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Löbl, Ivan (2023): Assessing Earth's biotic diversity in natural history museums - gone with the wind? Phegea 51, No. 2: 50-51, DOI: 10.6084/m9.figshare.22722733, URL: http://dx.doi.org/10.6084/m9.figshare.22722733

Members of the scaphidiine tribe Scaphisomatini collected recently in Tianmushan Biosphere Reserve (Zhejiang) and Wuyishan National Nature Reserve (part of Wuyishan Biosphere Reserve, Fujian), China, are studied. Two species, Scaphisoma krali Löbl, sp. nov. and Scaphisoma sekerkai Löbl, sp. nov., are described from Wuyishan. Baeocera franzi (Löbl, 1973) and Scaphisoma mutator Löbl, 2000 are recorded for the first time from Zhejiang province, and Scaphisoma binhanum (Pic, 1922) and Scaphoxium intermedium Löbl, 1984 are recorded for the first time from Fujian province.

Many members of our society ask in formal and informal meetings what is going on in natural history museums and especially in insect collections: little activity, little motivation, and little support. The editor-in-chief of Phegea asked an expert in museum insect collections for his opinion. Dr. Ivan Löbl is based at the natural history museum in Geneva and edited an impressive series of catalogues of Palaearctic Coleoptera https://brill.com/display/serial/CPC

Sapitia versicolor (Pic, 1920) Material examined. 474, Sabah, Batu Punggul Resort env., 24.vi.̅1.vii.1996, 11c, FIT (MHNG); 3, Sabah, Danum Valley, 14̅ 16.ii.1968, G. de Rougemont (MHNG); 1, Sabah, Poring Hot Springs, 500 m, 11.v.1987, Burckhard & Löbl (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05°02.682′ E 117°45.553′, 250 m, iii̅ iv.2005, FIT, primary dipterocarp forest, E. Slade & J. Villanuava lgt. OUMNH 2055 ­062 (OUMNH). Remarks. Thie species appears to be widely distributed and has been reported from Indonesia: Sumatra; East Malaysia: Sabah and Thailand (see LÖBL 2018). It is very similar to S. sumatrana Löbl, 1978 while it differs notably from S. lombokina by the nearly fused mesepimera and the stoud legs., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 16, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2018: Coleoptera: Staphylinidae: Scaphidiinae. World Catalogue of Insects. Volume XVI: i ‾ xvi + 1 ‾ 418 pp."]}

Scaphicoma rufa (Pic, 1923) Material examined. 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 450 m, 22̅ 23.xi.2005, primary dipterocarp forest, west trail area, first inferior FIT, D. Mann, E. Slade & J. Villanuava lgt., OUMNH 2006 ­051 (OUMNH); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, E. Slade & J. Villanuava lgt., selectively logged dipterocarp forest, Yayasan Sabah Logging Concesion, OUMNH 2005 ­052 (MHNG). Remarks. These specimens exhibit characters congruent with the redescription given by LÖBL & LESCHEN (2010). Notable are the widely interrupted basal striae of the elytra, the mesoventrite with distinct mesal ridge, the lateral parts of the metaventrite lacking coarser punctures, and the lateral parts of ventrite I, each bearing a subbasal patch of distinct punctures, in combination with the uniformly rufous, about 1.9 mm long body. The species is very similar to S. arcuatum Champion, 1927 and S. pulex Heller, 1917., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 17, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & LESCHEN R. A. B. 2010: Notes on the Toxidium group (Coleoptera: Staphylinidae: Scaphidiinae). Folia Heyrovskyana Series A 18 (1 - 3): 71 ‾ 93."]}

