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Large amounts of methane (CH4) could be released as a result of the gradual or abrupt thawing of Arctic permafrost due to global warming. Once available, this potent greenhouse gas is emitted into the atmosphere or transported laterally into aquatic ecosystems via hydrologic connectivity at the surface or via groundwaters. While high northern latitudes contribute up to 5 % of total global CH4 emissions, the specific contribution of Arctic rivers and streams is largely unknown. We analyzed high-resolution continuous CH4 concentrations measured between 15 and 17 June 2019 (late freshet) in a ∼120 km transect of the Kolyma River in northeast Siberia. The average partial pressure of CH4 (pCH4) in tributaries (66.8–206.8 µatm) was 2–7 times higher than in the main river channel (28.3 µatm). In the main channel, CH4 was up to 1600 % supersaturated with respect to atmospheric equilibrium. Key sites along the riverbank and at tributary confluences accounted for 10 % of the navigated transect and had the highest pCH4 (41 ± 7 µatm) and CH4 emissions (0.03 ± 0.004 mmolm-2d-1) compared to other sites in the main channel, contributing between 14 % to 17 % of the total CH4 flux in the transect. These key sites were characterized by warm waters (T>14.5 ∘C) and low specific conductivities (κ µS cm−1). The distribution of CH4 in the river could be linked statistically to T and κ of the water and to their proximity to the shore z, and these parameters served as predictors of CH4 concentrations in unsampled river areas. The abundance of CH4-consuming bacteria and CH4-producing archaea in the river was similar to those previously detected in nearby soils and was also strongly correlated to T and κ. These findings imply that the source of riverine CH4 is closely related with sites near land. The average total CH4 flux density in the river section was 0.02 ± 0.006 mmolm-2d-1, equivalent to an annual CH4 flux of 1.24×107 g CH4 yr−1 emitted during a 146 d open water season. Our study highlights the importance of high-resolution continuous CH4 measurements in Arctic rivers for identifying spatial and temporal variations, as well as providing a glimpse of the magnitude of riverine CH4 emissions in the Arctic and their potential relevance to regional CH4 budgets.

JJL received funding from the Research Foundation Flanders (grant nr. 12P1819N). The project received funding from the Research Foundation Flanders (grants nrs, G018919N, W001919N). JVDH and TWC received funding from DOB Ecology. JA received funding from the University of Helsinki, Faculty of Science (MICROCLIM, grant nr. 7510145) and Academy of Finland Flagship (grant no. 337552). PDF, CM and PV received funding from the European Research Council (ERC) under the European Union's Horizon 2020 research and innovation programme (ERC Starting Grant FORMICA 757833). JK received funding from the Arctic Interactions at the University of Oulu and Academy of Finland (318930, Profi 4), Maaja vesitekniikan tuki ry., Tiina and Antti Herlin Foundation, Nordenskiold Samfundet and Societas pro Fauna et Flora Fennica. MK received funding from the Czech Science Foundation (grant nr. 20-28119S) and the Czech Academy of Sciences (grant nr. RVO 67985939). TWC received funding from National Geographic Society grant no. 9480-14 and WW-240R-17. MA received funding from CISSC (program ICRP (grant nr:2397) and INSF (grant nr: 96005914). The Royal Botanic Garden Edinburgh is supported by the Scottish Government's Rural and Environment Science and Analytical Services Division. JMA received funding from the Funding Org. Qatar Petroleum (grant nr. QUEX-CAS-QP-RD-18/19). JMA received funding from the European Union's Horizon 2020 research and innovation program (grant no. 678841) and from the Swiss National Science Foundation (grant no. 31003A_176044). JA was supported by research grants LTAUSA19137 (program INTER-EXCELLENCE, subprogram INTER-ACTION) provided by Czech Ministry of Education, Youth and Sports and 20-05840Y of the Czech Science Foundation. AA was supported by the Ministry of Science and Higher Education of the Russian Federation (grant FSRZ-2020-0014). SN, UAT, JJA, and JvO received funding from the Independent Research Fund Denmark (7027-00133B). LvdB, KT, MYB and RC acknowledge funding from the German Research Foundation within the Priority Program SPP-1803 'EarthShape: Earth Surface Shaping by Biota' (grant TI 338/14-1&2 and BA 3843/6-1). PB was supported by grant project VEGA of the Ministry of Education of the Slovak Republic and the Slovak Academy of Sciences No. 2/0132/18. Forest Research received funding from the Forestry Commission (climate change research programme). JCB acknowledges the support of Universidad Javeriana. JLBA received funding from the Direccion General de Cambio Climatico del Gobierno de Aragon; JLBA acknowledges fieldwork assistance by Ana Acin, the Ordesa y Monte Perdido National Park, and the Servicio de Medio Ambiente de Soria de la Junta de Castilla y Leon. RGB and MPB received funding from BECC - Biodiversity and Ecosystem services in a Changing Climate. MPB received funding from The European Union's Horizon 2020 research and innovation program under the Marie Skodowska-Curie Grant Agreement No. 657627 and The Swedish Research Council FORMAS - future research leaders No. 2016-01187. JB received funding from the Czech Academy of Sciences (grant nr. RVO 67985939). NB received funding from the SNF (grant numbers 40FA40_154245, 20FI21_148992, 20FI20_173691, 407340_172433) and from the EU (contract no. 774124). ICOS EU research infrastructure. EU FP7 NitroEurope. EU FP7 ECLAIRE. The authors from Biological Dynamics of Forest Fragments Project, PDBFF, Instituto Nacional de Pesquisas da Amazonia, Brazil were supported by the MCTI/CNPq/FNDCT - AcAo Transversal no68/2013 - Programa de Grande Escala da Biosfera-Atmosfera na Amazonia - LBA; Project 'Como as florestas da Amazonia Central respondem as variacoes climaticas? Efeitos sobre dinamica florestal e sinergia com a fragmentacAo florestal'. This is the study 829 of the BDFFP Technical Series. to The EUCFLUX Cooperative Research Program and Forest Science and Research Institute-IPEF. NC acknowledges funding by Stelvio National Park. JC was funded by the Spanish government grant CGL2016-78093-R. ANID-FONDECYT 1181745 AND INSTITUTO ANTARTICO CHILENO (INACH FR-0418). SC received funding from the German Research Foundation (grant no. DFG- FZT 118, 202548816). The National Science Foundation, Poland (grant no. UMO-2017/27/B/ST10/02228), within the framework of the 'Carbon dioxide uptake potential of sphagnum peatlands in the context of atmospheric optical parameters and climate changes' (KUSCO2) project. SLC received funding from the South African National Research Foundation and the Australian Research Council. FM, M, KU and MU received funding from Slovak Research and Development Agency (no. APVV-19-0319). Instituto Antartico Chileno (INACH_RT-48_16), Iniciativa Cientifica Milenio Nucleo Milenio de Salmonidos Invasores INVASAL, Institute of Ecology and Biodiversity (IEB), CONICYT PIA APOYO CCTE AFB170008. PC is supported by NERC core funding to the BAS 'Biodiversity, Evolution and Adaptation Team. EJC received funding from the Norwegian Research Council (grant number 230970). GND was supported by NERC E3 doctoral training partnership grant (NE/L002558/1) at the University of Edinburgh and the Carnegie Trust for the Universities of Scotland. Monitoring stations on Livingston Island, Antarctica, were funded by different research projects of the Gobern of Spain (PERMAPLANET CTM2009-10165-E; ANTARPERMA CTM2011-15565-E; PERMASNOW CTM2014-52021-R), and the PERMATHERMAL arrangement between the University of Alcala and the Spanish Polar