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Contrasting results in visual and auditory working memory studies suggest that the mechanisms of association between location and identity of stimuli depend on the sensory modality of the input. In this auditory study, we tested whether the association of two features both encoded in the “what” stream is different from the association between a “what” and a “where” feature. In an old–new recognition task, blindfolded participants were presented with sequences of sounds varying in timbre, pitch and location. They were required to judge if either the timbre, pitch or location of a single-probe stimulus was identical or different to the timbre, pitch or location of one of the sounds of the previous sequence. Only variations in one of the three features were relevant for the task, whereas the other two features could vary, with task-irrelevant changes. Results showed that task-irrelevant variations in the “what” features (either timbre or pitch) caused an impaired recognition of sound location and in the othe...