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Hoffmannina libita Brennan and Goff, 1977 (Fig. 24) Hoffmannina libita Brennan and Goff, 1977: 911, figs. 23–33; Bassini-Silva et al. 2021: 774. Diagnosis. SIF = SIF = 7BS-N(b)-3-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/B/NNB; fSc: PL> AM ≥ AL; Ip = 941–1064; fD = (6–8)H-(6–11)-(7–10)-(7–10)-(6–8)+(4–7); DS = 38–50; V = 34–44; NDV = 72–91. Galeal seta nude or having 1 branch; scutum nearly pentagonal, densely punctated, with rounded, greatly projected posterior margin; AM posterior to level of AL; two PPL setae present on scutal margin posterior to PL; sensilla flagelliform, nude or having few cilia; scutal and dorsal idiosomal setae thick, sparsely covered with small scalelike barbs. Standard measurements of examined specimens given in Table 14. Distribution. Panama (Brennan & Goff 1977) and Costa Rica (this study). Hosts. Reithrodontomys sumichrasti (Saussure), Reithrodontomys creper Bangs, Peromyscus mexicanus (Saussure) (= P. nudipes) (Rodentia: Cricetidae) (Brennan & Goff 1977). New host records: Catharus gracilirostris, Chlorospingus pileatus, Momotus lessonii. Localization on hosts. Near vent (C. gracilirostris Nos BV 179, BV197, BV218). Material examined. Seven larvae (ZIN 9154, 9158, 9160, 9161, 9164, 9165, 9192) ex C. gracilirostris Nos BV 179, BV197, BV218, COSTA RICA: Braulio Carillo Nat. Park, Sector Barva, 4 and 6 August 2010; 1 larva (ZIN 9201) ex C. pileatus No CM 60, Tapanti Nat. Park, Sector Cerro de la Muerte, 12 August 2010; 1 larva (ZIN 9228) ex M. lessonii No LT 108, Cordillera de Talamanсa, Las Tablas, 20 August 2010., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 540-541, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054, {"references":["Brennan, J. M. & Goff, M. L. (1977) The Neotropical genus Hoffmannina: four new species and other records from Mexico, Panama and Venezuela (Acarina: Trombiculidae). Journal of Parasitology, 63, 908 - 914. https: // doi. org / 10.2307 / 3279906","Bassini-Silva, R., Takatsu, J. C., Bermudez, S. E., Miranda, R. J., Welbourn, C., Ochoa, R., Barros-Battesti, D. M. & Jacinavicius, F. C. (2021) A checklist of chiggers (Trombidiformes: Trombiculidae and Leeuwenhoekiidae) of Panama. Acarologia, 61, 763 - 789. https: // doi. org / 10.24349 / ach 8 - Eqk 3"]}

Ascoschoengastia tragulata (Brennan and Jones, 1961) (Fig. 22) Euschöngastia tragulata Brennan and Jones, 1961b: 110, fig. 6. Ascoschoengastia (Ascoschoengastia) tragulata: Vercammen-Grandjean 1965: 92. Euschoengastia tragulata: Brennan & Junker 1966: 235; Brennan 1970: 811. Microtrombicula tragulata: Webb & Loomis 1970: 141, fig. 4; Brennan & Reed 1975: 56; Hoffmann 1990: 94, fig. 59; Bassini-Silva et al. 2021: 777. Ascoschoengastia tragulata: Kudryashova 1998: 30. Microtrombicula (Microtrombicula) tragulata: Nielsen et al. 2021: 156. Diagnosis. SIF = 6B(N)-N-3-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/b/N(b)N(b)B(f); fSc: PL> AM> AL; Ip = 549–571; fD = 2H-6-6-6-4+(2–8); DS = 26–32; V = 23–30; NDV = 55–56. Palpal femoral seta with small cilia, palpal genual seta with 1–2 branches, dorsal and lateral palpal tibial setae nude or having 1–2 branches, ventral palpal tibial seta branched or forked; scutum nearly pentagonal, punctated, with anterolateral shoulders, with rounded, slightly projected posterior margin; AM anterior to level of AL; sensilla fusiform, with longitudinal rows of setules, far anterior to level of PL (PSB—P-PL = 8–11 µm). Standard measurements of examined specimens given in Table 11. Distribution. Bolivia, Mexico, Nicaragua, Panama, Venezuela (Bassini-Silva et al. 2021), and Costa Rica (this study). “ Microtrombicula nr. tragulata ” was also recorded by Brennan & Lukoschus (1971) from Surinam. Hosts. Bassariscus astutus (Lichtenstein), Nasua narica (L.), Potos flavus (Schreber) (Carnivora: Procyonidae), Coendou quichua Thomas (= C. rothschildi) (Rodentia: Erethizontidae), Rattus rattus (L.) (Rodentia: Muridae), Hylaeamus megacephalus (Fischer) (= Oryzomys capito) (Rodentia: Cricetidae), Didelphis marsupialis L., Marmosa robinsoni Bangs (Didelphimorphia: Didelphidae) (Brennan 1970; Brennan & Reed 1975). New host records: Glyphorynchus spirurus, Xiphorhynchus erythropygius, Myrmotherula schisticolor. Localization on hosts. In ears (X. erythropygius Nos T 071 and T297), around vent (G. spirurus No T 016), in ears and around vent (G. spirurus No T 250), in axilla and surrounding area (M. schisticolor No T 014). Material examined. 