Committee. GN received funding from the Autonomous Province of Bolzano (ITA). The infrastructure, part of the UK Environmental Change Network, was funded historically in part by ScotNature and NERC National Capability LTS-S: UK-SCAPE; NE/R016429/1). JD was supported by the Czech Science Foundation (GA17-19376S) and MSMT (LTAUSA18007). ED received funding from the Kempe Foundation (JCK-1112 and JCK-1822). The infrastructure was supported by the Ministry of Education, Youth and Sports of the Czech Republic within the National Sustainability Programme I (NPU I), grant number LO1415 and by the project for national infrastructure support CzeCOS/ICOS Reg. No. LM2015061. NE received funding from the German Research Foundation (DFG- FZT 118, 202548816). BE received funding from the GLORIA-EU project no EVK2-CT2000-00056, the Autonomous Province of Bolzano (ITA), from the Tiroler Wissenschaftsfonds and from the University of Innsbruck. RME was supported by funding to the SAFE Project from the Sime Darby Foundation. OF received funding from the German Research Foundation (DFG- FZT 118, 202548816). EFP was supported by the Jardin Botanico Atlantico (SV-20-GIJON-JBA). MF was funded by the German Federal Ministry of Education and Research (BMBF) in the context of The Future Okavango (Grant No. 01LL0912) and SASSCAL (01LG1201M; 01LG1201N) projects. EFL received funding from ANID PIA / BASAL FB210006. RAG received funding from Fondecyt 11170516, CONICYT PIA AFB170008 and ANID PIA / BASAL FB210006. MBG received funding from National Parks (DYNBIO, #1656/2015) and The Spanish Research Agency (VULBIMON, #CGL2017-90040-R). MG received funding from the Swiss National Science Foundation (ICOS-CH Phase 2 20FI20_173691). FG received funding from the German Research Foundation (DFG- FZT 118, 202548816). KG and TS received funding from the UK Biotechnology and Biological Research Council (grant = 206/D16053). SG was supported by the Research Foundation Flanders (FWO) (project G0H1517N). KJ and PH received funding from the EU Horizon2020 INFRAIA project eLTER-PLUS (871128), the project LTER-CWN (FFG, F&E Infrastrukturforderung, project number 858024) and the Austrian Climate Research Program (ACRP7 - CentForCSink - KR14AC7K11960). SH and ARB received funding through iDiv funded by the German Research Foundation (DFG- FZT 118, 202548816). LH received funding from the Czech Science Foundation (grant nr. 20-28119S) and the Czech Academy of Sciences (grant nr. RVO 67985939). MH received funding from the Baden-Wurttemberg Ministry of Science, Research and Arts via the project DRIeR (Drought impacts, processes and resilience: making the in-visible visible). LH received funding from International Polar Year, Weston Foundation, and ArcticNet. DH received funding from Natural Sciences and Engineering Council (Canada) (RGPIN-06691). TTH received funding from Independent Research Fund Denmark (grant no. 8021-00423B) and Villum Foundation (grant no. 17523). Ministry of Education, Youth and Sports of the Czech Republic (projects LM2015078, VAN2020/01 and CZ.02.1.01/0.0/0.0/16_013/0001708). KH, CG and CJD received funding from Bolin Centre for Climate Research, Stockholm University and from the Swedish research council Formas [grant n:o 2014-00530 to KH]. JJ received funding from the Funding Org. Swedish Forest Society Foundation (grant nr. 2018-485-Steg 2 2017) and Swedish Research Council FORMAS (grant nr. 2018-00792). AJ received funding from the German Federal Ministry of Education and Research BMBF (Grant Nr. FKZ 031B0516C SUSALPS) and the Oberfrankenstiftung (Grant Nr. OFS FP00237). ISJ received funding from the Energy Research Fund (NYR-11 - 2019, NYR-18 - 2020). TJ was supported by a UK NERC Independent Research Fellowship (grant number: NE/S01537X/1). RJ received funding from National Science Centre of Poland (grant number: 2016/21/B/ST10/02271) and Polish National Centre for Research and Development (grant number: Pol-Nor/203258/31/2013). VK received funding from the Czech Academy of Sciences (grant nr. RVO 67985939). AAK received funding from MoEFCC, Govt of India (AICOPTAX project F. No. 22018/12/2015/RE/Tax). NK received funding from FORMAS (grants nr. 2018-01781, 2018-02700, 2019-00836), VR, support from the research infrastructure ICOS-SE. BK received funding from the National Research, Development and Innovation Fund of Hungary (grant nr. K128441). Ministry of Education, Youth and Sports of the Czech Republic (projects LM2015078 and CZ.02.1.01/0.0/0.0/16_013/0001708). Project B1-RNM-163-UGR-18-Programa Operativo FEDER 2018, partially funded data collection. Norwegian Research Council (NORKLIMA grants #184912 and #244525) awarded to Vigdis Vandvik. MM received funding from the Czech Science Foundation (grant nr. 20-28119S) and the Czech Academy of Sciences (grant nr. RVO 67985939). Project CONICYT-PAI 79170119 and ANID-MPG 190029 awarded to Roy Mackenzie. This work was partly funded by project MIUR PON Cluster OT4CLIMA. RM received funding from the SNF project number 407340_172433. FM received funding from the Stelvio National Park. PM received funding from AIAS-COFUND fellowship programme supported by the Marie Skodowska- Curie actions under the European Union's Seventh Framework Pro-gramme for Research, Technological development and Demonstration (grant agreement no 609033) and the Aarhus University Research Foundation, Denmark. RM received funding from the Ministry of Education, Youth and Sports of the Czech Republic (project LTT17033). SM and VM received funding from EU FP6 NitroEurope (grant nr. 17841), EU FP7 ECLAIRE (grant nr. 282910), the Ministry of Education and Science of Ukraine (projects nr. 505, 550, 574, 602), GEF-UNEP funded "Toward INMS" project (grant nr. NEC05348) and ENI CBC BSB PONTOS (grant nr. BSB 889). The authors from Biological Dynamics of Forest Fragments Project, PDBFF, Instituto Nacional de Pesquisas da Amazonia, Brazil were supported by the MCTI/CNPq/FNDCT - AcAo Transversal no68/2013 - Programa de Grande Escala da Biosfera-Atmosfera na Amazonia - LBA; Project 'Como as florestas da Amazonia Central respondem as variacoes climaticas? Efeitos sobre dinamica florestal e sinergia com a fragmentacAo florestal'. FJRM was financially supported by the Netherlands Organization for Scientific Research (VICI grant 016.VICI.170.072) and Research Foundation Flanders (FWO-SBO grant S000619N). STM received funding from New Frontiers in Research Fund-Exploration (grant nr. NFRF-2018-02043) and NSERC Discovery. MMR received funding from the Australian Research Council Discovery Early Career Research Award (grant nr. DE180100570). JAM received funding from the National Science Foundation (DEB 1557094), International Center for Advanced Renewable Energy and Sustainability (I-CARES) at Washington University in St. Louis, ForestGEO, and Tyson Research Center. IM-S was funded by the UK Natural Environment Research Council through the ShrubTundra Project (NE/M016323/1). MBN received funding from FORMAS, VR, Kempe Foundations support from the research infrastructures ICOS and SITES. MDN received funding from CONICET (grant nr. PIP 112-201501-00609). Spanish Ministry of Science grant PID2019-110521GB-I00 and Catalan government grant 2017-1005. French National Research Agency (ANR) in the frame of the Cluster of Excellence COTE (project HydroBeech, ANR-10-LABX-45). VLIR-OUS, under the Institutional University Coorperation programme (IUC) with Mountains of the Moon University. Project LAS III 77/2017/B entitled: \"Estimation of net carbon dioxide fluxes exchanged between the forest ecosystem on post-agricultural land and between the tornado-damaged forest area and the atmosphere using spectroscopic and numerical methods\", source of funding: General Directorate of State Forests, Warsaw, Poland. Max