Pseudobironium rapax sp. nov. (Figs 8 ̅10) Type material. Holotype male, Malaysia, Sabah, Crocker Range, Mawar Waterfall env. 17.VI.1996, 9c, vegetation debris and forest floor litter accumulated along large fallen trees near river (MHNG). Description. Length 1.95 mm, width 1.30 mm. Head and most of body dark reddishbrown, apex of abdomen lighter, yellowish; femora and tibiae roughly as body, tarsi and antennae light brown. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows: III 15: IV 22: V 26: VI 21: VII 26: VIII 26: IX 27: X: 26: XI 35; antennomere XI about 3.5 times as long as wide. Pronotal punctation very fine, similar to that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 4 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation similar to pronotal. Hypomera impunctate, impressed posteriad. Mesoventrite lacking mesal ridge and lacking distinct setiferous punctures. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite lacking microsculpture; median area nearly flat, lacking impressions and tubercle, with two irregular rows of distinct admesal punctures. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, horizontal. Submesocoxal area short, about as fourth shortest interval to metacoxa. Submesocoxal line finely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Ventrite Ilacking microsculpture, with very fine and scattered punctures; following ventrites and exposed tergites with punctulate microsculpture. Male. Aedeagus (Figs 8 ̅10) 0.74 mm long. Median lobe with basal bulb shorter than apical process, the latter slightly inflexed, with sinuate ventral margin (lateral view), and blunt tip; apical part of median lobe lacking distinct valves. Parameres nearly evenly wide, bent in dorsal view, straight in lateral view. Internal sac with pair of strongly sclerotized denticles and mesal sclerite, lacking flagellum and lacking apical spinose structures. Etymology. The species epithet is a Latin noun, referring to the canine­like denticles of the internal sac. Differential diagnosis. The species is as P. irregulare less than 2 mm long and thus conspicuous by its small body­size. It may be easily distinguished from P. irregulare by the evenly very fine elytral punctation, the narrow submesocoxal areas and ventrite I lacking microsculpture. The aedeagal characters suggest relationship with members of the P. subovatum group (see LÖBL & TANG 2013), though the shape of the admesal sclerites of the internal sac is unique. The species falls in the key species to couplet 27, to P. schuhi Löbl & Tang, 2013 from which it differs notably by the parameres evenly narrow, except at their basis. Remarks. The single available specimen has its forelegs broken, probably lost when collected., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 14-15, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Mordelloscaphium Pic, 1915 The genus is monospecific. It was established to accommodate a species possessing characters typical for the termitophilous members of the Baeoceridium group (see LESCHEN & LÖBL 2005) and thus is assumed to live on fungus gardens of termites., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LESCHEN R. A. B. & LOBL I. 2005: Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1 ‾ 63."]}

Bironium borneense Löbl, 1987 Material examined. 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 27̅ 28.x.2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, West trail 10 FIT, OUMNH 2006­051 (OUMNH); 1, with the same data but 21.X.2005, West trail area (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, primary forest, Slade & J. Villanuava lgt. lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005­062 (OUMNH). Remarks. The species is known only from East Malaysia (LÖBL et al. 2020). It was based on specimens found in the Gunung Mulu National Park in Sarawak and subsequently recorted from the Mount Kinabalu National Park, the vicinity of Tambunan and the Batu Punggul Resort in Sabah., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I., LESCHEN R. A. B. & KODADA J. 2020: Review of the Asian species and cladistic analysis of Bironium Csiki (Coleoptera: Staphylinidae: Scaphidiinae) with comments on biogeography. Annales Zoologici (Warszawa) 70 (4): 711 ‾ 736."]}

Vituratella Reitter, 1908 The genus encompasses 31 species, all Afrotropical but the Asian V. termitophila (Champion, 1927) and V. kistneri (Löbl, 1979)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 18, DOI: 10.5281/zenodo.7501705

Scaphicoma Motschulsky, 1863 The genus comprises 19 Afrotropical, Oriental, Melanesian and Australian species. Several, described by J. Achard and M. Pick, still lack revisions. Asingle species is present in samples from Sabah., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 17, DOI: 10.5281/zenodo.7501705

