected also in C. griseopubens. The species was collected at elevations 0���1,700 m a. s. l. from April to June. Distribution. Greece, Macedonia, Bulgaria. Non-type specimens examined. We examined 32 specimens. GREECE: Peloponnesos (Kardamyli 2 ♀♀ BN; Menalon Mt. 2 ♂ 1 ♀ ME, 1 ♀ PY, 2 ♂♂ 3 ♀♀ coll. Angelini; Chelmos Mts. 2 ♂ BN; Kandila 1 ♂ KO, 1 ♂ BMNH; Pirgos 1 ♂ WM; Sparta 1 ♀ KY; Kalavryta 1 ♂, AI, 1 ♀ TI; Arcadias, Menalon Mt. 2 ♂♂ 1 ♀ ME), Thessalia (M. Pieria 1 ♂ OA), West Macedonia (Florina 1 ♂ OA), Epiros (Igoumenitsa 1 ♂ OA), Makedonia (Falakron Mts. 1 ♂ WM, Thessaloniki 1 ♀ PY); MACEDONIA: Galičica (1 ♂ 2 ♀♀ BMNH); BULGARIA: Sandanski (1 ♂ BN), Rodopi, Mugla (1 ♀ ZS)., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 77-78, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Wingelmuller, A. (1914) Monographie der palaarktischen Arten der Tribus Cionini. Munchen, pp. 166 - 237.","Wingelmuller, A. (1921) Bestimmungstabelle der palaarktischen Cionini (Curculionidae) nebst Beschreibungen neuer Arten. Koleopterologische Rundschau, 9, 101 - 124.","Wingelmuller, A. (1937) Monografie der palaarktischen Arten der Tribus Cionini. Koleopterologische Rundschau, 23, 143 - 221.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp."]}

7. Cionus armeniacus sp. n. Figs 7 a���f. http://zoobank.org/ urn:lsid:zoobank.org:act: ECB6C4B6-7BD2-4F1F-B2F1-5D9683191F6C Type locality. Vayots pr. Getap (Armenia). Type series. Holotype: well-preserved, 3.34 mm long male ��� ARMENIA, Vayots Dzor pr. Getap, 21.5.2016 39��46��16.43˝ N 45��18��53.61˝ E Petr Kresl leg. alt. 1140 m / HOLOTYPUS Cionus armeniacus sp. n. M.Ko��ť��l et R.Caldara des. 2017 [printed red label]��� (NMPC). Description. Male (holotype). Clearly suboval. Head: rostrum moderately stout, medium long (l/ w 4.8, Rl/Pl 1.42), blackish-brown; in lateral view evenly slightly curved except concave dorsal outline of its apical part, at antennal insertion very slightly broadened at lower outline, basal part same width, apical part moderately tapered to apex; in dorsal view moderately broadened to antennal insertion, then moderately narrowed to apex, in basal third laterally constricted, apical part moderately dorsoventrally flattened; basal part very densely, longitudinally punctured to finely ribbed, in anterior half of apical part nearly without punctures, shiny; basal part semidensely covered with mostly backwardly oriented, subrecumbent blackish short scales, apical part with forwardly oriented, subrecumbent to suberect, whitish and yellowish long seta-like scales. Head between eyes narrow, of 0.4 rostrum width at base. Eyes large, rounded, very slightly protruding from head outline. Antennae reddish, inserted at 2/3 of rostrum length; funicle of 0.7 of scape length, segment 1 moderately wider than segment 2, of 0.7 its length, segments 1 and 2 less than twice as long as wide, segments 3���5 subglobose; club strikingly elongate, 2.8 �� as long as wide, subcylindrical, 1.6 �� as long as funicle, completely covered with recumbent thin whitish hairs and several erect long whitish sensilla. Pronotum: dark brown, markedly wider than long (Pl/Pw 0.59), densely evenly punctured, punctures small, subrotund, similar in size; in median part sparsely, at sides densely covered with mostly medially oriented, subrecumbent, elongate (l/w 4���6), whitish, yellow and orange intermixed scales, in lateral view yellow scales clustered into two parallel longitudinal stripes reaching from pronotal base to 3/4 of pronotal length; widest at base, conically narrowed to half of its length, then concavely constricted to narrow subparallel anterior neck, in lateral view in basal half flat on disc, then moderately falling to anterior margin. Prosternum: anterior margin with deep, sharply incised emargination bounded by two tiny tubercles, not reaching coxae. Scutellum: triangular, bluntly rounded at apex, covered with short, backwardly oriented suberect black scales, punctured. Elytra: dark brown, until 0.7 of their length subparallel to slightly rounded, in apical part broadly evenly rounded, moderately elongate (El/Ew 1.27), widest in mid-length, at base markedly wider than pronotum (Ew/Pw 1.72), humeri bluntly broadly rounded but visibly prominent, with shallow posthumeral impression, in lateral view until half of their length flat on disc, then evenly moderately rounded to apex; interstria 1 from about 1/4 to half of medial length very strikingly, in preapical part in much less extent broadened, interstria 2 at same lengths markedly narrowed, in dorsal part very strikingly, in preapical part moderately constricted laterally encompassing large, evenly round dorsal and somewhat oblong preapical black tomentous maculae, dorsal macula sparsely surrounded by reddish elongate scales, with indistinct patch of whitish scales at posterior margin, preapical macula with larger distal and smaller proximal patches of reddish scales, odd interstriae broader than even ones, convex, very finely textured; odd interstriae with alternating larger black and smaller whitish patches of scales; striae relatively deep, formed by single rows of round, large, deep punctures; entire integument semidensely covered with recumbent to subrecumbent, thin, moderately elongate (l/w 4���8) scales leaving majority of integument visible. Venter: mesosternum, metasternum laterally, metepisternum entirely covered with densely arranged recumbent yellow and orange intermixed elongate scales, ventrites and median longitudinal part of metasternum with recumbent thin, relatively short whitish hair-like scales, margins of ventrites 3 and 4 with small clusters of reddish and whitish, shortly elongate scales; mesosternal process flat, subtriangular, blunt at posterior margin, finely punctured; metasternum flat to concave in midline, unevenly transversally, very densely punctured, ventrite 1 with moderate median impression, ventrite 2 flat, both ventrites with similar punctation as metasternum; ventrite 1 1.7 �� as long as ventrite 2, ventrites 1���2 combined 4.4 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.7 length of ventrite 5. Legs: brown, tarsi lighter, metafemora darker; profemora with blunt, medium-sized teeth, meso- and metafemora with large triangular sharp teeth, profemoral teeth with small cluster of erect elongate yellowish scales; femora and tibiae with suberect to subrecumbent whitish, yellowish and black elongate scales, tarsomeres 1���3 with suberect whitish hair-like scales, onychia sparsely covered with recumbent whitish long hairs, femora with unevenly indistinct transverse bands formed by whitish scales; onychia of anterior legs of normal length, of 0.7 length of tarsomeres 1���3 combined, protarsal tarsomere 3 slightly wider than long; lateral protarsal claws by approximately one third shorter than medial ones, clearly separated, medial claws of meso- and metatarsi slightly but visibly shorter than lateral ones. Penis: Figs 7 d���f, its body with subparallel sides, bluntly tapered at apex. Female. Unknown. Variability. We know only the holotype of this species. Diagnosis. This species is characterised by small size, long antennal club, partially well visible elytral integument, round and large dorsal macula, lateral pronotal longitudinal yellow stripes and lateral posthumeral yellow patch. Comparative notes. This species is most closely related to C. caucasicus and C. ponticus, with which it shares the small size and the elytral pattern (except for dorsal macula), and from which it clearly differs by elongate antennal club, large round dorsal macula, and shorter body of penis. Biological notes. P. Kresl (pers. comm.) collected this species in Armenia in a warm steppe-forest. Host plant unknown. Distribution. Armenia. Etymology. The species was named after its only known country of distribution. Non-type material examined. None., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 22-23, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117

24. Cionus tamazo K��no, 1930 Figs 24 a���f. Cionus tamazo K��no, 1930: 149. Zumpt, 1937: 222. Caldara, 2013: 124. Alonso-Zarazaga et al., 2017: 186. Type locality. Hokkaido (Japan). Type series. K��no described the species based on 16 specimens collected on Kuriles Islands, Hokkaido and Honshu without designating the holotype. In coll. K��no (EIHU), there is a well-preserved pinned male, which was photographed by S. Sejima and designated by Sejima and Yoshitake as the lectotype of C. tamazo K��no. We have seen high resolution photographs of the specimen and its labels. Pictures (dorsal and lateral) sufficiently allow to identify the specimen and to confirm the identification of our material. The lectotype is labeled ��� Hokkaido H. Kono / ♂ / Cionus tamazo K��no Type [red label] / SYNTYPE The label attached by OHARA and HIRONAGA 2006 / 0000000805 Sys. Ent Hokkaido Univ. Japan [SEHU] / [LECTOTYPE] ♂ Cionus tamazo K��no, 1930 des. Shouma SEJIMA & Hiraku YOSHITAKE, 2017���. Before the present designation, there was no formal recognition of this status. At the moment of preparation of our manuscript this designation was not yet published. The specimen proposed for the lectotype is conspecific with C. tamazo K��no as currently understood. We based the following redescription on other Japanese specimens undoubtedly conspecific with this specimen. Synonyms. None. Redescription. Male. Body medium stout, subrotund. Head: rostrum moderately slender, medium long (l/ w 5.6, Rl/Pl 1.23), black; in lateral view slightly evenly curved, same width from base to antennal insertion, then markedly tapered to apex; in dorsal view slightly broadened from base to antennal insertion, then parallel or very slightly narrowed to apex, basal part considerably laterally constricted, apical part dorsoventrally flattened; basal part very densely, longitudinally punctured with a few ribs, apical part semidensely punctured, shortly before apex smooth, shiny; basal part covered with up- and backwardly oriented, recumbent, small, thin, elongate whitish scales, apical part with subrecumbent to suberect, forwardly oriented longer hair-like scales. Head between eyes and eyes as in C. hortulanus. Antennae reddish-brown, club somewhat darkened, inserted at 0.65 of rostrum length; funicle of less than 0.7 scape length, segment 1 slightly wider than segment 2, segment 1 twice, segment 2 three times as long as wide, segments 3���4 as long as wide, segment 5 moderately transverse; club elongate, about three times as long as wide, of same length as funicle, completely covered with recumbent, tiny brownish hairs and sparse erect, longer, light brown sensilla. Pronotum: black, somewhat wider than long (Pl/Pw 0.66), finely evenly densely punctured, punctures round to subrotund, spaces between punctures equal to or smaller than puncture diameter; covered with unequally densely distributed, forwardly and medially oriented, recumbent to subrecumbent, elongate (l/w 4���8) yellowish scales; widest at base, then almost evenly conically narrowed to anterior margin, without apparent constriction, in lateral view almost flat in basal half, then falling to