Planck Society (Germany), RFBR, Krasnoyarsk Territory and Krasnoyarsk Regional Fund of Science, project number 20-45-242908. Estonian Research Council (PRG609), and the European Regional Development Fund (Centre of Excellence EcolChange). Canada-Denmark Arctic Research Station Early Career Scientist Exchange Program, from Polar knowledge Canada (POLAR) and the Danish Agency for Science and Higher Education. AP received funding from Fondecyt 1180205, CONICYT PIA AFB170008 and ANID PIA / BASAL FB210006. MP received funding from the Funding Org. Knut and Alice Wallenberg Foundation (grant nr. 2015.0047), and acknowledges funding from the Swedish Research Council (VR) with contributing research institutes to both the SITES and ICOS Sweden infrastructures. JP and RO were funded by the Spanish Ministry of Science grant PID2019-110521GB-I00, the fundacion Ramon Areces grant ELEMENTAL-CLIMATE, and the Catalan government grant 2017-1005. MPB received funding from the Svalbard Environmental Protection Fund (grant project number 15/128) and the Research Council of Norway (Arctic Field Grant, project number 269957). RP received funding from the Ministry of Education, Youth and Sports of the Czech Republic (grant INTER-TRANSFER nr. LTT20017). LTSER Zone Atelier Alpes; Federation FREE-Alpes. RP received funding from a Humboldt Fellowship for Experienced Researchers. Prokushkin AS and Zyryanov VI contribution has been supported by the RFBR grant #18-05-60203-Arktika. RPu received founding from the Polish National Science Centre (grant project number 2017/27/B/NZ8/00316). ODYSSEE project (ANR-13-ISV7-0004, PN-II-ID-JRP-RO-FR-2012). KR was supported through an Australian Government Research Training Program Scholarship. Fieldwork was supported by the Global Challenges program at the University of Wollongong, the ARC the Australian Antarctic Division and INACH. DR was funded by the project SUBANTECO IPEV 136 (French Polar Institute Paul-Emile Victor), Zone Atelier CNRS Antarctique et Terres Australes, SAD Region Bretagne (Project INFLICT), BiodivERsa 2019-2020 BioDivClim call 'ASICS' (ANR-20-EBI5-0004). SAR received funding from the Australian Research Council. NSF grant #1556772 to the University of Notre Dame. Pavia University (Italy). OR received funding from EU-LEAP-Agri (RAMSES II), EU-DESIRA (CASSECS), EU-H2020 (SustainSahel), AGROPOLIS and TOTAL Foundations (DSCATT), CGIAR (GLDC). AR was supported by the Russian Science Foundation (Grant 18-74-10048). Parc national des Ecrins. JS received funding from Vetenskapsradet grant nr (No: 2014-04270), ALTER-net multi-site grant, River LIFE project (LIFE08 NAT/S/000266), Flexpeil. Helmholtz Association long-term research program TERENO (Terrestrial Environmental Observatories). PS received funding from the Polish Ministry of Science and Higher Education (grant nr. N N305 304840). AS acknowledges funding by ETH Zurich project FEVER ETH-27 19-1. LSC received funding from NSERC Canada Graduate Scholarship (Doctoral) Program; LSC was also supported by ArcticNet-NCE (insert grant #). Conselho Nacional de Desenvolvimento Cientifico e Tecnologico (141513/2017-9); FundacAo Carlos Chagas Filho de Amparo a Pesquisa do Estado do Rio de Janeiro (E26/200.84/2019). ZS received funding from the SRDA (grants nos. APVV-16-0325 and APVV-20-0365) and from the ERDF (grant no. ITMS 313011S735, CE LignoSilva). JS, MB and CA received funding from core budget of ETH Zurich. State excellence Program M-V \"WETSCAPES\". AfricanBioServices project funded by the EU Horizon 2020 grant number 641918. The authors from KIT/IMK-IFU acknowledge the funding received within the German Terrestrial Environmental Observatories (TERENO) research program of the Helmholtz Association and from the Bavarian Ministry of the Environment and Public Health (UGV06080204000). Deutsche Forschungsgemeinschaft (DFG, German Research Foundation), project number 192626868, in the framework of the collaborative German-Indonesian research project CRC 990 (SFB): 'EFForTS, Ecological and Socioeconomic Functions of Tropical Lowland Rainforest Transformation Systems (Sumatra, Indonesia)'. MS received funding from the Ministry of Education, Youth and Sports of the Czech Republic (grant nr. INTER-TRANSFER LTT19018). TT received funding from the Swedish National Space Board (SNSB Dnr 95/16) and the CASSECS project supported by the European Union. HJDT received funding from the UK Natural Environment Research Council (NERC doctoral training partnership grant NE/L002558/1). German Science Foundation (DFG) GraKo 2010 \"Response\". PDT received funding from the MEMOIRE project (PN-III-P1-1.1-PD2016-0925). Arctic Challenge for Sustainability II (ArCS II; JPMXD1420318865). JU received funding from Czech Science Foundation (grant nr. 21-11487S). TU received funding from the Romanian Ministry of Education and Research (CCCDI - UEFISCDI -project PN-III-P2-2.1-PED-2019-4924 and PN2019-2022/19270201-Ctr. 25N BIODIVERS 3-BIOSERV). AV acknowledge funding from RSF, project 21-14-00209. GFV received funding from the Dutch Research Council NWO (Veni grant, no. 863.14.013). Australian Research Council Discovery Early Career Research Award DE140101611. FGAV received funding from the Portuguese Science Foundation (FCT) under CEECIND/02509/2018, CESAM (UIDP/50017/2020+UIDB/50017/2020), FCT/MCTES through national funds, and the co-funding by the FEDER, within the PT2020 Partnership Agreement and Compete 2020. Ordesa y Monte Perdido National Park. MVI received funding from the Spanish Ministry of Science and Innovation through a doctoral grant (FPU17/05869). JW received funding from the Czech Science Foundation (grant nr. 20-28119S) and the Czech Academy of Sciences (grant nr. RVO 67985939). CR and SW received funding from the Swiss Federal Office for the Environment (FOEN) and the de Giacomi foundation. YY received funding from the National Natural Science Foundation of China (Grant no. 41861134039 and 41941015). ZY received funding from the National Natural Science Foundation of China (grant nr. 41877458). FZ received funding from the Swiss National Science Foundation (grant nr. 172198 and 193645). PZ received funding from the Funding Org. Knut and Alice Wallenberg Foundation (grant no. 2015.0047). JL received funding from (i) the Agence Nationale de la Recherche (ANR), under the framework of the young investigators (JCJC) funding instrument (ANR JCJC Grant project NoANR-19-CE32-0005-01: IMPRINT) (ii) the Centre National de la Recherche Scientifique (CNRS) (Defi INFINITI 2018: MORFO); and the Structure Federative de Recherche (SFR) Condorcet (FR CNRS 3417: CREUSE). Fieldwork in the Arctic got facilitated by funding from the EU INTERACT program. SN, UAT, JJA and JvO would like to thank the field team of the Vegetation Dynamics group for their efforts and hard work. We acknowledge Dominique Tristan for letting access to the field. For the logistic support the crew of INACH and Gabriel de Castilla Station team on Deception Island. We thank the Inuvialuit and Kluane First Nations for the opportunity to work on their land. MAdP acknowledges fieldwork assistance and logistics support to Unidad de Tecnologia Marina CSIC, and the crew of Juan Carlos I and Gabriel de Castilla Spanish Antarctic Stations, as well as to the different colleagues from UAH that helped on the instrument maintenance. ERF acknowledges fieldwork assistance by Martin Heggli. MBG acknowledges fieldwork and technical assistance by P Abadia, C Benede, P Bravo, J Gomez, M Grasa, R Jimenez, H Miranda, B Ponz, J Revilla and P Tejero and the Ordesa and Monte Perdido National Park staff. LH acknowledges field assistance by John Jacobs, Andrew Trant, Robert Way, Darroch Whitaker; we acknowledge the Inuit of Nunatsiavut, and the Co-management Board of Torngat Mountains National