28 larvae (ZIN 9305–9314, 9318, 9319, 9325, 9486, 9488-9490, 9494, 9533-9542) ex X. erythropygius Nos T 071 and T 297, M. schisticolor No T 014, G. spirurus Nos T 016 and T 250, COSTA RICA: Tapanti Nat. Park, Sector Tapanti, 31 July–9 August 2009. Remarks. Based on the presence of fusiform sensilla, this species should be placed in the genus Ascoschoengastia Ewing, 1946 rather than Microtrombicula Ewing, 1950, to which it is referred by most modern authors (Webb & Loomis 1970; Brennan & Reed 1975; Hoffmann 1990; Bassini-Silva et al. 2021). These two genera are morphologically identical, except for the shape of sensilla—inflated in Ascoschoengastia and flagelliform in Microtrombicula (Nadchatram & Dohany 1974)., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 535-537, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054, {"references":["Brennan, J. M. & Jones, E. K. (1961 b) New genera and species of chiggers from Panama (Acarina: Trombiculidae). Journal of Parasitology, 47, 105 - 124. https: // doi. org / 10.2307 / 3274993","Vercammen-Grandjean, P. H. (1965) Trombiculinae of the world, synopsis with generic, subgeneric, and group diagnoses (Acarina, Trombiculidae). George Williams Hooper Foundation, University of California, San Francisco, California, 191 pp.","Brennan, J. M. (1970) Chiggers from the Bolivian-Brazilian border (Acarina: Trombiculidae). Journal of Parasitology, 56, 807 - 812. https: // doi. org / 10.2307 / 3277728","Webb, J. P. Jr. & Loomis, R. B. (1970) Four species of Microtrombicula (Acarina: Trombiculidae) from Mexico and Nicaragua. Bulletin of the Southern California Academy of Sciences, 69, 133 - 144.","Brennan, J. M. & Reed, J. T. (1975) A list of Venezuela chiggers, particularly of small mammalian hosts (Acarina: Trombiculidae). Brigham Young University Science Bulletin, Biological Series, 20, 45 - 75.","Hoffmann, A (1990) Los Trombiculidos de Mexico (Acarida: Trombiculidae). Parte taxonomica. Publicaciones especiales del Instituto de Biologia. Vol. 2. Universidad Nacional Autonoma de Mexico, Mexico, 275 pp.","Kudryashova, N. I. (1998) Chigger mites (Acariformes, Trombiculidae) of East Palaearctics. KMK Scientific Press, Moscow, 342 pp. [in Russian]","Nielsen, D. H., Robbins, R. G. & Rueda, L. M. (2021) Annotated world checklist of the Trombiculidae and Leeuwenhoekiidae (1758 - 2021) (Acari: Trombiculoidea), with notes on nomenclature, taxonomy, and distribution. Zootaxa, 4967 (1), 1 - 243 https: // doi. org / 10.11646 / zootaxa. 4967.1.1","Bassini-Silva, R., Takatsu, J. C., Bermudez, S. E., Miranda, R. J., Welbourn, C., Ochoa, R., Barros-Battesti, D. M. & Jacinavicius, F. C. (2021) A checklist of chiggers (Trombidiformes: Trombiculidae and Leeuwenhoekiidae) of Panama. Acarologia, 61, 763 - 789. https: // doi. org / 10.24349 / ach 8 - Eqk 3","Brennan, J. M. & Lukoschus, F. (1971) Parasitic mites of Surinam. VIII. A new genus and species of chigger, Fauranius atecmartus, and additional records of species (Acarina: Trombiculidae). Bulletin of the Southern California Academy of Sciences, 70, 42 - 45.","Nadchatram, M. & Dohany, A. L. (1974) A pictorial key to the subfamilies, genera and subgenera of Southeast Asian chiggers (Acari, Prostigmata, Trombiculidae). Bulletin - Institute for Medical Research, Malaysia, 16, 1 - 67."]}

Eutrombicula cathari Stekolnikov sp. nov. (Figs. 4–6) Diagnosis. SIF = 7BS-N-2-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/B(N)/NNB; fSc: PL> AM> AL; Ip = 1085–1186; fD = 2H-6-6-4-2(2-4); DS = 20; V = 11–13; NDV = 31–33. Inner prong of palpal claw slightly shorter than outer prong; scutal and dorsal idiosomal setae moderately covered with short barbs; scutum nearly pentagonal, densely covered with small puncta, without transverse striae; sensilla (trichobothria) with ca. 10 branches in distal half; sensillary bases far anterior to level of PL (PSB—P-PL = 8–13 µm). Standard measurements of type series given in Table 3. Description (larva) (based on holotype and 7 paratypes). IDIOSOMA (Figs. 4, 5A–E). Eyes 2 + 2; 20 dorsal idiosomal setae, including one pair of humeral setae, moderately covered with short barbs, by 6 setae in 1 st and 2 nd (D) posthumeral rows, 4 setae in 3 rd (E) row, 2 setae in 4 th (F) row, setae in rows E and F can be also counted as 2 and 4, respectively; 4 sternal setae; 11–13 ventral setae; NDV = 31–33. GNATHOSOMA (Fig. 5F, G). Cheliceral blade with tricuspid cap; cheliceral base with small dense puncta basally and sparse large puncta in distal part; gnathobase with small dense puncta, transverse striae, and 1 pair of branched gnathocoxal (tritorostral) setae; galeal (deutorostral) seta nude; palpal claw with 2 prongs, inner prong slightly shorter than outer prong; palpal femur with dense small puncta and branched seta; palpal genu with sparse small puncta and branched (in holotype and five paratypes) or nude (in two paratypes) seta; palpal tibia with few small puncta, nude dorsal and lateral and branched ventral seta; palpal tarsus with 7 branched setae, nude subterminala (ζ) and basal tarsala (ω). SCUTUM (Figs. 4C, 5A). Nearly pentagonal, densely covered with small puncta, without transverse striae, with anterolateral shoulders, anterior scutal margin sinuous, lateral margins concave, posterior margin rounded; AM at level of AL, sensillary (trichobothrial) bases far anterior to level of PL (PSB—P-PL = 8–13 µm); all scutal setae uniform with dorsal idiosomal setae; PL> AM> AL; flagelliform sensilla (trichobothria), with ca. 10 branches in distal half. Rudimentary additional AM present in one paratype (Fig. 4B). LEGS (Fig. 6). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 3 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) proximal to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 14B, mastitarsala. Hosts. Catharus frantzii, Catharus gracilirostris. Localization on hosts. Near vent (C. frantzii Nos BV 206 and BV213, C. gracilirostris Nos BV 179, BV193, BV196, B218, BV255), in ear (C. gracilirostris No BV 217). Type material. Holotype larva (ZIN 9167) ex C. gracilirostris No BV 196, COSTA RICA: Braulio Carillo National Park, Sector Barva, 5 August 2010. Paratypes: 6 larvae (ZIN 9168–9173) with same data; 36 larvae (interval of numbers: ZIN 9151–9200) ex C. frantzii Nos BV 117, BV206, BV211, BV213 (15 larvae) and C. gracilirostris Nos BV 179, BV193, BV212, BV217, BV218, BV255 (21 larvae), Braulio Carillo National Park, Sector Barva, 2–7 August 2010. Etymology. The species epithet cathari is derived from the generic name of the type host. Differential diagnosis. Eutrombicula cathari is similar to E. tinami and differs from it by the presence of relatively longer inner prong of palpal claw, which is only slightly shorter than the outer prong, vs. less than the two thirds as long as the outer prong in E. tinami; by the greater scutum (AW = 85–96, PW = 98–115, and SD = 67–76 vs. 76–77, 92, and 64, respectively); by longer PL (68–79 vs. 62–63) and longer D max (61–70 vs. 56) (Fuller 1952). The new species differs from the E. tinami form described by Bassini-Silva et al. (2019), in addition to the relatively longer inner prong of palpal claw, by the relatively wider scutum (SD = 67–76 vs. 74–85, while PW is similar: 98–115, mean 105 vs. 93–116, mean 104) without transverse striae (vs. with transverse striae), longer ventral idiosomal setae (V min = 45–52 vs 32–33 and V max = 58–67 vs. 39–45), longer legs (TaIIIL = 99–110 vs. 89–97; total lengths of legs are incomparable with our data, as noted above).

Eutrombicula helleri (Oudemans, 1911) (Figs. 13–15) Microthrombidium helleri Oudemans, 1911: 120; 1912: 15, fig. C. Eutrombicula helleri: Fuller 1952: 139. Diagnosis. SIF = 7BS-N-2-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/N(B)/NNN; fSc: PL> AM> AL; Ip = 812–934; fD = 2H-6-6-4-2-(0-2); DS = 19–22; V = 10–14; NDV = 32–35. Inner prong of palpal claw twice as short as outer prong and located medially on the latter; scutal and dorsal idiosomal setae thin, moderately covered with short barbs; scutum nearly pentagonal, with small dense puncta and transverse striae; sensilla (trichobothria) with ca. 8–10 branches in distal half; sensillary bases far anterior to level of PL (PSB—P-PL = 9–14 µm). Standard measurements of examined specimens given in Table 5. Redescription (larva) (based on 21 specimens). IDIOSOMA (Figs. 13, 14A–G). Eyes 2 + 2; 19–22 dorsal idiosomal setae, including one pair of humeral setae, moderately covered with short barbs, by 6 setae in 1 st and 2 nd (D) posthumeral rows, 4 setae in 3 rd (E) row, 2 setae in 4 th (F) row, 5 th (H) row includes 2 setae (half of specimens), 1 (one specimen) or absent; 4 sternal setae; 10–14 ventral setae; NDV = 32–35. GNATHOSOMA (Fig. 14H, I). Cheliceral blade with tricuspid cap; cheliceral base with small dense puncta basally and more sparse puncta in distal part; gnathobase with small dense puncta, transverse striae, and 1 pair of branched gnathocoxal (tritorostral) setae; galeal (deutorostral) seta nude; palpal claw with 2 prongs, inner prong twice as short as outer prong and located medially on the latter; palpal femur with puncta and branched seta; palpal genu with puncta and nude seta (having 1–2 branches in two specimens); palpal tibia with three nude setae; palpal tarsus with 7 branched setae, nude subterminala (ζ) and basal tarsala (ω). SCUTUM (Figs. 13C, 14A, B). Nearly pentagonal, densely covered with small puncta, with transverse striae, with small anterolateral shoulders, anterior scutal margin lightly sinuous, lateral margins concave, posterior margin broadly rounded; AM at level of AL, sensillary (trichobothrial) bases far anterior to level of PL (PSB—P-PL = 9–14 µm); all scutal setae uniform with dorsal idiosomal setae; PL> AM> AL; flagelliform sensilla (trichobothria), with ca. 8–10 branches in distal half. LEGS (Fig. 15). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 3 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) proximal to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 14B, mastitarsala. Distribution. Surinam, Panama (Fuller 1952), Paraguay, and Peru (this study). Hosts. Arthropoda: Passalus sp. (Coleoptera: Passalidae); Reptilia: Ameiva praesignis (Baird and Girard) (Squamata: Teiidae); Mammalia: Didelphis marsupialis L. (Didelphimorphia: Didelphidae), Dasyprocta punctata (Gray) (Rodentia: Dasyproctidae), “rat” (Rodentia: Muridae), Homo sapiens L. (Primates: Hominidae), Sylvilagus brasiliensis gabbi (J. Allen) (= S. gabbi) (Lagomorpha: Leporidae), Leopardus pardalis (L.) (= Felis pardalis) (Carnivora: Felidae); Aves: Gallus gallus domesticus (L.) (Galliformes: Phasianidae), Leptotila verreauxi (Columbiformes: Columbidae), Tyrannidae gen. sp. (Fuller 1952). New host records: Campylorhamphus trochilirostris, Cantorchilus guarayanus, Casiornis rufus, Glyphorynchus spirurus, Gymnopithys leucaspis, Lepidocolaptes angustirostris, Leptotila rufaxilla, Myiarchus tyrannulus, Paroaria capitata, Microspingus melanoleucus, Ramphocaenus melanurus, Saltator coerulescens, Taraba major, Thamnophilus doliatus, Thamnophilus schistaceus, Troglodytes aedon, Turdus amaurochalinus, Turdus rufiventris. Material examined. Three larvae (ZIN 9111–9113) ex C. trochilirostris No PG 400, 1 larva (ZIN 9130) ex C. rufus No PG 444, 4 larvae (ZIN 9020, 9034, 9134, 9135) ex L. angustirostris Nos PG 328, PG345, PG446, 2 larvae (ZIN 9035, 9137) ex M. tyrannulus Nos PG 346 and PG456, 1 larva (ZIN 9117) ex P. capitata No PG 440, one larva (ZIN 9115) ex M. melanoleucus No PG 431, 3 larvae (ZIN 9098, 9128, 9150) ex S. coerulescens Nos PG 376, PG443, PG473, 9 larvae (ZIN 9049–9057) ex T. major No PG 367, 4 larvae (ZIN 9103, 9106–9108) ex T. doliatus No PG 394, 15 larvae (ZIN 9021–9023, 9038–9041, 9092–9097, 9120, 9133) ex T. aedon Nos PG 337, PG363, PG372, PG442, PG445, 6 larvae (ZIN 9079–9081, 9083, 9088, 9089) ex C. guarayanus Nos PG 370 and PG371, 8 larvae (ZIN 9018, 9019, 9042–9047) ex T. amaurochalinus Nos PG 326 and PG366, 7 larvae (ZIN 9058–9064) ex T. rufiventris No PG 368, PARAGUAY: Tres Gigantes Biological Station, 6–9 September 2012; 2 larvae (ZIN 8831, 8832) ex T. schistaceus No TR 2, 8 larvae (ZIN 8833–8840) ex R. melanurus No TR 5, PERU: San Martin Province, Tarapoto, Centro URKU, 8 August 2011; 18 larvae (ZIN 8841–8858) ex G. leucaspis Nos IQ 1 and IQ2, 1 larva (ZIN 8876) ex L. rufaxilla No IQ 33, 2 larvae ex G. spirurus Nos IQ 32 and IQ5, PERU: Iquitos, National Reserve Allpahuayo Mishana, 13–16 August 2011. Variation. Specimens from Paraguay slightly differ from specimens collected in Peru by the shape of scutum and sparser scutal puncta (Fig. 14A, B). According to the results of Mann-Whitney test, measurements characterizing width (AW, PW) and length (ASB, PSB, SD, AP) of the scutum were significantly higher in the sample from Paraguay (Table 6), whereas the length of humeral setae (H) was lesser in these specimens. Remarks. Eutrombicula helleri differs from E. goeldii by significantly shorter setae (AM–V max in Table 5), shorter legs (pa–TaIIIL), and longer relative distance of mastitarsala from the base of leg tarsus III (m-t). The posterior scutal margin of E. goeldii is straight in the middle part and obliquely cut at the edges, whereas in E. helleri it is evenly arquate, bow-shaped (compare Fig. 11A and 14A, B). Scutal puncta are denser in E. goeldii than in E. helleri. Intervals of all scutal measurements for these species overlap, but differences in some of these characters are statistically significant. Thus, E. helleri has the scutum narrower (lesser AW and PW), ASB and AP lesser, and P-PL greater than those features in E. goeldii (Table 5)., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 522-525, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054, {"references":["Oudemans, A. C. (1911) Acarologische Aanteekeningen XXXV. Entomologische Berichten, Amsterdam, 3, 118 - 126.","Oudemans, A. C. (1912) Die bis jetzt bekannten Larven von Thrombidiidae und Erythraeidae. Zoologische Jahrbucher, Supplement 14, 1 - 230.","Fuller, H. S. (1952) The mite-larvae of the family Trombiculidae in the Oudemans collection: taxonomy and medical importance. Zoologische Verhandelingen, 18, 1 - 261."]}

Odontacarus robbinsi Loomis and Spath, 1969 (Fig. 27) Odontacarus robbinsi Loomis and Spath, 1969: 187, fig. 1. Odontacarus (Odontacarus) robbinsi: Goff & Loomis 1977: 373; Hoffmann 1990: 180, fig. 148. Diagnosis. SIF = 7B-B-3-2111.1000; fsp = 6.6.6; fCx = 2.1.1; fSt = 0.2; fPp = B/B/BNN; fSc: PL> AM> AL; Ip = 1238–1283; fD = 2H-[5-8]-8-10-8-6(7)-5; DS = 52–53; V = 49; NDV = 101–102. Cheliceral blade with 5 dorsal teeth; stigmata situated at base of gnathosoma, tracheae extend to caudal third of idiosoma; scutum nearly pentagonal, with pointed nasus on anterior margin and angulate posterior margin, with dense puncta; two AM at level of AL; sensilla flagelliform, with ca. 6 long branches in distal half, far anterior to level of PL (PSB—P-PL = 7–11 µm); onychotriches present; parasubterminala (z) branched; f 1 proximal to S 1, f 2 proximal to S 2; microgenuala II present; mastitarsala ciliated. Standard measurements of two examined specimens (ZIN 9004 and ZIN 9037): AW 64, 68; PW 84, 86; SB 27, 27; ASB 28, 35; PSB 29, 29; SD 57, 64; P-PL 18, 22; AA 12, 13; NL 16, 18; NW 8, 9; AP 35, 36; AM 39, 37; AL 36, 34; PL 55, 58; S -, 50; H 52, 56; D min 32, 32; D max 50, 41; V min 23, 25; V max 35, 40; pa 452, 430; pm 380, 371; pp 452, 437; Ip 1283, 1238; DS 52, 53; V 49, 49; NDV 101, 102; TaIIIL 119, 110; TaIIIW 16, 19. Distribution. Mexico (Loomis & Spath 1969) and Paraguay (this study). Hosts. Sceloporus zosteromus Cope (= S. magister zosteromus), S. licki Van Denburgh (= S. orcutti licki), Urosaurus nigricaudus (Cope) (Squamata: Phrynosomatidae) (Loomis & Spath 1969). New host records: Coryphistera alaudina and Zonotrichia capensis. Material examined. One larva (ZIN 9004) ex C. alaudina No PG 303, PARAGUAY: Teniente Agripino Enciso Nat. Park, 2 September 2012; 1 larva (ZIN 9037) ex Z. capensis, Tres Gigantes Biological Station, 6 September 2012. Remarks. According to the original description, this species has 4 dorsal and 3 ventral teeth on the cheliceral blade. We did not observe ventral teeth, but recorded 5 dorsal teeth; insufficient sample size does not allow estimation of the reliability of this difference. The difference in the dorsal setal formula of idiosoma (10-8-6-10-8+27, according to the original description) was caused most probably by differently traced borders between setal rows; however, the total number of setae in our specimens is almost the same (NDV = 101–102 vs. 103). Branched parasubterminala (z) of this species was evidently treated by the authors of original description as a non-specialized seta and thus was omitted. We should note that there is no way to distinguish “branched subterminala” from “subterminala replaced with a non-specialized branched seta” by a morphological examination. Finally, our measurements of the scutum and setae are in good agreement with those given in the original description., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 544-546, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054, {"references":["Loomis, R. B. & Spath, L. C. (1969) A new species of Odontacarus Ewing (Acarina, Trombiculidae) from lizards of Baja Cali- fornia Sur, Mexico. Bulletin of the Southern California Academy of Sciences, 68, 187 - 190.","Goff, M. L. & Loomis, R. B. (1977) The genus Odontacarus (Acari: Trombiculidae) in North America. Journal of Medical Entomology, 14, 370 - 373 https: // doi. org / 10.1093 / jmedent / 14.3.370","Hoffmann, A (1990) Los Trombiculidos de Mexico (Acarida: Trombiculidae). Parte taxonomica. Publicaciones especiales del Instituto de Biologia. Vol. 2. Universidad Nacional Autonoma de Mexico, Mexico, 275 pp."]}

Eutrombicula gonzalezi Stekolnikov sp. nov. (Figs. 7–9) Diagnosis. SIF = 7BS-N-2-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/B/NNB; fSc: PL> AL> AM; Ip = 954–1003; fD = 2H-6-6-4-2(2-4)-(0–2); DS = 20–22; V = 10–12; NDV = 32. Inner prong of palpal claw significantly shorter that outer prong; scutal and dorsal idiosomal setae thin and sparsely covered with short appressed barbs; scutum nearly pentagonal, covered with puncta of different size, without transverse striae; sensilla (trichobothria) with ca. 7–9 branches in distal half; sensillary bases anterior to level of PL (PSB—P-PL = 4–11 µm). Standard measurements of type series given in Table 4. Description (larva) (based on holotype and 9 paratypes). IDIOSOMA (Figs. 7, 8A–F). Eyes 2 + 2; 20–22 dorsal idiosomal setae, including one pair of humeral setae, thin and sparsely covered with short appressed barbs, by 6 setae in 1 st and 2 nd (D) posthumeral rows, 4 setae in 3 rd (E) row, 2 setae in 4 th (F) row (setae in rows E and F can be also counted as 2-4); 2 more caudal setae can be counted among ventral setae; arrangement of setae in holotype 2H-6-6-4-2; 4 sternal setae; 10–12 ventral setae; NDV = 32 (no variation recorded). GNATHOSOMA (Fig. 8G, H). Cheliceral blade with tricuspid cap; cheliceral base with small dense puncta basally and more sparse puncta in distal part; gnathobase with small puncta, transverse striae, and 1 pair of branched gnathocoxal (tritorostral) setae; galeal (deutorostral) seta nude; palpal claw with 2 prongs, inner prong significantly shorter that outer prong; palpal femur with sparse puncta different by size, with branched seta; palpal genu with puncta and branched seta; dorsal and lateral palpal tibial setae nude, ventral palpal tibial seta branched; palpal tarsus with 7 branched setae, nude subterminala (ζ) and basal tarsala (ω). SCUTUM (Figs. 7C, 8A). Nearly pentagonal, covered with puncta of two types: 1) of moderate size, sparsely covering all scutal surface, 2) very small, forming dense groups in some parts of scutum; anterolateral shoulders small, anterior scutal margin lightly sinuous, lateral margins concave, posterior margin angulate; AM at level of AL, sensillary (trichobothrial) bases anterior to level of PL (PSB —P-PL = 4–11 µm); all scutal setae uniform with dorsal idiosomal setae; PL> AL> AM; flagelliform sensilla (trichobothria), with ca. 7–9 branches in distal half. LEGS (Fig. 9). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 3 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) proximal to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 14B, mastitarsala. Hosts. Myiophobus fasciatus, Turdus albicollis, Schiffornis virescens. Type material. Holotype larva (ZIN 8940) ex T. albicollis No PG 212, PARAGUAY: San Rafael National Park, Kanguery, 22 August 2012. Paratypes: 9 larvae (ZIN 8941–8949) with same data; 3 larvae (ZIN 8905–8907) ex S. virescens No PG 52 and 2 larvae (ZIN 8983, 8984) ex M. fasciatus No PG 248, Kanguery, 19 and 24 August 2012, respectively. Etymology. The species is named after Dr. Daniel González-Acuña (1963–2020), a Chilean parasitologist who made an important contribution to the chigger studies in South America. Differential diagnosis. Eutrombicula gonzalezi sp. nov. is similar to E. picunche Stekolnikov and González-Acuña, 2015 described from a single specimen (holotype) collected from a lizard in Chile, in having the shape of scutum and setation of palps and idiosoma similar, but differs from it by the relatively shorter inner prong of palpal claw, the presence of nude mastitarsala (vs. ciliated), larger scutum (PW = 88–100 vs. 72, SD = 56–64 vs. 53) with more prominent posterior margin (PSB = 29–34 vs. 22, P-PL = 21–29 vs. 17), longer setae (e.g. AM = 30–36, AL = 35–41, PL = 56–63, and V min = 38–43 vs. 23, 23, 35, and 29, respectively), and much longer legs (Ip = 954–1003 vs. 720)., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 514-517, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054

Neoschoengastia americana (Hirst, 1921) (Fig. 26) Schöngastia americana Hirst, 1921: 37. Neoschoengastia americana: Wharton & Hardcastle 1946: 289; Domrow & Nadchatram 1960: 191, figs. 36–42; Domrow & Lester 1985: 32, figs. 116, 124–126; Stekolnikov 2021: 89. Diagnosis. SIF = 7BS-B-3-2111.0000; fsp = 7.7.7; fCx = 1.1.3; fSt = 2.2; fPp = B/B/BNB; fSc: PL> AL> AM; Ip = 1244–1264; fD = 2H-8(9)-6-6(4)-4(2)-6-2; DS = 32–33; V = 28–30; NDV = 61–62. Idiosoma pear-shaped in engorged specimens; scutum subquadrate, with anterolateral shoulders and bilobate posterior margin, with few large puncta and circular striae around sensillary bases; AM anterior to level of AL; AM and AL densely covered with very long barbs; sensilla capitate, covered with setules, far anterior to level of PL (closer to level of AL than to level of PL); leg claws and empodia with onychotriches; parasubterminala (z) branched; tibialae II situated in distal part of segment. Standard measurements of two examined specimens (ZIN 8995 and ZIN 9011): AW 49, 49; PW 70, 72; SB 33, 28; ASB 18, 22; PSB 23, 29; SD 41, 50; P-PL 7, 11; AP 28, 31;AM 37, -; AL 47, -; PL 54, 58; S 22, -; H 50, 54; D min 29, 32; D max 50, 52; V min 25, 32; V max 32, 40; pa 421, 430; pm 380, 387; pp 443, 446; Ip 1244, 1264; TaIIIL 131, 131; TaIIIW 15, 15; DS 32, 33; V 30, 28; NDV 62, 61. Distribution. Australia, Malaysia, Taiwan, USA (California, South Carolina, Texas, Virginia) (Wharton & Hardcastle 1946; Stekolnikov 2021), and Paraguay (this study). Hosts. Various species of birds (McClure 1987; Stekolnikov 2021) and, occasionally, Sylvilagus sp. (Lagomorpha: Leporidae) (Domrow & Lester 1985). Material examined. 18 larvae (ZIN 8995–9003, 9005–9013) ex Hemitriccus margaritaceiventer Nos PG 261 and PG323, Coryphospingus cucullatus No PG 267, Coryphistera alaudina No PG 303, PARAGUAY: Teniente Agripino Enciso National Park, 31 August–3 September 2012. Remarks. Records from Australia, Malaysia, and Taiwan can actually pertain to other species of Neoschoengastia Ewing, 1929, for example, Neoschoengastia solomonis Wharton and Hardcastle, 1946, which was described as a subspecies of N. americana (Stekolnikov 2021)., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 543-544, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054, {"references":["Wharton, G. W. & Hardcastle, A. B. (1946) The genus Neoschongastia (Acarinida: Trombiculidae) in the Western Pacific area. Journal of Parasitology, 32, 286 - 322. https: // doi. org / 10.2307 / 3272682","Domrow, R. & Nadchatram, M. (1960) Malaysian Parasites - XLIII. Neoschoengastia in Malaya (Acarina, Trombiculidae). Studies from the Institute for Medical Research, Federation of Malaya, 29, 185 - 193.","Domrow, R. & Lester, L. N. (1985) Chiggers of Australia (Acari: Trombiculidae): an annotated checklist, keys and bibliography. Australian Journal of Zoology, Supplementary Series, 114, 1 - 111. https: // doi. org / 10.1071 / AJZS 114","McClure, H. E. (1987) The occurrence of chiggers, Neoschoengastia americana, among chaparral birds of southern California. North American Bird Bander, 12, 148 - 150."]}

Eutrombicula goeldii (Oudemans, 1910) (Figs. 10–12) Microthrombidium göldii Oudemans, 1910: 84; 1912: 13, fig. B. Trombicula (Eutrombicula) göldii: Jenkins 1949: 312, figs. 8, 9 (larva), 11 (adult). Eutrombicula göldii: Fuller 1952: 136. Eutrombicula goeldii: Brennan & Reed 1974: 710, figs. 4, 19; Loomis & Wrenn 1984: 158; Jacinavicius et al. 2018: 9. Diagnosis. SIF = 7BS-N-2-3111.1000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/N/NNN; fSc: PL> AM> AL; Ip = 934–1001; fD = 2H-6-6-4-2-2(2-4-2); DS = 20–22; V = 12–14; NDV = 34. Inner prong of palpal claw twice as short as outer prong and located medially on the latter; scutal and dorsal idiosomal setae thin, sparsely covered with small barbs; scutum nearly trapezoidal, with small dense puncta and transverse striae; sensilla (trichobothria) with ca. 8 branches in distal two thirds; sensillary bases far anterior to level of PL (PSB—P-PL = 10–15 µm); leg coxae with longitudinal striae. Standard measurements of examined specimens given in Table 5. Note. P-values for estimation of differences between Eutrombicula species are given by the Mann-Whitney test. Redescription (larva) (based on 7 specimens). IDIOSOMA (Figs. 10, 11A–E). Eyes 2 + 2; 20–22 dorsal idiosomal setae, including one pair of humeral setae, sparsely covered with small barbs, by 6 setae in 1 st and 2 nd (D) posthumeral rows, 4 setae in 3 rd (E) row, by 2 setae in 4 th (F) and 5 th (H) rows (setae in last 3 rows can be also counted as 2-4-2); 4 sternal setae; 12–14 ventral setae; NDV = 34. GNATHOSOMA (Fig. 11F, G). Cheliceral blade with tricuspid cap; cheliceral base with small dense puncta basally and more sparse puncta in distal part; gnathobase with small dense puncta, transverse striae, and 1 pair of branched gnathocoxal (tritorostral) setae; galeal (deutorostral) seta nude; palpal claw with 2 prongs, inner prong twice as short as outer prong and located medially on the latter; palpal femur with numerous puncta and branched seta; palpal genu with puncta and nude seta; palpal tibia with three nude setae; palpal tarsus with 7 branched setae, nude subterminala (ζ) and basal tarsala (ω). SCUTUM (Figs. 10C, 11A). Nearly trapezoidal, densely covered with small puncta, with transverse striae, without anterolateral shoulders, anterior scutal margin almost straight, lateral margins slightly concave, posterior margin almost straight in middle part and obliquely cut at edges; AM at level of AL, sensillary (trichobothrial) bases far anterior to level of PL (PSB—P-PL = 10–15 µm); all scutal setae uniform with dorsal idiosomal setae; PL> AM> AL; flagelliform sensilla (trichobothria), with ca. 8 branches in distal half. LEGS (Fig. 12). All 7-segmented, with 1 pair of claws and claw-like empodium. Leg I: coxa with 1 branched seta (1B); trochanter 1B; basifemur 1B; telofemur 5B; genu 4B, 3 genualae (σ), microgenuala (κ); tibia 8B, 2 tibialae (φ), microtibiala (κ); tarsus 22B, tarsala (ω), microtarsala (ε) distal to tarsala, subterminala (ζ), parasubterminala (z), pretarsala (ζ). Leg II: coxa 1B; trochanter 1B; basifemur 2B; telofemur 4B; genu 3B, genuala; tibia 6B, 2 tibialae (φ); tarsus 16B, tarsala (ω), microtarsala (ε) proximal to tarsala, pretarsala (ζ). Leg III: coxa 1B; trochanter 1B; basifemur 2B; telofemur 3B; genu 3B, genuala; tibia 6B, tibiala; tarsus 14B, mastitarsala. Distribution. Brazil, Bolivia, Colombia, Costa Rica, Dominica, Panama, Surinam, Trinidad, Venezuela (Jacinavicius et al. 2018; Bassini-Silva et al. 2021), and Peru (this study). Hosts. Various species of mammals and birds, three species of lizards and one snake (Bassini-Silva et al. 2021). Material examined. Total 29 larvae (interval of numbers: ZIN 8821–8894) ex Leptotila rufaxilla Nos IQ 20, IQ33, IQ37, PERU: Iquitos, National Reserve Allpahuayo Mishana, 15–16 August 2011., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 518-521, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054, {"references":["Oudemans, A. C. (1910) Acarologische Aanteekeningen XXXIII. Entomologische Berichten, Amsterdam, 3, 83 - 90.","Oudemans, A. C. (1912) Die bis jetzt bekannten Larven von Thrombidiidae und Erythraeidae. Zoologische Jahrbucher, Supplement 14, 1 - 230.","Jenkins, D. W. (1949) Trombiculid mites affecting man. IV. Revision of Eutrombicula in the American Hemisphere. Annals of the Entomological Society of America, 42, 289 - 318 https: // doi. org / 10.1093 / aesa / 42.3.289","Fuller, H. S. (1952) The mite-larvae of the family Trombiculidae in the Oudemans collection: taxonomy and medical importance. Zoologische Verhandelingen, 18, 1 - 261.","Brennan, J. M. & Reed, J. T. (1974) The genus Eutrombicula in Venezuela (Acarina: Trombiculidae). Journal of Parasitology, 60, 699 - 711. https: // doi. org / 10.2307 / 3278741","Loomis, R. B. & Wrenn, W. J. (1984) Systematics of the pest chigger genus Eutrombicula (Acari: Trombiculidae). In: Griffiths, D. A. & Bowman, C. E. (Eds.), Acarology VI. 1. Ellis Horwood Limited, Chichester, pp. 152 - 159.","Jacinavicius, F. C., Bassini-Silva, R., Mendoza-Roldan, J. A., Pepato, A. R., Ochoa, R., Welbourn, C., Barros-Battesti, D. M. (2018) A checklist of chiggers from Brazil, including new records (Acari: Trombidiformes: Trombiculidae and Leeuwenhoekiidae). ZooKeys, 743, 1 - 41. https: // doi. org / 10.3897 / zookeys. 743.22675","Bassini-Silva, R., Takatsu, J. C., Bermudez, S. E., Miranda, R. J., Welbourn, C., Ochoa, R., Barros-Battesti, D. M. & Jacinavicius, F. C. (2021) A checklist of chiggers (Trombidiformes: Trombiculidae and Leeuwenhoekiidae) of Panama. Acarologia, 61, 763 - 789. https: // doi. org / 10.24349 / ach 8 - Eqk 3"]}