anterior margin. Prosternum: as in C. hortulanus. Scutellum: as in C. hortulanus. Elytra: black, in anterior 2/3 slightly rounded to subparallel, in apical third very broadly rounded, short (El/Ew 1.13); widest before half of their length, at base markedly wider than pronotum (Ew/Pw 1.70), humeri rounded, prominent, with only indistinct posthumeral impression; moderately convex on disc; interstria 1 from about 1/4 to 1/2 of medial length distinctly broadened, in preapical area slightly broadened, interstria 2 at same lengths on dorsum narrowed and constricted laterally, in preapical area of unchanged width encompassing medium large dorsal and preapical subrotund, black tomentous maculae without differently colored scale border; interstriae except perimacular areas of approximately equal width; odd interstriae with strikingly contrasting alternating black and yellowish patches of scales; striae shallow, formed by somewhat uneven single rows of densely arranged punctures; entire surface covered with semidensely arranged, recumbent, elongate (l/w 4���6), small yellowish scales concealing vast majority of integument. Venter: mesosternal process moderately narrow, without shallow incision at posterior margin, otherwise as in C. hortulanus. Legs: black, tarsi dark brown, onychia lighter, tibiae longer and slender, otherwise as in C. hortulanus. Penis: Figs 24 d���f, its body medium long, broadly rounded at apex. Female. Rostrum slightly longer (Rl/Pl 1.29), in distal half of apical part shiny, with very sparse tiny punctures, antennal insertion closer to mid-length, in 0.57 of rostrum length. Ventrites 1 and 2 without impression, convex. Claws equally long. Variability. Length ♂♂ 4.08���4.61 mm, ♀♀ 3.98���4.82 mm. Similarly to C. hortulanus, it is a very variable species in elytral pattern, color of antennae, length of club, shape of tibiae, and width of the body of penis. The number of white spots on odd interstriae varies considerably, but they are always present. The color of antennae varies from almost black to reddish-brown. In some specimens, tibiae at their protibio-femoral junction are bent inwardly. There are even specimens, where one protibia is bent and the other one straight. The body of penis varies rather considerably in its width. Ventral characters do not show this degree of variability. Diagnosis. Tis species is recognizable by hardly visible elytral integument, in lateral view apical part of rostrum in both sexes tapered, shiny in females, deep impression on ventrites 1 and 2 in males, and penis shape. Comparative notes. This species is mostly related to C. hortulanus from which it differs by elongate antennal club, black elytra, striking elytral ���black and white��� pattern, darker to black tarsi and usually evenly conically narrowed pronotum. Biological notes. Chuj�� collected this species in Kuwadaira (Shikoku, Japan) on Paulownia tomentosa Kanitz. Distribution. This is a very widely distributed species. Japan (Hokkaido, Honshu, Kyushu, Shikoku), South Korea, North Korea, Russian Federation (Primorie, Kuril Islands, South-Eastern Siberia), eastern China (Fujian, Nei Mongol, Yunnan). Non-type specimens examined. We examined 56 specimens from the above countries. CHINA: Fujian, Shaowu. JAPAN: Chuzenji Lake, Kuwadaira, Nikko, Shikoku, Tochigi. SOUTH KOREA: Seishin., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 45-46, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Kono, H. (1930) Langrussler aus dem japanischen Reich. Insecta Matsumurana, 4, 145 - 162.","Zumpt, F. (1937) Curculioniden-Studien XXVI. Erganzungen zur \" Monographie der palaarktischen Arten der Tribus Cionini \" von Alois Wingelmuller in Wien. Koleopterologische Rundschau, 23, 222 - 228.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp."]}

48. Cionus longicollis C. N. F. Brisout de Barneville, 1863 Figs 48 a���f. Cionus longicollis C. N. F. Brisout de Barneville, 1863:114. Reitter, 1904: 55. Wingelm��ller, 1914: 194; 1921: 112; 1937: 175. Hustache, 1932: 342. A. Hoffmann, 1958: 1220. Caldara, 2013: 123. Alonso-Zarazaga et al., 2017: 185. Type locality. Vernet Valley (eastern Pyrenees, France). Type series. This species was described based on more specimens collected in Vernet valley, Eastern Pyrenees. In coll. C. N. F. Brisout (MNHN), there are a couple on a single pin and two males on another pinned card. A 4.75 mm long, well-preserved male from the card with a couple labeled ��� longicollis Vernet / Mus��um Paris S. E. F. Coll. Ch. Brisout��� was designated as the lectotype by adding the label ��� LECTOTYPUS Cionus longicollis C. Brisout ♂ Michael Ko��ť��l des. 2014 [printed red label]���. Two males and one female with the same labels were labeled as paralectotypes accordingly. Synonyms. None. Redescription. Male. Body stout, subparallel. Head: rostrum moderately stout, medium long (l/ w 5.0, Rl/Pl 1.30), black except very apical part brown; in lateral view slightly curved, indistinctly enlarged at antennal insertion, of approximately same width from base to shortly before apex, at apex dorsal side in same line evenly curved to apex, ventral side abruptly beveled upwards to apex; in dorsal view of same width from base to apex, in basal part round in cross-section, in apical part slightly dorsoventrally flattened; in basal and proximal apical part very densely, moderately longitudinally punctured, at apex punctures sparser, well separated, with small shiny area; basal part with up- and backwardly oriented, subrecumbent, elongate whitish and light brown intermixed scales, apical part with subrecumbent to suberect, forwardly oriented, yellowish, long seta-like scales. Head between eyes narrow, of 0.4 rostrum width at base. Eyes large, moderately rounded, not protruding from head outline. Antennae reddishbrown except moderately darkened club, inserted between 0.6 and 0.7 of rostrum length; funicle of 3/4 scape length, segment 1 wider than segment 2, segment 1 twice, segment 2 about 2.5 �� as long as wide, segments 3���5 as long as wide; club spindle-shaped, 2.7 �� as long as wide, of funicle length, completely covered with recumbent brownish and yellowish thin hairs and sparse erect, relatively short light brown sensilla. Pronotum: black, moderately wider than long (Pl/Pw 0.73), densely evenly punctured, punctures medium-sized, subrotund, approximately of equal size, spaces between punctures smaller than puncture diameter; covered with somewhat unevenly distributed, variously oriented, recumbent and subrecumbent, elongate (l/w 5���8), yellowish and on anterior margin very sparse blackish scales; widest at base, subparallel to indistinctly convergent in basal half, then slightly rounded and conically narrowed to anterior margin, without constriction, in lateral view flat in basal half, then abruptly strongly falling to anterior margin. Prosternum: anterior margin with relatively shallow, semicircular, sharply incised emargination, anteriorly bounded by shiny tubercles, separated from coxae by very narrow strip of prosternum. Scutellum: subtriangular with rounded apex, covered with backwardly oriented, subrecumbent to suberect scales, very densely punctured to rugulose. Elytra: black to brown, in basal 2/3 subparallel, in apical third moderately rounded, then broadly rounded at apex, moderately elongate (El/Ew 1.25), widest at about half of their length, at base moderately wider than pronotum (Ew/Pw 1.53), humeri subrotund, moderately prominent, with only indistinct posthumeral impression; almost flat on disc; interstria 1 from approximately 1/4 to more than 1/2 of medial length and in preapical area very strongly broadened, interstria 2 at same lengths very strongly narrowed to concealed, constricted laterally, interstria 3 at same lengths moderately constricted laterally encompassing very large round black tomentous maculae with narrow border formed by gingery scales; interstriae except perimacular areas of approximately equal width; odd interstriae with unevenly distributed, on dorsum indistinct, on sides clearly visible patches of whitish scales, patches of black scales very sparse, indistinct; striae very shallow, formed by even rows of large shallow round, very densely arranged punctures; entire surface covered with slightly unevenly, densely distributed, recumbent and subrecumbent, elongate (l/w 4���8) whitish and sparsely intermixed gingery scales of unequal width almost fully concealing integument. Venter: semidensely covered on sides with broader scales, in median part with very thin to hair-like elongate scales, only indistinctly denser on metepisternum and margins of ventrites 3���4; mesosternal process flat, subquadrate, blunt at posterior margin, with hairs, very densely punctured; metasternum very slightly concave to flat, transversally ribbed; ventrites 1 and 2 with relatively deep and broad impression, in proximal part rugulose, in distal part punctured; ventrite 1 1.5 �� as long as ventrite 2, ventrites 1���2 combined 5.3 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.7 length of ventrite 5. Legs: brown to dark brown, tarsi lighter, profemora thick, with small teeth emphasized by erect scales, meso- and metafemora with large subtriangular sharp teeth; femora covered with recumbent to subrecumbent whitish and scattered gingery scales, tibiae covered with subrecumbent to suberect, whitish and sparse black elongate scales, tarsi covered with suberect whitish hair-like scales, onychia with recumbent, long, thin whitish hairs, on femora scales clustered into indistinct transverse bands; protarsal onychia of normal length, as long as tarsomeres 1���3 combined, protarsal tarsomere 3 wider than long; protarsal lateral, meso- and metatarsal medial claws of about 0.6 length of their pair-claws, clearly thinner. Penis: Figs 48 d���f, its body long, parallel-sided, broadly tapered to rounded at apex. Female. Rostrum considerably longer (Rl/Pl 1.52), apical part subparallel to very slightly concave in its midlength, antennae inserted at half of rostrum length. Ventrites 1 and 2 without impression, convex. Claws of approximately equal length. Variability. Length ♂♂ 3.89���4.78 mm, ♀♀ 4.30���5.25 mm. This species shows no special degree of variability except for the body size. In some specimens, the dorsal elytral macula can be slightly longitudinally elongate. Diagnosis. This species is recognizable by almost concealed elytral integument, subparallel elytra, very large dorsal and large preapical elytral maculae with border of gingery scales, relatively stout rostrum, in lateral view of same width from base to shortly before apex, and relatively deep and broad impression on ventrites 1 and 2 in males. Comparative notes. Cionus longicollis is most closely related to C. montanus and C. atlanticus. It differs from former by the presence of a border of gingery scales around both elytral maculae, and from latter by larger body size and large round dorsal elytral macula. Biological notes. The first author collected this species in Spain (Castilla-Le��n, Besande) in series on a large Verbascum sp. resembling V. densiflorum. Hoffmann (1958) reported V. lychnitis