Park for their support of this project and acknowledge that the field research was conducted on their traditional lands. We thank our many bear guides, especially Boonie, Eli, Herman, John and Maria Merkuratsuk. AAK acknowledges field support of Akhtar Malik, Rameez Ahmad. Part of microclimatic records from Saxony was funded by the Saxon Switzerland National Park Administration. Tyson Research Center. JP acknowledges field support of Emmanuel Malet (Edytem) and Rangers of Reserves Naturelles de Haute-Savoie (ASTERS). Practical help: Roel H. Janssen, N. Huig, E. Bakker, Schools in the tepaseforsoket, Forskar fredag, Erik Herberg. The support by the Bavarian Forest National Park administration is highly appreciated. LvdB acknowledges CONAF and onsite support from the park rangers from PN Pan de Azucar, PN La Campana, PN Nahuelbuta and from communidad agricola Quebrada de Talca. JL and FS acknowledge Manuel Nicolas and all forest officers from the Office National des Forets (ONF) who are in charge of the RENECOFOR network and who provided help and local support for the installation and maintenance of temperature loggers in the field., Research in global change ecology relies heavily on global climatic grids derived from estimates of air temperature in open areas at around 2 m above the ground. These climatic grids do not reflect conditions below vegetation canopies and near the ground surface, where critical ecosystem functions occur and most terrestrial species reside. Here, we provide global maps of soil temperature and bioclimatic variables at a 1-km2 resolution for 0–5 and 5–15 cm soil depth. These maps were created by calculating the difference (i.e. offset) between in situ soil temperature measurements, based on time series from over 1200 1-km2 p ixels ( summarized f rom 8 519 u nique t emperature sensors) across all the world's major terrestrial biomes, and coarse-grained air temperature estimates from ERA5-Land (an atmospheric reanalysis by the European Centre for Medium-Range Weather Forecasts). We show that mean annual soil temperature differs markedly from the corresponding gridded air temperature, by up to 10°C (mean = 3.0 ± 2.1°C), with substantial variation across biomes and seasons. Over the year, soils in cold and/or dry biomes are substantially warmer (+3.6 ± 2.3°C) than gridded air temperature, whereas soils in warm and humid environments are on average slightly cooler (−0.7 ± 2.3°C). The observed substantial and biome-specific offsets emphasize that the projected impacts of climate and climate change on near-surface biodiversity and ecosystem functioning are inaccurately assessed when air rather than soil temperature is used, especially in cold environments. The global soil-related bioclimatic variables provided here are an important step forward for any application in ecology and related disciplines. Nevertheless, we highlight the need to fill remaining geographic gaps by collecting more in situ measurements of microclimate conditions to further enhance the spatiotemporal resolution of global soil temperature products for ecological applications., FWO G018919N W001919N 12P1819N, DOB Ecology, University of Helsinki, Faculty of Science (MICROCLIM) 7510145, European Research Council (ERC) FORMICA 757833, Arctic Interactions at the University of Oulu, Academy of Finland 318930 337552, Maaja vesitekniikan tuki ry., Tiina and Antti Herlin Foundation, Nordenskiold Samfundet, Societas pro Fauna et Flora Fennica, Grant Agency of the Czech Republic 20-28119S 20-05840Y GA17-19376S 21-11487S, Czech Academy of Sciences RVO 67985939, National Geographic Society 9480-14 WW-240R-17, CISSC (program ICRP) 2397, Iran National Science Foundation (INSF) 96005914, Scottish Government's Rural and Environment Science and Analytical Services Division, Qatar Petroleum QUEX-CAS-QP-RD-18/19, European Union's Horizon 2020 research and innovation program 678841, Swiss National Science Foundation (SNSF), European Commission 172198 193645 31003A_176044, Ministry of Education, Youth & Sports - Czech Republic LTAUSA19137, Ministry of Science and Higher Education of the Russian Federation FSRZ-2020-0014, Independent Research Fund Denmark 8021-00423B 7027-00133B, German Research Foundation (DFG) DFG- FZT 118 202548816 TI 338/14-1 TI 338/14-2 BA 3843/6-1, grant project VEGA of the Ministry of Education of the Slovak Republic Slovak Academy of Sciences 2/0132/18, Forestry Commission, Universidad Javeriana, Direccion General de Cambio Climatico del Gobierno de Aragon, European Union's Horizon 2020 research and innovation program under the Marie Skodowska-Curie Grant 657627 SNF 407340_172433 40FA40_154245 20FI21_148992 20FI20_173691, European Commission 17841 774124, MCTI/CNPq/FNDCT 68/2013, Project 'Como as florestas da Amazonia Central respondem as variacoes climaticas? Efeitos sobre dinamica florestal e sinergia com a fragmentacAo florestal', Spanish Government, European Commission CGL2016-78093-R, ANID-FONDECYT 1181745, National Science Foundation, Poland UMO-2017/27/B/ST10/02228, National Research Foundation - South Africa, Australian Research Council, Slovak Research and Development Agency APVV-19-0319, Instituto Antartico Chileno INACH_RT-48_16 INACH FR-0418, Comision Nacional de Investigacion Cientifica y Tecnologica (CONICYT) PIA APOYO CCTE AFB170008 PIA AFB170008, UK Research & Innovation (UKRI), Natural Environment Research Council (NERC), Research Council of Norway, European Commission 230970, NERC E3 doctoral training partnership grant at the University of Edinburgh NE/L002558/1, Carnegie Trust for the Universities of Scotland, Gobern of Spain PERMAPLANET CTM2009-10165-E ANTARPERMA CTM2011-15565-E PERMASNOW CTM2014-52021-R, University of Alcala, Spanish Polar Committee, Autonomous Province of Bolzano (ITA), ScotNature, NERC National Capability LTS-S: UK-SCAPE NE/R016429/1, Ministry of Education, Youth & Sports - Czech Republic LTAUSA18007, Kempe Foundation JCK-1112 JCK-1822, Ministry of Education, Youth and Sports of the Czech Republic within the National Sustainability Programme I (NPU I) LO1415, project for national infrastructure support CzeCOS/ICOS LM2015061 GLORIA-EU EVK2-CT2000-00056, Tiroler Wissenschaftsfonds, University of Innsbruck, Sime Darby Foundation, Jardin Botanico Atlantico SV-20-GIJON-JBA, Federal Ministry of Education & Research (BMBF) 01LL0912 01LG1201M 01LG1201N, Comision Nacional de Investigacion Cientifica y Tecnologica (CONICYT) CONICYT FONDECYT 11170516 1180205, ANID PIA / BASAL FB210006, National Parks (DYNBIO) 1656/2015, Spanish Research Agency (VULBIMON) CGL2017-90040-R, Swiss National Science Foundation (SNSF) 20FI20_173691, Biotechnology and Biological Sciences Research Council (BBSRC) 206/D16053 FWO G0H1517N, EU Horizon2020 INFRAIA project eLTER-PLUS 871128, project LTER-CWN (FFG, F&E Infrastrukturforderung) 858024, Austrian Climate Research Program ACRP7 - CentForCSink - KR14AC7K11960, iDiv by the German Research Foundation DFG- FZT 118 202548816, Baden-Wurttemberg Ministry of Science, Research and Arts, Weston Foundation, ArcticNet, Natural Sciences and Engineering Research Council of Canada (NSERC) RGPIN-06691, Villum Foundation 17523, Ministry of Education, Youth & Sports - Czech Republic LM2015078 VAN2020/01 CZ.02.1.01/0.0/0.0/16_013/0001708 LTT17033 LTT20017 INTER-TRANSFER LTT19018, Bolin Centre for Climate Research, Stockholm University, Swedish Research Council Swedish Research Council Formas 2014-00530 2018-00792 2016-01187, Swedish Forest Society Foundation 2018-485-Steg 2 2017, Federal Ministry of Education & Research (BMBF) FKZ 031B0516C SUSALPS, Oberfrankenstiftung OFS FP00237, Energy Research Fund NYR-11 - 2019 NYR-18 - 2020, UK NERC Independent Research Fellowship NE/S01537X/1, National Science Centre, Poland 2016/21/B/ST10/02271, Polish