Baeocera incisa (Löbl, 1973) Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57′ E 117°48′, 450 m, 1̅ 5.ix.2005, Slade & Villanuava lgt., lowland mixed dipterocarp forest. Nature trail area 1, FIT, OUMNH (OUMNH). Remarks. The species is widely distributed in Sabah and seems to be common in samples of forest floor litter (see LÖBL 2021: 28)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53 (1): 85 ‾ 89."]}

Baeocera doriae (Pic, 1020) Material examined. 1 male, Sabah, Mt. Kinabalu N. P., 1700 m, 16.vii.02, Kurbatov & Zimina (MHNG); 1 male, Sabah, rd Kota Kinabalu ̅ Tambunan, km 52, 1600̅ 1800 m, 2̅ 5.vii.02, Kurbatov & Zimina (MHNG). Remarks. This species is known from Sarawak and Sabah only. It is poorly represented in collections, a single specimen has been reported from each of these states., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 9, DOI: 10.5281/zenodo.7501705

Baeocera hutan sp. nov. (Figs 1 ̅4) Type material. Holotype male, Sabah: Lahad Datu Ulu Segama forest Reserve, Danum Valley For. Centre, N 04°57.9′ E 117°48.1′, 200 m, 21.xi.2005, D. Mann, E. Slade & J. Villanuava lgt., lowland mix. dipterocarp forest. West trail area 1, FIT, OUMNH 2006­051 (OUMNH). Description. Length 1.28 mm, width 0.76 mm. Head and body nearly evenly reddishbrown, elytra hardly darker than pronotum, apical abdominal segments slightly lighter than pronotum, femora and tibiae as elytra, tarsi and antennae lighter. Head with interocular distance nearly as dorsoventral eye diameter. Head, thorax and elytra lacking microsculpture. Length/width ratios of antennomeres as follows: III 20 /5: IV 20 /6: V 30 /7: VI 27 /7: VII 33 /8: VIII 27 /7: IX 36 /11: X: 37/15: XI 48 /17. Pronotal and elytral punctation even, very fine. Lateral contours of pronotum and elytra continuously, very weakly arcuate. Pronotum with anterior margin bead broadly interrupted, lateral margins sinuate in lateral view, lateral margin striae impunctate. Margin of exposed part of scutellum rounded. Elytra with lateral margins nearly parallel, lateral margins in lateral view slightly sinuate, lateral margin carinae visible in dorsal view, lateral margin striae impunctate; sutural striae starting at base, parallel, deeply impressed, curved along pronotal lobe to form basal striae extended to humeral areas, broadly separated from lateral striae; discal punctation sparse, near base with puncture intervals about 8 to 10 larger than puncture diameters. Hind wing fully developed. Hypomera and mesanepisterna appearing impunctate. Mesoventrite with mesal ridge, punctate along lateral margins. Mesepimeron about 4 times as long as large and 5 times as long as interval to mesocoxa. Median part of metaventrite nearly flat, impunctate mesally, with dense, fairly coarse punctures on posterior third and laterally smooth mesal area, bearing conspicuous, long pubescence. Lateral parts of metaventrite with few scattered extremely fine punctures. Submesocoxal line parallel, margined with row of coarse elongate punctures extended posterior the inner part of mesepimeron. Submesocoxal area about 0.02 mm long. Metanepisterna large, about 0.10 mm wide, twice as long as wide, slightly convex, with inner margin broadly rounded anteriorly, nearly straight posteriad; inner suture impunctate. Protibiae straight. Mesotibiae and metatibiae slightly bent. Ventrite I lacking striae, with row of elongate basal punctures not interrupted in middle, extended laterally behind tip of metepimeron; punctation posterior basal row very fine and sparse. Male. Protarsomeres Ito III strongly widened, bearing long tenent setae; protarsomere I wider than apex of protibia. Ventrite VI with apical lobe subtriangular, about 0.04 mm long. Aedeagus (Figs 1 ̅4) 0.60 mm long. Median lobe and parameres strongly sclerotized, symmetrical, dorsal valves excepted. Articular process not prominent, distinct. Apical process of median lobe somewhat shorter than basal bulb, moderately inflexed, with weakly concave ventral margin, gradually narrowed apically, tip acute in lateral view, obtuse in dorsal view. Parameres robust, gradually narrowed apically in dorsal view, widened apical in lateral view, with broadly rounded apex. Internal sac with complex sclerites, lacking spinose and denticulate structures, with apical section of ejaculatory duct wide, weakly membranous, extruded in repos. Etymology. The species epithet is a noun meaning "forest̏ in Malay. Differential diagnosis. This species is unique in having broadly interrupted pronotal bead. Therefore, it is placed in Baeocera tentatively (the mouth­parts and thorax of the unique available specimen were not dissected). In addition, its aedeagal characters are unusual, in particular by the structure of the internal sac and the asymmetrical valves of the median lobe. The internal sac lacking a flagellar guide sclerite and possessing a membranous ejaculatory duct extruded in repos is shared with the Chinese B. xichangana Löbl, 1999. It differs notably from B. xichangana by the not prominent articular process, the two asymmetrical dorsal valves, and the much shorter, robust parameres. Both species are also very distinct in external characters, in particular by their body­shape and size (B. xichangana is 2.0̅ 2.15 mm long). Remarks. As to the Sabah species, only B. barbara Löbl, 1990 and B. doriae (Pic, 1920) share the wide metanepisterna. These two species have asymmetrical aedeagus and may be easily distinguished by their external characters. The antitheses 7 and 8 have been inversed in key to the Sabah Baeocera given by LÖBL & SMETANA (2021): the thesis 6 (Metanepisternum fused,...) should lead to ̤ 8., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 8-9, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & SMETANA A. 2021: On the Baeocera Erichson (Coleoptera: Staphylinidae: Scaphidiinae) of Sabah, Malaysia, and a tale on mystified biodiversity. Journal of Insect Biodiversity 23 (2): 23 ‾ 42."]}