as the host plant which is unlikely due to the first author��s observations. Distribution. This species is widespread in the European part of West Mediterranean from southern France (Alps, Provence, Garonne, Pyren��es) to Spain (Catalu��a, Castilla y Le��n, Navarra, Aragon, Andaluc��a: Puerto de la Ragua). We did not examine specimens from Portugal, and know only one record from southern Spain. Countries and areas of distribution like western Siberia, Finland, Germany etc. (Caldara, 2013) as well as record from Great Britain by Morris (2012) are surely based on data concerning C. montanus, which was previously regarded as a subspecies of C. longicollis. Non-type specimens examined. We examined 105 specimens from the above mentioned regions. FRANCE: Abries, Ariege, Eastern Pyrenees, Embrun, Hospice de France, Vernet; SPAIN: Andalusia, Aragon, Cantabria, Castilla-Le��n (Besande), Catalunya, Girona (La Molina), Huesca (Torla), Lerida, Navarra., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 81-83, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Brisout de Barneville, C. N. F. (1863) In: Grenier, A. J. F., Catalogue des coleopteres de France et materiaux pour servir a la faune des coleopteres francais, par MM. E. Allard, Dr. Ch. Aube, Ch. Brisout de Barneville, A. Chevrolat, L. Fairmaire, Al. Gauvel, Dr. A. Grienier, Dr. Kraatz, J. Linder, L. Reiche et Felicien de Saulcy. A. J. F. Grenier, Paris, pp. 114.","Reitter, E. (1904) Bestimmungs-Tabelle der Coleopteren-Gattung Cionus Clairv. aus Europa und angrenzenden Landern. Wiener Entomologische Zeitung, 23, 47 - 64. https: // doi. org / 10.5962 / bhl. part. 27188","Wingelmuller, A. (1914) Monographie der palaarktischen Arten der Tribus Cionini. Munchen, pp. 166 - 237.","Wingelmuller, A. (1921) Bestimmungstabelle der palaarktischen Cionini (Curculionidae) nebst Beschreibungen neuer Arten. Koleopterologische Rundschau, 9, 101 - 124.","Wingelmuller, A. (1937) Monografie der palaarktischen Arten der Tribus Cionini. Koleopterologische Rundschau, 23, 143 - 221.","Hustache, A. (1932) Curculionidae gallo-rhenans. Annales de la Societe Entomologique de France, 100, 153 - 470. [1931]","Hoffmann, A. (1958) Faune de France 62 Coleopteres curculionides. Troisieme Partie. Lechevalier, Paris, pp. 1209 - 1839.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp.","Morris, M. G. (2012) True Weevils 3: Coleoptera: Curculionidae (Curculioninae, Baridinae, Orobitinae) (Handbooks for the Identification of British Insects 5 / 17 d). Royal Entomological Society, London, 136 pp."]}

28. Cionus leonhardi Wingelm��ller, 1914 Figs 28 a���f. Cionus leonhardi Wingelm��ller, 1914: 212; 1921: 119; 1937: 193. Smreczyński, 1976: 59. Caldara, 2013: 123. Alonso-Zarazaga et al., 2017: 185. Cionus inexspectatus Temp��re, 1961: 98. Caldara, 2013: 123. Alonso-Zarazaga et al., 2017: 185. syn. n. Type locality. Oberweiden (Austria). Type series. In coll. Wingelm��ller (NHMW), there are a male of Cionus labeled ���Marchfeld Wingelm. / ♂ / Leonhardi m. det. Wingelm��ll. ��� and a female with the same data except for ��� ♀ ���. In the original description, one of localities is reported as ���Oberweiden��� in Lower Austria. This place is situated in Marchfeld area, therefore both specimens are syntypes. The male was remounted, dissected for the genitalia and designated as the lectotype of Cionus leonhardi Wingelm��ller by adding the printed red label ��� LECTOTYPUS Cionus leonhardi Wingelm��ller M. Ko��ť��l et R. Caldara des. 2011���. The lectotype is perfectly preserved, 3.64 mm long, the penis is slightly damaged on its side, but the typical apex is preserved. The other specimen was labeled as paralectotype. In the collections of SDEI, there are 17 specimens labeled as ��� Syntypus ��� of C. leonhardi Wingelm��ller. Fourteen (4 ♂♂, 10 ♀♀) specimens come from ���Kronstadt��� (modern Brasov, Romania), a couple from ���Bulg. Rhodope Geb.��� and one male from ���Bulg. Rila Kloster���, both last localities in Bulgaria. All these 17 specimens were labeled as paralectotypes of Cionus leonhardi Wingelm��ller by the printed red label ��� PARALECTOTYPUS Cionus leonhardi Wingelm��ller Michael Ko��ť��l des. 2012���, and all belong to the taxon as currently understood. Lastly, in the collections of MTD, there is a male labeled ���Siebenb��rgen Kronstadt / Leonhardi Wingelm. det. / PARATYPE / Samml. K. A. Penecke Geschenk 1940.20 / Staatliche Museum f��r Tierkunde Dresden���. This male was also labeled as paralectotype by adding the printed red label ��� PARALECTOTYPUS Cionus leonhardi Wingelm��ller Michael Ko��ť��l des. 2014���. Synonyms. Cionus inexspectatus was described based on several specimens from French eastern Pyrenees (Corsavy, Targassone) referring to its similarity to C. nigritarsis but differing by its peculiar penis shape (���P��nis tr��s remarquable...���). In the collection Temp��re (MNHN), there is a single 3.87 mm long, well-preserved male specimen with dissected genitalia labeled ���PYR��N��ES ORIENTALES Targasonne 27-7-57 G. TEMPERE / C. inexspectatus Temp. HOLOTYPE [red label] / MUSEM PARIS 1984 Coll. G. TEMPERE���. This specimen has darker integument and more striking dark spots on elytra, but in all important characters corresponds entirely in habitus and penis to the lectotype of C. leonhardi. We added an identification label ��� Cionus leonhardi Wingelm. M. Ko��ť��l det. 2014��� accordingly. Redescription. Male. Body medium stout, suboval. Head: rostrum medium stout, medium long (l/ w 3.5, Rl/Pl 1.17), brown; in lateral view slightly unevenly curved, with convexity on upper outline shortly before antennal insertion, of approximately same width from base to convexity, at antennal insertion rather abruptly narrowed and then almost of same with to shortly before apex; in dorsal view same width from base to antennal insertion, then slightly narrowed to apex, basal part in cross-section round, apical part dorsoventrally flattened; confluently longitudinally punctured, at apex punctures densely arranged; basal part with up- and backwardly oriented, recumbent yellowish scales, apical part with forwardly oriented, subrecumbent whitish longer scales. Head between eyes narrow, of 0.4 rostrum width at base. Eyes large, rounded, not protruding from head outline. Antennae reddish-brown, inserted beyond 0.7 of rostrum length; funicle of 2/3 scape length, segment 1 moderately wider than segment 2, segment 1 less than twice, segment 2 twice as long as wide, segment 3 slightly longer than wide, segments 4���5 as long as wide; club spindle-shaped, slightly more than twice as long as wide, completely covered with recumbent, thin yellowish-gray to reddish hairs and sparse long erect pale sensilla. Pronotum: black, markedly wider than long (Pl/Pw 0.60), very densely, finely evenly punctured, punctures subrotund, slightly unequal in size, spaces between punctures smaller than puncture diameter; covered with evenly distributed, variously oriented, recumbent, elongate (l/w 4���8) yellow scales; widest at base, in basal half moderately conically, in anterior half more distinctly conically narrowed to anterior margin, without constriction, in lateral view flat in basal half, then moderately falling to anterior margin. Prosternum: anterior margin with sharply incised semicircular to subquadrate emargination, bounded by two indistinct tubercles, separated from coxae by narrow prosternal area. Scutellum: subtriangular with blunt apex, covered with scales similar to those on elytra, punctured. Elytra: dark brown to reddish-brown, in basal 2/3 slightly rounded to subparallel, in apical part broadly rounded, moderately elongate (El/Ew 1.20), widest at about 1/4 of their length, at base markedly wider than pronotum (Ew/Pw 1.73), humeri subrotund, slightly prominent, without visible posthumeral impression; slightly convex on disc; interstria 1 at 1/3 of medial length and in preapical area moderately broadened, interstria 2 at same lengths moderately narrowed and constricted laterally encompassing subrotund to irregular black tomentous maculae without differently colored scale border; interstriae except perimacular areas of approximately same width; odd interstriae, especially in posterior half, with unevenly distributed, large black scale patches, and only indistinct alternating patches of clustered yellow scales; striae very shallow, formed by even single rows of very densely arranged, large punctures; entire surface covered with recumbent and subrecumbent, elongate (l/w 4���6) yellow scales, often truncated distally, almost completely concealing integument. Venter: densely, evenly covered with recumbent, light yellow, elongate, variously oriented thick scales; mesosternal process flat, broad, blunt at posterior margin, densely scaled, punctured; metasternum moderately concave, punctured, at posterior margin transversally ribbed, ventrite 1 with deep, ventrite 2 with shallow impression, punctured; ventrite 1 2.1 �� as long as ventrite 2, ventrites 1���2 combined 5.3 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.6 length of ventrite 5. Legs: reddish-brown to brown, profemora with very small teeth emphasized by erect scales, meso- and metafemora with large sharp subtriangular teeth; femora covered with recumbent yellow scales feebly clustered into indistinct transverse bands, tibiae covered with subrecumbent yellowish and whitish elongate scales, tarsomeres 1���3 with suberect whitish seta-like scales, onychia with recumbent, thin whitish hairs; protarsal onychia long, 1.2 �� as long as tarsomeres 1���3 combined, tarsomere 3 as long as wide, with narrow lobes; protarsal lateral, meso- and metatarsal medial claws by approximately 1/3 shorter than their pairclaws, markedly thinner. Penis: Figs 28 d���f, its body long and thin, with slightly sinuate sides, at apex extended into long thin downwards hooked process. Female. Rostrum markedly longer (Rl/Pl 1.50), black to dark brown, antennal insertion between 0.5 and 0.6 of rostrum length. Ventrites 1 and 2 without impression. Claws of approximately equal length. Variability. Length ♂♂ 3.42���3.89 mm, ♀♀ 3.59���4.08 mm. This species does not show remarkable variability. The shape and size of both elytral maculae, the elytral pattern on odd interstriae and the brightness of yellow scales on the entire body vary to some extent. In some specimens, the antennal club is markedly darkened. Diagnosis. Cionus leonhardi is recognizable by almost concealed elytral integument, antennal insertion in males closer to apex, at distinctly more than 0.6 of rostrum length, thin tarsi, narrow lobes of tarsomere 3, long protarsal onychium in males, and penis shape. Comparative notes. Cionus leonhardi is most closely related to C. hypsibatus and C. donckieri. It differs from the former by the more or less distinct elytral pattern, and the narrow lobes of tarsomere 3, from the latter by the lack of elytral erect seta-like scales. Biological notes. The first author collected this species in Hungary in primeval sandy steppe on Verbascum lychnitis. Smreczyński (1976) reported this plant and V. chaixii austriacum as host plants. Distribution. This species is widespread in southern part of Europe except for Portugal, reaching from Spain and France across Germany to east Ukraine and Asian part of Turkey. It does not occur in Poland and northwards. Dedyukhin (2014) reported also central European Russia and Wingelm��ller (1914) also Syria without detailed locality, hence, it is not clear if this record concerns the current territory of Syria or modern Turkey (Akbes). Non-type specimens examined. We examined more than 170 specimens from all above mentioned countries and regions except Russia., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 51-53, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Wingelmuller, A. (1914) Monographie der palaarktischen Arten der Tribus Cionini. Munchen, pp. 166 - 237.","Wingelmuller, A. (1921) Bestimmungstabelle der palaarktischen Cionini (Curculionidae) nebst Beschreibungen neuer Arten. Koleopterologische Rundschau, 9, 101 - 124.","Wingelmuller, A. (1937) Monografie der palaarktischen Arten der Tribus Cionini. Koleopterologische Rundschau, 23, 143 - 221.","Smreczynski, S. (1976) Ryjkowce-Curculionidae: Podrodzina Curculioninae. Klucze do Oznaczania Owadow Polski, 87 (XIX, 98 f), 1 - 115.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp.","Tempere, G. (1961) Sur quelques Curculionidae de la faune francaise (Col.). Cinquieme note. Bulletin de la Societe Entomologique de France, 66, 94 - 100.","Dedyukhin, S. V. (2014) New data on the fauna and ecology of the weevils (Coleoptera, Curculionoidea) Vyatka-Kama region and Middle Preduralye. Vestnik Udmurtskovo Universiteta, 1, 73 - 84."]}