National Centre for Research and Development Pol-Nor/203258/31/2013, MoEFCC, Govt of India (AICOPTAX project) 22018/12/2015/RE/Tax, Swedish Research Council Formas 2018-01781 2018-02700 2019-00836, research infrastructure ICOS-SE, National Research, Development and Innovation Fund of Hungary K128441, Programa Operativo FEDER 2018 B1-RNM-163-UGR-18, Norwegian Research Council (NORKLIMA grants) 184912 244525, CONICYT-PAI 79170119, ANID-MPG 190029, project MIUR PON Cluster OT4CLIMA, Stelvio National Park, AIAS-COFUND fellowship programme - Marie Skodowska- Curie actions under the European Union's Seventh Framework Pro-gramme for Research, Technological development and Demonstration 609033, Aarhus University Research Foundation, Denmark, EU FP6 NitroEurope 17841, EU FP7 ECLAIRE 282910, Ministry of Education and Science of Ukraine 505 550 574 602, GEF-UNEP NEC05348, ENI CBC BSB PONTOS BSB 889, Netherlands Organization for Scientific Research (NWO) 016.VICI.170.072, New Frontiers in Research Fund-Exploration NFRF-2018-02043, Natural Sciences and Engineering Research Council of Canada (NSERC), Australian Research Council DE180100570, National Science Foundation (NSF) DEB 1557094, International Center for Advanced Renewable Energy and Sustainability (I-CARES) at Washington University in St. Louis, Smithsonian Institution Smithsonian Tropical Research Institute, Tyson Research Center, UK Natural Environment Research Council through the ShrubTundra Project NE/M016323/1, Swedish Research Council Formas Swedish Research Council, Kempe Foundations - research infrastructure ICOS Kempe Foundations - research infrastructure SITES, Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET) PIP 112-201501-00609, Spanish Government PID2019-110521GB-I00, Catalan government 2017-1005, French National Research Agency (ANR) ANR-10-LABX-45, General Directorate of State Forests, Warsaw, Poland, Max Planck Society, Russian Foundation for Basic Research (RFBR), Krasnoyarsk Territory Krasnoyarsk Regional Fund of Science 20-45-242908, Estonian Research Council PRG609, Knut & Alice Wallenberg Foundation 2015.0047, Swedish Research Council, fundacion Ramon Areces grant ELEMENTAL-CLIMATE, Svalbard Environmental Protection Fund 15/128, Research Council of Norway 269957, Humboldt Fellowship for Experienced Researchers, Russian Foundation for Basic Research (RFBR) 18-05-60203-Arktika, Polish National Science Centre 2017/27/B/NZ8/00316, ODYSSEE project (PN-II-ID-JRP-RO-FR-2012) ANR-13-ISV7-0004, Australian Government, Department of Industry, Innovation and Science, Global Challenges program at the University of Wollongong, ARC the Australian Antarctic Division, INACH, project SUBANTECO IPEV 136 (French Polar Institute Paul-Emile Victor), Zone Atelier CNRS Antarctique et Terres Australes, SAD Region Bretagne (Project INFLICT), BiodivERsa 2019-2020 BioDivClim call 'ASICS' ANR-20-EBI5-0004, National Science Foundation (NSF) 1556772, EU-LEAP-Agri (RAMSES II) EU-DESIRA (CASSECS) EU-H2020 (SustainSahel), AGROPOLIS, Total SA, CGIAR, Russian Science Foundation (RSF) 18-74-10048, Swedish Research Council 2014-04270, ALTER-net multi-site grant, River LIFE project LIFE08 NAT/S/000266, Flexpeil, Ministry of Science and Higher Education, Poland N N305 304840, ETH Zurich FEVER ETH-27 19-1, NSERC Canada Graduate Scholarship (Doctoral) Program, ArcticNet-NCE, Conselho Nacional de Desenvolvimento Cientifico e Tecnologico (CNPQ) 141513/2017-9, Fundacao Carlos Chagas Filho de Amparo a Pesquisa do Estado do Rio De Janeiro (FAPERJ) E26/200.84/2019, SRDA APVV-16-0325 APVV-20-0365, ERDF (CE LignoSilva) ITMS 313011S735, ETH Zurich, EU Horizon 2020 641918, German Terrestrial Environmental Observatories (TERENO) research program of the Helmholtz Association, Bavarian Ministry of the Environment and Public Health UGV06080204000 German Research Foundation (DFG) 192626868, Swedish National Space Board (SNSB) 95/16, CASSECS project by the European Union, Natural Environment Research Council (NERC) NE/L002558/1, MEMOIRE project PN-III-P1-1.1-PD2016-0925, Arctic Challenge for Sustainability II (ArCS II) JPMXD1420318865, Consiliul National al Cercetarii Stiintifice (CNCS), Unitatea Executiva pentru Finantarea Invatamantului Superior, a Cercetarii, Dezvoltarii si Inovarii (UEFISCDI) PN-III-P2-2.1-PED-2019-4924 PN2019-2022/19270201, 25N BIODIVERS 3-BIOSERV, Russian Science Foundation (RSF) 21-14-00209., Netherlands Organization for Scientific Research (NWO) 863.14.013, Australian Research Council DE140101611, Portuguese Foundation for Science and Technology CEECIND/02509/2018 CESAM UIDP/50017/2020+UIDB/50017/2020, Portuguese Foundation for Science and Technology European Commission, FEDER, within the PT2020 Partnership Agreement, Compete 2020, Spanish Government FPU17/05869, Swiss Federal Office for the Environment (FOEN), Giacomi foundation, National Natural Science Foundation of China (NSFC) 41861134039 41941015 41877458, French National Research Agency (ANR) ANR-19-CE32-0005-01 Centre National de la Recherche Scientifique (CNRS), Structure Federative de Recherche (SFR) Condorcet (FR CNRS 3417: CREUSE), EU INTERACT program, Inuit of Nunatsiavut, Co-management Board of Torngat Mountains National Park, Saxon Switzerland National Park Administration, Bavarian Forest National Park administration, BECC - Biodiversity and Ecosystem services in a Changing Climate, Research Foundation Flanders (FWO-SBO) S000619N

The following authors were omitted from the original version of this Data Descriptor: Markus Reichstein and Nicolas Vuichard. Both contributed to the code development and N. Vuichard contributed to the processing of the ERA-Interim data downscaling. Furthermore, the contribution of the co-author Frank Tiedemann was re-evaluated relative to the colleague Corinna Rebmann, both working at the same sites, and based on this re-evaluation a substitution in the co-author list is implemented (with Rebmann replacing Tiedemann). Finally, two affiliations were listed incorrectly and are corrected here (entries 190 and 193). The author list and affiliations have been amended to address these omissions in both the HTML and PDF versions.

Research in environmental science relies heavily on global climatic grids derived from estimates of air temperature at around 2 meter above ground1-3. These climatic grids however fail to reflect conditions near and below the soil surface, where critical ecosystem functions such as soil carbon storage are controlled and most biodiversity resides4-8. By using soil temperature time series from over 8500 locations across all of the world’s terrestrial biomes4, we derived global maps of soil temperature-related variables at 1 km resolution for the 0–5 and 5–15 cm depth horizons. Based on these maps, we show that mean annual soil temperature differs markedly from the corresponding 2 m gridded air temperature, by up to 10°C, with substantial variation across biomes and seasons. Soils in cold and/or dry biomes are annually substantially warmer (3.6°C ± 2.3°C) than gridded air temperature, whereas soils in warm and humid environments are slightly cooler (0.7 ± 2.3°C). As a result, annual soil temperature varies less (by 17%) across the globe than air temperature. The effect of macroclimatic conditions on the difference between soil and air temperature highlights the importance of considering that macroclimate warming may not result in the same level of soil temperature warming. Similarly, changes in precipitation could alter the relationship between soil and air temperature, with implications for soil-atmosphere feedbacks9. Our results underpin that the impacts of climate and climate change on biodiversity and ecosystem functioning are inaccurately assessed when air rather than soil temperature is used, especially in cold environments.