4. Key to the Baeocera species of the People’s Republic of China 1 Lateral parts of metaventrite impunctate, or very finely and sparsely punctate...................2 - Lateral parts of metaventrite coarsely, usually densely punctate, or at least with several conspicuous, coarse punctures...........................................................................................14 2 Elytra with basal striae interrupted, not joined to lateral striae. Exposed part of scutellum large....................................................................................................................3 - Elytral with basal striae entire, joined to lateral striae. Scutellum concealed or minute, except in B. breveapicalis...................................................................................................6 3 Apical part of median lobe large, not overlapped, about as long as basal bulb. Male metatibiae each with subapical denticle........................................ B. satana NAKANE, 1963 - Apical part of median lobe small, to part or entirely overlapped, much shorter than basal bulb. Male metatibiae lacking denticles.....................................................................4 4 Aedeagus with parameres simple, hook-like and curved at apices....... B. gilloghyi LÖBL, 1973 - Aedeagus with parameres lobed, not hook-like at apices....................................................5 5 Left parameres with small dorsal lobe and ventrally inflexed apex; right parameres lobed ventrally........................................................................... B. yunnanensis LÖBL, 1999 - Left parameres with large dorsal lobe and hardly inflexed apex; right parameres not lobed............................................................................................ B. pubiventris LÖBL, 1990 6 Antennomere VI short and wide, about 2 times as long as wide and as long as half of antennomere VII..................................................................................................................7 - Antennomere VI long and narrow, about 3 to 5 times as longs as wide, slightly shorter than antennomere VII..........................................................................................................8 7 Aedeagus with outer margin of left paramere sinuate and inner lobe subtriangular............................................................................................................... B. breveapicalis (PIC, 1926) - Aedeagus with outer margin of left paramere convexly rounded and inner lobe blunt................................................................................................................ B. takizawai LÖBL, 1984 8 Aedeagus with parameres strongly expanded apico-dorsally to form large, strongly sclerotized lobes................................................................................. B. kubani LÖBL, 1999 - Aedeagus with parameres not lobed, or each with small subapical lobe.............................9 9 Aedeagus with apices of parameres distinctly widened (dorsal view)....................................................................................................................................... B. huanshana LÖBL, 1999 - Aedeagus with apices of parameres not or hardly widened (dorsal view).........................10 10 Aedeagus lacking strongly sclerotized guide-sclerite, parameres slightly shorter than median lobe................................................................................ B. xichanagna LÖBL, 1999 - Aedeagus lacking with strongly sclerotized guide-sclerite, parameres much shorter than median lobe.......................................................................................................................11 11 Aedeagus with guide-sclerite narrow, angulate basally...................... B. callida LÖBL, 1986 - Aedeagus with guide-sclerite robust, not angulate basally................................................12 12 Aedeagus with guide-sclerite tapering apicad........................................ B. freyi LÖBL, 1966 - Aedeagus with guide-sclerite not tapering apicad.............................................................13 13 Aedeagus with guide-sclerite straight, becoming stouter apicad........ B. cooteri LÖBL, 1999 - Aedeagus with guide-sclerite abruptly curved in middle, widened subapically, acute at tip................................................................................................. B. hammondi LÖBL, 1984 14 Elytra with sutural striae shortened, starting well behind level of scutellar tip, far from elytral base........................................................................................................................15 - Elytra with sutural striae starting at elytral base, usually curved along pronotal lobe and forming basal striae...........................................................................................................16 15 Pronotal punctation coarse and dense, centre of metaventrite with mesal row of coarse punctures and narrow impunctate admesal areas................................ B. karenovicsi nov.sp. - Pronotal punctation very fine and sparse, hardly visible, metaventrite lacking mesal row of coarse punctures....................................................................... B. proclinata nov.sp. 16 Elytra with basal striae widely separated from lateral striae.............................................17 - Elytra with basal striae entire, joined to lateral striae........................................................19 17 Elytral punctation coarse anterior mid-length, very fine posterior mid-length or in apical thirds. Aedeagus with notched parameres...............................................................18 - Elytral punctation coarse on entire discal surfaces. Aedeagus with straight, not notched parameres.................................................................................................. B. repleta nov.sp. 18 Internal sac of aedeagus bearing tuft of spines.................................. B. franzi (LÖBL, 1973) Internal sac of aedeagus lacking tuft of spines.......................... B. proseminata LÖBL, 2013 19 Pronotal and elytral punctation similar, very fine..............................................................20 - Pronotal punctation very fine, elytral punctation in part or entirely coarse.......................21 20 Aedeagus with flagellum curved and thickened basally. Lateroposterior part of metaventrite coarsely punctate................................................... B. sordidoides LÖBL, 1992 - Aedeagus with flagellum almost straight in dorsal view, not thicken basally. Lateroposterior part of metaventrite very finely punctate.................. B. coalita LÖBL, 2013 21 Metanepisterna separated from metaventrite by distinct sutures.......................................................................................................................................... B. serendibense (LÖBL, 1971) - Metanepisterna fused with metaventrite............................................................................22 22 Punctation coarse on most of basal half to two thirds of elytral disc, apical third to half of elytral disc very finely punctate, appearing impunctate................................................23 - Elytra entirely or nearly entirely coarsely punctate...........................................................24 23 Parameres narrow, not folded apically............................................... B. erroris LÖBL, 1990 - Parameres wide, folded apically................................................ B. pseudinculta LÖBL, 1990 24 Ventrite 1 very finely punctate posterior basal puncture row, appearing impunctate............................................................................................................. B. longicornis (LÖBL, 1971) - Ventrite 1 coarsely punctate posterior basal puncture row................................................25 25 Apical halves of parameres distinctly bent and with weakly sclerotized inner margins................................................................................................................... B. pigra (LÖBL, 1971) - - Parameres straight, with inner and outer margins nearly evenly sclerotized...................26 26 Parameres almost as long as median lobe. Internal sac without narrow lateral rod............................................................................................................................ B. vidua LÖBL, 1990 - Parameres much shorter than median lobe. Internal sac with narrow lateral rod.............................................................................................................................. B. lasciva LÖBL, 2013, Published as part of Löbl, Ivan, 2018, Supplement to the knowledge of the genera Baeocera ERICHSON, 1845 and Scaphobaeocera CSIKI, 1909 (Coleoptera, Staphylinidae, Scaphidiinae) of the People's Republic of China, pp. 1295-1303 in Linzer biologische Beiträge 50 (2) on pages 1299-1300, DOI: 10.5281/zenodo.5275102, {"references":["LOBL I. (1999): A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of China, I. - Revue suisse zool. 106: 691 - 744.","LOBL I. (1992): The Scaphidiidae (Coleoptera) of the Nepal Himalaya. - Revue suisse zool. 99: 471 - 627.","LOBL I. (1971): Die Scaphidiidae (Coleoptera) Sudindiens. - Revue s uisse zool. 86: 77 - 129."]}