35. Cionus boroveci sp. n. Figs 35 a���f. http://zoobank.org/ urn:lsid:zoobank.org:act: 99487B25-DA0E-4F1B-8F9A-95658917B9C4 Type locality. Omalos (Crete, Greece). Type series. Holotype: a completely preserved, 3.37 mm long male ��� CRETA occ. Lefka Ori Mts. M. Ko��ť��l leg. / Omalos 5 km S Gingilos Mt. 1300 m 13.vii.1992 / HOLOTYPUS Cionus boroveci sp.n. M.Ko��ť��l et R.Caldara des. 2012 [printed red label]��� (NMPC). Paratypes: the same labeling as holotype but instead of ��� HOLOTYPUS ��� ��� PARATYPUS ��� (1 ♂, 1 ♀ KO, 1 ♂ CA); ��� CRETE occ. Lefka ��ri Mts. Omalos-env., 4 km S 11.4.1990 1400 m R.Borovec lgt. / Cionus n.sp. pr. ungulatus R.Borovec det. 1991��� (1 ♂ BO). Description. Male. Body medium stout, subparallel. Head: rostrum moderately stout, medium long (l/ w 4.67, Rl/Pl 1.33), dark brown; in lateral view evenly curved, same width from base to antennal insertion, then moderately tapered to apex; in dorsal view slightly broadened to antennal insertion, then parallel to apex, in most basal part slightly laterally constricted, in about middle third in cross-section nearly round, beyond antennal insertion flattened dorsoventrally; basal part until antennal insertion very densely to confluent longitudinally punctured, apical part with round, sparsely distributed punctures, at apex glabrous, shiny; basal part covered with subrecumbent, backwardly oriented whitish and light brownish intermixed moderately elongate scales, apical part with suberect, forwardly oriented, long whitish and light brownish seta-like scales. Head between eyes narrow, of about 0.3 rostrum width at base. Eyes large, evenly rounded, not protruding from head outline. Antennae reddish-brown, inserted at 0.7 of rostrum length; funicle of about 0.7 scape length, segment 1 wider than, and as long as segment 2, segment 1 twice, segment 2 more than twice as long as wide, segments 3���5 subquadrate; club oval, about 2.4 �� as long as wide, of 0.8 funicle length, densely covered with recumbent, tiny, thin yellowish to whitish setae and sparse long thin whitish seta-like sensilla. Pronotum: reddish-brown, with darkened anterior margin, markedly wider than long (Pl/Pw 0.61), very densely, unevenly punctured, covered with subrecumbent, on disc forwardly oriented, reddish -yellow and white intermixed, elongate (l/w 5���7) scales and very few shorter, suberect black seta-like scales; widest at base, until half of length subparallel, in anterior half conically narrowed to anterior margin, very slightly convex on disc. Prosternum: anterior margin with marked, sharply incised subrotund emargination not reaching coxae. Scutellum: black, triangular, covered with evenly distributed, recumbent, backwardly oriented, small thin whitish and sporadic reddish scales. Elytra: reddish-brown, subparallel in anterior 2/3 of their length, then broadly rounded, moderately elongate (El/Ew 1.24), widest approximately at half of their length, at base markedly wider than pronotum (Ew/Pw 1.78), humeri prominent, broadly rounded; moderately convex on disc; interstriae convex, except perimacular areas of equal width, interstria 1at 1/3 of medial length and in preapical area strongly broadened, interstria 2 at same lengths narrowed and constricted laterally encompassing round, black tomentous maculae with border formed by sparsely arranged, reddish to gingery scales, dorsal macula larger than preapical one; odd interstriae with unevenly distributed, small, indistinct black and whitish alternating patches of scales; striae shallow and narrow, formed by single rows of subrotund shallow punctures; entire surface covered with semidensely arranged recumbent to subrecumbent, relatively short elongate (l/w 3���5), whitish and reddish scales, especially in posterior part, additionally with short suberect, clearly visible black seta-like scales concealing major part of integument. Venter: densely covered with subrecumbent, elongate, whitish scales, more densely arranged on lateral sides of ventrite 5, sides of metasternum with intermixed yellow elongate scales being on metepisternum very densely arranged to overlapping; mesosternal process short, straightly truncated at apex; metasternum slightly concave, with very densely arranged, transverse elongate punctures forming small ribs, at posterior margin of metasternum small bare round impression; ventrite 1 and 2 with deep median longitudinal impression; ventrite 1 1.5 �� as long as ventrite 2, ventrites 1���2 combined 5.4 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.6 length of ventrite 5. Legs: brown to reddish-brown, onychia at apices slightly darkened; profemora with forwardly protruding small triangular sharp teeth, meso- and metafemora with large, triangular sharp teeth; entire legs covered with recumbent to subrecumbent (on tarsi and tibial apices suberect), whitish, strongly elongate, thin seta-like scales; protarsal onychia long, of same length as tarsomeres 1���3 combined; protarsal lateral claws small and thin, meso- and metatarsal medial claws of about 2/3���3/4 length of their pair-claws. Penis: Figs 35 d���f, its body similar to that in C. thapsus. Female. Rostrum longer (Rl/Pl 1.53), antennae inserted at 0.6 of rostrum length. Ventrites 1 and 2 without impression. Protarsal onychia of normal length, of about 0.85 length of tarsomeres 1���3 combined. Claws of almost equal length. Variability. Length: ♂♂ 3.25���3.62 mm, ♀ 3.55 mm. The type series does not show considerable variability. In one specimen the legs, especially tarsi, are darker (it is a specimen collected in April i. e. overwintered). The reddish scale border, size and shape of both maculae vary from narrow, small and round to broad, larger and subrotund. Diagnosis. This species is characterised by reddish outlined maculae, small body size, in the ungulatus -group relatively short protarsal onychia, and body of penis similar to that of C. thapsus. Comparative notes. Cionus boroveci differs from the most closely related species C. balianii by clear reddish outline of elytral maculae, smaller body size, and completely different shape of penis. Distribution. Crete. Etymology. We dedicate the species to our friend Roman Borovec, who collected the first specimen more than 25 years ago, and already at that time supposed that the species might be new to science. Non-type specimens examined. None., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 61-63, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117

18. Cionus harani sp. n. Figs 18 a���f. http://zoobank.org/ urn:lsid:zoobank.org:act: 701CCEE1-9C2D-4A9A-B809-1EEC799432AF Type locality. Pushkar (Rajasthan, India). Type series. Holotype: well-preserved, 3.89 mm long dissected male with missing right mesotarsal onychium labeled ���PUSHKAR INDIA 16/X/2011 J. H. leg. / coll. J. Haran / HOLOTYPUS Cionus harani sp. n. M.Ko��ť��l et R.Caldara des. 2017 [printed red label]��� (MNHN). Paratype: same labeling as holotype except for ��� PARATYPUS ��� (1 ♀ HA). Description. Male (holotype). Body stout, subrotund to suboval. Head: rostrum moderately slender, medium long (l/ w 5.2, Rl/Pl 1.28), reddish-brown; in lateral view very slightly evenly curved, same width from base to antennal insertion, then moderately tapered to apex; in dorsal view same width from base to apex, basal part slightly laterally constricted, apical part moderately dorsoventrally flattened; texture and vestiture similar to that of C. hortulanus. Head between eyes and eyes as in C. hortulanus. Antennae reddish, inserted at 0.6 of rostrum length; funicle of 0.6 scape length, segment 1 slightly wider than segment 2, 1.5 �� as long as wide, segment 2 twice as long as wide, segments 3���5 subglobose to transverse; club oval, slightly more than twice as long as wide, slightly longer than funicle, almost completely covered with recumbent whitish hairs and sparse, relatively short whitish sensilla. Pronotum: brown, somewhat wider than long (Pl/Pw 0.69), texture and vestiture similar to that of C. hortulanus except punctures smaller and median part with semidensely distributed, subrecumbent elongate black scales; widest at base, moderately narrowed to half of its length, then conically narrowed to anterior margin, constricted as in C. hortulanus, in lateral view as in C. hortulanus. Prosternum: anterior margin with bluntly incised shallow semicircular incision bounded by flat shiny tubercles, separated from coxae by very narrow prosternal area. Scutellum: moderately convex, otherwise as in C. hortulanus. Elytra: reddish-brown, outline as in C. hortulanus, moderately elongate (El/Ew 1.16), widest in mid-length, at base strikingly wider than pronotum (Ew/Pw 1.79), humeri subrotund, apparently prominent, with slight posthumeral impression; almost flat on disc; interstriae moderately convex, otherwise texture and vestiture including black tomentous maculae as in C. hortulanus excepting large patches of black scales on humeri; entire surface covered with semidensely arranged, recumbent to subrecumbent, markedly elongate (l/w 6���10), whitish scales concealing majority of integument. Venter: slightly unevenly covered with recumbent, whitish elongate to hair-like scales, wider scales clustered especially on metepisterna, less strikingly on metasternum and ventrite sides; mesosternal process flat, broad, blunt at posterior margin, scaled, densely punctured; metasternum concave, transversally ribbed, ventrite 1 and 2 with deep, transversally ribbed to punctured impression; ventrite 1 1.7 �� as long as ventrite 2, ventrites 1���2 combined 6.5 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of half length of ventrite 5. Legs: reddish-brown; as in C. hortulanus except for all claws of approximately same length and profemora with more distinct transverse bands. Penis: Figs 18 d���f, its body shorter and wider, rounded at apex. Female. Rostrum longer (Rl/Pl 1.43), antennae inserted shortly beyond mid-length, apical part of rostrum with sparsely distributed tiny punctures, shiny. Ventrites 1 and 2 without impression. Claws equally long. Variability. Length ♂ 3.89 mm, ♀ 4.15 mm. The type couple does not show any variability. Diagnosis. This species is recognizable by almost completely concealed elytral integument, apical part of rostrum flattened in both sexes, in female long and shiny, medium large black tomentous maculae, deep impression on ventrites 1 and 2, antennae with oval club and short funicle, almost equally long claws in males, and penis shape. Comparative notes. Cionus harani is most closely related to C. hortulanus from which it differs by antennae with oval club and short funicle in both sexes, equally long claws and penis shape in males, punctured apical part of rostrum in females. Biological notes. Biology unknown. Distribution. India (Rajasthan). Etymology. The species is named after its collector and our colleague Julien Haran. Non-type specimens examined. None., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 37-38, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117