Parasecia fundata (Brennan, 1969) (Fig. 19) Fonsecia (Parasecia) fundata Brennan, 1969: 664, fig. 3. Parasecia fundata: Goff 1992b: 967; Stekolnikov et al. 2007: 66; Bassini-Silva et al. 2020: 160, figs. 7, 8. Diagnosis. SIF = 7BS-N-3-3111.0000; fsp = 7.7.7; fCx = 1.1.1; fSt = 2.2; fPp = B/B/NNB(f); fSc: PL> AM> AL; Ip = 578–675; fD = 2H-6-6-2(4)-(2–6)-(2–4); DS = 22–26; V = 17–20; NDV = 39–46. Scutum nearly trapezoidal, with anterolateral shoulders and with few puncta; sensilla with ca. 8–12 branches in distal half; sensillary bases slightly anterior to level of PL (PSB—P-PL = 4–6 µm). Standard measurements of one examined specimen from Peru (ZIN 8873) and two from Costa Rica (ZIN 7069 and ZIN 7083) given in Table 8. Distribution. Brazil, Costa Rica (Stekolnikov et al. 2007), and Peru (this study). Hosts. Mammalia: Caluromys philander (L.), Didelphis marsupialis L. (Didelphimorphia: Didelphidae), Proechimys guyannensis (É. Geoffroy) (Rodentia: Echimyidae); Aves: Glyphorynchus spirurus, Xiphorhynchus susurrans (Jardine) (Passeriformes: Furnariidae), Melanerpes pucherani (Malherbe) (Piciformes: Picidae) (Stekolnikov et al. 2007). New host record: Dendrocincla fuliginosa. Localization on hosts. Ear (D. fuliginosa No IQ 9). Material examined. One larva (ZIN 8859) ex. D. fuliginosa No IQ 9 and 1 larva (ZIN 8873) ex G. spirurus No IQ 32, PERU: Iquitos, National Reserve Allpahuayo Mishana, 15 and 16 August 2011; 2 larvae (ZIN 7069, 7083) ex G. spirurus Nos B 57 and HC321, COSTA RICA: Barbilla National Park, 1 July 2004 (Stekolnikov et al. 2007). Remarks. The ventral palpal tibial seta is branched or forked in our material that is in accordance with the original description, but Bassini-Silva et al. (2020) drew and described this seta as nude. Probably, this character is variable in P. fundata. Brennan (1969) also designated the state of lateral palpal tibial seta as Nb, i.e. nude or having one small cilium, whereas Bassini-Silva et al. (2020) stated that this seta (erroneously designated as “lateral seta of the palptarsus” in the text) is nude in the type series. Thus, P. fundata cannot be reliably discriminated by the setation of palps from Parasecia manueli (Brennan and Jones, 1960). The latter usually has fPp = B/B/NNN, but the ventral palpal tibial setae can be forked or branched (Brennan & Jones 1960; Daniel & Stekolnikov 2003a). In their key to species, Bassini-Silva et al. (2020) discriminated P. fundata and P. manueli by the number of setae in 3 rd posthumeral row (E) (6 setae in P. fundata vs. 2 setae in P. manueli) and by NDV or “total opisthosomal setae” (48–54 in P. fundata vs. 38 in P. manueli). However, according to our data, the number of setae in 3 rd row can be 2 or 4 both in P. fundata (our material from Costa Rica and Peru) and P. manueli from Cuba (Daniel & Stekolnikov 2003a). Note also that according to the original description of P. manueli, arrangement of dorsal idiosomal setae is variable in this species and 3 rd row includes 4 or 6 setae (Brennan & Jones 1960). Our samples of P. fundata and P. manueli also do not differ by NDV (Table 8). In fact, the characters allowing a precise discrimination of these two species are measurements. Parasecia fundata has definitely smaller scutum (AW–SD and AP in Table 8), shorter setae (AM–V max in Table 8), shorter and thinner legs (pa–TaIIIW in Table 8) than P. manueli. Unfortunately, the cited revision of the genus Parasecia (Bassini-Silva et al. 2020) includes measurements for only one new species that significantly lowers usefulness of this work., Published as part of Stekolnikov, Alexandr A., Capek, Miroslav & Literák, Ivan, 2022, New species and records of chiggers (Acariformes: Trombiculidae) from birds of the Neotropics, pp. 501-552 in Zootaxa 5141 (6) on pages 529-530, DOI: 10.11646/zootaxa.5141.6.1, http://zenodo.org/record/6598054, {"references":["Brennan, J. M. (1969) Three new species of subgenus Parasecia Loomis (genus Fonsecia) from northeastern Brazil and a key to the included species (Acarina: Trombiculidae). Journal of Parasitology, 55, 662 - 666. https: // doi. org / 10.2307 / 3277312","Goff, M. L. (1992 b) The genus Parasecia (Acari: Trombiculidae), with the description of a new species from Mexico. Journal of Medical Entomology, 29, 965 - 967 https: // doi. org / 10.1093 / jmedent / 29.6.965","Stekolnikov, A. A., Literak, I., Capek, M. & Havlicek, M. (2007) Chigger mites (Acari: Trombiculidae) from wild birds in Costa Rica, with a description of three new species. Folia Parasitologica, 54, 59 - 67. https: // doi. org / 10.14411 / fp. 2007.008","Bassini-Silva, R., Jacinavicius, F. C., Oliveira, M., Peinado, L. C., Faxina, C., Moreira-Lima, L., Welbourn, C., Ochoa, R., Fisch- er, E., Hingst-Zaher, E., Famadas, K. M., Faccini, J. L. H. & Barros-Battesti, D. M. (2020) A Revision of Parasecia (Trombidiformes: Trombiculidae) with a description of a new species, a new genus and a key to species. Journal of Medical Entomology, 58, 146 - 181. https: // doi. org / 10.1093 / jme / tjaa 149","Daniel, M. & Stekolnikov, A. A. (2003 a) Chigger mites (Acari: Trombiculidae) new to the fauna of Cuba, with the description of two new species. Folia Parasitologica, 50 (2), 143 - 150. https: // doi. org / 10.14411 / fp. 2003.025","Brennan, J. M. & Jones, E. K. (1960) Chiggers of Trinidad, BWI (Acarina: Trombiculidae). Acarologia, 2, 493 - 540."]}