Pseudobironium subovatum Pic, 1920 Material examined. 1 male, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04°59.738′ E 117°50.165′, 250 m, iii̅ iv.2005, FIT no 2, primary forest, E. Slade & J. Villanuava lgt. Lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005­062 (OUMNH). Remarks. The species seems to be rather widely distributed. It is currently known from Indonesia (Sumatra and Kalimantan), Malaysia (West Malaysia, Sabah and Sarawak), and Thailand, though only a single Sabah record is published from the Gunung Emas in the Crocket Range (LÖBL & TANG 2013). The species may be easily distinguished by the long protibial setae in males., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 11, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Mordelloscaphium testaceimembre Pic, 1915 Material examined. 184, Sabah, Batu Punggul Resort env., 24.vi. ̅ 1.vii.1996, FIT (MHNG); 1, with the same data but in debris accumulates around large trees near river (MHNG); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°57.9′ E 117°48.1′, 200 m, 22̅ 23.x.2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, west trail area, FIT, OUMNH 2006­051 (OUMNH); 1, with the same data but 3̅13.xi., D.J. Mann & E. Slade lgt,, forest natural trail area OUMNH 2065­051 (OUMNH); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05°02.682′ E 117°45.553′, 250 m, iii̅ iv.2005, E. Slade & J. Villaneuve lgt., FIT, primary dipterocarp forest, OUMNH 2065­062 (OUMNH). Remarks. The currently known specimens of this species are the syntypes from east Kalimantan, Indonesia., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 10-11, DOI: 10.5281/zenodo.7501705