33. Cionus balianii F. Solari, 1932 Figs 33 a���f. Cionus balianii F. Solari, 1932: 88. Zumpt, 1937: 223. Caldara, 2013: 123. Alonso-Zarazaga et al., 2017: 185. Cionus galanus Angelov, 1978: 68. Dieckmann & Behne, 1994: 295 (syn. n.). Caldara, 2013: 123. Alonso-Zarazaga et al., 2017: 185. Type locality. Gran Sasso (Abruzzo, central Italy). Type series. According to the original description, C. balianii was described from several places in Italy. Gran Sasso is reported as ���loc. class.��� and the other localities in the original description are: Modena province (Emilia), Turlago (Massa Carrara province, Toscana) and S. Biase di Ceraso (Salerno province, Campania). In coll. Solari (MSNM), there are six specimens under the name C. balianii: a couple from ���Gran Sasso���, one female from ���Turlago���, a couple from ���Emilia Modenese��� and one male from ���S. Biase Ceraso���. The original description corresponds to all localities on labels. Therefore, we designated a male from Gran Sasso labeled ���Gran Sasso Abruzzo VII. 1894 A. Fiori / C. balianii m. typus! ♂ det. F. Solari��� as the lectotype of Cionus balianii F. Solari by adding the printed red label ��� LECTOTYPUS Cionus balianii F. Solari M.Ko��ť��l et R.Caldara des. 2016���. The lectotype is 4.07 mm long, perfectly preserved, with originally dissected penis and spiculum gastrale on a separate card. All five above mentioned specimens were labeled as paralectotypes accordingly. Synonyms. Angelov (1978) described C. galanus based on several specimens not giving the sex and the number of paratypes. In the original description, the holotype and the allotype were clearly indicated. The holotype comes from Wratza (Balkan Mts.) and the allotype from the same place. Additionally, the paratypes come from different localities in Bulgaria (Balkan Mts., Wratza, Twarditza, Rodopi Mts., Momtschilgrad, Parwenetz, Golo Bardo Mts., L��lin Mts., Kalotina, K��stendil, Boboschewo), Turkey (G��kceada) and Greece (Kephallenia, Korfu). In coll. Angelov (MZHF), there is one male with missing posterior leg from tibia including, with dissected genitalia labeled as ��� Holotypus ��� of C. galanus Angelov coming from ���Wraza���. Moreover, 16 paratypes (9 ♂♂, 7 ♀♀) are present in this collection. In the collections of MSDEI, there are six specimens labeled as ��� Paratypus ��� of C. galanus Angelov. One male is from ���Corfu���, other three specimens (2 ♂♂, 1 ♀) from ���Kephallenia���. All these four specimens were previously identified by Wingelm��ller as C. ungulatus. In the collections of MSDEI, there are two additional paratypes, a male from ���Wraza��� and a female from ���Twardiza���. Holotype and all paratypes are conspecific with the lectotype of C. balianii F. Solari. Redescription. Male. Body medium stout, suboval. Head: rostrum moderately stout, medium long (l/ w 4.6, Rl/Pl 1.22), dark brown; in lateral view upper margin at antennal insertion slightly abruptly curved, lower margin almost evenly curved, same width from base to antennal insertion, then visibly tapered to apex; in dorsal view very slightly broadened to antennal insertion, then parallel-sided, basal part very slightly laterally constricted, apical part beyond antennal insertion markedly dorsoventrally flattened; basal part evenly, moderately densely covered with subrecumbent, backwardly oriented, yellowish and light brown elongate scales, in distal part with suberect and erect, forwardly and laterally oriented, long whitish seta-like scales being very close to apex sparse, with bare shiny median area. Head between eyes narrow, of about 0.3 rostrum width at base. Eyes large, not protruding from head outline. Antennae reddish brown, inserted at 0.6 of rostrum length; funicle of about 0.7 scape length, segment 1 very slightly wider than, and as long as segment 2, segments 1 and 2 twice as long as wide, segments 3���5 as long as wide, subquadrate; club oval, about 2.1 �� as long as wide, of about 0.6 funicle length, covered with short, densely arranged, whitish and brownish setae and sparse erect whitish long sensilla. Pronotum: dark brown to reddish brown, somewhat wider than long (Pl/Pw 0.65), with very densely arranged, subrotund punctures; covered with unevenly densely arranged and variously oriented, subrecumbent to suberect, elongate (l/w 5���7) yellowish scales; widest at about 1/4 of its length, in basal half rounded, then unevenly narrowed to anterior margin, with indistinct broad constriction, convex on disc. Prosternum: anterior margin with rounded, sharply deep incised emargination not reaching coxae. Scutellum: subtriangular, at apex broadly rounded, at margins densely covered with recumbent, backwardly oriented whitish and intermixed brown elongate scales. Elytra: brown, in anterior 2/3 very slightly rounded, in posterior 1/3 broadly rounded, slightly elongate (El/Ew 1.15), widest at about 1/3 of their length, at base markedly wider than pronotum (Ew/Pw 1,74), humeri prominent, broadly rounded; very slightly convex to flat on disc; interstria 1 broadened and interstria 2 considerably narrowed and constricted laterally at about 1/3 of medial length and in preapical area encompassing black tomentous subrotund maculae with border formed by sparsely arranged, yellowish to gingery scales, interstriae of about equal width; odd interstriae with evenly distributed patches of yellowish and whitish-yellow elongate scales alternating with black oblong patches of subrecumbent scales, interstriae convex; striae shallow, formed by densely arranged punctures, stria 5 at humeri equally deep as other striae; entire surface, especially even interstriae, covered with subrecumbent, backwardly oriented, whitish, very thin seta-like scales (l/w 5���9) leaving less than half of integument visible, yellowish elongate scales, similar to those patched on pronotum, feebly thicker on humeral base, and more markedly densely arranged in posthumeral part of elytra not forming distinct macula. Venter: covered with recumbent to subrecumbent, elongate, whitish and yellowish seta-like scales and elongate, pale yellowish scales, latter ones thicker at sides forming small patches at lateral margins of ventrites 2���4; mesosternal process very short, almost straight at apex; metasternum with broad shallow longitudinal furrow, transversally ribbed; ventrite 1 with deep impression, ventrite 2 with shallow impression, semidensely covered with variously oriented, whitish long hairs; ventrite 1 2.1 �� as long as ventrite 2, ventrites 1���2 combined 5.2 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.6 length of ventrite 5. Legs: brown, onychia and distal part of tarsi lighter; profemora with small teeth, meso- and metafemora with large sharp teeth; femora covered with recumbent yellowish elongate scales, tibiae and tarsi except onychia with suberect yellowish elongate scales on tibiae intermixed with suberect blackish seta-like scales, onychia covered with recumbent, very thin, elongate whitish hairs; protarsal onychia considerably long, about 1.2 �� as long as tarsomeres 1���3 combined; protarsal lateral claws by 1/4, meso- and metatarsal medial claws only slightly shorter than their pair-claws. Penis: Figs 33 d���f, its body with sharply tapered straight apex. Female. Rostrum longer (Rl/Pl 1.29), antennal insertion at about 0.6���0.7 of rostrum length. Ventrites 1 and 2 without impression. Protarsal onychia short, of about 0.7 length of tarsomeres 1���3 combined, claws only very slightly unequally long. Variability. Length ♂♂ 3.55���4.15 mm, ♀♀ 3.30���4.27 mm. This species does not show marked variability. In some specimens, especially from the Balkan Peninsula, the protarsal onychia in male are slightly longer in relation to tarsomeres 1���3 combined. Sporadically, the dorsal black macula is somewhat elongate instead of round. Diagnosis. This species is characterised by small body size, clearly vaulted interstriae, markedly rounded convex pronotum with densely, unevenly clustered yellowish scales in both sexes, and by long protarsal male onychia, and tapered sharp apex of body of penis. Comparative notes. Cionus balianii differs from the closely related C. ungulatus by the unevenly distributed scales on pronotum, and especially by the shape of the body of penis, which is tapered at apex and in lateral view straight in C. balianii, and subtruncate at apex, in lateral view clearly bent dorsally in C. ungulatus. C. winkelmanni has bigger teeth on profemora and on average larger body size. Biological notes. There are no data on host plant in the original description. However, the species is being regularly collected on species of Scrophularia in the S. canina -group. The first author collected a series of specimens on Scrophularia sp. of the above group in Bulgaria. According to data on specimens studied, the species was collected from May to July at elevations 250���1,400 m a. s. l. Distribution. Italy, Croatia (1 ♂ 40 km N Split (BN)), Bulgaria, Greece. The paratypes of C. galanus from Turkey belong probably to C. winkelmanni. Non-type specimens examined. We examined more than 280 specimens in large series from above mentioned countries except for Croatia, where only one specimen was recorded., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 58-60, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Solari, F. (1932) Una nuova specie italiana di \" Cionus \" (Col. Curcul.). Bollettino della Societa Entomologica Italiana, 64, 88 - 89.","Zumpt, F. (1937) Curculioniden-Studien XXVI. Erganzungen zur \" Monographie der palaarktischen Arten der Tribus Cionini \" von Alois Wingelmuller in Wien. Koleopterologische Rundschau, 23, 222 - 228.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp.","Angelov, P. (1978) Eine neue Art der Gattung Cionus Clairvillei (Coleoptera: Curculionidae). Acta Zoologica Bulgarica, 10, 68 - 70.","Dieckmann, L. & Behne, L. (1994) Curculionidae. In: Lohse, G. A. & Lucht, W. H. (Eds.), Die Kafer Mitteleuropas. 3. Supple- mentband. Goecke & Evers Verlag, Krefeld, pp. 246."]}