The FLUXNET2015 dataset provides ecosystem-scale data on CO2, water, and energy exchange between the biosphere and the atmosphere, and other meteorological and biological measurements, from 212 sites around the globe (over 1500 site-years, up to and including year 2014). These sites, independently managed and operated, voluntarily contributed their data to create global datasets. Data were quality controlled and processed using uniform methods, to improve consistency and intercomparability across sites. The dataset is already being used in a number of applications, including ecophysiology studies, remote sensing studies, and development of ecosystem and Earth system models. FLUXNET2015 includes derived-data products, such as gap-filled time series, ecosystem respiration and photosynthetic uptake estimates, estimation of uncertainties, and metadata about the measurements, presented for the first time in this paper. In addition, 206 of these sites are for the first time distributed under a Creative Commons (CC-BY 4.0) license. This paper details this enhanced dataset and the processing methods, now made available as open-source codes, making the dataset more accessible, transparent, and reproducible. Fil: Pastorello, Gilberto. Lawrence Berkeley National Laboratory. Computational Research Division; Estados Unidos Fil: Trotta, Carlo. University of Tuscia. DiBAf; Italia Fil: Canfora, Eleonora. University of Tuscia. DiBAf; Italia. Euro-Mediterranean Centre on Climate Change Foundation (CMCC); Italia Fil: Chu, Housen. Lawrence Berkeley national Laboratory. Climate & Ecosystem Sciences Division; Estados Unidos Fil: Christianson, Danielle. Lawrence Berkeley National Laboratory. Computational Research Division; Estados Unidos Fil: Cheah, You - Wei. Lawrence Berkeley National Laboratory. Computational Research Division; Estados Unidos Fil: Poindexter, Cristina. California State University. Department of Civil Engineering; Estados Unidos Fil: Chen, Jiquan. Michigan State University. Department of Geography, Environment, and Spatial Sciences; Estados Unidos Fil: Elbashandy, Abdelrah man. Lawrence Berkeley National Laboratory. Computational Research Division; Estados Unidos Fil: Humphrey, Marty. University of Virginia. Department of Computer Science; Estados Unidos Fil: Isaac, Peter. TeRn Ecosystrem Processes; Australia Fil: Polidori, Diego. University of Tuscia. DiBAf; Italia. Euro-Mediterranean Centre on Climate Change Foundation (CMCC); Italia Fil: Riveca, Alessio. University of Tuscia. DiBAf; Italia. Euro-Mediterranean Centre on Climate Change Foundation (CMCC); Italia Fil: van Ingen, Catharine. Lawrence Berkeley National Laboratory. Computational Research Division; Estados Unidos Fil: Zhang, Leiming. Chinese Academy of Sciences. Institute of Geographic Sciences and Natural Resources Research. Key Laboratory of Ecosystem Network Observation and Modeling; China Fil: Amiro, Brian. University of Manitoba. Department of Soil Science; Canadá Fil: Ammann, Christof. Agroscope Research Institute. Department of Agroecology and Environment; Suiza Fil: Altaf Arain, M. McMaster University. School of Geography and Earth Sciences; Canadá. Fil: Ardo, Jonas. Lund University. Department of Physical Geography and Ecosystem Science; Suecia Fil: Arkebauer, Timothy. University of Nebraska-Lincoln. Department of Agronomy and Horticulture; Estados Unidos Fil: Arndt, Stefan K. The University of Melbourne. School of Ecosystem and Forest Sciences; Australia Fil: Arriga, Nicola. University of Antwerp. Department of Biology, Research Group PLECO; Bélgica. European Commission. Joint Research Centre; Italia Fil: Aubinet, Marc. University of Liege. TeRRA Teaching and Research Center; Bélgica Fil: Aurela, Mika. Finnish Meteorological Institute; Finlandia Fil: Baldocchi, Dennis. University of California Berkeley. ESPM; Estados Unidos Fil: Barr, Alan. University of Saskatchewan. Global Institute for Water Security; Canadá. Environment and Climate Change Canada. Climate Research Division; Canadá. Fil: Beamesderfer, Eric. McMaster University. School of Geography and Earth Sciences; Canadá. Fil: Belelli Marchesini, Luca. Fondazione Edmund Mach. Research and Innovation Centre. Department of Sustainable Agro-ecosystems and Bioresources; Italia. RUDN University. Agrarian-Technological Institute. Department of Landscape Design and Sustainable Ecosystems; Rusia Fil: Bergeron, Onil. Ministère du Développement durable de l’Environnement et de la Lutte contre les Changements Climatiques. Direction du Marché du Carbone; Canadá. Fil: Beringer, Jason. University of Western Australia. School of Agriculture and Environment; Australia. Fil: Bernhofer, Christia. Technische Universität Dresden. Institute of Hydrology and Meteorology; Alemania Fil: Berveiller, Daniel. Université Paris-Saclay, CNRS, AgroParisTech, Ecologie Systématique et Evolution; Francia Fil: Billesbach, Dave. University of Nebraska-Lincoln. Biological Systems Engineering; Estados Unidos Fil: Black, Tomas Andrew. University of British Columbia. Faculty of Land and Food Systems; Canadá. Fil: Blanken, Peter D. University of Colorado. Department of Geography; Estados Unidos Fil: Bohrer, Gil. Ohio State University. Department of Civil, Environmental & Geodetic Engineering; Estados Unidos Fil: Boike, Julia Alfred. Wegener Institute. Helmholtz Centre for Polar and Marine Research; Alemania. Humboldt-Universität zu Berlin. Geography Department; Alemania Fil: Bolstad, Paul V. University of Minnesota. Forest Resources; Estados Unidos Fil: Bonal, Damien. Université de Lorraine, AgroParisTech, INRAE, UMR Silva; Francia Fil: Bonnefond, Jean - Marc. ISPA, Bordeaux Sciences Agro, INRAE; Francia Fil: Bowling, David R. University of Utah. School of Biological Sciences; Estados Unidos Fil: Bracho, Rosuel. University of Florida.School of Forest Resources and Conservation; Estados Unidos Fil: Brodeur, Jason. McMaster University. McMaster University Library; Estados Unidos Fil: Brummer, Christian. Federal Research Institute of Rural Areas, Forestry and Fisheries. Thünen Institute of Climate-Smart Agriculture; Alemania Fil: Buchmann, Nina. ETH. Department of Environmental Systems Science; Suiza Fil: Burban, Benoit. INRAE UMR ECOFOG; Guyana Francesa Fil: Burns, Sean P. University of Colorado. Department of Geography; Estados Unidos. National Center for Atmospheric Research. Mesoscale and Microscale Meteorology Laboratory; Estados Unidos Fil: Buysse, Pauline. Université Paris-Saclay, INRAE, AgroParisTech, UMR ECOSYS; Francia Fil: Cale, Peter. Australian Landscape Trust; Australia Fil: Cavagna, Mauro. Fondazione Edmund Mach. Research and Innovation Centre. Department of Sustainable Agro-ecosystems and Bioresources; Italia Fil: Cellier, Pierre. Université Paris-Saclay. INRAE, AgroParisTech, UMR ECOSYS; Francia Fil: Che, Shiping. Chinese Academy of Sciences. Institute of Botany. State Key Laboratory of Vegetation and Environmental Change; China Fil: Chini, Issac. Fondazione Edmund Mach. Research and Innovation Centre. Department of Sustainable Agro-ecosystems and Bioresources; Italia Fil: Christensen, Storben. Aarhus University. Arctic Research Center. Department of Bioscience; Dinamarca Fil: Cleverly, James. University of Technology. School of Life Sciences; Australia. University of Technology. Terrestrial Ecosystem Research Network; Australia. Fil: Collalti, Alessio. University of Tuscia. DiBAf; Italia. National Research Council of Italy. Institute for Agricultural and Forestry Systems in the Mediterranean; Italia Fil: Consalvo, Claudia. University of Tuscia. DiBAf; Italia. National Research Council of Italy. Research Institute on Terrestrial Ecosystems; Italia Fil: Cook, Bruce. NASA Goddard Space Flight Center. Biospheric Sciences Laboratory; Estados Unidos Fil: Cook, David. Argonne National Laboratory. Environmental Science Division; Estados Unidos Fil: Coursolle, Carole. Natural Resources Canada. Canadian Forest Service; Canadá. Université Laval. Faculté de Foresterie, de Géographie et de Géomatique. Centre d’étude de la Forêt; Canadá Fil: Cremonose, Edoardo. Environmental Protection Agency of Aosta Valley. Climate Change Unit; Italia Fil: Curtis, Peter. Ohio State University. Department of Evolution, Ecology, and