Pseudobironium Pic, 1920 The genus is Asian in distribution, with most members found in the tropics and subtropics, remaining unknown south of the Wallace line. Currently, six species are known to occur in Sabah (LÖBL & TANG 2013): P. convexum Löbl & Tang, 2013, P. horaki Löbl & Tang, 2013, P. pubiventer Löbl & Tang, 2013, P. sparsepunctatum (Pic, 1915), P. subovatum Pic, 1920, and P. vitalisi (Achard, 1920). Anew record is given, and two additional species are described below., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 11, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734."]}

Scaphoxium sinuatum sp. nov. Figs 11 ̅13 Type material. Holotype male, Malaysia, Sabah, Batu Punggul Resort env., 24.VI. ̅ 1.VII.1996, 11c. vegetation debris and forest floor litter accumulated around large trees near river (MHNG). Description. Length 1.18 mm, width 0.55 mm, dorsoventral diameter 0.62 mm. Thorax, elytra and most of abdomen uniformly reddish­brown, head somewhat lighter, apical abdominal segments light, nearly yellowish, femora and tibiae roughly as thorax, tarsi lighter, antennae light brown. Length/width ratios of antennomeres as follows: III 22 /5: IV 17 /4: V 22 /4: VI 23 /4: VII 28 /7: VIII 20 /7: IX 28/11: X: 28/11: XI 38 /12. Pronotal and elytral punctation similar, very fine, hardly visible at 40 times magnification. Scutellum concealed. Elytra with sutural striae parallel, shortened, starting about 0.15 mm posterior of margin of pronotal lobe, adsutural areas hardly raised. Hypomeron with scattered, very fine punctured and short oblique stria. Mesoventrite with mesal impression shallow, narrowed posteriad, lacking mesal ridge and lacking microsculpture. Margin of mesocoxal process rounded. Metaventrite lacking microsculpture, very finely punctate, convex in middle. Lateral stria oblique, nearly reaching convex, finely punctate submesocoxal line. Submesocoxal area about 0.03 mm, as long as half of the shortest interval to margin of metacoxa. Metanepisternum flat, with suture fairly narrow, punctate, somewhat rounded. Abdomen very finely punctate, median areas of ventrites with punctulate microsculpture. Male. Segments Ito III of protarsi hardly widened. Aedeagus (Figs 11 ̅13) 0.44 mm long. Parameres narrowed posterior of basal section, gradually weakly widened toward narrow apical section, not forming subapical lobe or process; outer margins of parameres concave and inner margins strongly sclerotized posterior of narrowed apical section. Internal sac bearing two pairs of sclerotized pieces, outer larger and more sclerotized than inner pair, expanded apically, inner pair widened proximally, with margins becoming indistinct apically; scale­like, spine­like or filamentous structures absent. Etymology. The species epithet is a Latin adjective meaning "sinuatȅ, referring to the outer margins of the parameres. Differential diagnosis. The body­size of this new species is significantly smaller than that of other Bornean species and the shape of its parameres is diagnostic for it. The new species is similar to S. bilobum Löbl, 2015 from Lombok. It shares with the latter the simple internal sac of the aedeagus lacking spine­like, denticulate or striate membranous structures in combination with the presence of two pairs of admesal sclerites. However, the outer pair of sclerites is much more expanded and the inner sclerites are much shorter and broader in S. sinuatum, which may be distinguished from B. bilobum also by its wide impression of the mesoventrite, finer punctation of the lateral parts of the metaventrite and shorter submesocoxal areas., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on pages 17-18, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2015: On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands. Revue suisse de zoologie 122 (1): 75 ‾ 120."]}

Baeocera sarawakensis Löbl, 1987 Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04°58′ E 117°42′, 450 m, viii.1998, D. Bebber lgt., lowland mixed dipterocarp forest, FIT, OUMNH 2003­050 (OUMNH). Remarks. As B. incisa, this species is widely distributed in Sabah and seems to be common in forest floor litter (see LÖBL 2021: 31)., Published as part of Löbl, Ivan, 2022, Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae), pp. 7-19 in Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2) on page 10, DOI: 10.5281/zenodo.7501705, {"references":["LOBL I. 2021: On the Bornean species of Scaphoxium Lobl, 1979 (Coleoptera, Staphylinidae, Scaphidiinae). Linzer biologische Beitage 53 (1): 85 ‾ 89."]}

Sapitia Achard, 1920 The genus includes three South-East Asian species, all reported also from Indonesia though only one of them from the Lesser Sundas. Members of Sapitia are presumably associated with termites (Leschen & Löbl, 2005)., Published as part of Löbl, Ivan, 2015, On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands, pp. 75-120 in Revue suisse de Zoologie 122 (1) on page 92, DOI: 10.5281/zenodo.14582, {"references":["Achard J. 1920. Notes sur les Scaphidiidae de la faune indomalaise. Annales de la Societe Entomologique de Belgique 60: 123 - 136.","Leschen R. A. B., Lobl I. 2005. Phylogeny and classification of Scaphisomatini (Staphylinidae: Scaphidiinae) with notes on mycophagy, termitophily and functional morphology. Coleopterists Society Monographs, Patricia Vaurie series no. 3: 63 pp."]}