46. Cionus neglectus sp. n. Figs 46 a���f. http://zoobank.org/ urn:lsid:zoobank.org:act: 57FA0E84-D61A-4FCD-8311-B7DE30EB0F28 Type locality. Rila-Kloster (Bulgaria). Type series. Holotype: a well-preserved, 4.12 mm long male labeled ���Bulg. Rila - Geb. Rila - Kloster 1200 m 15.VI.1987 leg. Zerche &Behne / sp. griseopubens Wnglm. Gruppe Dieckmann det. 1987 / coll. Be / HOLOTYPUS Cionus neglectus sp. n. M.Ko��ť��l et R.Caldara des. 2017 [printed red label]��� (SDEI). Paratypes (the same type labeling as holotype but instead of ��� HOLOTYPUS ��� ��� PARATYPUS ���): ��� Bulg. Rila - Geb. Rila - Kloster 1200 m 15.VI.1987 leg. Zerche & Behne / Cionus sp. griseopubens Wnglm. Gruppe Dieckmann det. 1987 / coll. Be ��� (1 ♂ BN); ���Bulg., Stara Pl. 8 km N Kalofer 800 m 6.VI.1987 leg. Zerche & Behne / spec. Gruppe um griseopubens Wnglm. Dieckmann det. 1987 / coll. Be ��� (1 ♀ BN); ��� Bulg. mer.occ. 20.6.73 Mt.Pirin-Begovica 1000-1600 m, Hor��k leg. / BMNH {E} 2010/26 O. Voř����ek ��� (1 ♀ BMNH); ��� Macedonia MK 2010 Kichevo, Malo Crsko 1.5. 41 o 23��N, 21 o 01��E 800 m Leg. L. Bure�� ��� (1 ♂ coll. Bure��). Description. Male (holotype). Body stout, suboval. Head: rostrum moderately stout, medium long (l/ w 4.6, Rl/Pl 1.33), black; in lateral view similar to that of C. ganglbaueri, except apical part longer, evenly tapered to apex; in dorsal view near base moderately laterally constricted, in apical part slightly dorsoventrally flattened; texture and vestiture as in C. ganglbaueri. Head between eyes and eyes as in C. griseopubens. Antennae as in C. griseopubens except segments 3���4 apparently longer than wide, segment 5 slightly longer than wide; club markedly elongate, more than three times as long as wide, otherwise as in C. griseopubens. Pronotum: black, somewhat wider than long (Pl/Pw 0.65), texture and vestiture as in C. ganglbaueri, widest at base, outline as in C. griseopubens, without constriction, in lateral view as in C. griseopubens except curvature between basal and apical part less abrupt. Prosternum: similar to that in C. griseopubens. Scutellum: as in C. ganglbaueri. Elytra: black, in basal 2/3 very slightly rounded to subparallel, in apical part broadly, somewhat irregularly rounded, moderately elongate (El/Ew 1.26), widest in about mid-length, at base somewhat wider than pronotum (Ew/Pw 1.63), humeri rounded, moderately prominent, with indistinct posthumeral impression; almost flat on disc; pattern, width of interstriae and striae as in C. ganglbaueri except for both black tomentous maculae noticeably smaller, vestiture almost completely concealing integument. Venter: vestiture as in C. griseopubens but hair-like scales slightly thicker on mes- and metepisterna; mesosternal process as in C. griseopubens; metasternum moderately concave, transversally punctured to ribbed; ventrite 1 with deep, semidensely roundly punctured impression, ventrite 2 with shallow, transversally ribbed impression; ventrite 1 1.6 �� as long as ventrite 2, ventrites 1���2 combined 4.4 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.8 length of ventrite 5. Legs: vestiture as in C. griseopubens, profemoral teeth and other characters as in C. ganglbaueri. Penis: Figs 46 d���f, very similar to that of C. ganglbaueri but its body slightly longer. Female. Rostrum longer (Rl/Pl 1.57), antennal insertion closer to rostrum mid-length, shortly beyond 0.6 of rostrum length. Ventrites 1 and 2 without impression. Protarsal onychia slightly shorter, claws equally long. Variability. Length ♂♂ 3.79���4.11 mm, ♀♀ 3.85���3.98 mm. The type series does not show much variability. Diagnosis. This species is recognizable by concealed elytral integument, alternating black patchy pattern on odd interstriae, suboval elytra, moderately convex and subrotund pronotum, grayish vestiture, elongate antennal club, normally long protarsal onychia in both sexes, and penis shape. Comparative notes. This species is most closely related to C. ganglbaueri and C. griseopubens. It differs from the former by its elongate antennal club, more convex and less conically narrowed pronotum, suboval elytra and smaller dorsal elytral macula, and from the latter by the suboval elytra, and penis shape. Biological notes. Unknown. Verbascum species are suspected to be host plants because some related species in the group (C. ganglbaueri and C. colonnellii) develop on Verbascum. Montane species collected at 800���1,600 m a. s. l. Distribution. Bulgaria, Macedonia. Etymology. The species was being overlooked and confused mostly with C. griseopubens and C. ganglbaueri for a long time. The adjective ���neglectus��� means ���overlooked��� in Latin. Non-type specimens examined. None., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on page 79, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117

53. Cionus zonovi Korotyaev, 1984 Figs 53 a���f. Cionus zonovi Korotyaev, 1984: 347. Caldara, 2013: 124. Alonso-Zarazaga et al., 2017: 186. Type locality. Sagly (Tuva Republic, Russian Federation). Type series. The species was described based on a large series of 54 specimens, five including the holotype from Tuva, 49 from Ulaanbaatar district and Bayankhongor province (Mongolia). We had an opportunity to study two female paratypes from coll. ZIN labeled ��� Mongolia, Bayan-Khong. a. 19 KM Yu Shine-Dzinsta 21.VII.81 Korotyaev [handwritten in Russian] / Paratypus / Cionus zonovi Korotyaev, 1984 / Zool. Inst. St. Petersburg / coll. Mus. Petersburg���. The locality and morphology of both paratypes fully correspond to the original description. Synonyms. None. Redescription. Male. Body moderately stout, subparallel. Head: rostrum medium stout, medium long (l/ w 4.1, Rl/Pl 1.16), reddish brown; in lateral view slightly curved, on upper outline at antennal insertion somewhat unevenly curved, in basal part of same width from base to antennal insertion, in apical part, especially on upper outline tapered to apex; in dorsal view slightly broadened from base to antennal insertion, from antennal insertion slightly narrowed to apex, in basal part almost round in cross-section, in apical part dorsoventrally flattened; very densely longitudinally punctured except apex being punctured more sparsely, with bare shiny median area very close to apex; basal part with up- and backwardly oriented, subrecumbent and recumbent brownish scales, in apical part with suberect, whitish, longer seta-like scales. Head between eyes moderately narrow, of 0.6 rostrum width at base. Eyes large, round, not protruding from head outline. Antennae reddish-brown, inserted at 2/3 of rostrum length, funicle of 0.7 scape length, segment 1 wider than segment 2, segment 1 1.5 ��, segment 2 almost twice as long as wide, segments 3���5 as long as wide to transverse; club spindle-shaped, twice as long as wide, completely covered with recumbent grayish and reddish hairs, and sparse erect, relatively short, light sensilla. Pronotum: reddish-brown, with brown irregular areas on disc, strikingly wider than long (Pl/Pw 0.54), very densely punctured, punctures subrotund, of approximately equal size, spaces between punctures smaller than puncture diameter; densely covered with subrecumbent, mostly forwardly oriented, moderately elongate (l/w 3���5), intermixed yellowish and reddish scales; widest at base, parallel to more than half of its length, then abruptly narrowed to anterior margin, with distinct, relatively deep constriction, in lateral view in basal 0.6 of its length flat, than moderately falling to anterior margin. Prosternum: anterior margin with shallowly incised, semicircular emargination without distinct bound, separated from coxae by very narrow prosternal area. Scutellum: triangular, covered with same type of scales as on elytra, densely to confluently punctured. Elytra: reddish brown, in basal 2/3 parallel, in apical third broadly evenly rounded, elongate (El/Ew 1.40), widest at about half of their length, at base moderately wider than pronotum (Ew/Pw 1.54), humeri subquadrate, prominent, with posthumeral impression; flat on disc; interstria 1 at about 0.4 of medial length moderately, in preapical area very slightly broadened, interstria 2 dorsally slightly narrowed and constricted laterally encompassing longitudinally elongate dorsal and rudimentary preapical black tomentous maculae without differently colored scale border; interstriae except perimacular areas of approximately equal width; odd interstriae with only indistinct alternating clusters of whitish and brown scales; striae shallow, formed by single rows of slightly unevenly arranged, round punctures; entire surface densely covered with recumbent to subrecumbent, relatively shortly elongate (l/w 3���5), whitish and reddish intermixed scales fully concealing integument. Venter: almost evenly covered with recumbent, elongate, whitish and yellowish scales without apparent clusters; mesosternal process flat, medium broad, blunt at posterior margin, scaled, densely punctured; metasternum moderately concave, transversally ribbed to punctured, ventrite 1 with medium deep impression, ventrite 2 in anterior part with very shallow impression, both punctured; ventrite 1 1.7 �� as long as ventrite 2, ventrites 1���2 combined 4.4 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of almost 0.9 length of ventrite 5, ventrite 5 transverse, slightly emarginate at apex. Legs: brown except reddish-brown tarsi, with darkened apices of onychia, profemora with only indistinct teeth emphasized by erect scales, meso- and metafemora with medium large subtriangular sharp teeth; femora and tibiae covered with recumbent to subrecumbent, shortly elongate, evenly, semidensely distributed whitish scales, tarsi on tarsomeres 1���3 with subrecumbent to suberect, onychia with recumbent whitish hair-like scales, tarsomere 3 wider than long; protarsal onychia moderately shorter than tarsomeres 1���3 combined; claws of equal length. Penis: Figs 53 d���f, its body medium long, with sinuate sides, broadly tapered at apex, with small incisure on tip. Female. Rostrum longer (Rl/Pl 1.58), antennal insertion closer to rostrum mid-length, at 0.6 of rostrum length. Ventrites 1 and 2 without impression. Variability. Length ♂♂ 3.70���3.83 mm, ♀♀ 3.74���4.04 mm. This species does not show a remarkable variability. In some specimens, the brown areas on pronotum can be reduced or missing. Diagnosis. Cionus zonovi is recognizable by completely concealed elytral integument, markedly transverse pronotum, elongate subrectangular elytra, indistinct elytral pattern with elongate dorsal macula, short spindle-shaped antennal club, equally long claws in males, and penis shape. Comparative notes. Cionus zonovi is a very distinct species apparently unrelated to any other known species of the genus. Biological notes. All specimens from the original description were collected on Scrophularia incisa Weinmann. Distribution. Siberia (Tuva), Mongolia. Non-type specimens examined. We examined 9 specimens from Mongolia (Gurvan Sajchan 2,7.87 O. Majzlan lgt., 3 ♂♂, 1 ♀ MN, 1 ♀ KO; Gobi-Altai. 17.vi.1988, leg. U. G��llner, 2 ♂♂ ZMHB; Bayankhongor, 8.vii.2004, 1,500 m a. s. l., leg. Straka, 1 ♂, 1 ♀ SC)., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 89-90, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Korotyaev, B. A. (1984) K poznanyiu fauny zhukov-dolgonosikov (Coleoptera, Apionidae, Curculionidae) Mongolii i sopredel'nych s ney territoriy. II. Nasekomye Mongolii, 9, 311 - 355.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp."]}