Organismal Biology; Estados Unidos Fil: D'Andrea, Ettore. National Research Council of Italy. Institute for Agricultural and Forestry Systems in the Mediterranean; Italia Fil: da Rocha, Humberto. Universidade de São Paulo. Instituto de Astronomia, Geofísica e Ciências Atmosféricas; Brasil Fil: Dai, Xiaoqin. Chinese Academy of Sciences. Institute of Geographic Sciences and Natural Resources Research. Key Laboratory of Ecosystem Network Observation and Modeling; China. Fil: Davis, Kenneth. The Pennsylvania State University. Department of Meteorology and Atmospheric Science; Estados Unidos Fil: De Cinti, Bruno. National Research Council of Italy. Institute of Research on Terrestrial Ecosystems; Italia Fil: de Grandcourt, Agnes. UMR Eco&Sols, CIRAD; Francia Fil: De Ligne, Anne. University of Liege. TeRRA Teaching and Research Center; Bélgica Fil: De Oliveira, Raimundo C. Pedology, Embrapa Amazonia Oriental; Brasil. Fil: Delpierre, Nicolas. Université Paris-Saclay. CNRS, AgroParisTech, Ecologie Systématique et Evolution; Francia Fil: Desai, Ankur R. University of Wisconsin-Madison. Atmospheric and Oceanic Sciences; Estados Unidos Fil: Di Bella, Carlos Marcelo. Universidad de Buenos Aires. Facultad de Agronomía. Departamento de Métodos Cuantitativos y Sistemas de Información; Argentina Fil: di Tommasi, Paul. National Research Council of Italy. Institute for Agricultural and Forestry Systems in the Mediterranean; Italia Fil: Dolman, Han. Vrije Universiteit Amsterdam. Department of Earth Sciences; Holanda Fil: Domingo, Francisco. CSIC. Experimental Station of Arid Zones. Desertification and Geoecology Department; España Fil: Dong, Gang. Shanxi University. School of Life Science; China. Fil: Dore, Sabina. HydroFocus; Estados Unidos Fil: Duce, Pierpaolo. National Research Council of Italy. Institute of Bioeconomy; Italia Fil: Dufrêne, Eric. Université Paris-Saclay, CNRS., AgroParisTech. Ecologie Systématique et Evolution; Francia Fil: Dunn, Allison. Worcester State University. Department of Earth, Environment, and Physics; Estados Unidos Fil: Dušek, Jiri. Global Change Research Institute of the Czech Academy of Sciences. Department of Matter and Energy Fluxes; República Checa Fil: Eamus Derek. University of Technology. School of Life Sciences; Australia. Fil: Eichelmann, Uwe.Technische Universität Dresden. Institute of Hydrology and Meteorology; Alemania Fil: ElKhidir, Hatim abdalla M. elObeid. Agricultural Research Corporation. Research Station; Sudán Fil: Eugster, Wener. ETH. Department of Environmental Systems Science; Suiza Fil: Ewenz, Cacilia M. Airborne Research Australia. TERN Ecosystem Processes Central Node; Australia Fil: Ewers, Brent. University of Wyoming. Department of Botany. Program in Ecology; Estados Unidos Fil: Famulari, Daniela. National Research Council of Italy. Institute for Agricultural and Forestry Systems in the Mediterranean; Italia Fil: Fares, Silvano. National Research Council of Italy. Institute of BioEconomy; Italia Fil: Feigenwinter, Iris. ETH. Department of Environmental Systems Science; Suiza Fil: Feitz, Andrew. Geoscience Australia; Australia Fil: Fensholt, Rasmus. University of Copenhagen. Department of Geosciences and Natural Resource Management; Dinamarca Fil: Filippa, Gianluca. Environmental Protection Agency of Aosta Valley. Climate Change Unit; Italia Fil: Fischer, Marc. Lawrence Berkeley National Laboratory. Energy Analysis & Environmental Impacts Division; Estados Unidos Fil: Frank, John. USDA Forest Service. Rocky Mountain Research Station; Estados Unidos Fil: Galvagno, Marta. Environmental Protection Agency of Aosta Valley. Climate Change Unit; Italia Fil: Gharun, Mana. ETH. Department of Environmental Systems Science; Suiza Fil: Gianelle, Damiano. Fondazione Edmund Mach. Research and Innovation Centre. Department of Sustainable Agro-ecosystems and Bioresources; Italia Fil: Gielen, Bert. University of Antwerp. Department of Biology. Research Group PLECO; Bélgica Fil: Gioli, Beniamino. National Research Council of Italy. Institute of BioEconomy; Italia Fil: Gitelson, Anatoly. University of Nebraska-Lincoln. School of natural Resources; Estados Unidos Fil: Goded, Ignacio. Joint Research Centre, European Commission; Italia Fil: Goeckede, Mathias. Max Planck Institute for Biogeochemistry; Alemania Fil: Goldstein. Allen H. University of California Berkeley. ESPM; Estados Unidos Fil: Gough, Christopher M. Virginia Commonwealth University. Department of Biology; Estados Unidos Fil: Goulden, Michael L. University of California Irvine. Department of Earth System Science; Estados Unidos Fil: Graf, Alexander. Agrosphere. (IBG3), Forschungszentrum Jülich; Alemania Fil: Griebel, Anne. The University of Melbourne. School of Ecosystem and Forest Sciences; Australia Fil: Gruening, Carsten. Joint Research Centre, European Commission; Italia Fil: Grünwald, Thomas. Technische Universität Dresden. Institute of Hydrology and Meteorology; Alemania Fil: Hammerle, Albin. University of Innsbruck. Department of Ecology; Austria. Fil: Han, Shijie. Henan University. School of Life Sciences. International Joint Research Laboratory for Global Change Ecology; China. Chinese Academy of Sciences. Institute of Applied Ecology; China. Fil: Han, Xingguo. Chinese Academy of Sciences.Institute of Botany. State Key Laboratory of Vegetation and Environmental Change; China Fil: Hansen, Birger Ulf. University of Copenhagen. Department of Geosciences and Natural Resource Management; Dinamarca Fil: Hanson, Chad. Oregon State University. Department of Forest Ecosystems and Society; Estados Unidos Fil: , Juha Hatakka, Juha. Finnish Meteorological Institute; Finlandia Fil: He, Yongtao. Chinese Academy of Sciences. Institute of Geographic Sciences and Natural Resources Research. Key Laboratory of Ecosystem Network Observation and Modeling; China. University of Chinese Academy of Sciences. College of Resources and Environment; China Fil: Hehn, Markus. Technische Universität Dresden. Institute of Hydrology and Meteorology; Alemania Fil: Heinesch, Bernard. University of Liege. TeRRA Teaching and Research Center; Bélgica Fil: Hinko-Najera, Nina. The University of Melbourne. School of Ecosystem and Forest Sciences; Australia. Fil: Hörtnagl, Lukas. ETH. Department of Environmental Systems Science; Suiza. Fil: Hutley, Lindsay. Charles Darwin University. Research Institute for the Environment and Livelihoods; Australia Fil: Ibrom, Andreas. Technical University of Denmark. Department of Environmental Engineering; Dinamarca Fil: Ikawa, Hiroki. National Agriculture and Food Research Organization. Institute for Agro-Environmental Sciences; Japón Fil: Jackowicz-Korczynski, Marcin. Lund University. Department of Physical Geography and Ecosystem Science; Suecia. Aarhus University. Arctic Research Center. Department of Bioscience; Dinamarca Fil: Janouš, Dalibor. Global Change Research Institute of the Czech Academy of Sciences. Department of Matter and Energy Fluxes; República Checa Fil: Jans, Wilma. Wageningen University and Research. Wageningen Environmental Research; Holanda Fil: Jassal, Rachhpal. University of British Columbia. Faculty of Land and Food Systems; Canadá. Fil: Jiang, Shicheng. Ministry of Education, Northeast Normal University. Key Laboratory of Vegetation Ecology; China Fil: Kato, Tomomichi. Hokkaido University. Research Faculty of Agriculture; Japón. Hokkaido University. GICore; Japón Fil: Khomik, Myroslava. Geography and Environmental Management; Canadá. Fil: Klatt, Janina. Karlsruhe Institute of Technology. Institute of Meteorology and Climate Research; Alemania Fil: Knohl, Alexander. University of Goettingen. Bioclimatology; Alelmania. University of Goettingen. Centre of Biodiversity and Sustainable Land Use; Alemania Fil: Knox, Sara. The University of British Columbia. Department of Geography; Canadá Fil: Kobayashi, Hideki. Japan Agency for Marine-Earth Science and Technology. Research Institute for Global change, Institute of Arctic Climate and Environment Research; Japón Fil: Koerber, Georgia. University of Adelaide. Biological Sciences; Australia. Fil: Kolle, Olaf. Max Planck Institute for Biogeochemistry; Alemania Fil: Kosugi, Yoshiko. Kyoto University. Graduate School of Agriculture; Japón Fil: Kotani, Ayumi. Nagoya University. Graduate School of Bioagricultural Sciences; Japón Fil: Kowalski, Andrew. University of Granada. Department of Applied Physics; España Fil: Kruijt, Bart. Wageningen University, Wageningen. Water systems and Global Change group; Holanda Fil: Kurbatova; Julia A. Russian Academy of Sciences. Severtsov institute of Ecology and Evolution; Rusia Fil: Kutsch, Werner L. Integrated Carbon Observation System (ICOS ERIC). Head Office; Finlandia Fil: Kwon, Hyojung. Oregon State University. Department of Forest Ecosystems and Society; Estados Unidos Fil: Launiainen, Samuli. Natural Resources Institute Finland; Finlandia Fil: Laurila, Tuomas. Finnish Meteorological Institute; Finlandia Fil: Law, Bev. Oregon State University. Department of Forest Ecosystems and Society; Estados Unidos Fil: Leuning. Ray. Lawrence Berkeley National Laboratory. Computational Research Division; Estados Unidos Fil: Li, Yingnian. Chinese Academy of Sciences. Northwest institute of Plateau Biology. Key Laboratory of Adaptation and Evolution of Plateau Biota; China Fil: Liddell, Michael. James Cook University. Centre for Tropical Environmental Sustainability Studies; Australia. Fil: Limousin, Jean-Marc. CEFE, CNRS, Univ Montpellier: Francia Fil: Lion, Marryanna. Forest Research Institute Malaysia. Forestry and Environment Division; Malasia Fil: Lohila, Annalea. University of Helsinki. Institute for Atmosphere and Earth System Research/Physics; Finlandia Fil: López-Ballesteros, Ana. Trinity College Dublin. School of Natural Sciences. Department of Botany; Irlanda Fil: López-Blanco, Efren. Aarhus University. Arctic Research Center. Department of Bioscience; Dinamarca Fil: Loubet, Benjamin. Université Paris-Saclay. INRAE, AgroParisTech, UMR ECOSYS; Francia Fil: Lucas-Moffat, Antje. Centre for Agrometeorological Research. German Meteorological Service; Alemania Fil: Lüers, Johannes. University of Bayreuth. Micrometeorology; Alemania. Bayreuth Center of Ecology and Environmental Research; Alemania Fil: Ma, Siyan. University of California Berkeley. ESPM; Estados Unidos Fil: Macfarlane, Craig. CSIRO Land and Water; Australia Fi: Magliulo, Vincenzo. Lawrence Berkeley National Laboratory. Computational Research Division; Estados Unidos Fil: Mammarella, Ivan. University of Helsinki . Institute for Atmosphere and Earth System Research/Physics; Finlandia Fil: Manca, Giovanni. Joint Research Centre, European Commission; Italia Fil: Marras, Serena. University of Sassari. Department of Agriculture; Italia Fil: Massman, William. USDA Forest Service. Rocky Mountain Research Station; Estados Unidos Fil: Mastepanov, Mikhail. University of Oulu. Oulanka Research Station; Finlandia Fil: Matamala, Roser. Argonne National Laboratory. Environmental Science Division; Estados Unidos Fil: Matthes, Jaclyn Hatala. Wellesley College. Department of Biological Sciences; Estados Unidos Fil: , Mazzenga, Francesco. National Research Council of Italy. Research Institute on Terrestrial Ecosystems; Italia Fil: McCaughey, Harry. Queen’s University. Department of Geography and Planning; Canadá. Fil: McHugh, Ian. The University of Melbourne. School of Ecosystem and Forest Sciences; Australia. Fil: McMillan, Andrew M.S. Environmental Analytics nZ, Ltd.; Nueva Zelanda Fil: Merbold, Lutz. International Livestock Research Institute. Mazingira Centre; Kenia. Fil: Meyer, Wayne. University of Adelaide. Biological Sciences; Australia. Fil: Meyers, Tilden. NOAA/OAR/Air Resources Laboratory; Estados Unidos Fil: Miller, Scott D. State University of New York at Albany. Atmospheric Sciences Research Center; Estados Unidos Fil: Minerbi, Stefano. Forest Department of South Tyrol; Italia Fil: Monson, Russell K. University of Arizona. Department of Ecology and Evolutionary Biology; Estados Unidos Fil: Montagnani, Leonardo. Forest Department of South Tyrol; Italia. Free University of Bolzano. Faculty of Science and Technology; Italia Fil: Moore, Caitlin E. University of Illinois at Urbana-Champaign. Department of Plant Biology; Estados Unidos Fil: Moors, Eddy. IHE Delft: Holanda. VU Amsterdam. Faculty of Science; Holanda Fil: Moreaux, Virginie. University Grenoble Alpes; Francia Fil: Moureaux, Christine. University of Liege. TeRRA Teaching and Research Center; Bélgica Fil: Munger, J. William. Harvard University. School of engineering and Applied Sciences; Estados Unidos. Harvard University. Department of Earth and Planetary Sciences; Estados Unidos Fil: Nakai, Taro. National Taiwan University. School of forestry and Resource conservation; Taiwan. University of Alaska Fairbanks. International Arctic Research Center; Estados Unidos Fil: Neirynck, Johan. Research Institute for Nature and Forest. Environment and Climate; Bélgica Fil: Nesic, Zoran. University of British Columbia. 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Agrosphere (IBG3), Forschungszentrum Jülich; Alemania Fil: Schneider, Karl. University of Cologne. Geographical Institute; Alemania Fil: Schrader, Frederik Thünen. Federal Research Institute of Rural Areas, Forestry and Fisheries. Institute of Climate-Smart Agriculture; Alemania Fil: Schroder, Ivan. Geoscience Australia. Department of industry, Innovation and Science; Australia. Fil: Scott, Russell L. USDA-ARS. Southwest Watershed Research Center; Estados Unidos Fil: Sedlák, Pavel. Global Change Research Institute of the Czech Academy of Sciences. Department of Matter and Energy Fluxes; República Checa. Institute of Atmospheric Physics of the Czech Academy of Sciences; República Checa Fil: Serrano-Ortíz, Penélope. Andalusian Institute for Earth System Research (CEAMA-IISTA); España. University of Granada. Department of Ecology; España Fil: Shao, Changliang. Chinese Academy of Agricultural Sciences. National Hulunber Grassland Ecosystem Observation and Research Station & Institute of Agricultural Resources and Regional Planning; China. Fil: Shi, Peili. Chinese Academy of Sciences. Institute of Geographic Sciences and Natural Resources Research. Key Laboratory of Ecosystem Network Observation and Modeling; China. Fil: Shironya, Ivan A.n. Russian Academy of Sciences. Severtsov Institute of Ecology and Evolution; Rusia Fil: Siebicke, Lukas. University of Goettingen. Bioclimatology; Alemania Fil: Šigut, Ladislav. Global Change Research Institute of the Czech Academy of Sciences. Department of Matter and Energy Fluxes; República Checa Fil: Silberstein, Richard. University of Western Australia. School of Agriculture and Environment; Australia. Edith Cowan University. School of Science; Australia. Fil: Sirca, Costantino. Euro-Mediterranean Centre on Climate Change Foundation (CMCC); Italia. University of Sassari. Department of Agriculture; Italia Fil: Spano, Donatella. 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The Integrated Carbon Observation System Re-search Infrastructure aims to provide long-term, continuous observations of sources and sinks of greenhouse gases such as carbon dioxide, methane, nitrous oxide, and water vapour. At ICOS ecosystem stations, the principal technique for measurements of ecosystem-atmosphere exchange of GHGs is the eddy-covariance technique. The establishment and setup of an eddy-covariance tower have to be carefully reasoned to ensure high quality flux measurements being representative of the investigated ecosystem and comparable to measurements at other stations. To fulfill the requirements needed for flux determination with the eddy-covariance technique, variations in GHG concentrations have to be measured at high frequency, simultaneously with the wind velocity, in order to fully capture turbulent fluctuations. This requires the use of high-frequency gas analysers and ultrasonic anemometers. In addition, to analyse flux data with respect to environmental conditions but also to enable corrections in the post-processing procedures, it is necessary to measure additional abiotic variables in close vicinity to the flux measurements. Here we describe the standards the ICOS ecosystem station network has adopted for GHG flux measurements with respect to the setup of instrumentation on towers to maximize measurement precision and accuracy while allowing for flexibility in order to observe specific ecosystem features.