31. Cionus wanati Białooki, 2006 Figs 31 a���f. Cionus wanati Białooki, 2006: 58. Caldara, 2013: 124. Alonso-Zarazaga et al., 2017: 186. Type locality. Camliyayla (Turkey). Type series. The species was described based on 33 specimens collected in a single locality. We had an opportunity to study three male paratypes (BI), all labeled ��� 26.05.2001. S Turkey Camliyayla env. N Icel leg. P. Bialooki / Cionus pr. donckieri Pic sp.n. det. P. Bialooki 2001��� bearing original red paratype labels. Synonyms. None. Redescription. Male. Body medium stout, subparallel to suboval. Head: rostrum medium stout, medium long (l/ w 4.2, Rl/Pl 1.11), in lateral and dorsal view texture and vestiture as in C. donckieri. Head, eyes and antennae as in C. donckieri. Pronotum: as in C. donckieri except in lateral view more flat. Prosternum: as in C. donckieri. Scutellum: as in C. donckieri. Elytra: color and shape similar to those in C. donckieri but somewhat shorter (El/Ew 1.23); interstria 1 at same medial lengths as in C. donckieri distinctly broadened, interstria 2 at same lengths apparently narrowed and constricted laterally encompassing medium large black tomentous maculae without differently colored scale border; width of interstriae as in C. donckieri; odd interstriae with unevenly distributed, variously large patches of black scales and alternating indistinct patches of yellow scales; entire surface as in C. donckieri but erect black seta-like scales more numerous. Venter: recumbent scales distributed without apparent clusters on margins; metasternum transversally ribbed, impression on ventrites 1 and 2 densely transversally punctured, with transverse ribs, otherwise as in C. donckieri. Legs: as in C. donckieri except missing fringe in apical part of metatibiae, tarsomere 3 only slightly wider than long, with relatively narrow lobes. Penis: Figs 31 d���f, very similar to that of C. donckieri. Female. Rostrum longer (Rl/Pl 1.48), antennae inserted at 0.6 of rostrum length. Ventrites 1 and 2 without impression. Claws approximately equally long. Variability. Length ♂♂ 3.98���4.41 mm, ♀♀ 4.20���4.53 mm. This species does not show remarkable variability. The elytral pattern may be more or less defined but always clearly apparent. The size of both elytral maculae does not show remarkable variability: in some specimens the dorsal macula is elongate. Diagnosis. Cionus wanati is recognizable by almost concealed elytral integument, antennal insertion in males close to apex, at about 0.7 of rostrum length, slender lobes of tarsomere 3, medium-sized elytral maculae, distinct elytral pattern of patches of black scales, lack of metatibial apical fringe of scales in males, erect seta-like scales on elytra, and penis shape. Comparative notes. Cionus wanati is very closely related to C. donckieri, from which it differs by more distinct elytral pattern, shorter rostrum in females and lack of fringe of scales on male metatibial apices. It differs from the related C. leonhardi and C. hypsibatus by erect seta-like scales on elytra. Biological notes. Białooki (2006) collected this species on Verbascum sp. It was collected at elevations 500��� 2,000 m a. s. l. from May to July. Distribution. Turkey. Non-type specimens examined. We examined 27 specimens from the following localities. TURKEY: Ankara (PY), Ardaham (PY), Boganbeyli (BMNH), Iskenderun (SDEI), Karacay env. (MNHN), Namrun (BMNH), Sultan Dagh (SDEI)., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 55-56, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Bialooki, P. (2006) Two new species of Cionus Clairville, 1798 (Coleoptera: Curculionidae: Cionini) from Turkey. Polskie Pismo Entomologiczne, 75, 55 - 64.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp."]}

22. Cionus rabinovitchi A. Hoffmann, 1938 Figs 22 a���f. Cionus rabinovitchi A. Hoffmann, 1938b:128. Caldara, 2013: 124. Alonso-Zarazaga et al., 2017: 186. Type locality. Wasgiya (Egypt). Type series. The species was described based on two females collected by A. Rabinovitch in Wasgiya, Egypt. In coll. Hofmann (MNHN), there are two females with label data corresponding to those reported in the original description. One bears a red label ���TYPE���, the other one ���COTYPE���. Because the author mentioned the existence of the type in his collection, the former specimen, completely preserved female, 4.08 mm long, labeled ��� EGYPTE Wasgiya 23.4.1935 A.RABINOVITCH / Cionus Rabinovitchi (♀) A.Hoffmann / TYPE [red label] / coll. M.Paris Hoffmann��� was regarded as the holotype, and the latter specimen labeled with the same locality and collection labels as the paratype. For clarity, we marked the former specimen by adding the label ��� HOLOTYPUS Cionus rabinovitchi Hoffmann M.Ko��ť��l et R.Caldara vid. 2017 [printed red label]���, the latter as the paratype accordingly. Synonyms. None. Redescription. Female. Body stout, subrotund to round. Head: rostrum moderately slender, medium long (l/ w 5.5, Rl/Pl 1.22), black to dark brown; in lateral view very slightly, somewhat unevenly curved, apical part almost narrow, tapered from base to apex, subparallel shortly before apex; in dorsal view same width from base to apex, basal part in cross-section almost round, apical part moderately flattened dorsoventrally; basal part densely punctured, punctures longitudinally elongate, distal half of apical part with semidensely, towards apex sparsely distributed, small punctures; basal part with up- and backwardly oriented, elongate reddish scales, apical part bare, apex with a few erect seta-like scales. Head between eyes narrow, of about 0.4 rostrum width at base. Eyes large, broadly rounded, not protruding from head outline. Antennae reddish-brown, inserted shortly beyond half of rostrum length; funicle of 3/4 scape length, segment 1 wider than segment 2, segment 1 twice, segment 2 2.5 �� as long as wide, segments 3���5 as long as wide, subquadrate; club oval, more than twice as long as wide, of 0.8 funicle length, completely covered with recumbent, tiny light hairs and relatively numerous, erect, long pale sensilla. Pronotum: reddish-brown, somewhat wider than long (Pl/Pw 0.63), very densely finely evenly punctured, punctures round or subrotund, approximately of equal size, spaces between punctures smaller than puncture diameter; covered with evenly densely arranged, in middle forwardly, on sides inwardly oriented, recumbent to subrecumbent, elongate (l/w 4���8) reddish and whitish scales; widest at base, until half of pronotum length moderately conically, then abruptly more conically narrowed to anterior margin, without apparent constriction, in lateral view flat in basal half, then falling to anterior margin. Prosternum: anterior margin with sharply incised emargination of medium depth, not bounded by tubercles, separated from coxae by narrow prosternal area. Scutellum: triangular, covered with backwardly oriented, elongate yellowish and reddish scales, finely punctured. Elytra: reddish-brown, in basal 2/3 slightly rounded, in apical third broadly rounded, moderately elongate (El/Ew 1.20), widest approximately at one third of their length, at base somewhat wider than pronotum (Ew/Pw 1.59), humeri rounded, moderately prominent, without posthumeral impression; moderately convex on disc; interstria 1 from 0.25 to 0.4 of medial length moderately broadened, in preapical area slightly broadened, interstria 2 in dorsal area slightly narrowed and visibly constricted laterally encompassing medium large, subrotund dorsal and small irregular preapical black tomentous maculae with indistinct border formed by reddish scales; interstriae except dorsal perimacular area of approximately same width, flat, with sparsely scattered round punctures; odd interstriae, especially in posterior half of elytra, with very indistinct, unevenly distributed small patches of dark scales; striae shallow, formed by single rows of slightly unevenly distributed round punctures; entire surface covered with dense, recumbent to subrecumbent, backwardly tipped, elongate (l/w 4���7) yellowish and sparsely intermixed reddish scales almost completely concealing integument. Venter: densely covered with yellowish and reddish scales of similar type as on elytra without clusters on ventrite sides; mesosternal process flat, broad, almost straight at posterior margin, with dense scales; metasternum convex, densely punctured; ventrites 1���2 convex, densely punctured; ventrite 1 1.6 �� as long as ventrite 2, ventrites 1���2 combined 4.3 �� as long as ventrites 3���4 combined, ventrites 3���4 combined slightly shorter than ventrite 5. Legs: reddish-brown, profemora with tiny teeth emphasized by erect scales, meso- and metafemora with large sharp teeth; excepting onychia covered with recumbent to suberect elongate scales being on femora reddish and sparsely yellowish, on tibiae reddish and yellowish, on tarsomeres 1���3 whitish, onychia covered with recumbent long whitish hairs, femora and tibiae without bands of scales; protarsal onychia as long as tarsomeres 1���3 combined, protarsal tarsomere 3 slightly wider than long; claws of approximately equal length. Male. Unknown. Variability. Length ♀♀ 3.90���4.08 mm. The two type specimens do not show any significant differences. Diagnosis. This species is recognizable by almost concealed elytral integument, tapered apical part of rostrum in lateral view, reddish-brown integument, yellowish and reddish intermixed tipped scales on elytra, and dark rostrum. Comparative notes. In some extent, especially in rostrum shape, C. rabinovitchi is similar to C. rufescens and C. hortulanus but clearly distinguishable from them by most of the above reported characters in the diagnosis. Biological notes. Biology unknown. Distribution. Egypt. Non-type specimens examined. None., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 42-43, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Hoffmann, A. (1938 b) Curculionidae (Col.) noveaux du Maroc. Bulletin de la Societe Entomologique de France, 43, 100 - 104.","Caldara, R. (2013) Curculioninae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 8. Leiden, Brill, pp. 117.","Alonso-Zarazaga, M. A., Barrios, H., Borovec, R., Bouchard, P., Caldara, R., Colonnelli, E., Gultekin, L., Hlavac, P., Korotyaev, B., Lyal, C. H. C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp."]}

51. Cionus schoenherri C. N. F. Brisout de Barneville, 1863 Figs 51 a���f. Cionus schoenherri C. N. F. Brisout de Barneville, 1863: 115. Reitter, 1904: 55. Wingelm��ller, 1914: 193; 1921: 105; 1937: 174. Hustache, 1932: 342. A. Hoffmann, 1958: 1220. Caldara, 2013: 124. Alonso-Zarazaga et al., 2017: 186. Cionus schoenherri laportei A. Hoffmann, 1953: 193. Caldara, 2013: 124. Alonso-Zarazaga et al., 2017: 186. syn. n. Type locality. France (Eastern Pyrenees), Spain. As both lectotype and paralectotype bear no locality label, and the lectotype coming from one of both type localities was designated, both localities should be regarded as type localities. According to Articles 73.2.3 and 76 of the ICZN (1999), more type localities for one species are admissible. Type series. The species was described based on specimens from Eastern Pyrenees and Spain. In coll. C. N. F. Brisout (MNHN), there are a couple on a single pinned card. A 3.77 mm long, well-preserved male, corresponding to the original description, labeled ���Schoenheri / Mus��um Paris S. E. F. Coll. Ch. Brisout��� was designated as the lectotype by adding the label ��� LECTOTYPUS Cionus schoenherri C. Brisout ♂ Michael Ko��ť��l des. 2014 [printed red label]���. The female was labeled as paralectotype accordingly. Synonyms. Cionus schoenherri var. laportei was described based on two males from Algeria. In coll. A. Hoffmann (MNHN), there is one male labeled ���Sakamody I-VI-52 (Algerie) M.Laporte / Cionus Schonherri Ch. Bris var. a...[illegible]... discale oblongue et etroite. A. Hoffmann / Cionus Schonherri Ch. Bris. v. Laportei m. A. Hoffmann det. / TYPE [red label] / MUSEUM PARIS 1968 Col. A. HOFFMANN���. According to the original description, this specimen is a syntype. We dissected this well-preserved, 4.15 mm long specimen for genitalia and designated it as the lectotype of Cionus schoenherri var. laportei A. Hoffmann by adding the printed red label ��� LECTOTYPUS Cionus schoenherri var. laportei Hoffmann M.Ko��ť��l et R.Caldara des.2018���. We compared it with the lectotype of C. schoenherri as well as with the non-type specimens of C. schoenherri from North Africa. As the only difference is the shape of the dorsal elytral macula and in the material from North Algeria (Setif) (coll. SMDEI), there are specimens with both round and oval dorsal macula, we consider this taxon consubspecific with C. schoenherri. Redescription. Male. Body stout, subrotund. Head: rostrum moderately slender, medium long (l/ w 4.5, Rl/Pl 1.17), dark brown; in lateral view evenly moderately curved, in basal part of same width, from antennal insertion to apex slightly tapered, ventral outline not beveled at apex; in dorsal view slightly broadened from base to antennal insertion, then of same width to apex, in basal part slightly constricted laterally, in apical part moderately flattened; texture as in C. longicollis; basal part with up- and backwardly oriented, subrecumbent, blackish and sparser yellowish elongate scales, apical part with relatively short, for- and inwardly oriented, suberect, yellowish seta-like scales. Head between eyes very narrow, of less than 0.4 rostrum width at base. Eyes large, broadly rounded, very slightly protruding from head outline. Antennae reddish-brown, club slightly darkened, inserted at 0.6 of rostrum length; funicle of 0.7 scape length, segment 1 wider than segment 2, segment 1 twice, segment 2 almost three times as long as wide, segments 3���5 as long as wide; club spindle-shaped, about 2.5 �� as long as wide, of about 0.8 funicle length, completely covered with thin, recumbent reddish hairs and sparse erect, relatively short, light brown sensilla. Pronotum: brown, markedly wider than long (Pl/Pw 0.60), very densely unevenly punctured, punctures rather variable in size, subrotund, spaces between punctures much smaller than puncture diameter; unevenly densely covered with recumbent, up- and forwardly oriented, variously large, elongate (l/w 4���6), yellowish, on sides gingery scales, mediobasal area with more sparsely distributed scales; widest at base, roundly narrowed to anterior margin, without constriction, in lateral view flat in basal half, then falling to anterior margin. Prosternum: anterior margin with moderately deep, semicircular emargination bounded by less distinct tubercles, separated from coxae by narrow prosternal area. Scutellum: as in C. longicollis. Elytra: brown to reddish-brown, in basal 2/3 slightly rounded to subparallel, in apical third broadly evenly rounded, moderately elongate (El/Ew 1.17), widest at 1/3 of their length, at base somewhat wider than pronotum (Ew/Pw 1.65), humeri subrotund, moderately prominent, with posthumeral impression; almost flat on disc; interstria 1 from 0.2 to almost half of medial length and in preapical area very strongly broadened, interstria 2 at same lengths very strongly narrowed to invisible, considerably constricted laterally, interstria 3 on dorsum clearly constricted laterally encompassing very large round black tomentous macula with medium broad border of gingery and clustered whitish scales; interstriae except perimacular areas of approximately equal width, in basal part of elytra slightly vaulted; odd interstriae with unevenly distributed small, indistinct patches of whitish to yellowish scales and sparser patches of dark brown scales; striae shallow, formed by densely arranged regular rows of large punctures; entire surface covered with unevenly densely distributed, variously large and thin, moderately elongate (l/w 3���5), recumbent whitish scales and sparsely intermixed gingery scales almost completely concealing integument; gingery and yellowish broad scales densely clustered in sub- and posthumeral area forming large orange irregular macula. Venter: semidensely covered with thin, backwardly oriented, subrecumbent, grayish hairs, at mes- and metepisternum, and lateral parts of metasternum covered with very densely distributed, broad, intermixed whitish and gingery scales forming confluent area with lateral pronotal and subhumeral macula; in posterior parts of ventrites 1 and 2 paramedially, and on margins of ventrites 3���4 clusters of broader, subrecumbent whitish scales; mesosternal process very slightly convex, subquadrate, moderately emarginate at posterior margin, with hairs, very densely punctured to rugulose; metasternum slightly concave, transversally punctured to ribbed; ventrites 1 and 2 with relatively deep and broad, transversally punctured to ribbed impression; ventrite 1 1.9 �� as long as ventrite 2, ventrites 1���2 combined 4.7 �� as long as ventrites 3���4 combined, ventrites 3���4 combined of 0.8 length of ventrite 5. Legs: reddish-brown, with darkened apices of onychia, profemora with small teeth, meso- and metafemora with large triangular sharp teeth; scales similar to those in C. longicollis but denser; protarsal onychia short, of 0.8 length of tarsomeres 1���3 combined, protarsal tarsomere 3 wider than long; claws as in C. longicollis except protarsal lateral claws less reduced, of about 3/4 length of their pair-claws. Penis: Figs 51 d���f, its body medium long, with slightly convex sides, bluntly tapered at apex. Female. Rostrum moderately longer (Rl/Pl 1.30), antennae inserted as in male. Ventrites 1 and 2 without impression, convex. Claws of approximately equal length. Variability. Length ♂♂ 3.74���4.27 mm, ♀♀ 3.98���4.61 mm. This species shows variability in the size and shape of the dorsal black tomentous elytral macula. This macula can be somewhat reduced, apparently longitudinally elongate, especially in specimens from North Africa (described as C. schoenherri ssp. laportei). Diagnosis. Cionus schoenherri is distinguishable by only partially visible, in perimacular areas concealed elytral integument, subrotund elytra, very large to large dorsal elytral macula with whitish and gingery border, large sub- and posthumeral patches of gingery to reddish scales, short onychia also in males, relatively deep and large impression on ventrites 1 and 2 in males. Comparative notes. Cionus schoenherri shows a relationship with C. longicollis and C. distinctus. From the former it differs by subrotund elytra, elytral subhumeral macula, evenly roundly narrowed pronotum, shorter rostrum in both sexes, and shorter onychia. From the latter it differs by larger body size, round or longitudinally elongate dorsal elytral macula, and elytral pattern. Biological notes. The first author collected series of specimens in Spain (Castilla-Le��n, Aldealpozo) and in Morocco (High Atlas Mts. Touama, N 31��30.3��W 07��31.7��, 900 m a. s. l.) on Scropularia spp. Hoffmann (1958) reported S. canina, S. nodosa and S. lucida as host plants. The second author found this species in northern Italy on S. canina and S. nodosa. There is no doubt that there are more Scrophularia -species which are host plants of this species. The species is xerothermophilous, inhabiting lower elevations or sun-exposed montane habitats in southern Europe, and reaches higher elevations, often up to 1,200 m a. s. l. in southern Morocco (above mentioned locality and six specimens from Morocco, Imlil, 1,200 m a. s. l., BMNH). Distribution. This species is widespread in Portugal, Spain, southern France, Italy, Morocco and Algeria. Germann (2010) reported this species also from Switzerland. Non-type specimens examined. We examined 170 specimens from all the above mentioned countries except Switzerland., Published as part of Ko������l, Michael & Caldara, Roberto, 2019, Revision of Palaearctic species of the genus Cionus Clairville (Coleoptera: Curculionidae: Cionini), pp. 1-144 in Zootaxa 4631 (1) on pages 86-87, DOI: 10.11646/zootaxa.4631.1.1, http://zenodo.org/record/3294117, {"references":["Brisout de Barneville, C. N. F. (1863) In: Grenier, A. J. F., Catalogue des coleopteres de France et materiaux pour servir a la faune des coleopteres francais, par MM. E. Allard, Dr. Ch. Aube, Ch. Brisout de Barneville, A. Chevrolat, L. Fairmaire, Al. Gauvel, Dr. A. Grienier, Dr. Kraatz, J. Linder, L. Reiche et Felicien de Saulcy. A. J. F. Grenier, Paris, pp. 114.","Reitter, E. (1904) Bestimmungs-Tabelle der Coleopteren-Gattung Cionus Clairv. aus Europa und angrenzenden Landern. Wiener Entomologische Zeitung, 23, 47 - 64. https: // doi. org / 10.5962 / bhl. part. 27188","Wingelmuller, A. 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C., Machado, A., Meregalli, M., Pierotti, H., Ren, L., Sanchez-Ruiz, M., Sforzi, A., Silfverberg, H., Skuhr- ovec, J., Tryzna, M., Velazquez de Castro, A. J. & Yunakov, N. N. (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8. Sociedad Entomologica Aragonesa S. E. A., Zaragosa, 729 pp.","ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition, adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, BMNH, London, 306 pp. https: // doi. org / 10.5962 / bhl. title. 50608","Germann, C. (2010) Die Russelkafer (Coleoptera, Curculionoidea) der Schweiz-Checkliste mit Verbreitungsangaben nach biogeografischen Regionen. Mitteilungen der Schweizerischen Entomologischen Gesselschaft, 83, 41 - 118."]}